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<document id="0A545AAF7EB1BD936D6AFCA5584A685F" ID-DOI="10.5962/bhl.part.117802" ID-ISSN="0035-418X" ID-Zenodo-Dep="6311706" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645021050174" checkinUser="carolina" docAuthor="Cuccodoro, Giulio" docDate="2011" docId="03F7D871AF725D5AFF10AA8174E9DFA7" docLanguage="en" docName="RevSuisseZool.118.107-148.pdf.imf" docOrigin="Revue suisse de Zoologie 118 (1)" docStyle="DocumentStyle:0846B485C0C87F6EB7648DEEBF34B77F.2:RevSuisseZool.2011-2014.journal_article" docStyleId="0846B485C0C87F6EB7648DEEBF34B77F" docStyleName="RevSuisseZool.2011-2014.journal_article" docStyleVersion="2" docTitle="Megarthrus solitarius Sharp 1887" docType="treatment" docVersion="4" lastPageNumber="120" masterDocId="FFCEA009AF7E5D57FF98AE43766CDA37" masterDocTitle="Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae)" masterLastPageNumber="147" masterPageNumber="107" pageNumber="119" updateTime="1699296264638" updateUser="plazi">
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<mods:title id="FA527C07AD96040E9659BEB73F8A4D27">Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae)</mods:title>
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<mods:namePart id="DC11739C3D8AB095A829CDD9BF99B67E">Cuccodoro, Giulio</mods:namePart>
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<mods:title id="E839297471842DFC2341544E059D2BDE">Revue suisse de Zoologie</mods:title>
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<mods:date id="ABA966FAEE1BB39937EB479325D0BAD2">2011</mods:date>
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<mods:identifier id="382BD630CF8BF09F4EC703A5611CD49D" type="DOI">10.5962/bhl.part.117802</mods:identifier>
<mods:identifier id="F9F349941698EA598C3B7C2513D55C6F" type="ISSN">0035-418X</mods:identifier>
<mods:identifier id="C113E6FBA880986A9A20E569B6612A57" type="Zenodo-Dep">6311706</mods:identifier>
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<treatment id="03F7D871AF725D5AFF10AA8174E9DFA7" ID-DOI="http://doi.org/10.5281/zenodo.6311373" ID-Zenodo-Dep="6311373" LSID="urn:lsid:plazi:treatment:03F7D871AF725D5AFF10AA8174E9DFA7" httpUri="http://treatment.plazi.org/id/03F7D871AF725D5AFF10AA8174E9DFA7" lastPageId="13" lastPageNumber="120" pageId="12" pageNumber="119">
<subSubSection id="C3443AECAF725D5BFF10AA81724EDEED" box="[136,1058,1218,1242]" pageId="12" pageNumber="119" type="nomenclature">
<paragraph id="8BE16967AF725D5BFF10AA81724EDEED" blockId="12.[136,1058,1218,1242]" box="[136,1058,1218,1242]" pageId="12" pageNumber="119">
<heading id="D0A9DE0BAF725D5BFF10AA81724EDEED" box="[136,1058,1218,1242]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="119" reason="2">
<taxonomicName id="4C5E12E4AF725D5BFF10AA81746DDEED" ID-CoL="3YXCG" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" box="[136,513,1218,1242]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="119" phylum="Arthropoda" rank="species" species="solitarius">
<emphasis id="B92AB575AF725D5BFF10AA817714DEED" bold="true" box="[136,376,1218,1242]" italics="true" pageId="12" pageNumber="119">Megarthrus solitarius</emphasis>
<bibRefCitation id="EFCF1496AF725D5BFEE7AA80746DDEED" author="SHARP, D." box="[383,513,1219,1242]" pageId="12" pageNumber="119" pagination="673 - 824" refId="ref12839" refString="SHARP, D. 1887. Staphylinidae (pp. 673 - 824). In: Biologia Centrali-Americana. Insecta, Coleoptera 1 (2). Taylor &amp; Francis, London, XVI + 824 pp. + 18 pl." type="book chapter" year="1887">Sharp, 1887</bibRefCitation>
</taxonomicName>
Figs 3, 154-170
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF725D5BFF10AAAF7455DF35" box="[136,569,1259,1283]" pageId="12" pageNumber="119" type="reference_group">
<paragraph id="8BE16967AF725D5BFF10AAAF7455DF35" blockId="12.[136,1059,1259,1422]" box="[136,569,1259,1283]" pageId="12" pageNumber="119">
<treatmentCitationGroup id="AB4E4E49AF725D5BFF10AAAF7455DF35" box="[136,569,1259,1283]" pageId="12" pageNumber="119">
<treatmentCitation id="0AFF4F76AF725D5BFF10AAAF7459DF35" author="SHARP, D." box="[136,565,1259,1283]" page="743" pageId="12" pageNumber="119" year="1887">
<taxonomicName id="4C5E12E4AF725D5BFF10AAAF7459DF35" ID-CoL="3YXCG" authority="Sharp, 1887: 743" authorityName="Sharp" authorityPageNumber="743" authorityYear="1887" box="[136,565,1259,1283]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="119" phylum="Arthropoda" rank="species" species="solitarius">
<emphasis id="B92AB575AF725D5BFF10AAAF771EDF34" box="[136,370,1260,1283]" italics="true" pageId="12" pageNumber="119">Megarthrus solitarius</emphasis>
<bibRefCitation id="EFCF1496AF725D5BFEE1AAA87459DF35" author="SHARP, D." box="[377,565,1259,1282]" pageId="12" pageNumber="119" pagination="673 - 824" refId="ref12839" refString="SHARP, D. 1887. Staphylinidae (pp. 673 - 824). In: Biologia Centrali-Americana. Insecta, Coleoptera 1 (2). Taylor &amp; Francis, London, XVI + 824 pp. + 18 pl." type="book chapter" year="1887">Sharp, 1887: 743</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF725D5BFF4CAB4D75A9DFB9" pageId="12" pageNumber="119" type="materials_examined">
<paragraph id="8BE16967AF725D5BFF4CAB4D77C7DF61" blockId="12.[136,1059,1259,1422]" pageId="12" pageNumber="119">
<materialsCitation id="3B36633AAF725D5BFF4CAB4D77C7DF61" collectionCode="TYPE, MATERIAL, BMNH" collectorName="Type D. S. Purula &amp; B. C. A. Col." country="Guatemala" location="Guatemala" pageId="12" pageNumber="119" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectionCode id="ED4FF1A2AF725D5BFF4CAB4D7704DF15" box="[212,360,1294,1315]" pageId="12" pageNumber="119">TYPE MATERIAL</collectionCode>
:
<typeStatus id="54E5D7C5AF725D5BFEEFAB4D77BCDF14" box="[375,464,1294,1315]" pageId="12" pageNumber="119" type="holotype">Holotype</typeStatus>
(
<specimenCount id="9D58A2EEAF725D5BFE78AB4E7798DF14" box="[480,500,1293,1315]" pageId="12" pageNumber="119" type="female">♀</specimenCount>
, in
<collectionCode id="ED4FF1A2AF725D5BFD86AB4D7407DF14" box="[542,619,1294,1315]" pageId="12" pageNumber="119">BMNH</collectionCode>
): “
<taxonomicName id="4C5E12E4AF725D5BFD10AB4C7530DF13" authorityName="Sharp" authorityYear="1887" box="[648,860,1295,1316]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="119" phylum="Arthropoda" rank="species" species="solitarius">
<emphasis id="B92AB575AF725D5BFD10AB4C7530DF13" box="[648,860,1295,1316]" italics="true" pageId="12" pageNumber="119">Megarthrus solitarius</emphasis>
</taxonomicName>
.
<collectorName id="26AB0CB1AF725D5BFCF2AB4D7273DF14" box="[874,1055,1294,1315]" pageId="12" pageNumber="119">Type D. S. Purula</collectorName>
,
<collectingCountry id="F34929F7AF725D5BFF10AB6B769EDF0A" box="[136,242,1320,1341]" name="Guatemala" pageId="12" pageNumber="119">Guatemala</collectingCountry>
, Champion [Sharp’s handwritting] / Purula, Vera Paz, Champion /
<collectorName id="26AB0CB1AF725D5BFCE5AB6B759DDF0A" box="[893,1009,1320,1341]" pageId="12" pageNumber="119">B.C.A. Col.</collectorName>
1. 2.
<taxonomicName id="4C5E12E4AF725D5BFF10AB0277F1DF61" authority="Sharp" authorityName="Sharp" authorityYear="1887" box="[136,413,1345,1366]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="119" phylum="Arthropoda" rank="species" species="solitarius">
<emphasis id="B92AB575AF725D5BFF10AB027737DF61" box="[136,347,1345,1366]" italics="true" pageId="12" pageNumber="119">Megarthrus solitarius</emphasis>
Sharp
</taxonomicName>
”.
</materialsCitation>
</paragraph>
<paragraph id="8BE16967AF725D5BFF4CAB2375A9DFB9" blockId="12.[136,1059,1259,1422]" pageId="12" pageNumber="119">
ADDITIONAL
<collectionCode id="ED4FF1A2AF725D5BFED5AB2777C4DF43" box="[333,424,1380,1396]" pageId="12" pageNumber="119">MATERIAL</collectionCode>
:
<materialsCitation id="3B36633AAF725D5BFE2DAB2375ADDFB9" collectingDate="1993-06-29" collectionCode="MHNG, SEMC" collectorName="Ashe &amp; Brooks" country="Guatemala" elevation="1650" location="Bja Verapaz" pageId="12" pageNumber="119" specimenCount="2" specimenCount-female="2">
<collectingCountry id="F34929F7AF725D5BFE2DAB237421DF42" box="[437,589,1376,1397]" name="Guatemala" pageId="12" pageNumber="119">GUATEMALA</collectingCountry>
,
<location id="8E813FBCAF725D5BFDCFAB2374A2DF42" LSID="urn:lsid:plazi:treatment:03F7D871AF725D5AFF10AA8174E9DFA7:8E813FBCAF725D5BFDCFAB2374A2DF42" box="[599,718,1376,1397]" country="Guatemala" name="Bja Verapaz" pageId="12" pageNumber="119">Bja Verapaz</location>
,
<locationDeviation id="9A537045AF725D5BFD4FAB237505DF42" box="[727,873,1376,1397]" location="Purulha" pageId="12" pageNumber="119">
<quantity id="4CA6C482AF725D5BFD4FAB23756EDF42" box="[727,770,1376,1397]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="119" unit="km" value="8.0">8km</quantity>
S Purulhá
</locationDeviation>
,
<quantity id="4CA6C482AF725D5BFCEBAB2375D9DF42" box="[883,949,1376,1397]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="12" pageNumber="119" unit="m" value="1650.0">
<elevation id="00738E54AF725D5BFCEBAB2375D9DF42" box="[883,949,1376,1397]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="12" pageNumber="119" unit="m" value="1650.0">1650m</elevation>
</quantity>
<date id="FFE04FA7AF725D5BFC24AB23724FDF42" box="[956,1059,1376,1397]" pageId="12" pageNumber="119" value="1993-06-29">
<collectingDate id="EFA4B64FAF725D5BFC24AB23724FDF42" box="[956,1059,1376,1397]" pageId="12" pageNumber="119" value="1993-06-29">29.vi.1993</collectingDate>
</date>
(
<collectorName id="26AB0CB1AF725D5BFF17AB3A76AEDFB9" box="[143,194,1401,1422]" pageId="12" pageNumber="119">Ashe</collectorName>
&amp;
<collectorName id="26AB0CB1AF725D5BFF79AB3A7744DFB9" box="[225,296,1401,1422]" pageId="12" pageNumber="119">Brooks</collectorName>
) #149, ex
<taxonomicName id="4C5E12E4AF725D5BFE17AB3A7458DFB9" box="[399,564,1401,1422]" class="Magnoliopsida" family="Urticaceae" genus="Cecropia" kingdom="Plantae" order="Rosales" pageId="12" pageNumber="120" phylum="Tracheophyta" rank="species" species="treefall">
<emphasis id="B92AB575AF725D5BFE17AB3A778BDFB9" box="[399,487,1401,1422]" italics="true" pageId="12" pageNumber="119">Cecropia</emphasis>
treefall
</taxonomicName>
, 2 ƋƋ and
<specimenCount id="9D58A2EEAF725D5BFD31AB3A7488DFB9" box="[681,740,1400,1422]" pageId="12" pageNumber="119" type="female">2 ♀♀</specimenCount>
in
<collectionCode id="ED4FF1A2AF725D5BFC9BAB3A7520DFB9" box="[771,844,1401,1422]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="12" pageNumber="119" type="Museum">MHNG</collectionCode>
and
<collectionCode id="ED4FF1A2AF725D5BFCE5AB3A75ADDFB9" box="[893,961,1401,1422]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="119" type="Museum">SEMC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF725D5AFF4CABDC77A7DEAD" lastPageId="13" lastPageNumber="120" pageId="12" pageNumber="119" type="description">
<paragraph id="8BE16967AF725D5AFF4CABDC75BBDB0D" blockId="12.[136,1059,1439,1654]" lastBlockId="13.[151,1074,227,1178]" lastPageId="13" lastPageNumber="120" pageId="12" pageNumber="119">
DESCRIPTION: Habitus as in Fig. 3. Combined length of pronotum and elytra =
<quantity id="4CA6C482AF725D5BFF3FABFC7743DFE1" box="[167,303,1471,1494]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="12" pageNumber="119" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3-1.5 mm</quantity>
; maximal pronotal width =
<quantity id="4CA6C482AF725D5BFD1BABFC7567DFE1" box="[643,779,1471,1494]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="12" pageNumber="119" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9-1.1 mm</quantity>
. Body dark brown wih appendages slightly paler; antennomeres 10-11 paler than antennomeres 1-9. Dorsal pubescence fairly uniform, denser on pronotum than on elytra; frontal setae directed forward; elytral and pronotal setae slightly arcuate, recumbent; metasternal setae uniform, shorter than prosternal; pubescence on abdominal tergites parallel; that on sternites IV-VII uniform, except for a pair of subapical macrosetae on each sternite. Frons, pronotum, elytra and anterior portion of prohypomera granulate; frontal granulation conspicuous, with granula about as high as their diameter, or higher; metasternum with granulofossulation becoming denser and finer posteriomedially.
</paragraph>
<paragraph id="8BE16967AF735D5AFF7BAF00759FDB8D" blockId="13.[151,1074,227,1178]" pageId="13" pageNumber="120">Frons forming above clypeus a sharp ridge, the latter finely evenly carinate, evenly arcuate in dorsal view; mesal portion of disc weakly evenly convex in lateral view; U-shaped frontal impression deep in middle, shallow laterally. Temples strongly convex in dorsal view. Antennae (Fig. 165) 2.1-2.3 times longer than pronotum.</paragraph>
<paragraph id="8BE16967AF735D5AFF7BAF807737D8AD" blockId="13.[151,1074,227,1178]" pageId="13" pageNumber="120">Pronotum (Fig. 169) with center moderately convex in frontal view; pronotal disc deeply depressed near middle of lateral edges, shallowly depressed along anterior margin, posterior margins and posterior portion of medial groove; the latter slightly arcuate in lateral view, deep, parallel-sided; hypomera ridged from anterior margin to laterobasal angle, disc without pit. Prosternal medial ridge absent. Scutellum with anterior margin subangulate in middle and posterior margin weakly arcuate toward right-angled apex.</paragraph>
<paragraph id="8BE16967AF735D5AFF7BACE075E6D8CD" blockId="13.[151,1074,227,1178]" pageId="13" pageNumber="120">Elytra gradually widened (Fig. 3); humeral callus low, moderately convex; disc with low swellings, moderately depressed posteriorly along lateral edge, the latter finely carinate, very finely denticulate, in dorsal view slightly arcuate.</paragraph>
<paragraph id="8BE16967AF735D5AFF7BAD4074D5D90D" blockId="13.[151,1074,227,1178]" pageId="13" pageNumber="120">Abdominal sternites II and III with medial processes as in Fig. 170, posterior portion of process of sternite III widened, or trifid.</paragraph>
<paragraph id="8BE16967AF735D5AFF7BAD00741FDE0D" blockId="13.[151,1074,227,1178]" pageId="13" pageNumber="120">Male: Frontoclypeal area not modified. Protarsomeres 1 lacking tenent setae. Mesofemora (Fig. 158) as long as metafemora. Mesotibiae (Fig. 159) shorter than metatibiae. Metatarsomeres 1 about as long as combined length of metatarsomeres 2-4. Peg-like setae arranged in a single row on mesotrochanters (Fig. 158), grouped in a field on mesotibiae, and absent from protrochanters, profemora, protibiae, mesofemora, metatrochanters, metafemora and metatibiae. Apex of abdominal tergite VIII as in Figs 156, 157. Sternite VIII as in Figs 160-161. Sternite IX lacking subbasal protuberance. Aedeagus as in Figs 154, 155.</paragraph>
<paragraph id="8BE16967AF735D5AFF7BAA0077A7DEAD" blockId="13.[151,1074,227,1178]" pageId="13" pageNumber="120">Female: Abdominal tergite VIII (Figs 166-167) with a medioapical projection. Sternite VIII as in Fig. 168. Genital segments as in Figs 162-164; gonocoxal plate bearing a mediodorsal ridge.</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF735D5AFF7BAAED7469DF32" pageId="13" pageNumber="120" type="distribution">
<paragraph id="8BE16967AF735D5AFF7BAAED7469DF32" blockId="13.[151,1074,1198,1285]" pageId="13" pageNumber="120">
DISTRIBUTION AND NATURAL HISTORY:
<taxonomicName id="4C5E12E4AF735D5AFD02AAED75E2DEF2" authorityName="Sharp" authorityYear="1887" box="[666,910,1198,1221]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="120" phylum="Arthropoda" rank="species" species="solitarius">
<emphasis id="B92AB575AF735D5AFD02AAED75E2DEF2" box="[666,910,1198,1221]" italics="true" pageId="13" pageNumber="120">Megarthrus solitarius</emphasis>
</taxonomicName>
is apparently endemic to
<collectingCountry id="F34929F7AF735D5AFE83AA8D77E2DED2" box="[283,398,1230,1253]" name="Guatemala" pageId="13" pageNumber="120">Guatemala</collectingCountry>
(Dept de
<collectingRegion id="499AA785AF735D5AFD9BAA8D74F4DED2" box="[515,664,1230,1253]" country="Guatemala" name="Baja Verapaz" pageId="13" pageNumber="120">Baja Verapaz</collectingRegion>
), where it was collected in
<taxonomicName id="4C5E12E4AF735D5AFC57AA8D768FDF32" class="Magnoliopsida" family="Urticaceae" genus="Cecropia" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="120" phylum="Tracheophyta" rank="species" species="treefall">
<emphasis id="B92AB575AF735D5AFC57AA8D768FDF32" italics="true" pageId="13" pageNumber="120">Cecropia treefall</emphasis>
</taxonomicName>
at an elevation of
<quantity id="4CA6C482AF735D5AFE34AAAD746DDF32" box="[428,513,1262,1285]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="13" pageNumber="120" unit="m" value="1650.0">1650 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF735D5AFF7BAB5A74E9DFA7" pageId="13" pageNumber="120" type="discussion">
<paragraph id="8BE16967AF735D5AFF7BAB5A74E9DFA7" blockId="13.[151,1073,1305,1424]" pageId="13" pageNumber="120">
COMMENTS:
<taxonomicName id="4C5E12E4AF735D5AFEF3AB5A7434DF07" authorityName="Sharp" authorityYear="1887" box="[363,600,1305,1328]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="120" phylum="Arthropoda" rank="species" species="solitarius">
<emphasis id="B92AB575AF735D5AFEF3AB5A7434DF07" box="[363,600,1305,1328]" italics="true" pageId="13" pageNumber="120">Megarthrus solitarius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5E12E4AF735D5AFD0CAB5A7549DF07" authorityName="Sharp. These" authorityYear="1887" box="[660,805,1305,1328]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="120" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="B92AB575AF735D5AFD0CAB5A7549DF07" box="[660,805,1305,1328]" italics="true" pageId="13" pageNumber="120">M. zunilensis</emphasis>
</taxonomicName>
are apparently the only Neotropical
<taxonomicName id="4C5E12E4AF735D5AFEBCAB7A77CEDF67" authorityName="Curtis" authorityYear="1829" box="[292,418,1337,1360]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B92AB575AF735D5AFEBCAB7A77CEDF67" box="[292,418,1337,1360]" italics="true" pageId="13" pageNumber="120">Megarthrus</emphasis>
</taxonomicName>
with the abdominal tergite VIII forming a medioapical projection in the female. These two species differ, however, in most of the other sexual characters. See discussion under
<taxonomicName id="4C5E12E4AF735D5AFE62AB3A7412DFA7" authorityName="Bruch" authorityYear="1940" box="[506,638,1401,1424]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="120" phylum="Arthropoda" rank="species" species="ogloblini">
<emphasis id="B92AB575AF735D5AFE62AB3A7412DFA7" box="[506,638,1401,1424]" italics="true" pageId="13" pageNumber="120">M. ogloblini</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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