1548 lines
201 KiB
XML
1548 lines
201 KiB
XML
<document id="8DAD794C7FF15106408539AF5DAADB84" ID-CLB-Dataset="31287" ID-DOI="10.11646/zootaxa.4358.2.1" ID-GBIF-Dataset="d226f633-d454-4e32-b12f-6d00c30bddfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="1067996" ID-ZooBank="8861BCC0-022F-4803-98E8-D28B90F666E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511949109352" checkinUser="plazi" docAuthor="Dow, Rory A., Stokvis, Frank & Ngiam, Robin W. J." docDate="2017" docId="03C3A90C9855FF87FF6A88FDFC0EEE3A" docLanguage="en" docName="zootaxa.4358.2.1.pdf.imf" docOrigin="Zootaxa 4358 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptogomphus coomansi Laidlaw 1936" docType="treatment" docVersion="9" lastPageNumber="213" masterDocId="FFFAD1749852FF8AFFFD8823FF9DEF7A" masterDocTitle="Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae)" masterLastPageNumber="257" masterPageNumber="201" pageNumber="208" updateTime="1698505057438" updateUser="ExternalLinkService">
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<mods:title id="E2B13A56FA0EEC7BE6C1328B6FCDE7B1">Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae)</mods:title>
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<mods:namePart id="9E5A846193FBB9B8A40EF982D0B6DE57">Dow, Rory A.</mods:namePart>
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<mods:namePart id="58F15C53894325D25E9C0F0194544DCB">Stokvis, Frank</mods:namePart>
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<mods:namePart id="1EDFA6F64C1D8CB164AF0203D28A6F56">Ngiam, Robin W. J.</mods:namePart>
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<mods:title id="4DC759A57FAA3BF5E06357F994B6B7EB">Zootaxa</mods:title>
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<mods:date id="316D6AB96A45057616FED00EDD362AC9">2017</mods:date>
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<mods:number id="A7BD88E0AF395ECC90BD70C8AED9FEBA">4358</mods:number>
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<treatment id="03C3A90C9855FF87FF6A88FDFC0EEE3A" ID-DOI="http://doi.org/10.5281/zenodo.5631246" ID-GBIF-Taxon="137580775" ID-Zenodo-Dep="5631246" LSID="urn:lsid:plazi:treatment:03C3A90C9855FF87FF6A88FDFC0EEE3A" httpUri="http://treatment.plazi.org/id/03C3A90C9855FF87FF6A88FDFC0EEE3A" lastPageId="13" lastPageNumber="213" pageId="7" pageNumber="208">
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<paragraph id="8BD5181A9855FF8DFF6A88FDFDE6EF82" blockId="7.[151,1437,222,318]" box="[151,635,222,248]" pageId="7" pageNumber="208">
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<heading id="D09DAF769855FF8DFF6A88FDFDE6EF82" bold="true" box="[151,635,222,248]" fontSize="11" level="1" pageId="7" pageNumber="208" reason="1">
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<taxonomicName id="4C6A63999855FF8DFF6A88FDFDE6EF82" ID-CoL="3TQ48" authority="Laidlaw, 1936" authorityName="Laidlaw" authorityYear="1936" box="[151,635,222,248]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="coomansi">
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<emphasis id="B91EC4089855FF8DFF6A88FDFDE6EF82" bold="true" box="[151,635,222,248]" pageId="7" pageNumber="208">
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<emphasis id="B91EC4089855FF8DFF6A88FDFE58EF82" bold="true" box="[151,453,222,248]" italics="true" pageId="7" pageNumber="208">Leptogomphus coomansi</emphasis>
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<bibRefCitation id="EFFB65EB9855FF8DFE3188FDFDE6EF82" author="Laidlaw" box="[460,635,222,248]" pageId="7" pageNumber="208" refString="Laidlaw, F. F. (1936) On some examples of the genus Leptogomphus (Odon., Gomphidae). Treubia, 15 (3), 267 - 270." type="journal article" year="1936">Laidlaw, 1936</bibRefCitation>
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<paragraph id="8BD5181A9855FF8DFF6A8922FE9FEE44" blockId="7.[151,1437,222,318]" pageId="7" pageNumber="208">
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(
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<figureCitation id="1351049F9855FF8DFF628922FF7BEE63" box="[159,230,257,282]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="7" pageNumber="208">Figs 1</figureCitation>
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,
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<figureCitation id="1351049F9855FF8DFF088922FE98EE63" box="[245,261,257,281]" captionStart="FIGURE 2" captionStartId="5.[151,250,1294,1316]" captionTargetBox="[243,1384,200,1237]" captionTargetId="figure@5.[151,1436,193,1273]" captionTargetPageId="5" captionText="FIGURE 2. ITS gene tree for 20 specimens of Leptogomphus and three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068000/files/figure.png" pageId="7" pageNumber="208">2</figureCitation>
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,
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<figureCitation id="1351049F9855FF8DFEEF8922FEBFEE63" box="[274,290,257,281]" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1893]" captionTargetBox="[209,1373,710,1811]" captionTargetId="figure@6.[151,1436,700,1850]" captionTargetPageId="6" captionText="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068002/files/figure.png" pageId="7" pageNumber="208">3</figureCitation>
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,
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<figureCitation id="1351049F9855FF8DFED28922FEA2EE63" box="[303,319,257,281]" captionStart="FIGURES 4–9" captionStartId="9.[151,264,1713,1736]" captionTargetBox="[151,1436,193,1692]" captionTargetId="figure@9.[151,1436,193,1692]" captionTargetPageId="9" captionText="FIGURES 4–9. Male head dorsal: (4) L. coomansi SAR09_10_GOM4; (5) L. pendleburyi SAR13_14_GOM13; (6) L. pasia SAB12_GOM1; (7) L. sii holotype; (8) L.schieli holotype; (9) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068004/files/figure.png" pageId="7" pageNumber="208">4</figureCitation>
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,
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<figureCitation id="1351049F9855FF8DFEB18922FE09EE63" box="[332,404,257,281]" captionStart="FIGURES10–13" captionStartId="14.[151,264,1864,1887]" captionTargetBox="[151,1436,772,1843]" captionTargetId="figure@14.[151,1436,772,1843]" captionTargetPageId="14" captionText="FIGURES10–13. Male head frontal: (10) L.coomansi SAR09_10_GOM4; (11) L.coomansi holotype; (12) L.sp. cf coomansi SAB12_GOM4; (13) L.pendleburyi SAR13_14_GOM13." httpUri="https://zenodo.org/record/1068006/files/figure.png" pageId="7" pageNumber="208">10, 11</figureCitation>
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,
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<figureCitation id="1351049F9855FF8DFE5C8922FE77EE63" box="[417,490,257,281]" captionStart="FIGURES18–23" captionStartId="18.[151,264,1895,1918]" captionTargetBox="[183,1420,198,1847]" captionTargetId="figure@18.[183,1420,198,1847]" captionTargetPageId="18" captionText="FIGURES18–23.Female headdorsal:(18) L.coomansi SAR11_12_GOM48;(19)L.mariaeholotype;(20) L.sp.cf coomansi PomS260; (21) L. pasia holotype; (22) L.sii paratype SAR13_14_GOM44; (23) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068010/files/figure.png" pageId="7" pageNumber="208">18, 19</figureCitation>
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<figureCitation id="1351049F9855FF8DFE0A8922FDDDEE63" box="[503,576,257,281]" captionStart="FIGURES28–31" captionStartId="22.[151,264,1357,1380]" captionTargetBox="[151,1435,266,1336]" captionTargetId="figure@22.[151,1436,265,1336]" captionTargetPageId="22" captionText="FIGURES28–31. Female head dorsaldetail; h—occipital horn; ot—occipital tube; p—pit; s—occipital spur; t—tubercle;thtubercle horn: (28) L. coomansi SAR11_12_GOM48; (29) L. mariae holotype; (30) L. sp. cf coomansi PomS260; (31) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068014/files/figure.png" pageId="7" pageNumber="208">28, 29</figureCitation>
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<figureCitation id="1351049F9855FF8DFDB08922FD0AEE63" box="[589,663,257,282]" captionStart="FIGURES 36–41" captionStartId="25.[151,264,1973,1996]" captionTargetBox="[204,1387,183,1958]" captionTargetId="figure@25.[204,1387,183,1958]" captionTargetPageId="25" captionText="FIGURES 36–41. Female head frontal: (36) L. coomansi SAR11_12_GOM48; (37) L. mariae holotype; (38) L. sp. cf coomansi PomS260; (39) L.pasia holotype; (40) L.sii paratype SAR13_14_GOM44; (41) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068018/files/figure.png" pageId="7" pageNumber="208">36, 37</figureCitation>
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<figureCitation id="1351049F9855FF8DFD5E8922FD5FEE60" box="[675,706,257,282]" captionStart="FIGURES 46–51" captionStartId="29.[151,264,1672,1695]" captionTargetBox="[156,1432,375,1651]" captionTargetId="figure@29.[156,1432,375,1651]" captionTargetPageId="29" captionText="FIGURES 46–51. Female head frontal detail; h—occipital horn; ot—occipital tube; t—tubercle; th—tubercle horn: (46) L. coomansi SAR11_12_GOM48; (47) L pendleburyi SAR16_GOM8; (48) L. schieli paratype SAR15_GOM6; (49) L. species SAR11_12_GOM33; (50) L. williamsoni SAR13_14_GOM51; (51) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068022/files/figure.png" pageId="7" pageNumber="208">46</figureCitation>
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<figureCitation id="1351049F9855FF8DFD338922FC8AEE60" box="[718,791,257,282]" captionStart="FIGURES 52–55" captionStartId="31.[151,264,1501,1524]" captionTargetBox="[151,1436,409,1479]" captionTargetId="figure@31.[151,1436,409,1480]" captionTargetPageId="31" captionText="FIGURES 52–55. Male synthorax dorsal: (52) L. coomansi SAR09_10_GOM4; (53) L. coomansi holotype; (54) L. sp. cf coomansi SAB12_GOM4; (55) L. pendleburyi SAR13_14_GOM13." httpUri="https://zenodo.org/record/1068024/files/figure.png" pageId="7" pageNumber="208">52, 53</figureCitation>
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,
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<figureCitation id="1351049F9855FF8DFCD98922FCDFEE60" box="[804,834,257,282]" captionStart="FIGURES 60–63" captionStartId="34.[151,264,1601,1624]" captionTargetBox="[156,1432,517,1580]" captionTargetId="figure@34.[156,1432,517,1580]" captionTargetPageId="34" captionText="FIGURES 60–63. Male synthorax lateral: (60) L. coomansi SAR09_10_GOM4; (61) L. sp. cf coomansi SAB12_GOM4; (62) L. schieli holotype; (63) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068028/files/figure.png" pageId="7" pageNumber="208">60</figureCitation>
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<figureCitation id="1351049F9855FF8DFCB28922FC05EE60" box="[847,920,257,282]" captionStart="FIGURES 68–73" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[156,1432,193,1894]" captionTargetId="figure@36.[156,1432,193,1894]" captionTargetPageId="36" captionText="FIGURES 68–73. Female synthorax dorsal: (68) L. coomansi SAR11_12_GOM48; (69) L. mariae holotype; (70) L. sp. cf coomansi PomS260; (71) L.pasia holotype; (72) L.sii paratype SAR13_14_GOM44; (73) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068032/files/figure.png" pageId="7" pageNumber="208">68, 69</figureCitation>
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<figureCitation id="1351049F9855FF8DFC588922FC73EE63" box="[933,1006,257,281]" captionStart="FIGURES 78–83" captionStartId="38.[151,264,1952,1975]" captionTargetBox="[169,1406,182,1934]" captionTargetId="figure@38.[169,1406,182,1934]" captionTargetPageId="38" captionText="FIGURES 78–83. Female synthorax lateral: (78) L. coomansi SAR11_12_GOM48; (79) L. mariae holotype, right hand side, flipped for consistency; (80) L. sp. cf coomansi PomS260; (81) L. pasia holotype; (82) L. sii paratype SAR13_14_GOM44; (83) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068036/files/figure.png" pageId="7" pageNumber="208">78, 79</figureCitation>
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<figureCitation id="1351049F9855FF8DFC078922FB85EE63" box="[1018,1048,257,281]" captionStart="FIGURES 88–93" captionStartId="40.[151,264,1280,1303]" captionTargetBox="[156,1432,195,1258]" captionTargetId="figure@40.[156,1432,195,1258]" captionTargetPageId="40" captionText="FIGURES 88–93. Male accessory genitalia, lateral view, all scale bars 1mm: (88) L. coomansi SAR09_10_GOM4; (89) L. pasia SAB12_GOM1; (90) L. pendleburyi SAR13_14_GOM13; (91) L. schieli holotype; (92) L. sii holotype; (93) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068040/files/figure.png" pageId="7" pageNumber="208">88</figureCitation>
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<figureCitation id="1351049F9855FF8DFBD88922FBD9EE63" box="[1061,1092,257,281]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="7" pageNumber="208">94</figureCitation>
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,
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<figureCitation id="1351049F9855FF8DFBAC8922FBE0EE63" box="[1105,1149,257,281]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="7" pageNumber="208">100</figureCitation>
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,
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<figureCitation id="1351049F9855FF8DFB778922FB2BEE60" box="[1162,1206,257,282]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="7" pageNumber="208">106</figureCitation>
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,
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<figureCitation id="1351049F9855FF8DFB3F8922FB70EE63" box="[1218,1261,257,281]" captionStart="FIGURES 112–117" captionStartId="44.[151,264,1808,1831]" captionTargetBox="[156,1432,193,1787]" captionTargetId="figure@44.[156,1432,193,1787]" captionTargetPageId="44" captionText="FIGURES 112–117. Male anal appendages in ventral view (all scale bars 0.5mm): (112) L. coomansi SAR09_10_GOM4; (113) L.pasia SAB12_GOM1;(114) L.pendleburyi SAR13_14_GOM13; (115) L. schieli holotype; (116) L.sii holotype; (117) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068048/files/figure.png" pageId="7" pageNumber="208">112</figureCitation>
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|
||
<figureCitation id="1351049F9855FF8DFB078922FAB8EE63" box="[1274,1317,257,281]" captionStart="FIGURES 118–119" captionStartId="45.[151,264,604,627]" captionTargetBox="[156,1432,193,583]" captionTargetId="figure@45.[156,1432,193,583]" captionTargetPageId="45" captionText="FIGURES 118–119. Ventral view female abdominal S8 (all scale bars 0.5mm): (118) L. coomansi SAR11_12_GOM48; (119) L. pasia SAB12_GOM2." httpUri="https://zenodo.org/record/1068050/files/figure.png" pageId="7" pageNumber="208">118</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9855FF8DFACF8922FAC3EE63" box="[1330,1374,257,281]" captionStart="FIGURES 120–125" captionStartId="46.[151,264,1891,1914]" captionTargetBox="[151,1435,372,1869]" captionTargetId="figure@46.[151,1436,371,1870]" captionTargetPageId="46" captionText="FIGURES 120–125. Female vulvar scale (all scale bars 0.5mm): (120) L. coomansi SAR11_12_GOM48; (121) L. pasia holotype; (122) L. pendleburyi SAR13_14_GOM15; (123) L. schieli paratype SAR15_GOM6; (124) L. sii paratype SAR13_14_GOM44; (125) L.williamsoni SAR13_14_GOM51." httpUri="https://zenodo.org/record/1068052/files/figure.png" pageId="7" pageNumber="208">120</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9855FF8DFA968922FA0AEE60" box="[1387,1431,257,282]" captionStart="FIGURE 126" captionStartId="48.[151,250,1801,1823]" captionTargetBox="[183,1420,421,1756]" captionTargetId="figure@48.[183,1420,421,1756]" captionTargetPageId="48" captionText="FIGURE 126. Distribution of L. coomansi, confirmed (yellow square) and unconfirmed (blue triangle) records, and L. sp. cf coomansi (black circle)." httpUri="https://zenodo.org/record/1068054/files/figure.png" pageId="7" pageNumber="208">126</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9855FF8DFF6A8905FF67EE44" box="[151,250,294,318]" captionStart="FIGURES 133–137" captionStartId="53.[151,264,1171,1194]" captionTargetBox="[151,1436,193,1150]" captionTargetId="figure@53.[151,1436,193,1150]" captionTargetPageId="53" captionText="FIGURES 133–137. Photographs in life of some Bornean Leptogomphus species: (133) L. coomansi male (left hand side, flippedto right for consistency with other images), Sungai Sbong, Kapit Division, Sarawak, photograph by G.T.Reels; (134) L. coomansi female, Ranchan Recreational Park, Serian, Serian Division, Sarawak; (135) L. pendleburyi female, Bukit Tanggan, Ulu Mujok, Lanjak Entimau WildlifeSanctuary, Sarikei Division, Sarawak, photograph by Robin Ngiam; (136) L. pendleburyi male, Sungai Kahei area, Ulu Balui, Kapit Division, Sarawak, photograph by Robin Ngiam; (137) L. williamsoni immature male, semi-teneral, Sungai Kahei area, Ulu Balui, Kapit Division,Sarawak, photograph by Robin Ngiam." httpUri="https://zenodo.org/record/1068064/files/figure.png" pageId="7" pageNumber="208">133, 134</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B919855FF8DFF6A894EFC23ECF6" pageId="7" pageNumber="208" type="reference_group">
|
||
<paragraph id="8BD5181A9855FF8DFF6A894EFE03EDED" blockId="7.[151,1436,365,663]" pageId="7" pageNumber="208">
|
||
<treatmentCitationGroup id="AB7A3F349855FF8DFF6A894EFE03EDED" pageId="7" pageNumber="208">
|
||
<taxonomicName id="4C6A63999855FF8DFF6A894EFDFFEEF9" ID-CoL="3TQ48" authority="Laidlaw 1936: 267" authorityName="Laidlaw" authorityPageNumber="267" authorityYear="1936" box="[151,610,365,387]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC4089855FF8DFF6A894EFE08EEF9" box="[151,405,365,387]" italics="true" pageId="7" pageNumber="208">Leptogomphus coomansi</emphasis>
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFE60894EFDFFEEF9" author="Laidlaw" box="[413,610,365,387]" page="267" pageId="7" pageNumber="208" year="1936">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFE60894EFDB2EEF9" author="Laidlaw" box="[413,559,365,387]" pageId="7" pageNumber="208" refString="Laidlaw, F. F. (1936) On some examples of the genus Leptogomphus (Odon., Gomphidae). Treubia, 15 (3), 267 - 270." type="journal article" year="1936">Laidlaw 1936</bibRefCitation>
|
||
: 267
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
–269,
|
||
<figureCitation id="1351049F9855FF8DFD5F894EFD43EEF9" box="[674,734,365,387]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="7" pageNumber="208">Fig. 1</figureCitation>
|
||
(original description, holotype ♂, allotype ♀, Singkawang area);—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFF3A89AFFE06EED8" author="Lieftinck" box="[199,411,396,418]" page="245" pageId="7" pageNumber="208" year="1948">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFF3A89AFFEF8EED8" author="Lieftinck" box="[199,357,396,418]" pageId="7" pageNumber="208" refString="Lieftinck, M. A. (1948) Descriptions and records of South - east Asiatic Odonata. Treubia, 19, 221 - 278." type="journal article" year="1948">Lieftinck 1948</bibRefCitation>
|
||
: 245
|
||
</treatmentCitation>
|
||
–247, Plate 8,
|
||
<figureCitation id="1351049F9855FF8DFDC089AFFD17EED8" box="[573,650,396,418]" captionStart="FIGURES10–13" captionStartId="14.[151,264,1864,1887]" captionTargetBox="[151,1436,772,1843]" captionTargetId="figure@14.[151,1436,772,1843]" captionTargetPageId="14" captionText="FIGURES10–13. Male head frontal: (10) L.coomansi SAR09_10_GOM4; (11) L.coomansi holotype; (12) L.sp. cf coomansi SAB12_GOM4; (13) L.pendleburyi SAR13_14_GOM13." httpUri="https://zenodo.org/record/1068006/files/figure.png" pageId="7" pageNumber="208">Fig. 10</figureCitation>
|
||
(types re-examined);—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFC7989AFFBD6EED8" author="Lieftinck" box="[900,1099,396,418]" page="81" pageId="7" pageNumber="208" year="1954">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFC7989AFFBBBEED8" author="Lieftinck" box="[900,1062,396,418]" pageId="7" pageNumber="208" refString="Lieftinck, M. A. (1954) Handlist of Malaysian Odonata. A catalogue of the dragonflies of the Malay Peninsula, Sumatra, Java and Borneo, including the adjacent small islands. Treubia, 22 (Supplement), i - xiii + 1 - 202." type="journal article" year="1954">Lieftinck 1954</bibRefCitation>
|
||
: 81
|
||
</treatmentCitation>
|
||
;—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFB9789AFFAABEED8" author="Lieftinck" box="[1130,1334,396,418]" page="81" pageId="7" pageNumber="208" year="1971">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFB9789AFFA93EED8" author="Lieftinck" box="[1130,1294,396,418]" pageId="7" pageNumber="208" refString="Lieftinck, M. A. (1971) A catalogue of type - specimens of Odonata preserved in The Netherlands, with a supplementary list of the Odonata types described by Dutch scientists deposited in foreign institutional collections. Tijdschrift voor Entomologie, 114, 65 - 139." type="journal article" year="1971">Lieftinck 1971</bibRefCitation>
|
||
: 81
|
||
</treatmentCitation>
|
||
(note on holotype and “first described” female);—?
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFD848989FC02EEBA" author="Tsuda" box="[633,927,426,448]" page="38" pageId="7" pageNumber="208" year="1989">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFD848989FCE6EEBA" author="Tsuda" box="[633,891,426,448]" pageId="7" pageNumber="208" refString="Tsuda, S. and Kitagawa, K. (1989) Odonata of Southeast Asia collected by the late Mr. M. Iwasaki. Part 3. Odonata of North Borneo (Sabah, Malaysia). Gracile, 40, 37 - 39." type="journal article" year="1989">Tsuda & Kitagawa 1989</bibRefCitation>
|
||
: 38
|
||
</treatmentCitation>
|
||
(♀, Poring, Sabah);—van
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFB4B8989FAA6EEBA" author="Tol" box="[1206,1339,426,448]" page="98" pageId="7" pageNumber="208" year="1990">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFB4B8989FA88EEBA" author="Tol" box="[1206,1301,426,448]" pageId="7" pageNumber="208" refString="van Tol, J. (1990) Key to the Malesian species of Leptogomphus Selys, with the description of a new species from Sabah (Odonata, Gomphidae). Tijdschrift voor Entomologie, 133, 97 - 105." type="journal article" year="1990">Tol 1990</bibRefCitation>
|
||
: 98
|
||
</treatmentCitation>
|
||
, 99, 101, 104,
|
||
<figureCitation id="1351049F9855FF8DFF0789EAFED5EEA5" box="[250,328,457,479]" captionStart="FIGURES18–23" captionStartId="18.[151,264,1895,1918]" captionTargetBox="[183,1420,198,1847]" captionTargetId="figure@18.[183,1420,198,1847]" captionTargetPageId="18" captionText="FIGURES18–23.Female headdorsal:(18) L.coomansi SAR11_12_GOM48;(19)L.mariaeholotype;(20) L.sp.cf coomansi PomS260; (21) L. pasia holotype; (22) L.sii paratype SAR13_14_GOM44; (23) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068010/files/figure.png" pageId="7" pageNumber="208">Figs 18</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9855FF8DFEA989EAFE0AEEA5" box="[340,407,457,479]" captionStart="FIGURES 24–27" captionStartId="20.[151,264,1441,1464]" captionTargetBox="[163,1439,268,1402]" captionTargetId="figure@20.[163,1439,268,1402]" captionTargetPageId="20" captionText="FIGURES 24–27. Female head dorsal: (24) L pendleburyi SAR16_GOM8; (25) L. schieli paratype SAR15_GOM6; (26) L. williamsoni SAR13_14_GOM51; (27) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068012/files/figure.png" pageId="7" pageNumber="208">26, 27</figureCitation>
|
||
,
|
||
<tableCitation id="C6E82DA19855FF8DFE5F89EAFE73EEA5" box="[418,494,457,479]" captionStart="TABLE 1" captionStartId="2.[151,239,358,381]" captionText="TABLE 1. Primer combinations used for amplification of ITS and COI." pageId="7" pageNumber="208">Table 1</tableCitation>
|
||
,
|
||
<tableCitation id="C6E82DA19855FF8DFE0689EAFD95EEA5" box="[507,520,457,479]" captionStart="TABLE 2" captionStartId="3.[151,239,256,279]" captionTargetBox="[159,1375,443,1403]" captionText="TABLE 2. Leptogomphus from Borneo used in the molecular analyses; collectors and more details of location given in material lists under each species. Collection codes and BOLD Process IDs are given along with basic location details. The collection codes can be used to locate the sequences on the BOLD website, and also appear as BOLD Sample IDs there." pageId="7" pageNumber="208">2</tableCitation>
|
||
(key);—van
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFD6D89EAFC8EEEA5" author="Tol" box="[656,787,457,479]" page="72" pageId="7" pageNumber="208" year="1992">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFD6D89EAFD72EEA5" author="Tol" box="[656,751,457,479]" pageId="7" pageNumber="208" refString="van Tol, J. (1992) An annotated index to names of Odonata used in publications by M. A. Lieftinck. Zoologische Verhandelingen, 279, 1 - 263." type="journal article" year="1992">Tol 1992</bibRefCitation>
|
||
: 72
|
||
</treatmentCitation>
|
||
;—Thompson & van
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFC1389EAFBEDEEA5" author="Tol" box="[1006,1136,457,479]" page="70" pageId="7" pageNumber="208" year="1993">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFC1389EAFBD1EEA5" author="Tol" box="[1006,1100,457,479]" pageId="7" pageNumber="208" refString="Thompson, D. J. & van Tol, J. (1993) Damselflies and dragonflies from four forest types in Brunei. Brunei Museum Journal, 8, 57 - 72." type="journal article" year="1993">Tol 1993</bibRefCitation>
|
||
: 70
|
||
</treatmentCitation>
|
||
(Belait, Brunei);—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFACB89EAFF71EE84" author="Orr" page="198" pageId="7" pageNumber="208" year="2001">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFACB89EAFA05EEA5" author="Orr" box="[1334,1432,457,479]" pageId="7" pageNumber="208" refString="Orr, A. G. (2001) An annotated checklist of the Odonata of Brunei with ecological notes and descriptions of hitherto unknown males and larvae. International Journal of Odonatology, 4, 167 - 220. https: // doi. org / 10.1080 / 13887890.2001.9748168" type="journal article" year="2001">Orr 2001</bibRefCitation>
|
||
: 198
|
||
</treatmentCitation>
|
||
–199 (Brunei);—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFE6089CBFDB6EE84" author="Orr" box="[413,555,488,510]" page="40" pageId="7" pageNumber="208" year="2003">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFE6089CBFD9FEE84" author="Orr" box="[413,514,488,510]" pageId="7" pageNumber="208" refString="Orr, A. G. (2003) A guide to the dragonflies of Borneo: their identification and biology. Natural History Publications (Borneo), Kota Kinabalu, x + 195 pp." type="book" year="2003">Orr 2003</bibRefCitation>
|
||
: 40
|
||
</treatmentCitation>
|
||
, 96–97, Figs 141, 142, plate 12c (discussed);—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFBDF89CBFA84EE84" author="Dow" box="[1058,1305,488,510]" page="15" pageId="7" pageNumber="208" year="2010">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFBDF89CBFB6EEE84" author="Dow" box="[1058,1267,488,510]" pageId="7" pageNumber="208" refString="Dow, R. A. & Reels, G. T. (2010) The Odonata of three national parks in Sarawak. Agrion, 14 (1), 14 - 19." type="journal article" year="2010">Dow & Reels 2010</bibRefCitation>
|
||
: 15
|
||
</treatmentCitation>
|
||
(Kubah NP, Sarawak);—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFEBB8A25FD83ED66" author="Dolny" box="[326,542,518,541]" page="73" pageId="7" pageNumber="208" year="2011">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFEBB8A25FE64ED66" author="Dolny" box="[326,505,518,541]" pageId="7" pageNumber="208" refString="Dolny, A., Barta, D., Lhota, S., Rusdianto & Drozd, P. (2011) Dragonflies (Odonata) in the Bornean rain forest as indicators of changes in biodiversity resulting from forest modifications and destruction. Tropical Zoology, 24, 63 - 86." type="journal article" year="2011">
|
||
Dolný
|
||
<emphasis id="B91EC4089855FF8DFE718A2BFE22ED66" box="[396,447,518,541]" italics="true" pageId="7" pageNumber="208">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
: 73
|
||
</treatmentCitation>
|
||
, 85 (Sungai Wain, Kailmantan Timur);—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFC3C8A25FB2BED66" author="Barta" box="[961,1206,518,540]" page="99" pageId="7" pageNumber="208" year="2013">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFC3C8A25FB09ED66" author="Barta" box="[961,1172,518,540]" pageId="7" pageNumber="208" refString="Barta, D. & Dolny, A. (2013) Dragonflies of Sungai Wain. Ecological Field Guide to the Dragonflies of Lowland Mixed Dipterocarp Forest of Southeastern Kalimantan. Taita Publishers, Hradec Kralove, 168 pp." type="book" year="2013">Bárta & Dolný 2013</bibRefCitation>
|
||
: 99
|
||
</treatmentCitation>
|
||
(images of female);— Dow & Reels 2013: 20,
|
||
<figureCitation id="1351049F9855FF8DFE3C8A06FD97ED41" box="[449,522,549,571]" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1933,1956]" captionTargetBox="[151,1436,841,1912]" captionTargetId="figure@16.[151,1436,841,1912]" captionTargetPageId="16" captionText="FIGURES 14–17. Male head frontal: (14) L. pasia SAB12_GOM1; (15) L. sii holotype; (16) L. schieli holotype; (17) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068008/files/figure.png" pageId="7" pageNumber="208">Fig. 16</figureCitation>
|
||
(Ranchan Recreational Park, Sarawak);—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFC468A06FB0BED41" author="Dow" box="[955,1174,549,571]" page="19" pageId="7" pageNumber="208" year="2013">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFC468A06FBEEED41" author="Dow" box="[955,1139,549,571]" pageId="7" pageNumber="208" refString="Dow, R. A., Reels, G. T. & Butler, S. G. (2013 b) Previously unpublished Odonata records from Sarawak, Borneo. Part II. Kubah National Park. Faunistic Studies in South-East Asian and Pacific Island Odonata, 6, 1 - 21." type="journal article" year="2013" yearSuffix="b">
|
||
Dow
|
||
<emphasis id="B91EC4089855FF8DFC088A05FBB4ED41" box="[1013,1065,549,571]" italics="true" pageId="7" pageNumber="208">et al.</emphasis>
|
||
2013b
|
||
</bibRefCitation>
|
||
: 19
|
||
</treatmentCitation>
|
||
(Kubah NP, Sarawak);— Dow & Ngiam 2015: 24 (Sungai Sii, upper Baram, Sarawak);—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFCAB8A67FBADED20" author="Dow" box="[854,1072,580,602]" page="10" pageId="7" pageNumber="208" year="2015">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFCAB8A67FB96ED20" author="Dow" box="[854,1035,580,602]" pageId="7" pageNumber="208" refString="Dow, R. A., Reels, G. T. & Ngiam, R. W. J. (2015 a) Previously unpublished Odonata records from Sarawak, Borneo, Part III. Sri Aman, Sibu and Kapit Divisions. Faunistic Studies in South-East Asian and Pacific Island Odonata, 9, 1 - 34." type="journal article" year="2015" yearSuffix="a">
|
||
Dow
|
||
<emphasis id="B91EC4089855FF8DFC728A66FC5EED20" box="[911,963,580,602]" italics="true" pageId="7" pageNumber="208">et al.</emphasis>
|
||
2015a
|
||
</bibRefCitation>
|
||
: 10
|
||
</treatmentCitation>
|
||
, 27 (Batang Ai NP and locations in Kapit Division, Sarawak);—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFE0C8A41FD2EED02" author="Steinhoff" box="[497,691,610,632]" page="8" pageId="7" pageNumber="208" year="2015">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFE0C8A41FD0BED02" author="Steinhoff" box="[497,662,610,632]" pageId="7" pageNumber="208" refString="Steinhoff, P. O. M. (2015) Results of Odonata larval rearing in the Gunung Mulu National Park, Sarawak, Malaysia from April to August 2014. International Dragonfly Fund Report, 78, 1 - 11." type="journal article" year="2015">Steinhoff 2015</bibRefCitation>
|
||
: 8
|
||
</treatmentCitation>
|
||
,
|
||
<figureCitation id="1351049F9855FF8DFD388A41FC8EED02" box="[709,787,610,632]" captionStart="FIGURES 4–9" captionStartId="9.[151,264,1713,1736]" captionTargetBox="[151,1436,193,1692]" captionTargetId="figure@9.[151,1436,193,1692]" captionTargetPageId="9" captionText="FIGURES 4–9. Male head dorsal: (4) L. coomansi SAR09_10_GOM4; (5) L. pendleburyi SAR13_14_GOM13; (6) L. pasia SAB12_GOM1; (7) L. sii holotype; (8) L.schieli holotype; (9) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068004/files/figure.png" pageId="7" pageNumber="208">Fig. 5b</figureCitation>
|
||
(GMNP, larva reared);—Dow 2016: 8 (Samunsam Wildlife Sanctuary, Sarawak).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9855FF8DFF6A8A9DFC74EDAE" blockId="7.[151,1001,702,725]" box="[151,1001,702,725]" pageId="7" pageNumber="208">
|
||
<treatmentCitationGroup id="AB7A3F349855FF8DFF6A8A9DFC74EDAE" box="[151,1001,702,725]" pageId="7" pageNumber="208">
|
||
<taxonomicName id="4C6A63999855FF8DFF6A8A9DFEB1EDAE" ID-CoL="5D3K" authorityName="Selys in Borneo" authorityYear="1878" box="[151,300,702,724]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EC4089855FF8DFF6A8A9DFEB1EDAE" box="[151,300,702,724]" italics="true" pageId="7" pageNumber="208">Leptogomphus</emphasis>
|
||
</taxonomicName>
|
||
cf
|
||
<taxonomicName id="4C6A63999855FF8DFEB18AE3FD95EDAE" ID-CoL="3TQ48" authority="Laidlaw" authorityName="Laidlaw" authorityYear="1936" box="[332,520,702,725]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC4089855FF8DFEB18AE3FE30EDAF" box="[332,429,704,725]" italics="true" pageId="7" pageNumber="208">coomansi</emphasis>
|
||
Laidlaw
|
||
</taxonomicName>
|
||
;—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFDDB8A9DFC8DEDAE" author="Cleary" box="[550,784,702,725]" page="445" pageId="7" pageNumber="208" year="2004">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFDDB8A9DFD7CEDAE" author="Cleary" box="[550,737,702,725]" pageId="7" pageNumber="208" refString="Cleary, D. F. R., Moers, A. O., Eichhorn, K. A. O., van Tol, J., de Jong, R. & Menken, S. B. J. (2004) Diversity and community composition of butterflies and odonates in an ENSO-induced fire affected habitat mosaic: a case study from East Kalimantan, Indonesia. Oikos, 105, 426 - 446. https: // doi. org / 10.1111 / j. 0030 - 1299.2004.12219. x" type="journal article" year="2004">
|
||
Cleary
|
||
<emphasis id="B91EC4089855FF8DFD8C8AE3FD39EDAE" box="[625,676,702,725]" italics="true" pageId="7" pageNumber="208">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
: 445
|
||
</treatmentCitation>
|
||
(
|
||
<collectingRegion id="49AED6F89855FF8DFCE18A9DFC43EDAE" box="[796,990,702,724]" country="Indonesia" name="Kalimantan Timur" pageId="7" pageNumber="208">Kalimantan Timur</collectingRegion>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9855FF8DFF6A8ADFFC0AEC34" blockId="7.[151,1436,764,847]" pageId="7" pageNumber="208">
|
||
<treatmentCitationGroup id="AB7A3F349855FF8DFF6A8ADFFC0AEC34" pageId="7" pageNumber="208">
|
||
<taxonomicName id="4C6A63999855FF8DFF6A8ADFFDC0EC68" ID-CoL="3TQ4H" authority="Lieftinck 1948: 249" authorityName="Lieftinck" authorityPageNumber="249" authorityYear="1948" box="[151,605,764,786]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC4089855FF8DFF6A8ADFFEE2EC68" box="[151,383,764,786]" italics="true" pageId="7" pageNumber="208">Leptogomphus mariae</emphasis>
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFE778ADFFDC0EC68" author="Lieftinck" box="[394,605,764,786]" page="249" pageId="7" pageNumber="208" year="1948">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFE778ADFFDBAEC68" author="Lieftinck" box="[394,551,764,786]" pageId="7" pageNumber="208" refString="Lieftinck, M. A. (1948) Descriptions and records of South - east Asiatic Odonata. Treubia, 19, 221 - 278." type="journal article" year="1948">Lieftinck 1948</bibRefCitation>
|
||
: 249
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
–251, Plate 8,
|
||
<figureCitation id="1351049F9855FF8DFD068ADFFCD5EC68" box="[763,840,764,786]" captionStart="FIGURES10–13" captionStartId="14.[151,264,1864,1887]" captionTargetBox="[151,1436,772,1843]" captionTargetId="figure@14.[151,1436,772,1843]" captionTargetPageId="14" captionText="FIGURES10–13. Male head frontal: (10) L.coomansi SAR09_10_GOM4; (11) L.coomansi holotype; (12) L.sp. cf coomansi SAB12_GOM4; (13) L.pendleburyi SAR13_14_GOM13." httpUri="https://zenodo.org/record/1068006/files/figure.png" pageId="7" pageNumber="208">Fig. 10</figureCitation>
|
||
(original description
|
||
<specimenCount id="9D6CD3939855FF8DFBCD8ADEFBDEEC68" box="[1072,1091,765,786]" pageId="7" pageNumber="208" type="female">♀</specimenCount>
|
||
, Kutai);—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFB4F8ADFFAE1EC68" author="Lieftinck" box="[1202,1404,764,786]" page="82" pageId="7" pageNumber="208" year="1954">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFB4F8ADFFAC9EC68" author="Lieftinck" box="[1202,1364,764,786]" pageId="7" pageNumber="208" refString="Lieftinck, M. A. (1954) Handlist of Malaysian Odonata. A catalogue of the dragonflies of the Malay Peninsula, Sumatra, Java and Borneo, including the adjacent small islands. Treubia, 22 (Supplement), i - xiii + 1 - 202." type="journal article" year="1954">Lieftinck 1954</bibRefCitation>
|
||
: 82
|
||
</treatmentCitation>
|
||
;—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFF3A8B39FE1FEC4A" author="Lieftinck" box="[199,386,794,816]" page="99" pageId="7" pageNumber="208" year="1971">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFF3A8B39FEFDEC4A" author="Lieftinck" box="[199,352,794,816]" pageId="7" pageNumber="208" refString="Lieftinck, M. A. (1971) A catalogue of type - specimens of Odonata preserved in The Netherlands, with a supplementary list of the Odonata types described by Dutch scientists deposited in foreign institutional collections. Tijdschrift voor Entomologie, 114, 65 - 139." type="journal article" year="1971">Lieftinck 1971</bibRefCitation>
|
||
: 99
|
||
</treatmentCitation>
|
||
–100 (note on
|
||
<typeStatus id="54D1A6B89855FF8DFDE98B39FDF2EC4A" box="[532,623,794,816]" pageId="7" pageNumber="208" type="holotype">holotype</typeStatus>
|
||
);—van
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFD3D8B39FCDFEC4A" author="Tol" box="[704,834,794,816]" page="99" pageId="7" pageNumber="208" year="1990">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFD3D8B39FC83EC4A" author="Tol" box="[704,798,794,816]" pageId="7" pageNumber="208" refString="van Tol, J. (1990) Key to the Malesian species of Leptogomphus Selys, with the description of a new species from Sabah (Odonata, Gomphidae). Tijdschrift voor Entomologie, 133, 97 - 105." type="journal article" year="1990">Tol 1990</bibRefCitation>
|
||
: 99
|
||
</treatmentCitation>
|
||
, 104,
|
||
<tableCitation id="C6E82DA19855FF8DFC838B39FC54EC4A" box="[894,969,794,816]" captionStart="TABLE 1" captionStartId="2.[151,239,358,381]" captionText="TABLE 1. Primer combinations used for amplification of ITS and COI." pageId="7" pageNumber="208">Table 1</tableCitation>
|
||
,
|
||
<tableCitation id="C6E82DA19855FF8DFC288B39FC7CEC4A" box="[981,993,794,816]" captionStart="TABLE 2" captionStartId="3.[151,239,256,279]" captionTargetBox="[159,1375,443,1403]" captionText="TABLE 2. Leptogomphus from Borneo used in the molecular analyses; collectors and more details of location given in material lists under each species. Collection codes and BOLD Process IDs are given along with basic location details. The collection codes can be used to locate the sequences on the BOLD website, and also appear as BOLD Sample IDs there." pageId="7" pageNumber="208">2</tableCitation>
|
||
(female in key);—van
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFB368B39FACBEC4A" author="Tol" box="[1227,1366,794,816]" page="148" pageId="7" pageNumber="208" year="1992">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFB368B39FAB4EC4A" author="Tol" box="[1227,1321,794,816]" pageId="7" pageNumber="208" refString="van Tol, J. (1992) An annotated index to names of Odonata used in publications by M. A. Lieftinck. Zoologische Verhandelingen, 279, 1 - 263." type="journal article" year="1992">Tol 1992</bibRefCitation>
|
||
: 148
|
||
</treatmentCitation>
|
||
;—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFA8B8B39FEB0EC35" author="Orr" page="199" pageId="7" pageNumber="208" year="2001">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFA8B8B39FF60EC35" author="Orr" pageId="7" pageNumber="208" refString="Orr, A. G. (2001) An annotated checklist of the Odonata of Brunei with ecological notes and descriptions of hitherto unknown males and larvae. International Journal of Odonatology, 4, 167 - 220. https: // doi. org / 10.1080 / 13887890.2001.9748168" type="journal article" year="2001">Orr 2001</bibRefCitation>
|
||
: 199
|
||
</treatmentCitation>
|
||
(females,
|
||
<collectingCountry id="F37D588A9855FF8DFE6A8B1AFE43EC35" box="[407,478,825,847]" name="Brunei" pageId="7" pageNumber="208">Brunei</collectingCountry>
|
||
);—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFDFE8B1AFD17EC35" author="Orr" box="[515,650,825,847]" page="40" pageId="7" pageNumber="208" year="2003">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFDFE8B1AFDF9EC35" author="Orr" box="[515,612,825,847]" pageId="7" pageNumber="208" refString="Orr, A. G. (2003) A guide to the dragonflies of Borneo: their identification and biology. Natural History Publications (Borneo), Kota Kinabalu, x + 195 pp." type="book" year="2003">Orr 2003</bibRefCitation>
|
||
: 40
|
||
</treatmentCitation>
|
||
, 96 (discussed).
|
||
<emphasis id="B91EC4089855FF8DFCCF8B1BFC0AEC34" bold="true" box="[818,919,824,847]" pageId="7" pageNumber="208">Syn. Nov.</emphasis>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9855FF8DFF6A8B55FC23ECF6" blockId="7.[151,958,886,908]" box="[151,958,886,908]" pageId="7" pageNumber="208">
|
||
<treatmentCitationGroup id="AB7A3F349855FF8DFF6A8B55FC23ECF6" box="[151,958,886,908]" pageId="7" pageNumber="208">
|
||
<taxonomicName id="4C6A63999855FF8DFF6A8B55FEB1ECF6" ID-CoL="5D3K" authorityName="Selys in Borneo" authorityYear="1878" box="[151,300,886,908]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="208" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EC4089855FF8DFF6A8B55FEB1ECF6" box="[151,300,886,908]" italics="true" pageId="7" pageNumber="208">Leptogomphus</emphasis>
|
||
</taxonomicName>
|
||
spec. A;—
|
||
<treatmentCitation id="0ACB3E0B9855FF8DFE668B55FDD2ECF6" author="Steinhoff" box="[411,591,886,908]" page="8" pageId="7" pageNumber="208" year="2015">
|
||
<bibRefCitation id="EFFB65EB9855FF8DFE668B55FDA4ECF6" author="Steinhoff" box="[411,569,886,908]" pageId="7" pageNumber="208" refString="Steinhoff, P. O. M. (2015) Results of Odonata larval rearing in the Gunung Mulu National Park, Sarawak, Malaysia from April to August 2014. International Dragonfly Fund Report, 78, 1 - 11." type="journal article" year="2015">Steinhoff 2015</bibRefCitation>
|
||
: 8
|
||
</treatmentCitation>
|
||
(female reared from larva, GMNP).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B919855FF8DFF6A8B9AFD41E968" pageId="7" pageNumber="208" type="materials_examined">
|
||
<paragraph id="8BD5181A9855FF8DFF6A8B9AFDAEEBD3" blockId="7.[151,1437,953,2017]" pageId="7" pageNumber="208">
|
||
<materialsCitation id="3B0212479855FF8DFF6A8B9AFCC2EB63" ID-GBIF-Occurrence="1702634979" collectingDate="1932-04-03" collectionCode="RMNH" collectorName="L. C. de Ruiter" country="Indonesia" county="Pandjoa" latitude="0.75725" location="Material" longLatPrecision="5" longitude="109.1381" municipality="Montrado" pageId="7" pageNumber="208" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan Barat" typeStatus="holotype">
|
||
<emphasis id="B91EC4089855FF8DFF6A8B9AFDBAECA8" bold="true" box="[151,551,953,978]" pageId="7" pageNumber="208">
|
||
<location id="8EB54EC19855FF8DFF6A8B9AFF62ECA8" LSID="urn:lsid:plazi:treatment:03C3A90C9855FF87FF6A88FDFC0EEE3A:8EB54EC19855FF8DFF6A8B9AFF62ECA8" box="[151,255,953,978]" country="Indonesia" county="Pandjoa" latitude="0.75725" longLatPrecision="5" longitude="109.1381" municipality="Montrado" name="Material" pageId="7" pageNumber="208" stateProvince="Kalimantan Barat">Material</location>
|
||
studied.
|
||
<typeStatus id="54D1A6B89855FF8DFE848B9AFE29ECA8" box="[377,436,953,978]" pageId="7" pageNumber="208">Type</typeStatus>
|
||
material
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="54D1A6B89855FF8DFDC18B9AFD34ECA8" box="[572,681,953,978]" pageId="7" pageNumber="208" type="holotype">
|
||
<emphasis id="B91EC4089855FF8DFDC18B9AFD34ECA8" bold="true" box="[572,681,953,978]" pageId="7" pageNumber="208">Holotype</emphasis>
|
||
</typeStatus>
|
||
<specimenCount id="9D6CD3939855FF8DFD4B8B99FD7AECAB" box="[694,743,954,978]" pageId="7" pageNumber="208" type="male">1 ♂</specimenCount>
|
||
(JvT number 3460),
|
||
<location id="8EB54EC19855FF8DFC0D8B9AFB65ECA8" LSID="urn:lsid:plazi:treatment:03C3A90C9855FF87FF6A88FDFC0EEE3A:8EB54EC19855FF8DFC0D8B9AFB65ECA8" box="[1008,1272,953,978]" country="Indonesia" county="Pandjoa" latitude="0.75725" longLatPrecision="5" longitude="109.1381" municipality="Montrado" name="Pakmiongtheo-Pandjoa" pageId="7" pageNumber="208" stateProvince="Kalimantan Barat">Pakmiongtheo-Pandjoa</location>
|
||
,
|
||
<location id="8EB54EC19855FF8DFAF68B9AFA05ECA8" LSID="urn:lsid:plazi:treatment:03C3A90C9855FF87FF6A88FDFC0EEE3A:8EB54EC19855FF8DFAF68B9AFA05ECA8" box="[1291,1432,953,978]" country="Indonesia" county="Pandjoa" latitude="0.75725" longLatPrecision="5" longitude="109.1381" municipality="Montrado" name="Singkawang" pageId="7" pageNumber="208" stateProvince="Kalimantan Barat">Singkawang</location>
|
||
,
|
||
<collectingRegion id="49AED6F89855FF8DFF6A8BFEFEFCEC8F" box="[151,353,989,1014]" country="Indonesia" name="Kalimantan Barat" pageId="7" pageNumber="208">Kalimantan Barat</collectingRegion>
|
||
,
|
||
<collectingRegion id="49AED6F89855FF8DFE938BFEFE69EC8C" box="[366,500,989,1014]" country="Indonesia" name="Kalimantan Utara" pageId="7" pageNumber="208">Kalimantan</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37D588A9855FF8DFDFD8BFEFDEDEC8C" box="[512,624,989,1014]" name="Indonesia" pageId="7" pageNumber="208">Indonesia</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFD818BFEFD73EC8C" box="[636,750,989,1014]" direction="north" orientation="latitude" pageId="7" pageNumber="208" precision="1" value="0.75725">0.75725N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFD078BFDFCE9EC8C" box="[762,884,990,1014]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="5" value="109.1381">109.1381E</geoCoordinate>
|
||
(coordinates for
|
||
<collectingMunicipality id="6BB182609855FF8DFBC58BFEFB3BEC8C" box="[1080,1190,989,1014]" pageId="7" pageNumber="208">Montrado</collectingMunicipality>
|
||
which is supposed to be close to
|
||
<collectingCounty id="62B460969855FF8DFEE48C22FEE5EB60" box="[281,376,1025,1050]" pageId="7" pageNumber="208">Pandjoa</collectingCounty>
|
||
),
|
||
<collectingDate id="EF90C7329855FF8DFE718C21FE66EB60" box="[396,507,1025,1050]" pageId="7" pageNumber="208" value="1932-04-03">
|
||
<date id="FFD43EDA9855FF8DFE718C21FE66EB60" box="[396,507,1025,1050]" pageId="7" pageNumber="208" value="1932-04-03">3 iv 1932</date>
|
||
</collectingDate>
|
||
, leg.
|
||
<collectorName id="269F7DCC9855FF8DFDC48C21FD40EB60" box="[569,733,1025,1050]" pageId="7" pageNumber="208">L.C. de Ruiter</collectorName>
|
||
, in
|
||
<collectionCode id="ED7B80DF9855FF8DFCFA8C21FCC2EB63" box="[775,863,1026,1049]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="208">RMNH</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B0212479855FF8DFC978C22FC05EB1B" ID-GBIF-Occurrence="1702634977" collectingDate="1937-06" collectionCode="RMNH" collectorName="M. E. Walsh" country="Indonesia" county="Sangkulirang" latitude="0.977" location="Other" longLatPrecision="55" longitude="117.9803" municipality="Sangkulirang District" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur" typeStatus="holotype">
|
||
<emphasis id="B91EC4089855FF8DFC978C22FBFFEB60" bold="true" box="[874,1122,1025,1050]" pageId="7" pageNumber="208">
|
||
<location id="8EB54EC19855FF8DFC978C22FC2FEB60" LSID="urn:lsid:plazi:treatment:03C3A90C9855FF87FF6A88FDFC0EEE3A:8EB54EC19855FF8DFC978C22FC2FEB60" box="[874,946,1025,1050]" country="Indonesia" county="Sangkulirang" latitude="0.977" longLatPrecision="55" longitude="117.9803" municipality="Sangkulirang District" name="Other" pageId="7" pageNumber="208" stateProvince="Kalimantan Timur">Other</location>
|
||
<typeStatus id="54D1A6B89855FF8DFC478C22FC73EB60" box="[954,1006,1025,1050]" pageId="7" pageNumber="208">type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<specimenCount id="9D6CD3939855FF8DFB978C21FB08EB63" box="[1130,1173,1026,1050]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
,
|
||
<typeStatus id="54D1A6B89855FF8DFB598C22FA9BEB60" box="[1188,1286,1025,1050]" pageId="7" pageNumber="208" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6A63999855FF8DFAD38C21FA01EB63" authorityName="Lieftinck" authorityYear="1948" box="[1326,1436,1026,1049]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC4089855FF8DFAD38C21FA01EB63" box="[1326,1436,1026,1049]" italics="true" pageId="7" pageNumber="208">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
(JvT number 3463),
|
||
<location id="8EB54EC19855FF8DFE628C05FD8EEB44" LSID="urn:lsid:plazi:treatment:03C3A90C9855FF87FF6A88FDFC0EEE3A:8EB54EC19855FF8DFE628C05FD8EEB44" box="[415,531,1062,1086]" country="Indonesia" county="Sangkulirang" latitude="0.977" longLatPrecision="55" longitude="117.9803" municipality="Sangkulirang District" name="Daguanan" pageId="7" pageNumber="208" stateProvince="Kalimantan Timur">Daguanan</location>
|
||
,
|
||
<collectingMunicipality id="6BB182609855FF8DFDD78C06FCB5EB44" box="[554,808,1061,1086]" pageId="7" pageNumber="208">Sangkulirang District</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49AED6F89855FF8DFCC38C06FB87EB44" box="[830,1050,1061,1086]" country="Indonesia" name="Kalimantan Timur" pageId="7" pageNumber="208">Kalimantan Timur</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37D588A9855FF8DFBCD8C06FB3DEB44" box="[1072,1184,1061,1086]" name="Indonesia" pageId="7" pageNumber="208">Indonesia</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFB4A8C05FA90EB44" box="[1207,1293,1062,1086]" direction="north" orientation="latitude" pageId="7" pageNumber="208" precision="55" value="0.977">0.977N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFADE8C05FA01EB44" box="[1315,1436,1062,1086]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="5" value="117.9803">117.9803E</geoCoordinate>
|
||
(coordinates for
|
||
<collectingCounty id="62B460969855FF8DFEAF8C6AFE74EB18" box="[338,489,1097,1122]" pageId="7" pageNumber="208">Sangkulirang</collectingCounty>
|
||
)
|
||
<collectingDate id="EF90C7329855FF8DFE058C6AFDCDEB18" box="[504,592,1097,1122]" pageId="7" pageNumber="208" value="1937-06">
|
||
<date id="FFD43EDA9855FF8DFE058C6AFDCDEB18" box="[504,592,1097,1122]" pageId="7" pageNumber="208" value="1937-06">vi 1937</date>
|
||
</collectingDate>
|
||
, leg.
|
||
<collectorName id="269F7DCC9855FF8DFD738C69FC8BEB18" box="[654,790,1097,1122]" pageId="7" pageNumber="208">M.E. Walsh</collectorName>
|
||
, in
|
||
<collectionCode id="ED7B80DF9855FF8DFCBD8C69FC05EB1B" box="[832,920,1098,1121]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="208">RMNH</collectionCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFC598C69FDB2EBD3" ID-GBIF-Occurrence="1702634981" collectingDate="1933-06-10" collectionCode="RMNH" collectorName="L. C. de Ruiter" country="Indonesia" latitude="0.841492" location="Gunung Poteng" longLatPrecision="1" longitude="109.056145" municipality="Sungai Bagak" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur" typeStatus="allotype">
|
||
<specimenCount id="9D6CD3939855FF8DFC598C69FC52EB1B" box="[932,975,1098,1122]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
,
|
||
<typeStatus id="54D1A6B89855FF8DFC238C6AFBA5EB18" box="[990,1080,1097,1122]" pageId="7" pageNumber="208" type="allotype">allotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6A63999855FF8DFBA28C69FEB8EBFC" authority=", Sungai Bagak, Singkawang" authorityName="Sungai Bagak, Singkawang" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC4089855FF8DFBA28C69FB75EB1B" box="[1119,1256,1098,1121]" italics="true" pageId="7" pageNumber="208">L. coomansi</emphasis>
|
||
, Sungai Bagak, Singkawang
|
||
</taxonomicName>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFED28C4DFE32EBFC" box="[303,431,1134,1158]" direction="north" orientation="latitude" pageId="7" pageNumber="208" precision="1" value="0.841492">0.841492N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFE448C4EFDCDEBFC" box="[441,592,1133,1158]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="1" value="109.056145">109.056146E</geoCoordinate>
|
||
(coordinates for
|
||
<location id="8EB54EC19855FF8DFCF38C4DFC23EBFC" LSID="urn:lsid:plazi:treatment:03C3A90C9855FF87FF6A88FDFC0EEE3A:8EB54EC19855FF8DFCF38C4DFC23EBFC" box="[782,958,1134,1158]" country="Indonesia" latitude="0.841492" longLatPrecision="1" longitude="109.056145" municipality="Sungai Bagak" name="Gunung Poteng" pageId="7" pageNumber="208" stateProvince="Kalimantan Timur">Gunung Poteng</location>
|
||
,
|
||
<collectingMunicipality id="6BB182609855FF8DFC348C4EFBF8EBFC" box="[969,1125,1133,1158]" pageId="7" pageNumber="208">Sungai Bagak</collectingMunicipality>
|
||
assumed near to this),
|
||
<collectingDate id="EF90C7329855FF8DFA998C4DFF4FEBD0" pageId="7" pageNumber="208" value="1933-06-10">
|
||
<date id="FFD43EDA9855FF8DFA998C4DFF4FEBD0" pageId="7" pageNumber="208" value="1933-06-10">10 vi 1933</date>
|
||
</collectingDate>
|
||
, leg.
|
||
<collectorName id="269F7DCC9855FF8DFEF08CB1FE32EBD0" box="[269,431,1169,1194]" pageId="7" pageNumber="208">L.C. de Ruiter</collectorName>
|
||
, in
|
||
<collectionCode id="ED7B80DF9855FF8DFE2A8CB1FDB2EBD3" box="[471,559,1170,1193]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="208">RMNH</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9855FF8DFF3A8C96FB36EBB7" blockId="7.[151,1437,953,2017]" box="[199,1195,1205,1230]" pageId="7" pageNumber="208">
|
||
<materialsCitation id="3B0212479855FF8DFF3A8C96FB36EBB7" ID-GBIF-Occurrence="1702634967" box="[199,1195,1205,1230]" collectingDate="1937-06" collectionCode="RMNH" collectorName="M. E. Walsh" country="Indonesia" county="Sangkulirang" latitude="0.977" location="Other" longLatPrecision="55" longitude="117.9803" municipality="Sangkulirang District" pageId="7" pageNumber="208" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan Barat">
|
||
<emphasis id="B91EC4089855FF8DFF3A8C96FEE7EBB4" bold="true" box="[199,378,1205,1230]" pageId="7" pageNumber="208">
|
||
<location id="8EB54EC19855FF8DFF3A8C96FE92EBB4" LSID="urn:lsid:plazi:treatment:03C3A90C9855FF87FF6A88FDFC0EEE3A:8EB54EC19855FF8DFF3A8C96FE92EBB4" box="[199,271,1205,1230]" country="Indonesia" name="Other" pageId="7" pageNumber="208" stateProvince="Kalimantan Barat">Other</location>
|
||
material
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B91EC4089855FF8DFE758C96FD72EBB4" bold="true" box="[392,751,1205,1230]" pageId="7" pageNumber="208">
|
||
<collectingRegion id="49AED6F89855FF8DFE758C96FDF9EBB4" box="[392,612,1205,1230]" country="Indonesia" name="Kalimantan Barat" pageId="7" pageNumber="208">Kalimantan Barat</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37D588A9855FF8DFD8C8C96FD77EBB4" box="[625,746,1205,1230]" name="Indonesia" pageId="7" pageNumber="208">Indonesia</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D6CD3939855FF8DFD0B8C95FCBFEBB7" box="[758,802,1206,1230]" pageId="7" pageNumber="208" type="male">1 ♂</specimenCount>
|
||
, same data as holotype, in
|
||
<collectionCode id="ED7B80DF9855FF8DFBB28C95FB3AEBB7" box="[1103,1191,1206,1229]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="208">RMNH</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9855FF8DFF3A8CFAFD41E968" blockId="7.[151,1437,953,2017]" pageId="7" pageNumber="208">
|
||
<materialsCitation id="3B0212479855FF8DFF3A8CFAFBB4EA6C" ID-GBIF-Occurrence="1702634965" collectingDate="1956-09-01" collectionCode="RMNH" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="3" specimenCount-female="3" stateProvince="Kalimantan Timur">
|
||
<emphasis id="B91EC4089855FF8DFF3A8CFAFDB2EB88" bold="true" box="[199,559,1241,1266]" pageId="7" pageNumber="208">
|
||
<collectingRegion id="49AED6F89855FF8DFF3A8CFAFE30EB88" box="[199,429,1241,1266]" country="Indonesia" name="Kalimantan Timur" pageId="7" pageNumber="208">Kalimantan Timur</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37D588A9855FF8DFE468CFAFDB2EB88" box="[443,559,1241,1266]" name="Indonesia" pageId="7" pageNumber="208">Indonesia</collectingCountry>
|
||
</emphasis>
|
||
(in
|
||
<collectionCode id="ED7B80DF9855FF8DFD9E8CF9FD2AEB8B" box="[611,695,1242,1265]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="208">RMNH</collectionCode>
|
||
unless explicitly otherwise noted):
|
||
<specimenCount id="9D6CD3939855FF8DFBA98CF9FB0AEB8B" box="[1108,1175,1242,1266]" pageId="7" pageNumber="208" type="female">3 ♀♀</specimenCount>
|
||
, Tabang, Kali Bengin,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFF6A8CDEFF66EA6C" box="[151,251,1277,1302]" direction="north" orientation="latitude" pageId="7" pageNumber="208" precision="5" value="0.5667">0.5667N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFEFB8CDEFEE2EA6C" box="[262,383,1277,1302]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="5" value="116.0333">116.0333E</geoCoordinate>
|
||
(approximate coordinates for Tabang),
|
||
<elevation id="0047FF299855FF8DFCC08CDEFCE2EA6C" box="[829,895,1277,1302]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="7" pageNumber="208" unit="m" value="125.0">
|
||
<quantity id="4C92B5FF9855FF8DFCC08CDEFCE2EA6C" box="[829,895,1277,1302]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="7" pageNumber="208" unit="m" value="125.0">125m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EF90C7329855FF8DFC778CDDFC65EA6C" box="[906,1016,1277,1302]" pageId="7" pageNumber="208" value="1956-09-01">
|
||
<date id="FFD43EDA9855FF8DFC778CDDFC65EA6C" box="[906,1016,1277,1302]" pageId="7" pageNumber="208" value="1956-09-01">1 ix 1956</date>
|
||
</collectingDate>
|
||
, leg
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B0212479855FF8DFBCE8CDDFECEEA40" ID-GBIF-Occurrence="1702634964" collectingDate="1956-09-02" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur">
|
||
<collectorName id="269F7DCC9855FF8DFBCE8CDDFB74EA6C" box="[1075,1257,1278,1302]" pageId="7" pageNumber="208">A.M.R. Wegner</collectorName>
|
||
;
|
||
<specimenCount id="9D6CD3939855FF8DFB098CDDFA82EA6F" box="[1268,1311,1278,1302]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFF1B8D01FECEEA40" box="[230,339,1313,1338]" pageId="7" pageNumber="208" value="1956-09-02">
|
||
<date id="FFD43EDA9855FF8DFF1B8D01FECEEA40" box="[230,339,1313,1338]" pageId="7" pageNumber="208" value="1956-09-02">2 ix 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFEA38D01FD63EA40" ID-GBIF-Occurrence="1702634966" box="[350,766,1313,1338]" collectingDate="1956-09-03" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFEA38D01FE17EA43" box="[350,394,1314,1338]" pageId="7" pageNumber="208" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D6CD3939855FF8DFE688D01FE22EA43" box="[405,447,1314,1338]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFD6C8D01FD63EA40" box="[657,766,1313,1338]" pageId="7" pageNumber="208" value="1956-09-03">
|
||
<date id="FFD43EDA9855FF8DFD6C8D01FD63EA40" box="[657,766,1313,1338]" pageId="7" pageNumber="208" value="1956-09-03">3 ix 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFCF48D01FBEFEA40" ID-GBIF-Occurrence="1702634968" box="[777,1138,1313,1338]" collectingDate="1956-09-04" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFCF48D01FCAEEA43" box="[777,819,1314,1338]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFBF88D01FBEFEA40" box="[1029,1138,1313,1338]" pageId="7" pageNumber="208" value="1956-09-04">
|
||
<date id="FFD43EDA9855FF8DFBF88D01FBEFEA40" box="[1029,1138,1313,1338]" pageId="7" pageNumber="208" value="1956-09-04">4 ix 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFB808D01FF72EA24" ID-GBIF-Occurrence="1702634986" collectingDate="1956-09-08" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="3" specimenCount-female="3" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFB808D01FB20EA43" box="[1149,1213,1314,1338]" pageId="7" pageNumber="208" type="female">3 ♀♀</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFA738D01FF72EA24" pageId="7" pageNumber="208" value="1956-09-08">
|
||
<date id="FFD43EDA9855FF8DFA738D01FF72EA24" pageId="7" pageNumber="208" value="1956-09-08">8 ix 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFF068D65FDE8EA24" ID-GBIF-Occurrence="1702634963" box="[251,629,1349,1374]" collectingDate="1956-09-10" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFF068D65FEBAEA27" box="[251,295,1350,1374]" pageId="7" pageNumber="208" type="male">1 ♂</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFE078D65FDE8EA24" box="[506,629,1349,1374]" pageId="7" pageNumber="208" value="1956-09-10">
|
||
<date id="FFD43EDA9855FF8DFE078D65FDE8EA24" box="[506,629,1349,1374]" pageId="7" pageNumber="208" value="1956-09-10">10 ix 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFD7C8D65FC61EA24" ID-GBIF-Occurrence="1702634978" box="[641,1020,1349,1374]" collectingDate="1956-09-18" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFD7C8D65FD31EA27" box="[641,684,1350,1374]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFC7C8D65FC61EA24" box="[897,1020,1349,1374]" pageId="7" pageNumber="208" value="1956-09-18">
|
||
<date id="FFD43EDA9855FF8DFC7C8D65FC61EA24" box="[897,1020,1349,1374]" pageId="7" pageNumber="208" value="1956-09-18">18 ix 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFBF58D65FA1FEA24" ID-GBIF-Occurrence="1702634983" box="[1032,1410,1349,1374]" collectingDate="1956-09-21" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFBF58D65FBAFEA27" box="[1032,1074,1350,1374]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFAFA8D65FA1FEA24" box="[1287,1410,1349,1374]" pageId="7" pageNumber="208" value="1956-09-21">
|
||
<date id="FFD43EDA9855FF8DFAFA8D65FA1FEA24" box="[1287,1410,1349,1374]" pageId="7" pageNumber="208" value="1956-09-21">21 ix 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFA738D65FD9CEAF8" ID-GBIF-Occurrence="1702634984" collectingDate="1956-09-26" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFA738D65FF33EAFB" pageId="7" pageNumber="208" type="male">1 ♂</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFE798D4AFD9CEAF8" box="[388,513,1385,1410]" pageId="7" pageNumber="208" value="1956-09-26">
|
||
<date id="FFD43EDA9855FF8DFE798D4AFD9CEAF8" box="[388,513,1385,1410]" pageId="7" pageNumber="208" value="1956-09-26">26 ix 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFDF38D49FC12EAF8" ID-GBIF-Occurrence="1702634972" box="[526,911,1385,1410]" collectingDate="1956-09-28" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFDF38D49FDA4EAFB" box="[526,569,1386,1410]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFCEF8D49FC12EAF8" box="[786,911,1385,1410]" pageId="7" pageNumber="208" value="1956-09-28">
|
||
<date id="FFD43EDA9855FF8DFCEF8D49FC12EAF8" box="[786,911,1385,1410]" pageId="7" pageNumber="208" value="1956-09-28">28 ix 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFC618D49FA80EAF8" ID-GBIF-Occurrence="1702634976" box="[924,1309,1385,1410]" collectingDate="1956-09-29" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFC618D49FC5AEAFB" box="[924,967,1386,1410]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFB628D49FA80EAF8" box="[1183,1309,1385,1410]" pageId="7" pageNumber="208" value="1956-09-29">
|
||
<date id="FFD43EDA9855FF8DFB628D49FA80EAF8" box="[1183,1309,1385,1410]" pageId="7" pageNumber="208" value="1956-09-29">29 ix 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFAD48D49FE0AEADC" ID-GBIF-Occurrence="1702634980" collectingDate="1956-09-30" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFAD48D49FAC9EAFB" box="[1321,1364,1386,1410]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFEE18DADFE0AEADC" box="[284,407,1421,1446]" pageId="7" pageNumber="208" value="1956-09-30">
|
||
<date id="FFD43EDA9855FF8DFEE18DADFE0AEADC" box="[284,407,1421,1446]" pageId="7" pageNumber="208" value="1956-09-30">30 ix 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFE5F8DADFC84EADC" ID-GBIF-Occurrence="1702634973" box="[418,793,1421,1446]" collectingDate="1956-10-02" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="2" specimenCount-female="2" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFE5F8DADFE7FEADF" box="[418,482,1422,1446]" pageId="7" pageNumber="208" type="female">2 ♀♀</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFD498DADFC84EADC" box="[692,793,1421,1446]" pageId="7" pageNumber="208" value="1956-10-02">
|
||
<date id="FFD43EDA9855FF8DFD498DADFC84EADC" box="[692,793,1421,1446]" pageId="7" pageNumber="208" value="1956-10-02">2 x 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFCD98DADFB1AEADC" ID-GBIF-Occurrence="1702634985" box="[804,1159,1421,1446]" collectingDate="1956-10-05" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFCD98DADFCD3EADF" box="[804,846,1422,1446]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFBDC8DAEFB1AEADC" box="[1057,1159,1421,1446]" pageId="7" pageNumber="208" value="1956-10-05">
|
||
<date id="FFD43EDA9855FF8DFBDC8DAEFB1AEADC" box="[1057,1159,1421,1446]" pageId="7" pageNumber="208" value="1956-10-05">5 x 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFB6C8DADFF75EAB0" ID-GBIF-Occurrence="1702634982" collectingDate="1956-10-06" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFB6C8DADFB26EADF" box="[1169,1211,1422,1446]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFA738DAEFF75EAB0" pageId="7" pageNumber="208" value="1956-10-06">
|
||
<date id="FFD43EDA9855FF8DFA738DAEFF75EAB0" pageId="7" pageNumber="208" value="1956-10-06">6 x 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFF098D91FD34EAB0" ID-GBIF-Occurrence="1702634971" box="[244,681,1457,1482]" collectingDate="1956-10-07" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFF098D91FEBDEAB3" box="[244,288,1458,1482]" pageId="7" pageNumber="208" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D6CD3939855FF8DFED08D91FEF0EAB3" box="[301,365,1458,1482]" pageId="7" pageNumber="208" type="female">2 ♀♀</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFDBF8D91FD34EAB0" box="[578,681,1457,1482]" pageId="7" pageNumber="208" value="1956-10-07">
|
||
<date id="FFD43EDA9855FF8DFDBF8D91FD34EAB0" box="[578,681,1457,1482]" pageId="7" pageNumber="208" value="1956-10-07">7 x 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFD488D91FA82EAB3" ID-GBIF-Occurrence="1702634974" box="[693,1311,1457,1482]" collectingDate="1956-09-06" collectionCode="BMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFD488D91FD42EAB3" box="[693,735,1458,1482]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, same location and collector,
|
||
<collectingDate id="EF90C7329855FF8DFBCD8D92FB03EAB0" box="[1072,1182,1457,1482]" pageId="7" pageNumber="208" value="1956-09-06">
|
||
<date id="FFD43EDA9855FF8DFBCD8D92FB03EAB0" box="[1072,1182,1457,1482]" pageId="7" pageNumber="208" value="1956-09-06">6 ix 1956</date>
|
||
</collectingDate>
|
||
, in
|
||
<collectionCode id="ED7B80DF9855FF8DFB3A8D91FA82EAB3" box="[1223,1311,1458,1481]" pageId="7" pageNumber="208">BMNH</collectionCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFAD78D91FD9DEA97" ID-GBIF-Occurrence="1702634991" collectingDate="1956-10-07" collectionCode="BMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFAD78D91FACBEAB3" box="[1322,1366,1458,1482]" pageId="7" pageNumber="208" type="male">1 ♂</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFEE18DF5FE1CEA94" box="[284,385,1493,1518]" pageId="7" pageNumber="208" value="1956-10-07">
|
||
<date id="FFD43EDA9855FF8DFEE18DF5FE1CEA94" box="[284,385,1493,1518]" pageId="7" pageNumber="208" value="1956-10-07">7 x 1956</date>
|
||
</collectingDate>
|
||
, in
|
||
<collectionCode id="ED7B80DF9855FF8DFE548DF5FD9DEA97" box="[425,512,1494,1517]" pageId="7" pageNumber="208">BMNH</collectionCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFDF68DF5FCF3EA94" ID-GBIF-Occurrence="1702634989" box="[523,878,1493,1518]" collectingDate="1956-10-09" collectionCode="BMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFDF68DF5FDAAEA97" box="[523,567,1494,1518]" pageId="7" pageNumber="208" type="male">1 ♂</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFCF48DF5FCF3EA94" box="[777,878,1493,1518]" pageId="7" pageNumber="208" value="1956-10-09">
|
||
<date id="FFD43EDA9855FF8DFCF48DF5FCF3EA94" box="[777,878,1493,1518]" pageId="7" pageNumber="208" value="1956-10-09">9 x 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFC848DF5FB77EA94" ID-GBIF-Occurrence="1702634988" box="[889,1258,1493,1518]" collectingDate="1956-10-17" collectionCode="BMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFC848DF5FC38EA97" box="[889,933,1494,1518]" pageId="7" pageNumber="208" type="male">1 ♂</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFB8A8DF5FB77EA94" box="[1143,1258,1493,1518]" pageId="7" pageNumber="208" value="1956-10-17">
|
||
<date id="FFD43EDA9855FF8DFB8A8DF5FB77EA94" box="[1143,1258,1493,1518]" pageId="7" pageNumber="208" value="1956-10-17">17 x 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFB0B8DF5FEC6E968" ID-GBIF-Occurrence="1702634990" collectingDate="1956-10-24" collectionCode="BMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFB0B8DF5FABCEA97" box="[1270,1313,1494,1518]" pageId="7" pageNumber="208" type="male">1 ♂</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFF1A8DD9FEC6E968" box="[231,347,1529,1554]" pageId="7" pageNumber="208" value="1956-10-24">
|
||
<date id="FFD43EDA9855FF8DFF1A8DD9FEC6E968" box="[231,347,1529,1554]" pageId="7" pageNumber="208" value="1956-10-24">24 x 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0212479855FF8DFE9B8DD9FD4AE968" ID-GBIF-Occurrence="1702634994" box="[358,727,1529,1554]" collectingDate="1956-10-27" collectionCode="BMNH" collectorName="A. M. R. Wegner" country="Indonesia" elevation="125" latitude="0.5667" location="Kalimantan Timur" longLatPrecision="7" longitude="116.0333" pageId="7" pageNumber="208" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur">
|
||
<specimenCount id="9D6CD3939855FF8DFE9B8DD9FE0DE96B" box="[358,400,1530,1554]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, same data except
|
||
<collectingDate id="EF90C7329855FF8DFD998DD9FD4AE968" box="[612,727,1529,1554]" pageId="7" pageNumber="208" value="1956-10-27">
|
||
<date id="FFD43EDA9855FF8DFD998DD9FD4AE968" box="[612,727,1529,1554]" pageId="7" pageNumber="208" value="1956-10-27">27 x 1956</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B919855FF86FF3A8E3EFD4BEAE6" lastPageId="12" lastPageNumber="213" pageId="7" pageNumber="208" type="description">
|
||
<paragraph id="8BD5181A9855FF8DFF3A8E3EFF4EE907" blockId="7.[151,1437,953,2017]" pageId="7" pageNumber="208">
|
||
<emphasis id="B91EC4089855FF8DFF3A8E3EFDD7E94C" bold="true" box="[199,586,1565,1590]" pageId="7" pageNumber="208">
|
||
<collectingRegion id="49AED6F89855FF8DFF3A8E3EFE26E94C" box="[199,443,1565,1590]" country="Indonesia" name="Kalimantan Selatan" pageId="7" pageNumber="208">Kalimantan Selatan</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37D588A9855FF8DFE318E3EFDD8E94C" box="[460,581,1565,1590]" name="Indonesia" pageId="7" pageNumber="208">Indonesia</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D6CD3939855FF8DFDA88E3DFD1EE94F" box="[597,643,1566,1590]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, Sungai Kalaan,
|
||
<quantity id="4C92B5FF9855FF8DFCAD8E3DFC10E94C" box="[848,909,1565,1590]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="208" unit="km" value="3.0">3 km</quantity>
|
||
SE of Belangian village, Riam Kanan Lake, Aranio District, Banjarbaru, -
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFE178E62FDCDE920" box="[490,592,1601,1626]" direction="north" orientation="latitude" pageId="7" pageNumber="208" precision="5" value="3.5059">3.5059N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFDA28E62FD4AE920" box="[607,727,1601,1626]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="5" value="115.0923">115.0923E</geoCoordinate>
|
||
(Riam Kanan),
|
||
<quantity id="4C92B5FF9855FF8DFC6B8E61FC44E920" box="[918,985,1602,1626]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="208" unit="m" value="100.0">100m</quantity>
|
||
,
|
||
<date id="FFD43EDA9855FF8DFC158E62FBF4E920" box="[1000,1129,1601,1626]" pageId="7" pageNumber="208" value="1996-11-15">15 xi 1996</date>
|
||
, leg. M. Bedjanic, in coll. Dow.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9855FF82FF3A8EAAFED4EAD6" blockId="7.[151,1437,953,2017]" lastBlockId="8.[151,1437,151,2021]" lastPageId="8" lastPageNumber="209" pageId="7" pageNumber="208">
|
||
<emphasis id="B91EC4089855FF8DFF3A8EAAFE21E9D8" bold="true" box="[199,444,1673,1698]" pageId="7" pageNumber="208">
|
||
<collectingRegion id="49AED6F89855FF8DFF3A8EAAFEAEE9D8" box="[199,307,1673,1698]" country="Malaysia" name="Sarawak" pageId="7" pageNumber="208">
|
||
<collectingCountry id="F37D588A9855FF8DFF3A8EAAFEAEE9D8" box="[199,307,1673,1698]" name="Malaysia" pageId="7" pageNumber="208">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37D588A9855FF8DFEB88EAAFE2BE9D8" box="[325,438,1673,1698]" name="Malaysia" pageId="7" pageNumber="208">Malaysia</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D6CD3939855FF8DFE378EA9FE61E9DB" box="[458,508,1674,1698]" pageId="7" pageNumber="208" type="male">1 ♂</specimenCount>
|
||
, Sungai Assam, Samunsam Wildlife Sanctuary, Kuching Division,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFADB8EAAFA05E9D8" box="[1318,1432,1673,1698]" direction="north" orientation="latitude" pageId="7" pageNumber="208" precision="1" value="1.92705">1.92705N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFF6A8E8FFEBCE9BF" box="[151,289,1708,1733]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="1" value="109.60133">109.60133E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA9855FF8DFED38E8EFE21E9BF" box="[302,444,1708,1733]" pageId="7" pageNumber="208" value="2015-08-18">18 viii 2015</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD3939855FF8DFCA58E8EFC1EE9BE" box="[856,899,1709,1733]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, last stream on Summit Trail, Gunung Gading, Gunung Gading NP, Kuching Division,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFDA28EF3FD4CE992" box="[607,721,1744,1768]" direction="north" orientation="latitude" pageId="7" pageNumber="208" precision="1" value="1.70242">1.70242N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFD238EF3FCFBE993" box="[734,870,1744,1769]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="1" value="109.83627">109.83627E</geoCoordinate>
|
||
(Gunung Gading),
|
||
<date id="FFD43EDA9855FF8DFBBB8EF3FB5BE993" box="[1094,1222,1744,1769]" pageId="7" pageNumber="208" value="2016-08-03">3 viii 2016</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD3939855FF8DFEA98ED7FEE2E871" box="[340,383,1780,1804]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, Tebang, Kuching Division,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFD348ED0FCA7E876" box="[713,826,1779,1804]" direction="north" orientation="latitude" pageId="7" pageNumber="208" precision="1" value="1.11905">1.11905N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFCBB8ED7FC50E876" box="[838,973,1780,1804]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="1" value="110.22793">110.22793E</geoCoordinate>
|
||
(Gunung Penrissen, Tebang supposed to be somewhere on this mountain),
|
||
<date id="FFD43EDA9855FF8DFDE88F3BFD1FE84A" box="[533,642,1815,1840]" pageId="7" pageNumber="208" value="1958-09-07">7 ix 1958</date>
|
||
, leg. T.C. Maa, in RMNH;
|
||
<specimenCount id="9D6CD3939855FF8DFC4A8F34FC7CE855" box="[951,993,1815,1839]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, same data except
|
||
<date id="FFD43EDA9855FF8DFB458F34FABBE84A" box="[1208,1318,1815,1840]" pageId="7" pageNumber="208" value="1958-09-08">8 ix 1958</date>
|
||
;
|
||
<specimenCount id="9D6CD3939855FF8DFACF8F34FAC3E855" box="[1330,1374,1815,1839]" pageId="7" pageNumber="208" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D6CD3939855FF8DFA968F34FA08E855" box="[1387,1429,1815,1839]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
, streams on Belian trail, Kubah NP, Kuching Division,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFD008F19FCCFE829" box="[765,850,1850,1875]" direction="north" orientation="latitude" pageId="7" pageNumber="208" precision="55" value="1.617">1.617N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFCA28F18FC51E829" box="[863,972,1851,1875]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="55" value="110.192">110.192E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA9855FF8DFC258F19FBC9E829" box="[984,1108,1850,1875]" pageId="7" pageNumber="208" value="2008-09-15">15 ix 2008</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD3939855FF8DFF238F7CFE97E80C" box="[222,266,1887,1911]" pageId="7" pageNumber="208" type="female">1 ♀</specimenCount>
|
||
(SAR11_12_GOM48, RMNH.INS.506897, used for description and illustrations), perched on road up Gunung Serapi, same NP,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFE198FA2FDA7E8E0" box="[484,570,1921,1946]" direction="north" orientation="latitude" pageId="7" pageNumber="208" precision="55" value="1.613">1.613N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFDB28FA1FD27E8E0" box="[591,698,1922,1946]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="55" value="110.197">110.197E</geoCoordinate>
|
||
(Kubah NP),
|
||
<date id="FFD43EDA9855FF8DFC898FA1FC69E8E0" box="[884,1012,1921,1946]" pageId="7" pageNumber="208" value="2012-09-03">3 ix 2012</date>
|
||
, leg. R.A. Dow, in RMNH;
|
||
<specimenCount id="9D6CD3939855FF8DFA958FA1FA01E8E3" box="[1384,1436,1922,1946]" pageId="7" pageNumber="208" type="male">1 ♂</specimenCount>
|
||
(SAR09_10_GOM4, RMNH.INS.503526, used for description below and illustrations), ponded section of small stream very near confluence with Sungai Rayu, Matang Wildlife Centre, Kuching Division,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFB318FEBFABFE89B" box="[1228,1314,1992,2017]" direction="north" orientation="latitude" pageId="7" pageNumber="208" precision="55" value="1.609">1.609N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9855FF8DFACC8FEBFA01E89B" box="[1329,1436,1992,2017]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="55" value="110.161">110.161E</geoCoordinate>
|
||
(Matang Wildlife Centre),
|
||
<date id="FFD43EDA985AFF82FE3788BBFDA6EFCA" box="[458,571,151,176]" pageId="8" pageNumber="209" value="2009-09-09">9 ix 2009</date>
|
||
, leg. R.A. Dow, in RMNH;
|
||
<specimenCount id="9D6CD393985AFF82FC7988BBFC2CEFD5" box="[900,945,152,176]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
(RMNH.INS.506264), same location,
|
||
<date id="FFD43EDA985AFF82FA9288BBFF4FEFAE" pageId="8" pageNumber="209" value="2012-04-01">1 iv 2012</date>
|
||
, leg. R.A. Dow, in RMNH;
|
||
<specimenCount id="9D6CD393985AFF82FDD8889FFDC9EFA9" box="[549,596,188,212]" pageId="8" pageNumber="209" type="female">1 ♀</specimenCount>
|
||
, on road to Hindu Temple on Mount Matang, Matang Range, Kuching Division,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FEFA88FDFEE7EF8D" box="[263,378,222,247]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="1" value="1.59632">1.59632N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FE7B88FCFD90EF8D" box="[390,525,223,247]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="1" value="110.20949">110.20949E</geoCoordinate>
|
||
(Mount Matang),
|
||
<date id="FFD43EDA985AFF82FD1F88FDFCC7EF8D" box="[738,858,222,247]" pageId="8" pageNumber="209" value="2015-07-05">5 vii 2015</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FB0488FCFAB8EF8D" box="[1273,1317,223,247]" pageId="8" pageNumber="209" type="female">1 ♀</specimenCount>
|
||
, stream at Ranchan Recreational Park, Serian, Serian Division,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FD078920FCDFEE61" box="[762,834,259,283]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="555" value="1.14">1.14N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FCAD8921FC23EE61" box="[848,958,258,283]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="55" value="110.581">110.581E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FC318920FBDBEE61" box="[972,1094,258,283]" pageId="8" pageNumber="209" value="2008-02-14">14 ii 2008</date>
|
||
, leg. G.T. Reels, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FF1D8905FE93EE44" box="[224,270,294,318]" pageId="8" pageNumber="209" type="female">1 ♀</specimenCount>
|
||
, Sungai Kepayang, Ulu Sebuyau, Samarahan Division,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FC608905FB92EE44" box="[925,1039,294,318]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="1" value="1.27712">1.27712N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FBE38905FB35EE44" box="[1054,1192,294,318]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="1" value="110.94812">110.94812E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FB4A8905FADCEE44" box="[1207,1345,293,318]" pageId="8" pageNumber="209" value="2015-07-10">10 vii 2015</date>
|
||
, leg. L. Southwell, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FE0D896AFD86EE1B" box="[496,539,329,353]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
(SAR07_8_GOM18, used in
|
||
<figureCitation id="1351049F985AFF82FC90896AFC5DEE1B" box="[877,960,329,354]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="8" pageNumber="209">Fig. 94</figureCitation>
|
||
), tributary to Sungai Bebiong Mit, Batang Ai NP, Sri Aman Division,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FE28894EFDB6EEFF" box="[469,555,365,389]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="55" value="1.202">1.202N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FDC5894FFD3EEEFF" box="[568,675,364,389]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="55" value="112.058">112.058E</geoCoordinate>
|
||
(approximate coordinates in Batang Ai NP),
|
||
<date id="FFD43EDA985AFF82FB52894FFAB5EEFF" box="[1199,1320,364,389]" pageId="8" pageNumber="209" value="2007-12-05">5 xii 2007</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FE4889B3FE7FEED2" box="[437,482,400,424]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, same location and date, leg. G.T. Reels, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FB6D89B3FB23EED2" box="[1168,1214,400,424]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, Sungai Segak Mit, Ulu Mujok area, inside LEWS, Sarikei Division,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FD3D8990FCAEEEB6" box="[704,819,435,460]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="1" value="1.70636">1.70636N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FCC38997FC55EEB6" box="[830,968,436,460]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="1" value="112.08934">112.08934E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FC2E8997FBCBEEB6" box="[979,1110,435,460]" pageId="8" pageNumber="209" value="2015-07-30">30 vii 2015</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FF2389F4FEBFEE95" box="[222,290,471,495]" pageId="8" pageNumber="209" type="male">2 ♂♂</specimenCount>
|
||
, Sungai Segak Besai, same area,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FD5B89F4FC85EE8A" box="[678,792,471,496]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="1" value="1.68253">1.68253N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FCD889F4FC32EE8A" box="[805,943,471,496]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="1" value="112.15868">112.15868E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FC4189F4FBD1EE8A" box="[956,1100,471,496]" pageId="8" pageNumber="209" value="2016-08-14">14 viii 2016</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FF2089D8FE95ED68" box="[221,264,507,531]" pageId="8" pageNumber="209" type="female">1 ♀</specimenCount>
|
||
(teneral), Sungai Mujok, same area,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FD5389D9FC8FED69" box="[686,786,506,531]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="5" value="1.6825">1.6825N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FCE389D9FC38ED69" box="[798,933,506,531]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="1" value="112.15791">112.15791E</geoCoordinate>
|
||
(Sungai Mujok at LEWS boundary),
|
||
<date id="FFD43EDA985AFF82FAAE89D9FF4FED4C" pageId="8" pageNumber="209" value="2016-08-15">15 viii 2016</date>
|
||
, leg. L. Southwell, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FD7A8A3DFD2CED4C" box="[647,689,542,566]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
(teneral), Sungai Selabi, same area,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FBB38A3EFB2FED4C" box="[1102,1202,541,566]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="5" value="1.6929">1.6929N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FB438A3EFADAED4C" box="[1214,1351,541,566]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="1" value="112.16291">112.16291E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FAAE8A3EFF4FED20" pageId="8" pageNumber="209" value="2016-08-16">16 viii 2016</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FD998A61FD13ED23" box="[612,654,578,602]" pageId="8" pageNumber="209" type="female">1 ♀</specimenCount>
|
||
, Sungai Tekalit, same area,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FC368A62FBA0ED20" box="[971,1085,577,602]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="1" value="1.67589">1.67589N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FBB58A61FB4FED20" box="[1096,1234,578,602]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="1" value="112.17488">112.17488E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FB208A61FAF5ED20" box="[1245,1384,577,602]" pageId="8" pageNumber="209" value="2016-08-17">17 viii 2016</date>
|
||
, leg. B. Megong & N. Mengiring, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FD448A46FD79ED07" box="[697,740,613,637]" pageId="8" pageNumber="209" type="female">1 ♀</specimenCount>
|
||
, Sungai Sengkadan, same area,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FBB38A46FB2FED04" box="[1102,1202,613,638]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="5" value="1.6817">1.6817N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FB438A46FAD5ED04" box="[1214,1352,613,638]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="1" value="112.15192">112.15192E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FAA98A46FF4FEDDB" pageId="8" pageNumber="209" value="2016-08-17">17 viii 2016</date>
|
||
, leg. R.W.J. Ngiam, to be deposited in ZRC;
|
||
<specimenCount id="9D6CD393985AFF82FD1C8AAAFC90EDDA" box="[737,781,649,673]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
(teneral), same location,
|
||
<date id="FFD43EDA985AFF82FBC98AAAFB5EEDDB" box="[1076,1219,648,673]" pageId="8" pageNumber="209" value="2016-08-18">18 viii 2016</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FEAD8A8FFEE1EDBE" box="[336,380,684,708]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, Sungai Temurok, same area but outside LEWS boundary,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FBF18A8FFBE3EDBF" box="[1036,1150,684,709]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="1" value="1.69165">1.69165N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FB748A8FFA8FEDBE" box="[1161,1298,684,708]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="1" value="112.12991">112.12991E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FAE08A8FFA05EDBF" box="[1309,1432,684,709]" pageId="8" pageNumber="209" value="2015-08-04">4 viii 2015</date>
|
||
, leg. N. Masil, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FDE18AF3FDD7ED9D" box="[540,586,720,744]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, Sungai Lingga, same area,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FC6C8AECFB9EED92" box="[913,1027,719,744]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="1" value="1.70369">1.70369N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FBED8AF3FB04ED92" box="[1040,1177,720,744]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="1" value="112.11807">112.11807E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FB5A8AECFAB5ED92" box="[1191,1320,719,744]" pageId="8" pageNumber="209" value="2015-08-05">5 viii 2015</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FE528AD0FE47EC71" box="[431,474,755,779]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, Sungai Mujok, same area,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FCF08AD0FC1DEC76" box="[781,896,755,780]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="1" value="1.71235">1.71235N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FC778AD0FB8CEC71" box="[906,1041,755,779]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="1" value="112.08381">112.08381E</geoCoordinate>
|
||
(coordinates by Sungai Mujok),
|
||
<date id="FFD43EDA985AFF82FA7D8AD0FF60EC55" pageId="8" pageNumber="209" value="2016-08-17">17 viii 2016</date>
|
||
(after 6 pm), leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FCC98B34FCFFEC54" box="[820,866,791,815]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, Sungai Sbong, Kapit Town area, Kapit Division,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FF6A8B19FF70EC28" box="[151,237,826,850]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="55" value="2.009">2.009N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FF078B19FEFBEC28" box="[250,358,826,850]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="55" value="113.118">113.118E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FE8E8B19FE77EC28" box="[371,490,825,850]" pageId="8" pageNumber="209" value="2008-02-11">11 ii 2008</date>
|
||
, leg. G,T. Reels, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FC748B19FC28EC2B" box="[905,949,826,850]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
(RMNH.INS.509952), stream in disturbed forest, foot of Hose Mountains, Tunoh district, Kapit Division,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FCA28B7EFC5EEC0C" box="[863,963,861,886]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="5" value="2.0698">2.0698N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FC338B7EFBD7EC0C" box="[974,1098,861,886]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="5" value="113.6474">113.6474E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FBA88B7EFB54EC0C" box="[1109,1225,861,886]" pageId="8" pageNumber="209" value="2010-05-16">16 v 2010</date>
|
||
, leg. G.T. Reels, in RMNH;
|
||
<specimenCount id="9D6CD393985AFF82FF018BA2FEBAECE2" box="[252,295,897,921]" pageId="8" pageNumber="209" type="female">1 ♀</specimenCount>
|
||
(RMNH.INS.509950), tributary to Sungai Sii, Upper Baram, Miri Division,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FB6A8BA2FA95ECE3" box="[1175,1288,897,921]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="1" value="2.99113">2.99113N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FAE88BA3FA01ECE3" box="[1301,1436,896,921]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="1" value="114.90581">114.90581E</geoCoordinate>
|
||
(Sungai Sii),
|
||
<date id="FFD43EDA985AFF82FED18B87FE33ECC7" box="[300,430,932,957]" pageId="8" pageNumber="209" value="2014-07-16">16 vii 2014</date>
|
||
, leg. R.A. Dow, in RMNH;
|
||
<specimenCount id="9D6CD393985AFF82FD158B86FC89ECC6" box="[744,788,933,957]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, stream on Oil Well trail, Lambir Hills NP, Miri Division,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FF6A8BEBFF66EC9A" box="[151,251,968,992]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="5" value="4.2004">4.2004N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FEFB8BEBFE1CEC9A" box="[262,385,968,992]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="5" value="114.0303">114.0303E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FE708BEBFD9BEC9A" box="[397,518,967,992]" pageId="8" pageNumber="209" value="2011-04-22">22 iv 2011</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FC6B8BEBFC5DEC9A" box="[918,960,968,992]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
(RMNH.INS.506807), tributary to Sungai Liku, extension area to same NP,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FDEC8BC8FD1FEB79" box="[529,642,1003,1027]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="1" value="4.23313">4.23313N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FD728BC8FC85EB7E" box="[655,792,1003,1028]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="1" value="114.06127">114.06127E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FCD98BC8FC35EB79" box="[804,936,1003,1028]" pageId="8" pageNumber="209" value="2012-07-16">16 vii 2012</date>
|
||
, leg. L. Southwell, in RMNH;
|
||
<specimenCount id="9D6CD393985AFF82FAF98BC8FAB2EB79" box="[1284,1327,1003,1027]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
(teneral),
|
||
<specimenCount id="9D6CD393985AFF82FF6A8C2CFF5CEB5C" box="[151,193,1039,1063]" pageId="8" pageNumber="209" type="female">1 ♀</specimenCount>
|
||
, Sungai Melinau, GMNP, Miri Division,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FD658C2DFD66EB5D" box="[664,763,1038,1063]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="5" value="4.0559">4.0559N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FCF58C2DFC1EEB5D" box="[776,899,1038,1063]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="5" value="114.8257">114.8257E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FC6D8C2DFB9BEB5D" box="[912,1030,1038,1063]" pageId="8" pageNumber="209" value="2006-02-16">16 ii 2006</date>
|
||
, leg. J. Simun, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FA738C2CFF31EB30" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
(teneral), Bat Observatory Stream, same NP,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FD528C11FC9BEB30" box="[687,774,1074,1098]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="55" value="4.024">4.024N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FCEC8C11FCE2EB30" box="[785,895,1074,1098]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="55" value="114.819">114.819E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FC768C11FB93EB30" box="[907,1038,1074,1099]" pageId="8" pageNumber="209" value="2007-12-28">28 xii 2007</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FF6A8C75FF5CEB17" box="[151,193,1110,1134]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
(teneral), same location,
|
||
<date id="FFD43EDA985AFF82FE1C8C75FDFBEB14" box="[481,614,1109,1134]" pageId="8" pageNumber="209" value="2007-12-30">30 xii 2007</date>
|
||
; leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FC008C75FBB4EB17" box="[1021,1065,1110,1134]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, same location,
|
||
<date id="FFD43EDA985AFF82FB1D8C75FAC4EB14" box="[1248,1369,1109,1134]" pageId="8" pageNumber="209" value="2014-04-21">21 iv 2014</date>
|
||
(6:15 pm), leg. P.O.M. Steinhoff, in collection Steinhoff;
|
||
<specimenCount id="9D6CD393985AFF82FD238C5AFCBDEBEB" box="[734,800,1145,1170]" pageId="8" pageNumber="209" type="male">5 ♂♂</specimenCount>
|
||
, same location,
|
||
<date id="FFD43EDA985AFF82FC248C5AFBD6EBEB" box="[985,1099,1145,1169]" pageId="8" pageNumber="209" value="2014-05-22">22 v 2014</date>
|
||
(6 pm and after), leg. P.O.M. Steinhoff, in collection Steinhoff;
|
||
<specimenCount id="9D6CD393985AFF82FDE48CBEFDDBEBCE" box="[537,582,1181,1205]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, Sungai Paku, same area,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FC8C8CBEFC5AEBCF" box="[881,967,1181,1205]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="55" value="4.048">4.048N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FC2E8CBFFBA2EBCF" box="[979,1087,1180,1205]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="55" value="114.836">114.836E</geoCoordinate>
|
||
(coordinates on Sungai Paku),
|
||
<date id="FFD43EDA985AFF82FF6A8CE3FE8EEBA2" box="[151,275,1215,1240]" pageId="8" pageNumber="209" value="2014-04-21">21 iv 2014</date>
|
||
(6:15 pm), leg. P.O.M. Steinhoff, in collection Steinhoff;
|
||
<specimenCount id="9D6CD393985AFF82FC538CE3FC41EBA2" box="[942,988,1216,1240]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, second stream on old trail to
|
||
<collectingRegion id="49AED6F8985AFF82FAC18C9CFA01EBA2" box="[1340,1436,1215,1240]" country="Malaysia" name="Sarawak" pageId="8" pageNumber="209">
|
||
<collectingCountry id="F37D588A985AFF82FAC18C9CFA01EBA2" box="[1340,1436,1215,1240]" name="Malaysia" pageId="8" pageNumber="209">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
Chamber,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FEF08CC0FEFEEB86" box="[269,355,1251,1276]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="55" value="4.059">4.059N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FE8D8CC0FE43EB86" box="[368,478,1251,1276]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="55" value="114.861">114.861E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA985AFF82FE118CC7FDC0EB86" box="[492,605,1251,1276]" pageId="8" pageNumber="209" value="2008-01-13">13 i 2008</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FC028CC7FBB0EB81" box="[1023,1069,1252,1276]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, open area on summit trail near Camp 1, same NP,
|
||
<date id="FFD43EDA985AFF82FE938D24FE77EA65" box="[366,490,1286,1311]" pageId="8" pageNumber="209" value="2008-09-13">13 ix 2008</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FC7D8D24FC30EA65" box="[896,941,1287,1311]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, Lubang
|
||
<collectingCountry id="F37D588A985AFF82FBEA8D25FBD2EA65" box="[1047,1103,1286,1311]" name="China" pageId="8" pageNumber="209">Cina</collectingCountry>
|
||
, Headhunters Trail, same NP, Limbang Division,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FE8E8D08FE54EA39" box="[371,457,1323,1347]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="55" value="4.198">4.198N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FE288D09FDA2EA39" box="[469,575,1322,1347]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="55" value="114.895">114.895E</geoCoordinate>
|
||
(approximate coordinates on Headhunters Trail),
|
||
<date id="FFD43EDA985AFF82FB8E8D08FB75EA39" box="[1139,1256,1322,1347]" pageId="8" pageNumber="209" value="2006-02-11">11 ii 2006</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FE918D6DFE05EA1C" box="[364,408,1358,1382]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, same location,
|
||
<date id="FFD43EDA985AFF82FDB68D6DFD23EA1C" box="[587,702,1357,1382]" pageId="8" pageNumber="209" value="2006-02-14">14 ii 2006</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393985AFF82FBB68D6DFBEBEA1C" box="[1099,1142,1358,1382]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, Mentawei boundary trail, same national park and Division,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FDD78D51FD1DEAF0" box="[554,640,1394,1418]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="55" value="4.239">4.239N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FD6D8D51FD66EAF0" box="[656,763,1394,1418]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="55" value="114.877">114.877E</geoCoordinate>
|
||
(approximate coordinates),
|
||
<date id="FFD43EDA985AFF82FBBA8D51FB5BEAF0" box="[1095,1222,1393,1418]" pageId="8" pageNumber="209" value="2006-02-13">13 ii 2006</date>
|
||
, leg. J. Simun, in collection Dow.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A985AFF82FF3A8D9BFDB9E9DF" blockId="8.[151,1437,151,2021]" pageId="8" pageNumber="209">
|
||
<emphasis id="B91EC408985AFF82FF3A8D9BFEBEEAAB" bold="true" box="[199,291,1464,1489]" pageId="8" pageNumber="209">
|
||
<collectingCountry id="F37D588A985AFF82FF3A8D9BFE83EAAB" box="[199,286,1464,1489]" name="Brunei" pageId="8" pageNumber="209">Brunei</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D6CD393985AFF82FED78D9BFEC9EAAA" box="[298,340,1464,1488]" pageId="8" pageNumber="209" type="female">1 ♀</specimenCount>
|
||
, Sungai Ingei,
|
||
<collectingRegion id="49AED6F8985AFF82FE028D9BFD07EAAB" box="[511,666,1464,1489]" country="Brunei" name="Belait" pageId="8" pageNumber="209">Belait district</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FD588D9BFC94EAAB" box="[677,777,1464,1489]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="5" value="4.1535">4.1535N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FCE98D9BFC10EAAA" box="[788,909,1464,1488]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="5" value="114.7201">114.7201E</geoCoordinate>
|
||
(Sungai Ingei),
|
||
<date id="FFD43EDA985AFF82FBB98D9BFB22EAAA" box="[1092,1215,1464,1489]" pageId="8" pageNumber="209" value="1992-09-30">30 ix 1992</date>
|
||
, leg. D. Thompson, in RMNH;
|
||
<specimenCount id="9D6CD393985AFF82FEE08DFFFED7EA89" box="[285,330,1500,1524]" pageId="8" pageNumber="209" type="female">1 ♀</specimenCount>
|
||
, track in freshwater swamp forest, along Ingei river, surroundings Ingei base camp, same area,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FF6A8DDCFF66E962" box="[151,251,1535,1560]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="5" value="4.1535">4.1535N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FEF48E23FE1FE962" box="[265,386,1536,1560]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="5" value="114.7201">114.7201E</geoCoordinate>
|
||
(Sungai Ingei), same date and collector, in BMNH;
|
||
<specimenCount id="9D6CD393985AFF82FC178E23FB84E96D" box="[1002,1049,1536,1560]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
, same location and collector,
|
||
<date id="FFD43EDA985AFF82FA8B8E23FF4FE941" pageId="8" pageNumber="209" value="1992-10-02">2 x 1992</date>
|
||
, in RMNH;
|
||
<specimenCount id="9D6CD393985AFF82FEA68E00FE19E940" box="[347,388,1571,1595]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
(RMNH.5008295), camp at Sungai Ingei,
|
||
<collectingRegion id="49AED6F8985AFF82FC9F8E01FC61E941" box="[866,1020,1570,1595]" country="Brunei" name="Belait" pageId="8" pageNumber="209">Belait district</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FBFB8E01FBF6E941" box="[1030,1131,1570,1595]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="5" value="4.1535">4.1535N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FB888E00FB73E941" box="[1141,1262,1571,1595]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="5" value="114.7201">114.7201E</geoCoordinate>
|
||
(Sungai Ingei),
|
||
<date id="FFD43EDA985AFF82FF6A8E64FE92E925" box="[151,271,1606,1631]" pageId="8" pageNumber="209" value="2014-02-24">24 ii 2014</date>
|
||
, leg. R.A. Dow;
|
||
<specimenCount id="9D6CD393985AFF82FE298E64FD9DE924" box="[468,512,1607,1631]" pageId="8" pageNumber="209" type="male">1 ♂</specimenCount>
|
||
(RMNH.5008363), same location,
|
||
<date id="FFD43EDA985AFF82FC688E64FB8BE925" box="[917,1046,1606,1631]" pageId="8" pageNumber="209" value="2014-03-10">10 iii 2014</date>
|
||
, leg. R.A. Dow, in RMNH;
|
||
<specimenCount id="9D6CD393985AFF82FAA68E65FA01E924" box="[1371,1436,1606,1631]" pageId="8" pageNumber="209" type="male">5 ♂♂</specimenCount>
|
||
(including RMNH.INS.500716), Sungai Lumut,
|
||
<collectingRegion id="49AED6F8985AFF82FD438E4AFCC7E9F8" box="[702,858,1641,1666]" country="Brunei" name="Belait" pageId="8" pageNumber="209">Belait district</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FC9B8E4AFC56E9F8" box="[870,971,1641,1666]" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="5" value="4.6167">4.6167N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD985AFF82FC2A8E4AFBCDE9F8" box="[983,1104,1641,1666]" direction="east" orientation="longitude" pageId="8" pageNumber="209" precision="5" value="114.5167">114.5167E</geoCoordinate>
|
||
(Sungai Lumut),
|
||
<date id="FFD43EDA985AFF82FAE68E49FA05E9F8" box="[1307,1432,1641,1666]" pageId="8" pageNumber="209" value="2004-11-17">17 xi 2004</date>
|
||
, leg. V.J. Kalkman
|
||
<emphasis id="B91EC408985AFF82FE988EADFE00E9DC" box="[357,413,1677,1702]" italics="true" pageId="8" pageNumber="209">et al.</emphasis>
|
||
, in RMNH.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A985AFF82FF3A8E93FE57E89F" blockId="8.[151,1437,151,2021]" pageId="8" pageNumber="209">
|
||
<emphasis id="B91EC408985AFF82FF3A8E93FB27E9B3" bold="true" box="[199,1210,1712,1737]" pageId="8" pageNumber="209">Description of male (based on SAR09_10_GOM4, RMNH.INS.503526 Kubah NP).</emphasis>
|
||
<emphasis id="B91EC408985AFF82FB388E93FA9FE9B3" box="[1221,1282,1712,1737]" italics="true" pageId="8" pageNumber="209">Head</emphasis>
|
||
(
|
||
<figureCitation id="1351049F985AFF82FAE98E93FAFDE9B3" box="[1300,1376,1712,1737]" captionStart="FIGURES 4–9" captionStartId="9.[151,264,1713,1736]" captionTargetBox="[151,1436,193,1692]" captionTargetId="figure@9.[151,1436,193,1692]" captionTargetPageId="9" captionText="FIGURES 4–9. Male head dorsal: (4) L. coomansi SAR09_10_GOM4; (5) L. pendleburyi SAR13_14_GOM13; (6) L. pasia SAB12_GOM1; (7) L. sii holotype; (8) L.schieli holotype; (9) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068004/files/figure.png" pageId="8" pageNumber="209">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F985AFF82FA8D8E92FA12E9B3" box="[1392,1423,1713,1737]" captionStart="FIGURES10–13" captionStartId="14.[151,264,1864,1887]" captionTargetBox="[151,1436,772,1843]" captionTargetId="figure@14.[151,1436,772,1843]" captionTargetPageId="14" captionText="FIGURES10–13. Male head frontal: (10) L.coomansi SAR09_10_GOM4; (11) L.coomansi holotype; (12) L.sp. cf coomansi SAB12_GOM4; (13) L.pendleburyi SAR13_14_GOM13." httpUri="https://zenodo.org/record/1068006/files/figure.png" pageId="8" pageNumber="209">10</figureCitation>
|
||
). Median lobe labium black, lateral lobes pale, hooks black. Labrum black with pale central transverse basal mark, not quite divided centrally. Mandible bases pale, genae brownish adjacent to mandible bases, rest black. Anteclypeus black. Postclypeus mostly black with pale basal transverse mark occupying most of width, in centre extending almost to anteclypeus. Ante- and postfrons not sharply divided, whole mostly greenish, very narrowly divided by black centrally, black to rear postfrons (
|
||
<figureCitation id="1351049F985AFF82FD268F42FCBFE800" box="[731,802,1889,1914]" captionStart="FIGURES 4–9" captionStartId="9.[151,264,1713,1736]" captionTargetBox="[151,1436,193,1692]" captionTargetId="figure@9.[151,1436,193,1692]" captionTargetPageId="9" captionText="FIGURES 4–9. Male head dorsal: (4) L. coomansi SAR09_10_GOM4; (5) L. pendleburyi SAR13_14_GOM13; (6) L. pasia SAB12_GOM1; (7) L. sii holotype; (8) L.schieli holotype; (9) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068004/files/figure.png" pageId="8" pageNumber="209">Fig. 4</figureCitation>
|
||
). Vertex and occiput black. Pair of ridge-like transverse tubercles behind lateral ocelli, narrowly divided centrally and extending almost to compound eye margins, remainder of vertex smooth, ocelli orange. Free margin of occipital plate raised to form narrow ridge, narrowly extended forward centrally.
|
||
</paragraph>
|
||
<caption id="DF154892985BFF83FF6A8E92FB18E99C" httpUri="https://zenodo.org/record/1068004/files/figure.png" pageId="9" pageNumber="210" targetBox="[151,1436,193,1692]" targetPageId="9">
|
||
<paragraph id="8BD5181A985BFF83FF6A8E92FB18E99C" blockId="9.[151,1436,1713,1766]" pageId="9" pageNumber="210">
|
||
<emphasis id="B91EC408985BFF83FF6A8E92FEA1E9BC" bold="true" box="[151,316,1713,1736]" pageId="9" pageNumber="210">FIGURES 4–9.</emphasis>
|
||
Male head dorsal: (4)
|
||
<taxonomicName id="4C6A6399985BFF83FDD58E91FD38E9BD" box="[552,677,1714,1735]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985BFF83FDD58E91FD38E9BD" box="[552,677,1714,1735]" italics="true" pageId="9" pageNumber="210">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
SAR09_10_GOM4; (5)
|
||
<taxonomicName id="4C6A6399985BFF83FC5B8E91FBA1E9BD" authorityName="Laidlaw" authorityYear="1934" box="[934,1084,1713,1735]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="pendleburyi">
|
||
<emphasis id="B91EC408985BFF83FC5B8E91FBA1E9BD" box="[934,1084,1713,1735]" italics="true" pageId="9" pageNumber="210">L. pendleburyi</emphasis>
|
||
</taxonomicName>
|
||
SAR13_14_GOM13; (6)
|
||
<taxonomicName id="4C6A6399985BFF83FAB48E91FA01E9BD" authorityName="van Tol" authorityYear="1990" box="[1353,1436,1714,1735]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="pasia">
|
||
<emphasis id="B91EC408985BFF83FAB48E91FA01E9BD" box="[1353,1436,1714,1735]" italics="true" pageId="9" pageNumber="210">L. pasia</emphasis>
|
||
</taxonomicName>
|
||
SAB12_GOM1; (7)
|
||
<taxonomicName id="4C6A6399985BFF83FE9A8EF2FE07E99C" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[359,410,1745,1766]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="sii">
|
||
<emphasis id="B91EC408985BFF83FE9A8EF2FE07E99C" box="[359,410,1745,1766]" italics="true" pageId="9" pageNumber="210">L. sii</emphasis>
|
||
</taxonomicName>
|
||
holotype; (8)
|
||
<taxonomicName id="4C6A6399985BFF83FDD78EF2FD1BE99C" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[554,646,1744,1766]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="schieli">
|
||
<emphasis id="B91EC408985BFF83FDD78EF2FD1BE99C" box="[554,646,1744,1766]" italics="true" pageId="9" pageNumber="210">L. schieli</emphasis>
|
||
</taxonomicName>
|
||
holotype; (9)
|
||
<taxonomicName id="4C6A6399985BFF83FCEB8EF2FC3AE99C" authorityName="Laidlaw" authorityYear="1912" box="[790,935,1744,1766]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC408985BFF83FCEB8EF2FC3AE99C" box="[790,935,1744,1766]" italics="true" pageId="9" pageNumber="210">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
SAR11_12_GOM56.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD5181A985BFF80FF3A8F2EFC41EEAA" blockId="9.[151,1436,1805,2010]" lastBlockId="10.[151,1437,151,2013]" lastPageId="10" lastPageNumber="211" pageId="9" pageNumber="210">
|
||
<emphasis id="B91EC408985BFF83FF3A8F2EFE85E85C" box="[199,280,1805,1830]" italics="true" pageId="9" pageNumber="210">Thorax</emphasis>
|
||
. Prothorax dark brown, most of anterior pronotal lobe pale yellow, pale yellow transverse marking to rear of middle lobe, deepest laterally, and centrally where narrowly divided, a small isolated yellow spot centrally on dorsum. Posterior pronotal lobe with tiny central yellow spot. Propleuron mottled brow and black. Synthorax dark brown with pale yellow-green markings as follows (
|
||
<figureCitation id="1351049F985BFF83FCDF8F5AFCE7E8E8" box="[802,890,1913,1938]" captionStart="FIGURES 52–55" captionStartId="31.[151,264,1501,1524]" captionTargetBox="[151,1436,409,1479]" captionTargetId="figure@31.[151,1436,409,1480]" captionTargetPageId="31" captionText="FIGURES 52–55. Male synthorax dorsal: (52) L. coomansi SAR09_10_GOM4; (53) L. coomansi holotype; (54) L. sp. cf coomansi SAB12_GOM4; (55) L. pendleburyi SAR13_14_GOM13." httpUri="https://zenodo.org/record/1068024/files/figure.png" pageId="9" pageNumber="210">Figs 52</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F985BFF83FC7A8F5AFC3AE8E8" box="[903,935,1913,1938]" captionStart="FIGURES 60–63" captionStartId="34.[151,264,1601,1624]" captionTargetBox="[156,1432,517,1580]" captionTargetId="figure@34.[156,1432,517,1580]" captionTargetPageId="34" captionText="FIGURES 60–63. Male synthorax lateral: (60) L. coomansi SAR09_10_GOM4; (61) L. sp. cf coomansi SAB12_GOM4; (62) L. schieli holotype; (63) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068028/files/figure.png" pageId="9" pageNumber="210">60</figureCitation>
|
||
): short, narrow mesothoracic collar, broadly divided at middorsal carina, joined to narrow dorsal thoracic stripes that extend beyond level of apex of antealar crest, terminating just short of antealar carina, almost parallel on their inner margins. Small spot below dorsal thoracic stripe on mesepisternum, near antealar carina. Stripe running most of length of mesepimeron. Metepisternum with irregular stripe running from metakatepisternum, surrounding spiracle, narrowly separated from subcircular spot at antealar carina (
|
||
<figureCitation id="1351049F9858FF80FD5388FCFC8DEF82" box="[686,784,223,248]" captionStart="FIGURES 60–63" captionStartId="34.[151,264,1601,1624]" captionTargetBox="[156,1432,517,1580]" captionTargetId="figure@34.[156,1432,517,1580]" captionTargetPageId="34" captionText="FIGURES 60–63. Male synthorax lateral: (60) L. coomansi SAR09_10_GOM4; (61) L. sp. cf coomansi SAB12_GOM4; (62) L. schieli holotype; (63) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068028/files/figure.png" pageId="10" pageNumber="211">Fig. 60</figureCitation>
|
||
). A broad stripe occupying most of metepimeron. Mesokatepisternum dark brown and pale, metakatepisternum mostly pale. Venter pale. Legs (posterior pair removed for DNA extraction) robust and relatively short. Anterior and middle pairs with coxae mottled pale and dark grey, remainder mostly black. Wings: sectors of arculus separated at origin with 6 (right) or 5 (left) cross veins up to and at first bifurcation of superior sector in Fw,
|
||
<quantity id="4C92B5FF9858FF80FCFA8953FCA9EEF2" box="[775,820,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="211" unit="in" value="3.0">3 in</quantity>
|
||
Hw. Discoidal field with 2 rows of cells from origin, transitioning to three rows before level of nodus in both wings. 17 (left) or 18 (right) Ax in Fw, 12 (left) or 13 (right) in Hw, 12 Px in all wings. Pt brown, covering ca 5 underlying cells.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9858FF80FF3A89FFFA11EB4F" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="211">
|
||
<emphasis id="B91EC4089858FF80FF3A89FFFEACEE8F" box="[199,305,476,501]" italics="true" pageId="10" pageNumber="211">Abdomen</emphasis>
|
||
. Slender after base of S3, expanding moderately from base of S7, maximum width and height reached apical part of S8, then almost constant. Dorsum of S10 with pair of prominent ca conical protuberances basally (
|
||
<figureCitation id="1351049F9858FF80FF058A07FEC1ED47" box="[248,348,548,573]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="10" pageNumber="211">Fig. 106</figureCitation>
|
||
), bearing many small tubercles on posterior faces. Dark brown to black with pale markings as follows: S1 mostly pale laterally, pale and dark dorsally. S2 pale lateral stripe running from base below and including auricle, except free margin of auricle, another pale lateral mark placed posteriorly, narrow yellowish middorsal stripe. S3 with small basal lateral pale marking, S3–6 with very narrow and rather irregular middorsal yellow line. Cerci (
|
||
<figureCitation id="1351049F9858FF80FE908A97FE4CEDB6" box="[365,465,692,717]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="10" pageNumber="211">Figs 100</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9858FF80FE208A97FD95EDB7" box="[477,520,692,717]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="10" pageNumber="211">106</figureCitation>
|
||
) broad at base in lateral view (
|
||
<figureCitation id="1351049F9858FF80FCA28A97FC5DEDB7" box="[863,960,692,717]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="10" pageNumber="211">Fig. 106</figureCitation>
|
||
), tapering to sharp tip, along upper margin sloping down from base to just before mid-length, then more gently to apex, along lower margin sloping down from base to ca one-third length, then gently up to apex, small teeth present along much of lower margin. In dorsal view (
|
||
<figureCitation id="1351049F9858FF80FF228B3CFEDDEC42" box="[223,320,799,824]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="10" pageNumber="211">Fig. 100</figureCitation>
|
||
) relatively slender, outer margin curving slightly out from base, then running gently inward until just before apex, where turned slightly outward. Black, brown at base laterally, this more extensive dorsally. Epiproct (
|
||
<figureCitation id="1351049F9858FF80FEF78B44FEEEECFA" box="[266,371,871,896]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="10" pageNumber="211">Figs 100</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9858FF80FE7E8B4BFE33ECFA" box="[387,430,872,896]" captionStart="FIGURES 112–117" captionStartId="44.[151,264,1808,1831]" captionTargetBox="[156,1432,193,1787]" captionTargetId="figure@44.[156,1432,193,1787]" captionTargetPageId="44" captionText="FIGURES 112–117. Male anal appendages in ventral view (all scale bars 0.5mm): (112) L. coomansi SAR09_10_GOM4; (113) L.pasia SAB12_GOM1;(114) L.pendleburyi SAR13_14_GOM13; (115) L. schieli holotype; (116) L.sii holotype; (117) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068048/files/figure.png" pageId="10" pageNumber="211">112</figureCitation>
|
||
) black, and dark brown, just shorter than cerci, deeply divided in a “U” shape, outer margins gently convex, apices rounded. In lateral view (
|
||
<figureCitation id="1351049F9858FF80FCF68BAFFCF7ECDF" box="[779,874,908,933]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="10" pageNumber="211">Fig. 106</figureCitation>
|
||
) curved gently down then up. Accessory genitalia as shown in
|
||
<figureCitation id="1351049F9858FF80FED98B8CFE00ECB2" box="[292,413,943,968]" captionStart="FIGURES 88–93" captionStartId="40.[151,264,1280,1303]" captionTargetBox="[156,1432,195,1258]" captionTargetId="figure@40.[156,1432,195,1258]" captionTargetPageId="40" captionText="FIGURES 88–93. Male accessory genitalia, lateral view, all scale bars 1mm: (88) L. coomansi SAR09_10_GOM4; (89) L. pasia SAB12_GOM1; (90) L. pendleburyi SAR13_14_GOM13; (91) L. schieli holotype; (92) L. sii holotype; (93) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068040/files/figure.png" pageId="10" pageNumber="211">Figures 88</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9858FF80FE578B93FE55ECB2" box="[426,456,944,968]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="10" pageNumber="211">94</figureCitation>
|
||
, anterior hamule moderate sized, ca three-quarters height of posterior hamule, tapering toward rounded apex, hooked to rear in apical ca one-third. Posterior hamule large, ca parallel sided over much of length and directed postero-ventrally, curving gently toward sharp tip along posterior margin, abruptly hooked back to tip along front anterior margin, apex pointed slightly out. Penis missing (see below). Penis vesicle low, simple.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9858FF80FF3A8C63FBF6EB22" blockId="10.[151,1437,151,2013]" box="[199,1131,1087,1112]" pageId="10" pageNumber="211">Measurements (mm). Hw 28, abdomen excluding anal appendages 35.5, cerci ca 1.</paragraph>
|
||
<paragraph id="8BD5181A9858FF80FF3A8C47FE84EA2F" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="211">
|
||
<emphasis id="B91EC4089858FF80FF3A8C47FACBEB07" bold="true" box="[199,1366,1124,1149]" pageId="10" pageNumber="211">Description of female (based on SAR11_12_GOM48, RMNH.INS.506897 Kubah National Park).</emphasis>
|
||
<emphasis id="B91EC4089858FF80FAA28C47FA06EB07" box="[1375,1435,1124,1149]" italics="true" pageId="10" pageNumber="211">Head</emphasis>
|
||
(
|
||
<figureCitation id="1351049F9858FF80FF628CA4FF67EBDA" box="[159,250,1159,1184]" captionStart="FIGURES18–23" captionStartId="18.[151,264,1895,1918]" captionTargetBox="[183,1420,198,1847]" captionTargetId="figure@18.[183,1420,198,1847]" captionTargetPageId="18" captionText="FIGURES18–23.Female headdorsal:(18) L.coomansi SAR11_12_GOM48;(19)L.mariaeholotype;(20) L.sp.cf coomansi PomS260; (21) L. pasia holotype; (22) L.sii paratype SAR13_14_GOM44; (23) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068010/files/figure.png" pageId="10" pageNumber="211">Figs 18</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9858FF80FEF78CABFEB4EBDA" box="[266,297,1160,1184]" captionStart="FIGURES28–31" captionStartId="22.[151,264,1357,1380]" captionTargetBox="[151,1435,266,1336]" captionTargetId="figure@22.[151,1436,265,1336]" captionTargetPageId="22" captionText="FIGURES28–31. Female head dorsaldetail; h—occipital horn; ot—occipital tube; p—pit; s—occipital spur; t—tubercle;thtubercle horn: (28) L. coomansi SAR11_12_GOM48; (29) L. mariae holotype; (30) L. sp. cf coomansi PomS260; (31) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068014/files/figure.png" pageId="10" pageNumber="211">28</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9858FF80FEC58CA4FECAEBDA" box="[312,343,1159,1184]" captionStart="FIGURES 36–41" captionStartId="25.[151,264,1973,1996]" captionTargetBox="[204,1387,183,1958]" captionTargetId="figure@25.[204,1387,183,1958]" captionTargetPageId="25" captionText="FIGURES 36–41. Female head frontal: (36) L. coomansi SAR11_12_GOM48; (37) L. mariae holotype; (38) L. sp. cf coomansi PomS260; (39) L.pasia holotype; (40) L.sii paratype SAR13_14_GOM44; (41) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068018/files/figure.png" pageId="10" pageNumber="211">36</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9858FF80FE9A8CA4FE1AEBDA" box="[359,391,1159,1184]" captionStart="FIGURES 46–51" captionStartId="29.[151,264,1672,1695]" captionTargetBox="[156,1432,375,1651]" captionTargetId="figure@29.[156,1432,375,1651]" captionTargetPageId="29" captionText="FIGURES 46–51. Female head frontal detail; h—occipital horn; ot—occipital tube; t—tubercle; th—tubercle horn: (46) L. coomansi SAR11_12_GOM48; (47) L pendleburyi SAR16_GOM8; (48) L. schieli paratype SAR15_GOM6; (49) L. species SAR11_12_GOM33; (50) L. williamsoni SAR13_14_GOM51; (51) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068022/files/figure.png" pageId="10" pageNumber="211">46</figureCitation>
|
||
). Pale marks yellow. Labrum pale central transverse basal mark divided centrally. Genae yellowish adjacent to mandible bases. Postclypeus with additional pair of tiny yellow marks widely spaced near anteclypeus, that on the right very faint. Marking on post frons not divided centrally. On each side behind the ridgelike transverse tubercles and adjacent to the compound eyes is a distinctive pit. Occipital plate bearing pair of occipital horns placed near eye margins, centrally with shallow notch to rear, slightly raised immediately to each side of this.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9858FF80FF3A8D7CFDC0E92A" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="211">
|
||
<emphasis id="B91EC4089858FF80FF3A8D7CFE85EA02" box="[199,280,1375,1400]" italics="true" pageId="10" pageNumber="211">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="1351049F9858FF80FED48D7CFE1FEA02" box="[297,386,1375,1400]" captionStart="FIGURES 68–73" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[156,1432,193,1894]" captionTargetId="figure@36.[156,1432,193,1894]" captionTargetPageId="36" captionText="FIGURES 68–73. Female synthorax dorsal: (68) L. coomansi SAR11_12_GOM48; (69) L. mariae holotype; (70) L. sp. cf coomansi PomS260; (71) L.pasia holotype; (72) L.sii paratype SAR13_14_GOM44; (73) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068032/files/figure.png" pageId="10" pageNumber="211">Figs 68</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9858FF80FE6C8D43FE2CEA02" box="[401,433,1376,1400]" captionStart="FIGURES 78–83" captionStartId="38.[151,264,1952,1975]" captionTargetBox="[169,1406,182,1934]" captionTargetId="figure@38.[169,1406,182,1934]" captionTargetPageId="38" captionText="FIGURES 78–83. Female synthorax lateral: (78) L. coomansi SAR11_12_GOM48; (79) L. mariae holotype, right hand side, flipped for consistency; (80) L. sp. cf coomansi PomS260; (81) L. pasia holotype; (82) L. sii paratype SAR13_14_GOM44; (83) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068036/files/figure.png" pageId="10" pageNumber="211">78</figureCitation>
|
||
). No small isolated yellow spot centrally on dorsum of pronotal middle lobe. Posterior pronotal lobe almost entirely yellow. Stripe running most of length of mesepimeron partially and narrowly intruded by brown on left side (
|
||
<figureCitation id="1351049F9858FF80FE5F8D84FE64EABA" box="[418,505,1447,1472]" captionStart="FIGURES 78–83" captionStartId="38.[151,264,1952,1975]" captionTargetBox="[169,1406,182,1934]" captionTargetId="figure@38.[169,1406,182,1934]" captionTargetPageId="38" captionText="FIGURES 78–83. Female synthorax lateral: (78) L. coomansi SAR11_12_GOM48; (79) L. mariae holotype, right hand side, flipped for consistency; (80) L. sp. cf coomansi PomS260; (81) L. pasia holotype; (82) L. sii paratype SAR13_14_GOM44; (83) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068036/files/figure.png" pageId="10" pageNumber="211">Fig. 78</figureCitation>
|
||
). Mesokatepisternum dark brown with yellow triangular marking based by coxa. Venter pale. Legs: right middle removed for DNA extraction, left anterior detached below coxa, left middle detached below trochanter. A small, obscure pale area in the upper interior flexor surface of the anterior femur. Wings: 17 (right) or 19 (left) Ax in Fw, 12 (left) or (13) in Hw, 12 Px in all wings except left Fw, where 13. Pt brown, covering ca 4-5 underlying cells.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9858FF80FF3A8E7FFD0DE836" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="211">
|
||
<emphasis id="B91EC4089858FF80FF3A8E7FFEACE90F" box="[199,305,1628,1653]" italics="true" pageId="10" pageNumber="211">Abdomen</emphasis>
|
||
. Only expanding slightly from base of S7. S1 large lateral yellow mark, separated from broad dorsal mark. S2 pale lateral stripe running from base, including small rounded auricle, to posterior carina, broad dorsal stripe, widest at base. Middorsal line on S3–6 better developed than in male, but still narrow. Cerci conical, longer than S10, tapering to sharp tip, tips slightly divaricate, a short but prominent, rounded, epiproct in between. Vulvar scale (as in
|
||
<figureCitation id="1351049F9858FF80FEE18ECFFEE7E87E" box="[284,378,1772,1797]" captionStart="FIGURES 120–125" captionStartId="46.[151,264,1891,1914]" captionTargetBox="[151,1435,372,1869]" captionTargetId="figure@46.[151,1436,371,1870]" captionTargetPageId="46" captionText="FIGURES 120–125. Female vulvar scale (all scale bars 0.5mm): (120) L. coomansi SAR11_12_GOM48; (121) L. pasia holotype; (122) L. pendleburyi SAR13_14_GOM15; (123) L. schieli paratype SAR15_GOM6; (124) L. sii paratype SAR13_14_GOM44; (125) L.williamsoni SAR13_14_GOM51." httpUri="https://zenodo.org/record/1068052/files/figure.png" pageId="10" pageNumber="211">Fig. 120</figureCitation>
|
||
) ca two thirds length of S9, divided at between one third and one half of its length from base. The sternite of S8, is strongly depressed for a short distance at around two-thirds length, immediately after this there is a prominent tubercle (as in
|
||
<figureCitation id="1351049F9858FF80FDDE8F17FD1EE836" box="[547,643,1844,1869]" captionStart="FIGURES 118–119" captionStartId="45.[151,264,604,627]" captionTargetBox="[156,1432,193,583]" captionTargetId="figure@45.[156,1432,193,583]" captionTargetPageId="45" captionText="FIGURES 118–119. Ventral view female abdominal S8 (all scale bars 0.5mm): (118) L. coomansi SAR11_12_GOM48; (119) L. pasia SAB12_GOM2." httpUri="https://zenodo.org/record/1068050/files/figure.png" pageId="10" pageNumber="211">Fig. 118</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9858FF80FF3A8F7AFC6FE80A" blockId="10.[151,1437,151,2013]" box="[199,1010,1879,1904]" pageId="10" pageNumber="211">Measurements (mm). Hw 32, abdomen excluding anal appendages 36.5.</paragraph>
|
||
<paragraph id="8BD5181A9858FF81FF3A8F5FFAD6EED6" blockId="10.[151,1437,151,2013]" lastBlockId="11.[151,1437,151,2013]" lastPageId="11" lastPageNumber="212" pageId="10" pageNumber="211">
|
||
<emphasis id="B91EC4089858FF80FF3A8F5FFEA2E8EF" bold="true" box="[199,319,1916,1941]" pageId="10" pageNumber="211">Variation.</emphasis>
|
||
<taxonomicName id="4C6A63999858FF80FEBA8F5FFDDEE8EE" authorityName="Lieftinck" authorityYear="1948" box="[327,579,1916,1941]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC4089858FF80FEBA8F5FFDDEE8EE" box="[327,579,1916,1941]" italics="true" pageId="10" pageNumber="211">Leptogomphus mariae</emphasis>
|
||
</taxonomicName>
|
||
is discussed separately below. There is considerable variation in the extent of pale markings on the labrum and clypeus. In a number of males, mostly from GMNP in
|
||
<collectingRegion id="49AED6F89858FF80FB688F83FB65E8C3" box="[1173,1272,1952,1977]" country="Malaysia" name="Sarawak" pageId="10" pageNumber="211">
|
||
<collectingCountry id="F37D588A9858FF80FB688F83FB65E8C3" box="[1173,1272,1952,1977]" name="Malaysia" pageId="10" pageNumber="211">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
, the clypeus is entirely black, but various intermediate conditions between an entirely black clypeus and the condition seen in the male described here (and in the
|
||
<typeStatus id="54D1A6B89859FF81FDF488B4FDF0EFCA" box="[521,621,151,176]" pageId="11" pageNumber="212" type="holotype">holotype</typeStatus>
|
||
) occur as well. No female examined, except the
|
||
<typeStatus id="54D1A6B89859FF81FB5F88B4FA99EFCA" box="[1186,1284,151,176]" pageId="11" pageNumber="212" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6A63999859FF81FAD088BAFA06EFCA" authorityName="Lieftinck" authorityYear="1948" box="[1325,1435,153,176]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC4089859FF81FAD088BAFA06EFCA" box="[1325,1435,153,176]" italics="true" pageId="11" pageNumber="212">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
(see below) and a teneral female from Sungai Ingei in collection Orr (A.G. Orr personal communication) has an entirely black clypeus, but a few others have only small pale marks there. Nearly all specimens examined (both sexes) have either most of the antefrons pale, or an extensive intrusion of pale colour from postfrons onto antefrons (not just narrowly along the upper part). However three males from Mulu have no pale colour on the antefrons. The occipital horns, when present, vary in orientation and length; they are entirely absent in nine of the females examined (six from
|
||
<collectingRegion id="49AED6F89859FF81FE84894CFE62EEF2" box="[377,511,367,392]" country="Indonesia" name="Kalimantan Utara" pageId="11" pageNumber="212">Kalimantan</collectingRegion>
|
||
, three from
|
||
<collectingRegion id="49AED6F89859FF81FD7B894CFD75EEF2" box="[646,744,367,392]" country="Malaysia" name="Sarawak" pageId="11" pageNumber="212">
|
||
<collectingCountry id="F37D588A9859FF81FD7B894CFD75EEF2" box="[646,744,367,392]" name="Malaysia" pageId="11" pageNumber="212">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
) and only present on the right hand side in another three from
|
||
<collectingRegion id="49AED6F89859FF81FF6A89B7FE80EED7" box="[151,285,404,429]" country="Indonesia" name="Kalimantan Utara" pageId="11" pageNumber="212">Kalimantan</collectingRegion>
|
||
. The pits behind the tubercles are present in all specimens considered to belong to
|
||
<taxonomicName id="4C6A63999859FF81FB4689B6FAD9EED6" box="[1211,1348,405,428]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC4089859FF81FB4689B6FAD9EED6" box="[1211,1348,405,428]" italics="true" pageId="11" pageNumber="212">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9859FF81FF3A8994FF4BEDB7" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="212">There is variation in the exact width of the mesothoracic collar and the middorsal thoracic stripes; in one male from Lambir Hills the middorsal thoracic stripes are only connected to the mesothoracic collar on their inner side. In some individuals of both sexes there is a faint and very narrow section of antehumeral stripe present (never complete and never joined with the spot on the mesepisternum) around mid-length on the mesepisternum. There is variation in the exact shape of the stripe on the metepisternum and occasionally it is joined to the spot near the antealar carina on one or both sides of the synthorax. Small variations occur in wing venation and in the markings of the abdomen. The wing membrane varies between entirely hyaline to deeply brown tinted over much of the wing.</paragraph>
|
||
<paragraph id="8BD5181A9859FF81FF3A8AFBFE33EC97" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="212">
|
||
In males, there is some variation in how produced and how upturned the tip of the cercus is, but the main variation is in the curvature of the arms of the epiproct in lateral view; in some individuals they appear almost straight along the upper margin in its central part, in others the upper margin is strongly curved, almost semicircular. In females there is some variation in the exact shapes of the apices of the vulvar scale and its exact length. The peculiar tubercle on the sternite of S8 is present on all examined females where the condition of the specimen allows examination of this part, except that this character was only noticed after most older specimens from
|
||
<collectingRegion id="49AED6F89859FF81FF6A8B8CFE80ECB2" box="[151,285,943,968]" country="Indonesia" name="Kalimantan Utara" pageId="11" pageNumber="212">Kalimantan</collectingRegion>
|
||
,
|
||
<collectingRegion id="49AED6F89859FF81FEDA8B8CFEF6ECB2" box="[295,363,943,968]" country="Malaysia" name="Sabah" pageId="11" pageNumber="212">Sabah</collectingRegion>
|
||
and
|
||
<collectingCountry id="F37D588A9859FF81FE5E8B8CFE72ECB2" box="[419,495,943,968]" name="Brunei" pageId="11" pageNumber="212">Brunei</collectingCountry>
|
||
in RMNH and BMNH were examined by the first author, so it cannot be confirmed that it is present in these.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9859FF81FF3A8BD4FC69EB4F" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="212">
|
||
Considerable variation in size occurs in this species. Specimens from Sungai Ingei in
|
||
<collectingCountry id="F37D588A9859FF81FB6C8BD4FB40EB6A" box="[1169,1245,1015,1040]" name="Brunei" pageId="11" pageNumber="212">Brunei</collectingCountry>
|
||
are the smallest, but aside from this no clear geographical pattern is apparent to this variation.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9859FF81FF3A8C62FE92EA02" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="212">
|
||
<emphasis id="B91EC4089859FF81FF3A8C62FE98EB22" box="[199,261,1089,1112]" italics="true" pageId="11" pageNumber="212">Penis</emphasis>
|
||
. Unfortunately the penis of the specimen used in the description above was lost during extraction. The penis has been examined in a number of other specimens (e.g.
|
||
<figureCitation id="1351049F9859FF81FC948C47FC27EB06" box="[873,954,1124,1149]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="11" pageNumber="212">Fig. 94</figureCitation>
|
||
which shows that of SAR07_8_GOM18). The penultimate segment is extended to rear beyond the join with the terminal segment, and is bifurcated in this part, apices broadly triangular in lateral view, sometimes bent outwards so looking almost square ended in lateral view. The terminal segment is large at base, with a slightly constricted section before the apical part, giving the upper margin a bi-lobed appearance in lateral view. The apical part of the terminal segment bears two short cornua, just before these is a short ventrally directed flap, this can appear like a spine in lateral view. There is some variation in the depth of the constriction of the terminal segment, it is frequently shallower than in the individual illustrated.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9859FF81FF3A8DA0FB7EE956" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="212">
|
||
<emphasis id="B91EC4089859FF81FF3A8DA0FE4BEAE7" bold="true" box="[199,470,1411,1437]" pageId="11" pageNumber="212">
|
||
<taxonomicName id="4C6A63999859FF81FF3A8DA0FE52EAE6" authorityName="Lieftinck" authorityYear="1948" box="[199,463,1411,1436]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC4089859FF81FF3A8DA0FE52EAE6" bold="true" box="[199,463,1411,1436]" italics="true" pageId="11" pageNumber="212">Leptogomphus mariae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<bibRefCitation id="EFFB65EB9859FF81FE1E8DA7FD3DEAE6" author="Lieftinck" box="[483,672,1412,1437]" pageId="11" pageNumber="212" refString="Lieftinck, M. A. (1948) Descriptions and records of South - east Asiatic Odonata. Treubia, 19, 221 - 278." type="journal article" year="1948">Lieftinck (1948)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C6A63999859FF81FCD88DA6FC04EAE6" authorityName="Lieftinck" authorityYear="1948" box="[805,921,1413,1436]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC4089859FF81FCD88DA6FC04EAE6" box="[805,921,1413,1436]" italics="true" pageId="11" pageNumber="212">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
based on a single female, noting that it is “probably most closely related to
|
||
<taxonomicName id="4C6A63999859FF81FDD68D8AFD0BEABA" box="[555,662,1449,1472]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC4089859FF81FDD68D8AFD0BEABA" box="[555,662,1449,1472]" italics="true" pageId="11" pageNumber="212">coomansi</emphasis>
|
||
</taxonomicName>
|
||
”, but stating that it differs from
|
||
<taxonomicName id="4C6A63999859FF81FBD88D8AFB2FEABA" box="[1061,1202,1449,1472]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC4089859FF81FBD88D8AFB2FEABA" box="[1061,1202,1449,1472]" italics="true" pageId="11" pageNumber="212">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
(of which only the
|
||
<typeStatus id="54D1A6B89859FF81FF6A8DEFFF64EA9F" box="[151,249,1484,1509]" pageId="11" pageNumber="212" type="holotype">holotype</typeStatus>
|
||
male and one female were available at the time) “in its superior size, the structural features of the head, the much denser venation and longer pterostigma, and in details of colouration.” Examination of the
|
||
<typeStatus id="54D1A6B89859FF81FB038DCCFAFDE972" box="[1278,1376,1519,1544]" pageId="11" pageNumber="212" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6A63999859FF81FA788DD2FF7BE956" authorityName="Lieftinck" authorityYear="1948" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC4089859FF81FA788DD2FF7BE956" italics="true" pageId="11" pageNumber="212">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
has revealed that here are a number of inaccuracies in Lieftinck’s description of
|
||
<taxonomicName id="4C6A63999859FF81FB928E36FB46E956" authorityName="Lieftinck" authorityYear="1948" box="[1135,1243,1557,1580]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC4089859FF81FB928E36FB46E956" box="[1135,1243,1557,1580]" italics="true" pageId="11" pageNumber="212">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9859FF81FF3A8E14FDFFE9E2" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="212">
|
||
Lieftinck mentions a “small greenish spot placed upon the middle of the occipital plate” but no such spot is evident (
|
||
<figureCitation id="1351049F9859FF81FF058E7FFED6E90E" box="[248,331,1628,1653]" captionStart="FIGURES18–23" captionStartId="18.[151,264,1895,1918]" captionTargetBox="[183,1420,198,1847]" captionTargetId="figure@18.[183,1420,198,1847]" captionTargetPageId="18" captionText="FIGURES18–23.Female headdorsal:(18) L.coomansi SAR11_12_GOM48;(19)L.mariaeholotype;(20) L.sp.cf coomansi PomS260; (21) L. pasia holotype; (22) L.sii paratype SAR13_14_GOM44; (23) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068010/files/figure.png" pageId="11" pageNumber="212">Fig. 19</figureCitation>
|
||
); there is a slightly lighter region (dark brown rather than black) in this position but it is difficult to imagine that this ever appeared greenish.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9859FF81FF3A8E87FBE4E87F" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="212">
|
||
On the pits behind the tubercles on the vertex, Lieftinck writes “Just behind each of these protuberances are two minute smooth concavities, one placed in front of the other.” In fact only a single pit is evident behind each tubercle (
|
||
<figureCitation id="1351049F9859FF81FEFD8ECFFEC9E87E" box="[256,340,1772,1797]" captionStart="FIGURES28–31" captionStartId="22.[151,264,1357,1380]" captionTargetBox="[151,1435,266,1336]" captionTargetId="figure@22.[151,1436,265,1336]" captionTargetPageId="22" captionText="FIGURES28–31. Female head dorsaldetail; h—occipital horn; ot—occipital tube; p—pit; s—occipital spur; t—tubercle;thtubercle horn: (28) L. coomansi SAR11_12_GOM48; (29) L. mariae holotype; (30) L. sp. cf coomansi PomS260; (31) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068014/files/figure.png" pageId="11" pageNumber="212">Fig. 29</figureCitation>
|
||
), falling well within the range of variation seen in
|
||
<taxonomicName id="4C6A63999859FF81FC768ECEFB89E87E" box="[907,1044,1773,1796]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC4089859FF81FC768ECEFB89E87E" box="[907,1044,1773,1796]" italics="true" pageId="11" pageNumber="212">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
females.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9859FF81FF3A8F2CFA00E8A7" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="212">
|
||
Lieftinck states “Occipital plate with two distinct, divergent, roundish lateral ridges which are widest basally, originating from a point a little distant from the eye margin near the rounded occipital margin, but soon curving cephalad and laterad, stopping short a short distance ahead along the mesial border of the eye, sloping gently down without forming a projecting spine or protuberance”. These “ridges” are actually horns (
|
||
<figureCitation id="1351049F9859FF81FB648F5FFB6DE8EE" box="[1177,1264,1916,1941]" captionStart="FIGURES28–31" captionStartId="22.[151,264,1357,1380]" captionTargetBox="[151,1435,266,1336]" captionTargetId="figure@22.[151,1436,265,1336]" captionTargetPageId="22" captionText="FIGURES28–31. Female head dorsaldetail; h—occipital horn; ot—occipital tube; p—pit; s—occipital spur; t—tubercle;thtubercle horn: (28) L. coomansi SAR11_12_GOM48; (29) L. mariae holotype; (30) L. sp. cf coomansi PomS260; (31) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068014/files/figure.png" pageId="11" pageNumber="212">Fig. 29</figureCitation>
|
||
), as seen in
|
||
<taxonomicName id="4C6A63999859FF81FA788F5EFE9EE8C2" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC4089859FF81FA788F5EFE9EE8C2" italics="true" pageId="11" pageNumber="212">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1351049F9859FF81FEEC8F83FEF9E8C2" box="[273,356,1952,1977]" captionStart="FIGURES28–31" captionStartId="22.[151,264,1357,1380]" captionTargetBox="[151,1435,266,1336]" captionTargetId="figure@22.[151,1436,265,1336]" captionTargetPageId="22" captionText="FIGURES28–31. Female head dorsaldetail; h—occipital horn; ot—occipital tube; p—pit; s—occipital spur; t—tubercle;thtubercle horn: (28) L. coomansi SAR11_12_GOM48; (29) L. mariae holotype; (30) L. sp. cf coomansi PomS260; (31) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068014/files/figure.png" pageId="11" pageNumber="212">Fig. 28</figureCitation>
|
||
), but very strongly bent forward; possibly the position of the horns is not natural but the result of pressure applied to the head of the specimen, the top of the basal parts looks somewhat flattened and squashed.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A985EFF86FF6A88BBFEE1EFAE" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="213">
|
||
Apart from the extreme angle to the occipital ridge of these horns, they fall within the variation in shape and size seen in
|
||
<taxonomicName id="4C6A6399985EFF86FF11889EFEE8EFAE" box="[236,373,189,212]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FF11889EFEE8EFAE" box="[236,373,189,212]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A985EFF86FF3A88FCFB9FEE1F" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="213">Lieftinck states that “Prothorax entirely green above, only the sides of the median portion somewhat obscured.” A casual glance at the specimen conveys the impression that Lieftinck’s statement is more-or-less accurate, but a detailed examination reveals more extensive dark areas anterio-centrally on the middle pronotal lobe and in fact the colouration is much like that of the other Bornean species.</paragraph>
|
||
<paragraph id="8BD5181A985EFF86FF3A8953FB01ED47" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="213">
|
||
For the synthorax, Lieftinck refers the reader to his plate 8, where that of the female of
|
||
<taxonomicName id="4C6A6399985EFF86FB438952FAD4EEF2" box="[1214,1353,369,392]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FB438952FAD4EEF2" box="[1214,1353,369,392]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
is also illustrated. Differences in the dorsal markings in
|
||
<taxonomicName id="4C6A6399985EFF86FD4589B6FCB9EED6" authorityName="Lieftinck" authorityYear="1948" box="[696,804,405,428]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FD4589B6FCB9EED6" box="[696,804,405,428]" italics="true" pageId="12" pageNumber="213">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
fall within the variation seen in
|
||
<taxonomicName id="4C6A6399985EFF86FB7189B6FA88EED6" box="[1164,1301,405,428]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FB7189B6FA88EED6" box="[1164,1301,405,428]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
. Lieftinck’s illustration of the right hand side of the synthorax shows a significant difference in the pale marking on the metepisternum from that illustrated for
|
||
<taxonomicName id="4C6A6399985EFF86FDAB89FEFD7DEE8E" box="[598,736,477,500]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FDAB89FEFD7DEE8E" box="[598,736,477,500]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
in the same plate; the illustration of
|
||
<taxonomicName id="4C6A6399985EFF86FB7F89FEFB6CEE8E" authorityName="Lieftinck" authorityYear="1948" box="[1154,1265,477,500]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FB7F89FEFB6CEE8E" box="[1154,1265,477,500]" italics="true" pageId="12" pageNumber="213">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
is accurate but the marking on the left hand side (shown in
|
||
<figureCitation id="1351049F985EFF86FD6D89DCFD7DED62" box="[656,736,511,536]" captionStart="FIGURES 78–83" captionStartId="38.[151,264,1952,1975]" captionTargetBox="[169,1406,182,1934]" captionTargetId="figure@38.[169,1406,182,1934]" captionTargetPageId="38" captionText="FIGURES 78–83. Female synthorax lateral: (78) L. coomansi SAR11_12_GOM48; (79) L. mariae holotype, right hand side, flipped for consistency; (80) L. sp. cf coomansi PomS260; (81) L. pasia holotype; (82) L. sii paratype SAR13_14_GOM44; (83) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068036/files/figure.png" pageId="12" pageNumber="213">Fig. 79</figureCitation>
|
||
here) is typical for
|
||
<taxonomicName id="4C6A6399985EFF86FC3F8A22FBD1ED62" box="[962,1100,513,536]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FC3F8A22FBD1ED62" box="[962,1100,513,536]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
; and the marking of the right hand side falls within the variation seen in the
|
||
<taxonomicName id="4C6A6399985EFF86FD5C8A06FCB4ED46" box="[673,809,549,572]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FD5C8A06FCB4ED46" box="[673,809,549,572]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
females examined for this study.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A985EFF86FF3A8A64FDADEC4D" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="213">
|
||
Differences in density of wing venation and the length of the pterostigma between
|
||
<taxonomicName id="4C6A6399985EFF86FB908A6AFB6AED1A" box="[1133,1271,585,608]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FB908A6AFB6AED1A" box="[1133,1271,585,608]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6A6399985EFF86FAD38A6AFA01ED1A" authorityName="Lieftinck" authorityYear="1948" box="[1326,1436,585,608]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FAD38A6AFA01ED1A" box="[1326,1436,585,608]" italics="true" pageId="12" pageNumber="213">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
stated by Lieftinck are merely the result of the very few specimens examined;
|
||
<taxonomicName id="4C6A6399985EFF86FBE48A4EFB15EDFE" authorityName="Lieftinck" authorityYear="1948" box="[1049,1160,621,644]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FBE48A4EFB15EDFE" box="[1049,1160,621,644]" italics="true" pageId="12" pageNumber="213">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
actually falls within the variation seen in
|
||
<taxonomicName id="4C6A6399985EFF86FEA78AB2FE7EEDD2" box="[346,483,657,680]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FEA78AB2FE7EEDD2" box="[346,483,657,680]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
in these characters (at the upper end of pterostigma length). Although not explicitly mentioned, Lieftinck’s illustrations (his
|
||
<figureCitation id="1351049F985EFF86FDA48A97FD34EDB6" box="[601,681,692,717]" captionStart="FIGURES10–13" captionStartId="14.[151,264,1864,1887]" captionTargetBox="[151,1436,772,1843]" captionTargetId="figure@14.[151,1436,772,1843]" captionTargetPageId="14" captionText="FIGURES10–13. Male head frontal: (10) L.coomansi SAR09_10_GOM4; (11) L.coomansi holotype; (12) L.sp. cf coomansi SAB12_GOM4; (13) L.pendleburyi SAR13_14_GOM13." httpUri="https://zenodo.org/record/1068006/files/figure.png" pageId="12" pageNumber="213">Fig. 10</figureCitation>
|
||
on page 250) seem to show some differences in the vulvar scale of
|
||
<taxonomicName id="4C6A6399985EFF86FF6A8AFAFE99ED8A" authorityName="Lieftinck" authorityYear="1948" box="[151,260,729,752]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FF6A8AFAFE99ED8A" box="[151,260,729,752]" italics="true" pageId="12" pageNumber="213">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
compared to
|
||
<taxonomicName id="4C6A6399985EFF86FE5D8AFAFDB4ED8A" box="[416,553,729,752]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FE5D8AFAFDB4ED8A" box="[416,553,729,752]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
, but again examination of more material has shown that the vulvar scale of
|
||
<taxonomicName id="4C6A6399985EFF86FA788AFAFF7BEC6E" authorityName="Lieftinck" authorityYear="1948" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FA788AFAFF7BEC6E" italics="true" pageId="12" pageNumber="213">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
falls within the variation in
|
||
<taxonomicName id="4C6A6399985EFF86FDD48ADEFD2FEC6E" box="[553,690,765,788]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FDD48ADEFD2FEC6E" box="[553,690,765,788]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
. The tubercle on the sternite of S8 of
|
||
<taxonomicName id="4C6A6399985EFF86FB998ADEFB70EC6E" box="[1124,1261,765,788]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FB998ADEFB70EC6E" box="[1124,1261,765,788]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
females is also present in the
|
||
<typeStatus id="54D1A6B8985EFF86FEC98B3CFE0BEC42" box="[308,406,799,824]" pageId="12" pageNumber="213" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6A6399985EFF86FE418B02FDB4EC42" authorityName="Lieftinck" authorityYear="1948" box="[444,553,801,824]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FE418B02FDB4EC42" box="[444,553,801,824]" italics="true" pageId="12" pageNumber="213">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A985EFF86FF3A8B67FB3BEB4F" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="213">
|
||
<taxonomicName id="4C6A6399985EFF86FF3A8B67FE5EEC26" authorityName="Lieftinck" authorityYear="1948" box="[199,451,836,861]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FF3A8B67FE5EEC26" box="[199,451,836,861]" italics="true" pageId="12" pageNumber="213">Leptogomphus mariae</emphasis>
|
||
</taxonomicName>
|
||
is slightly larger than any female of
|
||
<taxonomicName id="4C6A6399985EFF86FC9C8B66FC77EC26" box="[865,1002,837,860]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FC9C8B66FC77EC26" box="[865,1002,837,860]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
examined in this study, with abdomen and Hw each ca
|
||
<quantity id="4C92B5FF985EFF86FEAC8B4BFE16ECFA" box="[337,395,872,896]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="213" unit="mm" value="1.0">1mm</quantity>
|
||
longer than in the largest
|
||
<taxonomicName id="4C6A6399985EFF86FD4D8B4AFCA4ECFA" box="[688,825,873,896]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FD4D8B4AFCA4ECFA" box="[688,825,873,896]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
specimens (note that Lieftinck gives the Hw length of
|
||
<taxonomicName id="4C6A6399985EFF86FF6A8BAEFE99ECDE" authorityName="Lieftinck" authorityYear="1948" box="[151,260,909,932]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FF6A8BAEFE99ECDE" box="[151,260,909,932]" italics="true" pageId="12" pageNumber="213">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
as
|
||
<quantity id="4C92B5FF985EFF86FED68BAFFEEFECDF" box="[299,370,908,933]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="213" unit="mm" value="35.0">35mm</quantity>
|
||
but the first author of this paper measured it as
|
||
<quantity id="4C92B5FF985EFF86FC6F8BAFFC42ECDE" box="[914,991,908,932]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="213" unit="mm" value="34.0">34mm</quantity>
|
||
). The entirely black clypeus is unusual, but does occur in some
|
||
<taxonomicName id="4C6A6399985EFF86FE598B92FDB3ECB2" box="[420,558,945,968]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FE598B92FDB3ECB2" box="[420,558,945,968]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
males and one female in collection Orr; this female is at the lower end of the size range of
|
||
<taxonomicName id="4C6A6399985EFF86FED18BF6FE2BEC96" box="[300,438,981,1004]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FED18BF6FE2BEC96" box="[300,438,981,1004]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
(A.G. Orr personal communication), in contrast to the
|
||
<taxonomicName id="4C6A6399985EFF86FBE48BF6FB18EC96" authorityName="Lieftinck" authorityYear="1948" box="[1049,1157,981,1004]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FBE48BF6FB18EC96" box="[1049,1157,981,1004]" italics="true" pageId="12" pageNumber="213">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54D1A6B8985EFF86FB708BF7FB6EEC97" box="[1165,1267,980,1005]" pageId="12" pageNumber="213" type="holotype">holotype</typeStatus>
|
||
. The
|
||
<taxonomicName id="4C6A6399985EFF86FAD28BF6FA06EC96" authorityName="Lieftinck" authorityYear="1948" box="[1327,1435,981,1004]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FAD28BF6FA06EC96" box="[1327,1435,981,1004]" italics="true" pageId="12" pageNumber="213">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54D1A6B8985EFF86FF6A8BD4FF64EB6A" box="[151,249,1015,1040]" pageId="12" pageNumber="213" type="holotype">holotype</typeStatus>
|
||
has a reduced pale marking on the frons compared with typical
|
||
<taxonomicName id="4C6A6399985EFF86FBF88BDAFB0EEB6A" box="[1029,1171,1017,1040]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FBF88BDAFB0EEB6A" box="[1029,1171,1017,1040]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
but there is sufficient variation in this character in
|
||
<taxonomicName id="4C6A6399985EFF86FE258C3EFDFDEB4E" box="[472,608,1053,1076]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FE258C3EFDFDEB4E" box="[472,608,1053,1076]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
to be sure that it is not a reliable diagnostic feature.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A985EFF86FF3A8C63FCB5EA76" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="213">
|
||
On balance there seems insufficient reason to keep
|
||
<taxonomicName id="4C6A6399985EFF86FCFA8C62FCE9EB22" authorityName="Lieftinck" authorityYear="1948" box="[775,884,1089,1112]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FCFA8C62FCE9EB22" box="[775,884,1089,1112]" italics="true" pageId="12" pageNumber="213">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
separate from
|
||
<taxonomicName id="4C6A6399985EFF86FBE18C62FB38EB22" box="[1052,1189,1089,1112]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FBE18C62FB38EB22" box="[1052,1189,1089,1112]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
at this time. The only characters in which it falls outside of the variation seen in typical
|
||
<taxonomicName id="4C6A6399985EFF86FC578C46FBAAEB06" box="[938,1079,1125,1148]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FC578C46FBAAEB06" box="[938,1079,1125,1148]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
females are the entirely black clypeus (
|
||
<figureCitation id="1351049F985EFF86FF018CA4FED0EBDA" box="[252,333,1159,1184]" captionStart="FIGURES 36–41" captionStartId="25.[151,264,1973,1996]" captionTargetBox="[204,1387,183,1958]" captionTargetId="figure@25.[204,1387,183,1958]" captionTargetPageId="25" captionText="FIGURES 36–41. Female head frontal: (36) L. coomansi SAR11_12_GOM48; (37) L. mariae holotype; (38) L. sp. cf coomansi PomS260; (39) L.pasia holotype; (40) L.sii paratype SAR13_14_GOM44; (41) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068018/files/figure.png" pageId="12" pageNumber="213">Fig. 37</figureCitation>
|
||
) and slightly greater size. However some female
|
||
<taxonomicName id="4C6A6399985EFF86FC8A8CAAFB9DEBDA" box="[887,1024,1161,1184]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FC8A8CAAFB9DEBDA" box="[887,1024,1161,1184]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
have very reduced pale markings on the clypeus (entirely black in one, as mentioned above), and some males also have an entirely black clypeus; the size difference alone is not great enough to justify the separation of
|
||
<taxonomicName id="4C6A6399985EFF86FC538CF2FB80EB92" authorityName="Lieftinck" authorityYear="1948" box="[942,1053,1233,1256]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FC538CF2FB80EB92" box="[942,1053,1233,1256]" italics="true" pageId="12" pageNumber="213">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C6A6399985EFF86FB9A8CF2FB6FEB92" box="[1127,1266,1233,1256]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FB9A8CF2FB6FEB92" box="[1127,1266,1233,1256]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
. Therefore we consider
|
||
<taxonomicName id="4C6A6399985EFF86FF018CD6FEF4EA76" authorityName="Lieftinck" authorityYear="1948" box="[252,361,1269,1292]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FF018CD6FEF4EA76" box="[252,361,1269,1292]" italics="true" pageId="12" pageNumber="213">L. mariae</emphasis>
|
||
</taxonomicName>
|
||
to be a junior synonym of
|
||
<taxonomicName id="4C6A6399985EFF86FD658CD6FCBDEA76" box="[664,800,1269,1292]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FD658CD6FCBDEA76" box="[664,800,1269,1292]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A985EFF86FF3A8D3BFD4BEAE6" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="213">
|
||
Measurements (mm). Males: Hw 27.5–32, abdomen excluding anal appendages 32–38.5, 15–19 Ax in Fw, 11– 14 Ax in Hw, 10–14 Px in Fw, 9–13 Px in Hw. Females: Typical
|
||
<taxonomicName id="4C6A6399985EFF86FC928D1EFBB2EA2E" authority=": Hw" authorityName="Hw" box="[879,1071,1341,1364]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FC928D1EFC6AEA2E" box="[879,1015,1341,1364]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
: Hw
|
||
</taxonomicName>
|
||
30–33, abdomen excluding anal appendages 34.5–39, 14–19 Ax in Fw, 11–14 Ax in Hw, 10–14 Px in Fw, 9–13 Px in Hw;
|
||
<typeStatus id="54D1A6B8985EFF86FB608D7CFB62EA02" box="[1181,1279,1375,1400]" pageId="12" pageNumber="213" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6A6399985EFF86FADB8D42FF5DEAE6" authority=": Hw" authorityName="Hw" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B91EC408985EFF86FADB8D42FA09EA02" box="[1318,1428,1377,1400]" italics="true" pageId="12" pageNumber="213">L. mariae</emphasis>
|
||
: Hw
|
||
</taxonomicName>
|
||
34, abdomen excluding anal appendages ca 40.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B91985EFF87FF3A8D84FC0EEE3A" lastPageId="13" lastPageNumber="214" pageId="12" pageNumber="213" type="discussion">
|
||
<paragraph id="8BD5181A985EFF86FF3A8D84FB99E852" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="213">
|
||
<emphasis id="B91EC408985EFF86FF3A8D84FEA6EABA" bold="true" box="[199,315,1447,1472]" pageId="12" pageNumber="213">Remarks.</emphasis>
|
||
The COI gene tree (
|
||
<figureCitation id="1351049F985EFF86FDCB8D84FDE6EABA" box="[566,635,1447,1472]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="12" pageNumber="213">Fig. 1</figureCitation>
|
||
) shows
|
||
<taxonomicName id="4C6A6399985EFF86FD1E8D8AFCEDEABA" box="[739,880,1449,1472]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FD1E8D8AFCEDEABA" box="[739,880,1449,1472]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
divided into two clades. One clade consists of specimens from
|
||
<collectingRegion id="49AED6F8985EFF86FEAC8DEFFE77EA9F" box="[337,490,1484,1509]" country="Brunei" name="Belait" pageId="12" pageNumber="213">Belait district</collectingRegion>
|
||
in
|
||
<collectingCountry id="F37D588A985EFF86FDF28DEFFDC6EA9F" box="[527,603,1484,1509]" name="Brunei" pageId="12" pageNumber="213">Brunei</collectingCountry>
|
||
and Lambir Hills in
|
||
<collectingRegion id="49AED6F8985EFF86FCB48DEFFC37EA9F" box="[841,938,1484,1509]" country="Malaysia" name="Sarawak" pageId="12" pageNumber="213">
|
||
<collectingCountry id="F37D588A985EFF86FCB48DEFFC37EA9F" box="[841,938,1484,1509]" name="Malaysia" pageId="12" pageNumber="213">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
(not far from
|
||
<collectingCountry id="F37D588A985EFF86FBB68DEFFB06EA9F" box="[1099,1179,1484,1509]" name="Brunei" pageId="12" pageNumber="213">Brunei</collectingCountry>
|
||
), and also including
|
||
<emphasis id="B91EC408985EFF86FA788DEEFA01EA9E" box="[1413,1436,1485,1508]" italics="true" pageId="12" pageNumber="213">L.</emphasis>
|
||
sp. cf
|
||
<taxonomicName id="4C6A6399985EFF86FF278DD2FED8E972" box="[218,325,1521,1544]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FF278DD2FED8E972" box="[218,325,1521,1544]" italics="true" pageId="12" pageNumber="213">coomansi</emphasis>
|
||
</taxonomicName>
|
||
. The other clade consists of specimens from the upper Baram in Sarawak’s Miri Division, the Hose Mountains in Kapit Division and Kubah National Park in Kuching Division; these locations are well spread across
|
||
<collectingCountry id="F37D588A985EFF86FF6A8E14FF66E92A" box="[151,251,1591,1616]" name="Malaysia" pageId="12" pageNumber="213">
|
||
<collectingRegion id="49AED6F8985EFF86FF6A8E14FF66E92A" box="[151,251,1591,1616]" country="Malaysia" name="Sarawak" pageId="12" pageNumber="213">Sarawak</collectingRegion>
|
||
</collectingCountry>
|
||
. However the ITS gene tree (
|
||
<figureCitation id="1351049F985EFF86FDB18E14FD0CE92A" box="[588,657,1591,1616]" captionStart="FIGURE 2" captionStartId="5.[151,250,1294,1316]" captionTargetBox="[243,1384,200,1237]" captionTargetId="figure@5.[151,1436,193,1273]" captionTargetPageId="5" captionText="FIGURE 2. ITS gene tree for 20 specimens of Leptogomphus and three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068000/files/figure.png" pageId="12" pageNumber="213">Fig. 2</figureCitation>
|
||
) tells a somewhat different story, with
|
||
<emphasis id="B91EC408985EFF86FBAC8E1AFBF5E92A" box="[1105,1128,1593,1616]" italics="true" pageId="12" pageNumber="213">L.</emphasis>
|
||
sp. cf
|
||
<taxonomicName id="4C6A6399985EFF86FB4A8E1AFABFE92A" box="[1207,1314,1593,1616]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FB4A8E1AFABFE92A" box="[1207,1314,1593,1616]" italics="true" pageId="12" pageNumber="213">coomansi</emphasis>
|
||
</taxonomicName>
|
||
separated, albeit weakly, from
|
||
<taxonomicName id="4C6A6399985EFF86FE818E7EFD9AE90E" box="[380,519,1629,1652]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FE818E7EFD9AE90E" box="[380,519,1629,1652]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
, and little separation between the
|
||
<taxonomicName id="4C6A6399985EFF86FC6E8E7EFB83E90E" box="[915,1054,1629,1652]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FC6E8E7EFB83E90E" box="[915,1054,1629,1652]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
samples. The combined analyses (
|
||
<figureCitation id="1351049F985EFF86FF628E5CFF79E9E2" box="[159,228,1663,1688]" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1893]" captionTargetBox="[209,1373,710,1811]" captionTargetId="figure@6.[151,1436,700,1850]" captionTargetPageId="6" captionText="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068002/files/figure.png" pageId="12" pageNumber="213">Fig. 3</figureCitation>
|
||
) retains the separation of
|
||
<emphasis id="B91EC408985EFF86FDE58EA2FDB3E9E2" box="[536,558,1665,1688]" italics="true" pageId="12" pageNumber="213">L.</emphasis>
|
||
sp. cf
|
||
<taxonomicName id="4C6A6399985EFF86FD7F8EA2FD70E9E2" box="[642,749,1665,1688]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FD7F8EA2FD70E9E2" box="[642,749,1665,1688]" italics="true" pageId="12" pageNumber="213">coomansi</emphasis>
|
||
</taxonomicName>
|
||
as the sister of
|
||
<taxonomicName id="4C6A6399985EFF86FC518EA2FBAAE9E2" box="[940,1079,1665,1688]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FC518EA2FBAAE9E2" box="[940,1079,1665,1688]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
seen in the ITS gene tree, but relationships within
|
||
<taxonomicName id="4C6A6399985EFF86FE878E86FD9FE9C6" box="[378,514,1701,1724]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FE878E86FD9FE9C6" box="[378,514,1701,1724]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
are dominated by the COI so that the same two clades seen in
|
||
<figureCitation id="1351049F985EFF86FB468E87FB61E9C6" box="[1211,1276,1700,1725]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="12" pageNumber="213">Fig. 1</figureCitation>
|
||
appear in
|
||
<figureCitation id="1351049F985EFF86FA8D8E87FF3AE99A" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1893]" captionTargetBox="[209,1373,710,1811]" captionTargetId="figure@6.[151,1436,700,1850]" captionTargetPageId="6" captionText="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068002/files/figure.png" pageId="12" pageNumber="213">Fig. 3</figureCitation>
|
||
. No consistent morphological difference has been found between the two clades in
|
||
<taxonomicName id="4C6A6399985EFF86FBB48EEAFB4CE99A" box="[1097,1233,1737,1760]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FBB48EEAFB4CE99A" box="[1097,1233,1737,1760]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
seen in
|
||
<figureCitation id="1351049F985EFF86FAD18EE4FA08E99A" box="[1324,1429,1735,1760]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="12" pageNumber="213">Figures 1</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F985EFF86FF6A8ECFFF38E87E" box="[151,165,1772,1796]" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1893]" captionTargetBox="[209,1373,710,1811]" captionTargetId="figure@6.[151,1436,700,1850]" captionTargetPageId="6" captionText="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068002/files/figure.png" pageId="12" pageNumber="213">3</figureCitation>
|
||
and in the absence of further evidence we do not believe that they represent separate, cryptic species, and are merely the result of variation in the COI marker within one biological species.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A985EFF86FF3A8F17FBB6E8C2" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="213">
|
||
As noted above, the pits behind the tubercles are present in all females examined and appear to offer a more reliable diagnostic character than the occipital horns, which are variably present. The peculiar tubercle on the sternum of S8 also appears to offer a diagnostic character for the female; swellings are present in this position in some of the other named species, but are not as well developed as in
|
||
<taxonomicName id="4C6A6399985EFF86FC668F82FBBEE8C2" box="[923,1059,1953,1976]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FC668F82FBBEE8C2" box="[923,1059,1953,1976]" italics="true" pageId="12" pageNumber="213">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A985EFF87FF3A8FE7FC0EEE3A" blockId="12.[151,1437,151,2013]" lastBlockId="13.[151,1436,151,320]" lastPageId="13" lastPageNumber="214" pageId="12" pageNumber="213">
|
||
<taxonomicName id="4C6A6399985EFF86FF3A8FE7FE7DE8A6" box="[199,480,1988,2013]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985EFF86FF3A8FE7FE7DE8A6" box="[199,480,1988,2013]" italics="true" pageId="12" pageNumber="213">Leptogomphus coomansi</emphasis>
|
||
</taxonomicName>
|
||
occurs at forest streams in the lowlands of Borneo, and has been recorded from a wide range of stream sizes, from very small closed canopy streams to broad open canopy ones. It is widely distributed in
|
||
<collectingCountry id="F37D588A985FFF87FECE889FFE1FEFAF" box="[307,386,188,213]" name="Brunei" pageId="13" pageNumber="214">Brunei</collectingCountry>
|
||
,
|
||
<collectingRegion id="49AED6F8985FFF87FE70889FFD8DEFAF" box="[397,528,188,213]" country="Indonesia" name="Kalimantan Utara" pageId="13" pageNumber="214">Kalimantan</collectingRegion>
|
||
and
|
||
<collectingRegion id="49AED6F8985FFF87FDBA889FFD3AEFAF" box="[583,679,188,213]" country="Malaysia" name="Sarawak" pageId="13" pageNumber="214">
|
||
<collectingCountry id="F37D588A985FFF87FDBA889FFD3AEFAF" box="[583,679,188,213]" name="Malaysia" pageId="13" pageNumber="214">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
(
|
||
<figureCitation id="1351049F985FFF87FD4A889FFC87EFAF" box="[695,794,188,213]" captionStart="FIGURE 126" captionStartId="48.[151,250,1801,1823]" captionTargetBox="[183,1420,421,1756]" captionTargetId="figure@48.[183,1420,421,1756]" captionTargetPageId="48" captionText="FIGURE 126. Distribution of L. coomansi, confirmed (yellow square) and unconfirmed (blue triangle) records, and L. sp. cf coomansi (black circle)." httpUri="https://zenodo.org/record/1068054/files/figure.png" pageId="13" pageNumber="214">Fig. 126</figureCitation>
|
||
). The only record of
|
||
<taxonomicName id="4C6A6399985FFF87FBFB889EFB12EFAE" box="[1030,1167,189,212]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408985FFF87FBFB889EFB12EFAE" box="[1030,1167,189,212]" italics="true" pageId="13" pageNumber="214">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49AED6F8985FFF87FB28889FFA84EFAF" box="[1237,1305,188,213]" country="Malaysia" name="Sabah" pageId="13" pageNumber="214">Sabah</collectingRegion>
|
||
that we are aware of is that of a female from Poring in
|
||
<bibRefCitation id="EFFB65EB985FFF87FD8888FCFC0BEF82" author="Tsuda" box="[629,918,223,248]" pageId="13" pageNumber="214" refString="Tsuda, S. and Kitagawa, K. (1989) Odonata of Southeast Asia collected by the late Mr. M. Iwasaki. Part 3. Odonata of North Borneo (Sabah, Malaysia). Gracile, 40, 37 - 39." type="journal article" year="1989">Tsuda & Kitagawa (1989)</bibRefCitation>
|
||
without any further details apart from the date of collection. We consider this record as unconfirmed, at least until more material of this species from
|
||
<collectingRegion id="49AED6F8985FFF87FAA58927FA01EE67" box="[1368,1436,260,285]" country="Malaysia" name="Sabah" pageId="13" pageNumber="214">Sabah</collectingRegion>
|
||
becomes available. However, the species is likely to occur in
|
||
<collectingRegion id="49AED6F8985FFF87FCBA8904FC12EE3A" box="[839,911,295,320]" country="Malaysia" name="Sabah" pageId="13" pageNumber="214">Sabah</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |