537 lines
83 KiB
XML
537 lines
83 KiB
XML
<document id="A9CAD6297D680AA464396195356499E6" ID-CLB-Dataset="53164" ID-DOI="10.11646/zootaxa.5124.1.1" ID-GBIF-Dataset="9666d599-2c7d-4a52-bce2-a8794f03265b" ID-ISSN="1175-5326" ID-Zenodo-Dep="6404674" ID-ZooBank="C015F8CB-799B-4A92-90AE-02B4C576089E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648795871367" checkinUser="plazi" docAuthor="O’Hara, Timothy D. & Thuy, Ben" docDate="2022" docId="C72D87A15342FF8FFEA385DCFEBCFBF7" docLanguage="en" docName="zootaxa.5124.1.1.pdf" docOrigin="Zootaxa 5124 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ophiocomina arnaudi O’Hara & Thuy 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="32" masterDocId="3B14FFD9535EFF90FE348503FFA8FFAE" masterDocTitle="Biogeography and taxonomy of Ophiuroidea (Echinodermata) from the Îles Saint- Paul and Amsterdam in the southern Indian Ocean" masterLastPageNumber="49" masterPageNumber="1" pageNumber="29" updateTime="1699310728982" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F248ABD6576C1C0DC1A6FFE2FBE95E16">Biogeography and taxonomy of Ophiuroidea (Echinodermata) from the Îles Saint- Paul and Amsterdam in the southern Indian Ocean</mods:title>
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<mods:name id="D3084255B3B95514D713E3A05640B629" type="personal">
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<mods:namePart id="0FE7C374C0C76D47A1796C304C7328E0">O’Hara, Timothy D.</mods:namePart>
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<mods:affiliation id="8C7F4794A70F3C8D319794782299C8C0">Museums Victoria, GPO Box 666 E, Melbourne, 3001, AUSTRALIA,</mods:affiliation>
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<mods:namePart id="935D728ED7FA2B2DA5C79D797776F01F">Thuy, Ben</mods:namePart>
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<mods:affiliation id="3AC9B713423CFE81B2C7B4D69F4A6F9C">Natural History Museum of Luxembourg, 24 Rue Münster, 2160 Luxembourg</mods:affiliation>
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<mods:title id="9ADDA49622A8C20936AE748B166E229A">Zootaxa</mods:title>
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<mods:part id="0BE198D09F6C31A7ABACACF98BB49C3C">
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<mods:date id="F1529E1CE2E7BB282EED0AC4E7DFC68C">2022</mods:date>
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<treatment id="C72D87A15342FF8FFEA385DCFEBCFBF7" ID-DOI="http://doi.org/10.5281/zenodo.6404726" ID-GBIF-Taxon="194442346" ID-Zenodo-Dep="6404726" LSID="urn:lsid:plazi:treatment:C72D87A15342FF8FFEA385DCFEBCFBF7" httpUri="http://treatment.plazi.org/id/C72D87A15342FF8FFEA385DCFEBCFBF7" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29">
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<subSubSection id="079E653C5342FF8CFEA385DCFDA0FF57" box="[151,520,223,250]" pageId="28" pageNumber="29" type="nomenclature">
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<paragraph id="4F3B36B75342FF8CFEA385DCFDA0FF57" blockId="28.[151,1197,223,322]" box="[151,520,223,250]" pageId="28" pageNumber="29">
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<heading id="147381DB5342FF8CFEA385DCFDA0FF57" bold="true" box="[151,520,223,250]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
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<emphasis id="7DF0EAA55342FF8CFEA385DCFDA0FF57" bold="true" box="[151,520,223,250]" pageId="28" pageNumber="29">
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<taxonomicName id="88844D345342FF8CFEA385DCFE0BFF54" ID-CoL="8ZDNS" authority="O’Hara & Thuy, 2022" authorityName="O’Hara & Thuy" authorityYear="2022" box="[151,419,223,250]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiocomina" kingdom="Animalia" order="Ophiacanthida" pageId="28" pageNumber="29" phylum="Echinodermata" rank="species" species="arnaudi" status="sp. nov.">
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<emphasis id="7DF0EAA55342FF8CFEA385DCFE0BFF54" bold="true" box="[151,419,223,250]" italics="true" pageId="28" pageNumber="29">Ophiocomina arnaudi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="66C357DE5342FF8CFF9F85DCFDA0FF57" box="[427,520,223,249]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="079E653C5342FF8CFEA38400FB05FEEC" pageId="28" pageNumber="29" type="description">
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<paragraph id="4F3B36B75342FF8CFEA38400FED9FEB3" blockId="28.[151,1197,223,322]" box="[151,369,259,285]" pageId="28" pageNumber="29">
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<figureCitation id="D7BF2A325342FF8CFEA38400FEBFFEB3" box="[151,279,259,285]" captionStart="FIGURE 11" captionStartId="28.[151,250,1801,1825]" captionTargetBox="[152,1435,921,1777]" captionTargetId="figure-264@28.[151,1436,921,1777]" captionTargetPageId="28" captionText="FIGURE 11. (A–E) Ophiocomina arnaudi sp. nov., holotype, MNHN IE.2019.4747,A: dorsal outline, B: dorsal view, C: ventral view; D: inset of marginal disc granules, E: inset of central disc granules. (F) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599, lateral view of two jaws. Abbreviations: T=tooth, TPa=tooth papillae, IPa=infradental papillae, LOPa=lateral oral papillae, AdShSp=adoral shield spine." figureDoi="http://doi.org/10.5281/zenodo.6404706" httpUri="https://zenodo.org/record/6404706/files/figure.png" pageId="28" pageNumber="29">Fig. 11A–F</figureCitation>
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,
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<figureCitation id="D7BF2A325342FF8CFF178400FED9FEB3" box="[291,369,259,285]" captionStart="FIGURE 12" captionStartId="30.[151,250,1809,1833]" captionTargetBox="[183,1405,181,1785]" captionTargetId="figure-26@30.[183,1405,181,1785]" captionTargetPageId="30" captionText="FIGURE 12. (A–S) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599.A–B: proximal lateral arm plate in external (A) and internal (B) views. C: distal lateral arm plate in external view. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N–O: teeth. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404710" httpUri="https://zenodo.org/record/6404710/files/figure.png" pageId="28" pageNumber="29">12A–S</figureCitation>
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</paragraph>
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<paragraph id="4F3B36B75342FF8CFEA38424FB05FEEC" blockId="28.[151,1197,223,322]" box="[151,1197,295,322]" pageId="28" pageNumber="29">
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="079E653C5342FF8FFEA3846CFC53FEEF" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29" type="materials_examined">
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<paragraph id="4F3B36B75342FF8CFEA3846CFBFEFE00" blockId="28.[151,1437,367,898]" pageId="28" pageNumber="29">
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<emphasis id="7DF0EAA55342FF8CFEA3846CFE96FE27" bold="true" box="[151,318,367,393]" pageId="28" pageNumber="29">Type material</emphasis>
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.
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<materialsCitation id="FFEC3CEA5342FF8CFF78846CFEA3FE03" ID-GBIF-Occurrence="3746276312" collectingDate="1986-07-18" collectionCode="MD" elevation="485" latitude="-38.813835" longLatPrecision="11" longitude="77.594666" pageId="28" pageNumber="29" specimenCode="MD50" specimenCount="1">
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<specimenCode id="1F229ECC5342FF8CFF78846CFE3DFE24" box="[332,405,367,394]" collectionCode="MD" country="Germany" name="Museum Donaueschingen" pageId="28" pageNumber="29">MD50</specimenCode>
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DC108, south-east of Île Saint-Paul,
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,
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,
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<quantity id="887C9B525342FF8CFA57846CFB76FE24" box="[1123,1246,367,394]" metricMagnitude="2" metricUnit="m" metricValue="4.85" metricValueMax="5.1" metricValueMin="4.6" pageId="28" pageNumber="29" unit="m" value="485.0" valueMax="510.0" valueMin="460.0">
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</quantity>
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, Charcot dredge,
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<date id="3B3A10775342FF8CFEA38490FEA3FE03" box="[151,267,403,429]" pageId="28" pageNumber="29" value="1986-07-18">
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<collectingDate id="2B7EE99F5342FF8CFEA38490FEA3FE03" box="[151,267,403,429]" pageId="28" pageNumber="29" value="1986-07-18">18/7/1986</collectingDate>
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</date>
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,
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<materialsCitation id="FFEC3CEA5342FF8CFF238497FD0EFE00" ID-GBIF-Occurrence="3746276315" box="[279,678,403,430]" collectionCode="MNHN, IE" pageId="28" pageNumber="29" specimenCount="1" typeStatus="holotype">
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<collectionCode id="2995AE725342FF8CFF238497FEC5FE00" box="[279,365,404,430]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">MNHN</collectionCode>
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<collectionCode id="2995AE725342FF8CFF478490FE26FE03" box="[371,398,403,429]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="28" pageNumber="29">IE</collectionCode>
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.202019.4747 (
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<typeStatus id="903F88155342FF8CFC038490FD37FE00" box="[567,671,403,430]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
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)
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</materialsCitation>
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,
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<materialsCitation id="FFEC3CEA5342FF8CFC858497FBFAFE00" ID-GBIF-Occurrence="3746276308" box="[689,1106,403,430]" collectionCode="MNHN, IE" pageId="28" pageNumber="29" specimenCount="52" typeStatus="paratype">
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<collectionCode id="2995AE725342FF8CFC858497FCAFFE00" box="[689,775,404,430]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">MNHN</collectionCode>
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<collectionCode id="2995AE725342FF8CFD3A8490FC81FE00" box="[782,809,403,430]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="28" pageNumber="29">IE</collectionCode>
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.2009.1599 (
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<specimenCount id="5982FD3E5342FF8CFD818490FBE3FE00" box="[949,1099,403,430]" pageId="28" pageNumber="29" type="generic" typeStatus="paratypes">52 paratypes</specimenCount>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="4F3B36B75342FF8CFEF384B4FABAFD93" blockId="28.[151,1437,367,898]" pageId="28" pageNumber="29">
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<emphasis id="7DF0EAA55342FF8CFEF384B4FE55FE7F" bold="true" box="[199,509,439,465]" pageId="28" pageNumber="29">Other material examined.</emphasis>
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<collectionCode id="2995AE725342FF8CFC3184B4FD9AFE7C" box="[517,562,439,466]" country="Germany" name="Museum Donaueschingen" pageId="28" pageNumber="29">MD</collectionCode>
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50 DC71,
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<collectionCode id="2995AE725342FF8CFC9384BBFD55FE7C" box="[679,765,440,466]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">MNHN</collectionCode>
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<collectionCode id="2995AE725342FF8CFD3184B4FC88FE7C" box="[773,800,439,466]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="28" pageNumber="29">IE</collectionCode>
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.2009.1594 (2).
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<collectionCode id="2995AE725342FF8CFDE084B4FBA9FE7C" box="[980,1025,439,466]" country="Germany" name="Museum Donaueschingen" pageId="28" pageNumber="29">MD</collectionCode>
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50 DC79,
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<collectionCode id="2995AE725342FF8CFA4384BBFB65FE7C" box="[1143,1229,440,466]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">MNHN</collectionCode>
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<collectionCode id="2995AE725342FF8CFAE084B4FB47FE7C" box="[1236,1263,439,466]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="28" pageNumber="29">IE</collectionCode>
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.2009.1595 (2).
|
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<collectionCode id="2995AE725342FF8CFEA384D8FF6CFE58" box="[151,196,475,502]" country="Germany" name="Museum Donaueschingen" pageId="28" pageNumber="29">MD</collectionCode>
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50 CP80,
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<collectionCode id="2995AE725342FF8CFF1A84DFFE2CFE58" box="[302,388,476,502]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">MNHN</collectionCode>
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<collectionCode id="2995AE725342FF8CFFBC84D8FE0BFE58" box="[392,419,475,502]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="28" pageNumber="29">IE</collectionCode>
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.2009.1596 (6).
|
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<collectionCode id="2995AE725342FF8CFC6484D8FDD5FE58" box="[592,637,475,502]" country="Germany" name="Museum Donaueschingen" pageId="28" pageNumber="29">MD</collectionCode>
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50 DC82,
|
||
<collectionCode id="2995AE725342FF8CFCD884DFFCEAFE58" box="[748,834,476,502]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">MNHN</collectionCode>
|
||
<collectionCode id="2995AE725342FF8CFD7284D8FCC9FE58" box="[838,865,475,502]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="28" pageNumber="29">IE</collectionCode>
|
||
.2009.1597 (3).
|
||
<collectionCode id="2995AE725342FF8CFA3A84D8FB93FE58" box="[1038,1083,475,502]" country="Germany" name="Museum Donaueschingen" pageId="28" pageNumber="29">MD</collectionCode>
|
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50 DC99,
|
||
<collectionCode id="2995AE725342FF8CFA9E84DFFAA8FE58" box="[1194,1280,476,502]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">MNHN</collectionCode>
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<collectionCode id="2995AE725342FF8CFB3084D8FAB7FE58" box="[1284,1311,475,502]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="28" pageNumber="29">IE</collectionCode>
|
||
.2009.1598 (2).
|
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<collectionCode id="2995AE725342FF8CFEFD84FCFF5EFDB4" box="[201,246,511,538]" country="Germany" name="Museum Donaueschingen" pageId="28" pageNumber="29">MD</collectionCode>
|
||
50 DC114,
|
||
<collectionCode id="2995AE725342FF8CFF4A8703FE7CFDB4" box="[382,468,512,538]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">MNHN</collectionCode>
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<collectionCode id="2995AE725342FF8CFFEA84FCFE51FDB7" box="[478,505,511,537]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="28" pageNumber="29">IE</collectionCode>
|
||
.2009.1600 (1).
|
||
<collectionCode id="2995AE725342FF8CFC8684FCFD77FDB4" box="[690,735,511,538]" country="Germany" name="Museum Donaueschingen" pageId="28" pageNumber="29">MD</collectionCode>
|
||
50 CP115,
|
||
<collectionCode id="2995AE725342FF8CFD568703FC10FDB4" box="[866,952,512,538]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">MNHN</collectionCode>
|
||
<collectionCode id="2995AE725342FF8CFDF684FCFC75FDB7" box="[962,989,511,537]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="28" pageNumber="29">IE</collectionCode>
|
||
.2009.1601 (1).
|
||
<collectionCode id="2995AE725342FF8CFAA384FCFB6CFDB4" box="[1175,1220,511,538]" country="Germany" name="Museum Donaueschingen" pageId="28" pageNumber="29">MD</collectionCode>
|
||
50 CP116,
|
||
<collectionCode id="2995AE725342FF8CFB738703FA35FDB4" box="[1351,1437,512,538]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">MNHN</collectionCode>
|
||
<collectionCode id="2995AE725342FF8CFEA38720FF1AFD93" box="[151,178,547,573]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="28" pageNumber="29">IE</collectionCode>
|
||
.2009.1602 (9).
|
||
<collectionCode id="2995AE725342FF8CFF518720FE3AFD90" box="[357,402,547,574]" country="Germany" name="Museum Donaueschingen" pageId="28" pageNumber="29">MD</collectionCode>
|
||
50 DC127,
|
||
<collectionCode id="2995AE725342FF8CFC208727FDC2FD90" box="[532,618,548,574]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">MNHN</collectionCode>
|
||
<collectionCode id="2995AE725342FF8CFC458720FD24FD93" box="[625,652,547,573]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="28" pageNumber="29">IE</collectionCode>
|
||
.2009.1603 (1).
|
||
<collectionCode id="2995AE725342FF8CFD0B8720FCC4FD90" box="[831,876,547,574]" country="Germany" name="Museum Donaueschingen" pageId="28" pageNumber="29">MD</collectionCode>
|
||
50 DC146,
|
||
<collectionCode id="2995AE725342FF8CFDDA8727FBECFD90" box="[1006,1092,548,574]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">MNHN</collectionCode>
|
||
<collectionCode id="2995AE725342FF8CFA7E8720FBCDFD93" box="[1098,1125,547,573]" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="28" pageNumber="29">IE</collectionCode>
|
||
.2009.1604 (1).
|
||
</paragraph>
|
||
<paragraph id="4F3B36B75342FF8CFEF38744FD30FD63" blockId="28.[151,1437,367,898]" pageId="28" pageNumber="29">
|
||
<emphasis id="7DF0EAA55342FF8CFEF38744FE79FDCF" bold="true" box="[199,465,583,609]" pageId="28" pageNumber="29">Comparative material</emphasis>
|
||
.
|
||
<taxonomicName id="88844D345342FF8CFFD48744FC37FDCF" authority="(Abildgaard, 1789)" baseAuthorityName="Abildgaard" baseAuthorityYear="1789" box="[480,927,583,609]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiocomina" kingdom="Animalia" order="Ophiacanthida" pageId="28" pageNumber="29" phylum="Echinodermata" rank="species" species="nigra">
|
||
<emphasis id="7DF0EAA55342FF8CFFD48744FD15FDCF" box="[480,701,583,609]" italics="true" pageId="28" pageNumber="29">Ophiocomina nigra</emphasis>
|
||
(
|
||
<bibRefCitation id="2B154B465342FF8CFCFF8744FC3FFDCF" author="Abildgaard, P. C." box="[715,919,583,609]" pageId="28" pageNumber="29" pagination="81 - 120" refId="ref28769" refString="Abildgaard, P. C. (1789) s. n. In: Muller, O. F. (Ed.), Zoologia Danica seu animalium Daniae et Norvegiae rariorum ac minus notorum descriptiones et historia. N. Moller, Havniae [Copenhague], pp. 1 - 71, pls. 81 - 120." type="book chapter" year="1789">Abildgaard, 1789</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
:
|
||
<materialsCitation id="FFEC3CEA5342FF8CFD9E8744FEE4FD28" ID-GBIF-Occurrence="3746276316" accessionNumber="F92216, F91624" collectingDate="1969-08-05" collectingDateMax="2004-07-07" collectingDateMin="1969-08-05" collectionCode="MV" country="Saint Helena, Ascension and Tristan da Cunha" county="The Limestone" elevation="20" latitude="-37.336334" location="Roscoff" longLatPrecision="735" longitude="-12.5" municipality="County Down" pageId="28" pageNumber="29" specimenCode="F167538" specimenCount="1">
|
||
<location id="4A5B606C5342FF8CFD9E8744FBADFDCF" LSID="urn:lsid:plazi:treatment:C72D87A15342FF8FFEA385DCFEBCFBF7:4A5B606C5342FF8CFD9E8744FBADFDCF" box="[938,1029,583,609]" country="Saint Helena, Ascension and Tristan da Cunha" county="The Limestone" latitude="-37.336334" longLatPrecision="735" longitude="-12.5" municipality="County Down" name="Roscoff" pageId="28" pageNumber="29">Roscoff</location>
|
||
,
|
||
<location id="4A5B606C5342FF8CFA258744FBC6FDCF" LSID="urn:lsid:plazi:treatment:C72D87A15342FF8FFEA385DCFEBCFBF7:4A5B606C5342FF8CFA258744FBC6FDCF" box="[1041,1134,583,609]" country="Saint Helena, Ascension and Tristan da Cunha" county="The Limestone" latitude="-37.336334" longLatPrecision="735" longitude="-12.5" municipality="County Down" name="Brittany" pageId="28" pageNumber="29">Brittany</location>
|
||
,
|
||
<geoCoordinate id="2AB050705342FF8CFA4E8744FB4DFDCF" box="[1146,1253,583,609]" degrees="48" direction="north" minutes="43" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="48.716667">48° 43´N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2AB050705342FF8CFAC58744FAFAFDCC" box="[1265,1362,583,610]" degrees="3" direction="west" minutes="59" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="-3.9833333">3° 59´W</geoCoordinate>
|
||
, 1997,
|
||
<collectionCode id="2995AE725342FF8CFEA3876FFF6CFD28" box="[151,196,620,646]" country="0" name="University of Montana Museum" pageId="28" pageNumber="29">MV</collectionCode>
|
||
<accessionNumber id="50D7AB545342FF8CFEFE8768FEB7FD2B" box="[202,287,619,645]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/F92216" pageId="28" pageNumber="29" type="EnaNcbi">F92216</accessionNumber>
|
||
(5).
|
||
</materialsCitation>
|
||
<materialsCitation id="FFEC3CEA5342FF8CFF668768FDF4FD04" ID-GBIF-Occurrence="3746276309" accessionNumber="F92216, F91624" collectingDate="1969-08-05" collectingDateMax="2004-07-07" collectingDateMin="1969-08-05" collectionCode="MV" country="Saint Helena, Ascension and Tristan da Cunha" county="The Limestone" elevation="20" latitude="-37.336334" location="Roscoff" longLatPrecision="735" longitude="-12.5" municipality="County Down" pageId="28" pageNumber="29" specimenCode="F167538" specimenCount="1">
|
||
<location id="4A5B606C5342FF8CFF668768FDB4FD2B" LSID="urn:lsid:plazi:treatment:C72D87A15342FF8FFEA385DCFEBCFBF7:4A5B606C5342FF8CFF668768FDB4FD2B" box="[338,540,619,645]" country="Saint Helena, Ascension and Tristan da Cunha" county="The Limestone" latitude="-37.336334" longLatPrecision="735" longitude="-12.5" municipality="County Down" name="Strangford Lough" pageId="28" pageNumber="29">Strangford Lough</location>
|
||
,
|
||
<collectingMunicipality id="AF5FACCD5342FF8CFC128768FD6DFD28" box="[550,709,619,646]" pageId="28" pageNumber="29">County Down</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="A65A4E3B5342FF8CFCFB8768FCD1FD2B" box="[719,889,619,645]" pageId="28" pageNumber="29">The Limestone</collectingCounty>
|
||
,
|
||
<geoCoordinate id="2AB050705342FF8CFDB78768FC45FD2B" box="[899,1005,619,646]" degrees="54" direction="north" minutes="30" orientation="latitude" pageId="28" pageNumber="29" precision="925" value="54.5">54° 30´N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2AB050705342FF8CFDC38768FBFDFD2B" box="[1015,1109,619,646]" degrees="5" direction="west" minutes="40" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="-5.6666665">5° 40´W</geoCoordinate>
|
||
,
|
||
<quantity id="887C9B525342FF8CFA6B8768FB31FD28" box="[1119,1177,619,646]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="29" unit="m" value="20.0">
|
||
<elevation id="C4A9D1845342FF8CFA6B8768FB31FD28" box="[1119,1177,619,646]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="29" unit="m" value="20.0">20 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="3B3A10775342FF8CFA908768FAA2FD28" box="[1188,1290,619,646]" pageId="28" pageNumber="29" value="1969-08-05">
|
||
<collectingDate id="2B7EE99F5342FF8CFA908768FAA2FD28" box="[1188,1290,619,646]" pageId="28" pageNumber="29" value="1969-08-05">5/8/1969</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="2995AE725342FF8CFB20876FFAE9FD28" box="[1300,1345,620,646]" country="0" name="University of Montana Museum" pageId="28" pageNumber="29">MV</collectionCode>
|
||
<accessionNumber id="50D7AB545342FF8CFB738768FA34FD2B" box="[1351,1436,619,645]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/F91624" pageId="28" pageNumber="29" type="EnaNcbi">F91624</accessionNumber>
|
||
(5).
|
||
<taxonomicName id="88844D345342FF8CFEFF878CFEDBFD07" box="[203,371,655,681]" class="Ophiuroidea" family="Clarkcomidae" genus="Clarkcoma" kingdom="Animalia" order="Ophiacanthida" pageId="28" pageNumber="29" phylum="Echinodermata" rank="species" species="undetermined">
|
||
<emphasis id="7DF0EAA55342FF8CFEFF878CFEDBFD07" box="[203,371,655,681]" italics="true" pageId="28" pageNumber="29">Clarkcoma sp.</emphasis>
|
||
</taxonomicName>
|
||
ICEFISH/85-BT45,
|
||
</materialsCitation>
|
||
<materialsCitation id="FFEC3CEA5342FF8CFC53878CFD3BFD63" ID-GBIF-Occurrence="3746276310" accessionNumber="F92216, F91624" collectingDate="1969-08-05" collectingDateMax="2004-07-07" collectingDateMin="1969-08-05" collectionCode="MV" country="Saint Helena, Ascension and Tristan da Cunha" county="The Limestone" elevation="20" latitude="-37.336334" location="Roscoff" longLatPrecision="735" longitude="-12.5" municipality="County Down" pageId="28" pageNumber="29" specimenCode="F167538" specimenCount="1">
|
||
<collectingCountry id="379376275342FF8CFC53878CFC9BFD07" box="[615,819,655,681]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="28" pageNumber="29">Tristan da Cunha</collectingCountry>
|
||
,
|
||
<geoCoordinate id="2AB050705342FF8CFD77878CFC78FD07" box="[835,976,655,681]" degrees="37" direction="south" minutes="20.18" orientation="latitude" pageId="28" pageNumber="29" precision="9" value="-37.336334">37° 20.18´S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2AB050705342FF8CFDD4878CFBFBFD07" box="[992,1107,655,681]" degrees="12" direction="west" minutes="30" orientation="longitude" pageId="28" pageNumber="29" precision="925" value="-12.5">12° 30´W</geoCoordinate>
|
||
,
|
||
<quantity id="887C9B525342FF8CFA57878CFB41FD04" box="[1123,1257,655,682]" metricMagnitude="2" metricUnit="m" metricValue="2.165" metricValueMax="2.23" metricValueMin="2.1" pageId="28" pageNumber="29" unit="m" value="216.5" valueMax="223.0" valueMin="210.0">
|
||
<elevation id="C4A9D1845342FF8CFA57878CFB41FD04" box="[1123,1257,655,682]" metricMagnitude="2" metricUnit="m" metricValue="2.165" metricValueMax="2.23" metricValueMin="2.1" pageId="28" pageNumber="29" unit="m" value="216.5" valueMax="223.0" valueMin="210.0">210–223 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="3B3A10775342FF8CFACD878CFAC8FD07" box="[1273,1376,655,681]" pageId="28" pageNumber="29" value="2004-07-07">
|
||
<collectingDate id="2B7EE99F5342FF8CFACD878CFAC8FD07" box="[1273,1376,655,681]" pageId="28" pageNumber="29" value="2004-07-07">7/7/2004</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="2995AE725342FF8CFB448793FA35FD04" box="[1392,1437,656,682]" country="0" name="University of Montana Museum" pageId="28" pageNumber="29">MV</collectionCode>
|
||
<specimenCode id="1F229ECC5342FF8CFEA387B0FF53FD60" box="[151,251,691,718]" collectionCode="MV" country="0" name="University of Montana Museum" pageId="28" pageNumber="29">F167538</specimenCode>
|
||
(DNA code=TOH_0050 & OTC1a)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B75342FF8CFEF387D4FB9EFC2F" blockId="28.[151,1437,367,898]" pageId="28" pageNumber="29">
|
||
<emphasis id="7DF0EAA55342FF8CFEF387D4FE68FD5F" bold="true" box="[199,448,727,753]" pageId="28" pageNumber="29">
|
||
<typeStatus id="903F88155342FF8CFEF387D4FE9CFD5F" box="[199,308,727,753]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
|
||
description
|
||
</emphasis>
|
||
. Disc is
|
||
<quantity id="887C9B525342FF8CFC1487D4FDDDFD5C" box="[544,629,727,754]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="28" pageNumber="29" unit="mm" value="8.5">8.5 mm</quantity>
|
||
dd, pentagonal, covered in dense granules
|
||
<quantity id="887C9B525342FF8CFA6787D4FB1EFD5C" box="[1107,1206,727,754]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="28" pageNumber="29" unit="mm" value="0.07">0.07 mm</quantity>
|
||
dia, as high as wide, cylindrical with a granular surface, ~132 per mm
|
||
<quantity id="887C9B525342FF8CFC8387F8FD74FCBB" box="[695,732,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="28" pageNumber="29" unit="in" value="2.0">
|
||
<superScript id="B8F19BFF5342FF8CFC8387F8FD17FCA7" attach="left" box="[695,703,763,777]" fontSize="6" pageId="28" pageNumber="29">2</superScript>
|
||
in
|
||
</quantity>
|
||
the centre of the disc (
|
||
<figureCitation id="D7BF2A325342FF8CFDE387F8FB91FCBB" box="[983,1081,763,789]" captionStart="FIGURE 11" captionStartId="28.[151,250,1801,1825]" captionTargetBox="[152,1435,921,1777]" captionTargetId="figure-264@28.[151,1436,921,1777]" captionTargetPageId="28" captionText="FIGURE 11. (A–E) Ophiocomina arnaudi sp. nov., holotype, MNHN IE.2019.4747,A: dorsal outline, B: dorsal view, C: ventral view; D: inset of marginal disc granules, E: inset of central disc granules. (F) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599, lateral view of two jaws. Abbreviations: T=tooth, TPa=tooth papillae, IPa=infradental papillae, LOPa=lateral oral papillae, AdShSp=adoral shield spine." figureDoi="http://doi.org/10.5281/zenodo.6404706" httpUri="https://zenodo.org/record/6404706/files/figure.png" pageId="28" pageNumber="29">Fig. 11E</figureCitation>
|
||
), slightly sparser near the radial margin (
|
||
<figureCitation id="D7BF2A325342FF8CFEC3861CFEC8FC97" box="[247,352,799,825]" captionStart="FIGURE 11" captionStartId="28.[151,250,1801,1825]" captionTargetBox="[152,1435,921,1777]" captionTargetId="figure-264@28.[151,1436,921,1777]" captionTargetPageId="28" captionText="FIGURE 11. (A–E) Ophiocomina arnaudi sp. nov., holotype, MNHN IE.2019.4747,A: dorsal outline, B: dorsal view, C: ventral view; D: inset of marginal disc granules, E: inset of central disc granules. (F) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599, lateral view of two jaws. Abbreviations: T=tooth, TPa=tooth papillae, IPa=infradental papillae, LOPa=lateral oral papillae, AdShSp=adoral shield spine." figureDoi="http://doi.org/10.5281/zenodo.6404706" httpUri="https://zenodo.org/record/6404706/files/figure.png" pageId="28" pageNumber="29">Fig. 11D</figureCitation>
|
||
), obscuring the thin underlying scales and radial shields (
|
||
<figureCitation id="D7BF2A325342FF8CFDC1861CFBF4FC97" box="[1013,1116,799,825]" captionStart="FIGURE 11" captionStartId="28.[151,250,1801,1825]" captionTargetBox="[152,1435,921,1777]" captionTargetId="figure-264@28.[151,1436,921,1777]" captionTargetPageId="28" captionText="FIGURE 11. (A–E) Ophiocomina arnaudi sp. nov., holotype, MNHN IE.2019.4747,A: dorsal outline, B: dorsal view, C: ventral view; D: inset of marginal disc granules, E: inset of central disc granules. (F) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599, lateral view of two jaws. Abbreviations: T=tooth, TPa=tooth papillae, IPa=infradental papillae, LOPa=lateral oral papillae, AdShSp=adoral shield spine." figureDoi="http://doi.org/10.5281/zenodo.6404706" httpUri="https://zenodo.org/record/6404706/files/figure.png" pageId="28" pageNumber="29">Fig. 11B</figureCitation>
|
||
). Ventral disc surfaces (
|
||
<figureCitation id="D7BF2A325342FF8CFB5B861CFF6DFCF3" captionStart="FIGURE 11" captionStartId="28.[151,250,1801,1825]" captionTargetBox="[152,1435,921,1777]" captionTargetId="figure-264@28.[151,1436,921,1777]" captionTargetPageId="28" captionText="FIGURE 11. (A–E) Ophiocomina arnaudi sp. nov., holotype, MNHN IE.2019.4747,A: dorsal outline, B: dorsal view, C: ventral view; D: inset of marginal disc granules, E: inset of central disc granules. (F) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599, lateral view of two jaws. Abbreviations: T=tooth, TPa=tooth papillae, IPa=infradental papillae, LOPa=lateral oral papillae, AdShSp=adoral shield spine." figureDoi="http://doi.org/10.5281/zenodo.6404706" httpUri="https://zenodo.org/record/6404706/files/figure.png" pageId="28" pageNumber="29">Fig. 11C</figureCitation>
|
||
) are covered in small glassy imbricating scales, with a wedge of granules extending from the margin to about 1/3 to 1/2 of the distance to the oral shield. There are no genital granules or spines.
|
||
</paragraph>
|
||
<caption id="1BFB663F5342FF8CFEA3820AFDE0F823" ID-DOI="http://doi.org/10.5281/zenodo.6404706" ID-Zenodo-Dep="6404706" httpUri="https://zenodo.org/record/6404706/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,250,1801,1825]" targetBox="[152,1435,921,1777]" targetPageId="28">
|
||
<paragraph id="4F3B36B75342FF8CFEA3820AFDE0F823" blockId="28.[151,1436,1801,1933]" pageId="28" pageNumber="29">
|
||
<emphasis id="7DF0EAA55342FF8CFEA3820AFEB5F88F" bold="true" box="[151,285,1801,1825]" pageId="28" pageNumber="29">FIGURE 11.</emphasis>
|
||
(A–E)
|
||
<taxonomicName id="88844D345342FF8CFF52820AFDECF88F" authority="O’Hara & Thuy, 2022" authorityName="O’Hara & Thuy" authorityYear="2022" box="[358,580,1801,1825]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiocomina" kingdom="Animalia" order="Ophiacanthida" pageId="28" pageNumber="29" phylum="Echinodermata" rank="species" species="arnaudi" status="sp. nov.">
|
||
<emphasis id="7DF0EAA55342FF8CFF52820AFDECF88F" box="[358,580,1801,1825]" italics="true" pageId="28" pageNumber="29">Ophiocomina arnaudi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7DF0EAA55342FF8CFC7D820AFD30F88F" bold="true" box="[585,664,1801,1825]" pageId="28" pageNumber="29">
|
||
<taxonomicNameLabel id="66C357DE5342FF8CFC7D820AFD30F88F" box="[585,664,1801,1825]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, MNHN IE.2019.4747, A: dorsal outline, B: dorsal view, C: ventral view; D: inset of marginal disc granules, E: inset of central disc granules. (F)
|
||
<taxonomicName id="88844D345342FF8CFD93822EFB2FF8EB" authority="O’Hara & Thuy, 2022" authorityName="O’Hara & Thuy" authorityYear="2022" box="[935,1159,1837,1861]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiocomina" kingdom="Animalia" order="Ophiacanthida" pageId="28" pageNumber="29" phylum="Echinodermata" rank="species" species="arnaudi" status="sp. nov.">
|
||
<emphasis id="7DF0EAA55342FF8CFD93822EFB2FF8EB" box="[935,1159,1837,1861]" italics="true" pageId="28" pageNumber="29">Ophiocomina arnaudi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7DF0EAA55342FF8CFABA822EFB76F8EB" bold="true" box="[1166,1246,1837,1861]" pageId="28" pageNumber="29">
|
||
<taxonomicNameLabel id="66C357DE5342FF8CFABA822EFB76F8EB" box="[1166,1246,1837,1861]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, MNHN IE.2009.1599, lateral view of two jaws. Abbreviations: T=tooth, TPa=tooth papillae, IPa=infradental papillae, LOPa=lateral oral papillae, AdShSp=adoral shield spine.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4F3B36B75342FF8DFEF382B8FCB0FE03" blockId="28.[151,1436,1979,2042]" lastBlockId="29.[151,1437,151,2013]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
|
||
Oral shields are spear-head shaped, 1.2 times as wide as long, with a hemispherical proximal edge and slightly lobed distal margin, slightly depressed near distal margin. Madreporite is as wide as long, with a convex rather than lobed distal margin. Adoral shields are long and narrow, fully contiguous proximally, sometimes extending around the oral shield to abut the first LAP, other adoral shields appear to finish at the lateral angle of the oral shield, perhaps with the distant extension lying underneath the oral shield and allowing the first LAP to be contiguous with the oral shield. Jaws are long and narrow, oral plates slightly protuberant near dental plate, with 6 rounded teeth (
|
||
<figureCitation id="D7BF2A325343FF8DFE948424FEA9FEEF" box="[160,257,295,321]" captionStart="FIGURE 11" captionStartId="28.[151,250,1801,1825]" captionTargetBox="[152,1435,921,1777]" captionTargetId="figure-264@28.[151,1436,921,1777]" captionTargetPageId="28" captionText="FIGURE 11. (A–E) Ophiocomina arnaudi sp. nov., holotype, MNHN IE.2019.4747,A: dorsal outline, B: dorsal view, C: ventral view; D: inset of marginal disc granules, E: inset of central disc granules. (F) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599, lateral view of two jaws. Abbreviations: T=tooth, TPa=tooth papillae, IPa=infradental papillae, LOPa=lateral oral papillae, AdShSp=adoral shield spine." figureDoi="http://doi.org/10.5281/zenodo.6404706" httpUri="https://zenodo.org/record/6404706/files/figure.png" pageId="29" pageNumber="30">Fig. 11F</figureCitation>
|
||
) and 4–5 conical tooth papillae in 1–2 rows, teeth not projecting further than the tooth papillae, lacking hylanated cap; 5–6 oral papillae around the jaw side, including one tall conical, proximately-directed infradental papilla, 1–2 slender lateral oral papillae, 1–2 flattened oval adoral shield spines, and a small extension of the ventral compartment ossicle that is situated near to the first VAP.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B75343FF8DFEF384B4FE0CFCBB" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="30">
|
||
DAPs are approximately
|
||
<quantity id="887C9B525343FF8DFFDC84B5FDE6FE7F" box="[488,590,438,465]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="29" pageNumber="30" unit="mm" value="1.05">1.05 mm</quantity>
|
||
wide and
|
||
<quantity id="887C9B525343FF8DFCF284B4FC83FE7F" box="[710,811,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="29" pageNumber="30" unit="mm" value="0.63">0.63 mm</quantity>
|
||
long at arm base, fan-shaped, with a straight proximal edge, tapered proximo-lateral sides and a convex distal margin, the distal margin is sometimes truncate in the middle where it meets the succeeding plates, contiguous for 1/5 of their width, becoming more kite-shaped distally, with longer proximal taper but still just contiguous, arms slightly noded and DAPs just separate at arm tip. LAPs with a few thorns on the proximal side of the pronounced arm spine ridge. VAPs on the first segments are heart-shaped, a little wider than long, with a rounded distal margin and a notched proximal edge marking the end of the jaw slit; 2nd VAPs are axe-head-shaped, 1.6 times as wide as long, with a straight to slightly concave ventral margin, concave proximolateral sides around the tentacle pore, and a wide convex uplifted distal edge; succeeding plates become larger and more quadrilateral, with a slightly convex proximal margin, and a less pronounced distal margin, plates touch but don’t overlap.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B75343FF8DFEF3861CFAA1FBBF" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="30">
|
||
Arm spines are long (
|
||
<figureCitation id="D7BF2A325343FF8DFF8A861CFD8DFC97" box="[446,549,799,825]" captionStart="FIGURE 11" captionStartId="28.[151,250,1801,1825]" captionTargetBox="[152,1435,921,1777]" captionTargetId="figure-264@28.[151,1436,921,1777]" captionTargetPageId="28" captionText="FIGURE 11. (A–E) Ophiocomina arnaudi sp. nov., holotype, MNHN IE.2019.4747,A: dorsal outline, B: dorsal view, C: ventral view; D: inset of marginal disc granules, E: inset of central disc granules. (F) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599, lateral view of two jaws. Abbreviations: T=tooth, TPa=tooth papillae, IPa=infradental papillae, LOPa=lateral oral papillae, AdShSp=adoral shield spine." figureDoi="http://doi.org/10.5281/zenodo.6404706" httpUri="https://zenodo.org/record/6404706/files/figure.png" pageId="29" pageNumber="30">Fig. 11A</figureCitation>
|
||
), flattened, oval in cross section, inner lumen less than half diameter, stereom very finely granulated, with a blunt or truncate tip with tiny thorns, 3 spines on first segment, 4 on next three segments under the disc, and 5–6 arm spines once free of the disc, with the upper one often being very long (
|
||
<quantity id="887C9B525343FF8DFB098664FA3CFC2F" box="[1341,1428,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="29" pageNumber="30" unit="mm" value="2.8">2.8 mm</quantity>
|
||
) on one side and short and lanceolate on the other (but not alternating in number), other spines in descending order are ~2.5, 1.8, 1.5 and
|
||
<quantity id="887C9B525343FF8DFFB986ACFE4EFC67" box="[397,486,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="29" pageNumber="30" unit="mm" value="1.3">1.3 mm</quantity>
|
||
, ventralmost spines can be a little curved; in the distal half of the arm the number of arm spines falls to 4–5; with 3 needle-like spines at the arm trip. Two oval tentacle scales occur on the LAPs, half the length of the VAP, subequal at first, but the inner one becoming smaller by about the 6th segment.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B75343FF8DFEF38118FAEDFA9F" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="30">
|
||
<emphasis id="7DF0EAA55343FF8DFEF38118FE14FB9A" bold="true" box="[199,444,1050,1077]" pageId="29" pageNumber="30">
|
||
<typeStatus id="903F88155343FF8DFEF38118FE9CFB9B" box="[199,308,1051,1077]" pageId="29" pageNumber="30" type="paratype">Paratype</typeStatus>
|
||
variations.
|
||
</emphasis>
|
||
<typeStatus id="903F88155343FF8DFFF18118FD8FFB9B" box="[453,551,1051,1077]" pageId="29" pageNumber="30" type="paratype">Paratype</typeStatus>
|
||
disc size varies from
|
||
<quantity id="887C9B525343FF8DFD1C8118FC17FB9B" box="[808,959,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="10.7" metricValueMin="4.8" pageId="29" pageNumber="30" unit="mm" value="7.75" valueMax="10.7" valueMin="4.8">4.8–10.7 mm</quantity>
|
||
dd. Largest with up to 8 arm spines. 1–2 small spines can be present on the side of the oral plate within the jaw slit. There are up to 6 flat blade-like teeth (
|
||
<figureCitation id="D7BF2A325343FF8DFB5B813CFF6CFBD3" captionStart="FIGURE 11" captionStartId="28.[151,250,1801,1825]" captionTargetBox="[152,1435,921,1777]" captionTargetId="figure-264@28.[151,1436,921,1777]" captionTargetPageId="28" captionText="FIGURE 11. (A–E) Ophiocomina arnaudi sp. nov., holotype, MNHN IE.2019.4747,A: dorsal outline, B: dorsal view, C: ventral view; D: inset of marginal disc granules, E: inset of central disc granules. (F) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599, lateral view of two jaws. Abbreviations: T=tooth, TPa=tooth papillae, IPa=infradental papillae, LOPa=lateral oral papillae, AdShSp=adoral shield spine." figureDoi="http://doi.org/10.5281/zenodo.6404706" httpUri="https://zenodo.org/record/6404706/files/figure.png" pageId="29" pageNumber="30">Fig. 11F</figureCitation>
|
||
), the most ventral are broadly triangular with a rounded proximal tip, the dorsal teeth and more quadrangular with rounded lateral corners, and up to 10 conical tooth papillae, a little larger near the teeth. Underlying disc scales are rounded, thin, glassy, 0.7–1.0 mm dia, often fragmented. Radial shields are small, with a beaded surface, approximately 4 times the size of the disc scales, separated by 3–4 scales, and covered in granules. Disc granules can occur on the first DAP. Peristomial plates are obscured by dried internal tissue. Colour is generally grey-yellow, with a hint of darker patches, arm tips pinkish, banded with 2–3 segments darker and 3–4 segments lighter.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B75343FF8DFEF38038FC41F913" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="30">
|
||
<emphasis id="7DF0EAA55343FF8DFEF38038FDB6FAFA" bold="true" box="[199,542,1338,1365]" pageId="29" pageNumber="30">
|
||
<typeStatus id="903F88155343FF8DFEF38038FE9CFAFB" box="[199,308,1339,1365]" pageId="29" pageNumber="30" type="paratype">Paratype</typeStatus>
|
||
ossicle description.
|
||
</emphasis>
|
||
Proximal LAPs (
|
||
<figureCitation id="D7BF2A325343FF8DFCD18038FCC4FAFB" box="[741,876,1339,1365]" captionStart="FIGURE 12" captionStartId="30.[151,250,1809,1833]" captionTargetBox="[183,1405,181,1785]" captionTargetId="figure-26@30.[183,1405,181,1785]" captionTargetPageId="30" captionText="FIGURE 12. (A–S) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599.A–B: proximal lateral arm plate in external (A) and internal (B) views. C: distal lateral arm plate in external view. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N–O: teeth. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404710" httpUri="https://zenodo.org/record/6404710/files/figure.png" pageId="29" pageNumber="30">Fig. 12A–B</figureCitation>
|
||
) are two times higher than long, strongly arched, relatively thin, with strongly protruding ventral portion; dorsal edge slightly convex; outer surface with finely meshed stereom and with a fine, regular vertical striation close to the ventral and dorsal edges and close to the raised distal portion; outer proximal edge with a band of more finely meshed stereom devoid of spurs. Distal plates becoming longer than high (
|
||
<figureCitation id="D7BF2A325343FF8DFFD680C8FDE5FA4B" box="[482,589,1483,1509]" captionStart="FIGURE 12" captionStartId="30.[151,250,1809,1833]" captionTargetBox="[183,1405,181,1785]" captionTargetId="figure-26@30.[183,1405,181,1785]" captionTargetPageId="30" captionText="FIGURE 12. (A–S) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599.A–B: proximal lateral arm plate in external (A) and internal (B) views. C: distal lateral arm plate in external view. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N–O: teeth. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404710" httpUri="https://zenodo.org/record/6404710/files/figure.png" pageId="29" pageNumber="30">Fig. 12C</figureCitation>
|
||
). Spine articulations (
|
||
<figureCitation id="D7BF2A325343FF8DFD7880C8FC1EFA4B" box="[844,950,1483,1509]" captionStart="FIGURE 12" captionStartId="30.[151,250,1809,1833]" captionTargetBox="[183,1405,181,1785]" captionTargetId="figure-26@30.[183,1405,181,1785]" captionTargetPageId="30" captionText="FIGURE 12. (A–S) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599.A–B: proximal lateral arm plate in external (A) and internal (B) views. C: distal lateral arm plate in external view. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N–O: teeth. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404710" httpUri="https://zenodo.org/record/6404710/files/figure.png" pageId="29" pageNumber="30">Fig. 12D</figureCitation>
|
||
) are freestanding in a continuous vertical row on a strongly raised distal portion of the LAP, ear-shaped, very large, composed of proximally merged dorsal and ventral lobes enclosing a large muscle opening and much smaller nerve opening and with a well-developed sigmoidal fold; dorsal and ventral lobes expanded dorsally and ventrally, respectively, in the three dorsalmost spine articulations. Inner side of LAPs have a large, prominent, well-defined, slender vertebral articular ridge reminiscent of a reversed sickle; no spurs on the inner distal edge; tentacle notch is relatively large, oblique, shallow. Small perforations occur in a vertical row, with a slender, shallow furrow dorsally.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B75343FF8DFEF383C4FBE6F8DF" blockId="29.[151,1437,151,2013]" pageId="29" pageNumber="30">
|
||
Vertebrae (
|
||
<figureCitation id="D7BF2A325343FF8DFF7083C4FE69F94F" box="[324,449,1735,1761]" captionStart="FIGURE 12" captionStartId="30.[151,250,1809,1833]" captionTargetBox="[183,1405,181,1785]" captionTargetId="figure-26@30.[183,1405,181,1785]" captionTargetPageId="30" captionText="FIGURE 12. (A–S) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599.A–B: proximal lateral arm plate in external (A) and internal (B) views. C: distal lateral arm plate in external view. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N–O: teeth. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404710" httpUri="https://zenodo.org/record/6404710/files/figure.png" pageId="29" pageNumber="30">Fig. 12F–J</figureCitation>
|
||
) are short, with very wide, rounded triangular dorsal fossae and with relatively small ventral muscle fossae on both distal and proximal faces; articulations small, typically zygospondylous, with a relatively large zygosphene on the distal face; very slender L-shaped lateral articulation structure corresponding to articular structure on the inner side of the LAP; dorso-distal muscle fossae are deeply carved into the lateral side of the vertebra; large ventral furrow, with moderately small, well-defined podial basins.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B75343FF8FFEF38278FC53FEEF" blockId="29.[151,1437,151,2013]" lastBlockId="31.[151,1437,151,1113]" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
|
||
Oral plates (
|
||
<figureCitation id="D7BF2A325343FF8DFF6E8278FE61F83B" box="[346,457,1915,1941]" captionStart="FIGURE 12" captionStartId="30.[151,250,1809,1833]" captionTargetBox="[183,1405,181,1785]" captionTargetId="figure-26@30.[183,1405,181,1785]" captionTargetPageId="30" captionText="FIGURE 12. (A–S) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599.A–B: proximal lateral arm plate in external (A) and internal (B) views. C: distal lateral arm plate in external view. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N–O: teeth. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404710" httpUri="https://zenodo.org/record/6404710/files/figure.png" pageId="29" pageNumber="30">Fig. 12M</figureCitation>
|
||
) are slightly wider than long, with adradial muscle fossa lining the lateral edge of articulation area. Dental plates (
|
||
<figureCitation id="D7BF2A325343FF8DFC36829CFDCCF817" box="[514,612,1951,1977]" captionStart="FIGURE 12" captionStartId="30.[151,250,1809,1833]" captionTargetBox="[183,1405,181,1785]" captionTargetId="figure-26@30.[183,1405,181,1785]" captionTargetPageId="30" captionText="FIGURE 12. (A–S) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599.A–B: proximal lateral arm plate in external (A) and internal (B) views. C: distal lateral arm plate in external view. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N–O: teeth. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404710" httpUri="https://zenodo.org/record/6404710/files/figure.png" pageId="29" pageNumber="30">Fig. 12P</figureCitation>
|
||
) are undivided, rounded, with single row of large, shallow tooth sockets surrounded by separate, weakly protruding ridges; ventral tip of dental plate widest, with a cluster of smaller, shallow tooth sockets each surrounded by a continuous ring. Teeth (
|
||
<figureCitation id="D7BF2A325341FF8FFD038594FC68FF1F" box="[823,960,151,177]" captionStart="FIGURE 12" captionStartId="30.[151,250,1809,1833]" captionTargetBox="[183,1405,181,1785]" captionTargetId="figure-26@30.[183,1405,181,1785]" captionTargetPageId="30" captionText="FIGURE 12. (A–S) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599.A–B: proximal lateral arm plate in external (A) and internal (B) views. C: distal lateral arm plate in external view. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N–O: teeth. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404710" httpUri="https://zenodo.org/record/6404710/files/figure.png" pageId="31" pageNumber="32">Fig. 12N, O</figureCitation>
|
||
) are dome-shaped to rounded rectangular, devoid of enamelled tip. Abradial genital plates are shorter than the adradial ones, slender, bar-like without ridges or groves and with a concave adradio-distal tip. Radial shields (
|
||
<figureCitation id="D7BF2A325341FF8FFD0B85DCFC08FF57" box="[831,928,223,249]" captionStart="FIGURE 12" captionStartId="30.[151,250,1809,1833]" captionTargetBox="[183,1405,181,1785]" captionTargetId="figure-26@30.[183,1405,181,1785]" captionTargetPageId="30" captionText="FIGURE 12. (A–S) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599.A–B: proximal lateral arm plate in external (A) and internal (B) views. C: distal lateral arm plate in external view. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N–O: teeth. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404710" httpUri="https://zenodo.org/record/6404710/files/figure.png" pageId="31" pageNumber="32">Fig. 12S</figureCitation>
|
||
) have an oblique isosceles-triangular outline, with a more thickly-meshed stereom in the middle part of the outer surface. A proximal dorsal arm spine (
|
||
<figureCitation id="D7BF2A325341FF8FFB1F8400FA27FEB3" box="[1323,1423,259,285]" captionStart="FIGURE 12" captionStartId="30.[151,250,1809,1833]" captionTargetBox="[183,1405,181,1785]" captionTargetId="figure-26@30.[183,1405,181,1785]" captionTargetPageId="30" captionText="FIGURE 12. (A–S) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599.A–B: proximal lateral arm plate in external (A) and internal (B) views. C: distal lateral arm plate in external view. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N–O: teeth. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404710" httpUri="https://zenodo.org/record/6404710/files/figure.png" pageId="31" pageNumber="32">Fig. 12E</figureCitation>
|
||
), dorsal arm plate (
|
||
<figureCitation id="D7BF2A325341FF8FFF6F8424FE6BFEEF" box="[347,451,295,321]" captionStart="FIGURE 12" captionStartId="30.[151,250,1809,1833]" captionTargetBox="[183,1405,181,1785]" captionTargetId="figure-26@30.[183,1405,181,1785]" captionTargetPageId="30" captionText="FIGURE 12. (A–S) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599.A–B: proximal lateral arm plate in external (A) and internal (B) views. C: distal lateral arm plate in external view. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N–O: teeth. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404710" httpUri="https://zenodo.org/record/6404710/files/figure.png" pageId="31" pageNumber="32">Fig. 12K</figureCitation>
|
||
), and ventral arm plate (
|
||
<figureCitation id="D7BF2A325341FF8FFCE78424FC9EFEEF" box="[723,822,295,321]" captionStart="FIGURE 12" captionStartId="30.[151,250,1809,1833]" captionTargetBox="[183,1405,181,1785]" captionTargetId="figure-26@30.[183,1405,181,1785]" captionTargetPageId="30" captionText="FIGURE 12. (A–S) Ophiocomina arnaudi sp. nov., paratype, MNHN IE.2009.1599.A–B: proximal lateral arm plate in external (A) and internal (B) views. C: distal lateral arm plate in external view. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N–O: teeth. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene." figureDoi="http://doi.org/10.5281/zenodo.6404710" httpUri="https://zenodo.org/record/6404710/files/figure.png" pageId="31" pageNumber="32">Fig. 12L</figureCitation>
|
||
) are also figured.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="1BFB663F5340FF8EFEA38212FB4BF873" ID-DOI="http://doi.org/10.5281/zenodo.6404710" ID-Zenodo-Dep="6404710" httpUri="https://zenodo.org/record/6404710/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,250,1809,1833]" targetBox="[183,1405,181,1785]" targetPageId="30">
|
||
<paragraph id="4F3B36B75340FF8EFEA38212FB4BF873" blockId="30.[151,1437,1809,2013]" pageId="30" pageNumber="31">
|
||
<emphasis id="7DF0EAA55340FF8EFEA38212FEB6F887" bold="true" box="[151,286,1809,1833]" pageId="30" pageNumber="31">FIGURE 12.</emphasis>
|
||
(A–S)
|
||
<taxonomicName id="88844D345340FF8EFF528212FDEDF887" authority="O’Hara & Thuy, 2022" authorityName="O’Hara & Thuy" authorityYear="2022" box="[358,581,1809,1833]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiocomina" kingdom="Animalia" order="Ophiacanthida" pageId="30" pageNumber="31" phylum="Echinodermata" rank="species" species="arnaudi" status="sp. nov.">
|
||
<emphasis id="7DF0EAA55340FF8EFF528212FDEDF887" box="[358,581,1809,1833]" italics="true" pageId="30" pageNumber="31">Ophiocomina arnaudi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7DF0EAA55340FF8EFC7F8212FD32F887" bold="true" box="[587,666,1809,1833]" pageId="30" pageNumber="31">
|
||
<taxonomicNameLabel id="66C357DE5340FF8EFC7F8212FD32F887" box="[587,666,1809,1833]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, MNHN IE.2009.1599.A–B: proximal lateral arm plate in external (A) and internal (B) views. C: distal lateral arm plate in external view. D: detail of spine articulations. E: arm spine. F–J: proximal vertebra in distal (F), proximal (G), lateral (H), dorsal (I) and ventral (J) views. K: dorsal arm plate. L: ventral arm plate. M: oral plate. N–O: teeth. P: dental plate. Q: abradial genital plate. R: adradial genital plate. S: radial shield. Abbreviations: AdMF = adradial muscle fossa; di = distal; DL = dorsal lobe; do = dorsal; MO = muscle opening; NO = nerve opening; P = perforation; R = vertebral articular ridge; TN = tentacle notch; VL = ventral lobe; ZC = zygocondyles; ZS = zygosphene.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="079E653C5341FF8FFEF38449FD9EFECB" box="[199,566,330,357]" pageId="31" pageNumber="32" type="distribution">
|
||
<paragraph id="4F3B36B75341FF8FFEF38449FD9EFECB" blockId="31.[151,1437,151,1113]" box="[199,566,330,357]" pageId="31" pageNumber="32">
|
||
<emphasis id="7DF0EAA55341FF8FFEF38449FEF1FECA" bold="true" box="[199,345,330,356]" pageId="31" pageNumber="32">Distribution</emphasis>
|
||
. SPA (
|
||
<quantity id="887C9B525341FF8FFF928449FD82FECB" box="[422,554,330,357]" metricMagnitude="2" metricUnit="m" metricValue="3.3" metricValueMax="5.25" metricValueMin="1.35" pageId="31" pageNumber="32" unit="m" value="330.0" valueMax="525.0" valueMin="135.0">135–525 m</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="079E653C5341FF8FFEF3846CFBC6FBBF" pageId="31" pageNumber="32" type="discussion">
|
||
<paragraph id="4F3B36B75341FF8FFEF3846CFB62FDCF" blockId="31.[151,1437,151,1113]" pageId="31" pageNumber="32">
|
||
<emphasis id="7DF0EAA55341FF8FFEF3846CFE9CFE27" bold="true" box="[199,308,367,393]" pageId="31" pageNumber="32">Remarks</emphasis>
|
||
. This species is referred to the genus
|
||
<taxonomicName id="88844D345341FF8FFCD1846CFCD5FE27" box="[741,893,367,393]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiocomina" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7DF0EAA55341FF8FFCD1846CFCD5FE27" box="[741,893,367,393]" italics="true" pageId="31" pageNumber="32">Ophiocomina</emphasis>
|
||
</taxonomicName>
|
||
on the basis of the granulated disc, presence of tooth papillae on the ventral dental plate, a series of oral papillae, adoral shields that meet interradially, long hollow arm spines, and two oval glassy tentacle scales. The only other
|
||
<taxonomicName id="88844D345341FF8FFD5B84B4FBCCFE7F" box="[879,1124,439,465]" pageId="31" pageNumber="32">
|
||
<emphasis id="7DF0EAA55341FF8FFD5B84B4FBAFFE7F" box="[879,1031,439,465]" italics="true" pageId="31" pageNumber="32">Ophiocomina</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88844D345341FF8FFA4484B4FB79FE7E" baseAuthorityName="Abildgaard" baseAuthorityYear="1789" box="[1136,1233,439,465]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiocomina" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="nigra">
|
||
<emphasis id="7DF0EAA55341FF8FFA4484B4FB79FE7E" box="[1136,1233,439,465]" italics="true" pageId="31" pageNumber="32">O. nigra</emphasis>
|
||
</taxonomicName>
|
||
, differs in having smaller more separated disc granules that often persist to the oral shields, more fragmented disc scales, and wider more oval DAPs with convex laterodistal and lateroproximal margins. The genus
|
||
<taxonomicName id="88844D345341FF8FFA1584FCFB11FDB7" box="[1057,1209,511,537]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiocomina" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7DF0EAA55341FF8FFA1584FCFB11FDB7" box="[1057,1209,511,537]" italics="true" pageId="31" pageNumber="32">Ophiocomina</emphasis>
|
||
</taxonomicName>
|
||
is now placed in the family
|
||
<taxonomicName id="88844D345341FF8FFED38720FE2FFD93" authorityName="Paterson" authorityYear="1985" box="[231,391,547,573]" class="Ophiuroidea" family="Ophiotomidae" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="family">Ophiotomidae</taxonomicName>
|
||
on the basis of DNA evidence (
|
||
<bibRefCitation id="2B154B465341FF8FFCD88720FC6CFD93" author="O'Hara, T. D. & Hugall, A. F. & Thuy, B. & Stohr, S. & Martynov, A. V." box="[748,964,547,573]" pageId="31" pageNumber="32" pagination="415 - 430" refId="ref34885" refString="O'Hara, T. D., Hugall, A. F., Thuy, B., Stohr, S. & Martynov, A. V. (2017) Restructuring higher taxonomy using broad-scale phylogenomics: the living Ophiuroidea. Molecular Phylogenetics and Evolution, 107, 415 - 430. https: // doi. org / 10.1016 / j. ympev. 2016.12.006" type="journal article" year="2017">
|
||
O’Hara
|
||
<emphasis id="7DF0EAA55341FF8FFD7D8720FC29FD93" box="[841,897,547,573]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
). The species
|
||
<taxonomicName id="88844D345341FF8FFA558720FEB3FDCF" authority="H. L. Clark, 1928" authorityName="H. L. Clark" authorityYear="1928" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiocomina" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="australis">
|
||
<emphasis id="7DF0EAA55341FF8FFA558720FACAFD93" box="[1121,1378,547,573]" italics="true" pageId="31" pageNumber="32">Ophiocomina australis</emphasis>
|
||
H.L.
|
||
<bibRefCitation id="2B154B465341FF8FFEA38744FEB3FDCF" author="Clark, H. L." box="[151,283,583,609]" pageId="31" pageNumber="32" pagination="361 - 482" refId="ref30116" refString="Clark, H. L. (1928) The sea-lilies, sea-stars, brittle-stars and sea-urchins of the South Australian Museum. Records of the South Australian Museum, 3, 361 - 482, figs. 108 - 142." type="journal article" year="1928">Clark, 1928</bibRefCitation>
|
||
</taxonomicName>
|
||
is now considered a
|
||
<taxonomicName id="88844D345341FF8FFC338744FD73FDCF" box="[519,731,583,609]" pageId="31" pageNumber="32">
|
||
<emphasis id="7DF0EAA55341FF8FFC338744FD2BFDCF" box="[519,643,583,609]" italics="true" pageId="31" pageNumber="32">Clarkcoma</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="2B154B465341FF8FFCDE8744FC74FDCF" author="Naughton, K. M. & O'Hara, T. D. & Appleton, B. & Cisternas, P. A." box="[746,988,583,609]" pageId="31" pageNumber="32" pagination="232 - 244" refId="ref34602" refString="Naughton, K. M., O'Hara, T. D., Appleton, B. & Cisternas, P. A. (2014) Antitropical distributions and species delimitation in a group of ophiocomid brittle stars (Echinodermata: Ophiuroidea: Ophiocomidae). Molecular Phylogenetics and Evolution, 78, 232 - 244. https: // doi. org / 10.1016 / j. ympev. 2014.05.020" type="journal article" year="2014">
|
||
Naughton
|
||
<emphasis id="7DF0EAA55341FF8FFD558744FC31FDCF" box="[865,921,583,609]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="2B154B465341FF8FFDD38744FB16FDCF" author="O'Hara, T. D. & Hugall, A. F. & Thuy, B. & Stohr, S. & Martynov, A. V." box="[999,1214,583,609]" pageId="31" pageNumber="32" pagination="415 - 430" refId="ref34885" refString="O'Hara, T. D., Hugall, A. F., Thuy, B., Stohr, S. & Martynov, A. V. (2017) Restructuring higher taxonomy using broad-scale phylogenomics: the living Ophiuroidea. Molecular Phylogenetics and Evolution, 107, 415 - 430. https: // doi. org / 10.1016 / j. ympev. 2016.12.006" type="journal article" year="2017">
|
||
O’Hara
|
||
<emphasis id="7DF0EAA55341FF8FFA768744FBD2FDCF" box="[1090,1146,583,609]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4F3B36B75341FF8FFEF38768FB31FC97" blockId="31.[151,1437,151,1113]" pageId="31" pageNumber="32">
|
||
<taxonomicName id="88844D345341FF8FFEF38768FEF7FD2B" box="[199,351,619,645]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiocomina" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7DF0EAA55341FF8FFEF38768FEF7FD2B" box="[199,351,619,645]" italics="true" pageId="31" pageNumber="32">Ophiocomina</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="88844D345341FF8FFFCB8768FD7FFD2B" box="[511,727,619,645]" pageId="31" pageNumber="32">
|
||
<emphasis id="7DF0EAA55341FF8FFFCB8768FDD3FD2B" box="[511,635,619,645]" italics="true" pageId="31" pageNumber="32">Clarkcoma</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(except
|
||
<taxonomicName id="88844D345341FF8FFD0B8768FC6EFD2B" authorityName="H. L. Clark" authorityYear="1928" box="[831,966,619,645]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiocomina" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="australis">
|
||
<emphasis id="7DF0EAA55341FF8FFD0B8768FC6EFD2B" box="[831,966,619,645]" italics="true" pageId="31" pageNumber="32">C. australis</emphasis>
|
||
</taxonomicName>
|
||
) in lacking the thick dense stereom cap on the teeth that makes them protrude further than the tooth papillae. This cap is present even on
|
||
<quantity id="887C9B525341FF8FFAD9878CFA87FD07" box="[1261,1327,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="31" pageNumber="32" unit="mm" value="8.0">8 mm</quantity>
|
||
juveniles of
|
||
<taxonomicName id="88844D345341FF8FFE8387B0FDBCFD60" authority="(Lutken 1869)" baseAuthorityName="Lutken" baseAuthorityYear="1869" box="[183,532,691,718]" class="Ophiuroidea" family="Clarkcomidae" genus="Clarkcoma" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="canaliculata">
|
||
<emphasis id="7DF0EAA55341FF8FFE8387B0FECEFD63" box="[183,358,691,717]" italics="true" pageId="31" pageNumber="32">C. canaliculata</emphasis>
|
||
(
|
||
<bibRefCitation id="2B154B465341FF8FFF4287B0FDA4FD60" author="Lutken, C. F." box="[374,524,691,718]" pageId="31" pageNumber="32" pagination="22 - 109" refId="ref32990" refString="Lutken, C. F. (1869) Additamenta ad historiam Ophiuridarum. 3. Beskrivende og kritiske Bidrag til Kundskab an Slangestjernerne. Kongelige Danske Videnskabernes Selskabs Skrifter, 5, 22 - 109. https: // doi. org / 10.5962 / bhl. title. 1408" type="journal article" year="1869">Lütken 1869</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88844D345341FF8FFC1587B0FC9FFD63" authority="(Clark 1928)" baseAuthorityName="Clark" baseAuthorityYear="1928" box="[545,823,691,717]" class="Ophiuroidea" family="Clarkcomidae" genus="Clarkcoma" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="pulchra">
|
||
<emphasis id="7DF0EAA55341FF8FFC1587B0FD33FD63" box="[545,667,691,717]" italics="true" pageId="31" pageNumber="32">C. pulchra</emphasis>
|
||
(
|
||
<bibRefCitation id="2B154B465341FF8FFC9F87B0FC87FD63" author="Clark, H. L." box="[683,815,691,717]" pageId="31" pageNumber="32" pagination="361 - 482" refId="ref30116" refString="Clark, H. L. (1928) The sea-lilies, sea-stars, brittle-stars and sea-urchins of the South Australian Museum. Records of the South Australian Museum, 3, 361 - 482, figs. 108 - 142." type="journal article" year="1928">Clark 1928</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88844D345341FF8FFD7187B0FB2BFD63" authority="(Farquhar 1908)" baseAuthorityName="Farquhar" baseAuthorityYear="1908" box="[837,1155,691,717]" class="Ophiuroidea" family="Clarkcomidae" genus="Clarkcoma" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="bollonsi">
|
||
<emphasis id="7DF0EAA55341FF8FFD7187B0FC69FD63" box="[837,961,691,717]" italics="true" pageId="31" pageNumber="32">C. bollonsi</emphasis>
|
||
(
|
||
<bibRefCitation id="2B154B465341FF8FFDE687B0FBD3FD63" author="Farquhar, H." box="[978,1147,691,717]" pageId="31" pageNumber="32" pagination="108" refId="ref30951" refString="Farquhar, H. (1908) Description of a new ophiuroid. Transactions and Proceedings of the New Zealand Institute, 40, 108." type="journal article" year="1908">Farquhar 1908</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and several undescribed Australian species in the genus (
|
||
<bibRefCitation id="2B154B465341FF8FFC3587D4FD5CFD5F" author="Naughton, K. M. & O'Hara, T. D. & Appleton, B. & Cisternas, P. A." box="[513,756,727,753]" pageId="31" pageNumber="32" pagination="232 - 244" refId="ref34602" refString="Naughton, K. M., O'Hara, T. D., Appleton, B. & Cisternas, P. A. (2014) Antitropical distributions and species delimitation in a group of ophiocomid brittle stars (Echinodermata: Ophiuroidea: Ophiocomidae). Molecular Phylogenetics and Evolution, 78, 232 - 244. https: // doi. org / 10.1016 / j. ympev. 2014.05.020" type="journal article" year="2014">
|
||
Naughton
|
||
<emphasis id="7DF0EAA55341FF8FFC4C87D4FD18FD5F" box="[632,688,727,753]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
). These species have numerous tooth papillae (up to
|
||
<quantity id="887C9B525341FF8FFB7787D4FAD4FD5F" box="[1347,1404,727,753]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="31" pageNumber="32" unit="in" value="30.0">30 in</quantity>
|
||
<taxonomicName id="88844D345341FF8FFBB787D4FF59FCBB" baseAuthorityName="Farquhar" baseAuthorityYear="1908" class="Ophiuroidea" family="Clarkcomidae" genus="Clarkcoma" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="bollonsi">
|
||
<emphasis id="7DF0EAA55341FF8FFBB787D4FF59FCBB" italics="true" pageId="31" pageNumber="32">C. bollonsi</emphasis>
|
||
</taxonomicName>
|
||
) and swollen fleshy tips (sacculae) that occur on many arm spines. The diminutive
|
||
<taxonomicName id="88844D345341FF8FFABB87F8FAB9FCBB" authorityName="H. L. Clark" authorityYear="1928" box="[1167,1297,763,789]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiocomina" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="australis">
|
||
<emphasis id="7DF0EAA55341FF8FFABB87F8FAB9FCBB" box="[1167,1297,763,789]" italics="true" pageId="31" pageNumber="32">C. australis</emphasis>
|
||
</taxonomicName>
|
||
differs from all other
|
||
<taxonomicName id="88844D345341FF8FFECF861CFE3BFC97" box="[251,403,799,825]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiocomina" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="7DF0EAA55341FF8FFECF861CFE3BFC97" box="[251,403,799,825]" italics="true" pageId="31" pageNumber="32">Ophiocomina</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="88844D345341FF8FFFFE861CFD36FC97" box="[458,670,799,825]" pageId="31" pageNumber="32">
|
||
<emphasis id="7DF0EAA55341FF8FFFFE861CFDEEFC97" box="[458,582,799,825]" italics="true" pageId="31" pageNumber="32">Clarkcoma</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in having only one tentacle scale as an adult.
|
||
</paragraph>
|
||
<paragraph id="4F3B36B75341FF8FFEF38640FBC6FBBF" blockId="31.[151,1437,151,1113]" pageId="31" pageNumber="32">
|
||
<bibRefCitation id="2B154B465341FF8FFEF38640FE65FCF3" author="Naughton, K. M. & O'Hara, T. D. & Appleton, B. & Cisternas, P. A." box="[199,461,835,861]" pageId="31" pageNumber="32" pagination="232 - 244" refId="ref34602" refString="Naughton, K. M., O'Hara, T. D., Appleton, B. & Cisternas, P. A. (2014) Antitropical distributions and species delimitation in a group of ophiocomid brittle stars (Echinodermata: Ophiuroidea: Ophiocomidae). Molecular Phylogenetics and Evolution, 78, 232 - 244. https: // doi. org / 10.1016 / j. ympev. 2014.05.020" type="journal article" year="2014">
|
||
Naughton
|
||
<emphasis id="7DF0EAA55341FF8FFF0B8640FED1FCF3" box="[319,377,835,861]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
|
||
(2014)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="2B154B465341FF8FFC338640FD5AFCF3" author="O'Hara, T. D. & Hugall, A. F. & Thuy, B. & Stohr, S. & Martynov, A. V." box="[519,754,835,861]" pageId="31" pageNumber="32" pagination="415 - 430" refId="ref34885" refString="O'Hara, T. D., Hugall, A. F., Thuy, B., Stohr, S. & Martynov, A. V. (2017) Restructuring higher taxonomy using broad-scale phylogenomics: the living Ophiuroidea. Molecular Phylogenetics and Evolution, 107, 415 - 430. https: // doi. org / 10.1016 / j. ympev. 2016.12.006" type="journal article" year="2017">
|
||
O’Hara
|
||
<emphasis id="7DF0EAA55341FF8FFC508640FD37FCF3" box="[612,671,835,861]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
|
||
(2017)
|
||
</bibRefCitation>
|
||
also sequenced specimens of an undescribed species in the
|
||
<taxonomicName id="88844D345341FF8FFEA38664FE9CFC2F" authorityName="O'Hara, Stohr, Hugall, Thuy & Martynov" authorityYear="2018" box="[151,308,871,897]" class="Ophiuroidea" family="Clarkcomidae" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="family">Clarkcomidae</taxonomicName>
|
||
from
|
||
<collectingCountry id="379376275341FF8FFF428664FD9BFC2F" box="[374,563,871,897]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="31" pageNumber="32">Tristan da Cunha</collectingCountry>
|
||
in the South Atlantic collected by the ICEFISH expedition. This is possibly the same species as specimens from Gough Island previously reported (
|
||
<bibRefCitation id="2B154B465341FF8FFDA58688FB98FC0B" author="Koehler, R." box="[913,1072,907,933]" pageId="31" pageNumber="32" pagination="193 - 313" refId="ref32294" refString="Koehler, R. (1908) Asteries, ophiures et echinides de l'Expedition Antarctique Nationale Ecossaise Report on the Scientific Results of the Voyage of the S. Y. Scotia Zoology, 13, 193 - 313, 16 pls." type="journal article" year="1908">Koehler 1908</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="2B154B465341FF8FFA088688FB5CFC08" author="Mortensen, T." box="[1084,1268,907,934]" pageId="31" pageNumber="32" pagination="199 - 348" refId="ref34461" refString="Mortensen, T. (1936) Echinoidea and Ophiuroidea. Discovery Reports, 12, 199 - 348, 9 pls. https: // doi. org / 10.5962 / bhl. part. 8051" type="journal article" year="1936">Mortensen 1936</bibRefCitation>
|
||
) as
|
||
<taxonomicName id="88844D345341FF8FFB168688FE48FC67" authority="(Lyman, 1879)" baseAuthorityName="Lyman" baseAuthorityYear="1879" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiotreta" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="valenciennesi">
|
||
<emphasis id="7DF0EAA55341FF8FFB168688FE87FC67" italics="true" pageId="31" pageNumber="32">Ophiotreta valenciennesi</emphasis>
|
||
(
|
||
<bibRefCitation id="2B154B465341FF8FFF0A86ACFE70FC67" author="Lyman, T." box="[318,472,943,969]" pageId="31" pageNumber="32" pagination="17 - 83" refId="ref33210" refString="Lyman, T. (1879) Ophiuridae and Astrophytidae of the exploring voyage of H. M. S. Challenger under Prof. Sir Wyville Thomson, F. R. S. Part 2. Bulletin of the Museum of Comparative Zoology, Harvard University, 6, 17 - 83, pls. 11 - 19." type="journal article" year="1879">Lyman, 1879</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The ICEFISH specimens, up to
|
||
<quantity id="887C9B525341FF8FFD6486ACFC37FC67" box="[848,927,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="31" pageNumber="32" unit="mm" value="18.0">18 mm</quantity>
|
||
dd, differ from
|
||
<taxonomicName id="88844D345341FF8FFA6586ACFB64FC67" authority="O’Hara & Thuy, 2022" authorityName="O’Hara & Thuy" authorityYear="2022" box="[1105,1228,943,969]" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiocomina" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="arnaudi" status="sp. nov.">
|
||
<emphasis id="7DF0EAA55341FF8FFA6586ACFB64FC67" box="[1105,1228,943,969]" italics="true" pageId="31" pageNumber="32">O. arnaudi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="7DF0EAA55341FF8FFAE786ACFA84FC67" bold="true" box="[1235,1324,943,969]" pageId="31" pageNumber="32">
|
||
<taxonomicNameLabel id="66C357DE5341FF8FFAE786ACFA84FC67" box="[1235,1324,943,969]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having dark markings on the circular disc, small spines on the basal DAPs, and shorter, more curved arm spines.
|
||
<taxonomicName id="88844D345341FF8FFB1686D0FE87FBBF" baseAuthorityName="Lyman" baseAuthorityYear="1879" class="Ophiuroidea" family="Ophiotomidae" genus="Ophiotreta" kingdom="Animalia" order="Ophiacanthida" pageId="31" pageNumber="32" phylum="Echinodermata" rank="species" species="valenciennesi">
|
||
<emphasis id="7DF0EAA55341FF8FFB1686D0FE87FBBF" italics="true" pageId="31" pageNumber="32">Ophiotreta valenciennesi</emphasis>
|
||
</taxonomicName>
|
||
differs in having hemispherical disc granules and only 1–2 tooth papillae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="079E653C5341FF8FFEF38118FEBCFBF7" pageId="31" pageNumber="32" type="etymology">
|
||
<paragraph id="4F3B36B75341FF8FFEF38118FEBCFBF7" blockId="31.[151,1437,151,1113]" pageId="31" pageNumber="32">
|
||
<emphasis id="7DF0EAA55341FF8FFEF38118FEE0FB9B" bold="true" box="[199,328,1051,1077]" pageId="31" pageNumber="32">Etymology</emphasis>
|
||
. Named after Patrick M. Arnaud who led the 1986 MD50 expedition to the Île Amsterdam and Saint-Paul.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |