treatments-xml/data/79/BF/C9/79BFC9D3526F5AEAB9A17253EDCDC6F3.xml

<document ID-CLB-Dataset="276890" ID-DOI="http://dx.doi.org/10.3897/phytokeys.235.110525" ID-GBIF-Dataset="825a76cd-8667-407c-9d17-f8eca655e813" ID-PMC="PMC10656614" ID-Pensoft-Pub="1314-2003-235-31" ID-Pensoft-UUID="A22591C65BDA5F3EAC53EE06CC2BBD8A" ID-PubMed="38020473" ModsDocID="1314-2003-235-31" checkinTime="1699693859738" checkinUser="pensoft" docAuthor="Clark, John L., Fierro-Minda, Alisson, Exe, Nolan, Johnson, Mia, Tribble, Carrie M. &amp; Jost, Lou" docDate="2023" docId="79BFC9D3526F5AEAB9A17253EDCDC6F3" docLanguage="en" docName="PhytoKeys 235: 31-42" docOrigin="PhytoKeys 235" docPubDate="2023-11-10" docSource="http://dx.doi.org/10.3897/phytokeys.235.110525" docTitle="Bomarea pastazensis J. L. Clark, Fierro-Minda &amp; N. Exe 2023, sp. nov." docType="treatment" docVersion="6" id="A22591C65BDA5F3EAC53EE06CC2BBD8A" lastPageNumber="31" masterDocId="A22591C65BDA5F3EAC53EE06CC2BBD8A" masterDocTitle="Bomarea pastazensis (Alstroemeriaceae), an exceptionally small new species from the eastern Andean slopes of Ecuador" masterLastPageNumber="42" masterPageNumber="31" pageNumber="31" updateTime="1720552701392" updateUser="ExternalLinkService">
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<mods:title id="9D1929941EFF2D575FC5DCC8CEB40DD3">Bomarea pastazensis (Alstroemeriaceae), an exceptionally small new species from the eastern Andean slopes of Ecuador</mods:title>
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<mods:namePart id="5F9628B4175A94D153EA49E75539BAA1">Clark, John L.</mods:namePart>
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<mods:affiliation id="8A93440480AA26FE1E2EFC7A86AF8DE6">Marie Selby Botanical Gardens, 1534 Mound Street, Sarasota, FL 34236, USA</mods:affiliation>
<mods:nameIdentifier id="48981B98F2395C5294DD925410D933C5" type="email">jclark@selby.org</mods:nameIdentifier>
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<mods:namePart id="BB618453411E473CA574C12D84A6C96F">Fierro-Minda, Alisson</mods:namePart>
<mods:nameIdentifier id="538EF844143C834F810B8D3A6B043B3E" type="ORCID">https://orcid.org/0000-0001-7169-2989</mods:nameIdentifier>
<mods:affiliation id="93FD2E9D00984EC97FFABCF15EB9F1B9">Herbario QUSF, Colegio de Ciencias Biologicas y Ambientales, Universidad San Francisco de Quito, USFQ, Quito, Ecuador &amp; Herbario QCA, Escuela de Ciencias Biologicas, Pontificia Universidad Catolica de Ecuador, Quito, Ecuador &amp; Reserva: The Youth Land Trust, Washington DC, USA</mods:affiliation>
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<mods:namePart id="B7CF1B19C54B98C02D7A8839224AF452">Exe, Nolan</mods:namePart>
<mods:nameIdentifier id="1C1638C83FD92E0C7F7E6A7CDB918419" type="ORCID">https://orcid.org/0000-0001-8847-7978</mods:nameIdentifier>
<mods:affiliation id="0A5D12FC68E9355528E1625C36DF4807">Reservas de Fundacion EcoMinga, Fundacion EcoMinga, Banos, Tungurahua, Ecuador</mods:affiliation>
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<mods:namePart id="B88E5FE4D06AD25F50C64456912B97B3">Johnson, Mia</mods:namePart>
<mods:nameIdentifier id="5B1F9BC481FD444C1BAF54885803F1C8" type="ORCID">https://orcid.org/0000-0002-4822-0606</mods:nameIdentifier>
<mods:affiliation id="A80CFAB2B853DDAD051CA2E822D7EA56">Reservas de Fundacion EcoMinga, Fundacion EcoMinga, Banos, Tungurahua, Ecuador</mods:affiliation>
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<mods:namePart id="3A57B12ABE94E5F89C75076213F2EDB4">Tribble, Carrie M.</mods:namePart>
<mods:nameIdentifier id="97195C928C52AA78DF1F6397094EB263" type="ORCID">https://orcid.org/0000-0001-7263-7885</mods:nameIdentifier>
<mods:affiliation id="DD1C434E473E595021608FF8DBF81346">School of Life Sciences, University of Hawai'i at Manoa, Honolulu, HI, USA</mods:affiliation>
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<mods:namePart id="C60D4A3F992A86A040987EE1D304AB7E">Jost, Lou</mods:namePart>
<mods:nameIdentifier id="3ABB633B53DC8F103E813F59C1BB977C" type="ORCID">https://orcid.org/0000-0002-5850-0716</mods:nameIdentifier>
<mods:affiliation id="099F0225E82C645D66BACC876701D2A0">Reservas de Fundacion EcoMinga, Fundacion EcoMinga, Banos, Tungurahua, Ecuador</mods:affiliation>
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<mods:date id="63A44A25F96B58B1B0B05D9B6BC2E802">2023</mods:date>
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<mods:number id="28BEF61CBDA1E5845CCA948D98A5D3AA">2023-11-10</mods:number>
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<mods:number id="15C93AC062CCE7CCF51CBE4FB65DEDD6">235</mods:number>
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<mods:identifier id="D4B0FF4EC32C46F03765C8BDEDB2A30B" type="DOI">http://dx.doi.org/10.3897/phytokeys.235.110525</mods:identifier>
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<treatment id="79BFC9D3526F5AEAB9A17253EDCDC6F3" ID-DOI="http://doi.org/10.5281/zenodo.10170526" ID-GBIF-Taxon="217559535" ID-Zenodo-Dep="10170526" LSID="urn:lsid:plazi:treatment:79BFC9D3526F5AEAB9A17253EDCDC6F3" httpUri="http://treatment.plazi.org/id/79BFC9D3526F5AEAB9A17253EDCDC6F3" lastPageNumber="31" pageId="0" pageNumber="31">
<subSubSection id="A2EA97CC4C655DA83EA6D18A8E75291A" pageId="0" pageNumber="31" type="nomenclature">
<paragraph id="98FAFDFC2BFAE8E27A1E56647088620A" pageId="0" pageNumber="31">
<taxonomicName id="6D48744A9036953A6B3CCFE426585F7B" ID-CoL="CCXC6" LSID="79BFC9D3-526F-5AEA-B9A1-7253EDCDC6F3" authority="J. L. Clark, Fierro-Minda &amp; N. Exe" authorityName="J. L. Clark, Fierro-Minda &amp; N. Exe" authorityYear="2023" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Bomarea pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis" status="sp. nov.">Bomarea pastazensis J.L.Clark, Fierro-Minda &amp; N.Exe</taxonomicName>
<taxonomicNameLabel id="04A0DBB5174E3178B13B4D745AB56E71" pageId="0" pageNumber="31">sp. nov.</taxonomicNameLabel>
</paragraph>
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<paragraph id="2BC7285A4AC663F826409911B9DC14A2" pageId="0" pageNumber="31">
<figureCitation id="A09CDE758D233B621D87339242CE725B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bomarea pastazensis A lateral view of flower B front view of flower featuring mature androecium C mature gynoecium D petal E adaxial leaf surface F mature flowers with foliage G habitat H tuberous roots. Photos A by J. L. Clark (J. L. Clark 14839), B-H by N. Exe (N. Exe et al. 2)." figureDoi="10.3897/phytokeys.235.110525.figure2" httpUri="https://binary.pensoft.net/fig/934580" pageId="0" pageNumber="31">Figs 2</figureCitation>
<figureCitation id="44ED0905B9DDC14D74A1C6B0F3172282" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Holotype of Bomarea pastazensis, J. L. Clark 14839 (QCA)." figureDoi="10.3897/phytokeys.235.110525.figure4" httpUri="https://binary.pensoft.net/fig/934582" pageId="0" pageNumber="31">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="611DE4E149AFB75E4A2649CA137DFA24" pageId="0" pageNumber="31" type="diagnosis">
<paragraph id="EB4739180517601FD1731B07B0F4647D" pageId="0" pageNumber="31">Diagnosis.</paragraph>
<paragraph id="CC4B71BEF2170094BE79AF28A78726B9" pageId="0" pageNumber="31">
Similar to 
<taxonomicName id="C95E275A99CB85F1C850DADDFDF3E2EF" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="02445F265DC7FEDFE74AE22E1973A27D" italics="true" pageId="0" pageNumber="31">Bomarea pumila</emphasis>
</taxonomicName>
Griseb. ex Baker, differing in plant height reaching 10-14 cm (vs. 5-8 cm high in 
<taxonomicName id="66B0A7FA4AAD53441D05B3B3B722C137" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="B1121BD0A13ABC47C8D0FB2820C77418" italics="true" pageId="0" pageNumber="31">B. pumila</emphasis>
</taxonomicName>
), pubescent stem (vs. glabrous stem in 
<taxonomicName id="57C57AC8B0CA6DAB3066F33BF178FFB4" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="7BE15A4F3EDD6F5FE5494223E452CC99" italics="true" pageId="0" pageNumber="31">B. pumila</emphasis>
</taxonomicName>
), the presence of ciliate leaf margin (vs. glabrous leaf margin in 
<taxonomicName id="FD6E8A31EE52633EFFC2EC9D6BAD3911" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="6FB3BB6A02C5B30BFA6BF135252FD378" italics="true" pageId="0" pageNumber="31">B. pumila</emphasis>
</taxonomicName>
), and the presence of dark spots on the petals (vs. no spotting on the petals in 
<taxonomicName id="2C54DF339C769254F0F5808DA4AE0E8E" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="587BE158FC7E9178A1A6551292FA0EC3" italics="true" pageId="0" pageNumber="31">B. pumila</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="6298CCB27E1517614F40C71E717DA880" pageId="0" pageNumber="31" type="materials_examined">
<paragraph id="C3B2EA93E1B5230DDABA879FCC89F7A2" pageId="0" pageNumber="31">
<typeStatus id="E43D5768B9387385DDEA906F005A82E7">Type</typeStatus>
.
</paragraph>
<paragraph id="F560A857697BD6D671F169D613B77C57" pageId="0" pageNumber="31">
<materialsCitation id="09F5C5BC26CBE017EF5D926C5F366408" collectingDate="2016-03-10" collectorName="J. L. Clark" country="Ecuador" county="Canton Banos" elevation="3642" latitude="-1.4775918" location="Rio Verde" longLatPrecision="1" longitude="-78.298225" municipality="Parroquia Rio Verde" specimenCount="1" stateProvince="Tungurahua" typeStatus="holotype">
<emphasis id="19E617D0354C7122284036588A9793C2" bold="true" pageId="0" pageNumber="31">
<collectingCountry id="AC922DA0FE8CF7C06C7C8A22FBEC2D31" name="Ecuador">Ecuador</collectingCountry>
. 
<collectingRegion id="0ECFFDD738595FDAD4061D6ECB079BFA" country="Ecuador" name="Tungurahua">Tungurahua</collectingRegion>
</emphasis>
: 
<collectingCounty id="A8536101473B12C990FE4D6F4858A55D">
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, 
<collectingMunicipality id="52E32D9A9F8DF8866759466442B05FAB">
Parroquia 
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<normalizedToken id="7C6D1ABB715DE560405F44DF141DE2AE" originalValue="Río">Rio</normalizedToken>
Verde
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, 
<location id="3B206BFA6E2073FE767D3AFB1E2D4171" LSID="urn:lsid:plazi:treatment:79BFC9D3526F5AEAB9A17253EDCDC6F3:3B206BFA6E2073FE767D3AFB1E2D4171" country="Ecuador" county="Canton Banos" latitude="-1.4775918" longLatPrecision="1" longitude="-78.298225" municipality="Parroquia Rio Verde" name="Cerro Candelaria Reserve" stateProvince="Tungurahua">Cerro Candelaria Reserve</location>
(
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EcoMinga), upper 
<collectingRegion id="BDB885BAEC81F1ABA5A050A8ABEE9074" country="Ecuador" name="Pastaza">Pastaza</collectingRegion>
watershed, 
<geoCoordinate id="8297AF62DAD165AC056164465FF1781A" degrees="1" direction="south" minutes="28" orientation="latitude" precision="1" seconds="39.33" value="-1.4775918">1°28'39.33&quot;S</geoCoordinate>
, 
<geoCoordinate id="4424504C0D700163810AECFD5CBFCC2F" degrees="78" direction="west" minutes="17" orientation="longitude" precision="1" seconds="53.61" value="-78.298225">78°17'53.61&quot;W</geoCoordinate>
, 
<quantity id="6D495215011A43FA7A7A62BC0726E841" metricMagnitude="3" metricUnit="m" metricValue="3.642" unit="m" value="3642.0">
<elevation id="7EBB9D4CAA7C6A701DE3FD8C97DA5320" metricMagnitude="3" metricUnit="m" metricValue="3.642" unit="m" value="3642.0">3642 m</elevation>
</quantity>
, 
<collectingDate id="C961D0543A2F2CD58253609A8F09CF86" value="2016-03-10">10 Mar 2016</collectingDate>
, 
<emphasis id="29104F0C7524E9CD420DCD9472558AA0" italics="true" pageId="0" pageNumber="31">
<collectorName id="25F666F4826418E0A58587BD1640AFC4">J.L. Clark</collectorName>
14839
</emphasis>
(
<typeStatus id="934B9F286FD76F781101682FB2E93A4B">holotype</typeStatus>
: QCA! [245371]; isotype: SEL! [079072])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3127C3955CAB66CE2F5263B8F71C87BF" pageId="0" pageNumber="31" type="description">
<paragraph id="E8DEFC638D43748A94DD216F852A4A3B" pageId="0" pageNumber="31">Description.</paragraph>
<paragraph id="9A47229B42E0CBE1626FEE3CA1B06281" pageId="0" pageNumber="31">
Terrestrial or epiphytic herb. 
<emphasis id="28FCAC850EB7B2C54C811DB16F1FBE55" bold="true" italics="true" pageId="0" pageNumber="31">Rhizome</emphasis>
short with multiple underground fusiform to globose root tubers, pale and heavily stippled with dull reddish-purple spots, 5-9 
<normalizedToken id="0539A152D10A69F3BDD10D4A8F1F256D" originalValue="×">x</normalizedToken>
3-4 mm, surface striate, yellowish-brown with red spots (Fig. 
<figureCitation id="F6A5A61D4C11F6FBB457241FD0912519" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bomarea pastazensis A lateral view of flower B front view of flower featuring mature androecium C mature gynoecium D petal E adaxial leaf surface F mature flowers with foliage G habitat H tuberous roots. Photos A by J. L. Clark (J. L. Clark 14839), B-H by N. Exe (N. Exe et al. 2)." figureDoi="10.3897/phytokeys.235.110525.figure2" httpUri="https://binary.pensoft.net/fig/934580" pageId="0" pageNumber="31">2H</figureCitation>
). 
<emphasis id="75FBF8C4A4124C0014A38E33C8D05914" bold="true" italics="true" pageId="0" pageNumber="31">Stem</emphasis>
erect, 10-14 cm long, ca. 0.13 cm in diameter, slender, terete, base pubescent, apex puberulous, internodes 0.6 cm long. 
<emphasis id="0C1BA902B7EFE82C86028A62E2090E0B" bold="true" italics="true" pageId="0" pageNumber="31">Leaves</emphasis>
alternate; blade ovate, 1.4-1.8 
<normalizedToken id="7A1576FF810835D532C2051DC8807852" originalValue="×">x</normalizedToken>
0.4-0.7 cm, base rounded to obtuse, apex acute, abaxially glabrous, adaxially light green, suffused with whitish-translucent trichomes, ca., 0.2 mm long, clustered along veins; blades with 5-7 prominent parallel veins, raised below and slightly raised above; margin hyaline, slightly revolute with unicellular trichomes to 0.2 mm long; petioles resupinate and canaliculate, 0.1-0.4 cm long, basal leaves reduced to scales. 
<emphasis id="75404C6E62383C31F7AA9D46A1DBD47C" bold="true" italics="true" pageId="0" pageNumber="31">Flowers</emphasis>
produced from the stem apex, usually one (rarely two) apical flower(s) per stem.; pedicels 1-1.4 cm long, terete, with brownish red to dark purple trichomes, bisexual, actinomorphic, epigynous. 
<emphasis id="D18C10D242F86EA894945AAC4F81CEF1" bold="true" italics="true" pageId="0" pageNumber="31">Sepals</emphasis>
3, 1.0-1.3 
<normalizedToken id="D51352508EBB8F40AC4415B3C106AAC9" originalValue="×">x</normalizedToken>
1.0-1.2 cm, each sepal with 6 parallel veins, broadly ovate, apically rounded, bright red to orange-red with a 1 mm green to black claw, inner and outer surfaces glabrous. 
<emphasis id="0BB62B46147E51CC97784E5F8173EE50" bold="true" italics="true" pageId="0" pageNumber="31">Petals</emphasis>
3, basally constricted and caniculate, distally obovate to broadly spathulate, 1.5-1.6 
<normalizedToken id="5A6A995C785F5957EC679AC4840CE333" originalValue="×">x</normalizedToken>
0.6-0.7 cm, greenish yellow, puberulous at base, inner surface with 3 large reddish-brown spots, outer surface uniformly yellow (occasionally yellow suffused with red from the inner spots). 
<emphasis id="75AE7691E88FFF768530F8D215AA13D6" bold="true" italics="true" pageId="0" pageNumber="31">Androecium</emphasis>
of 6 free stamens, 0.8-0.9 cm long, thickened near center of filaments; anthers pseudo-basifixed, 0.3 
<normalizedToken id="3BA99FE43EA2C13BC441CDB2B5153FFC" originalValue="×">x</normalizedToken>
0.1 cm, fusiform; pollen grains lilac. 
<emphasis id="FF45A4075FD2B5440E7482871363D168" bold="true" italics="true" pageId="0" pageNumber="31">Gynoecium</emphasis>
comprised of three fused carpels, ovary 0.5 
<normalizedToken id="0B488DFE0FAD7CA77229036D27FFEAB8" originalValue="×">x</normalizedToken>
0.4 cm with surface covered with glutinous trichomes, style ca. 1 cm long, stigma with three circinate lobes. 
<emphasis id="B3EDA7EDD07041DE15ED5F41BB52294C" bold="true" italics="true" pageId="0" pageNumber="31">Fruits</emphasis>
not observed.
</paragraph>
</subSubSection>
<subSubSection id="6C964A6890E59466979059F64B4AEE45" pageId="0" pageNumber="31" type="ecology">
<paragraph id="CF8C01557D08B862DE6FF0DF0631DB36" pageId="0" pageNumber="31">Ecology.</paragraph>
<paragraph id="E29105E7DC083F036E5CC2813A53A517" pageId="0" pageNumber="31">
Found growing on 
<taxonomicName id="90909C150340FD9C3562B624F242086B" authorityName="C.Linnaeus" authorityYear="1753" class="Bryopsida" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sphagnum" order="Sphagnales" pageId="0" pageNumber="31" phylum="Bryophyta" rank="genus">
<emphasis id="582CE4E56FC7DF03DFE3BE64983617A9" italics="true" pageId="0" pageNumber="31">Sphagnum</emphasis>
</taxonomicName>
and mossy cushions at ground level and epiphytically (up to 2.5 m high) in high elevation cloud forest and paramo (observed from 3235 to 3700 m). Plants growing in partially shaded areas to full sunlight. Stems erect to hanging, with flowers often found on or slightly above ground level. Flowers protandrous, commonly with one flower per stem but occasionally two.
</paragraph>
</subSubSection>
<subSubSection id="0EBB0A4EE3F891A8C121BBC14BB69017" pageId="0" pageNumber="31" type="phenology">
<paragraph id="5D026894002C58418266AB82C259ED39" pageId="0" pageNumber="31">Phenology.</paragraph>
<paragraph id="D6CFC69BD542842FEE3F2CE051909CCB" pageId="0" pageNumber="31">Observed in flower in January, March, July, and November. Approximately 20 individual plants with mature flowers were located during an expedition in November of 2022 and fewer individuals with mature flowers were observed in January of 2022. Fruits not documented.</paragraph>
</subSubSection>
<subSubSection id="CD51EAC013D3F2F3280B4DF5DC200B77" pageId="0" pageNumber="31" type="etymology">
<paragraph id="9D5F6E2C82A69EECC28B247010E02924" pageId="0" pageNumber="31">Etymology.</paragraph>
<paragraph id="B95C4B9059C1D644B97D88044745FA7F" pageId="0" pageNumber="31">
The specific epiphyte, 
<emphasis id="AD1D4E5E69342D8A1D57EC5A7A46211E" italics="true" pageId="0" pageNumber="31">pastazensis</emphasis>
, reflects the watershed of the type locality that includes 
<normalizedToken id="FC5C5EB8C4E525E0CF8D991B48EC4E37" originalValue="Río">Rio</normalizedToken>
Pastaza and adjacent tributaries.
</paragraph>
</subSubSection>
<subSubSection id="D8573E951D9CD8473B2DC3984A40268E" pageId="0" pageNumber="31" type="distribution">
<paragraph id="F13F713AEBBE5813742E22C999BFC0F2" pageId="0" pageNumber="31">Distribution and preliminary assessment of conservation status.</paragraph>
<paragraph id="1E6279F4CBAC40DC494679BF2C54BA20" pageId="0" pageNumber="31">
<taxonomicName id="6DE711126F4EADAE2D3DA2818C69C9BA" authorityName="J. L. Clark, Fierro-Minda &amp; N. Exe" authorityYear="2023" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="9BF11687334D277858CC103FA9F39C4D" italics="true" pageId="0" pageNumber="31">Bomarea pastazensis</emphasis>
</taxonomicName>
is endemic to the upper Pastaza watershed, located in the eastern Andean slopes of Ecuador. The first documented population was inside Cerro Candelaria Reserve in Tungurahua province (Fig. 
<figureCitation id="23E249FA54C0C76C6CCCE7BF4C4C17CD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution map of Bomarea pastazensis. Red dots indicate collections or observations." figureDoi="10.3897/phytokeys.235.110525.figure3" httpUri="https://binary.pensoft.net/fig/934581" pageId="0" pageNumber="31">3</figureCitation>
). The Cerro Candelaria Reserve is a private reserve of 2800+ hectares managed by 
<normalizedToken id="979E2056E035C2DB112D624DC3521BBD" originalValue="Fundación">Fundacion</normalizedToken>
EcoMinga. It is bordered on the north by 
<normalizedToken id="BC436422C4F083FA9DDC33DB9AD46DBA" originalValue="EcoMinga’s">EcoMinga's</normalizedToken>
Naturetrek Reserve, and the south by Parque Nacional Sangay. Along with 
<normalizedToken id="585ED8089F804CC82922B6AED3D800A2" originalValue="Fundación">Fundacion</normalizedToken>
<normalizedToken id="41CA585CCE85A78CD0580619DCFC2CD0" originalValue="EcoMinga’s">EcoMinga's</normalizedToken>
Machay and Naturetrek Reserves to the north, it forms a protected corridor between Parque Nacional Sangay and Parque Nacional Llanganates. Cerro Candelaria Reserve was founded by 
<normalizedToken id="FDC7648FA7205A9E7213BD8825D40690" originalValue="Fundación">Fundacion</normalizedToken>
EcoMinga in 2007 and financed by the World Land Trust (UK); the majority of the reserve is undisturbed by anthropogenic activity. A diverse flora and fauna has been recorded here, including many endemic species and species of high conservation priority (
<bibRefCitation id="F4C3E97B1589E221870F72A4C372466A" author="Jost, L" journalOrPublisher="Lyonia" pageId="0" pageNumber="31" pagination="41 - 47" refId="B16" refString="Jost, L, 2004. Explosive local radiation of the genus Teagueia (Orchidaceae) in the Upper Pastaza Watershed of Ecuador. Lyonia 7: 41 - 47" title="Explosive local radiation of the genus Teagueia (Orchidaceae) in the Upper Pastaza Watershed of Ecuador." volume="7" year="2004">Jost 2004</bibRefCitation>
; 
<bibRefCitation id="6F9288D9529C994C896F9BDE0EE45F40" author="Reyes-Puig, J" journalOrPublisher="Timber Press, Portland, Oregon, USA" pageId="0" pageNumber="31" refId="B20" refString="Reyes-Puig, J, Ramirez, S, Yanez-Munoz, M, Morales, M, Recalde, L, 2013. Bosque Protector Cerro Candelaria: Una isla de biodiversidad entre las nubes. In: MECN, JOCOTOCO y ECOMINGA. Herpetofauna en Areas prioritarias para la conservacion: El sistema de Reservas Jocotoco y EcoMinga. Monografia 6. Serie de publicaciones del Museo Ecuatoriano de Ciencias Naturales (MECN), Fundacion para la Conservacion Jocotoco, and Fundacion EcoMinga. Quito, Ecuador, 196-211." title="Bosque Protector Cerro Candelaria: Una isla de biodiversidad entre las nubes. In: MECN, JOCOTOCO y ECOMINGA. Herpetofauna en Areas prioritarias para la conservacion: El sistema de Reservas Jocotoco y EcoMinga. Monografia 6. Serie de publicaciones del Museo Ecuatoriano de Ciencias Naturales (MECN), Fundacion para la Conservacion Jocotoco, and Fundacion EcoMinga. Quito, Ecuador, 196 - 211." year="2013">Reyes-Puig et al. 2013</bibRefCitation>
), such as 
<taxonomicName id="E029A54683C052BEEBAEACAC19D56A08" class="Liliopsida" family="Orchidaceae" genus="Blakea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Blakea attenboroughii" order="Asparagales" pageId="0" pageNumber="31" phylum="Liliopsida" rank="species" species="attenboroughii">
<emphasis id="2CD96610CDDC3EFB75C47125F3C03983" italics="true" pageId="0" pageNumber="31">Blakea attenboroughii</emphasis>
</taxonomicName>
Penneys &amp; L.Jost (
<taxonomicName id="2307CABECC8F6D18EEA21CE237869E70" class="Dicotyledones" family="Melastomataceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="0" pageNumber="31" phylum="Angiospermae" rank="family">Melastomataceae</taxonomicName>
) which is endemic to the lower elevation forests of Bosque Protector Cerro Candelaria. Several recently described species in the 
<taxonomicName id="8B3C83EBC4A8C57A89F16BF3E66DF5F3" class="Liliopsida" family="Orchidaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Asparagales" pageId="0" pageNumber="31" phylum="Liliopsida" rank="family">Orchidaceae</taxonomicName>
share a similar geographic distribution to 
<taxonomicName id="998400A9579420EDC5CDDC791B8A1A4A" authorityName="J. L. Clark, Fierro-Minda &amp; N. Exe" authorityYear="2023" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="94D1972779094BF9F53AFA9FCC19FB52" italics="true" pageId="0" pageNumber="31">Bomarea pastazensis</emphasis>
</taxonomicName>
(Fig. 
<figureCitation id="AA27F1531C503B7117B6FC8697036C7D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution map of Bomarea pastazensis. Red dots indicate collections or observations." figureDoi="10.3897/phytokeys.235.110525.figure3" httpUri="https://binary.pensoft.net/fig/934581" pageId="0" pageNumber="31">3</figureCitation>
), include 
<taxonomicName id="8E47FED2014C67C358F8D09E91791DEB" authorityName="L.Jost &amp; Shepard" authorityYear="2011" class="Liliopsida" family="Orchidaceae" genus="Teagueia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Teagueia barbeliana" order="Asparagales" pageId="0" pageNumber="31" phylum="Liliopsida" rank="species" species="barbeliana">
<emphasis id="4E6539B7B03FA64F5B64EDA3EAC4214A" italics="true" pageId="0" pageNumber="31">Teagueia barbeliana</emphasis>
</taxonomicName>
L.Jost &amp; Shepard, 
<taxonomicName id="F3159BADB6A410FF596622AD84A5456F" lsidName="T. puroana" pageId="0" pageNumber="31" rank="species" species="puroana">
<emphasis id="CF294771693EAB9F03C3BDF0239B54B5" italics="true" pageId="0" pageNumber="31">T. puroana</emphasis>
</taxonomicName>
L.Jost &amp; Shepard, 
<taxonomicName id="FB8C8177A88F94C49983999E06D7F48F" lsidName="T. kostoglouana" pageId="0" pageNumber="31" rank="species" species="kostoglouana">
<emphasis id="B04356EE41715C95008A4946C1959269" italics="true" pageId="0" pageNumber="31">T. kostoglouana</emphasis>
</taxonomicName>
L.Jost &amp; Shepard, 
<taxonomicName id="67CFD9BFCF2545535DBD2E5AA524C51F" lsidName="T. lizziefinchiana" pageId="0" pageNumber="31" rank="species" species="lizziefinchiana">
<emphasis id="189ED7984DF7035DF31C05932602907D" italics="true" pageId="0" pageNumber="31">T. lizziefinchiana</emphasis>
</taxonomicName>
L.Jost &amp; Shepard, and 
<taxonomicName id="A31ABFC1D790DEDA7C6B120E7A07A1D6" lsidName="T. anitana" pageId="0" pageNumber="31" rank="species" species="anitana">
<emphasis id="C71A6C27FD384FD77FB7B0DF01A916A9" italics="true" pageId="0" pageNumber="31">T. anitana</emphasis>
</taxonomicName>
L.Jost &amp; Shepard (
<bibRefCitation id="5D053C8A66F367DC8592905D5B6E90A6" DOI="https://doi.org/10.15517/lank.v11i1.18310" author="Jost, L" journalOrPublisher="Lankesteriana" pageId="0" pageNumber="31" pagination="9 - 14" refId="B17" refString="Jost, L, Shepard, A, 2011. Two new species of Teagueia (Orchidaceae: Pleurothallidinae) from East-Central Ecuador. Lankesteriana 1 (11): 9 - 14, DOI: https://doi.org/10.15517/lank.v11i1.18310" title="Two new species of Teagueia (Orchidaceae: Pleurothallidinae) from East-Central Ecuador." url="https://doi.org/10.15517/lank.v11i1.18310" volume="1" year="2011">Jost and Shepard 2011</bibRefCitation>
; 
<bibRefCitation id="23821E97AB5B9DF8BF2EFD829F4D82E9" DOI="https://doi.org/10.15517/lank.v17i2.30159" author="Jost, L" journalOrPublisher="Lankesteriana" pageId="0" pageNumber="31" pagination="261 - 268" refId="B18" refString="Jost, L, Shepard, A, 2017. Four new Teagueia (Orchidaceae: Pleurothallidinae) from the Upper Rio Pastaza watershed of East-Central Ecuador. Lankesteriana 2 (17): 261 - 268, DOI: https://doi.org/10.15517/lank.v17i2.30159" title="Four new Teagueia (Orchidaceae: Pleurothallidinae) from the Upper Rio Pastaza watershed of East-Central Ecuador." url="https://doi.org/10.15517/lank.v17i2.30159" volume="2" year="2017">2017</bibRefCitation>
). Additional surveys in Cerro Candelaria and in the surrounding upper Rio Pastaza watershed are likely to yield many more undescribed species, and potentially additional populations of 
<taxonomicName id="AE83583ADA54789905D65041E0B11360" authorityName="J. L. Clark, Fierro-Minda &amp; N. Exe" authorityYear="2023" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="828CCAB1CA177D70B014ED975B0F5080" italics="true" pageId="0" pageNumber="31">Bomarea pastazensis</emphasis>
</taxonomicName>
. The distribution of this species highlights the importance of the Llanganates-Sangay corridor for allowing species gene flow and connectivity among populations (
<bibRefCitation id="595030C778BCB744EB9CE24A3F1BD17E" author="Rios Alvear, G" journalOrPublisher="Yachana Revista Cientifica" pageId="0" pageNumber="31" pagination="11 - 21" refId="B21" refString="Rios Alvear, G, Reyes-Puig, C, 2015. Corredor ecologico Llanganates-Sangay: Un acercamiento hacia su manejo y funcionalidad. Yachana Revista Cientifica 4: 11 - 21" title="Corredor ecologico Llanganates-Sangay: Un acercamiento hacia su manejo y funcionalidad." volume="4" year="2015">
<normalizedToken id="12E1C5C11D71F4DDD9020198B2A7CBAE" originalValue="Ríos">Rios</normalizedToken>
Alvear and Reyes-Puig 2015
</bibRefCitation>
). A second documented population of 
<taxonomicName id="D2605D6612A8C043045536004681204A" authorityName="J. L. Clark, Fierro-Minda &amp; N. Exe" authorityYear="2023" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="E20352412480323432D4777F38473113" italics="true" pageId="0" pageNumber="31">Bomarea pastazensis</emphasis>
</taxonomicName>
was provided by digital images from 
<normalizedToken id="B5B8625331A44CA79C8FC0EF08019032" originalValue="EcoMinga’s">EcoMinga's</normalizedToken>
park guard Eduardo 
<normalizedToken id="3EC0AE0A00FD1B2FCAE1E1627C56EE32" originalValue="Peña">Pena</normalizedToken>
(July 2023) in the Llanganates National Park, north of 
<normalizedToken id="DC22E24D01C7942B89214E3BD91756C5" originalValue="Río">Rio</normalizedToken>
Pastaza (Fig. 
<figureCitation id="C53FE6027FE56E8282DF46170A466A00" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution map of Bomarea pastazensis. Red dots indicate collections or observations." figureDoi="10.3897/phytokeys.235.110525.figure3" httpUri="https://binary.pensoft.net/fig/934581" pageId="0" pageNumber="31">3</figureCitation>
). We calculated the AOO=12 km2 from 
<normalizedToken id="603E2E9640181811D806E4B437E28491" originalValue="Peña’s">Pena's</normalizedToken>
observation and the type collection (Fig. 
<figureCitation id="C4D9C76BD91B2A7981679006662E44DA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution map of Bomarea pastazensis. Red dots indicate collections or observations." figureDoi="10.3897/phytokeys.235.110525.figure3" httpUri="https://binary.pensoft.net/fig/934581" pageId="0" pageNumber="31">3</figureCitation>
). Based on the available information and according to the IUCN Red List criteria (
<bibRefCitation id="68C5C36198F6F630499175C5ED767251" author="Hofreiter, A" journalOrPublisher="Feddes Repertorium" pageId="0" pageNumber="31" refId="B14" refString="IUCN (2012) IUCN Red List Categories and Criteria: Version 3.1. 2nd edn. IUCN, Gland, Switzerland and Cambridge." year="2002">IUCN 2012</bibRefCitation>
; 
<bibRefCitation id="F2E7114981F9370D468113ED90D9F184" author="Hofreiter, A" journalOrPublisher="Feddes Repertorium" pageId="0" pageNumber="31" publicationUrl="https://www.iucnredlist.org/resources/redlistguidelines" refId="B15" refString="2022. . https://www.iucnredlist.org/resources/redlistguidelines" url="https://www.iucnredlist.org/resources/redlistguidelines" year="2022">IUCN Standards and Petitions Committee 2022</bibRefCitation>
), 
<taxonomicName id="3CCAEB411E0B53650342DB26F90F0ED2" authorityName="J. L. Clark, Fierro-Minda &amp; N. Exe" authorityYear="2023" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="3F39A563C76EEBD6CDDD007A5EDBE10D" italics="true" pageId="0" pageNumber="31">Bomarea pastazensis</emphasis>
</taxonomicName>
is preliminarily assessed as Vulnerable (VU) based on a limited area of occupancy (IUCN criterion D2 where AOO &lt;20 km2) and dependence on conservation efforts for its continued survival. The only documented populations of 
<taxonomicName id="13CB9B2F7F6F1D8F3878257B0F99DCAF" authorityName="J. L. Clark, Fierro-Minda &amp; N. Exe" authorityYear="2023" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="2A6D3E69F012B99C8CF0A201290358DE" italics="true" pageId="0" pageNumber="31">Bomarea pastazensis</emphasis>
</taxonomicName>
are located inside protected areas, at elevations that are used for agriculture in unprotected parts of the Pastaza province. Effective conservation of this and the other unique species of the Pastaza watershed will require constant vigilance.
</paragraph>
</subSubSection>
<subSubSection id="500A8EEB579B405A0F4E39C177F479F2" pageId="0" pageNumber="31" type="comments">
<paragraph id="6DBE0F6375CEC002942AC3B599616C3A" pageId="0" pageNumber="31">Comments.</paragraph>
<paragraph id="7ABF8674006B4D4596DA0FFF9E9C2426" pageId="0" pageNumber="31">
<taxonomicName id="882B7946407672B3241B185BA586D1C7" authorityName="J. L. Clark, Fierro-Minda &amp; N. Exe" authorityYear="2023" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="88718F1AF45D4958502AE8FA27AB9F14" italics="true" pageId="0" pageNumber="31">Bomarea pastazensis</emphasis>
</taxonomicName>
differs from other congeners by a distinctive pattern of three dark spots on the petals (Fig. 
<figureCitation id="156081DD978A6DB0C1D9DE4705865E26" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bomarea pastazensis A lateral view of flower B front view of flower featuring mature androecium C mature gynoecium D petal E adaxial leaf surface F mature flowers with foliage G habitat H tuberous roots. Photos A by J. L. Clark (J. L. Clark 14839), B-H by N. Exe (N. Exe et al. 2)." figureDoi="10.3897/phytokeys.235.110525.figure2" httpUri="https://binary.pensoft.net/fig/934580" pageId="0" pageNumber="31">2D</figureCitation>
), widely opened flowers, and small size. 
<taxonomicName id="0355598B17A6350BBA0E250627720552" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="7F13FA0EFA78A766B9454ED0019F0AFC" italics="true" pageId="0" pageNumber="31">Bomarea pumila</emphasis>
</taxonomicName>
and 
<taxonomicName id="6C79C2E1BD5592974E4FAE18D662B066" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="9F68FEBF38A99C57EF5BFE517A99DE71" italics="true" pageId="0" pageNumber="31">B. pastazensis</emphasis>
</taxonomicName>
share similar traits such as their small size in comparison with the rest of 
<taxonomicName id="6783EE294383E9DD1C60F82D82311FFA" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="A1697D70140E22882269F914C2B23D6D" italics="true" pageId="0" pageNumber="31">Bomarea</emphasis>
</taxonomicName>
species, the presence of pubescence throughout their above-surface organs, and the color patterns of the flowers (Figs 
<figureCitation id="F6543A67B6509A69489CEBE4B7C2671D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bomarea pastazensis A lateral view of flower B front view of flower featuring mature androecium C mature gynoecium D petal E adaxial leaf surface F mature flowers with foliage G habitat H tuberous roots. Photos A by J. L. Clark (J. L. Clark 14839), B-H by N. Exe (N. Exe et al. 2)." figureDoi="10.3897/phytokeys.235.110525.figure2" httpUri="https://binary.pensoft.net/fig/934580" pageId="0" pageNumber="31">2</figureCitation>
, 
<figureCitation id="3F2C4D853B74F32167D23422727034BC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bomarea pumila A-D field images from Peru E syntype (K! [K 000400912]) of Bomarea pumila of W. Lechler 2240 from Sachapata, Peru. Photos A-D from Lucely L. Vilca Bustamente E from The Herbarium Catalogue, Royal Botanic Gardens, Kew. The Board of Trustees of the RBG, Kew." figureDoi="10.3897/phytokeys.235.110525.figure5" httpUri="https://binary.pensoft.net/fig/934583" pageId="0" pageNumber="31">5</figureCitation>
; Table 
<tableCitation id="19EFBB43311BB155F8CA17F982ED771A" captionStart="Table 1" captionStartId="T1" captionText="Table 1. General geographic distribution (names in parentheses indicate Ecuadorian province) and comparison of morphological characters between Bomarea pastazensis and B. pumila." httpUri="http://table.plazi.org/id/870C79E5CCECBCF058358784D67DC573" pageId="0" pageNumber="31" tableUuid="870C79E5CCECBCF058358784D67DC573">1</tableCitation>
). 
<taxonomicName id="C92691BA9DB364A26D58AB4BDEFDFC36" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="0C608FECD1F7FFA74773F95788EB33BA" italics="true" pageId="0" pageNumber="31">B. pastazensis</emphasis>
</taxonomicName>
is slightly bigger in size (10-14 cm tall) than 
<taxonomicName id="117B7FE6E2D3FD46CFD8B41ED02E8A1C" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="E4409DE55A112B68F53B5ABA8C5479DF" italics="true" pageId="0" pageNumber="31">B. pumila</emphasis>
</taxonomicName>
(5-9 cm tall). In contrast, 
<taxonomicName id="463FC692B61EF01F90BBC30BB90DA2FE" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="093600825FA3B56750D52FBC1480AC27" italics="true" pageId="0" pageNumber="31">B. pastazensis</emphasis>
</taxonomicName>
is sparsely pubescent at the base of the sepals, while the sepals of 
<taxonomicName id="EA960D34EEC6AE0768F0D0D4E0F4A0FC" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="BC3C539C06BACBAD4F9754165AB08E54" italics="true" pageId="0" pageNumber="31">B. pumila</emphasis>
</taxonomicName>
are uniformly densely pubescent (Fig. 
<figureCitation id="3D057FDCB519A64D6662C8E7DC29B724" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Bomarea pumila A-D field images from Peru E syntype (K! [K 000400912]) of Bomarea pumila of W. Lechler 2240 from Sachapata, Peru. Photos A-D from Lucely L. Vilca Bustamente E from The Herbarium Catalogue, Royal Botanic Gardens, Kew. The Board of Trustees of the RBG, Kew." figureDoi="10.3897/phytokeys.235.110525.figure5" httpUri="https://binary.pensoft.net/fig/934583" pageId="0" pageNumber="31">5</figureCitation>
). The petals of 
<taxonomicName id="741E2A08BE96A5A85BB29DBC40EAC9A7" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="35D60287B44BB8B2975098DB21068C02" italics="true" pageId="0" pageNumber="31">B. pumila</emphasis>
</taxonomicName>
are not spotted, which differentiates it from 
<taxonomicName id="83607A0C99EB5FEF7895D44C3E214BDF" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="407D1EF7B509D769A338D0BAB95B98B2" italics="true" pageId="0" pageNumber="31">B. pastazensis</emphasis>
</taxonomicName>
which has a three-spotted pattern on the petals (Fig. 
<figureCitation id="3630881969BCD4D1742490CE0329C041" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bomarea pastazensis A lateral view of flower B front view of flower featuring mature androecium C mature gynoecium D petal E adaxial leaf surface F mature flowers with foliage G habitat H tuberous roots. Photos A by J. L. Clark (J. L. Clark 14839), B-H by N. Exe (N. Exe et al. 2)." figureDoi="10.3897/phytokeys.235.110525.figure2" httpUri="https://binary.pensoft.net/fig/934580" pageId="0" pageNumber="31">2B</figureCitation>
). The two species are geographically isolated with 
<taxonomicName id="068B74BE2AC20370C865ECE5F7DFCB51" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="F5560FFC82F75F59C88798A0D3A30BE4" italics="true" pageId="0" pageNumber="31">B. pastazensis</emphasis>
</taxonomicName>
endemic to the Pastaza watershed on the eastern Andean slopes in central Ecuador (Tungurahua province) and 
<taxonomicName id="70EBDF92CAC77061A42D586872AE6E47" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="42E9BEAAA72ACA7BB647EFF03B359B7E" italics="true" pageId="0" pageNumber="31">B. pumila</emphasis>
</taxonomicName>
from Central Peru to northern Bolivia. Table 
<tableCitation id="3DDE4577E0D8E21DE3963FAFC3A0A931" captionStart="Table 1" captionStartId="T1" captionText="Table 1. General geographic distribution (names in parentheses indicate Ecuadorian province) and comparison of morphological characters between Bomarea pastazensis and B. pumila." httpUri="http://table.plazi.org/id/870C79E5CCECBCF058358784D67DC573" pageId="0" pageNumber="31" tableUuid="870C79E5CCECBCF058358784D67DC573">1</tableCitation>
provides a summary of distribution and useful characters for differentiating 
<taxonomicName id="CD4F762A25813F32CA59A7790CDA6087" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="D3C91A73CE12BE4627905114CCC20CF8" italics="true" pageId="0" pageNumber="31">B. pastazensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="46A556B45468228C4EC48D4E91BEFC80" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="EE8C378C184AEEC46E80D6EDBFF650A4" italics="true" pageId="0" pageNumber="31">B. pumila</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="870C79E5CCECBCF058358784D67DC573" ID-Table-UUID="870C79E5CCECBCF058358784D67DC573" httpUri="http://table.plazi.org/id/870C79E5CCECBCF058358784D67DC573" pageId="0" pageNumber="31" start="Table 1" startId="T1">
<paragraph id="FA83B60F30458425E32289AD97DF0FF5" pageId="0" pageNumber="31">
<emphasis id="AEF893EF3D703AAF59D04D57E5902F02" bold="true" pageId="0" pageNumber="31">Table 1.</emphasis>
General geographic distribution (names in parentheses indicate Ecuadorian province) and comparison of morphological characters between 
<taxonomicName id="28638B2A7B12A096D8E70ED8D378E068" authorityName="J. L. Clark, Fierro-Minda &amp; N. Exe" authorityYear="2023" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="4BB2205447BEFF4073439A5938CD398F" italics="true" pageId="0" pageNumber="31">Bomarea pastazensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="62BB8BF2C407EE6DBCA8E8CDE3CB00A8" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="B. pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="E25DCFE11FC6674A80243939D856F7CD" italics="true" pageId="0" pageNumber="31">B. pumila</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="3D53522327B9913AD82BCFAB0F6B1226" pageId="0" pageNumber="31">
<table id="D577B85D91C5D16959BD3EDA879C1BF5" pageId="0" pageNumber="31">
<tr id="35684767F0F5C3D9E0F970B40EBA72CE" pageId="0" pageNumber="31">
<th id="F55AED6EC3C1914B32FCDFA31EA6123B" colspan="1" isEmpty="true" pageId="0" pageNumber="31" rowspan="1">-</th>
<th id="BA50E72EC84560F0B41C164A44106684" colspan="1" pageId="0" pageNumber="31" rowspan="1">
<taxonomicName id="B68CC7D917228658B8D8F67D7EF0EAE8" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="7E40D025279A2BD97CF40D15F9D7B699" italics="true" pageId="0" pageNumber="31">Bomarea pumila</emphasis>
</taxonomicName>
</th>
<th id="7F658D8827BFA232077BF737AD111F92" colspan="1" pageId="0" pageNumber="31" rowspan="1">
<taxonomicName id="2C34783F5999740B4D3EAE947FE5827B" authorityName="J. L. Clark, Fierro-Minda &amp; N. Exe" authorityYear="2023" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="DDEAE0F37D827FF9B659E949DD4ECEFA" italics="true" pageId="0" pageNumber="31">Bomarea pastazensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="A22D2A52BEA51A3F6DF6D3A2EC2B20DB" pageId="0" pageNumber="31">
<td id="49C322535FF0E18EB733F58856575BB0" colspan="1" pageId="0" pageNumber="31" rowspan="1">Habit</td>
<td id="782B32ADF7150F59DBBBE662F285822F" colspan="1" pageId="0" pageNumber="31" rowspan="1">erect</td>
<td id="9DE191CA567F38A8064555F548CC8020" colspan="1" pageId="0" pageNumber="31" rowspan="1">erect to hanging</td>
</tr>
<tr id="EAE1A0115FFA6521D58077B036F5638D" pageId="0" pageNumber="31">
<td id="435270F1740051D8DDAE5E9B639CEFF7" colspan="1" pageId="0" pageNumber="31" rowspan="1">Plant height</td>
<td id="87DE510602A2738C287CD5CB9C07A05F" colspan="1" pageId="0" pageNumber="31" rowspan="1">5-9 cm long</td>
<td id="7B332E3A251A13B6DA36B76F2A63A58A" colspan="1" pageId="0" pageNumber="31" rowspan="1">10-14 cm long</td>
</tr>
<tr id="8366B33A3E5149CB2CB734CD7D56F820" pageId="0" pageNumber="31">
<td id="E3331B815FDF808DE1AAB9C1ED032885" colspan="1" pageId="0" pageNumber="31" rowspan="1">Stem surface</td>
<td id="CC4F41D29859AD6126F2168504F9309A" colspan="1" pageId="0" pageNumber="31" rowspan="1">glabrous</td>
<td id="D9552F015EE25DE4A95A62E27DAC2DB9" colspan="1" pageId="0" pageNumber="31" rowspan="1">pubescent</td>
</tr>
<tr id="12EF7B3C076A465B3AE59E14FB669395" pageId="0" pageNumber="31">
<td id="A151284F89FE09A1035966FABBDC1AE1" colspan="1" pageId="0" pageNumber="31" rowspan="1">Leaf - relative size</td>
<td id="B027378DEF1589ACAAD6485396CA68DF" colspan="1" pageId="0" pageNumber="31" rowspan="1">leaves not uniform on stem; central leaves relatively larger compared with basal and apical leaves</td>
<td id="D883225261D02DDD4FA2FD35194C2F38" colspan="1" pageId="0" pageNumber="31" rowspan="1">leaves relatively uniform on stem</td>
</tr>
<tr id="99798859407F32AB617AC1F60ED14DBF" pageId="0" pageNumber="31">
<td id="056BDB4382EE4455AE023E287DDA5203" colspan="1" pageId="0" pageNumber="31" rowspan="1">Leaf margin</td>
<td id="CF368B1B75505CBA0333D477F9EC8ACD" colspan="1" pageId="0" pageNumber="31" rowspan="1">glabrous</td>
<td id="4FC83BB315C51A7DCDDF2D7B2F60C7F4" colspan="1" pageId="0" pageNumber="31" rowspan="1">ciliate</td>
</tr>
<tr id="F1C4D71859CD8E0D03A0A2C8B8FEC925" pageId="0" pageNumber="31">
<td id="AE46E58F4032C73F78D5C7DFA2835078" colspan="1" pageId="0" pageNumber="31" rowspan="1">Pedicel length</td>
<td id="EAEE6FC5DB2C44947C7EAEC27688C80A" colspan="1" pageId="0" pageNumber="31" rowspan="1">1.5-4 cm long</td>
<td id="16B8D42D6DD3C353351BAE4059706BE3" colspan="1" pageId="0" pageNumber="31" rowspan="1">1-1.4 cm long</td>
</tr>
<tr id="D85264BA4F1CD60FC2DE605B7E40F3E4" pageId="0" pageNumber="31">
<td id="7DF1565F840115489CCC473B3597CB85" colspan="1" pageId="0" pageNumber="31" rowspan="1">Flower length</td>
<td id="A3DC8820E97DB43EF7B107A4F224FFA4" colspan="1" pageId="0" pageNumber="31" rowspan="1">0.8-1.5 cm long</td>
<td id="B428EB815689C22346F46FE0E1502CA1" colspan="1" pageId="0" pageNumber="31" rowspan="1">1.5-1.6 cm long</td>
</tr>
<tr id="F75BE4A9554E663D10A9928721FAE190" pageId="0" pageNumber="31">
<td id="6AF8CC92F214F3AF5F8014AC367A1F46" colspan="1" pageId="0" pageNumber="31" rowspan="1">Flower width during anthesis</td>
<td id="7824DC07A5D1C7FC16666F450A4BD46F" colspan="1" pageId="0" pageNumber="31" rowspan="1">2-3 cm wide</td>
<td id="F1EA2A54913F85694AF54BD5C70E7A12" colspan="1" pageId="0" pageNumber="31" rowspan="1">1.5-2 cm wide</td>
</tr>
<tr id="77CC16EC14E260E3950454C2BFCC3E19" pageId="0" pageNumber="31">
<td id="76A069655171E90EF305C9CDECA17BD1" colspan="1" pageId="0" pageNumber="31" rowspan="1">Sepal surface</td>
<td id="86F8AF8A437EE2927BF0E98FCC3C9C6E" colspan="1" pageId="0" pageNumber="31" rowspan="1">conspicuously pubescent</td>
<td id="D586E568FC5B93D998F1C90E03DEDC08" colspan="1" pageId="0" pageNumber="31" rowspan="1">sparsely pubescent at base only</td>
</tr>
<tr id="973D3F98F18620CEEA1982B3C1BCD9AC" pageId="0" pageNumber="31">
<td id="482304297B3265574AC3BE31B8F64B5C" colspan="1" pageId="0" pageNumber="31" rowspan="1">Sepal horn</td>
<td id="9916BADF61AA1389A37473448990A92E" colspan="1" pageId="0" pageNumber="31" rowspan="1">white to reddish horn at apex</td>
<td id="7DF6A1C9C89DB8E5585C713C8A0024E2" colspan="1" pageId="0" pageNumber="31" rowspan="1">green to black horn at apex</td>
</tr>
<tr id="C2EF3F43DE9AB4122CD0A5BC335FCD90" pageId="0" pageNumber="31">
<td id="ECE81EF2765112D86551553CE44A5157" colspan="1" pageId="0" pageNumber="31" rowspan="1">Petal coloration (=inner tepals)</td>
<td id="EE4043581654C66CCB04781249E3692E" colspan="1" pageId="0" pageNumber="31" rowspan="1">yellow with a red stripe and green tip.</td>
<td id="665B9A031B6A9A3BF4A6372581AA9D2F" colspan="1" pageId="0" pageNumber="31" rowspan="1">green suffused with yellow with three prominent red spots at the base</td>
</tr>
<tr id="B7C1CC7540BEDFC4E26B1398F207BA10" pageId="0" pageNumber="31">
<td id="3DBC1C4D4071D03F8B4F9CEFEAB70297" colspan="1" pageId="0" pageNumber="31" rowspan="1">Distribution</td>
<td id="4FB2954F465F3B0F73BAB99F5984E23A" colspan="1" pageId="0" pageNumber="31" rowspan="1">Peru and Bolivia</td>
<td id="8DBDC44E9E183647C6956DFB42F8A84F" colspan="1" pageId="0" pageNumber="31" rowspan="1">Ecuador (Tungurahua)</td>
</tr>
</table>
</paragraph>
<caption id="690F35EFC3AAD39A0340BDBCC40F9671" doi="10.3897/phytokeys.235.110525.figure4" httpUri="https://binary.pensoft.net/fig/934582" pageId="0" pageNumber="31" start="Figure 4" startId="F4">
<paragraph id="27304E195C5C2CC5B323BDE039E89D59" pageId="0" pageNumber="31">
<emphasis id="E7A55F1ECA9D46CDF6D227A2751904D6" bold="true" pageId="0" pageNumber="31">Figure 4.</emphasis>
Holotype of 
<taxonomicName id="614BD6F1D12F7D7F79BC32A91045C359" authorityName="J. L. Clark, Fierro-Minda &amp; N. Exe" authorityYear="2023" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pastazensis" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pastazensis">
<emphasis id="03C1D9C77A31CBB9729ED1A48CCE6215" italics="true" pageId="0" pageNumber="31">Bomarea pastazensis</emphasis>
</taxonomicName>
, 
<emphasis id="5A66DE9E42C5AA037C40201978C7B56B" italics="true" pageId="0" pageNumber="31">J.L. Clark 14839</emphasis>
(QCA).
</paragraph>
</caption>
<caption id="F56EE7D139BC2A516287068BD6D885A2" doi="10.3897/phytokeys.235.110525.figure5" httpUri="https://binary.pensoft.net/fig/934583" pageId="0" pageNumber="31" start="Figure 5" startId="F5">
<paragraph id="83B1C207A2D1ACA528244E0E0A984EA7" pageId="0" pageNumber="31">
<emphasis id="B89846436FD6EB483B53D58A4572818F" bold="true" pageId="0" pageNumber="31">Figure 5.</emphasis>
<taxonomicName id="AD8B70224E41B4FC01091CC3F2BFDA61" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="7CBCAB484AAF6ADB8E5EC3AAAC1E6BF2" italics="true" pageId="0" pageNumber="31">Bomarea pumila</emphasis>
</taxonomicName>
<emphasis id="6A9FF6B2E7E114C2F8960E5E4598B6A4" bold="true" pageId="0" pageNumber="31">A-D</emphasis>
field images from Peru 
<emphasis id="B883A6316C70E217A6786FACC824B510" bold="true" pageId="0" pageNumber="31">E</emphasis>
syntype (K! [K000400912]) of 
<taxonomicName id="BD45F243CD9DB43B91CEFA2E855780EE" class="Liliopsida" family="Alstroemeriaceae" genus="Bomarea" kingdom="Plantae" lsidName="Bomarea pumila" order="Liliales" pageId="0" pageNumber="31" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="55B6EEDA313EE43E0A498C3E883A1DDA" italics="true" pageId="0" pageNumber="31">Bomarea pumila</emphasis>
</taxonomicName>
of 
<emphasis id="34EED4B3B84A22E5B729581B4274B1E4" italics="true" pageId="0" pageNumber="31">W. Lechler 2240</emphasis>
from Sachapata, Peru. Photos 
<emphasis id="02284C470B5D6BB03AE150DC7D1EBC39" bold="true" pageId="0" pageNumber="31">A-D</emphasis>
from Lucely L. Vilca Bustamente 
<emphasis id="B9D901FE580E8BA6AEA8C29144315403" bold="true" pageId="0" pageNumber="31">E</emphasis>
from The Herbarium Catalogue, Royal Botanic Gardens, Kew. The Board of Trustees of the RBG, Kew.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="75E1507A9ABFD07EE68606EDD604A2CC" pageId="0" pageNumber="31" type="materials_examined">
<paragraph id="7E2BCC58C4A0ED2B1E4E6EF421B336B2" pageId="0" pageNumber="31">Additional specimens examined.</paragraph>
<paragraph id="87D7E0EEB4ED3315841D1B4BF7419040" pageId="0" pageNumber="31">
<materialsCitation id="CD293D51225F9162B1FD50647A812CA8" collectingDate="2022-01-01" collectorName="N. Exe, M. Johnson, A. Fierro-Minda" country="Ecuador" county="Canton Banos" elevation="3489" latitude="-1.4775918" location="Rio Verde" longLatPrecision="1" longitude="-78.298225" municipality="Parroquia Rio Verde" specimenCount="1" stateProvince="Tungurahua">
<emphasis id="A270F1D439F77A181F9F5A6BF9C37917" bold="true" pageId="0" pageNumber="31">
<collectingCountry id="DD865C187FD6CAC833A97548AD0EBF53" name="Ecuador">Ecuador</collectingCountry>
. 
<collectingRegion id="53C7E28D173DAD163CD6E38A2EF344A5" country="Ecuador" name="Tungurahua">Tungurahua</collectingRegion>
</emphasis>
: 
<collectingCounty id="F3773B7023F7B29133EFFA1322DAC222">
<normalizedToken id="E28A5D2C68CBE57A94210BF633BC7A90" originalValue="Cantón">Canton</normalizedToken>
<normalizedToken id="25BEFCAC0F29FC599A655E2ED9816BDD" originalValue="Baños">Banos</normalizedToken>
</collectingCounty>
, 
<collectingMunicipality id="8D845C27E91C0F5CDFDF68D19F13227F">
Parroquia 
<location id="5502BCB0AAC4C8294AAEE184E1F69721" LSID="urn:lsid:plazi:treatment:79BFC9D3526F5AEAB9A17253EDCDC6F3:5502BCB0AAC4C8294AAEE184E1F69721" country="Ecuador" county="Canton Banos" latitude="-1.4775918" longLatPrecision="1" longitude="-78.298225" municipality="Parroquia Rio Verde" name="Rio Verde" stateProvince="Tungurahua">
<normalizedToken id="5B9BFB01E10D3C16B2D8288F3DE7E3C7" originalValue="Río">Rio</normalizedToken>
Verde
</location>
</collectingMunicipality>
, 
<location id="36B547C06DC92F7DF50B3954F58BAE71" LSID="urn:lsid:plazi:treatment:79BFC9D3526F5AEAB9A17253EDCDC6F3:36B547C06DC92F7DF50B3954F58BAE71" country="Ecuador" county="Canton Banos" latitude="-1.4775918" longLatPrecision="1" longitude="-78.298225" municipality="Parroquia Rio Verde" name="Bosque Protector Cerro Candelaria" stateProvince="Tungurahua">Bosque Protector Cerro Candelaria</location>
(
<normalizedToken id="EBADC2CC6C9D23C68483B04842D18775" originalValue="Fundación">Fundacion</normalizedToken>
EcoMinga), upper 
<collectingRegion id="E982C646FFD2030D3C675AC6E981984B" country="Ecuador" name="Pastaza">Pastaza</collectingRegion>
watershed, 
<location id="F2198217DCC70C139A6FBA8585E2121A" LSID="urn:lsid:plazi:treatment:79BFC9D3526F5AEAB9A17253EDCDC6F3:F2198217DCC70C139A6FBA8585E2121A" country="Ecuador" county="Canton Banos" latitude="-1.4775918" longLatPrecision="1" longitude="-78.298225" municipality="Parroquia Rio Verde" name="Cerro Candelaria" stateProvince="Tungurahua">Cerro Candelaria</location>
summit trail, just below paramo, 
<geoCoordinate id="44665C72E783B2A58540A53D5073EA1F" degrees="1" direction="south" minutes="28" orientation="latitude" precision="1" seconds="39.33" value="-1.4775918">1°28'39.33&quot;S</geoCoordinate>
, 
<geoCoordinate id="4C90F494615E60A83928C9E38645AC87" degrees="78" direction="west" minutes="17" orientation="longitude" precision="1" seconds="53.61" value="-78.298225">78°17'53.61&quot;W</geoCoordinate>
, 
<quantity id="1120CAE81E9CA557B2210C6FF124640F" metricMagnitude="3" metricUnit="m" metricValue="3.4885" metricValueMax="3.8269999999999995" metricValueMin="3.15" unit="m" value="3488.5" valueMax="3827.0" valueMin="3150.0">
<elevation id="0D8FAD618DCD4A88AEC6DB6109D1DC04" metricMagnitude="3" metricUnit="m" metricValue="3.4885" metricValueMax="3.8269999999999995" metricValueMin="3.15" unit="m" value="3488.5" valueMax="3827.0" valueMin="3150.0">3150-3827 m</elevation>
</quantity>
, 
<collectingDate id="8A00AF998AFE87DBDA70681577DD7706" value="2022-01-01">1 Jan 2022</collectingDate>
, 
<emphasis id="F99E41125D9724F88744814A6F61C8E3" italics="true" pageId="0" pageNumber="31">
<collectorName id="E0CDFCDF328066464A6227EE169712A8">N. Exe</collectorName>
, 
<collectorName id="A0BE58FD352D55F7167F8552BD75F678">M. Johnson</collectorName>
&amp; 
<collectorName id="DABBC78E9C104D5590EF933C9C63C7D5">A. Fierro-Minda</collectorName>
2
</emphasis>
(GUAY)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>