318 lines
36 KiB
XML
318 lines
36 KiB
XML
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<mods:title id="0B2C48BCA0B3EFEC29481AC4B0DFE0D9">Taxonomy of Guatemalan water beetles in the genus Hydrochus (Coleoptera: Hydrochidae)</mods:title>
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<mods:title id="483827737C6617F07CF1D9022A36CD2A">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="DD04DE9A2577A96549128811F57CF700">2020</mods:date>
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<treatment id="060AF5509B263708FE971689FE95FC1D" ID-DOI="http://doi.org/10.5281/zenodo.3883395" ID-GBIF-Taxon="164481678" ID-Zenodo-Dep="3883395" LSID="urn:lsid:plazi:treatment:060AF5509B263708FE971689FE95FC1D" httpUri="http://treatment.plazi.org/id/060AF5509B263708FE971689FE95FC1D" lastPageId="4" lastPageNumber="295" pageId="2" pageNumber="293">
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<subSubSection id="C6B917CD9B26370EFE971689FDE5FEEE" box="[284,626,237,262]" pageId="2" pageNumber="293" type="nomenclature">
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<paragraph id="8E1C44469B26370EFE971689FDE5FEEE" blockId="2.[151,626,237,322]" box="[284,626,237,262]" pageId="2" pageNumber="293">
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<heading id="D554F32A9B26370EFE971689FDE5FEEE" box="[284,626,237,262]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="293" reason="3">
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<taxonomicName id="49A33FC59B26370EFE971689FDE5FEEE" ID-CoL="3N673" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[284,626,237,262]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="debilis">
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<emphasis id="BCD798549B26370EFE971689FDE5FEEE" bold="true" box="[284,626,237,262]" pageId="2" pageNumber="293">
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<emphasis id="BCD798549B26370EFE971689FE74FEED" bold="true" box="[284,483,237,261]" italics="true" pageId="2" pageNumber="293">Hydrochus debilis</emphasis>
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||
<bibRefCitation id="EA3239B79B26370EFE61168BFDE5FEEE" author="SHARP D." box="[490,626,239,262]" pageId="2" pageNumber="293" pagination="1 - 144" refId="ref6023" refString="SHARP D. 1882: Insecta. Coleoptera. Vol. 1, part 2 (Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae, Heteroceridae, Parnidae, Georissidae, Cyathoceridae, Staphylinidae). Pp 1 - 144. In: GODMAN F. D. & SALVIN O.: Biologia Centrali-Americana. Volume 16. Taylor and Francis, London, xv + 824 pp." type="book chapter" year="1882">Sharp, 1882</bibRefCitation>
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||
</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C6B917CD9B26370EFE1A1769FE69FEC8" box="[401,510,269,288]" pageId="2" pageNumber="293" type="description">
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<paragraph id="8E1C44469B26370EFE1A1769FE69FEC8" blockId="2.[151,626,237,322]" box="[401,510,269,288]" pageId="2" pageNumber="293">
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(
|
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<figureCitation id="169858C39B26370EFE1C1769FE5AFEC8" box="[407,461,269,288]" captionStart="Fig" captionStartId="1.[150,181,2064,2083]" captionTargetBox="[151,1437,799,2034]" captionTargetId="figure@1.[151,1437,798,2034]" captionTargetPageId="1" captionText="Fig. 1. Dorsal habitus of Guatemalan Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3879762" httpUri="https://zenodo.org/record/3879762/files/figure.png" pageId="2" pageNumber="293">Figs 1</figureCitation>
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||
,
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<figureCitation id="169858C39B26370EFE5C1769FE74FEC8" box="[471,483,269,288]" captionStart="Fig" captionStartId="2.[151,182,2059,2078]" captionTargetBox="[204,1384,841,2039]" captionTargetId="figure@2.[204,1384,841,2039]" captionTargetPageId="2" captionText="Fig. 2. Hydrochus debilis Sharp, 1882, habitus and male genitalia of lectotype." figureDoi="http://doi.org/10.5281/zenodo.3879766" httpUri="https://zenodo.org/record/3879766/files/figure.png" pageId="2" pageNumber="293">2</figureCitation>
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,
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<figureCitation id="169858C39B26370EFE671769FE60FEC8" box="[492,503,269,288]" captionStart="Fig" captionStartId="9.[150,181,2064,2083]" captionTargetBox="[266,1320,1467,2045]" captionTargetId="figure@9.[266,1320,1467,2045]" captionTargetPageId="9" captionText="Fig. 8 Hydrochus lectotype labels.A – H. obscurus Sharp, 1882; B – H. debilis Sharp, 1882." figureDoi="http://doi.org/10.5281/zenodo.3879778" httpUri="https://zenodo.org/record/3879778/files/figure.png" pageId="2" pageNumber="293">8</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C6B917CD9B26370EFF1C174BFE51FEAA" box="[151,454,303,322]" pageId="2" pageNumber="293" type="reference_group">
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||
<paragraph id="8E1C44469B26370EFF1C174BFE51FEAA" blockId="2.[151,626,237,322]" box="[151,454,303,322]" pageId="2" pageNumber="293">
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<treatmentCitationGroup id="AEB363689B26370EFF1C174BFE51FEAA" box="[151,454,303,322]" pageId="2" pageNumber="293">
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<treatmentCitation id="0F0262579B26370EFF1C174BFE55FEAA" author="SHARP D." box="[151,450,303,322]" page="89" pageId="2" pageNumber="293" year="1882">
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<taxonomicName id="49A33FC59B26370EFF1C174BFE55FEAA" ID-CoL="3N673" authority="Sharp, 1882: 89" authorityName="Sharp" authorityPageNumber="89" authorityYear="1882" box="[151,450,303,322]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="debilis">
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<emphasis id="BCD798549B26370EFF1C174BFEA5FEAA" box="[151,306,303,322]" italics="true" pageId="2" pageNumber="293">Hydrochus debilis</emphasis>
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<bibRefCitation id="EA3239B79B26370EFEBC174BFE55FEAA" author="SHARP D." box="[311,450,303,322]" pageId="2" pageNumber="293" pagination="1 - 144" refId="ref6023" refString="SHARP D. 1882: Insecta. Coleoptera. Vol. 1, part 2 (Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae, Heteroceridae, Parnidae, Georissidae, Cyathoceridae, Staphylinidae). Pp 1 - 144. In: GODMAN F. D. & SALVIN O.: Biologia Centrali-Americana. Volume 16. Taylor and Francis, London, xv + 824 pp." type="book chapter" year="1882">Sharp, 1882: 89</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C6B917CD9B26370EFF1C173DFC55FDBD" pageId="2" pageNumber="293" type="materials_examined">
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<paragraph id="8E1C44469B26370EFF1C173DFE57FD9E" blockId="2.[151,761,345,764]" pageId="2" pageNumber="293">
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<emphasis id="BCD798549B26370EFF1C173DFEE0FE84" bold="true" box="[151,375,345,364]" pageId="2" pageNumber="293">Type material examined.</emphasis>
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LൾǤŦƟŦYPൾ (hereby designated,
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<figureCitation id="169858C39B26370EFD00173DFD29FE84" box="[651,702,345,364]" captionStart="Fig" captionStartId="9.[150,181,2064,2083]" captionTargetBox="[266,1320,1467,2045]" captionTargetId="figure@9.[266,1320,1467,2045]" captionTargetPageId="9" captionText="Fig. 8 Hydrochus lectotype labels.A – H. obscurus Sharp, 1882; B – H. debilis Sharp, 1882." figureDoi="http://doi.org/10.5281/zenodo.3879778" httpUri="https://zenodo.org/record/3879778/files/figure.png" pageId="2" pageNumber="293">Fig. 8</figureCitation>
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): male (
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<collectionCode id="E8B2DC839B26370EFF161717FF43FE6E" box="[157,212,371,390]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="2" pageNumber="293" type="Herbarium">NHM</collectionCode>
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): ‘
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<taxonomicName id="49A33FC59B26370EFF6D1717FE11FE6E" box="[230,390,371,390]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="debilis">Hydrochus debilis</taxonomicName>
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Type D. S.
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||
<collectingRegion id="4C678AA49B26370EFE651717FDD1FE6E" box="[494,582,371,390]" country="Nicaragua" name="Chontales" pageId="2" pageNumber="293">Chontales</collectingRegion>
|
||
.
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||
<collectingCountry id="F6B404D69B26370EFDC51717FD31FE6E" box="[590,678,371,390]" name="Nicaragua" pageId="2" pageNumber="293">Nicaragua</collectingCountry>
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Janson // [male symbol] //
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<typeStatus id="5118FAE49B26370EFEAC17EAFE36FE49" box="[295,417,398,417]" pageId="2" pageNumber="293" type="lectotype">LECTOTYPE</typeStatus>
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[blue disc] //
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||
<collectingRegion id="4C678AA49B26370EFD9F17EAFDFBFE49" box="[532,620,398,417]" country="Nicaragua" name="Chontales" pageId="2" pageNumber="293">Chontales</collectingRegion>
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||
,
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||
<collectingCountry id="F6B404D69B26370EFDF817EAFD5AFE49" box="[627,717,398,417]" name="Nicaragua" pageId="2" pageNumber="293">Nicaragua</collectingCountry>
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||
. Janson. // B.C.A. Col. I. 2.
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<taxonomicName id="49A33FC59B26370EFED017CDFDA0FE54" authority=", Sharp." authorityName="Sharp." box="[347,567,425,444]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="debilis">Hydrochus debilis, Sharp.</taxonomicName>
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// HABITUS / GENIT. DIGITAL IMAGES captured 2020 P.D. Perkins // [red]
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<typeStatus id="5118FAE49B26370EFDF517A7FD6FFE3E" box="[638,760,451,470]" pageId="2" pageNumber="293" type="lectotype">LECTOTYPE</typeStatus>
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||
<taxonomicName id="49A33FC59B26370EFF1C17BAFEFDFE19" authority="Sharp" authorityName="Sharp" box="[151,362,478,497]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="debilis">
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||
<emphasis id="BCD798549B26370EFF1C17BAFEA4FE19" box="[151,307,478,497]" italics="true" pageId="2" pageNumber="293">Hydrochus debilis</emphasis>
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Sharp
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</taxonomicName>
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desig. P. D. Perkinsʼ. PARAĿൾǤŦƟŦYPൾ: female (
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<collectionCode id="E8B2DC839B26370EFF16179DFF43FDE4" box="[157,212,505,524]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="2" pageNumber="293" type="Herbarium">NHM</collectionCode>
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): ‘
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<taxonomicName id="49A33FC59B26370EFF6E179DFE12FDE4" box="[229,389,505,524]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="debilis">Hydrochus debilis</taxonomicName>
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Type D. S.
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<collectingRegion id="4C678AA49B26370EFE62179DFDD6FDE4" box="[489,577,505,524]" country="Nicaragua" name="Chontales" pageId="2" pageNumber="293">Chontales</collectingRegion>
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||
.
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<collectingCountry id="F6B404D69B26370EFDC2179DFD36FDE4" box="[585,673,505,524]" name="Nicaragua" pageId="2" pageNumber="293">Nicaragua</collectingCountry>
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// [female symbol] //
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<typeStatus id="5118FAE49B26370EFF7A1477FE35FDCE" box="[241,418,531,550]" pageId="2" pageNumber="293" type="paralectotype">PARALECTOTYPE</typeStatus>
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[blue disc] //
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<collectingRegion id="4C678AA49B26370EFD9F1477FDFBFDCE" box="[532,620,531,550]" country="Nicaragua" name="Chontales" pageId="2" pageNumber="293">Chontales</collectingRegion>
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||
,
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<collectingCountry id="F6B404D69B26370EFDF81477FD5AFDCE" box="[627,717,531,550]" name="Nicaragua" pageId="2" pageNumber="293">Nicaragua</collectingCountry>
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||
. Janson. // B.C.A. Col. I. 2.
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||
<taxonomicName id="49A33FC59B26370EFED1144AFDA0FDA9" authority=", Sharp." authorityName="Sharp." box="[346,567,558,577]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="debilis">Hydrochus debilis, Sharp.</taxonomicName>
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||
// LECTOPARATYPE
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<taxonomicName id="49A33FC59B26370EFF1C142DFF70FDB4" box="[151,231,585,604]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="debilis">H.debilis</taxonomicName>
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by J. L. Hellman 75 // [yellow]
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||
<typeStatus id="5118FAE49B26370EFE61142DFD0FFDB4" box="[490,664,585,604]" pageId="2" pageNumber="293" type="paralectotype">PARALECTOTYPE</typeStatus>
|
||
<taxonomicName id="49A33FC59B26370EFD17142DFE90FD9E" authority="Sharp" authorityName="Sharp" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="debilis">
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||
<emphasis id="BCD798549B26370EFD17142DFF47FD9E" italics="true" pageId="2" pageNumber="293">Hydrochus debilis</emphasis>
|
||
Sharp
|
||
</taxonomicName>
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||
desig. P. D. Perkinsʼ.
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||
</paragraph>
|
||
<paragraph id="8E1C44469B26370EFF1C141AFD66FD14" blockId="2.[151,761,345,764]" pageId="2" pageNumber="293">
|
||
<emphasis id="BCD798549B26370EFF1C141AFE0DFD79" bold="true" box="[151,410,638,657]" pageId="2" pageNumber="293">Additonal material examined</emphasis>
|
||
(
|
||
<specimenCount id="98A58FCF9B26370EFE2F141AFE7CFD79" box="[420,491,638,657]" pageId="2" pageNumber="293" type="generic">15 spec.</specimenCount>
|
||
).
|
||
<emphasis id="BCD798549B26370EFE7C141AFD18FD79" bold="true" box="[503,655,638,657]" pageId="2" pageNumber="293">
|
||
<collectingCountry id="F6B404D69B26370EFE7C141AFD1CFD79" box="[503,651,638,657]" name="Guatemala" pageId="2" pageNumber="293">GUATEMALA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<collectingRegion id="4C678AA49B26370EFD19141AFD50FD79" box="[658,711,638,657]" country="Guatemala" name="Izabal" pageId="2" pageNumber="293">Izabal</collectingRegion>
|
||
,
|
||
<quantity id="495BE9A39B26370EFD45141AFD6EFD79" box="[718,761,638,657]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="2" pageNumber="293" unit="mi" value="1.0">1 mi.</quantity>
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||
N. Morales.Km 239 Atlantic Hwy,
|
||
<date id="FA1D62869B26370EFE3514FDFDD1FD44" box="[446,582,665,684]" pageId="2" pageNumber="293" value="1965-08-16" valueMax="1965-08-18" valueMin="1965-08-16">16–18.viii.1965</date>
|
||
, leg. Paul J. Spangler (1:
|
||
<collectionCode id="E8B2DC839B26370EFF3914D7FF61FD2E" box="[178,246,691,710]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="293" type="Museum">NMNH</collectionCode>
|
||
);
|
||
<quantity id="495BE9A39B26370EFE8F14D7FEAEFD2E" box="[260,313,691,710]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="2" pageNumber="293" unit="mi" value="20.0">20 mi.</quantity>
|
||
SW Puerto Barrios,
|
||
<date id="FA1D62869B26370EFE6A14D7FDDFFD2E" box="[481,584,691,710]" pageId="2" pageNumber="293" value="1965-08-16">16.viii.1965</date>
|
||
, leg.Paul J. Spangler (4:
|
||
<collectionCode id="E8B2DC839B26370EFF3914AAFF61FD09" box="[178,246,718,737]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="293" type="Museum">NMNH</collectionCode>
|
||
);
|
||
<quantity id="495BE9A39B26370EFE8F14AAFEAEFD09" box="[260,313,718,737]" metricMagnitude="4" metricUnit="m" metricValue="4.5061632" pageId="2" pageNumber="293" unit="mi" value="28.0">28 mi.</quantity>
|
||
SW Puerto Barrios,
|
||
<date id="FA1D62869B26370EFE6A14AAFDDFFD09" box="[481,584,718,737]" pageId="2" pageNumber="293" value="1965-08-16">16.viii.1965</date>
|
||
, leg.Paul J. Spangler (1:
|
||
<collectionCode id="E8B2DC839B26370EFF3F148DFF6EFD14" box="[180,249,745,764]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="293" type="Museum">NMNH</collectionCode>
|
||
); Rio Dulce Fy.,
|
||
<date id="FA1D62869B26370EFE02148DFE75FD14" box="[393,482,745,764]" pageId="2" pageNumber="293" value="1974-07-02">2.vii.1974</date>
|
||
, leg. W. E. Steiner (9:
|
||
<collectionCode id="E8B2DC839B26370EFD29148DFD70FD14" box="[674,743,745,764]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="293" type="Museum">NMNH</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8E1C44469B26370EFCB716DAFC55FDBD" blockId="2.[828,1437,190,757]" pageId="2" pageNumber="293">
|
||
<materialsCitation id="3ECB4E1B9B26370EFCB716DAFC55FDBD" ID-GBIF-Occurrence="2632269302" collectorName="J. L. Hellman & This" country="Mexico" location="Mexico" pageId="2" pageNumber="293" specimenCount="8" typeStatus="lectotype">
|
||
<emphasis id="BCD798549B26370EFCB716DAFBA9FF3D" bold="true" box="[828,1086,190,213]" pageId="2" pageNumber="293">
|
||
<typeStatus id="5118FAE49B26370EFCB716DAFC3AFF3D" box="[828,941,190,213]" pageId="2" pageNumber="293" type="lectotype">Lectotype</typeStatus>
|
||
designation.
|
||
</emphasis>
|
||
The NHM kindly provided two
|
||
<typeStatus id="5118FAE49B26370EFCB716BAFC0EFF1D" box="[828,921,222,245]" pageId="2" pageNumber="293" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="49A33FC59B26370EFC3216BAFBEFFF1D" box="[953,1144,222,245]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="BCD798549B26370EFC3216BAFBEFFF1D" box="[953,1144,222,245]" italics="true" pageId="2" pageNumber="293">Hydrochus debilis</emphasis>
|
||
</taxonomicName>
|
||
, which are part of the Sharp
|
||
<emphasis id="BCD798549B26370EFCB7169AFBE3FEFD" box="[828,1140,254,277]" italics="true" pageId="2" pageNumber="293">Biologia Centrali-Americana</emphasis>
|
||
types collection. The
|
||
<typeStatus id="5118FAE49B26370EFAD1169AFCF7FEDD" pageId="2" pageNumber="293" type="syntype">syntypes</typeStatus>
|
||
consist of one male and one female. I have selected the male to serve as
|
||
<typeStatus id="5118FAE49B26370EFB93175AFBEDFEBD" box="[1048,1146,318,341]" pageId="2" pageNumber="293" type="lectotype">lectotype</typeStatus>
|
||
and labeled it as such, and provide images of the habitus, male genitalia, and labels of that specimen (
|
||
<figureCitation id="169858C39B26370EFC7D171AFBA0FE7D" box="[1014,1079,382,405]" captionStart="Fig" captionStartId="1.[150,181,2064,2083]" captionTargetBox="[151,1437,799,2034]" captionTargetId="figure@1.[151,1437,798,2034]" captionTargetPageId="1" captionText="Fig. 1. Dorsal habitus of Guatemalan Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3879762" httpUri="https://zenodo.org/record/3879762/files/figure.png" pageId="2" pageNumber="293">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="169858C39B26370EFBC8171AFBC5FE7D" box="[1091,1106,382,405]" captionStart="Fig" captionStartId="2.[151,182,2059,2078]" captionTargetBox="[204,1384,841,2039]" captionTargetId="figure@2.[204,1384,841,2039]" captionTargetPageId="2" captionText="Fig. 2. Hydrochus debilis Sharp, 1882, habitus and male genitalia of lectotype." figureDoi="http://doi.org/10.5281/zenodo.3879766" httpUri="https://zenodo.org/record/3879766/files/figure.png" pageId="2" pageNumber="293">2</figureCitation>
|
||
,
|
||
<figureCitation id="169858C39B26370EFBD6171AFBFBFE7D" box="[1117,1132,382,405]" captionStart="Fig" captionStartId="9.[150,181,2064,2083]" captionTargetBox="[266,1320,1467,2045]" captionTargetId="figure@9.[266,1320,1467,2045]" captionTargetPageId="9" captionText="Fig. 8 Hydrochus lectotype labels.A – H. obscurus Sharp, 1882; B – H. debilis Sharp, 1882." figureDoi="http://doi.org/10.5281/zenodo.3879778" httpUri="https://zenodo.org/record/3879778/files/figure.png" pageId="2" pageNumber="293">8</figureCitation>
|
||
). The other
|
||
<typeStatus id="5118FAE49B26370EFB62171AFAABFE7D" box="[1257,1340,382,405]" pageId="2" pageNumber="293" type="syntype">syntype</typeStatus>
|
||
has been labeled
|
||
<typeStatus id="5118FAE49B26370EFC1B17FAFBB4FE5D" box="[912,1059,414,437]" pageId="2" pageNumber="293" type="paralectotype">paralectotype</typeStatus>
|
||
. One additional specimen provided by the NHM as ‘Possible typeʼ is a different species, from
|
||
<collectingCountry id="F6B404D69B26370EFCB717BAFC06FE1D" box="[828,913,478,501]" name="Mexico" pageId="2" pageNumber="293">Mexico</collectingCountry>
|
||
. One of the
|
||
<typeStatus id="5118FAE49B26370EFB9617BAFBEDFE1D" box="[1053,1146,478,501]" pageId="2" pageNumber="293" type="syntype">syntypes</typeStatus>
|
||
bears a
|
||
<typeStatus id="5118FAE49B26370EFB5D17BAFAAFFE1D" box="[1238,1336,478,501]" pageId="2" pageNumber="293" type="lectotype">lectotype</typeStatus>
|
||
label, by
|
||
<collectorName id="235621909B26370EFCB7179AFC45FDFD" box="[828,978,510,533]" pageId="2" pageNumber="293">J. L. Hellman</collectorName>
|
||
, but this designation was never published.
|
||
<collectorName id="235621909B26370EFCB7147AFCFCFDDD" box="[828,875,542,565]" pageId="2" pageNumber="293">This</collectorName>
|
||
label has been reversed and “unpub.” written on the reverse side.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C6B917CD9B26370FFCB7143AFE2AFEDD" lastPageId="3" lastPageNumber="294" pageId="2" pageNumber="293" type="diagnosis">
|
||
<paragraph id="8E1C44469B26370FFCB7143AFE2AFEDD" blockId="2.[828,1437,190,757]" lastBlockId="3.[151,760,188,565]" lastPageId="3" lastPageNumber="294" pageId="2" pageNumber="293">
|
||
<emphasis id="BCD798549B26370EFCB7143AFBDFFD9D" bold="true" box="[828,1096,606,629]" pageId="2" pageNumber="293">Differential diagnosis.</emphasis>
|
||
Among Guatemalan species,
|
||
<taxonomicName id="49A33FC59B26370EFCB7141BFC32FD7D" box="[828,933,638,662]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="BCD798549B26370EFCB7141BFC32FD7D" box="[828,933,638,662]" italics="true" pageId="2" pageNumber="293">H. debilis</emphasis>
|
||
</taxonomicName>
|
||
is recognized by the combination of moderate size (ca.
|
||
<quantity id="495BE9A39B26370EFC1014FAFBABFD5D" box="[923,1084,670,693]" metricMagnitude="-3" metricUnit="m" metricValue="2.885" metricValueMax="3.08" metricValueMin="2.69" pageId="2" pageNumber="293" unit="mm" value="2.885" valueMax="3.08" valueMin="2.69">2.69–3.08 mm</quantity>
|
||
), the densely punctate pronotum with deep foveae, and the elytral sculpture, each elytron having four raised interstriae: 3
|
||
<superScript id="79D6E90E9B26370EFB0E14B8FB05FD02" attach="left" box="[1157,1170,732,746]" fontSize="6" pageId="2" pageNumber="293">rd</superScript>
|
||
slightly raised over basal ¼, 4
|
||
<superScript id="79D6E90E9B27370FFF4C16D8FF44FF22" attach="left" box="[199,211,188,202]" fontSize="6" pageId="3" pageNumber="294">th</superScript>
|
||
raised in area anterior to large callus of 5
|
||
<superScript id="79D6E90E9B27370FFD1C16D8FD34FF22" attach="none" box="[663,675,188,202]" fontSize="6" pageId="3" pageNumber="294">th</superScript>
|
||
, 5
|
||
<superScript id="79D6E90E9B27370FFD3416D8FD5CFF22" attach="left" box="[703,715,188,202]" fontSize="6" pageId="3" pageNumber="294">th</superScript>
|
||
raised from about anterior ¼ to midlength, and usual callus of 5
|
||
<superScript id="79D6E90E9B27370FFF4F1698FF47FEE2" attach="none" box="[196,208,252,266]" fontSize="6" pageId="3" pageNumber="294">th</superScript>
|
||
. The distinctive male genitalia (
|
||
<figureCitation id="169858C39B27370FFDB7169AFD16FEFD" box="[572,641,254,277]" captionStart="Fig" captionStartId="2.[151,182,2059,2078]" captionTargetBox="[204,1384,841,2039]" captionTargetId="figure@2.[204,1384,841,2039]" captionTargetPageId="2" captionText="Fig. 2. Hydrochus debilis Sharp, 1882, habitus and male genitalia of lectotype." figureDoi="http://doi.org/10.5281/zenodo.3879766" httpUri="https://zenodo.org/record/3879766/files/figure.png" pageId="3" pageNumber="294">Fig. 2</figureCitation>
|
||
; described below) are also diagnostic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DADC14CE9B26370EFF1C1E6FFCA0F7F6" ID-DOI="http://doi.org/10.5281/zenodo.3879766" ID-Zenodo-Dep="3879766" box="[151,823,2059,2078]" httpUri="https://zenodo.org/record/3879766/files/figure.png" pageId="2" pageNumber="293" startId="2.[151,182,2059,2078]" targetBox="[204,1384,841,2039]" targetPageId="2">
|
||
<paragraph id="8E1C44469B26370EFF1C1E6FFCA0F7F6" blockId="2.[151,823,2059,2078]" box="[151,823,2059,2078]" pageId="2" pageNumber="293">
|
||
Fig. 2.
|
||
<taxonomicName id="49A33FC59B26370EFF5F1E6FFE4AF7F6" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[212,477,2059,2078]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="BCD798549B26370EFF5F1E6FFEF8F7F6" box="[212,367,2059,2078]" italics="true" pageId="2" pageNumber="293">Hydrochus debilis</emphasis>
|
||
<bibRefCitation id="EA3239B79B26370EFEFF1E6FFE4AF7F6" author="SHARP D." box="[372,477,2059,2078]" pageId="2" pageNumber="293" pagination="1 - 144" refId="ref6023" refString="SHARP D. 1882: Insecta. Coleoptera. Vol. 1, part 2 (Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae, Heteroceridae, Parnidae, Georissidae, Cyathoceridae, Staphylinidae). Pp 1 - 144. In: GODMAN F. D. & SALVIN O.: Biologia Centrali-Americana. Volume 16. Taylor and Francis, London, xv + 824 pp." type="book chapter" year="1882">Sharp, 1882</bibRefCitation>
|
||
</taxonomicName>
|
||
, habitus and male genitalia of lectotype.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C6B917CD9B273708FF1C175AFE95FC1D" lastPageId="4" lastPageNumber="295" pageId="3" pageNumber="294" type="description">
|
||
<paragraph id="8E1C44469B27370FFF1C175AFB04FEDD" blockId="3.[151,760,188,565]" lastBlockId="3.[828,1437,190,565]" pageId="3" pageNumber="294">
|
||
<emphasis id="BCD798549B27370FFF1C175AFEA9FEBD" bold="true" box="[151,318,318,341]" pageId="3" pageNumber="294">Redescription.</emphasis>
|
||
Habitus as illustrated (
|
||
<figureCitation id="169858C39B27370FFDCF175AFD1DFEBD" box="[580,650,318,341]" captionStart="Fig" captionStartId="2.[151,182,2059,2078]" captionTargetBox="[204,1384,841,2039]" captionTargetId="figure@2.[204,1384,841,2039]" captionTargetPageId="2" captionText="Fig. 2. Hydrochus debilis Sharp, 1882, habitus and male genitalia of lectotype." figureDoi="http://doi.org/10.5281/zenodo.3879766" httpUri="https://zenodo.org/record/3879766/files/figure.png" pageId="3" pageNumber="294">Fig. 2</figureCitation>
|
||
). Size (in mm):
|
||
<typeStatus id="5118FAE49B27370FFF51173AFEDEFE9D" box="[218,329,350,373]" pageId="3" pageNumber="294" type="lectotype">Lectotype</typeStatus>
|
||
: body (length to elytral apices / width) 2.88 / 1.04; head width 0.69; pronotum length / width 0.75 / 0.71; PA 0.67; PB 0.56; elytra length / width 1.85 / 1.04. Body size moderate (ca.
|
||
<quantity id="495BE9A39B27370FFE1F17DAFDBDFE3D" box="[404,554,446,469]" metricMagnitude="-3" metricUnit="m" metricValue="2.885" metricValueMax="3.08" metricValueMin="2.69" pageId="3" pageNumber="294" unit="mm" value="2.885" valueMax="3.08" valueMin="2.69">2.69–3.08 mm</quantity>
|
||
in Guatemalan specimens). Dorsum dark grey to black, with very slight if any iridescence, elytra with black spots. Legs dark brown, with tibiofemoral joints slightly darker. Punctation on elytra ca. 2–4× that of pronotum. Elytra interstriae ca. 0.5–1.0× strial puncture diameter. Interstria 9
|
||
<superScript id="79D6E90E9B27370FFB4816B8FB58FF02" attach="left" box="[1219,1231,220,234]" fontSize="6" pageId="3" pageNumber="294">th</superScript>
|
||
more convex than others, overhanging 10
|
||
<superScript id="79D6E90E9B27370FFBBF1698FBD7FEE2" attach="left" box="[1076,1088,252,266]" fontSize="6" pageId="3" pageNumber="294">th</superScript>
|
||
interstria. Callus on 5
|
||
<superScript id="79D6E90E9B27370FFAB81698FAA8FEE2" attach="left" box="[1331,1343,252,266]" fontSize="6" pageId="3" pageNumber="294">th</superScript>
|
||
in usual area, large, very strongly raised.
|
||
</paragraph>
|
||
<paragraph id="8E1C44469B27370FFCD1175AFC37FE3D" blockId="3.[828,1437,190,565]" pageId="3" pageNumber="294">Pronotum length ca. equal width, widest at ca. anterior 1/3, narrowed at base, sides sinuate, minutely denticulate; anterior margin slightly bisinuate; depressions deep, reliefs sharply raised, depressions and reliefs densely punctate.</paragraph>
|
||
<paragraph id="8E1C44469B273708FCD117BAFEEEFEBD" blockId="3.[828,1437,190,565]" lastBlockId="4.[151,760,188,1013]" lastPageId="4" lastPageNumber="295" pageId="3" pageNumber="294">
|
||
Elytra with spaces between strial punctures narrow ridges to ca. 0.5× their diameter; 3
|
||
<superScript id="79D6E90E9B27370FFB061798FB0DFDE2" attach="left" box="[1165,1178,508,522]" fontSize="6" pageId="3" pageNumber="294">rd</superScript>
|
||
interstria slightly raised over basal fourth; 4
|
||
<superScript id="79D6E90E9B27370FFB981478FB88FDC2" attach="left" box="[1043,1055,540,554]" fontSize="6" pageId="3" pageNumber="294">th</superScript>
|
||
interstria raised in area anterior to callus of 5
|
||
<superScript id="79D6E90E9B203708FE8F16D8FE87FF22" attach="right" box="[260,272,188,202]" fontSize="6" pageId="4" pageNumber="295">th</superScript>
|
||
; 5
|
||
<superScript id="79D6E90E9B203708FEA116D8FEA1FF22" attach="left" box="[298,310,188,202]" fontSize="6" pageId="4" pageNumber="295">th</superScript>
|
||
interstria raised from about anterior fourth to midlength, raised area separated from callus of 5
|
||
<superScript id="79D6E90E9B203708FD4316B8FD43FF02" attach="left" box="[712,724,220,234]" fontSize="6" pageId="4" pageNumber="295">th</superScript>
|
||
by very slight depression; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with moderate angulation.
|
||
</paragraph>
|
||
<caption id="DADC14CE9B27370FFF1C1E6DFC8EF7F4" ID-DOI="http://doi.org/10.5281/zenodo.3879768" ID-Zenodo-Dep="3879768" box="[151,793,2057,2076]" httpUri="https://zenodo.org/record/3879768/files/figure.png" pageId="3" pageNumber="294" startId="3.[151,182,2057,2076]" targetBox="[220,1373,626,2023]" targetPageId="3">
|
||
<paragraph id="8E1C44469B27370FFF1C1E6DFC8EF7F4" blockId="3.[151,793,2057,2076]" box="[151,793,2057,2076]" pageId="3" pageNumber="294">
|
||
Fig. 3.
|
||
<taxonomicName id="49A33FC59B27370FFF5F1E6DFEEDF7F4" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[212,378,2057,2076]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="directus" status="sp. nov.">
|
||
<emphasis id="BCD798549B27370FFF5F1E6DFEEDF7F4" box="[212,378,2057,2076]" italics="true" pageId="3" pageNumber="294">Hydrochus directus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A7E4252F9B27370FFEF41E6DFE53F7F4" box="[383,452,2057,2076]" pageId="3" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, habitus and male genitalia of holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8E1C44469B203708FF3E173AFE17FCDD" blockId="4.[151,760,188,1013]" pageId="4" pageNumber="295">
|
||
Male genitalia (
|
||
<figureCitation id="169858C39B203708FEEA173AFE33FE9D" box="[353,420,350,373]" captionStart="Fig" captionStartId="2.[151,182,2059,2078]" captionTargetBox="[204,1384,841,2039]" captionTargetId="figure@2.[204,1384,841,2039]" captionTargetPageId="2" captionText="Fig. 2. Hydrochus debilis Sharp, 1882, habitus and male genitalia of lectotype." figureDoi="http://doi.org/10.5281/zenodo.3879766" httpUri="https://zenodo.org/record/3879766/files/figure.png" pageId="4" pageNumber="295">Fig. 2</figureCitation>
|
||
). General characters: paramere tips moderately large, extending beyond distal tip of aedeagus, sagittate, distal ends pointed, lateral margins between
|
||
<emphasis id="BCD798549B203708FF7217DAFEB0FE3D" box="[249,295,446,469]" italics="true" pageId="4" pageNumber="295">plsa</emphasis>
|
||
and distal end slightly emarginate, medial margins of tips very slightly arcuate toward midline and well separated from lateral margins of aedeagus; parameres in dorsal/ventral views constricted at base of tips and dorsally with low ridge; parameres in dorsal/ventral views much wider at widest point than tips; parameres in lateral view wide, sinuate, and tapering to sharp tips; aedeagus greatest width at
|
||
<emphasis id="BCD798549B203708FE3714FAFE7BFD5D" box="[444,492,670,693]" italics="true" pageId="4" pageNumber="295">adbl</emphasis>
|
||
, shaft (portion of aedeagus distal to
|
||
<emphasis id="BCD798549B203708FEAC14DAFEC0FD3D" box="[295,343,702,725]" italics="true" pageId="4" pageNumber="295">adbl</emphasis>
|
||
) in dorsal/ventral views very narrow, almost parallel-sided; aedeagus dorsal margin sinuate in lateral view; basal piece shorter than parameres, with rim of orifice sclerotized.
|
||
</paragraph>
|
||
<paragraph id="8E1C44469B203708FF3E155AFD71FC5D" blockId="4.[151,760,188,1013]" pageId="4" pageNumber="295">
|
||
Dorsal surface:
|
||
<emphasis id="BCD798549B203708FEEB155AFE1EFCBD" box="[352,393,830,853]" italics="true" pageId="4" pageNumber="295">adtl</emphasis>
|
||
located at about distal fifth of aedeagus;
|
||
<emphasis id="BCD798549B203708FF7B153AFEA6FC9D" box="[240,305,862,885]" italics="true" pageId="4" pageNumber="295">pdmm</emphasis>
|
||
very slightly narrowly overlapping
|
||
<emphasis id="BCD798549B203708FD3E153AFD4AFC9D" box="[693,733,862,885]" italics="true" pageId="4" pageNumber="295">alm</emphasis>
|
||
at about midlength of aedeagus;
|
||
<emphasis id="BCD798549B203708FE59151AFD96FC7D" box="[466,513,894,917]" italics="true" pageId="4" pageNumber="295">adbl</emphasis>
|
||
very large, much wider than paramere area lateral to
|
||
<emphasis id="BCD798549B203708FE4615FAFE6BFC5D" box="[461,508,926,949]" italics="true" pageId="4" pageNumber="295">adbl</emphasis>
|
||
, distinctively shaped.
|
||
</paragraph>
|
||
<paragraph id="8E1C44469B203708FF3E15DAFE95FC1D" blockId="4.[151,760,188,1013]" pageId="4" pageNumber="295">
|
||
Ventral surface:
|
||
<emphasis id="BCD798549B203708FEEC15DBFE30FC3E" box="[359,423,959,982]" italics="true" pageId="4" pageNumber="295">pvmm</emphasis>
|
||
overlapping
|
||
<emphasis id="BCD798549B203708FDBD15DAFDC9FC3D" box="[566,606,958,981]" italics="true" pageId="4" pageNumber="295">alm</emphasis>
|
||
at base;
|
||
<emphasis id="BCD798549B203708FD4A15DBFD6FFC3E" box="[705,760,959,982]" italics="true" pageId="4" pageNumber="295">pwsa</emphasis>
|
||
near base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |