treatments-xml/data/06/0A/F5/060AF5509B20370AFC3815B9FF69FDBD.xml
2024-06-21 12:28:37 +02:00

304 lines
34 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="3D91DD4E27598198413B83C93CB1B320" ID-CLB-Dataset="21739" ID-DOI="10.37520/aemnp.2020.016" ID-GBIF-Dataset="6f4e1328-0bba-4f1c-aa66-4c1fe1fda040" ID-ISSN="1804-6487" ID-Zenodo-Dep="3879760" ID-ZooBank="45563A23-8582-4C53-81F3-591948E7A74D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591387310617" checkinUser="tatiana" docAuthor="Perkins, Philip D." docDate="2020" docId="060AF5509B20370AFC3815B9FF69FDBD" docLanguage="en" docName="ActaEntMusNatPra.60.1.291-301.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 60 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Hydrochus obscurus Sharp 1882" docType="treatment" docVersion="6" lastPageNumber="297" masterDocId="FA338D289B24370CFF8B1664FF97FFE8" masterDocTitle="Taxonomy of Guatemalan water beetles in the genus Hydrochus (Coleoptera: Hydrochidae)" masterLastPageNumber="301" masterPageNumber="291" pageNumber="295" updateTime="1698842118860" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
<mods:mods id="F0BF306081AFA206CD7F5FB7A1AA9696" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B93AE3D8EB440007E5CBAB8313EDDC52">
<mods:title id="7B4354E9F68E105173B8004611896D88">Taxonomy of Guatemalan water beetles in the genus Hydrochus (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
<mods:name id="2F82D8CF02613622FBCF3E99A8D03B6D" type="personal">
<mods:role id="C08DE0B139387FF78C258DB56C1D20A6">
<mods:roleTerm id="F2F411844F893F9DA162499E8441954B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F0663E9CF5CAFF425959B94745E9FE4E">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D5BA8C0A2BEE35484598DFD13B6B052B">text</mods:typeOfResource>
<mods:relatedItem id="12D5DBF1C589C0F953A23E3D89AF510C" type="host">
<mods:titleInfo id="82CB106376E9AA658BB5E540F43B2EE5">
<mods:title id="E1C0997CB4BDD7FEDA53C90F8990322A">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="AD423DCF892D6CC7B3DFE34E2E5E9C17">
<mods:date id="8B440951426D4A252518517FD33B63C4">2020</mods:date>
<mods:detail id="0AEAD54E8840C0A9CE3F48971DE40D61" type="pubDate">
<mods:number id="F1BCEA38F6E1D3595A2B6895B4361B67">2020-04-23</mods:number>
</mods:detail>
<mods:detail id="4F07D233D75191593D9BB542246AC89D" type="volume">
<mods:number id="3E7A91BEEE04F1FDEE8A3B7E9B36E62A">60</mods:number>
</mods:detail>
<mods:detail id="9A4421E9A25E4F984E5E34B035E42D23" type="issue">
<mods:number id="DC3C35E1F2911597B51D4C4FC38C0F50">1</mods:number>
</mods:detail>
<mods:extent id="14E40BD4C0B076C6E71B859135EF170A" unit="page">
<mods:start id="2E4C476A071C08BC41C921AED7AA3272">291</mods:start>
<mods:end id="A81D9A5D6BA5F1D4977FD15D9AC1DA6F">301</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="995D4DE879E7EF3F85866E5A77B5E221">journal article</mods:classification>
<mods:identifier id="0BAD961B86D9D8F48AB57432E2A5DFC1" type="CLB-Dataset">21739</mods:identifier>
<mods:identifier id="B313F64AE08F71A8C859719CD1318C19" type="DOI">10.37520/aemnp.2020.016</mods:identifier>
<mods:identifier id="9C9AE4227699410D2247B616A9142A12" type="GBIF-Dataset">6f4e1328-0bba-4f1c-aa66-4c1fe1fda040</mods:identifier>
<mods:identifier id="AD4645249891EEED664E9199B4BE03F3" type="ISSN">1804-6487</mods:identifier>
<mods:identifier id="B236587DC1B115FB3E68969768565068" type="Zenodo-Dep">3879760</mods:identifier>
<mods:identifier id="51F51D95A999CD69F45E708BD6EB5D57" type="ZooBank">45563A23-8582-4C53-81F3-591948E7A74D</mods:identifier>
</mods:mods>
<treatment id="060AF5509B20370AFC3815B9FF69FDBD" ID-DOI="http://doi.org/10.5281/zenodo.3883401" ID-GBIF-Taxon="164481680" ID-Zenodo-Dep="3883401" LSID="urn:lsid:plazi:treatment:060AF5509B20370AFC3815B9FF69FDBD" httpUri="http://treatment.plazi.org/id/060AF5509B20370AFC3815B9FF69FDBD" lastPageId="6" lastPageNumber="297" pageId="4" pageNumber="295">
<subSubSection id="C6B917CD9B203708FC3815B9FAB2FC1D" box="[947,1317,989,1013]" pageId="4" pageNumber="295" type="nomenclature">
<paragraph id="8E1C44469B203708FC3815B9FAB2FC1D" blockId="4.[828,1436,989,1127]" box="[947,1317,989,1013]" pageId="4" pageNumber="295">
<heading id="D554F32A9B203708FC3815B9FAB2FC1D" box="[947,1317,989,1013]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="295" reason="3">
<taxonomicName id="49A33FC59B203708FC3815B9FAB2FC1D" ID-CoL="6MK96" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[947,1317,989,1013]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="295" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="BCD798549B203708FC3815B9FAB2FC1D" bold="true" box="[947,1317,989,1013]" pageId="4" pageNumber="295">
<emphasis id="BCD798549B203708FC3815B9FB01FC1D" bold="true" box="[947,1174,989,1013]" italics="true" pageId="4" pageNumber="295">Hydrochus obscurus</emphasis>
<bibRefCitation id="EA3239B79B203708FB1615BAFAB2FC1D" author="SHARP D." box="[1181,1317,990,1013]" pageId="4" pageNumber="295" pagination="1 - 144" refId="ref6023" refString="SHARP D. 1882: Insecta. Coleoptera. Vol. 1, part 2 (Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae, Heteroceridae, Parnidae, Georissidae, Cyathoceridae, Staphylinidae). Pp 1 - 144. In: GODMAN F. D. &amp; SALVIN O.: Biologia Centrali-Americana. Volume 16. Taylor and Francis, London, xv + 824 pp." type="book chapter" year="1882">Sharp, 1882</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C6B917CD9B203708FBBE1598FB34FBE7" box="[1077,1187,1020,1039]" pageId="4" pageNumber="295" type="description">
<paragraph id="8E1C44469B203708FBBE1598FB34FBE7" blockId="4.[828,1436,989,1127]" box="[1077,1187,1020,1039]" pageId="4" pageNumber="295">
(
<figureCitation id="169858C39B203708FBB01598FBE6FBE7" box="[1083,1137,1020,1039]" captionStart="Fig" captionStartId="1.[150,181,2064,2083]" captionTargetBox="[151,1437,799,2034]" captionTargetId="figure@1.[151,1437,798,2034]" captionTargetPageId="1" captionText="Fig. 1. Dorsal habitus of Guatemalan Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3879762" httpUri="https://zenodo.org/record/3879762/files/figure.png" pageId="4" pageNumber="295">Figs 1</figureCitation>
,
<figureCitation id="169858C39B203708FBF71598FB1FFBE7" box="[1148,1160,1020,1039]" captionStart="Fig" captionStartId="5.[151,182,2060,2079]" captionTargetBox="[150,1435,783,2035]" captionTargetId="figure@5.[150,1435,782,2035]" captionTargetPageId="5" captionText="Fig. 4. Hydrochus obscurus Sharp, 1882, habitus and male genitalia of lectotype." figureDoi="http://doi.org/10.5281/zenodo.3879770" httpUri="https://zenodo.org/record/3879770/files/figure.png" pageId="4" pageNumber="295">4</figureCitation>
,
<figureCitation id="169858C39B203708FB1A1598FB0BFBE7" box="[1169,1180,1020,1039]" captionStart="Fig" captionStartId="9.[150,181,2064,2083]" captionTargetBox="[266,1320,1467,2045]" captionTargetId="figure@9.[266,1320,1467,2045]" captionTargetPageId="9" captionText="Fig. 8 Hydrochus lectotype labels.A H. obscurus Sharp, 1882; B H. debilis Sharp, 1882." figureDoi="http://doi.org/10.5281/zenodo.3879778" httpUri="https://zenodo.org/record/3879778/files/figure.png" pageId="4" pageNumber="295">8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C6B917CD9B203708FCB7127AFC54FB8F" pageId="4" pageNumber="295" type="reference_group">
<paragraph id="8E1C44469B203708FCB7127AFBE9FBD9" blockId="4.[828,1436,989,1127]" box="[828,1150,1054,1073]" pageId="4" pageNumber="295">
<treatmentCitationGroup id="AEB363689B203708FCB7127AFBE9FBD9" box="[828,1150,1054,1073]" pageId="4" pageNumber="295">
<treatmentCitation id="0F0262579B203708FCB7127AFBECFBD9" author="SHARP D." box="[828,1147,1054,1073]" page="90" pageId="4" pageNumber="295" year="1882">
<taxonomicName id="49A33FC59B203708FCB7127AFBECFBD9" ID-CoL="6MK96" authority="Sharp, 1882: 90" authorityName="Sharp" authorityPageNumber="90" authorityYear="1882" box="[828,1147,1054,1073]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="295" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="BCD798549B203708FCB7127AFC7CFBD9" box="[828,1003,1054,1073]" italics="true" pageId="4" pageNumber="295">Hydrochus obscurus</emphasis>
<bibRefCitation id="EA3239B79B203708FC7B127AFBECFBD9" author="SHARP D." box="[1008,1147,1054,1073]" pageId="4" pageNumber="295" pagination="1 - 144" refId="ref6023" refString="SHARP D. 1882: Insecta. Coleoptera. Vol. 1, part 2 (Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae, Heteroceridae, Parnidae, Georissidae, Cyathoceridae, Staphylinidae). Pp 1 - 144. In: GODMAN F. D. &amp; SALVIN O.: Biologia Centrali-Americana. Volume 16. Taylor and Francis, London, xv + 824 pp." type="book chapter" year="1882">Sharp, 1882: 90</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8E1C44469B203708FCB7125DFC54FB8F" blockId="4.[828,1436,989,1127]" pageId="4" pageNumber="295">
<emphasis id="BCD798549B203708FCB7125DFBFBFBA4" box="[828,1132,1081,1100]" italics="true" pageId="4" pageNumber="295">
<taxonomicName id="49A33FC59B203708FCB7125DFC0AFBA4" ID-CoL="8VRRD" authorityName="Leach" authorityYear="1817" box="[828,925,1081,1100]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="295" phylum="Arthropoda" rank="genus">Hydrochus</taxonomicName>
vanbergehenegouweni
</emphasis>
<bibRefCitation id="EA3239B79B203708FBFF125DFB60FBA4" author="MAKHAN D." box="[1140,1271,1081,1100]" pageId="4" pageNumber="295" pagination="95 - 103" refId="ref5463" refString="MAKHAN D. 1992: Twelve new Hydrochus-species from South America. Phegea 202: 95 - 103." type="journal article" year="1992">Makhan, 1992</bibRefCitation>
(synonymized by OĿıඏA 1996)
</paragraph>
</subSubSection>
<subSubSection id="C6B917CD9B203708FCB7121AFB5FF893" pageId="4" pageNumber="295" type="materials_examined">
<paragraph id="8E1C44469B203708FCB7121AFB6AFA8E" blockId="4.[828,1438,1150,1515]" pageId="4" pageNumber="295">
<emphasis id="BCD798549B203708FCB7121AFB80FB79" bold="true" box="[828,1047,1150,1169]" pageId="4" pageNumber="295">Type material examined.</emphasis>
LൾǤŦƟŦYPൾ (hereby designated): male (
<collectionCode id="E8B2DC839B203708FAD6121AFA04FB79" box="[1373,1427,1150,1169]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="4" pageNumber="295" type="Herbarium">NHM</collectionCode>
):
<taxonomicName id="49A33FC59B203708FCB412FCFB01FB43" authority="D. S. Paso Antonio." authorityName="D. S. Paso Antonio." box="[831,1174,1176,1195]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="295" phylum="Arthropoda" rank="species" species="obscurus">Hydrochus obscurus D. S. Paso Antonio.</taxonomicName>
<collectingCountry id="F6B404D69B203708FB1212FCFB60FB43" box="[1177,1271,1176,1195]" name="Guatemala" pageId="4" pageNumber="295">Guatemala</collectingCountry>
.Champion [written on specimen card] // [male symbol] //
<typeStatus id="5118FAE49B203708FB0812D7FB6AFB2E" box="[1155,1277,1203,1222]" pageId="4" pageNumber="295" type="lectotype">LECTOTYPE</typeStatus>
[blue disc] // Paso Antonio,
<quantity id="495BE9A39B203708FC0212AAFC28FB09" box="[905,959,1230,1249]" metricMagnitude="2" metricUnit="m" metricValue="1.2192" pageId="4" pageNumber="295" unit="ft" value="400.0">400 ft.</quantity>
Champion.// B.C.A. Col. I. 2.
<taxonomicName id="49A33FC59B203708FB3212AAFA09FB09" authority="Sharp." authorityName="Sharp." box="[1209,1438,1230,1249]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="295" phylum="Arthropoda" rank="species" species="obscurus">Hydrochus obscurus Sharp.</taxonomicName>
// HABITUS/GENIT. DIGITAL IMAGES captured 2020 P.D. Perkins // [red]
<typeStatus id="5118FAE49B203708FCF01367FC62FAFE" box="[891,1013,1283,1302]" pageId="4" pageNumber="295" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="49A33FC59B203708FC711367FB77FAFE" authority="Sharp" authorityName="Sharp" box="[1018,1248,1283,1302]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="295" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="BCD798549B203708FC711367FB3EFAFE" box="[1018,1193,1283,1302]" italics="true" pageId="4" pageNumber="295">Hydrochus obscurus</emphasis>
Sharp
</taxonomicName>
desig. P. D. Perkins.ʼ. PARAĿൾǤŦƟŦYPൾඌ:
<specimenCount id="98A58FCF9B203708FC5E137AFBB1FAD9" box="[981,1062,1310,1329]" pageId="4" pageNumber="295" type="female">2 females</specimenCount>
(
<collectionCode id="E8B2DC839B203708FBBB137AFBF0FAD9" box="[1072,1127,1310,1329]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="4" pageNumber="295" type="Herbarium">NHM</collectionCode>
): label data same as
<typeStatus id="5118FAE49B203708FA85137AFAC9FAD9" box="[1294,1374,1310,1329]" pageId="4" pageNumber="295" type="lectotype">lectotype</typeStatus>
, except with blue
<typeStatus id="5118FAE49B203708FC1B135CFB94FAA3" box="[912,1027,1336,1355]" pageId="4" pageNumber="295" type="paralectotype">paralectotype</typeStatus>
labels, and yellow labels:
<typeStatus id="5118FAE49B203708FB6C135CFA0BFAA3" box="[1255,1436,1336,1355]" pageId="4" pageNumber="295" type="paralectotype">PARALECTOTYPE</typeStatus>
//
<taxonomicName id="49A33FC59B203708FCC61337FBA4FA8E" authority="Sharp" authorityName="Sharp" box="[845,1075,1363,1382]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="295" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="BCD798549B203708FCC61337FC6BFA8E" box="[845,1020,1363,1382]" italics="true" pageId="4" pageNumber="295">Hydrochus obscurus</emphasis>
Sharp
</taxonomicName>
// desig. P. D. Perkinsʼ.
</paragraph>
<paragraph id="8E1C44469B203708FCB7130AFA1FFA03" blockId="4.[828,1438,1150,1515]" pageId="4" pageNumber="295">
<emphasis id="BCD798549B203708FCB7130AFBDFFA69" bold="true" box="[828,1096,1390,1409]" pageId="4" pageNumber="295">Additional material examined</emphasis>
(
<specimenCount id="98A58FCF9B203708FBD9130AFB0BFA69" box="[1106,1180,1390,1409]" pageId="4" pageNumber="295" type="generic">26 spec.</specimenCount>
):
<emphasis id="BCD798549B203708FB23130AFA8EFA69" bold="true" box="[1192,1305,1390,1409]" pageId="4" pageNumber="295">GƲAŦFMAĿA:</emphasis>
<collectingRegion id="4C678AA49B203708FA97130AFAC5FA69" box="[1308,1362,1390,1409]" country="Guatemala" name="Izabal" pageId="4" pageNumber="295">Izabal</collectingRegion>
,
<quantity id="495BE9A39B203708FAD2130AFA12FA69" box="[1369,1413,1390,1409]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="4" pageNumber="295" unit="mi" value="1.0">1 mi.</quantity>
N. Morales. Km 239 Atlantic Hwy,
<date id="FA1D62869B203708FBC613ECFB45FA73" box="[1101,1234,1416,1435]" pageId="4" pageNumber="295" value="1965-08-16" valueMax="1965-08-18" valueMin="1965-08-16">16-18.viii.1965</date>
, leg. Paul J. Spangler (2
<collectionCode id="E8B2DC839B203708FCB713C7FC16FA5E" box="[828,897,1443,1462]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="295" type="Museum">NMNH</collectionCode>
);
<quantity id="495BE9A39B203708FC0513C7FC53FA5E" box="[910,964,1443,1462]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="4" pageNumber="295" unit="mi" value="20.0">20 mi.</quantity>
SW Puerto Barrios,
<date id="FA1D62869B203708FBE613C7FB41FA5E" box="[1133,1238,1443,1462]" pageId="4" pageNumber="295" value="1965-08-16">16.viii.1965</date>
, leg. Paul J. Spangler (5
<collectionCode id="E8B2DC839B203708FCB713DAFC16FA39" box="[828,897,1470,1489]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="295" type="Museum">NMNH</collectionCode>
);
<quantity id="495BE9A39B203708FC1B13DAFC5FFA39" box="[912,968,1470,1489]" metricMagnitude="4" metricUnit="m" metricValue="4.5061632" pageId="4" pageNumber="295" unit="mi" value="28.0">28 mi.</quantity>
SW Puerto Barrios,
<date id="FA1D62869B203708FBF113DAFB74FA39" box="[1146,1251,1470,1489]" pageId="4" pageNumber="295" value="1965-08-16">16.viii.1965</date>
, leg. Paul J. Spangler (18:
<collectionCode id="E8B2DC839B203708FCE813BCFC3FFA03" box="[867,936,1496,1515]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="295" type="Museum">NMNH</collectionCode>
); Quirigua,
<date id="FA1D62869B203708FB8613BCFBE0FA03" box="[1037,1143,1496,1515]" pageId="4" pageNumber="295" value="1965-08-14">14.viii.1965</date>
, leg. P. J. Spangler (1:
<collectionCode id="E8B2DC839B203708FAB213BCFAE9FA03" box="[1337,1406,1496,1515]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="295" type="Museum">NMNH</collectionCode>
).
</paragraph>
<paragraph id="8E1C44469B203708FCB71060FB5FF893" blockId="4.[828,1437,1540,2075]" pageId="4" pageNumber="295">
<materialsCitation id="3ECB4E1B9B203708FCB71060FB5FF893" ID-GBIF-Occurrence="2632269301" collectorName="J. L. Hellman &amp; This" country="Guatemala" location="Guatemala" pageId="4" pageNumber="295" specimenCount="8" typeStatus="lectotype">
<emphasis id="BCD798549B203708FCB71060FBA8F9F3" bold="true" box="[828,1087,1540,1563]" pageId="4" pageNumber="295">
<typeStatus id="5118FAE49B203708FCB71060FC39F9F3" box="[828,942,1540,1563]" pageId="4" pageNumber="295" type="lectotype">Lectotype</typeStatus>
designation.
</emphasis>
The NHM kindly provided three
<typeStatus id="5118FAE49B203708FCB71040FC0CF9D3" box="[828,923,1572,1595]" pageId="4" pageNumber="295" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="49A33FC59B203708FC481040FB33F9D3" box="[963,1188,1572,1595]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="295" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="BCD798549B203708FC481040FB33F9D3" box="[963,1188,1572,1595]" italics="true" pageId="4" pageNumber="295">Hydrochus obscurus</emphasis>
</taxonomicName>
, which are part of the Sharp
<emphasis id="BCD798549B203708FC031020FB42F9B3" box="[904,1237,1604,1627]" italics="true" pageId="4" pageNumber="295">Biologia Centrali-Americana</emphasis>
types collection. The
<typeStatus id="5118FAE49B203708FCE41000FC59F993" box="[879,974,1636,1659]" pageId="4" pageNumber="295" type="syntype">syntypes</typeStatus>
consist of one male and two females, all from Paso Antonio,
<collectingCountry id="F6B404D69B203708FB9E10E0FB18F973" box="[1045,1167,1668,1691]" name="Guatemala" pageId="4" pageNumber="295">Guatemala</collectingCountry>
. I have selected the male to serve as
<typeStatus id="5118FAE49B203708FC3C10C0FB8CF953" box="[951,1051,1700,1723]" pageId="4" pageNumber="295" type="lectotype">lectotype</typeStatus>
and labeled it as such, and provide images of the habitus, male genitalia, and labels of that specimen (
<figureCitation id="169858C39B203708FC381080FC6CF913" box="[947,1019,1764,1787]" captionStart="Fig" captionStartId="1.[150,181,2064,2083]" captionTargetBox="[151,1437,799,2034]" captionTargetId="figure@1.[151,1437,798,2034]" captionTargetPageId="1" captionText="Fig. 1. Dorsal habitus of Guatemalan Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.3879762" httpUri="https://zenodo.org/record/3879762/files/figure.png" pageId="4" pageNumber="295">Figs 1</figureCitation>
,
<figureCitation id="169858C39B203708FB801080FB8CF913" box="[1035,1051,1764,1787]" captionStart="Fig" captionStartId="5.[151,182,2060,2079]" captionTargetBox="[150,1435,783,2035]" captionTargetId="figure@5.[150,1435,782,2035]" captionTargetPageId="5" captionText="Fig. 4. Hydrochus obscurus Sharp, 1882, habitus and male genitalia of lectotype." figureDoi="http://doi.org/10.5281/zenodo.3879770" httpUri="https://zenodo.org/record/3879770/files/figure.png" pageId="4" pageNumber="295">4</figureCitation>
,
<figureCitation id="169858C39B203708FBA21080FBAFF913" box="[1065,1080,1764,1787]" captionStart="Fig" captionStartId="9.[150,181,2064,2083]" captionTargetBox="[266,1320,1467,2045]" captionTargetId="figure@9.[266,1320,1467,2045]" captionTargetPageId="9" captionText="Fig. 8 Hydrochus lectotype labels.A H. obscurus Sharp, 1882; B H. debilis Sharp, 1882." figureDoi="http://doi.org/10.5281/zenodo.3879778" httpUri="https://zenodo.org/record/3879778/files/figure.png" pageId="4" pageNumber="295">8</figureCitation>
). The other
<typeStatus id="5118FAE49B203708FB4F1080FAB4F913" box="[1220,1315,1764,1787]" pageId="4" pageNumber="295" type="syntype">syntypes</typeStatus>
have been labeled
<typeStatus id="5118FAE49B203708FC121160FBD5F8F3" box="[921,1090,1796,1819]" pageId="4" pageNumber="295" type="paralectotype">paralectotypes</typeStatus>
. One of the
<typeStatus id="5118FAE49B203708FB501160FAA8F8F3" box="[1243,1343,1796,1819]" pageId="4" pageNumber="295" type="syntype">syntypes</typeStatus>
bears a
<typeStatus id="5118FAE49B203708FCB71140FC36F8D3" box="[828,929,1828,1851]" pageId="4" pageNumber="295" type="lectotype">lectotype</typeStatus>
label by
<collectorName id="235621909B203708FB9B1140FB27F8D3" box="[1040,1200,1828,1851]" pageId="4" pageNumber="295">J. L. Hellman</collectorName>
, but this designation was never published.
<collectorName id="235621909B203708FBA51120FBC9F8B3" box="[1070,1118,1860,1883]" pageId="4" pageNumber="295">This</collectorName>
label has been reversed and unpub.ʼ written on the reverse side.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C6B917CD9B203709FCB711E0FE94FEDD" lastPageId="5" lastPageNumber="296" pageId="4" pageNumber="295" type="diagnosis">
<paragraph id="8E1C44469B203709FCB711E0FE94FEDD" blockId="4.[828,1437,1540,2075]" lastBlockId="5.[151,761,190,725]" lastPageId="5" lastPageNumber="296" pageId="4" pageNumber="295">
<emphasis id="BCD798549B203708FCB711E0FBDEF873" bold="true" box="[828,1097,1924,1947]" pageId="4" pageNumber="295">Differential diagnosis.</emphasis>
Among Guatemalan species,
<taxonomicName id="49A33FC59B203708FCB711C1FC2DF853" box="[828,954,1956,1980]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="295" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="BCD798549B203708FCB711C1FC2DF853" box="[828,954,1956,1980]" italics="true" pageId="4" pageNumber="295">H. obscurus</emphasis>
</taxonomicName>
is recognized by the combination of moderate size (ca.
<quantity id="495BE9A39B203708FC1E11A0FBA7F833" box="[917,1072,1988,2011]" metricMagnitude="-3" metricUnit="m" metricValue="2.96" metricValueMax="3.23" metricValueMin="2.69" pageId="4" pageNumber="295" unit="mm" value="2.96" valueMax="3.23" valueMin="2.69">2.693.23 mm</quantity>
), the moderately low and relatively impunctate pronotal reliefs, and the male genitalia (
<figureCitation id="169858C39B203708FAFA1180FCDAF7F3" captionStart="Fig" captionStartId="5.[151,182,2060,2079]" captionTargetBox="[150,1435,783,2035]" captionTargetId="figure@5.[150,1435,782,2035]" captionTargetPageId="5" captionText="Fig. 4. Hydrochus obscurus Sharp, 1882, habitus and male genitalia of lectotype." figureDoi="http://doi.org/10.5281/zenodo.3879770" httpUri="https://zenodo.org/record/3879770/files/figure.png" pageId="4" pageNumber="295">Fig. 4</figureCitation>
). This species has a generalized
<taxonomicName id="49A33FC59B203708FB4E1E60FAABF7F3" authorityName="Leach" authorityYear="1817" box="[1221,1340,2052,2075]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="BCD798549B203708FB4E1E60FAABF7F3" box="[1221,1340,2052,2075]" italics="true" pageId="4" pageNumber="295">Hydrochus</emphasis>
</taxonomicName>
habitus; reliable determinations will require dissection of males. The parameres are slightly spread in the
<typeStatus id="5118FAE49B213709FDCD16BAFD3CFF1D" box="[582,683,222,245]" pageId="5" pageNumber="296" type="lectotype">lectotype</typeStatus>
; see
<taxonomicName id="49A33FC59B213709FD5516BBFF62FEFD" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="296" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="BCD798549B213709FD5516BBFF62FEFD" italics="true" pageId="5" pageNumber="296">H. obscurus</emphasis>
</taxonomicName>
genitalia figure in PൾRĸıNඌ (2019b) for non-spread condition.
</paragraph>
</subSubSection>
<subSubSection id="C6B917CD9B21370AFF1C175AFD0DFE5D" lastPageId="6" lastPageNumber="297" pageId="5" pageNumber="296" type="description">
<paragraph id="8E1C44469B213709FF1C175AFEEFFD3D" blockId="5.[151,761,190,725]" pageId="5" pageNumber="296">
<emphasis id="BCD798549B213709FF1C175AFEAAFEBD" bold="true" box="[151,317,318,341]" pageId="5" pageNumber="296">Redescription.</emphasis>
Habitus as illustrated in
<figureCitation id="169858C39B213709FDDC175AFD0CFEBD" box="[599,667,318,341]" captionStart="Fig" captionStartId="5.[151,182,2060,2079]" captionTargetBox="[150,1435,783,2035]" captionTargetId="figure@5.[150,1435,782,2035]" captionTargetPageId="5" captionText="Fig. 4. Hydrochus obscurus Sharp, 1882, habitus and male genitalia of lectotype." figureDoi="http://doi.org/10.5281/zenodo.3879770" httpUri="https://zenodo.org/record/3879770/files/figure.png" pageId="5" pageNumber="296">Fig. 4</figureCitation>
. Size of
<typeStatus id="5118FAE49B213709FF1C173AFF6EFE9D" box="[151,249,350,373]" pageId="5" pageNumber="296" type="lectotype">lectotype</typeStatus>
(in mm): body (length to elytral apices / width) 2.92 / 0.98; head width 0.69; pronotum length / width 0.63 / 0.67; PA 0.63; PB 0.58; elytra length / width 1.81 / 0.98. Body size moderate (ca.
<quantity id="495BE9A39B213709FE6417DAFD1CFE3D" box="[495,651,446,469]" metricMagnitude="-3" metricUnit="m" metricValue="2.96" metricValueMax="3.23" metricValueMin="2.69" pageId="5" pageNumber="296" unit="mm" value="2.96" valueMax="3.23" valueMin="2.69">2.693.23 mm</quantity>
in Guatemalan specimens). Dorsum grey, with slight to moderately strong iridescence, elytra with diffuse dark grey to black spots. Legs testaceous, with tibiofemoral joints darker. Punctation on elytra ca. 23× that of pronotum. Elytra interstriae ca. 1.01.5× strial puncture diameter. Interstria 9
<superScript id="79D6E90E9B213709FF2F1418FF27FD62" attach="left" box="[164,176,636,650]" fontSize="6" pageId="5" pageNumber="296">th</superScript>
more convex than others, overhanging 10
<superScript id="79D6E90E9B213709FDF61418FD1EFD62" attach="left" box="[637,649,636,650]" fontSize="6" pageId="5" pageNumber="296">th</superScript>
interstria. Callus on 5
<superScript id="79D6E90E9B213709FE8714F8FE8FFD42" attach="left" box="[268,280,668,682]" fontSize="6" pageId="5" pageNumber="296">th</superScript>
interstria in usual location, moderately raised, moderately elongate.
</paragraph>
<paragraph id="8E1C44469B213709FCD116DAFC2DFE7D" blockId="5.[828,1437,190,725]" pageId="5" pageNumber="296">Pronotum length and width equal, or only very slightly wider than long, widest slightly behind anterior margin, narrowed at base, sides arcuate or very weakly sinuate, smooth; anterior margin slightly bisinuate; depressions deep to moderately deep, much more densely punctate than low reliefs between depressions, latter almost entirely impunctate.</paragraph>
<paragraph id="8E1C44469B213709FCD117FAFC23FDFD" blockId="5.[828,1437,190,725]" pageId="5" pageNumber="296">Elytra with spaces between strial punctures ca. 0.51.0× their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with slight angulation.</paragraph>
<paragraph id="8E1C44469B213709FCD1147AFA83FDBD" blockId="5.[828,1437,190,725]" pageId="5" pageNumber="296">Ventral characters: Mentum coarsely punctate, with median fovea. Submentum with two foveae.</paragraph>
<paragraph id="8E1C44469B21370AFCD1143AFD6EFEDD" blockId="5.[828,1437,190,725]" lastBlockId="6.[151,761,190,597]" lastPageId="6" lastPageNumber="297" pageId="5" pageNumber="296">
Male genitalia (
<figureCitation id="169858C39B213709FB8D143AFBDDFD9D" box="[1030,1098,606,629]" captionStart="Fig" captionStartId="5.[151,182,2060,2079]" captionTargetBox="[150,1435,783,2035]" captionTargetId="figure@5.[150,1435,782,2035]" captionTargetPageId="5" captionText="Fig. 4. Hydrochus obscurus Sharp, 1882, habitus and male genitalia of lectotype." figureDoi="http://doi.org/10.5281/zenodo.3879770" httpUri="https://zenodo.org/record/3879770/files/figure.png" pageId="5" pageNumber="296">Fig. 4</figureCitation>
). General characters: paramere tips in dorsal/ventral views very weakly sagittate, lateral margins between
<emphasis id="BCD798549B213709FC7114FAFBB1FD5D" box="[1018,1062,670,693]" italics="true" pageId="5" pageNumber="296">plsa</emphasis>
and distal end very slightly emarginate, medial margins arcuate toward midline, in lateral view tips narrow and tapering; aedeagus basically lanceolate shaped,
<emphasis id="BCD798549B22370AFEA716BBFECFFF1E" box="[300,344,223,246]" italics="true" pageId="6" pageNumber="297">agw</emphasis>
slightly before midlength; basal piece slightly shorter than parameres, with orifice sclerotized rim produced in small projection on apicoventral margin.
</paragraph>
<caption id="DADC14CE9B213709FF1C1E68FCDCF7F7" ID-DOI="http://doi.org/10.5281/zenodo.3879770" ID-Zenodo-Dep="3879770" box="[151,843,2060,2079]" httpUri="https://zenodo.org/record/3879770/files/figure.png" pageId="5" pageNumber="296" startId="5.[151,182,2060,2079]" targetBox="[150,1435,783,2035]" targetPageId="5">
<paragraph id="8E1C44469B213709FF1C1E68FCDCF7F7" blockId="5.[151,843,2060,2079]" box="[151,843,2060,2079]" pageId="5" pageNumber="296">
Fig. 4.
<taxonomicName id="49A33FC59B213709FF5F1E68FE65F7F7" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[212,498,2060,2079]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="296" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="BCD798549B213709FF5F1E68FE14F7F7" box="[212,387,2060,2079]" italics="true" pageId="5" pageNumber="296">Hydrochus obscurus</emphasis>
<bibRefCitation id="EA3239B79B213709FE031E68FE65F7F7" author="SHARP D." box="[392,498,2060,2079]" pageId="5" pageNumber="296" pagination="1 - 144" refId="ref6023" refString="SHARP D. 1882: Insecta. Coleoptera. Vol. 1, part 2 (Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae, Heteroceridae, Parnidae, Georissidae, Cyathoceridae, Staphylinidae). Pp 1 - 144. In: GODMAN F. D. &amp; SALVIN O.: Biologia Centrali-Americana. Volume 16. Taylor and Francis, London, xv + 824 pp." type="book chapter" year="1882">Sharp, 1882</bibRefCitation>
</taxonomicName>
, habitus and male genitalia of lectotype.
</paragraph>
</caption>
<paragraph id="8E1C44469B22370AFF3E175AFE72FE9D" blockId="6.[151,761,190,597]" pageId="6" pageNumber="297">
Dorsal surface:
<emphasis id="BCD798549B22370AFEEF175AFE1AFEBD" box="[356,397,318,341]" italics="true" pageId="6" pageNumber="297">adtl</emphasis>
at about distal third;
<emphasis id="BCD798549B22370AFDF5175AFD3AFEBD" box="[638,685,318,341]" italics="true" pageId="6" pageNumber="297">adbl</emphasis>
small;
<emphasis id="BCD798549B22370AFF1C173AFF4FFE9D" box="[151,216,350,373]" italics="true" pageId="6" pageNumber="297">pdmm</emphasis>
overlapping
<emphasis id="BCD798549B22370AFEEE173AFE1AFE9D" box="[357,397,350,373]" italics="true" pageId="6" pageNumber="297">alm</emphasis>
basally.
</paragraph>
<paragraph id="8E1C44469B22370AFF3E171AFD0DFE5D" blockId="6.[151,761,190,597]" pageId="6" pageNumber="297">
Ventral surface:
<emphasis id="BCD798549B22370AFEE5171AFE00FE7D" box="[366,407,382,405]" italics="true" pageId="6" pageNumber="297">avtl</emphasis>
at about proximal third;
<emphasis id="BCD798549B22370AFD3C171BFD6FFE7E" box="[695,760,383,406]" italics="true" pageId="6" pageNumber="297">pvmm</emphasis>
closely approximating aedeagus lateral margins.
</paragraph>
</subSubSection>
<subSubSection id="C6B917CD9B22370AFF1C17DAFF69FDBD" pageId="6" pageNumber="297" type="discussion">
<paragraph id="8E1C44469B22370AFF1C17DAFF69FDBD" blockId="6.[151,761,190,597]" pageId="6" pageNumber="297">
<emphasis id="BCD798549B22370AFF1C17DAFE91FE3D" bold="true" box="[151,262,446,469]" pageId="6" pageNumber="297">Remarks.</emphasis>
I have examined the
<typeStatus id="5118FAE49B22370AFE7717DAFDCDFE3D" box="[508,602,446,469]" pageId="6" pageNumber="297" type="holotype">holotype</typeStatus>
of
<emphasis id="BCD798549B22370AFD0817DAFE06FE1D" italics="true" pageId="6" pageNumber="297">
<taxonomicName id="49A33FC59B22370AFD0817DAFD60FE3D" authorityName="Leach" authorityYear="1817" box="[643,759,446,469]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="297" phylum="Arthropoda" rank="genus">Hydrochus</taxonomicName>
vanbergehenegouweni
</emphasis>
from
<collectingCountry id="F6B404D69B22370AFE5717BAFDC5FE1D" box="[476,594,478,501]" name="Venezuela" pageId="6" pageNumber="297">Venezuela</collectingCountry>
, including the male genitalia. I concur with OĿıඏA (1996) that
<emphasis id="BCD798549B22370AFD0F179AFE06FDDD" italics="true" pageId="6" pageNumber="297">
<taxonomicName id="49A33FC59B22370AFD0F179AFD6FFDFD" authorityName="Leach" authorityYear="1817" box="[644,760,510,533]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="297" phylum="Arthropoda" rank="genus">Hydrochus</taxonomicName>
vanbergehenegouweni
</emphasis>
is conspecific with
<taxonomicName id="49A33FC59B22370AFDF4147AFF60FDBD" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="297" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="BCD798549B22370AFDF4147AFF60FDBD" italics="true" pageId="6" pageNumber="297">Hydrochus obscurus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>