treatments-xml/data/03/C6/87/03C687E26F13D37BFE0DFA1AD522FC6C.xml

<document id="47C1954BA3BBE12E442EB5CF78C26DCC" ID-CLB-Dataset="6535" ID-DOI="10.1080/00222933.2014.902140" ID-GBIF-Dataset="58cf5eca-ad81-4c3d-97a3-c6b17ee9137b" ID-ISSN="1464-5262" ID-Zenodo-Dep="4631429" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616520937039" checkinUser="carolina" docAuthor="Dankittipakul, Pakawin &amp; Singtripop, Tippawan" docDate="2014" docId="03C687E26F13D37BFE0DFA1AD522FC6C" docLanguage="en" docName="JNATHIST.48.31-32.1923-1936.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 48 (31 - 32)" docSource="http://dx.doi.org/10.1080/00222933.2014.902140" docStyle="DocumentStyle:CD936EBDDF5274E0936345450CC2DAE4.5:JNatHist.2013-2015.journal_article.1cover" docStyleId="CD936EBDDF5274E0936345450CC2DAE4" docStyleName="JNatHist.2013-2015.journal_article.1cover" docStyleVersion="5" docTitle="Clubiona apiculata Dankittipakul &amp; Singtripop 2014, sp. nov." docType="treatment" docUuid="00CAE87E-8896-425C-A5ED-FC22C7293FFA" docUuidSource="ZooBank" docVersion="8" lastPageNumber="1930" masterDocId="FFFFFF9A6F15D373FFD8FF85D751FFC8" masterDocTitle="A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae)" masterLastPageNumber="1936" masterPageNumber="1923" pageNumber="1928" updateTime="1698934598660" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2E668C9C9815547E4C37C937A30D32F8">A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae)</mods:title>
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<mods:namePart id="15D20FAFCEC832AE9AF30F659A89ED9E">Dankittipakul, Pakawin</mods:namePart>
<mods:affiliation id="7C3BA3D3F9B3C547335CDE959FAB9715">Endocrinology Research Laboratory, Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand; &amp; Manukau Institute of Technology, Auckland, New Zealand</mods:affiliation>
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<mods:namePart id="B58A11C59BCF49FB346F04F657B1F09A">Singtripop, Tippawan</mods:namePart>
<mods:affiliation id="78DB92EE93347E2C186BBFFAAEB6C0DE">Endocrinology Research Laboratory, Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand;</mods:affiliation>
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<mods:title id="0726A5B56FEB815453809CFAB63BE4D8">Journal of Natural History</mods:title>
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<treatment id="03C687E26F13D37BFE0DFA1AD522FC6C" ID-DOI="http://doi.org/10.5281/zenodo.4783722" ID-GBIF-Taxon="182383859" ID-Zenodo-Dep="4783722" LSID="urn:lsid:zoobank.org:act:00CAE87E-8896-425C-A5ED-FC22C7293FFA" httpUri="http://treatment.plazi.org/id/03C687E26F13D37BFE0DFA1AD522FC6C" lastPageId="8" lastPageNumber="1930" pageId="6" pageNumber="1928">
<subSubSection id="C375657F6F13D375FE0DFA1AD45BFA70" box="[469,778,1439,1464]" pageId="6" pageNumber="1928" type="nomenclature">
<paragraph id="8BD036F46F13D375FE0DFA1AD45BFA70" blockId="6.[445,803,1439,1496]" box="[469,778,1439,1464]" pageId="6" pageNumber="1928">
<heading id="D09881986F13D375FE0DFA1AD45BFA70" box="[469,778,1439,1464]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="1928" reason="2">
<taxonomicName id="4C6F4D776F13D375FE0DFA1AD5F6FA7F" ID-CoL="5ZDBH" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[469,679,1439,1463]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1928" phylum="Arthropoda" rank="species" species="apiculata" status="sp. nov.">
<emphasis id="B91BEAE66F13D375FE0DFA1AD5F6FA7F" bold="true" box="[469,679,1439,1463]" italics="true" pageId="6" pageNumber="1928">Clubiona apiculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F13D375FD68FA25D45BFA70" box="[688,778,1440,1464]" pageId="6" pageNumber="1928" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C375657F6F13D375FE65FA45D472FA10" box="[445,803,1472,1496]" pageId="6" pageNumber="1928" type="description">
<paragraph id="8BD036F46F13D375FE65FA45D472FA10" blockId="6.[445,803,1439,1496]" box="[445,803,1472,1496]" pageId="6" pageNumber="1928">
(
<figureCitation id="13542A716F13D375FE1DFA45D52BFA10" box="[453,634,1472,1496]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="6" pageNumber="1928">Figures 1A,F,G</figureCitation>
, 
<figureCitation id="13542A716F13D375FD52FA45D594FA10" box="[650,709,1472,1496]" captionStart="Figure 2" captionStartId="4.[144,211,1288,1310]" captionTargetBox="[144,1104,168,1256]" captionTargetId="figure-110@4.[144,1104,168,1256]" captionTargetPageId="4" captionText="Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C) Clubiona conica sp. nov.; (D–I) Clubiona apiculata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4631433" httpUri="https://zenodo.org/record/4631433/files/figure.png" pageId="6" pageNumber="1928">2D–I</figureCitation>
, 
<figureCitation id="13542A716F13D375FD0DFA45D44BFA10" box="[725,794,1472,1496]" captionStart="Figure 3" captionStartId="5.[212,279,1578,1600]" captionTargetBox="[212,1172,511,1542]" captionTargetId="figure-138@5.[212,1172,511,1547]" captionTargetPageId="5" captionText="Figure 3. Clubiona apiculata sp. nov.; male holotype (A–E); female paratype (F,G). Male palp; (A) ventral view, (B) prolateral view, (C) retrolateral view, (D) dorsal view. (E) Apex of tegulum showing conductor, prolateral view. (F) Epigyne, ventral view. (G) Internal genitalia, dorsal view. Scale bars: A–D, 0.1 mm; F,G, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4631435" httpUri="https://zenodo.org/record/4631435/files/figure.png" pageId="6" pageNumber="1928">3A–G</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C375657F6F13D374FF48F999D30FFEE8" lastPageId="7" lastPageNumber="1929" pageId="6" pageNumber="1928" type="diagnosis">
<paragraph id="8BD036F46F13D375FF48F999D7ADF9FC" blockId="6.[144,1104,1564,1691]" box="[144,252,1564,1588]" pageId="6" pageNumber="1928">
<heading id="D09881986F13D375FF48F999D7ADF9FC" box="[144,252,1564,1588]" fontSize="10" level="3" pageId="6" pageNumber="1928" reason="5">
<emphasis id="B91BEAE66F13D375FF48F999D7ADF9FC" box="[144,252,1564,1588]" italics="true" pageId="6" pageNumber="1928">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036F46F13D374FF48F9C6D30FFEE8" blockId="6.[144,1104,1564,1691]" lastBlockId="7.[212,1172,168,288]" lastPageId="7" lastPageNumber="1929" pageId="6" pageNumber="1928">
Males of 
<taxonomicName id="4C6F4D776F13D375FF25F9C1D6D4F994" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[253,389,1604,1628]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1928" phylum="Arthropoda" rank="species" species="apiculata" status="sp. nov.">
<emphasis id="B91BEAE66F13D375FF25F9C1D6D4F994" box="[253,389,1604,1628]" italics="true" pageId="6" pageNumber="1928">C. apiculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F13D375FE56F9C6D6B9F993" box="[398,488,1603,1627]" pageId="6" pageNumber="1928" rank="species">sp. nov.</taxonomicNameLabel>
can be recognized by the presence of a basal tubercle on the dorso-retrolateral side of the cymbium (
<figureCitation id="13542A716F13D375FD6DF9E6D469F9B3" box="[693,824,1635,1659]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="6" pageNumber="1928">Figures 1G</figureCitation>
, 
<figureCitation id="13542A716F13D375FC91F9E6D4D6F9B3" box="[841,903,1635,1659]" captionStart="Figure 3" captionStartId="5.[212,279,1578,1600]" captionTargetBox="[212,1172,511,1542]" captionTargetId="figure-138@5.[212,1172,511,1547]" captionTargetPageId="5" captionText="Figure 3. Clubiona apiculata sp. nov.; male holotype (A–E); female paratype (F,G). Male palp; (A) ventral view, (B) prolateral view, (C) retrolateral view, (D) dorsal view. (E) Apex of tegulum showing conductor, prolateral view. (F) Epigyne, ventral view. (G) Internal genitalia, dorsal view. Scale bars: A–D, 0.1 mm; F,G, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4631435" httpUri="https://zenodo.org/record/4631435/files/figure.png" pageId="6" pageNumber="1928">3C,D</figureCitation>
), in combination with a colossal, rectangular, sclerotized VTA (
<figureCitation id="13542A716F13D375FD6FF906D467F953" box="[695,822,1667,1691]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="6" pageNumber="1928">Figures 1F</figureCitation>
, 
<figureCitation id="13542A716F13D375FC92F906D491F953" box="[842,960,1667,1691]" captionStart="Figure 3" captionStartId="5.[212,279,1578,1600]" captionTargetBox="[212,1172,511,1542]" captionTargetId="figure-138@5.[212,1172,511,1547]" captionTargetPageId="5" captionText="Figure 3. Clubiona apiculata sp. nov.; male holotype (A–E); female paratype (F,G). Male palp; (A) ventral view, (B) prolateral view, (C) retrolateral view, (D) dorsal view. (E) Apex of tegulum showing conductor, prolateral view. (F) Epigyne, ventral view. (G) Internal genitalia, dorsal view. Scale bars: A–D, 0.1 mm; F,G, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4631435" httpUri="https://zenodo.org/record/4631435/files/figure.png" pageId="6" pageNumber="1928">Figure 3B</figureCitation>
). Males are distinguished from other members of the species-group by the short triangular embolus, and by the heavily sclerotized RTA (compare 
<figureCitation id="13542A716F12D374FCF3FF4DD4F2FF28" box="[811,931,200,224]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="7" pageNumber="1929">Figure 1A</figureCitation>
with 
<figureCitation id="13542A716F12D374FC34FF4DD3D5FF28" box="[1004,1156,200,224]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="7" pageNumber="1929">Figure 1B–D</figureCitation>
). Females can be recognized by the oblong spermathecae and the distinctly larger spherical bursae connected to thick-walled insemination ducts (
<figureCitation id="13542A716F12D374FC7DFE8DD34AFEE8" box="[933,1051,264,288]" captionStart="Figure 2" captionStartId="4.[144,211,1288,1310]" captionTargetBox="[144,1104,168,1256]" captionTargetId="figure-110@4.[144,1104,168,1256]" captionTargetPageId="4" captionText="Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C) Clubiona conica sp. nov.; (D–I) Clubiona apiculata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4631433" httpUri="https://zenodo.org/record/4631433/files/figure.png" pageId="7" pageNumber="1929">Figure 2D</figureCitation>
, 
<figureCitation id="13542A716F12D374FBF3FE8DD31FFEE8" box="[1067,1102,264,288]" captionStart="Figure 3" captionStartId="5.[212,279,1578,1600]" captionTargetBox="[212,1172,511,1542]" captionTargetId="figure-138@5.[212,1172,511,1547]" captionTargetPageId="5" captionText="Figure 3. Clubiona apiculata sp. nov.; male holotype (A–E); female paratype (F,G). Male palp; (A) ventral view, (B) prolateral view, (C) retrolateral view, (D) dorsal view. (E) Apex of tegulum showing conductor, prolateral view. (F) Epigyne, ventral view. (G) Internal genitalia, dorsal view. Scale bars: A–D, 0.1 mm; F,G, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4631435" httpUri="https://zenodo.org/record/4631435/files/figure.png" pageId="7" pageNumber="1929">3G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C375657F6F12D374FF0CFEE5D4D8FDD7" pageId="7" pageNumber="1929" type="materials_examined">
<paragraph id="8BD036F46F12D374FF0CFEE5D621FEB0" blockId="7.[212,1172,352,479]" box="[212,368,352,376]" pageId="7" pageNumber="1929">
<heading id="D09881986F12D374FF0CFEE5D621FEB0" box="[212,368,352,376]" fontSize="10" level="3" pageId="7" pageNumber="1929" reason="5">
<emphasis id="B91BEAE66F12D374FF0CFEE5D621FEB0" box="[212,368,352,376]" italics="true" pageId="7" pageNumber="1929">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036F46F12D374FF0CFE0DD6BEFE17" blockId="7.[212,1172,352,479]" pageId="7" pageNumber="1929">
<materialsCitation id="3B073CA96F12D374FF0CFE0DD6BEFE17" ID-GBIF-Occurrence="3117074303" collectingDate="1987-05-01" collectorName="D. Burckhardt &amp; I. Lobl" country="Malaysia" elevation="2590" location="Kinabalu National Park" municipality="Borneo" pageId="7" pageNumber="1929" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B91BEAE66F12D374FF0CFE0DD613FE68" box="[212,322,392,416]" italics="true" pageId="7" pageNumber="1929">
<typeStatus id="54D488566F12D374FF0CFE0DD66FFE68" box="[212,318,392,416]" pageId="7" pageNumber="1929" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D69FD7D6F12D374FE97FE02D632FE57" box="[335,355,391,415]" pageId="7" pageNumber="1929" type="male">♂</specimenCount>
, 
<collectingCountry id="F37876646F12D374FEA0FE02D6B2FE57" box="[376,483,391,415]" name="Malaysia" pageId="7" pageNumber="1929">Malaysia</collectingCountry>
, 
<collectingMunicipality id="6BB4AC8E6F12D374FE2DFE02D51BFE57" box="[501,586,391,415]" pageId="7" pageNumber="1929">Borneo</collectingMunicipality>
, 
<collectingRegion id="49ABF8166F12D374FD85FE02D5BDFE57" box="[605,748,391,415]" country="Malaysia" name="Sabah" pageId="7" pageNumber="1929">Sabah State</collectingRegion>
, 
<location id="8EB0602F6F12D374FD26FE02D370FE57" LSID="urn:lsid:plazi:treatment:03C687E26F13D37BFE0DFA1AD522FC6C:8EB0602F6F12D374FD26FE02D370FE57" box="[766,1057,391,415]" country="Malaysia" municipality="Borneo" name="Kinabalu National Park" pageId="7" pageNumber="1929" stateProvince="Sabah">Kinabalu National Park</location>
, 
<quantity id="4C979B116F12D374FBEBFE02D3DEFE57" box="[1075,1167,391,415]" metricMagnitude="3" metricUnit="m" metricValue="2.59" pageId="7" pageNumber="1929" unit="m" value="2590.0">
<elevation id="0042D1C76F12D374FBEBFE02D3DEFE57" box="[1075,1167,391,415]" metricMagnitude="3" metricUnit="m" metricValue="2.59" pageId="7" pageNumber="1929" unit="m" value="2590.0">2590 m</elevation>
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, humid ravine below Layang Layang, cloud forest, 
<date id="FFD110346F12D374FCC3FE22D4F6FE77" box="[795,935,423,447]" pageId="7" pageNumber="1929" value="1987-05-01">
<collectingDate id="EF95E9DC6F12D374FCC3FE22D4F6FE77" box="[795,935,423,447]" pageId="7" pageNumber="1929" value="1987-05-01">1 May 1987</collectingDate>
</date>
, 
<collectorName id="269A53226F12D374FC6DFE22D30DFE77" box="[949,1116,423,447]" pageId="7" pageNumber="1929">D. Burckhardt</collectorName>
&amp; 
<collectorName id="269A53226F12D374FB5AFE22D65BFE17" pageId="7" pageNumber="1929">I. Löbl</collectorName>
leg. (MHNG, 10a).
</materialsCitation>
</paragraph>
<paragraph id="8BD036F46F12D374FF0CFD82D4D8FDD7" blockId="7.[212,905,519,543]" box="[212,905,519,543]" pageId="7" pageNumber="1929">
<materialsCitation id="3B073CA96F12D374FF0CFD82D4D4FDD7" ID-GBIF-Occurrence="3117074304" box="[212,901,519,543]" collectingDate="1987-05-01" collectorName="D. Burckhardt &amp; I. Lobl" country="Malaysia" elevation="2590" location="Kinabalu National Park" municipality="Borneo" pageId="7" pageNumber="1929" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="B91BEAE66F12D374FF0CFD82D611FDD7" box="[212,320,519,543]" italics="true" pageId="7" pageNumber="1929">
<typeStatus id="54D488566F12D374FF0CFD82D66CFDD7" box="[212,317,519,543]" pageId="7" pageNumber="1929" type="paratype">Paratype</typeStatus>
.
</emphasis>
One 
<specimenCount id="9D69FD7D6F12D374FE58FD82D6C5FDD7" box="[384,404,519,543]" pageId="7" pageNumber="1929" type="male">♂</specimenCount>
, 
<specimenCount id="9D69FD7D6F12D374FE7CFD82D694FDD7" box="[420,453,519,543]" pageId="7" pageNumber="1929" type="female">2♀</specimenCount>
, same data as holotype (MHNG, 10a)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C375657F6F12D374FF0CFDDAD3C5FD76" pageId="7" pageNumber="1929" type="etymology">
<paragraph id="8BD036F46F12D374FF0CFDDAD61CFDBF" blockId="7.[212,1172,607,702]" box="[212,333,607,631]" pageId="7" pageNumber="1929">
<heading id="D09881986F12D374FF0CFDDAD61CFDBF" box="[212,333,607,631]" fontSize="10" level="3" pageId="7" pageNumber="1929" reason="5">
<emphasis id="B91BEAE66F12D374FF0CFDDAD61CFDBF" box="[212,333,607,631]" italics="true" pageId="7" pageNumber="1929">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036F46F12D374FF0CFD02D3C5FD76" blockId="7.[212,1172,607,702]" pageId="7" pageNumber="1929">
The specific epithet is derived from the Latin adjective 
<emphasis id="B91BEAE66F12D374FCBBFD02D481FD57" box="[867,976,647,671]" italics="true" pageId="7" pageNumber="1929">apiculatus</emphasis>
, referring to the triangular embolus and its conical base, which are situated near the apex of the tegulum.
</paragraph>
</subSubSection>
<subSubSection id="C375657F6F12D37BFF0CFD7BD698FCCB" lastPageId="8" lastPageNumber="1930" pageId="7" pageNumber="1929" type="description">
<paragraph id="8BD036F46F12D374FF0CFD7BD602FCDE" blockId="7.[212,1172,766,1244]" box="[212,339,766,790]" pageId="7" pageNumber="1929">
<heading id="D09881986F12D374FF0CFD7BD602FCDE" box="[212,339,766,790]" fontSize="10" level="3" pageId="7" pageNumber="1929" reason="5">
<emphasis id="B91BEAE66F12D374FF0CFD7BD602FCDE" box="[212,339,766,790]" italics="true" pageId="7" pageNumber="1929">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036F46F12D374FF0CFCA3D316FB14" blockId="7.[212,1172,766,1244]" pageId="7" pageNumber="1929">
<emphasis id="B91BEAE66F12D374FF0CFCA3D65FFCF6" box="[212,270,806,830]" italics="true" pageId="7" pageNumber="1929">Male</emphasis>
(
<typeStatus id="54D488566F12D374FEC5FCA3D6D0FCF6" box="[285,385,806,830]" pageId="7" pageNumber="1929" type="holotype">holotype</typeStatus>
). Total length 6.9; prosoma 3.3 long, 2.0 wide; opisthosoma 3.6 long, 1.8 wide. Prosoma ovoid, in profile distinctly higher just in front longitudinal fovea; integument smooth; carapace uniform yellow; chelicerae yellowish brown, with three promarginal and three retromarginal teeth; labium and gnathocoxae pale brown; sternum yellow. Eye sizes and interdistances: AME 0.08, ALE 0.12, PME 0.18, PLE 0.16, AME–AME 0.08, AME–ALE 0.10, PME–PME 0.32, PME–PLE 0.18; MOQ: 0.42 long, 0.24 anterior width, 0.64 posterior width. Legs pale yellow, anterior metatarsi and tarsi brown, distinctly darker than other segments; scopulae and tufts indistinct. Leg formula 4213. Leg measurements: leg I 6.9 (1.9, 1.0, 1.8, 1.6, 0.6); leg II 7.6 (2.4, 1.0, 2.2, 1.4, 0.6); leg III 6.6 (1.6, 1.4, 1.4, 1.4, 0.6); leg IV 10.0 (2.8, 1.0, 2.4, 2.8, 1.0). Spines on anterior legs elongate and slender; anterior tibiae armed with three pairs of elongated ventral spines; anterior metatarsi armed with a pair of very long spines, originating proximally, apices almost reaching tarsus. Opisthosoma ovoid, pale, without colour marking; lightly sclerotized dorsal scutum extending half length of opisthosoma.
</paragraph>
<paragraph id="8BD036F46F12D374FF0CFA80D40FF9D4" blockId="7.[212,1172,1284,1564]" pageId="7" pageNumber="1929">
<emphasis id="B91BEAE66F12D374FF0CFA80D656FAD5" box="[212,263,1285,1309]" italics="true" pageId="7" pageNumber="1929">Palp</emphasis>
(
<figureCitation id="13542A716F12D374FEC4FA81D684FAD4" box="[284,469,1284,1308]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="7" pageNumber="1929">Figures 1A,F,G</figureCitation>
, 
<figureCitation id="13542A716F12D374FE31FA81D57BFAD4" box="[489,554,1284,1309]" captionStart="Figure 3" captionStartId="5.[212,279,1578,1600]" captionTargetBox="[212,1172,511,1542]" captionTargetId="figure-138@5.[212,1172,511,1547]" captionTargetPageId="5" captionText="Figure 3. Clubiona apiculata sp. nov.; male holotype (A–E); female paratype (F,G). Male palp; (A) ventral view, (B) prolateral view, (C) retrolateral view, (D) dorsal view. (E) Apex of tegulum showing conductor, prolateral view. (F) Epigyne, ventral view. (G) Internal genitalia, dorsal view. Scale bars: A–D, 0.1 mm; F,G, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4631435" httpUri="https://zenodo.org/record/4631435/files/figure.png" pageId="7" pageNumber="1929">3A–E</figureCitation>
). Palpal tibia with distinctly elevated VTA in ventral (
<figureCitation id="13542A716F12D374FF04FAA1D602FAF4" box="[220,339,1316,1340]" captionStart="Figure 3" captionStartId="5.[212,279,1578,1600]" captionTargetBox="[212,1172,511,1542]" captionTargetId="figure-138@5.[212,1172,511,1547]" captionTargetPageId="5" captionText="Figure 3. Clubiona apiculata sp. nov.; male holotype (A–E); female paratype (F,G). Male palp; (A) ventral view, (B) prolateral view, (C) retrolateral view, (D) dorsal view. (E) Apex of tegulum showing conductor, prolateral view. (F) Epigyne, ventral view. (G) Internal genitalia, dorsal view. Scale bars: A–D, 0.1 mm; F,G, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4631435" httpUri="https://zenodo.org/record/4631435/files/figure.png" pageId="7" pageNumber="1929">Figure 3A</figureCitation>
) and retrolateral views (
<figureCitation id="13542A716F12D374FDA1FAA1D5BEFAF4" box="[633,751,1316,1340]" captionStart="Figure 3" captionStartId="5.[212,279,1578,1600]" captionTargetBox="[212,1172,511,1542]" captionTargetId="figure-138@5.[212,1172,511,1547]" captionTargetPageId="5" captionText="Figure 3. Clubiona apiculata sp. nov.; male holotype (A–E); female paratype (F,G). Male palp; (A) ventral view, (B) prolateral view, (C) retrolateral view, (D) dorsal view. (E) Apex of tegulum showing conductor, prolateral view. (F) Epigyne, ventral view. (G) Internal genitalia, dorsal view. Scale bars: A–D, 0.1 mm; F,G, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4631435" httpUri="https://zenodo.org/record/4631435/files/figure.png" pageId="7" pageNumber="1929">Figure 3C</figureCitation>
), broad and massive rectangular in prolateral view (
<figureCitation id="13542A716F12D374FE5DFAC1D6AAFA94" box="[389,507,1348,1372]" captionStart="Figure 1" captionStartId="3.[212,279,1608,1630]" captionTargetBox="[212,1172,671,1576]" captionTargetId="figure-174@3.[212,1172,663,1576]" captionTargetPageId="3" captionText="Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E)." figureDoi="http://doi.org/10.5281/zenodo.4631431" httpUri="https://zenodo.org/record/4631431/files/figure.png" pageId="7" pageNumber="1929">Figures 1F</figureCitation>
, 
<figureCitation id="13542A716F12D374FDD0FAC1D577FA94" box="[520,550,1348,1372]" captionStart="Figure 3" captionStartId="5.[212,279,1578,1600]" captionTargetBox="[212,1172,511,1542]" captionTargetId="figure-138@5.[212,1172,511,1547]" captionTargetPageId="5" captionText="Figure 3. Clubiona apiculata sp. nov.; male holotype (A–E); female paratype (F,G). Male palp; (A) ventral view, (B) prolateral view, (C) retrolateral view, (D) dorsal view. (E) Apex of tegulum showing conductor, prolateral view. (F) Epigyne, ventral view. (G) Internal genitalia, dorsal view. Scale bars: A–D, 0.1 mm; F,G, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4631435" httpUri="https://zenodo.org/record/4631435/files/figure.png" pageId="7" pageNumber="1929">3B</figureCitation>
); RTA petal-shaped, heavily sclerotized, reaching almost half length of cymbium; cymbium apically with shallow dorsal excavation, covered with thick patch of hairs; digitiform tubercle originating on baso-retrolateral side of cymbium, directed horizontally, pointing ectad; tegulum oblong; conductor digitiform, translucent, originating on apico-retrolateral area of tegulum; sperm duct U-shaped; embolic base conical, situated apico-prolaterally; embolus triangular, minute, apex sharply pointed, directed anteriad, emerging subterminally on embolic base.
</paragraph>
<paragraph id="8BD036F46F12D37BFF0CF9C1D4ACFEC8" blockId="7.[212,1172,1603,1691]" lastBlockId="8.[144,1104,168,256]" lastPageId="8" lastPageNumber="1930" pageId="7" pageNumber="1929">
<emphasis id="B91BEAE66F12D374FF0CF9C1D675F994" box="[212,292,1604,1628]" italics="true" pageId="7" pageNumber="1929">Female</emphasis>
(
<typeStatus id="54D488566F12D374FEECF9C6D6C8F993" box="[308,409,1603,1627]" pageId="7" pageNumber="1929" type="paratype">paratype</typeStatus>
). Total length 5.6; prosoma 3.0 long, 2.6 wide; opisthosoma 2.6 long, 1.1 wide. General appearance as in male, including spination on anterior tibiae and metatarsi. AME 0.08, ALE 0.10, PME 0.18, PLE 0.14, AME–AME 0.08, AME–ALE 0.10, PME–PME 0.30, PME–PLE 0.16; MOQ: 0.38 long, 0.24 anterior width, 0.66 posterior width. Leg measurements: leg I 5.9 (1.7, 0.8, 1.6, 1.2, 0.6); leg II 7.0 (2.2, 0.8, 2.2, 1.2, 0.6); leg III 6.3 (1.6, 1.4, 1.2, 1.5, 0.6); leg IV 9.0 (2.6, 0.8, 2.2, 2.6, 0.8).
</paragraph>
<paragraph id="8BD036F46F1DD37BFF48FEAFD5A0FD89" blockId="8.[144,1104,298,577]" pageId="8" pageNumber="1930">
<emphasis id="B91BEAE66F1DD37BFF48FEAFD7A4FE8A" box="[144,245,298,322]" italics="true" pageId="8" pageNumber="1930">Genitalia</emphasis>
(
<figureCitation id="13542A716F1DD37BFEDDFEAFD6F2FE8A" box="[261,419,298,322]" captionStart="Figure 2" captionStartId="4.[144,211,1288,1310]" captionTargetBox="[144,1104,168,1256]" captionTargetId="figure-110@4.[144,1104,168,1256]" captionTargetPageId="4" captionText="Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C) Clubiona conica sp. nov.; (D–I) Clubiona apiculata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4631433" httpUri="https://zenodo.org/record/4631433/files/figure.png" pageId="8" pageNumber="1930">Figures 2D–F</figureCitation>
, 
<figureCitation id="13542A716F1DD37BFE6AFEAFD6BFFE8A" box="[434,494,298,322]" captionStart="Figure 3" captionStartId="5.[212,279,1578,1600]" captionTargetBox="[212,1172,511,1542]" captionTargetId="figure-138@5.[212,1172,511,1547]" captionTargetPageId="5" captionText="Figure 3. Clubiona apiculata sp. nov.; male holotype (A–E); female paratype (F,G). Male palp; (A) ventral view, (B) prolateral view, (C) retrolateral view, (D) dorsal view. (E) Apex of tegulum showing conductor, prolateral view. (F) Epigyne, ventral view. (G) Internal genitalia, dorsal view. Scale bars: A–D, 0.1 mm; F,G, 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4631435" httpUri="https://zenodo.org/record/4631435/files/figure.png" pageId="8" pageNumber="1930">3F,G</figureCitation>
). Epigynal region circular, yellow, weekly sclerotized; copulatory orifices situated underneath heavily sclerotized, V-shaped ridge protruding over epigastric furrow; insemination ducts thick-walled, twisted proximally to form semicircular loop, with funnel-shaped atrium internally, ducts ascending anteriorly and opening into base of spermathecae; spermathecae oblong, basolaterally with glandular head (
<figureCitation id="13542A716F1DD37BFE5AFE4CD6A5FE29" box="[386,500,457,481]" captionStart="Figure 2" captionStartId="4.[144,211,1288,1310]" captionTargetBox="[144,1104,168,1256]" captionTargetId="figure-110@4.[144,1104,168,1256]" captionTargetPageId="4" captionText="Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C) Clubiona conica sp. nov.; (D–I) Clubiona apiculata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4631433" httpUri="https://zenodo.org/record/4631433/files/figure.png" pageId="8" pageNumber="1930">Figure 2E</figureCitation>
); acicular fertilization ducts membranous, located on dorsal side near base of spermathecae (
<figureCitation id="13542A716F1DD37BFD9EFE6CD5E7FDC9" box="[582,694,489,513]" captionStart="Figure 2" captionStartId="4.[144,211,1288,1310]" captionTargetBox="[144,1104,168,1256]" captionTargetId="figure-110@4.[144,1104,168,1256]" captionTargetPageId="4" captionText="Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C) Clubiona conica sp. nov.; (D–I) Clubiona apiculata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4631433" httpUri="https://zenodo.org/record/4631433/files/figure.png" pageId="8" pageNumber="1930">Figure 2F</figureCitation>
); bursae spherical, hyaline, approximately twice diameter of spermathecae, connected to insemination ducts at mid length between copulatory orifices and spermathecae.
</paragraph>
<paragraph id="8BD036F46F1DD37BFF48FD06D66AFD53" blockId="8.[144,1104,643,771]" box="[144,315,643,667]" pageId="8" pageNumber="1930">
<heading id="D09881986F1DD37BFF48FD06D66AFD53" box="[144,315,643,667]" fontSize="10" level="3" pageId="8" pageNumber="1930" reason="5">
<emphasis id="B91BEAE66F1DD37BFF48FD06D66AFD53" box="[144,315,643,667]" italics="true" pageId="8" pageNumber="1930">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036F46F1DD37BFF48FD2ED698FCCB" blockId="8.[144,1104,643,771]" pageId="8" pageNumber="1930">
<taxonomicName id="4C6F4D776F1DD37BFF48FD2ED630FD0B" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[144,353,683,707]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="1930" phylum="Arthropoda" rank="species" species="apiculata" status="sp. nov.">
<emphasis id="B91BEAE66F1DD37BFF48FD2ED630FD0B" box="[144,353,683,707]" italics="true" pageId="8" pageNumber="1930">Clubiona apiculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F1DD37BFEB2FD2ED695FD0B" box="[362,452,683,707]" pageId="8" pageNumber="1930" rank="species">sp. nov.</taxonomicNameLabel>
is considered a high-altitude species. The 
<typeStatus id="54D488566F1DD37BFC7EFD2ED487FD0B" box="[934,982,683,707]" pageId="8" pageNumber="1930">type</typeStatus>
specimens were collected in a cloud forest (
<quantity id="4C979B116F1DD37BFDD1FD4ED531FD2B" box="[521,608,715,739]" metricMagnitude="3" metricUnit="m" metricValue="2.59" pageId="8" pageNumber="1930" unit="m" value="2590.0">2590 m</quantity>
) by sifting decomposing plant and organic material in a humid ravine.
</paragraph>
</subSubSection>
<subSubSection id="C375657F6F1DD37BFF48FCC0D522FC6C" pageId="8" pageNumber="1930" type="distribution">
<paragraph id="8BD036F46F1DD37BFF48FCC0D645FC95" blockId="8.[144,1104,837,933]" box="[144,276,837,861]" pageId="8" pageNumber="1930">
<heading id="D09881986F1DD37BFF48FCC0D645FC95" box="[144,276,837,861]" fontSize="10" level="3" pageId="8" pageNumber="1930" reason="5">
<emphasis id="B91BEAE66F1DD37BFF48FCC0D645FC95" box="[144,276,837,861]" italics="true" pageId="8" pageNumber="1930">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036F46F1DD37BFF48FCE8D522FC6C" blockId="8.[144,1104,837,933]" pageId="8" pageNumber="1930">
Known only from the 
<typeStatus id="54D488566F1DD37BFE53FCE8D6EAFC4D" box="[395,443,877,901]" pageId="8" pageNumber="1930">type</typeStatus>
locality (Kinabalu National Park, Borneo; separated from 
<taxonomicName id="4C6F4D776F1DD37BFF48FC08D7A8FC6D" authority="Dankittipakul &amp; Singtripop, 2014" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2014" box="[144,249,909,933]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="1930" phylum="Arthropoda" rank="species" species="conica" status="sp. nov.">
<emphasis id="B91BEAE66F1DD37BFF48FC08D7A8FC6D" box="[144,249,909,933]" italics="true" pageId="8" pageNumber="1930">C. conica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579D6F1DD37BFEDAFC09D60DFC6C" box="[258,348,908,932]" pageId="8" pageNumber="1930" rank="species">sp. nov.</taxonomicNameLabel>
by altitudinal gradient).
</paragraph>
</subSubSection>
</treatment>
</document>