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<document id="E6E51D707F1EC69E8351D3F99068C129" ID-DOI="10.5281/zenodo.156322" ID-GBIF-Dataset="4778cd2f-9478-4315-86b5-2d9c7c7a2b31" ID-ISSN="1175­5326" ID-Zenodo-Dep="156322" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459633653077" checkinUser="plazi" docAuthor="Smith, Andrew B. T." docDate="2003" docId="03965606FFFC340F6E70C6F9F95AFB66" docLanguage="en" docName="zt00316.pdf" docOrigin="Zootaxa 316" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Phalangogonia hawksi Smith, 2003, new species" docType="treatment" docVersion="11" lastPageNumber="4" masterDocId="FFAF2E7EFFFC340C6F78C010FC70FFC0" masterDocTitle="A new species of Phalangogonia Burmeister (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini) from Costa Rica" masterLastPageNumber="4" masterPageNumber="1" pageNumber="1" updateTime="1698204162534" updateUser="plazi">
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<mods:title id="E48C8A2D29DE69DA8DD82315BE8C4908">A new species of Phalangogonia Burmeister (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini) from Costa Rica</mods:title>
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<mods:namePart id="927341DAF0029A3EA6B38F9C7CAD22B1">Smith, Andrew B. T.</mods:namePart>
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<mods:date id="4D90B5F1EAE24DD7A64E9E59BE682016">2003</mods:date>
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<treatment id="03965606FFFC340F6E70C6F9F95AFB66" ID-DOI="http://doi.org/10.5281/zenodo.6276671" ID-GBIF-Taxon="119332226" ID-Zenodo-Dep="6276671" LSID="urn:lsid:zoobank.org:act:8F74FE41-6BBD-458C-8C0C-BA27907BF9D0" httpUri="http://treatment.plazi.org/id/03965606FFFC340F6E70C6F9F95AFB66" lastPageId="3" lastPageNumber="4" pageId="0" pageNumber="1">
<subSubSection id="C325B49BFFFC340C6E70C6F9FF4DF8C3" box="[264,829,1769,1795]" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="8B80E710FFFC340C6E70C6F9FF4DF8C3" blockId="0.[264,829,1769,1795]" box="[264,829,1769,1795]" pageId="0" pageNumber="1">
<heading id="D0C8507CFFFC340C6E70C6F9FF4DF8C3" bold="true" box="[264,829,1769,1795]" fontSize="11" level="1" pageId="0" pageNumber="1" reason="1">
<emphasis id="B94B3B02FFFC340C6E70C6F9FF4DF8C3" bold="true" box="[264,829,1769,1795]" pageId="0" pageNumber="1">
<taxonomicName id="4C3F9C93FFFC340C6E70C6F9FE6DF8C3" box="[264,541,1769,1795]" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hawksi" status="sp. nov.">
<emphasis id="B94B3B02FFFC340C6E70C6F9FE6DF8C3" bold="true" box="[264,541,1769,1795]" italics="true" pageId="0" pageNumber="1">Phalangogonia hawksi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2788679FFFC340C6D5DC6F9FEC5F8C3" box="[549,693,1769,1795]" pageId="0" pageNumber="1" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="1304FB95FFFC340C6DBDC6F9FF45F8C3" box="[709,821,1769,1795]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 3" captionStartId-0="1.[264,368,1446,1470]" captionStartId-1="1.[264,382,1800,1824]" captionTargetBox-1="[269,1300,1580,1786]" captionTargetId-1="figure@1.[782,1312,1588,1791]" captionTargetPageId-1="1" captionText-0="FIGURE 1. Habitus of male Phalangogonia hawksi." captionText-1="FIGURES 2 3. Male genitalia of Phalangogonia hawksi. 2. lateral view; 3. dorsal view." httpUri-1="https://zenodo.org/record/168786/files/figure.png" pageId="0" pageNumber="1">Figs. 13</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C325B49BFFFC340D6E70C739FDE0FE6E" lastPageId="1" lastPageNumber="2" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="8B80E710FFFC340D6E70C739FDE0FE6E" blockId="0.[264,1325,1833,1899]" lastBlockId="1.[264,1323,284,430]" lastPageId="1" lastPageNumber="2" pageId="0" pageNumber="1">
<emphasis id="B94B3B02FFFC340C6E70C739FDB6F883" bold="true" box="[264,454,1833,1859]" pageId="0" pageNumber="1">
<typeStatus id="548459B2FFFC340C6E70C739FD35F883" box="[264,325,1833,1859]" pageId="0" pageNumber="1">Type</typeStatus>
specimen
</emphasis>
. Male
<typeStatus id="548459B2FFFC340C6D4AC739FEE9F883" box="[562,665,1833,1859]" pageId="0" pageNumber="1" type="holotype">holotype</typeStatus>
deposited at the Instituto Nacional de Biodiversidad (
<collectionCode id="ED2E7FD5FFFC340C6E68C741FD2FF8AB" box="[272,351,1873,1899]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="0" pageNumber="1">INBio</collectionCode>
), Santo
<collectingCountry id="F328A780FFFC340C6EB2C741FE49F8AB" box="[458,569,1873,1899]" name="Dominican Republic" pageId="0" pageNumber="1">Domingo</collectingCountry>
de Heredia,
<collectingCountry id="F328A780FFFC340C6D99C741FF1BF8AB" box="[737,875,1873,1899]" name="Costa Rica" pageId="0" pageNumber="1">Costa Rica</collectingCountry>
.
<typeStatus id="548459B2FFFC340C6CF1C741FF86F8AB" box="[905,1014,1873,1899]" pageId="0" pageNumber="1" type="holotype">Holotype</typeStatus>
labeled “
<collectingCountry id="F328A780FFFC340C6B17C741F959F8AB" box="[1135,1321,1873,1899]" name="Costa Rica" pageId="0" pageNumber="1">COSTA RICA</collectingCountry>
: PUNTARENAS Rancho Quemado,
<quantity id="4CC74AF5FFFD340D6DCAC10CFE8DFEF6" box="[690,765,284,310]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="2" unit="m" value="300.0">300 m</quantity>
<date id="FF81C1D0FFFD340D6C7CC10CFFD2FEF6" box="[772,930,284,310]" pageId="1" pageNumber="2" value="2001-05-25">25 May 2001</date>
M. Posla” (typeface) and “PHA­ LANGOGONIA
<taxonomicName id="4C3F9C93FFFD340D6EA2C154FEC5FE9E" authority="Smith" authorityName="Smith" box="[474,693,324,350]" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="4" phylum="Arthropoda" rank="species" species="hawksi">
HAWKSI
<collectionCode id="ED2E7FD5FFFD340D6D22C154FEC5FE9E" box="[602,693,324,350]" pageId="1" pageNumber="2">SMITH</collectionCode>
</taxonomicName>
ɗ
<typeStatus id="548459B2FFFD340D6D9BC154FFF1FE9E" box="[739,897,324,350]" pageId="1" pageNumber="2" type="holotype">HOLOTYPE</typeStatus>
” (handwriting and typeface, black border).
<typeStatus id="548459B2FFFD340D6E0EC17CFDC0FE46" box="[374,432,364,390]" pageId="1" pageNumber="2">Type</typeStatus>
locality: Rancho Quemado (
<geoCoordinate id="EE0B81D7FFFD340D6C7FC17CFF2DFE46" box="[775,861,364,390]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="8.666667">8°40'N</geoCoordinate>
,
<geoCoordinate id="EE0B81D7FFFD340D6C12C17CFFA4FE46" box="[874,980,364,390]" direction="west" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="-83.566666">83°34'W</geoCoordinate>
; Osa Peninsula), Puntarenas,
<collectingCountry id="F328A780FFFD340D6E70C184FDFCFE6E" box="[264,396,404,430]" name="Costa Rica" pageId="1" pageNumber="2">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF40B798FFFD340D6E70C5B6FF35FA7E" box="[264,837,1446,1470]" pageId="1" pageNumber="2">
<paragraph id="8B80E710FFFD340D6E70C5B6FF35FA7E" blockId="1.[264,837,1446,1470]" box="[264,837,1446,1470]" pageId="1" pageNumber="2">
<emphasis id="B94B3B02FFFD340D6E70C5B6FDFAFA7E" bold="true" box="[264,394,1446,1470]" pageId="1" pageNumber="2">FIGURE 1.</emphasis>
Habitus of male
<taxonomicName id="4C3F9C93FFFD340D6D31C5B7FF4EFA7E" box="[585,830,1447,1470]" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="hawksi">
<emphasis id="B94B3B02FFFD340D6D31C5B7FF4EFA7E" box="[585,830,1447,1470]" italics="true" pageId="1" pageNumber="2">Phalangogonia hawksi</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF40B798FFFD340D6E70C718F8B2F8E0" box="[264,1218,1800,1824]" httpUri="https://zenodo.org/record/168786/files/figure.png" pageId="1" pageNumber="2" targetBox="[269,1300,1580,1786]" targetPageId="1">
<paragraph id="8B80E710FFFD340D6E70C718F8B2F8E0" blockId="1.[264,1218,1800,1824]" box="[264,1218,1800,1824]" pageId="1" pageNumber="2">
<emphasis id="B94B3B02FFFD340D6E70C718FDC4F8E0" bold="true" box="[264,436,1800,1824]" pageId="1" pageNumber="2">FIGURES 23.</emphasis>
Male genitalia of
<emphasis id="B94B3B02FFFD340D6D06C719FF0BF8E0" box="[638,891,1801,1824]" italics="true" pageId="1" pageNumber="2">
<taxonomicName id="4C3F9C93FFFD340D6D06C719FF07F8E0" box="[638,887,1801,1824]" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="hawksi">Phalangogonia hawksi</taxonomicName>
.
</emphasis>
2. lateral view; 3. dorsal view.
</paragraph>
</caption>
<subSubSection id="C325B49BFFFE340E6E40C10BFEADFE3E" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8B80E710FFFE340E6E40C10BFEADFE3E" blockId="2.[264,1324,283,1470]" pageId="2" pageNumber="3">
<emphasis id="B94B3B02FFFE340E6E40C10BFDC0FEF5" bold="true" box="[312,432,283,309]" pageId="2" pageNumber="3">Diagnosis</emphasis>
. This species is distinguished from all other species in the genus
<taxonomicName id="4C3F9C93FFFE340E6B8AC10CFDFDFE9D" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B94B3B02FFFE340E6B8AC10CFDFDFE9D" italics="true" pageId="2" pageNumber="3">Phalangogonia</emphasis>
</taxonomicName>
by the following combination of characters: dorsal colour tan with prominent black pattern on head, pronotum, and elytra (
<figureCitation id="1304FB95FFFE340E6C60C17CFF2DFE46" box="[792,861,364,390]" captionStart="FIGURE 1" captionStartId="1.[264,368,1446,1470]" captionText="FIGURE 1. Habitus of male Phalangogonia hawksi." pageId="2" pageNumber="3">Fig. 1</figureCitation>
); frons glabrous; eye bulbous, protruding from side of head in dorsal view; pronotum moderately punctate; pygidial disc shagreened to granular, setose; mesometasternal process robust, not declivous with respect to venter; male genitalia as in
<figureCitation id="1304FB95FFFE340E6D11C1F4FEA7FE3E" box="[617,727,484,510]" captionStart="FIGURES 2 3" captionStartId="1.[264,382,1800,1824]" captionTargetBox="[269,1300,1580,1786]" captionTargetId="figure@1.[782,1312,1588,1791]" captionTargetPageId="1" captionText="FIGURES 2 3. Male genitalia of Phalangogonia hawksi. 2. lateral view; 3. dorsal view." httpUri="https://zenodo.org/record/168786/files/figure.png" pageId="2" pageNumber="3">Figs. 23</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C325B49BFFFE340E6E40C21BF811FADE" pageId="2" pageNumber="3" type="description">
<paragraph id="8B80E710FFFE340E6E40C21BF805FB36" blockId="2.[264,1324,283,1470]" pageId="2" pageNumber="3">
<emphasis id="B94B3B02FFFE340E6E40C21BFE2EFDE5" bold="true" box="[312,606,523,549]" pageId="2" pageNumber="3">
Description of
<typeStatus id="548459B2FFFE340E6E8BC21BFE2EFDE5" box="[499,606,523,549]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
</emphasis>
. Male. Length 21.0 mm, width 11.0 mm. Colour tan with prominent black pattern on head, pronotum, and elytra (
<figureCitation id="1304FB95FFFE340E6CE0C224FFADFD8E" box="[920,989,564,590]" captionStart="FIGURE 1" captionStartId="1.[264,368,1446,1470]" captionText="FIGURE 1. Habitus of male Phalangogonia hawksi." pageId="2" pageNumber="3">Fig. 1</figureCitation>
); ventral surface and pygidium black.
<emphasis id="B94B3B02FFFE340E6EEAC24CFDABFDB5" box="[402,475,604,629]" italics="true" pageId="2" pageNumber="3">Head:</emphasis>
Dorsal surface densely punctate. Clypeus setose, weakly rounded, apex reflexed. Frontoclypeal suture complete, straight. Eye large, bulbous, length in lateral view 0.3 x head length in lateral view. Labrum with apex vertically produced with respect to clypeus, produced at middle, triangular. Mentum setose, apex strongly reflexed into oral cavity. Antenna with 10 antennomeres; club length greater than length of antennomeres 26.
<emphasis id="B94B3B02FFFE340E6EE2C334FE68FCFD" box="[410,536,804,829]" italics="true" pageId="2" pageNumber="3">Pronotum:</emphasis>
Surface glabrous, moderately punctate (densely punctate near lateral border). Lateral border distinct; apical, basal borders indistinct.
<emphasis id="B94B3B02FFFE340E6B1DC35CF8B6FCA5" box="[1125,1222,844,869]" italics="true" pageId="2" pageNumber="3">Elytron:</emphasis>
Surface glabrous. Striae weakly defined, punctate. Epipleuron with ventral surface flat.
<emphasis id="B94B3B02FFFE340E6BC8C364F95BFC4D" box="[1200,1323,884,909]" italics="true" pageId="2" pageNumber="3">Pygidium:</emphasis>
Surface shagreened, setose; setae long, cream­coloured.
<emphasis id="B94B3B02FFFE340E6CC1C38CF87BFC75" box="[953,1035,924,949]" italics="true" pageId="2" pageNumber="3">Ve nt er:</emphasis>
Thorax glabrous medially, setose laterally; setae long, yellowish­brown. Mesometasternal process projecting parallel to body, apex adjacent to procoxae. Abdominal sternites sparsely setose.
<emphasis id="B94B3B02FFFE340E6B91C3FCF95BFBC5" box="[1257,1323,1004,1029]" italics="true" pageId="2" pageNumber="3">Legs:</emphasis>
Protibia with 3 teeth. Mesotibia and metatibia with medial carina. Tarsomeres 14 wider than long, cup­shaped. Mesotarsomere and metatarsomere 13 with ventral pad of thick, yellowish­brown setae. Tarsomere 5 elongate, with weak ventrobasal tooth. Tarsal claws with modified claw thickened when compared to other claw, apex bifurcate.
<emphasis id="B94B3B02FFFE340E6BECC49CFD40FB0D" italics="true" pageId="2" pageNumber="3">Male genitalia:</emphasis>
<figureCitation id="1304FB95FFFE340E6E42C4A4FDDAFB0E" box="[314,426,1204,1230]" captionStart="FIGURES 2 3" captionStartId="1.[264,382,1800,1824]" captionTargetBox="[269,1300,1580,1786]" captionTargetId="figure@1.[782,1312,1588,1791]" captionTargetPageId="1" captionText="FIGURES 2 3. Male genitalia of Phalangogonia hawksi. 2. lateral view; 3. dorsal view." httpUri="https://zenodo.org/record/168786/files/figure.png" pageId="2" pageNumber="3">Figs. 23</figureCitation>
. Phallobase fused to parameres. Parameres fused together except at apex; dorsomedially without keel or swelling; apex constricted, curved ventrally.
</paragraph>
<paragraph id="8B80E710FFFE340E6E40C514F811FADE" blockId="2.[264,1324,283,1470]" box="[312,1121,1284,1310]" pageId="2" pageNumber="3">
<emphasis id="B94B3B02FFFE340E6E40C514FDDBFADE" bold="true" box="[312,427,1284,1310]" pageId="2" pageNumber="3">Variation</emphasis>
. Unknown. The
<typeStatus id="548459B2FFFE340E6DFAC514FE99FADE" box="[642,745,1284,1310]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
is the only specimen examined.
</paragraph>
</subSubSection>
<subSubSection id="C325B49BFFFE340E6E40C53CFED5FA56" pageId="2" pageNumber="3" type="etymology">
<paragraph id="8B80E710FFFE340E6E40C53CFED5FA56" blockId="2.[264,1324,283,1470]" pageId="2" pageNumber="3">
<emphasis id="B94B3B02FFFE340E6E40C53CFDCDFA86" bold="true" box="[312,445,1324,1350]" pageId="2" pageNumber="3">Etymology</emphasis>
. I am very pleased to name this species after David C. Hawks (University of California­Riverside). It was during a visit to Riverside when I discovered the
<typeStatus id="548459B2FFFE340E6BBCC544F95AFAAE" box="[1220,1322,1364,1390]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
of this species in Dave's collection.
</paragraph>
</subSubSection>
<subSubSection id="C325B49BFFFE340E6E40C5B4F831FA7E" box="[312,1089,1444,1470]" pageId="2" pageNumber="3" type="distribution">
<paragraph id="8B80E710FFFE340E6E40C5B4F831FA7E" blockId="2.[264,1324,283,1470]" box="[312,1089,1444,1470]" pageId="2" pageNumber="3">
<emphasis id="B94B3B02FFFE340E6E40C5B4FDA1FA7E" bold="true" box="[312,465,1444,1470]" pageId="2" pageNumber="3">Distribution</emphasis>
. Known only from the Osa Peninsula of
<collectingCountry id="F328A780FFFE340E6CC1C5B4F84DFA7E" box="[953,1085,1444,1470]" name="Costa Rica" pageId="2" pageNumber="3">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C325B49BFFFE340F6E70C60CF95AFB66" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="key">
<paragraph id="8B80E710FFFE340E6E70C60CF8C7F9F6" blockId="2.[264,1207,1564,1590]" box="[264,1207,1564,1590]" pageId="2" pageNumber="3">
<emphasis id="B94B3B02FFFE340E6E70C60CFEB8F9F6" bold="true" box="[264,712,1564,1590]" pageId="2" pageNumber="3">
Key to the species of
<taxonomicName id="4C3F9C93FFFE340E6D76C60CFEB8F9F6" box="[526,712,1564,1590]" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B94B3B02FFFE340E6D76C60CFEB8F9F6" bold="true" box="[526,712,1564,1590]" italics="true" pageId="2" pageNumber="3">Phalangogonia</emphasis>
</taxonomicName>
</emphasis>
(modified from
<bibRefCitation id="EFAE9AE1FFFE340E6CF1C60CF8DEF9F6" author="Smith" box="[905,1198,1564,1590]" pageId="2" pageNumber="4" refString="Smith, A. B. T. &amp; Moron, M. A. (2003) Revision and phylogenetic analysis of the Central American endemic genus Phalangogonia Burmeister (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini). Systematic Entomology, 28, 323 - 338." type="journal article" year="2003">Smith and Morón [2003]</bibRefCitation>
)
</paragraph>
<key id="FFAEB021FFFE340F6E70C677F95AFB66" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<keyStep id="30CBFFB5FFFE340E6E70C677F95CF8E1" pageId="2" pageNumber="3">
<paragraph id="8B80E710FFFE340E6E70C677F95BF911" blockId="2.[264,1324,1639,1905]" pageId="2" pageNumber="3">
<keyLead id="30CE4425FFFE340E6E70C677F95BF911" pageId="2" pageNumber="3">1. Pygidial disc appearing smooth, without noticeable microsculpturing (some scattered punctures and setae present); mesometasternal process robust, declivous (sloping away) with respect to venter........................................................................................ 2</keyLead>
</paragraph>
<paragraph id="8B80E710FFFE340E6E70C6CFF95CF8E1" blockId="2.[264,1324,1639,1905]" pageId="2" pageNumber="3">
<keyLead id="30CE4425FFFE340E6E70C6CFF95CF8E1" pageId="2" pageNumber="3">­ Pygidial disc shagreened, rugose or granular; mesometasternal process weak to robust, apex parallel with respect to venter................................................................ 3</keyLead>
</paragraph>
</keyStep>
<paragraph id="8B80E710FFFE340E6E70C73FF95AF8B1" blockId="2.[264,1324,1639,1905]" pageId="2" pageNumber="3">
<keyStep id="30CBFFB5FFFE340E6E70C73FF95AF8B1" pageId="2" pageNumber="3">
<keyLead id="30CE4425FFFE340E6E70C73FF95AF8B1" pageId="2" pageNumber="3">2(1). Metatarsomeres 13 with well developed pad of ventral spines and setae; male parameres with distinct dorsomedial keel, apices with strong tooth; Puebla, Veracruz,</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B80E710FFFF340F6E3EC10CF95BFE16" blockId="3.[264,1326,284,1190]" pageId="3" pageNumber="4">
Oaxaca and Veracruz,
<collectingCountry id="F328A780FFFF340F6D35C10CFED6FEF6" box="[589,678,284,310]" name="Mexico" pageId="3" pageNumber="4">México</collectingCountry>
.......................................................
<taxonomicName id="4C3F9C93FFFF340F6B3BC10CF95AFEF6" authority="Bates" authorityName="Bates" box="[1091,1322,284,310]" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lacordairei">
<emphasis id="B94B3B02FFFF340F6B3BC10CF822FEF5" box="[1091,1106,284,309]" italics="true" pageId="3" pageNumber="4">P</emphasis>
.
<emphasis id="B94B3B02FFFF340F6B18C10CF892FEF5" box="[1120,1250,284,309]" italics="true" pageId="3" pageNumber="4">lacordairei</emphasis>
Bates
</taxonomicName>
­ Metatarsomeres 13 with weak pad of ventral spines and setae; male parameres with poorly defined dorsomedial keel, apices with weak tooth; eastern
<collectingCountry id="F328A780FFFF340F6BFEC17CF975FE46" box="[1158,1285,364,390]" name="Guatemala" pageId="3" pageNumber="4">Guatemala</collectingCountry>
to northwestern
<collectingCountry id="F328A780FFFF340F6E9EC184FE28FE6E" box="[486,600,404,430]" name="Honduras" pageId="3" pageNumber="4">Honduras</collectingCountry>
<emphasis id="B94B3B02FFFF340F6D27C184F80FFE6D" box="[607,1151,404,429]" italics="true" pageId="3" pageNumber="4">
........................................................................
<taxonomicName id="4C3F9C93FFFF340F6B15C184F80FFE6D" authority="Ohaus" authorityName="Ohaus" box="[1133,1151,404,429]" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="dispar">P</taxonomicName>
</emphasis>
.
<emphasis id="B94B3B02FFFF340F6BF5C184F8A8FE6D" box="[1165,1240,404,429]" italics="true" pageId="3" pageNumber="4">dispar</emphasis>
Ohaus
<keyLead id="30CE4425FFFF340F6E70C1ACF95BFE16" box="[264,1323,444,470]" pageId="3" pageNumber="4">
<keyStep id="30CBFFB5FFFF340F6E70C1ACF95BFE16" box="[264,1323,444,470]" pageId="3" pageNumber="4">3(1). Eyes small, almost flush with side of head in dorsal view.......................................... 4</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B80E710FFFF340F6E70C1F4F95BFE3E" blockId="3.[264,1326,284,1190]" box="[264,1323,484,510]" pageId="3" pageNumber="4">
<keyLead id="30CE4425FFFF340F6E70C1F4F95BFE3E" box="[264,1323,484,510]" pageId="3" pageNumber="4">­ Eyes bulbous, distinctly protruding from side of head in dorsal view ........................ 5</keyLead>
</paragraph>
<keyStep id="30CBFFB5FFFF340F6E70C21CF95BFD06" pageId="3" pageNumber="4">
<paragraph id="8B80E710FFFF340F6E70C21CF95AFDB6" blockId="3.[264,1326,284,1190]" pageId="3" pageNumber="4">
<keyLead id="30CE4425FFFF340F6E70C21CF95AFDB6" pageId="3" pageNumber="4">
4(3). Pronotum with sparse, scattered punctures; clypeus with surface mainly glabrous (sometimes with sparse, scattered setae); southern
<collectingCountry id="F328A780FFFF340F6CEFC224FF81FD8E" box="[919,1009,564,590]" name="Mexico" pageId="3" pageNumber="4">México</collectingCountry>
to
<collectingCountry id="F328A780FFFF340F6B6EC224F8E5FD8E" box="[1046,1173,564,590]" name="Guatemala" pageId="3" pageNumber="4">Guatemala</collectingCountry>
...................
<emphasis id="B94B3B02FFFF340F6E31C24CF83EFDB5" box="[329,1102,604,629]" italics="true" pageId="3" pageNumber="4">
.......................................................................................................
<taxonomicName id="4C3F9C93FFFF340F6B38C24CF83EFDB5" authority="Burmeister" authorityName="Burmeister" box="[1088,1102,604,629]" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="obesa">P</taxonomicName>
</emphasis>
.
<emphasis id="B94B3B02FFFF340F6B24C24CF8D1FDB5" box="[1116,1185,604,629]" italics="true" pageId="3" pageNumber="4">obesa</emphasis>
Burmeister
</keyLead>
</paragraph>
<paragraph id="8B80E710FFFF340F6E70C294F95BFD06" blockId="3.[264,1326,284,1190]" pageId="3" pageNumber="4">
<keyLead id="30CE4425FFFF340F6E70C294F95BFD06" pageId="3" pageNumber="4">
­ Pronotum with dense, sometimes confluent punctures; clypeus with surface uniformally setose;
<collectingCountry id="F328A780FFFF340F6E9FC2BCFE01FD06" box="[487,625,684,710]" name="El Salvador" pageId="3" pageNumber="4">El Salvador</collectingCountry>
<emphasis id="B94B3B02FFFF340F6D0DC2BCF81BFD05" box="[629,1131,684,709]" italics="true" pageId="3" pageNumber="4">
..................................................................
<taxonomicName id="4C3F9C93FFFF340F6B25C2BCF81BFD05" authority="Franz" authorityName="Franz" box="[1117,1131,684,709]" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="punctata">P</taxonomicName>
</emphasis>
.
<emphasis id="B94B3B02FFFF340F6B02C2BCF891FD05" box="[1146,1249,684,709]" italics="true" pageId="3" pageNumber="4">punctata</emphasis>
Franz
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CBFFB5FFFF340F6E70C2C4F95BFCD6" pageId="3" pageNumber="4">
<paragraph id="8B80E710FFFF340F6E70C2C4F95EFD2E" blockId="3.[264,1326,284,1190]" box="[264,1326,724,750]" pageId="3" pageNumber="4">
<keyLead id="30CE4425FFFF340F6E70C2C4F95EFD2E" box="[264,1326,724,750]" pageId="3" pageNumber="4">5(3). Mesometasternal process produced to or surpassing base of procoxae ..................... 6</keyLead>
</paragraph>
<paragraph id="8B80E710FFFF340F6E70C2ECF95BFCD6" blockId="3.[264,1326,284,1190]" box="[264,1323,764,790]" pageId="3" pageNumber="4">
<keyLead id="30CE4425FFFF340F6E70C2ECF95BFCD6" box="[264,1323,764,790]" pageId="3" pageNumber="4">­ Mesometasternal process not reaching base of procoxae........................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CBFFB5FFFF340F6E70C334F95CFC4E" pageId="3" pageNumber="4">
<paragraph id="8B80E710FFFF340F6E70C334F95AFCA6" blockId="3.[264,1326,284,1190]" pageId="3" pageNumber="4">
<keyLead id="30CE4425FFFF340F6E70C334F95AFCA6" pageId="3" pageNumber="4">
6(5). Dorsal colour tan with prominent black markings on head, pronotum, and elytra (
<figureCitation id="1304FB95FFFF340F6B83C334FD24FCA6" captionStart="FIGURE 1" captionStartId="1.[264,368,1446,1470]" captionText="FIGURE 1. Habitus of male Phalangogonia hawksi." pageId="3" pageNumber="4">Fig. 1</figureCitation>
); male parameres as in
<figureCitation id="1304FB95FFFF340F6D14C35CFEAAFCA6" box="[620,730,844,870]" captionStart="FIGURES 2 3" captionStartId="1.[264,382,1800,1824]" captionTargetBox="[269,1300,1580,1786]" captionTargetId="figure@1.[782,1312,1588,1791]" captionTargetPageId="1" captionText="FIGURES 2 3. Male genitalia of Phalangogonia hawksi. 2. lateral view; 3. dorsal view." httpUri="https://zenodo.org/record/168786/files/figure.png" pageId="3" pageNumber="4">Figs. 23</figureCitation>
; Osa Peninsula,
<collectingCountry id="F328A780FFFF340F6CE4C35CF86DFCA6" box="[924,1053,844,870]" name="Costa Rica" pageId="3" pageNumber="4">Costa Rica</collectingCountry>
<emphasis id="B94B3B02FFFF340F6B66C35CF80DFCA5" box="[1054,1149,844,869]" italics="true" pageId="3" pageNumber="4">
..........
<taxonomicName id="4C3F9C93FFFF340F6B16C35CF80DFCA5" authority="Smith" authorityName="Smith" box="[1134,1149,844,869]" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="hawksi">P</taxonomicName>
</emphasis>
.
<emphasis id="B94B3B02FFFF340F6BF4C35CF8AEFCA5" box="[1164,1246,844,869]" italics="true" pageId="3" pageNumber="4">hawksi</emphasis>
Smith
</keyLead>
</paragraph>
<paragraph id="8B80E710FFFF340F6E70C364F95CFC4E" blockId="3.[264,1326,284,1190]" box="[264,1324,884,910]" pageId="3" pageNumber="4">
<keyLead id="30CE4425FFFF340F6E70C364F95CFC4E" box="[264,1324,884,910]" pageId="3" pageNumber="4">­ Dorsal surface with uniform coloration....................................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CBFFB5FFFF340F6E70C38CF95BFBC6" pageId="3" pageNumber="4">
<paragraph id="8B80E710FFFF340F6E70C38CF95AFC76" blockId="3.[264,1326,284,1190]" box="[264,1322,924,950]" pageId="3" pageNumber="4">
<keyLead id="30CE4425FFFF340F6E70C38CF95AFC76" box="[264,1322,924,950]" pageId="3" pageNumber="4">
7(6). Dorsal colour mostly pale green; Oaxaca,
<collectingCountry id="F328A780FFFF340F6C4BC38CFFFDFC76" box="[819,909,924,950]" name="Mexico" pageId="3" pageNumber="4">México</collectingCountry>
....
<taxonomicName id="4C3F9C93FFFF340F6CC2C38CF95AFC76" authority="Smith and Moron" authorityName="Smith and Moron" box="[954,1322,924,950]" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jamesonae">
<emphasis id="B94B3B02FFFF340F6CC2C38CFFB8FC75" box="[954,968,924,949]" italics="true" pageId="3" pageNumber="4">P</emphasis>
.
<emphasis id="B94B3B02FFFF340F6CAFC38CF824FC75" box="[983,1108,924,949]" italics="true" pageId="3" pageNumber="4">jamesonae</emphasis>
Smith and Morón
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B80E710FFFF340F6E70C3D4F95BFBC6" blockId="3.[264,1326,284,1190]" pageId="3" pageNumber="4">
<keyLead id="30CE4425FFFF340F6E70C3D4F95BFBC6" pageId="3" pageNumber="4">
­ Dorsal colour tan or light to dark yellowish­brown (sometimes creamy white when alive);
<collectingCountry id="F328A780FFFF340F6EE0C3FCFE7AFBC6" box="[408,522,1004,1030]" name="Honduras" pageId="3" pageNumber="4">Honduras</collectingCountry>
to
<collectingCountry id="F328A780FFFF340F6D48C3FCFEFDFBC6" box="[560,653,1004,1030]" name="Panama" pageId="3" pageNumber="4">Panama</collectingCountry>
<emphasis id="B94B3B02FFFF340F6DEBC3FCF808FBC5" box="[659,1144,1004,1029]" italics="true" pageId="3" pageNumber="4">
................................................................
<taxonomicName id="4C3F9C93FFFF340F6B12C3FCF808FBC5" authority="Sharp" authorityName="Sharp" box="[1130,1144,1004,1029]" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sperata">P</taxonomicName>
</emphasis>
.
<emphasis id="B94B3B02FFFF340F6BFFC3FCF8AFFBC5" box="[1159,1247,1004,1029]" italics="true" pageId="3" pageNumber="4">sperata</emphasis>
Sharp
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CBFFB5FFFF340F6E70C404F95AFB66" pageId="3" pageNumber="4">
<paragraph id="8B80E710FFFF340F6E70C404F95AFB96" blockId="3.[264,1326,284,1190]" pageId="3" pageNumber="4">
<keyLead id="30CE4425FFFF340F6E70C404F95AFB96" pageId="3" pageNumber="4">
8(5). Head dorsally glabrous; pygidium brown to black, granulate; length
<quantity id="4CC74AF5FFFF340F6BE7C404F956FBEE" box="[1183,1318,1044,1070]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="3" pageNumber="4" unit="mm" value="21.0" valueMax="24.0" valueMin="18.0">1824 mm</quantity>
; Quezaltenango,
<collectingCountry id="F328A780FFFF340F6D7CC42CFEF2FB96" box="[516,642,1084,1110]" name="Guatemala" pageId="3" pageNumber="4">Guatemala</collectingCountry>
...................................................................
<taxonomicName id="4C3F9C93FFFF340F6B00C42CF95AFB96" authority="Bates" authorityName="Bates" box="[1144,1322,1084,1110]" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="parilis">
<emphasis id="B94B3B02FFFF340F6B00C42CF8F6FB95" box="[1144,1158,1084,1109]" italics="true" pageId="3" pageNumber="4">P</emphasis>
.
<emphasis id="B94B3B02FFFF340F6BEDC42CF892FB95" box="[1173,1250,1084,1109]" italics="true" pageId="3" pageNumber="4">parilis</emphasis>
Bates
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B80E710FFFF340F6E70C474F95AFB66" blockId="3.[264,1326,284,1190]" pageId="3" pageNumber="4">
<keyLead id="30CE4425FFFF340F6E70C474F95AFB66" pageId="3" pageNumber="4">
­ Head dorsally setose; pygidium tan, not granulate but with obvious microsculpturing; length
<quantity id="4CC74AF5FFFF340F6EEFC49CFE67FB66" box="[407,535,1164,1190]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.1" metricValueMin="2.4" pageId="3" pageNumber="4" unit="mm" value="27.5" valueMax="31.0" valueMin="24.0">2431 mm</quantity>
; Oaxaca to Chiapas,
<collectingCountry id="F328A780FFFF340F6C76C49CFF17FB66" box="[782,871,1164,1190]" name="Mexico" pageId="3" pageNumber="4">México</collectingCountry>
<emphasis id="B94B3B02FFFF340F6C1FC49CFF91FB65" box="[871,993,1164,1189]" italics="true" pageId="3" pageNumber="4">
..............
<taxonomicName id="4C3F9C93FFFF340F6CAAC49CFF91FB65" authority="Smith and Moron" authorityName="Smith and Moron" box="[978,993,1164,1189]" class="Insecta" family="Rutelidae" genus="Phalangogonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ratcliffei">P</taxonomicName>
</emphasis>
.
<emphasis id="B94B3B02FFFF340F6C97C49CF824FB65" box="[1007,1108,1164,1189]" italics="true" pageId="3" pageNumber="4">ratcliffei</emphasis>
Smith and Morón
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>