303 lines
44 KiB
XML
303 lines
44 KiB
XML
<document id="15B6BB891153C34F9BACCA219D2521D6" ID-CLB-Dataset="22181" ID-DOI="10.5852/ejt.2018.477" ID-GBIF-Dataset="c447a6b6-36b5-40de-ab76-80a9c7d96d3a" ID-Zenodo-Dep="3825218" ID-ZooBank="935D0B09-0508-4771-9575-28C1FE761383" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589391210860" checkinUser="valdenar" docAuthor="Read, Helen J. & Enghoff, Henrik" docDate="2018" docId="03C3A16F9039FFD3F072FE8DFEE91B66" docLanguage="en" docName="ejt-477_read_enghoff.pdf.imf" docOrigin="European Journal of Taxonomy 477" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Columbianum nahvalr Read & Enghoff, 2018, sp. nov." docType="treatment" docUuid="FA3360D7-B4BD-4DB8-B248-97C92EF1517B" docUuidSource="ZooBank" docVersion="8" lastPageNumber="18" masterDocId="FFFAD9179037FFC2F22DFF86FF911806" masterDocTitle="Siphonophoridae from Brazilian Amazonia Part 1 - The genus Columbianum Verhoeff, 1941 (Diplopoda, Siphonophorida)" masterLastPageNumber="23" masterPageNumber="1" pageNumber="15" updateTime="1698839485074" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="059A0A0DA38CD4A644CB558406C70E2C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B91417FB975504BBB0A2E81CA1EF2567">
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<mods:title id="AC77874388E2CC047D69AB363C2371ED">Siphonophoridae from Brazilian Amazonia Part 1 - The genus Columbianum Verhoeff, 1941 (Diplopoda, Siphonophorida)</mods:title>
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<mods:name id="D83CDE392C8BFA93E597034DDD0C5C6F" type="personal">
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<mods:roleTerm id="F6755FC354FF6F26B9573B55C5DCEB6E">Author</mods:roleTerm>
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<mods:namePart id="5A9E26B2F50D2BCC13037AD89EA2317F">Read, Helen J.</mods:namePart>
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</mods:name>
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<mods:name id="CF3D614D5AB49495EE8551B807BEB3A1" type="personal">
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<mods:roleTerm id="288BCE60FD8FE27D90F62ECF4ADD8823">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CAF138EBD4FDDA588F2C083675D86583">Enghoff, Henrik</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="F261D1BA22AAF00249A2D3C7FFA541C6">text</mods:typeOfResource>
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<mods:title id="76A62501EDD7FD518AA2FE654EB0F7F7">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="6CA2DDEFC2538F2C78EA252F290F396A">
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<mods:date id="CA9D7B8E29182FF3C54658906E9A2C34">2018</mods:date>
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<mods:number id="BADF5AB83E90C9F3FBC7BADCEFBA3492">2018-11-20</mods:number>
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<mods:detail id="1E6716EE8F45FC6736DFDF2AEB816DCC" type="volume">
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<mods:number id="E325BE106D1BF5BC7B1FD3E8414BDC48">477</mods:number>
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<mods:classification id="408BB84CDC4A53B13AE28B0360168B6C">journal article</mods:classification>
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<mods:identifier id="BE0CC29ED3132A271A08358C0B9BBD9C" type="CLB-Dataset">22181</mods:identifier>
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<mods:identifier id="AD20198A40021EDD8AAD57021C30D053" type="DOI">10.5852/ejt.2018.477</mods:identifier>
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<mods:identifier id="54C15D858CE45BCF60E36D7E6C3F8F54" type="GBIF-Dataset">c447a6b6-36b5-40de-ab76-80a9c7d96d3a</mods:identifier>
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<mods:identifier id="BC4A06EEB224A869EE6C0417F5EF859D" type="Zenodo-Dep">3825218</mods:identifier>
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<mods:identifier id="FA0FC959B90CDF16F29695E15A98BF7E" type="ZooBank">935D0B09-0508-4771-9575-28C1FE761383</mods:identifier>
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<treatment id="03C3A16F9039FFD3F072FE8DFEE91B66" ID-DOI="http://doi.org/10.5281/zenodo.3845979" ID-GBIF-Taxon="164131185" ID-Zenodo-Dep="3845979" LSID="urn:lsid:zoobank.org:act:FA3360D7-B4BD-4DB8-B248-97C92EF1517B" httpUri="http://treatment.plazi.org/id/03C3A16F9039FFD3F072FE8DFEE91B66" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
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<subSubSection id="C37043F29039FFCCF072FE8DFB58194F" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="8BD510799039FFCCF072FE8DFC441920" blockId="14.[363,1225,267,365]" box="[607,981,267,294]" pageId="14" pageNumber="15">
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<heading id="D09DA7159039FFCCF072FE8DFC441920" box="[607,981,267,294]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
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<taxonomicName id="4C6A6BFA9039FFCCF072FE8DFCE51920" ID-CoL="8Q8T9" authorityName="Read & Enghoff" authorityYear="2018" box="[607,884,267,294]" class="Diplopoda" family="Siphonophoridae" genus="Columbianum" kingdom="Animalia" order="Siphonophorida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nahvalr" status="sp. nov.">
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<emphasis id="B91ECC6B9039FFCCF072FE8DFCE51920" bold="true" box="[607,884,267,294]" italics="true" pageId="14" pageNumber="15">Columbianum nahvalr</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22D71109039FFCCF157FE8AFC441920" box="[890,981,268,294]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BD510799039FFCCF346FEA8FB58194F" blockId="14.[363,1225,267,365]" box="[363,1225,302,329]" pageId="14" pageNumber="15">
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<uri id="FFFB1C7B9039FFCCF346FEA8FB58194F" box="[363,1225,302,329]" pageId="14" pageNumber="15">
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urn:lsid:zoobank.org:act:
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<uuid id="FFCC2AAC9039FFCCF0A6FEA8FB58194F" box="[651,1225,302,329]" pageId="14" pageNumber="15">FA3360D7-B4BD-4DB8-B248-97C92EF1517B</uuid>
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</uri>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C37043F29039FFCCF0F5FED4FCCA196B" box="[728,859,338,365]" pageId="14" pageNumber="15" type="description">
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<paragraph id="8BD510799039FFCCF0F5FED4FCCA196B" blockId="14.[363,1225,267,365]" box="[728,859,338,365]" pageId="14" pageNumber="15">
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<figureCitation id="13510CFC9039FFCCF0F5FED4FCB0196A" box="[728,801,338,364]" captionStart="Fig" captionStartId="11.[189,232,1945,1971]" captionTargetBox="[203,1385,344,1859]" captionTargetId="figure@11.[202,1393,333,1873]" captionTargetPageId="11" captionText="Fig. 1. Body width (mm) against the total number of tergites with legs for new species of Columbianum Verhoeff, 1941. A. Males. B. Females." figureDoi="http://doi.org/10.5281/zenodo.3825220" httpUri="https://zenodo.org/record/3825220/files/figure.png" pageId="14" pageNumber="15">Figs 1</figureCitation>
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,
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<figureCitation id="13510CFC9039FFCCF102FED4FCCA196B" box="[815,859,338,365]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="15.[189,232,1847,1873]" captionStartId-1="16.[189,232,1807,1833]" captionTargetBox-0="[195,1391,967,1802]" captionTargetBox-1="[197,1387,751,1756]" captionTargetId-0="figure@15.[189,1399,960,1810]" captionTargetId-1="figure@16.[189,1399,745,1767]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="Fig. 4. Columbianum nahvalr sp. nov. A. Head and anterior body rings of male, showing the appearance of gonopods in a ‘relaxed specimen’. Body width approximately 1 mm. B. Close up of gonopods in a ‘relaxed’ specimen. Body width approximately 1 mm. C. Head of holotype male in dorsal view. Note long setae originating from base of rostrum. D. Head and rostrum of female in dorsal view. E. Head and rostrum of female in lateral view. Scale bars = C = 0.2 mm; D–E = 0.1 mm." captionText-1="Fig. 5. Columbianum nahvalr sp. nov.A. Midbody rings from lateral view, showing pronounced difference between pro- and metazonite and shape of pleurites. B. ‘Channel’ between pro- and metazonite in dorsal view of male showing tubercles and line of setae. C. Tarsal claw and accessory claw of midbody leg from right side of male, in anterior dorsal view. D. Gonopods of holotype male, lateral view with anterior gonopods to right. E. Gonopods of small, presumably immature male, in lateral view, anterior gonopds to left. Scale bars = A, D–E = 0.2 mm; B–C = 0.01 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3825226" figureDoi-1="http://doi.org/10.5281/zenodo.3825230" httpUri-0="https://zenodo.org/record/3825226/files/figure.png" httpUri-1="https://zenodo.org/record/3825230/files/figure.png" pageId="14" pageNumber="15">4–5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37043F29039FFCCF290FE15FD121A8A" pageId="14" pageNumber="15" type="diagnosis">
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<paragraph id="8BD510799039FFCCF290FE15FEA719AB" blockId="14.[189,310,403,429]" box="[189,310,403,429]" pageId="14" pageNumber="15">
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<heading id="D09DA7159039FFCCF290FE15FEA719AB" bold="true" box="[189,310,403,429]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
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<emphasis id="B91ECC6B9039FFCCF290FE15FEA719AB" bold="true" box="[189,310,403,429]" pageId="14" pageNumber="15">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD510799039FFCCF290FE44FD121A8A" blockId="14.[189,1398,450,652]" pageId="14" pageNumber="15">Fairly large pale species. Rounded and domed head with sharp distinction between head and narwhallike rostrum. Tubercles and setae on metazonites abundant and irregular, setae quite long, giving a rather ragged appearance. Prozonites without tubercles. Channel between pro- and metazonite with horseshoeshaped tubercles. Limbus crenulated like posterior margin of pleurites. Accessory claws robust, roughly half the length of claw and situated on the dorso-anterior side of claw. Anterior gonopods resembling walking legs with a simple distal claw.</paragraph>
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</subSubSection>
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<subSubSection id="C37043F29039FFCCF290FD35FDA11CC4" pageId="14" pageNumber="15" type="etymology">
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<paragraph id="8BD510799039FFCCF290FD35FED51ACB" blockId="14.[189,324,691,717]" box="[189,324,691,717]" pageId="14" pageNumber="15">
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<heading id="D09DA7159039FFCCF290FD35FED51ACB" bold="true" box="[189,324,691,717]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
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<emphasis id="B91ECC6B9039FFCCF290FD35FED51ACB" bold="true" box="[189,324,691,717]" pageId="14" pageNumber="15">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD510799039FFCCF290FD64FBBC1B19" blockId="14.[189,1398,738,800]" pageId="14" pageNumber="15">
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Noun in apposition referring to the Old Norse name for the narwhal (
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<taxonomicName id="4C6A6BFA9039FFCCF62CFD64FF6D1B26" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Monodontidae" genus="Monodon" kingdom="Animalia" order="Cetacea" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="monoceros">
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<emphasis id="B91ECC6B9039FFCCF62CFD64FB6A1AFA" box="[1025,1275,738,764]" italics="true" pageId="14" pageNumber="15">Monodon monoceros</emphasis>
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Linnaeus, 1758
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</taxonomicName>
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). The head of our species strongly resembles the head of the narwhal.
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</paragraph>
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<paragraph id="8BD510799039FFCCF290FCC0FD391B67" blockId="14.[189,680,837,865]" box="[189,680,837,865]" pageId="14" pageNumber="15">
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<emphasis id="B91ECC6B9039FFCCF290FCC0FE1D1B66" bold="true" box="[189,396,838,864]" pageId="14" pageNumber="15">Material studied</emphasis>
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(6 ♂♂,
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<specimenCount id="9D6CDBF09039FFCCF3C2FCC0FDA11B66" box="[495,560,838,864]" pageId="14" pageNumber="15" type="generic">1 juv.</specimenCount>
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♂,
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<specimenCount id="9D6CDBF09039FFCCF076FCC0FD0A1B67" box="[603,667,838,865]" pageId="14" pageNumber="15" type="female">9 ♀♀</specimenCount>
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)
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</paragraph>
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<materialsCitation id="3B021A249039FFCCF2C7FCF3FAC01BD3" ID-GBIF-Occurrence="2617839304" collectingDate="1983-04-25" collectionCode="INPA" collectorName="Adis & Rodrigues" country="Brazil" latitude="-3.0333333" location="Amazonas Province" longLatPrecision="1307" longitude="-60.283333" municipality="Rio Taruma Mirim" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
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<paragraph id="8BD510799039FFCCF2C7FCF3FECD1B89" blockId="14.[189,1399,885,982]" box="[234,348,885,911]" pageId="14" pageNumber="15">
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<emphasis id="B91ECC6B9039FFCCF2C7FCF3FECD1B89" bold="true" box="[234,348,885,911]" pageId="14" pageNumber="15">
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<typeStatus id="54D1AEDB9039FFCCF2C7FCF3FECD1B89" box="[234,348,885,911]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="8BD510799039FFCCF290FC1EFAC01BD3" blockId="14.[189,1399,885,982]" pageId="14" pageNumber="15">
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<collectingCountry id="F37D50E99039FFCCF290FC1EFEBD1BB4" box="[189,300,920,946]" name="Brazil" pageId="14" pageNumber="15">BRAZIL</collectingCountry>
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: ♂,
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<location id="8EB546A29039FFCCF376FC1EFDDB1BB4" LSID="urn:lsid:plazi:treatment:03C3A16F9039FFD3F072FE8DFEE91B66:8EB546A29039FFCCF376FC1EFDDB1BB4" box="[347,586,920,946]" country="Brazil" latitude="-3.0333333" longLatPrecision="1307" longitude="-60.283333" municipality="Rio Taruma Mirim" name="Amazonas Province" pageId="14" pageNumber="15">Amazonas Province</location>
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,
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<collectingMunicipality id="6BB18A039039FFCCF07AFC1EFCA91BB4" box="[599,824,920,947]" pageId="14" pageNumber="15">Rio Taruma Mirim</collectingMunicipality>
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,
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<geoCoordinate id="EE5E76BE9039FFCCF168FC1EFC311BB4" box="[837,928,920,946]" degrees="3" direction="south" minutes="02" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="-3.0333333">3°02´S</geoCoordinate>
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,
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<geoCoordinate id="EE5E76BE9039FFCCF183FC1EFBB31BB5" box="[942,1058,920,947]" degrees="60" direction="west" minutes="17" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="-60.283333">60°17´W</geoCoordinate>
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, Capoeira, secondary upland forest, Terra Firma, not inundated,
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<quantity id="4C92BD9C9039FFCCF07AFC3DFD201BD0" box="[599,689,955,982]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="7.0" metricValueMin="0.0" pageId="14" pageNumber="15" unit="cm" value="3.5" valueMax="7.0" valueMin="0.0">0–7 cm</quantity>
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||
, K29 TU,
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<date id="FFD436B99039FFCCF101FC3DFC571BD3" box="[812,966,955,982]" pageId="14" pageNumber="15" value="1983-04-25">
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<collectingDate id="EF90CF519039FFCCF101FC3DFC571BD3" box="[812,966,955,982]" pageId="14" pageNumber="15" value="1983-04-25">25 Apr. 1983</collectingDate>
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</date>
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,
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<collectorName id="269F75AF9039FFCCF1FCFC3DFB991BD3" box="[977,1032,955,981]" pageId="14" pageNumber="15">Adis</collectorName>
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and
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<collectorName id="269F75AF9039FFCCF66CFC3DFB2B1BD3" box="[1089,1210,955,981]" pageId="14" pageNumber="15">Rodrigues</collectorName>
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leg. (
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<collectionCode id="ED7B88BC9039FFCCF6D1FC3DFAD41BD3" box="[1276,1349,955,981]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="14" pageNumber="15">INPA</collectionCode>
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).
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</paragraph>
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</materialsCitation>
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<paragraph id="8BD510799039FFCCF2C7FC7BFEF61C11" blockId="14.[189,1251,1021,1082]" box="[234,359,1021,1047]" pageId="14" pageNumber="15">
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||
<emphasis id="B91ECC6B9039FFCCF2C7FC7BFEF61C11" bold="true" box="[234,359,1021,1047]" pageId="14" pageNumber="15">
|
||
<typeStatus id="54D1AEDB9039FFCCF2C7FC7BFEF61C11" box="[234,359,1021,1047]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="8BD510799039FFCCF290FBA6FB721C3C" blockId="14.[189,1251,1021,1082]" box="[189,1251,1054,1082]" pageId="14" pageNumber="15">
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<collectingCountry id="F37D50E99039FFCCF290FBA6FEBD1C3C" box="[189,300,1056,1082]" name="Brazil" pageId="14" pageNumber="15">BRAZIL</collectingCountry>
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: 1 ♂,
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<specimenCount id="9D6CDBF09039FFCCF35CFBA6FE251C3C" box="[369,436,1056,1082]" pageId="14" pageNumber="15" type="female">2 ♀♀</specimenCount>
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,
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<specimenCount id="9D6CDBF09039FFCCF3EFFBA6FD921C3C" box="[450,515,1056,1082]" pageId="14" pageNumber="15" type="generic">1 juv.</specimenCount>
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♂, same data as for the
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<typeStatus id="54D1AEDB9039FFCCF132FBA6FC171C3C" box="[799,902,1056,1082]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
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(1 ♂ NHMD, the rest INPA).
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</paragraph>
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<paragraph id="8BD510799039FFCCF2C7FBE7FDF81C7A" blockId="14.[189,1398,1120,1218]" box="[234,617,1120,1148]" pageId="14" pageNumber="15">
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<emphasis id="B91ECC6B9039FFCCF2C7FBE7FE2A1C7D" bold="true" box="[234,443,1121,1147]" pageId="14" pageNumber="15">Other specimens</emphasis>
|
||
(4 ♂♂,
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<specimenCount id="9D6CDBF09039FFCCF030FBE4FDCC1C7A" box="[541,605,1122,1148]" pageId="14" pageNumber="15" type="female">7 ♀♀</specimenCount>
|
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)
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</paragraph>
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<paragraph id="8BD510799039FFCCF290FB02FDA11CC4" blockId="14.[189,1398,1120,1218]" pageId="14" pageNumber="15">
|
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All from the same locality with dates between
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<date id="FFD436B99039FFCCF0F4FB03FCDF1C99" box="[729,846,1157,1183]" pageId="14" pageNumber="15" value="1982-04">Apr. 1982</date>
|
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and
|
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<date id="FFD436B99039FFCCF1A9FB03FC6D1C99" box="[900,1020,1157,1183]" pageId="14" pageNumber="15" value="1983-04">Apr. 1983</date>
|
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. Specimens split between INPA, NHMUK, NHMD and ZMUM.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="C37043F29039FFD3F290FB6FFEA61ACB" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15" type="description">
|
||
<paragraph id="8BD510799039FFCCF290FB6FFEDF1D05" blockId="14.[189,334,1257,1283]" box="[189,334,1257,1283]" pageId="14" pageNumber="15">
|
||
<heading id="D09DA7159039FFCCF290FB6FFEDF1D05" bold="true" box="[189,334,1257,1283]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="B91ECC6B9039FFCCF290FB6FFEDF1D05" bold="true" box="[189,334,1257,1283]" pageId="14" pageNumber="15">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD510799039FFCCF290FA9EFBB81D53" blockId="14.[189,1398,1302,1366]" pageId="14" pageNumber="15">
|
||
MEASUREMENTS. Body length ♂:
|
||
<quantity id="4C92BD9C9039FFCCF017FA9EFD2B1D34" box="[570,698,1304,1330]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.9" metricValueMin="1.1" pageId="14" pageNumber="15" unit="mm" value="20.0" valueMax="29.0" valueMin="11.0">11–29 mm</quantity>
|
||
,
|
||
<specimenCount id="9D6CDBF09039FFCCF0EBFA9EFD4F1D34" box="[710,734,1304,1330]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
|
||
: 9–18. Body width ♂:
|
||
<quantity id="4C92BD9C9039FFCCF1C0FA9EFBEF1D34" box="[1005,1150,1304,1330]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.6" metricValueMin="0.8" pageId="14" pageNumber="15" unit="mm" value="1.2" valueMax="1.6" valueMin="0.8">0.8–1.6 mm</quantity>
|
||
,
|
||
<specimenCount id="9D6CDBF09039FFCCF6A4FA9EFB301D34" box="[1161,1185,1304,1330]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
|
||
: 0.8–1.2. Number of podous tergites ♂: 43–85,
|
||
<specimenCount id="9D6CDBF09039FFCCF03EFABDFDBA1D53" box="[531,555,1339,1365]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
|
||
: 41–106. Number of apodous tergites: 0–2.
|
||
</paragraph>
|
||
<paragraph id="8BD510799039FFCCF290FAFAFA9A1D90" blockId="14.[189,1291,1404,1431]" box="[189,1291,1404,1431]" pageId="14" pageNumber="15">COLOUR. Various shades of yellow/white. Perhaps with darker spots near repugnatorial glands.</paragraph>
|
||
<paragraph id="8BD510799039FFCCF290FA38FAF11DDE" blockId="14.[189,1376,1470,1496]" box="[189,1376,1470,1496]" pageId="14" pageNumber="15">BODY SHAPE. More or less parallel-sided in central trunk region. Attenuated anteriorly and posteriorly.</paragraph>
|
||
<paragraph id="8BD510799039FFCCF290F986FAE61EA0" blockId="14.[189,1399,1535,1703]" pageId="14" pageNumber="15">
|
||
HEAD (
|
||
<figureCitation id="13510CFC9039FFCCF323FA79FEC41E1F" box="[270,341,1535,1562]" captionStart="Fig" captionStartId="15.[189,232,1847,1873]" captionTargetBox="[195,1391,967,1802]" captionTargetId="figure@15.[189,1399,960,1810]" captionTargetPageId="15" captionText="Fig. 4. Columbianum nahvalr sp. nov. A. Head and anterior body rings of male, showing the appearance of gonopods in a ‘relaxed specimen’. Body width approximately 1 mm. B. Close up of gonopods in a ‘relaxed’ specimen. Body width approximately 1 mm. C. Head of holotype male in dorsal view. Note long setae originating from base of rostrum. D. Head and rostrum of female in dorsal view. E. Head and rostrum of female in lateral view. Scale bars = C = 0.2 mm; D–E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3825226" httpUri="https://zenodo.org/record/3825226/files/figure.png" pageId="14" pageNumber="15">Fig. 4</figureCitation>
|
||
C–D). Densely pilose all over and covered in regular, sub-circular tubercles, including the antennae. Width (between antennal sockets)
|
||
<quantity id="4C92BD9C9039FFCCF09EF9A4FCAF1E3B" box="[691,830,1570,1597]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="14" pageNumber="15" unit="mm" value="0.44999999999999996" valueMax="0.6" valueMin="0.3">0.3–0.6 mm</quantity>
|
||
. Head very sharply defined from rostrum in dorsal view, slightly rounded anteriorly. In lateral view head so domed that an abrupt ‘step’ is formed between head and rostrum. Length of rostrum to antennal base:
|
||
<quantity id="4C92BD9C9039FFCCF0DEF9EFFC111E85" box="[755,896,1641,1668]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="14" pageNumber="15" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.5–0.8 mm</quantity>
|
||
. Rostrum slightly downcurved, with several long setae at base, almost reaching tip of rostrum and smaller lateral ones on both sides of the whole length.
|
||
</paragraph>
|
||
<paragraph id="8BD510799039FFCCF290F948FE0C1F57" blockId="14.[189,1399,1741,1873]" pageId="14" pageNumber="15">ANTENNAE. About the same as the body width (1:1.06), long in relation to rostrum, clearly surpassing it by over 100% of the rostrum length. Length:width ratio of antennomeres 1–8: 0.82, 1.3, 1.12, 0.95, 0.96, 1.6, 0.45, sensory cones at apex not measured. Sensory pits on antennomeres 5 and 6, with a large number of sensilla.</paragraph>
|
||
<paragraph id="8BD510799039FFCCF290F8FFFCF11FDF" blockId="14.[189,1399,1912,2009]" pageId="14" pageNumber="15">COLLUM. Covered with tubercles and setae; anterior margin appearing sharply V-shaped in some specimens but in reality a thinner region anteriorly with a shallow posterior curve (also seen under SEM). Tergite 2 much less than half the length of collum.</paragraph>
|
||
<paragraph id="8BD510799038FFCDF290FE8AFAB11B33" blockId="15.[189,1399,267,822]" pageId="15" pageNumber="16">
|
||
MIDBODY SEGMENTS. When the number of tergites are plotted relative to body length this species forms an intermediate group on the graphs (
|
||
<figureCitation id="13510CFC9038FFCDF050FEA8FD52194F" box="[637,707,302,329]" captionStart="Fig" captionStartId="11.[189,232,1945,1971]" captionTargetBox="[203,1385,344,1859]" captionTargetId="figure@11.[202,1393,333,1873]" captionTargetPageId="11" captionText="Fig. 1. Body width (mm) against the total number of tergites with legs for new species of Columbianum Verhoeff, 1941. A. Males. B. Females." figureDoi="http://doi.org/10.5281/zenodo.3825220" httpUri="https://zenodo.org/record/3825220/files/figure.png" pageId="15" pageNumber="16">Fig. 1</figureCitation>
|
||
). No obvious paranota-type structure, although there may be slight bumps where the repugnatorial glands exit, appearing more pronounced by a ring of ‘palisadelike’ setae. In dorsal view it appears as if there are small paranota but these consist of the metazonital ridges. Clear differences in height of pro- and metazonite so that the metazonite stands proud, and the whole appearance when viewing the animal from the side is of a castellated dorsal margin (
|
||
<figureCitation id="13510CFC9038FFCDF724FE3DFAF819D0" box="[1289,1385,443,470]" captionStart="Fig" captionStartId="16.[189,232,1807,1833]" captionTargetBox="[197,1387,751,1756]" captionTargetId="figure@16.[189,1399,745,1767]" captionTargetPageId="16" captionText="Fig. 5. Columbianum nahvalr sp. nov.A. Midbody rings from lateral view, showing pronounced difference between pro- and metazonite and shape of pleurites. B. ‘Channel’ between pro- and metazonite in dorsal view of male showing tubercles and line of setae. C. Tarsal claw and accessory claw of midbody leg from right side of male, in anterior dorsal view. D. Gonopods of holotype male, lateral view with anterior gonopods to right. E. Gonopods of small, presumably immature male, in lateral view, anterior gonopds to left. Scale bars = A, D–E = 0.2 mm; B–C = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3825230" httpUri="https://zenodo.org/record/3825230/files/figure.png" pageId="15" pageNumber="16">Fig. 5A</figureCitation>
|
||
). Relative width of pro- and metazonite: 0.78–0.88. Prozonites with cytoscutes but without any tubercles and lacking setae. Metazonites with numerous tubercles and setae. Under SEM tubercles very uneven in shape. Setae rather long and uneven, giving the animal a slightly ragged look. Setae apparently of similar length regardless of the size of the individual so appearing longer in smaller specimens. ‘Channel’ between pro- and metazonite with tubercles appearing horseshoe-shaped; possibly due to the partial collapse of more blister-like structures (
|
||
<figureCitation id="13510CFC9038FFCDF101FD08FC1C1AAF" box="[812,909,654,681]" captionStart="Fig" captionStartId="16.[189,232,1807,1833]" captionTargetBox="[197,1387,751,1756]" captionTargetId="figure@16.[189,1399,745,1767]" captionTargetPageId="16" captionText="Fig. 5. Columbianum nahvalr sp. nov.A. Midbody rings from lateral view, showing pronounced difference between pro- and metazonite and shape of pleurites. B. ‘Channel’ between pro- and metazonite in dorsal view of male showing tubercles and line of setae. C. Tarsal claw and accessory claw of midbody leg from right side of male, in anterior dorsal view. D. Gonopods of holotype male, lateral view with anterior gonopods to right. E. Gonopods of small, presumably immature male, in lateral view, anterior gonopds to left. Scale bars = A, D–E = 0.2 mm; B–C = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3825230" httpUri="https://zenodo.org/record/3825230/files/figure.png" pageId="15" pageNumber="16">Fig. 5B</figureCitation>
|
||
). Setae 0.04–0.1 × body width. Limbus crenulated. Pleurites setose and with curved posterior margin, ventral margin bilobed, hind margin crenulated. Ozopores present from tergite 5 to the ultimate tergite, on tergal ridges, in anterior ¼ located more or less centrally on ridge, in posterior ¾ on posterior margin of ridge. Some appearing on very slight protuberance probably largely due to the slight change in angle of the tergite at this point.
|
||
</paragraph>
|
||
<paragraph id="8BD510799038FFD2F290FCDBFAB419D3" blockId="15.[189,1398,861,923]" lastBlockId="16.[189,1399,267,470]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
|
||
LEGS. Length 0.57 × body width. Length:width ratio of podomeres (prefemur to claw): 1.54, 1.69, 1.1, 1.47, 3.31, 4.25. Tarsus very long and narrow. Claw rather angular in shape and with slightly thickened region at base on anterior side. Accessory claw slightly shorter than claw but substantial in size and thickness, arising from anterior-dorsal side of the claw (
|
||
<figureCitation id="13510CFC9027FFD2F14FFEA8FC53194F" box="[866,962,302,329]" captionStart="Fig" captionStartId="16.[189,232,1807,1833]" captionTargetBox="[197,1387,751,1756]" captionTargetId="figure@16.[189,1399,745,1767]" captionTargetPageId="16" captionText="Fig. 5. Columbianum nahvalr sp. nov.A. Midbody rings from lateral view, showing pronounced difference between pro- and metazonite and shape of pleurites. B. ‘Channel’ between pro- and metazonite in dorsal view of male showing tubercles and line of setae. C. Tarsal claw and accessory claw of midbody leg from right side of male, in anterior dorsal view. D. Gonopods of holotype male, lateral view with anterior gonopods to right. E. Gonopods of small, presumably immature male, in lateral view, anterior gonopds to left. Scale bars = A, D–E = 0.2 mm; B–C = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3825230" httpUri="https://zenodo.org/record/3825230/files/figure.png" pageId="16" pageNumber="17">Fig. 5C</figureCitation>
|
||
). Accessory claw can be seen under light microscope and makes it appear as if there is a double claw at the end of the tarsus when seen in dorsal view, less obvious in anterior view because the accessory claw does not project beyond the claw. In females these ‘double claws’ only seem to be obvious at the anterior end of the body, in males they are apparent almost the whole length. Coxal pores present and large at least on some specimens.
|
||
</paragraph>
|
||
<caption id="DF1540F19038FFCDF290F8B1FBA51FD9" ID-DOI="http://doi.org/10.5281/zenodo.3825226" ID-Zenodo-Dep="3825226" httpUri="https://zenodo.org/record/3825226/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,232,1847,1873]" targetBox="[195,1391,967,1802]" targetPageId="15">
|
||
<paragraph id="8BD510799038FFCDF290F8B1FBA51FD9" blockId="15.[189,1399,1847,2015]" pageId="15" pageNumber="16">
|
||
<emphasis id="B91ECC6B9038FFCDF290F8B1FE981F57" bold="true" box="[189,265,1847,1873]" pageId="15" pageNumber="16">Fig. 4.</emphasis>
|
||
<taxonomicName id="4C6A6BFA9038FFCDF33DF8B1FD861F57" authorityName="Read & Enghoff" authorityYear="2018" box="[272,535,1847,1873]" class="Diplopoda" family="Siphonophoridae" genus="Columbianum" kingdom="Animalia" order="Siphonophorida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nahvalr" status="sp. nov.">
|
||
<emphasis id="B91ECC6B9038FFCDF33DF8B1FD861F57" box="[272,535,1847,1873]" italics="true" pageId="15" pageNumber="16">Columbianum nahvalr</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22D71109038FFCDF033F8BEFDE91F54" box="[542,632,1848,1874]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B91ECC6B9038FFCDF053F8B1FD021F57" bold="true" box="[638,659,1847,1873]" pageId="15" pageNumber="16">A</emphasis>
|
||
. Head and anterior body rings of male, showing the appearance of gonopods in a ‘relaxed specimen’. Body width approximately 1 mm.
|
||
<emphasis id="B91ECC6B9038FFCDF637F8DDFBBF1F73" bold="true" box="[1050,1070,1883,1909]" pageId="15" pageNumber="16">B</emphasis>
|
||
. Close up of gonopods in a ‘relaxed’ specimen. Body width approximately 1 mm.
|
||
<emphasis id="B91ECC6B9038FFCDF165F8FBFCCC1F91" bold="true" box="[840,861,1917,1943]" pageId="15" pageNumber="16">C</emphasis>
|
||
. Head of holotype male in dorsal view. Note long setae originating from base of rostrum.
|
||
<emphasis id="B91ECC6B9038FFCDF0E9F827FD481FBD" bold="true" box="[708,729,1953,1979]" pageId="15" pageNumber="16">D</emphasis>
|
||
. Head and rostrum of female in dorsal view.
|
||
<emphasis id="B91ECC6B9038FFCDF6CBF827FB6B1FBD" bold="true" box="[1254,1274,1953,1979]" pageId="15" pageNumber="16">E</emphasis>
|
||
. Head and rostrum of female in lateral view. Scale bars = C = 0.2 mm; D–E = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD510799027FFD2F290FE7BFD521A11" blockId="16.[189,707,509,535]" box="[189,707,509,535]" pageId="16" pageNumber="17">TELSON. Rounded, length:width ratio: 1–2.4.</paragraph>
|
||
<paragraph id="8BD510799027FFD2F290FDB9FD761A5E" blockId="16.[189,743,574,601]" box="[189,743,574,601]" pageId="16" pageNumber="17">MALE FIRST PAIRS OF LEGS (1–8). Appear normal.</paragraph>
|
||
<paragraph id="8BD510799027FFD2F290FD06FAE71ABB" blockId="16.[189,1399,639,701]" pageId="16" pageNumber="17">
|
||
ANTERIOR GONOPODS (
|
||
<figureCitation id="13510CFC9027FFD2F398FD06FD851A9C" box="[437,532,640,666]" captionStart="Fig" captionStartId="15.[189,232,1847,1873]" captionTargetBox="[195,1391,967,1802]" captionTargetId="figure@15.[189,1399,960,1810]" captionTargetPageId="15" captionText="Fig. 4. Columbianum nahvalr sp. nov. A. Head and anterior body rings of male, showing the appearance of gonopods in a ‘relaxed specimen’. Body width approximately 1 mm. B. Close up of gonopods in a ‘relaxed’ specimen. Body width approximately 1 mm. C. Head of holotype male in dorsal view. Note long setae originating from base of rostrum. D. Head and rostrum of female in dorsal view. E. Head and rostrum of female in lateral view. Scale bars = C = 0.2 mm; D–E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3825226" httpUri="https://zenodo.org/record/3825226/files/figure.png" pageId="16" pageNumber="17">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="13510CFC9027FFD2F00FFDF9FDBE1A9C" box="[546,559,639,666]" captionStart="Fig" captionStartId="16.[189,232,1807,1833]" captionTargetBox="[197,1387,751,1756]" captionTargetId="figure@16.[189,1399,745,1767]" captionTargetPageId="16" captionText="Fig. 5. Columbianum nahvalr sp. nov.A. Midbody rings from lateral view, showing pronounced difference between pro- and metazonite and shape of pleurites. B. ‘Channel’ between pro- and metazonite in dorsal view of male showing tubercles and line of setae. C. Tarsal claw and accessory claw of midbody leg from right side of male, in anterior dorsal view. D. Gonopods of holotype male, lateral view with anterior gonopods to right. E. Gonopods of small, presumably immature male, in lateral view, anterior gonopds to left. Scale bars = A, D–E = 0.2 mm; B–C = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3825230" httpUri="https://zenodo.org/record/3825230/files/figure.png" pageId="16" pageNumber="17">5</figureCitation>
|
||
D–E). With five segments (including coxa). Distally a simple claw, the whole structure very reminiscent of a normal walking leg but rather stouter and with especially stout
|
||
</paragraph>
|
||
<caption id="DF1540F19027FFD2F290F889FCDC1FDC" ID-DOI="http://doi.org/10.5281/zenodo.3825230" ID-Zenodo-Dep="3825230" httpUri="https://zenodo.org/record/3825230/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,232,1807,1833]" targetBox="[197,1387,751,1756]" targetPageId="16">
|
||
<paragraph id="8BD510799027FFD2F290F889FCDC1FDC" blockId="16.[189,1399,1807,2010]" pageId="16" pageNumber="17">
|
||
<emphasis id="B91ECC6B9027FFD2F290F889FE961F2C" bold="true" box="[189,263,1807,1834]" pageId="16" pageNumber="17">Fig. 5.</emphasis>
|
||
<taxonomicName id="4C6A6BFA9027FFD2F326F889FD811F2F" authorityName="Read & Enghoff" authorityYear="2018" box="[267,528,1807,1833]" class="Diplopoda" family="Siphonophoridae" genus="Columbianum" kingdom="Animalia" order="Siphonophorida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="nahvalr" status="sp. nov.">
|
||
<emphasis id="B91ECC6B9027FFD2F326F889FD811F2F" box="[267,528,1807,1833]" italics="true" pageId="16" pageNumber="17">Columbianum nahvalr</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22D71109027FFD2F039F896FDFD1F2C" box="[532,620,1808,1834]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B91ECC6B9027FFD2F042F889FD151F2F" bold="true" box="[623,644,1807,1833]" pageId="16" pageNumber="17">A</emphasis>
|
||
. Midbody rings from lateral view, showing pronounced difference between pro- and metazonite and shape of pleurites.
|
||
<emphasis id="B91ECC6B9027FFD2F10DF8B5FCA51F4B" bold="true" box="[800,820,1843,1869]" pageId="16" pageNumber="17">B</emphasis>
|
||
. ‘Channel’ between pro- and metazonite in dorsal view of male showing tubercles and line of setae.
|
||
<emphasis id="B91ECC6B9027FFD2F134F8D0FCBF1F76" bold="true" box="[793,814,1878,1904]" pageId="16" pageNumber="17">C</emphasis>
|
||
. Tarsal claw and accessory claw of midbody leg from right side of male, in anterior dorsal view.
|
||
<emphasis id="B91ECC6B9027FFD2F0CFF8FFFD661F95" bold="true" box="[738,759,1913,1939]" pageId="16" pageNumber="17">D</emphasis>
|
||
. Gonopods of holotype male, lateral view with anterior gonopods to right.
|
||
<emphasis id="B91ECC6B9027FFD2F3B6F81BFE3E1FB1" bold="true" box="[411,431,1949,1975]" pageId="16" pageNumber="17">E</emphasis>
|
||
. Gonopods of small, presumably immature male, in lateral view, anterior gonopds to left. Scale bars = A, D–E = 0.2 mm; B–C = 0.01 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD510799026FFD3F290FE8DFD6B196A" blockId="17.[189,1399,267,364]" pageId="17" pageNumber="18">and bent tibia. Abundant setae on terminal two podomeres, including a tuft of short setae just below the base of the last segment, on the posterior side and more setae on the top of the ‘knee’. Several prominent setae on anterior margin of basal three segments.</paragraph>
|
||
<paragraph id="8BD510799026FFD3F290FE12FD2919F5" blockId="17.[189,1399,403,500]" pageId="17" pageNumber="18">
|
||
POSTERIOR GONOPODS (
|
||
<figureCitation id="13510CFC9026FFD3F394FE15FD8719AB" box="[441,534,403,429]" captionStart="Fig" captionStartId="15.[189,232,1847,1873]" captionTargetBox="[195,1391,967,1802]" captionTargetId="figure@15.[189,1399,960,1810]" captionTargetPageId="15" captionText="Fig. 4. Columbianum nahvalr sp. nov. A. Head and anterior body rings of male, showing the appearance of gonopods in a ‘relaxed specimen’. Body width approximately 1 mm. B. Close up of gonopods in a ‘relaxed’ specimen. Body width approximately 1 mm. C. Head of holotype male in dorsal view. Note long setae originating from base of rostrum. D. Head and rostrum of female in dorsal view. E. Head and rostrum of female in lateral view. Scale bars = C = 0.2 mm; D–E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3825226" httpUri="https://zenodo.org/record/3825226/files/figure.png" pageId="17" pageNumber="18">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="13510CFC9026FFD3F00FFE15FDA019A8" box="[546,561,403,430]" captionStart="Fig" captionStartId="16.[189,232,1807,1833]" captionTargetBox="[197,1387,751,1756]" captionTargetId="figure@16.[189,1399,745,1767]" captionTargetPageId="16" captionText="Fig. 5. Columbianum nahvalr sp. nov.A. Midbody rings from lateral view, showing pronounced difference between pro- and metazonite and shape of pleurites. B. ‘Channel’ between pro- and metazonite in dorsal view of male showing tubercles and line of setae. C. Tarsal claw and accessory claw of midbody leg from right side of male, in anterior dorsal view. D. Gonopods of holotype male, lateral view with anterior gonopods to right. E. Gonopods of small, presumably immature male, in lateral view, anterior gonopds to left. Scale bars = A, D–E = 0.2 mm; B–C = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3825230" httpUri="https://zenodo.org/record/3825230/files/figure.png" pageId="17" pageNumber="18">5</figureCitation>
|
||
D–E). Number of segments may be 5–7 (including coxa) but distinction between segments unclear. Distal part a simple hyaline tip with slight notch and two small ‘nicks’ towards the tip at least on some specimens.
|
||
</paragraph>
|
||
<paragraph id="8BD510799026FFD3F290FD9DFF691A33" blockId="17.[189,248,539,565]" box="[189,248,539,565]" pageId="17" pageNumber="18">
|
||
<heading id="D09DA7159026FFD3F290FD9DFF691A33" bold="true" box="[189,248,539,565]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="B91ECC6B9026FFD3F290FD9DFF691A33" bold="true" box="[189,248,539,565]" pageId="17" pageNumber="18">Note</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD510799026FFD3F290FDCFFEA61ACB" blockId="17.[189,1399,585,717]" pageId="17" pageNumber="18">
|
||
The gonopods of this species appear from the drawings (
|
||
<figureCitation id="13510CFC9026FFD3F14AFDCFFC541A62" box="[871,965,585,612]" captionStart="Fig" captionStartId="16.[189,232,1807,1833]" captionTargetBox="[197,1387,751,1756]" captionTargetId="figure@16.[189,1399,745,1767]" captionTargetPageId="16" captionText="Fig. 5. Columbianum nahvalr sp. nov.A. Midbody rings from lateral view, showing pronounced difference between pro- and metazonite and shape of pleurites. B. ‘Channel’ between pro- and metazonite in dorsal view of male showing tubercles and line of setae. C. Tarsal claw and accessory claw of midbody leg from right side of male, in anterior dorsal view. D. Gonopods of holotype male, lateral view with anterior gonopods to right. E. Gonopods of small, presumably immature male, in lateral view, anterior gonopds to left. Scale bars = A, D–E = 0.2 mm; B–C = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3825230" httpUri="https://zenodo.org/record/3825230/files/figure.png" pageId="17" pageNumber="18">Fig. 5D</figureCitation>
|
||
) to be clearly distinct from those of
|
||
<taxonomicName id="4C6A6BFA9026FFD3F290FDEBFEB41A81" authorityName="Read & Enghoff" authorityYear="2018" box="[189,293,621,647]" class="Diplopoda" family="Siphonophoridae" genus="Columbianum" kingdom="Animalia" order="Siphonophorida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="major" status="sp. nov.">
|
||
<emphasis id="B91ECC6B9026FFD3F290FDEBFEB41A81" box="[189,293,621,647]" italics="true" pageId="17" pageNumber="18">C. major</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22D71109026FFD3F301FDEBFE171A81" box="[300,390,621,647]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13510CFC9026FFD3F3B8FDEBFE4A1A81" box="[405,475,621,647]" captionStart="Fig" captionStartId="13.[189,232,1845,1871]" captionTargetBox="[203,1371,913,1789]" captionTargetId="figure@13.[189,1399,910,1806]" captionTargetPageId="13" captionText="Fig. 3. Columbianum major sp. nov. A. Tarsal claw and accessory claw of midbody leg from right side of male, anterior view. B–F. Gonopods of holotype male. B. In situ (apex of posterior gonopod hidden behind the anterior gonopods). C. Anterior gonopod in lateral view. D. Anterior gonopod in mesal view. E. Posterior view in lateral view (apex broken). F. Posterior gonopods in mesal view (apex broken). Scale bars = A = 0.01 mm; B–F = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3825224" httpUri="https://zenodo.org/record/3825224/files/figure.png" pageId="17" pageNumber="18">Fig. 3</figureCitation>
|
||
). In reality this is because all specimens of
|
||
<taxonomicName id="4C6A6BFA9026FFD3F1FBFDEBFBC21A81" authorityName="Read & Enghoff" authorityYear="2018" box="[982,1107,621,647]" class="Diplopoda" family="Siphonophoridae" genus="Columbianum" kingdom="Animalia" order="Siphonophorida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="nahvalr" status="sp. nov.">
|
||
<emphasis id="B91ECC6B9026FFD3F1FBFDEBFBC21A81" box="[982,1107,621,647]" italics="true" pageId="17" pageNumber="18">C. nahvalr</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22D71109026FFD3F677FDEBFB251A81" box="[1114,1204,621,647]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are very relaxed and the gonopods much easier to see; those of
|
||
<taxonomicName id="4C6A6BFA9026FFD3F0CDFD16FCD91AAC" authorityName="Read & Enghoff" authorityYear="2018" box="[736,840,656,682]" class="Diplopoda" family="Siphonophoridae" genus="Columbianum" kingdom="Animalia" order="Siphonophorida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="major" status="sp. nov.">
|
||
<emphasis id="B91ECC6B9026FFD3F0CDFD16FCD91AAC" box="[736,840,656,682]" italics="true" pageId="17" pageNumber="18">C. major</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22D71109026FFD3F162FD16FC3B1AAC" box="[847,938,656,682]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are tightly compressed and also poorly preserved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37043F29026FFD3F290FD72FEE91B66" pageId="17" pageNumber="18" type="discussion">
|
||
<paragraph id="8BD510799026FFD3F290FD72FE5E1B09" blockId="17.[189,463,756,783]" box="[189,463,756,783]" pageId="17" pageNumber="18">
|
||
<heading id="D09DA7159026FFD3F290FD72FE5E1B09" bold="true" box="[189,463,756,783]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="B91ECC6B9026FFD3F290FD72FE5E1B09" bold="true" box="[189,463,756,783]" pageId="17" pageNumber="18">Comments on ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD510799026FFD3F290FCA5FEE91B66" blockId="17.[189,1399,803,864]" pageId="17" pageNumber="18">
|
||
All specimens of
|
||
<taxonomicName id="4C6A6BFA9026FFD3F3ABFCA2FD931B3B" authorityName="Read & Enghoff" authorityYear="2018" box="[390,514,803,830]" class="Diplopoda" family="Siphonophoridae" genus="Columbianum" kingdom="Animalia" order="Siphonophorida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="nahvalr" status="sp. nov.">
|
||
<emphasis id="B91ECC6B9026FFD3F3ABFCA2FD931B3B" box="[390,514,803,830]" italics="true" pageId="17" pageNumber="18">C. nahvalr</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22D71109026FFD3F02AFCA2FDF11B38" box="[519,608,804,830]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
were obtained from Kempson extractors suggesting that this species is soil dwelling.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |