treatments-xml/data/03/C0/FF/03C0FF7EFFB85856FF6B05A7FD89FB7A.xml

<document id="C7FDFE8F85B1F4C904DFD7C5FB86E339" ID-DOI="10.11646/zootaxa.5476.1.25" ID-ISSN="1175-5326" ID-Zenodo-Dep="12681792" ID-ZooBank="324743F9-D5D0-4FDE-A43B-5A1F4AD35C0F" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720375311343" checkinUser="plazi" docAuthor="Ng, Peter K. L. &amp; Naruse, Tohru" docDate="2024" docId="03C0FF7EFFB85856FF6B05A7FD89FB7A" docLanguage="en" docName="zootaxa.5476.1.25.pdf" docOrigin="Zootaxa 5476 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5476.1.25" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orcovita varungan Ng &amp; Naruse, 2024, n. sp." docType="treatment" docVersion="6" lastPageNumber="323" masterDocId="FFF98706FFB9585FFFFC0728FFB6FFAC" masterDocTitle="Orcovita varungan n. sp. (Crustacea: Decapoda: Varunidae) from Muna Island, southern Sulawesi, Indonesia" masterLastPageNumber="324" masterPageNumber="314" pageNumber="315" updateTime="1721067532891" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="140E90DA239C5BEC5ECD71949B4B13BE">Orcovita varungan n. sp. (Crustacea: Decapoda: Varunidae) from Muna Island, southern Sulawesi, Indonesia</mods:title>
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<mods:namePart id="811228A057F9FEF7FFDE987627121511">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="ADD812A2C13A3DB83883EA347724CA0E">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore.</mods:affiliation>
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<mods:namePart id="D183C17ED5C5EC41697C3115881F4B66">Naruse, Tohru</mods:namePart>
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<taxonomicName id="4C6935EBFFB8585EFF6B05A7FECAFD05" authorityName="Ng &amp; Naruse" authorityYear="2024" box="[151,380,655,681]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="315" phylum="Arthropoda" rank="species" species="varungan" status="sp. nov.">
<emphasis id="B91D927AFFB8585EFF6B05A7FECAFD05" bold="true" box="[151,380,655,681]" italics="true" pageId="1" pageNumber="315">Orcovita varungan</emphasis>
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<paragraph id="8BD64E68FFB8585EFF6B059BFEA4FD61" blockId="1.[151,454,655,717]" box="[151,274,691,717]" pageId="1" pageNumber="315">
(
<figureCitation id="135252EDFFB8585EFF63059BFEBFFD61" box="[159,265,691,717]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,250,1935,1959]" captionStartId-1="3.[151,250,1706,1730]" captionStartId-2="4.[151,250,1658,1682]" captionStartId-3="5.[151,250,1893,1917]" captionStartId-4="6.[151,250,1357,1381]" captionStartId-5="7.[151,250,1851,1875]" captionTargetBox-0="[151,1435,181,1911]" captionTargetBox-1="[151,1435,181,1682]" captionTargetBox-2="[158,1430,489,1634]" captionTargetBox-3="[151,1435,345,1870]" captionTargetBox-4="[151,1435,181,1333]" captionTargetBox-5="[177,1411,501,1810]" captionTargetId-0="figure-17@2.[151,1435,181,1911]" captionTargetId-1="figure-20@3.[151,1435,181,1682]" captionTargetId-2="figure-182@4.[158,1430,489,1634]" captionTargetId-3="figure-109@5.[151,1435,345,1870]" captionTargetId-4="figure-17@6.[151,1435,181,1333]" captionTargetId-5="figure-164@7.[151,1435,489,1827]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="FIGURE 1. Orcovita varungan n. sp., overall dorsal habitus. A, holotype male (22.9 × 17.6 mm) (SMF 5257a), Sulawesi; B, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; C, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi." captionText-1="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." captionText-2="FIGURE 3. Orcovita varungan n. sp., A, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; B, C, D, E, G, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, outer view of right chela; B, outer view of left chela; C, view of pulvinus at base of fingers of left chela; D, dorsal view of left cheliped; E, right P4 and P5; F, G, left third maxilliped." captionText-3="FIGURE 4. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, D, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; E, F, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, B, male anterior thoracic sternum and pleon; C–E, male thoracic sternum and pleon; F, male pleon." captionText-4="FIGURE 5. Orcovita varungan n. sp., A, B, E, F, paratype female (17.6 × 13.9 mm), ZRC 2023.0014, Sulawesi; C, D, paratype female (19.8 × 15.6 mm), SMF 5257b, Sulawesi. A, C, dorsal view of carapace; B, outer view of left chela; D, outer view of right chela; E, female anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." captionText-5="FIGURE 6. Gonopods of Orcovita varungan n. sp., holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi. A–D, with setae, before denuding; E–O, setae denuded. A–G, left G1; H–M, distal part of left G1 with corneous distal part; N, O. left G2. A, E, H, N, ventral view; B, I, subventral view; C, J, outer mesial view; D, G, K, O, dorsal view; L, subdorsal view; F, M, inner mesial view. Scales: E–G, N, O = 0.5 mm; H–M = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12681794" figureDoi-1="http://doi.org/10.5281/zenodo.12681796" figureDoi-2="http://doi.org/10.5281/zenodo.12681798" figureDoi-3="http://doi.org/10.5281/zenodo.12681800" figureDoi-4="http://doi.org/10.5281/zenodo.12681804" figureDoi-5="http://doi.org/10.5281/zenodo.12681806" httpUri-0="https://zenodo.org/record/12681794/files/figure.png" httpUri-1="https://zenodo.org/record/12681796/files/figure.png" httpUri-2="https://zenodo.org/record/12681798/files/figure.png" httpUri-3="https://zenodo.org/record/12681800/files/figure.png" httpUri-4="https://zenodo.org/record/12681804/files/figure.png" httpUri-5="https://zenodo.org/record/12681806/files/figure.png" pageId="1" pageNumber="315">Figs. 1–6</figureCitation>
)
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<emphasis id="B91D927AFFB8585EFF6B05D3FE8AFCB8" bold="true" box="[151,316,762,789]" pageId="1" pageNumber="315">Type material</emphasis>
. 
<materialsCitation id="3B014435FFB8585EFEB405D3FE3BFCF1" collectingDate="1908-08-10" collectionCode="SMF" collectorName="Sunda-Expedition &amp; J. Elbert" latitude="-4.82945" location="Muna" longLatPrecision="1" longitude="122.72825" municipality="Moena Raha" pageId="1" pageNumber="315" specimenCode="SMF 5257" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
<emphasis id="B91D927AFFB8585EFEB405D3FE19FCB9" box="[328,431,763,789]" italics="true" pageId="1" pageNumber="315">
<typeStatus id="54D2F0CAFFB8585EFEB405D3FE19FCB9" box="[328,431,763,789]" pageId="1" pageNumber="315" type="holotype">Holotype</typeStatus>
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: male (22.9 × 
<quantity id="4C91E38DFFB8585EFDB105D3FD05FCB9" box="[589,691,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="1" pageNumber="315" unit="mm" value="17.6">17.6 mm</quantity>
), 
<specimenCode id="DBCFE613FFB8585EFD3805D3FC8BFCB9" box="[708,829,762,789]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="315">SMF 5257</specimenCode>
a, 
<collectingMunicipality id="6BB2D412FFB8585EFCAE05D3FC69FCB9" box="[850,991,763,789]" pageId="1" pageNumber="315">Moena Raha</collectingMunicipality>
(
<location id="8EB618B3FFB8585EFC1005D3FB85FCB9" LSID="urn:lsid:plazi:treatment:03C0FF7EFFB85856FF6B05A7FD89FB7A:8EB618B3FFB8585EFC1005D3FB85FCB9" box="[1004,1075,763,789]" latitude="-4.82945" longLatPrecision="1" longitude="122.72825" municipality="Moena Raha" name="Muna" pageId="1" pageNumber="315" stateProvince="Sulawesi Utara">Muna</location>
), southern 
<collectingRegion id="49AD808AFFB8585EFB5605D3FAB9FCB9" box="[1194,1295,763,789]" country="Indonesia" name="Sulawesi Utara" pageId="1" pageNumber="315">Sulawesi</collectingRegion>
(= Celebes), 
<geoCoordinate id="EE5D28AFFFB8585EFF6B0437FEA4FC95" box="[151,274,799,825]" degrees="4" direction="south" minutes="49.767" orientation="latitude" pageId="1" pageNumber="315" precision="1" value="-4.82945">4°49.767'S</geoCoordinate>
<geoCoordinate id="EE5D28AFFFB8585EFEEB0436FE04FC95" box="[279,434,798,825]" degrees="122" direction="east" minutes="43.695" orientation="longitude" pageId="1" pageNumber="315" precision="1" value="122.72825">122°43.695'E</geoCoordinate>
, 
<collectorName id="269C2BBEFFB8585EFE400437FD30FC95" box="[444,646,799,825]" pageId="1" pageNumber="315">Sunda-Expedition</collectorName>
1909/1910, coll. 
<collectorName id="269C2BBEFFB8585EFCB90437FC15FC95" box="[837,931,799,825]" pageId="1" pageNumber="315">J. Elbert</collectorName>
, 
<date id="FFD768A8FFB8585EFC510437FBECFC95" box="[941,1114,799,825]" pageId="1" pageNumber="315" value="1908-08-10">
<collectingDate id="EF939140FFB8585EFC510437FBECFC95" box="[941,1114,799,825]" pageId="1" pageNumber="315" value="1908-08-10">10August 1908</collectingDate>
</date>
[identified as 
<taxonomicName id="4C6935EBFFB8585EFB0A0437FE31FCF1" authority="Heller, 1865" authorityName="Heller" authorityYear="1865" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="315" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B91D927AFFB8585EFB0A0437FF58FCF1" italics="true" pageId="1" pageNumber="315">Ptychognathus pusillus</emphasis>
<bibRefCitation id="EFF83399FFB8585EFF0A046BFE31FCF1" author="Heller, C." box="[246,391,834,861]" pageId="1" pageNumber="315" pagination="1 - 280" refId="ref4649" refString="Heller, C. (1865) Crustaceen. Reise der Osterreichischen Fregatte Novarra um die Erdre, in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wullerstorf-Urbair, Zoologischer Theil, Wein, 2 (3), 1 - 280, pls. 1 - 25." type="book chapter" year="1865">Heller, 1865</bibRefCitation>
</taxonomicName>
]
</materialsCitation>
. 
<materialsCitation id="3B014435FFB8585EFE64046BFC83FCF1" box="[408,821,834,861]" pageId="1" pageNumber="315">
<emphasis id="B91D927AFFB8585EFE64046BFDBCFCF0" box="[408,522,835,860]" italics="true" pageId="1" pageNumber="315">
<typeStatus id="54D2F0CAFFB8585EFE64046BFDBCFCF0" box="[408,522,835,860]" pageId="1" pageNumber="315" type="paratype">Paratypes</typeStatus>
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: 
<specimenCount id="9D6F85E1FFB8585EFDE5046BFDD2FCF1" box="[537,612,835,861]" count="1" pageId="1" pageNumber="315" type="male">1 male</specimenCount>
(19.5 × 
<quantity id="4C91E38DFFB8585EFD39046BFC9BFCF1" box="[709,813,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="1" pageNumber="315" unit="mm" value="14.9">14.9 mm</quantity>
)
</materialsCitation>
, 
<materialsCitation id="3B014435FFB8585EFCBC046BFE56FC2D" collectingDate="1908-08-10" collectionCode="ZRC, SMF" collectorName="Sunda-Expedition &amp; J. Elbert" latitude="-4.82945" location="Muna" longLatPrecision="1" longitude="122.72825" municipality="Moena Raha" pageId="1" pageNumber="315" specimenCode="ZRC 2023.0014, SMF 5257" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
<specimenCount id="9D6F85E1FFB8585EFCBC046BFC17FCF1" box="[832,929,835,861]" count="1" pageId="1" pageNumber="315" type="female">1 female</specimenCount>
(17.6 × 
<quantity id="4C91E38DFFB8585EFBFE046BFBDCFCF1" box="[1026,1130,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="1" pageNumber="315" unit="mm" value="13.9">13.9 mm</quantity>
), 
<specimenCode id="DBCFE613FFB8585EFB81046BFA84FCF1" box="[1149,1330,835,861]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="315">ZRC 2023.0014</specimenCode>
(ex 
<specimenCode id="DBCFE613FFB8585EFA98046BFF65FC2D" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="315">SMF 5257</specimenCode>
), same data as holotype
</materialsCitation>
; 
<materialsCitation id="3B014435FFB8585EFE16044FFCB5FC2D" box="[490,771,871,897]" pageId="1" pageNumber="315">
<specimenCount id="9D6F85E1FFB8585EFE16044FFD83FC2D" box="[490,565,871,897]" count="1" pageId="1" pageNumber="315" type="male">1 male</specimenCount>
(22.9 × 
<quantity id="4C91E38DFFB8585EFD6F044FFD4DFC2D" box="[659,763,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="1" pageNumber="315" unit="mm" value="17.7">17.7 mm</quantity>
)
</materialsCitation>
, 
<materialsCitation id="3B014435FFB8585EFCF1044FFE23FC09" collectingDate="1908-08-10" collectionCode="SMF" collectorName="Sunda-Expedition &amp; J. Elbert" latitude="-4.82945" location="Muna" longLatPrecision="1" longitude="122.72825" municipality="Moena Raha" pageId="1" pageNumber="315" specimenCode="SMF 5257" specimenCount="2" specimenCount-female="2" stateProvince="Sulawesi Utara" typeStatus="paratype">
<specimenCount id="9D6F85E1FFB8585EFCF1044FFCCFFC2D" box="[781,889,871,897]" count="2" pageId="1" pageNumber="315" type="female">2 females</specimenCount>
(16.6 × 
<quantity id="4C91E38DFFB8585EFC2B044FFB8BFC2D" box="[983,1085,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="1" pageNumber="315" unit="mm" value="13.1">13.1 mm</quantity>
, 19.8 × 
<quantity id="4C91E38DFFB8585EFB6A044EFB48FC2D" box="[1174,1278,870,897]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="1" pageNumber="315" unit="mm" value="15.6">15.6 mm</quantity>
), 
<specimenCode id="DBCFE613FFB8585EFAED044FFA3CFC2D" box="[1297,1418,870,897]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="315">SMF 5257</specimenCode>
b, same data as holotype
</materialsCitation>
; 
<materialsCitation id="3B014435FFB8585EFE5D04A3FBE0FC09" box="[417,1110,906,933]" pageId="1" pageNumber="315">
<specimenCount id="9D6F85E1FFB8585EFE5D04A3FDB0FC09" box="[417,518,907,933]" count="13" pageId="1" pageNumber="315" type="male">13 males</specimenCount>
(smallest 15.9 × 
<quantity id="4C91E38DFFB8585EFD3204A3FC83FC09" box="[718,821,907,933]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="1" pageNumber="315" unit="mm" value="12.6">12.6 mm</quantity>
, largest 23.6 × 
<quantity id="4C91E38DFFB8585EFC1904A3FBF8FC09" box="[997,1102,907,933]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="1" pageNumber="315" unit="mm" value="18.4">18.4 mm</quantity>
)
</materialsCitation>
, 
<materialsCitation id="3B014435FFB8585EFB9E04A3FC06FC65" collectingDate="1908-08-10" collectionCode="SMF" collectorName="Sunda-Expedition &amp; J. Elbert" latitude="-4.82945" location="Muna" longLatPrecision="1" longitude="122.72825" municipality="Moena Raha" pageId="1" pageNumber="315" specimenCode="SMF 5257" specimenCount="19" specimenCount-female="19" stateProvince="Sulawesi Utara" typeStatus="paratype">
<specimenCount id="9D6F85E1FFB8585EFB9E04A3FB68FC09" box="[1122,1246,907,933]" count="19" pageId="1" pageNumber="315" type="female">19 females</specimenCount>
(smallest 15.9 × 
<quantity id="4C91E38DFFB8585EFF6B0487FF4BFC65" box="[151,253,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="1" pageNumber="315" unit="mm" value="12.8">12.8 mm</quantity>
, largest 21.6 × 
<quantity id="4C91E38DFFB8585EFE550487FDA7FC65" box="[425,529,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="1.71" pageId="1" pageNumber="315" unit="mm" value="17.1">17.1 mm</quantity>
), 
<specimenCode id="DBCFE613FFB8585EFDD80487FD28FC65" box="[548,670,942,969]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="315">SMF 5257</specimenCode>
c, same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BD64E68FFB8585EFF3B04FAFBF7FBBD" blockId="1.[151,1437,762,2013]" pageId="1" pageNumber="315">
<emphasis id="B91D927AFFB8585EFF3B04FAFE43FC40" bold="true" box="[199,501,978,1004]" pageId="1" pageNumber="315">Other material examined</emphasis>
. 
<materialsCitation id="3B014435FFB8585EFDFF04FBFB88FBBD" collectingDate="2007-09-15" collectionCode="ZRC" collectorName="F. Brehner" location="Muna Island" municipality="Gua La Ode Panu" pageId="1" pageNumber="315" specimenCode="ZRC 2023.0015" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara">
<specimenCount id="9D6F85E1FFB8585EFDFF04FBFDF9FC41" box="[515,591,979,1005]" count="1" pageId="1" pageNumber="315" type="male">1 male</specimenCount>
(22.0 × 
<quantity id="4C91E38DFFB8585EFD5304FBFCA1FC41" box="[687,791,979,1005]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="1" pageNumber="315" unit="mm" value="16.2">16.2 mm</quantity>
), 
<specimenCode id="DBCFE613FFB8585EFCD604FBFC54FC41" box="[810,994,978,1005]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="315">ZRC 2023.0015</specimenCode>
, 
<collectingMunicipality id="6BB2D412FFB8585EFC1204FBFB0FFC41" box="[1006,1209,979,1005]" pageId="1" pageNumber="315">Gua La Ode Panu</collectingMunicipality>
, Walenkabola-nuna, 
<location id="8EB618B3FFB8585EFF6B04DFFE91FBBD" LSID="urn:lsid:plazi:treatment:03C0FF7EFFB85856FF6B05A7FD89FB7A:8EB618B3FFB8585EFF6B04DFFE91FBBD" box="[151,295,1015,1041]" municipality="Gua La Ode Panu" name="Muna Island" pageId="1" pageNumber="315" stateProvince="Sulawesi Utara">Muna Island</location>
, Pulau Buton, southern 
<collectingRegion id="49AD808AFFB8585EFDCF04DFFD2DFBBD" box="[563,667,1015,1041]" country="Indonesia" name="Sulawesi Utara" pageId="1" pageNumber="315">Sulawesi</collectingRegion>
, coll. 
<collectorName id="269C2BBEFFB8585EFD2204DFFCE3FBBD" box="[734,853,1015,1041]" pageId="1" pageNumber="315">F. Brehner</collectorName>
, 
<date id="FFD768A8FFB8585EFC9C04DEFB88FBBD" box="[864,1086,1014,1041]" pageId="1" pageNumber="315" value="2007-09-15">
<collectingDate id="EF939140FFB8585EFC9C04DEFB88FBBD" box="[864,1086,1014,1041]" pageId="1" pageNumber="315" value="2007-09-15">15 September 2007</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD64E68FFB8585EFF3B0332FDF5FB69" blockId="1.[151,1437,762,2013]" pageId="1" pageNumber="315">
<emphasis id="B91D927AFFB8585EFF3B0332FE6CFB98" bold="true" box="[199,474,1050,1076]" pageId="1" pageNumber="315">Comparative material.</emphasis>
<taxonomicName id="4C6935EBFFB8585EFE1F0333FBE8FB99" authority="P. K. L. Ng, Guinot &amp; Iliffe, 1996: 1" authorityName="P. K. L. Ng, Guinot &amp; Iliffe" authorityPageNumber="1" authorityYear="1996" box="[483,1118,1051,1077]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="315" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B91D927AFFB8585EFE1F0333FD08FB99" box="[483,702,1051,1077]" italics="true" pageId="1" pageNumber="315">Orcovita gracilipes</emphasis>
<materialsCitation id="3B014435FFB8585EFD3B0333FBE8FB99" box="[711,1118,1051,1077]" collectingDate="1998-02-06" collectionCode="T" collectorName="M. Illife" country="Niue" location="Vaikona Chasma" pageId="1" pageNumber="315" specimenCode="ZRC 1996.0096, 0097" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
P.K.L. 
<bibRefCitation id="EFF83399FFB8585EFCEB0333FBE8FB99" author="Ng, P. K. L. &amp; Guinot, D. &amp; Iliffe, T. M." box="[791,1118,1051,1077]" pageId="1" pageNumber="315" pagination="109 - 134" refId="ref5235" refString="Ng, P. K. L., Guinot, D. &amp; Iliffe, T. M. (1996) Revision of the anchialine varunine crabs of the genus Orcovita Ng &amp; Tomascik, 1994 (Crustacea: Decapoda: Brachyura: Grapsidae), with descriptions of four new species. Raffles Bulletin of Zoology, 44 (1), 109 - 134." type="journal article" year="1996">
Ng, Guinot &amp; Iliffe, 1996: 
<specimenCount id="9D6F85E1FFB8585EFBAC0333FBE8FB99" box="[1104,1118,1051,1077]" count="1" pageId="1" pageNumber="315" type="male" typeStatus="paratype">1</specimenCount>
</bibRefCitation>
</materialsCitation>
</taxonomicName>
paratype male (15.6 × 
<quantity id="4C91E38DFFB8585EFA900333FF71FBF5" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="1" pageNumber="315" unit="mm" value="11.3">11.3 mm</quantity>
), 
<specimenCount id="9D6F85E1FFB8585EFF250317FE29FBF5" box="[217,415,1087,1113]" count="1" pageId="1" pageNumber="315" type="female" typeStatus="paratype">1 paratype female</specimenCount>
(16.9 × 
<quantity id="4C91E38DFFB8585EFE060316FDD7FBF5" box="[506,609,1086,1113]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="1" pageNumber="315" unit="mm" value="12.5">12.5 mm</quantity>
), 
<specimenCode id="DBCFE613FFB8585EFD8F0317FC9EFBF5" box="[627,808,1087,1113]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="315">ZRC 1996.0096</specimenCode>
– 
<specimenCode id="DBCFE613FFB8585EFCCA0317FCD9FBF5" box="[822,879,1087,1113]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="315">0097</specimenCode>
, station 88-006, 
<location id="8EB618B3FFB8585EFBD80317FB55FBF5" LSID="urn:lsid:plazi:treatment:03C0FF7EFFB85856FF6B05A7FD89FB7A:8EB618B3FFB8585EFBD80317FB55FBF5" box="[1060,1251,1087,1113]" country="Niue" name="Vaikona Chasma" pageId="1" pageNumber="315">Vaikona Chasma</location>
, 
<collectingCountry id="F37E0EF8FFB8585EFB110317FA90FBF5" box="[1261,1318,1087,1113]" name="Niue" pageId="1" pageNumber="315">Niue</collectingCountry>
, coll. 
<collectionCode id="ED78D6ADFFB8585EFA9A0317FACFFBF5" box="[1382,1401,1087,1113]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="315">T</collectionCode>
. 
<collectorName id="269C2BBEFFB8585EFA810317FF66FBD1" pageId="1" pageNumber="315">M. Illife</collectorName>
, 
<date id="FFD768A8FFB8585EFF21034BFE2BFBD1" box="[221,413,1123,1149]" pageId="1" pageNumber="315" value="1998-02-06">
<collectingDate id="EF939140FFB8585EFF21034BFE2BFBD1" box="[221,413,1123,1149]" pageId="1" pageNumber="315" value="1998-02-06">6 February 1998</collectingDate>
</date>
, 
<materialsCitation id="3B014435FFB8585EFE56034BFD89FB69" collectingDate="1993-08-18" collectionCode="ZRC" collectorName="Loyalty" country="Niue" county="Loyalty Islands" elevation="13" location="Luengoni Cave" municipality="Lifou Island" pageId="1" pageNumber="315" specimenCode="ZRC 2001.1149" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
with plastic bottle baited with fish and left overnight at 
<elevation id="0044A95BFFB8585EFBD5034BFB19FBD1" box="[1065,1199,1123,1149]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="315" unit="m" value="13.0">
<quantity id="4C91E38DFFB8585EFBD5034BFBD2FBD1" box="[1065,1124,1123,1149]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="315" unit="m" value="13.0">13 m</quantity>
depth
</elevation>
. 
<taxonomicName id="4C6935EBFFB8585EFB47034BFE44FB0D" authority="N. K. Ng &amp; P. K. L. Ng, 2002: 1" authorityName="N. K. Ng &amp; P. K. L. Ng" authorityPageNumber="1" authorityYear="2002" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="315" phylum="Arthropoda" rank="species" species="mcneiceae">
<emphasis id="B91D927AFFB8585EFB47034BFA2AFBD0" box="[1211,1436,1123,1149]" italics="true" pageId="1" pageNumber="315">Orcovita mcneiceae</emphasis>
N.K. Ng &amp; P.K.L. 
<bibRefCitation id="EFF83399FFB8585EFE9003AFFE44FB0D" author="Ng, N. K. &amp; Ng, P. K. L." box="[364,498,1159,1185]" pageId="1" pageNumber="315" pagination="663 - 670" refId="ref5175" refString="Ng, N. K. &amp; Ng, P. K. L. (2002) Orcovita mcneiceae, a new species of anchialine crab (Brachyura, Grapsidae, Varuninae) from the Loyalty Islands. Crustaceana, 75 (5), 663 - 670. https: // doi. org / 10.1163 / 156854002760202660" type="journal article" year="2002">
Ng, 2002: 
<specimenCount id="9D6F85E1FFB8585EFE1803AFFE44FB0D" box="[484,498,1159,1185]" count="1" pageId="1" pageNumber="315" type="female" typeStatus="paratype">1</specimenCount>
</bibRefCitation>
</taxonomicName>
paratype female (13.5 × 
<quantity id="4C91E38DFFB8585EFCEC03AFFCCFFB0D" box="[784,889,1159,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="1" pageNumber="315" unit="mm" value="10.2">10.2 mm</quantity>
), 
<specimenCode id="DBCFE613FFB8585EFC7003AFFBF2FB0D" box="[908,1092,1159,1185]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="315">ZRC 2001.1149</specimenCode>
, 
<location id="8EB618B3FFB8585EFBB303AFFB48FB0D" LSID="urn:lsid:plazi:treatment:03C0FF7EFFB85856FF6B05A7FD89FB7A:8EB618B3FFB8585EFBB303AFFB48FB0D" box="[1103,1278,1159,1185]" country="Niue" county="Loyalty Islands" municipality="Lifou Island" name="Luengoni Cave" pageId="1" pageNumber="315">Luengoni Cave</location>
, 
<collectingMunicipality id="6BB2D412FFB8585EFAF703AFFA2EFB0D" box="[1291,1432,1159,1185]" pageId="1" pageNumber="315">Lifou Island</collectingMunicipality>
, 
<collectingCounty id="62B736E4FFB8585EFF6B0383FEF1FB69" box="[151,327,1195,1221]" pageId="1" pageNumber="315">
<collectorName id="269C2BBEFFB8585EFF6B0383FF58FB69" box="[151,238,1195,1221]" pageId="1" pageNumber="315">Loyalty</collectorName>
Islands
</collectingCounty>
, coll. 
<date id="FFD768A8FFB8585EFE770383FD89FB69" box="[395,575,1195,1221]" pageId="1" pageNumber="315" value="1993-08-18">
<collectingDate id="EF939140FFB8585EFE770383FD89FB69" box="[395,575,1195,1221]" pageId="1" pageNumber="315" value="1993-08-18">18 August 1993</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3731DE3FFB8585EFF3B03E6FE6CF871" pageId="1" pageNumber="315" type="diagnosis">
<paragraph id="8BD64E68FFB8585EFF3B03E6FE6CF871" blockId="1.[151,1437,762,2013]" pageId="1" pageNumber="315">
<emphasis id="B91D927AFFB8585EFF3B03E6FEF4FB44" bold="true" box="[199,322,1230,1256]" pageId="1" pageNumber="315">Diagnosis.</emphasis>
Carapace subrectangular, distinctly broader than long, CW 1.27–1.36 CL (mean 1.30, n = 7) (
<figureCitation id="135252EDFFB8585EFA9803E7FF6AFAA1" captionStart="FIGURE 1" captionStartId="2.[151,250,1935,1959]" captionTargetBox="[151,1435,181,1911]" captionTargetId="figure-17@2.[151,1435,181,1911]" captionTargetPageId="2" captionText="FIGURE 1. Orcovita varungan n. sp., overall dorsal habitus. A, holotype male (22.9 × 17.6 mm) (SMF 5257a), Sulawesi; B, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; C, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi." figureDoi="http://doi.org/10.5281/zenodo.12681794" httpUri="https://zenodo.org/record/12681794/files/figure.png" pageId="1" pageNumber="315">Figs. 1A–C</figureCitation>
, 
<figureCitation id="135252EDFFB8585EFF1B03DBFE99FAA1" box="[231,303,1267,1293]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="1" pageNumber="315">2A–D</figureCitation>
, 
<figureCitation id="135252EDFFB8585EFEC503DAFEC9FAA1" box="[313,383,1266,1293]" captionStart="FIGURE 5" captionStartId="6.[151,250,1357,1381]" captionTargetBox="[151,1435,181,1333]" captionTargetId="figure-17@6.[151,1435,181,1333]" captionTargetPageId="6" captionText="FIGURE 5. Orcovita varungan n. sp., A, B, E, F, paratype female (17.6 × 13.9 mm), ZRC 2023.0014, Sulawesi; C, D, paratype female (19.8 × 15.6 mm), SMF 5257b, Sulawesi. A, C, dorsal view of carapace; B, outer view of left chela; D, outer view of right chela; E, female anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681804" httpUri="https://zenodo.org/record/12681804/files/figure.png" pageId="1" pageNumber="315">5A, C</figureCitation>
); dorsal surface slightly convex longitudinally, transversely, relatively smooth (
<figureCitation id="135252EDFFB8585EFB0F03DBFADAFAA1" box="[1267,1388,1267,1293]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="1" pageNumber="315">Fig. 2E–H</figureCitation>
), Hshaped gastric groove distinct, other cristae and grooves undiscernible (
<figureCitation id="135252EDFFB8585EFC41023FFBF3FA9D" box="[957,1093,1303,1329]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="1" pageNumber="315">Figs. 2A–D</figureCitation>
, 
<figureCitation id="135252EDFFB8585EFBAC023EFB21FA9D" box="[1104,1175,1302,1329]" captionStart="FIGURE 5" captionStartId="6.[151,250,1357,1381]" captionTargetBox="[151,1435,181,1333]" captionTargetId="figure-17@6.[151,1435,181,1333]" captionTargetPageId="6" captionText="FIGURE 5. Orcovita varungan n. sp., A, B, E, F, paratype female (17.6 × 13.9 mm), ZRC 2023.0014, Sulawesi; C, D, paratype female (19.8 × 15.6 mm), SMF 5257b, Sulawesi. A, C, dorsal view of carapace; B, outer view of left chela; D, outer view of right chela; E, female anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681804" httpUri="https://zenodo.org/record/12681804/files/figure.png" pageId="1" pageNumber="315">5A, C</figureCitation>
). External orbital tooth very broad; anterolateral margin convex, lined with granules, with one low epibranchial tooth; posterolateral margins convergent posteriorly, mesobranchial region sloping posterolaterally (
<figureCitation id="135252EDFFB8585EFC490277FB8AFAD5" box="[949,1084,1375,1401]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="1" pageNumber="315">Figs. 2A–D</figureCitation>
, 
<figureCitation id="135252EDFFB8585EFBB40276FB39FAD5" box="[1096,1167,1374,1401]" captionStart="FIGURE 5" captionStartId="6.[151,250,1357,1381]" captionTargetBox="[151,1435,181,1333]" captionTargetId="figure-17@6.[151,1435,181,1333]" captionTargetPageId="6" captionText="FIGURE 5. Orcovita varungan n. sp., A, B, E, F, paratype female (17.6 × 13.9 mm), ZRC 2023.0014, Sulawesi; C, D, paratype female (19.8 × 15.6 mm), SMF 5257b, Sulawesi. A, C, dorsal view of carapace; B, outer view of left chela; D, outer view of right chela; E, female anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681804" httpUri="https://zenodo.org/record/12681804/files/figure.png" pageId="1" pageNumber="315">5A, C</figureCitation>
). Front truncate, frontal margin slightly sinuous, subfrontal triangular facet (low vertical area between frontal margin and superior part of antennular septum) present (
<figureCitation id="135252EDFFB8585EFE2E028FFDFDFA6D" box="[466,587,1447,1473]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="1" pageNumber="315">Fig. 2E–H</figureCitation>
). Suborbital margin with 19–21 rounded granules, mesial 10–12 larger, more prominent, lateral ones low and indistinct (
<figureCitation id="135252EDFFB8585EFD7A02E3FCB2FA49" box="[646,772,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="1" pageNumber="315">Fig. 2E–H</figureCitation>
). Antennule basal article subtriangular, large (
<figureCitation id="135252EDFFB8585EFAE802E3FA27FA49" box="[1300,1425,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="1" pageNumber="315">Fig. 2E–H</figureCitation>
). Antenna with basal article subquadrate, mobile, flagellum long, reaching beyond orbit and external orbital angle when stretched laterally (
<figureCitation id="135252EDFFB8585EFE4D013BFD9AF981" box="[433,556,1555,1581]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="1" pageNumber="315">Fig. 2E–H</figureCitation>
). Eye cylindrical, cornea distinct, narrower than base of ocular peduncle, distal end of eye reaching lateral fifth of orbit (
<figureCitation id="135252EDFFB8585EFD99011FFD57F9FD" box="[613,737,1591,1617]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="1" pageNumber="315">Fig. 2E–H</figureCitation>
). Third maxilliped subrectangular, leaving wide gape between endopods and between distal margin of merus and posterior margin of epistome; ischium narrow, distally concave; merus widened distally, distolaterally auriculate; exopod stout, wider than distal width of ischium (
<figureCitation id="135252EDFFB8585EFB210157FAE6F935" box="[1245,1360,1663,1689]" captionStart="FIGURE 3" captionStartId="4.[151,250,1658,1682]" captionTargetBox="[158,1430,489,1634]" captionTargetId="figure-182@4.[158,1430,489,1634]" captionTargetPageId="4" captionText="FIGURE 3. Orcovita varungan n. sp., A, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; B, C, D, E, G, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, outer view of right chela; B, outer view of left chela; C, view of pulvinus at base of fingers of left chela; D, dorsal view of left cheliped; E, right P4 and P5; F, G, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.12681798" httpUri="https://zenodo.org/record/12681798/files/figure.png" pageId="1" pageNumber="315">Fig. 3F, G</figureCitation>
). Male chelipeds relatively long, symmetrical, entire surface covered with microscopic granules, immovable finger with tuft of soft setae along proximal two-fifths of occlusal margins (
<figureCitation id="135252EDFFB8585EFCA301EFFC54F94D" box="[863,994,1735,1761]" captionStart="FIGURE 1" captionStartId="2.[151,250,1935,1959]" captionTargetBox="[151,1435,181,1911]" captionTargetId="figure-17@2.[151,1435,181,1911]" captionTargetPageId="2" captionText="FIGURE 1. Orcovita varungan n. sp., overall dorsal habitus. A, holotype male (22.9 × 17.6 mm) (SMF 5257a), Sulawesi; B, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; C, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi." figureDoi="http://doi.org/10.5281/zenodo.12681794" httpUri="https://zenodo.org/record/12681794/files/figure.png" pageId="1" pageNumber="315">Figs. 1A, C</figureCitation>
, 
<figureCitation id="135252EDFFB8585EFC1101EFFB81F94D" box="[1005,1079,1735,1761]" captionStart="FIGURE 3" captionStartId="4.[151,250,1658,1682]" captionTargetBox="[158,1430,489,1634]" captionTargetId="figure-182@4.[158,1430,489,1634]" captionTargetPageId="4" captionText="FIGURE 3. Orcovita varungan n. sp., A, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; B, C, D, E, G, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, outer view of right chela; B, outer view of left chela; C, view of pulvinus at base of fingers of left chela; D, dorsal view of left cheliped; E, right P4 and P5; F, G, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.12681798" httpUri="https://zenodo.org/record/12681798/files/figure.png" pageId="1" pageNumber="315">3A–D</figureCitation>
). Female chelipeds symmetrical or almost so, chelae with 1 row of granules from tip of immovable finger to base of palm, no tuft of setae present (
<figureCitation id="135252EDFFB8585EFF630027FEADF885" box="[159,283,1806,1833]" captionStart="FIGURE 5" captionStartId="6.[151,250,1357,1381]" captionTargetBox="[151,1435,181,1333]" captionTargetId="figure-17@6.[151,1435,181,1333]" captionTargetPageId="6" captionText="FIGURE 5. Orcovita varungan n. sp., A, B, E, F, paratype female (17.6 × 13.9 mm), ZRC 2023.0014, Sulawesi; C, D, paratype female (19.8 × 15.6 mm), SMF 5257b, Sulawesi. A, C, dorsal view of carapace; B, outer view of left chela; D, outer view of right chela; E, female anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681804" httpUri="https://zenodo.org/record/12681804/files/figure.png" pageId="1" pageNumber="315">Fig. 5A–E</figureCitation>
). Ambulatory legs subcylindrical in cross-section, articles long and slender (
<figureCitation id="135252EDFFB8585EFB8B0027FB4AF885" box="[1143,1276,1807,1833]" captionStart="FIGURE 1" captionStartId="2.[151,250,1935,1959]" captionTargetBox="[151,1435,181,1911]" captionTargetId="figure-17@2.[151,1435,181,1911]" captionTargetPageId="2" captionText="FIGURE 1. Orcovita varungan n. sp., overall dorsal habitus. A, holotype male (22.9 × 17.6 mm) (SMF 5257a), Sulawesi; B, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; C, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi." figureDoi="http://doi.org/10.5281/zenodo.12681794" httpUri="https://zenodo.org/record/12681794/files/figure.png" pageId="1" pageNumber="315">Figs. 1A–C</figureCitation>
, 
<figureCitation id="135252EDFFB8585EFAF40027FA9CF885" box="[1288,1322,1807,1833]" captionStart="FIGURE 3" captionStartId="4.[151,250,1658,1682]" captionTargetBox="[158,1430,489,1634]" captionTargetId="figure-182@4.[158,1430,489,1634]" captionTargetPageId="4" captionText="FIGURE 3. Orcovita varungan n. sp., A, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; B, C, D, E, G, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, outer view of right chela; B, outer view of left chela; C, view of pulvinus at base of fingers of left chela; D, dorsal view of left cheliped; E, right P4 and P5; F, G, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.12681798" httpUri="https://zenodo.org/record/12681798/files/figure.png" pageId="1" pageNumber="315">3E</figureCitation>
), ratios of P4, P5 articles as in 
<tableCitation id="C6EB7BD3FFB8585EFE86001BFE78F8E1" box="[378,462,1843,1869]" captionStart="TABLE 1" captionStartId="9.[151,239,151,175]" captionTargetPageId="9" captionText="TABLE 1. Length to width ratios of P4 and P5 merus, propodus and dactylus of Orcovita varungan n. sp." httpUri="http://table.plazi.org/id/DF161EE0FFB05856FF6B07BFFAB5FF1C" pageId="1" pageNumber="315" tableUuid="DF161EE0FFB05856FF6B07BFFAB5FF1C">Table 1</tableCitation>
. Male pleon narrow, lateral margins of somites 3–6 (except both proximal, distal ends) forming distally convergent straight lines; telson lateral margins parallel, distally rounded, longer than somite 6 (
<figureCitation id="135252EDFFB8585EFF630053FEAFF839" box="[159,281,1915,1941]" captionStart="FIGURE 4" captionStartId="5.[151,250,1893,1917]" captionTargetBox="[151,1435,345,1870]" captionTargetId="figure-109@5.[151,1435,345,1870]" captionTargetPageId="5" captionText="FIGURE 4. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, D, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; E, F, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, B, male anterior thoracic sternum and pleon; C–E, male thoracic sternum and pleon; F, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681800" httpUri="https://zenodo.org/record/12681800/files/figure.png" pageId="1" pageNumber="315">Fig. 4A–F</figureCitation>
). G1 relatively short, stout, distally with corneous beak directed obliquely proximo-dorsolaterally (
<figureCitation id="135252EDFFB8585EFA930053FE9AF815" captionStart="FIGURE 6" captionStartId="7.[151,250,1851,1875]" captionTargetBox="[177,1411,501,1810]" captionTargetId="figure-164@7.[151,1435,489,1827]" captionTargetPageId="7" captionText="FIGURE 6. Gonopods of Orcovita varungan n. sp., holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi. A–D, with setae, before denuding; E–O, setae denuded. A–G, left G1; H–M, distal part of left G1 with corneous distal part; N, O. left G2. A, E, H, N, ventral view; B, I, subventral view; C, J, outer mesial view; D, G, K, O, dorsal view; L, subdorsal view; F, M, inner mesial view. Scales: E–G, N, O = 0.5 mm; H–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12681806" httpUri="https://zenodo.org/record/12681806/files/figure.png" pageId="1" pageNumber="315">Fig. 6A–G, H–M</figureCitation>
). Vulvae small, positioned on proximal third of thoracic sternite 6, operculum rounded, prominent, slightly protruding (
<figureCitation id="135252EDFFB8585EFE8400EBFE7BF871" box="[376,461,1986,2013]" captionStart="FIGURE 5" captionStartId="6.[151,250,1357,1381]" captionTargetBox="[151,1435,181,1333]" captionTargetId="figure-17@6.[151,1435,181,1333]" captionTargetPageId="6" captionText="FIGURE 5. Orcovita varungan n. sp., A, B, E, F, paratype female (17.6 × 13.9 mm), ZRC 2023.0014, Sulawesi; C, D, paratype female (19.8 × 15.6 mm), SMF 5257b, Sulawesi. A, C, dorsal view of carapace; B, outer view of left chela; D, outer view of right chela; E, female anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681804" httpUri="https://zenodo.org/record/12681804/files/figure.png" pageId="1" pageNumber="315">Fig. 5F</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF161EE0FFBB585DFF6B00A7FA9DF867" ID-DOI="http://doi.org/10.5281/zenodo.12681794" ID-Zenodo-Dep="12681794" httpUri="https://zenodo.org/record/12681794/files/figure.png" pageId="2" pageNumber="316" startId="2.[151,250,1935,1959]" targetBox="[151,1435,181,1911]" targetPageId="2" targetType="figure">
<paragraph id="8BD64E68FFBB585DFF6B00A7FA9DF867" blockId="2.[151,1437,1935,1996]" pageId="2" pageNumber="316">
<emphasis id="B91D927AFFBB585DFF6B00A7FEB8F80B" bold="true" box="[151,270,1935,1959]" pageId="2" pageNumber="316">FIGURE 1</emphasis>
. 
<taxonomicName id="4C6935EBFFBB585DFEE700B8FE6AF80B" authorityName="Ng &amp; Naruse" authorityYear="2024" box="[283,476,1936,1959]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="316" phylum="Arthropoda" rank="species" species="varungan" status="sp. nov.">
<emphasis id="B91D927AFFBB585DFEE700B8FE6AF80B" box="[283,476,1936,1959]" italics="true" pageId="2" pageNumber="316">Orcovita varungan</emphasis>
</taxonomicName>
<emphasis id="B91D927AFFBB585DFE1F00B8FDABF804" bold="true" box="[483,541,1936,1960]" pageId="2" pageNumber="316">
<taxonomicNameLabel id="A22E2F01FFBB585DFE1F00B8FDABF804" box="[483,541,1936,1960]" pageId="2" pageNumber="316" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, overall dorsal habitus. A, holotype male (22.9 × 17.6 mm) (SMF 5257a), Sulawesi; B, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; C, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi.
</paragraph>
</caption>
<caption id="DF161EE0FFBA585CFF6B0182FB22F8A6" ID-DOI="http://doi.org/10.5281/zenodo.12681796" ID-Zenodo-Dep="12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="3" pageNumber="317" startId="3.[151,250,1706,1730]" targetBox="[151,1435,181,1682]" targetPageId="3" targetType="figure">
<paragraph id="8BD64E68FFBA585CFF6B0182FB22F8A6" blockId="3.[151,1436,1706,1803]" pageId="3" pageNumber="317">
<emphasis id="B91D927AFFBA585CFF6B0182FEBAF96E" bold="true" box="[151,268,1706,1730]" pageId="3" pageNumber="317">FIGURE 2</emphasis>
. 
<taxonomicName id="4C6935EBFFBA585CFEEB0183FE60F96E" authorityName="Ng &amp; Naruse" authorityYear="2024" box="[279,470,1707,1730]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="317" phylum="Arthropoda" rank="species" species="varungan" status="sp. nov.">
<emphasis id="B91D927AFFBA585CFEEB0183FE60F96E" box="[279,470,1707,1730]" italics="true" pageId="3" pageNumber="317">Orcovita varungan</emphasis>
</taxonomicName>
<emphasis id="B91D927AFFBA585CFE200182FDA2F96E" bold="true" box="[476,532,1706,1730]" pageId="3" pageNumber="317">
<taxonomicNameLabel id="A22E2F01FFBA585CFE200182FDA2F96E" box="[476,532,1706,1730]" pageId="3" pageNumber="317" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax.
</paragraph>
</caption>
<subSubSection id="C3731DE3FFBA585BFF3B0010FA80FEC9" lastPageId="4" lastPageNumber="318" pageId="3" pageNumber="317" type="description">
<paragraph id="8BD64E68FFBA585BFF3B0010FA80FEC9" blockId="3.[151,1437,1848,2019]" lastBlockId="4.[151,1437,150,465]" lastPageId="4" lastPageNumber="318" pageId="3" pageNumber="317">
<emphasis id="B91D927AFFBA585CFF3B0010FE8EF8FE" bold="true" box="[199,312,1848,1874]" pageId="3" pageNumber="317">Variation</emphasis>
. The carapace shape varies slightly, most specimens have the posterolateral margin gently converging towards the posterior carapace margin (
<figureCitation id="135252EDFFBA585CFDAE0074FD7CF8DB" box="[594,714,1884,1911]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="3" pageNumber="317">Fig. 2A–C</figureCitation>
) but in the most recent specimen (ZRC 2023.0015), the margins are more convergent making the carapace appear more hexagonal (
<figureCitation id="135252EDFFBA585CFC7200A8FC5CF837" box="[910,1002,1920,1947]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="3" pageNumber="317">Fig. 2D</figureCitation>
). The dorsal surface of the carapace is gently convex in the type series (
<figureCitation id="135252EDFFBA585CFDEA008CFDDBF813" box="[534,621,1956,1983]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="3" pageNumber="317">Fig. 2E</figureCitation>
) but in the most recent specimen (ZRC 2023.0015), it is slightly more convex, especially on the branchial regions (
<figureCitation id="135252EDFFBA585CFD6400E0FD43F84F" box="[664,757,1992,2019]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="3" pageNumber="317">Fig. 2G</figureCitation>
). While the external orbital tooth is always very broad and clearly separated from the rest of the anterolateral margin by a distinct cleft (
<figureCitation id="135252EDFFBD585BFC1707BFFB26FF1D" box="[1003,1168,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="4" pageNumber="318">Figs. 2A, B, D</figureCitation>
, 
<figureCitation id="135252EDFFBD585BFB6707BEFB69FF1D" box="[1179,1247,150,177]" captionStart="FIGURE 5" captionStartId="6.[151,250,1357,1381]" captionTargetBox="[151,1435,181,1333]" captionTargetId="figure-17@6.[151,1435,181,1333]" captionTargetPageId="6" captionText="FIGURE 5. Orcovita varungan n. sp., A, B, E, F, paratype female (17.6 × 13.9 mm), ZRC 2023.0014, Sulawesi; C, D, paratype female (19.8 × 15.6 mm), SMF 5257b, Sulawesi. A, C, dorsal view of carapace; B, outer view of left chela; D, outer view of right chela; E, female anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681804" httpUri="https://zenodo.org/record/12681804/files/figure.png" pageId="4" pageNumber="318">5A, C</figureCitation>
), the cleft can be shallow in some specimens and may even be absent, with the anterolateral margin appearing entire (
<figureCitation id="135252EDFFBD585BFAFF0793FAEBFF79" box="[1283,1373,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="4" pageNumber="318">Fig. 2C</figureCitation>
). The proportions of male pleonal somite 6 also vary slightly in width, being narrower in the 
<typeStatus id="54D2F0CAFFBD585BFB9F07F7FB73FF55" box="[1123,1221,223,249]" pageId="4" pageNumber="318" type="holotype">holotype</typeStatus>
male (
<figureCitation id="135252EDFFBD585BFAEE07F7FADFFF55" box="[1298,1385,223,249]" captionStart="FIGURE 4" captionStartId="5.[151,250,1893,1917]" captionTargetBox="[151,1435,345,1870]" captionTargetId="figure-109@5.[151,1435,345,1870]" captionTargetPageId="5" captionText="FIGURE 4. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, D, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; E, F, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, B, male anterior thoracic sternum and pleon; C–E, male thoracic sternum and pleon; F, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681800" httpUri="https://zenodo.org/record/12681800/files/figure.png" pageId="4" pageNumber="318">Fig. 4A</figureCitation>
) but slightly wider in the 
<typeStatus id="54D2F0CAFFBD585BFE79062BFE46FEB1" box="[389,496,259,285]" pageId="4" pageNumber="318" type="paratype">paratypes</typeStatus>
and other specimen (
<figureCitation id="135252EDFFBD585BFD1B062BFC32FEB1" box="[743,900,259,285]" captionStart="FIGURE 4" captionStartId="5.[151,250,1893,1917]" captionTargetBox="[151,1435,345,1870]" captionTargetId="figure-109@5.[151,1435,345,1870]" captionTargetPageId="5" captionText="FIGURE 4. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, D, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; E, F, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, B, male anterior thoracic sternum and pleon; C–E, male thoracic sternum and pleon; F, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681800" httpUri="https://zenodo.org/record/12681800/files/figure.png" pageId="4" pageNumber="318">Fig. 4B, D, F</figureCitation>
). The granules lining the suborbital margin are relatively smaller in smaller males and females, with the lateral granules very low and sometimes barely visible. In larger males (e.g., the 
<typeStatus id="54D2F0CAFFBD585BFE6C0663FE44FEC9" box="[400,498,331,357]" pageId="4" pageNumber="318" type="holotype">holotype</typeStatus>
male), these granules are clearly visible even in the dorsal view (
<figureCitation id="135252EDFFBD585BFB330663FA9FFEC9" box="[1231,1321,331,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="4" pageNumber="318">Fig. 2A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3731DE3FFBD585AFF3B0647FB89FEED" lastPageId="5" lastPageNumber="319" pageId="4" pageNumber="318" type="etymology">
<paragraph id="8BD64E68FFBD585BFF3B0647FDDDFE7D" blockId="4.[151,1437,150,465]" pageId="4" pageNumber="318">
<emphasis id="B91D927AFFBD585BFF3B0647FEFEFE25" bold="true" box="[199,328,367,393]" pageId="4" pageNumber="318">Etymology</emphasis>
. The species name is an arbitrary combination of the family name 
<taxonomicName id="4C6935EBFFBD585BFBC30647FB07FE25" authorityName="H. Milne Edwards" authorityYear="1853" box="[1087,1201,367,393]" class="Malacostraca" family="Varunidae" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="318" phylum="Arthropoda" rank="family">Varunidae</taxonomicName>
with the first part of Ngan Kee’s name, alluding to her expertise and love for this group of crabs that she spent so many years studying. The name is used as a noun in apposition.
</paragraph>
<caption id="DF161EE0FFBD585BFF6B0152FB7DF976" ID-DOI="http://doi.org/10.5281/zenodo.12681798" ID-Zenodo-Dep="12681798" httpUri="https://zenodo.org/record/12681798/files/figure.png" pageId="4" pageNumber="318" startId="4.[151,250,1658,1682]" targetBox="[158,1430,489,1634]" targetPageId="4" targetType="figure">
<paragraph id="8BD64E68FFBD585BFF6B0152FB7DF976" blockId="4.[151,1436,1658,1754]" pageId="4" pageNumber="318">
<emphasis id="B91D927AFFBD585BFF6B0152FEB8F93E" bold="true" box="[151,270,1658,1682]" pageId="4" pageNumber="318">FIGURE 3</emphasis>
. 
<taxonomicName id="4C6935EBFFBD585BFEE70152FE6AF93E" authorityName="Ng &amp; Naruse" authorityYear="2024" box="[283,476,1658,1682]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="318" phylum="Arthropoda" rank="species" species="varungan" status="sp. nov.">
<emphasis id="B91D927AFFBD585BFEE70152FE6AF93E" box="[283,476,1658,1682]" italics="true" pageId="4" pageNumber="318">Orcovita varungan</emphasis>
</taxonomicName>
<emphasis id="B91D927AFFBD585BFE1F0152FDAAF93E" bold="true" box="[483,540,1658,1682]" pageId="4" pageNumber="318">
<taxonomicNameLabel id="A22E2F01FFBD585BFE1F0152FDAAF93E" box="[483,540,1658,1682]" pageId="4" pageNumber="318" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, A, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; B, C, D, E, G, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, outer view of right chela; B, outer view of left chela; C, view of pulvinus at base of fingers of left chela; D, dorsal view of left cheliped; E, right P4 and P5; F, G, left third maxilliped.
</paragraph>
</caption>
<paragraph id="8BD64E68FFBD585AFF3B002AFB89FEED" blockId="4.[151,1437,1794,2037]" lastBlockId="5.[151,1437,151,321]" lastPageId="5" lastPageNumber="319" pageId="4" pageNumber="318">
<emphasis id="B91D927AFFBD585BFF3B002AFE95F8B0" bold="true" box="[199,291,1794,1820]" pageId="4" pageNumber="318">Habitat</emphasis>
. While all species of 
<taxonomicName id="4C6935EBFFBD585BFDD5002BFD3AF8B1" authorityName="P. K. L. Ng &amp; Tomascik" authorityYear="1994" box="[553,652,1795,1821]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="B91D927AFFBD585BFDD5002BFD3AF8B1" box="[553,652,1795,1821]" italics="true" pageId="4" pageNumber="318">Orcovita</emphasis>
</taxonomicName>
are known to have anchialine habits, there was no ecological data associated with the large series of specimens collected by J. Elbert in 1909–1910. The Sunda Expedition (1909– 1910), led by German geographer and geologist Johannes Elbert (1878–1915), explored several parts of 
<collectingRegion id="49AD808AFFBD585BFAC60063FADAF8C9" box="[1338,1388,1867,1893]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="318">Java</collectingRegion>
and 
<collectingRegion id="49AD808AFFBD585BFF6B0047FF41F825" box="[151,247,1903,1929]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="318">Sumatra</collectingRegion>
, the 
<collectingRegion id="49AD808AFFBD585BFED60047FDA5F825" box="[298,531,1903,1929]" country="Indonesia" name="Nusa Tenggara Timur" pageId="4" pageNumber="318">Lesser Sunda Islands</collectingRegion>
(including Lombok and 
<collectingCountry id="F37E0EF8FFBD585BFCDA0047FCC6F825" box="[806,880,1903,1929]" name="Indonesia" pageId="4" pageNumber="318">Flores</collectingCountry>
), and southeastern 
<collectingRegion id="49AD808AFFBD585BFBBE0047FB11F825" box="[1090,1191,1903,1929]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="318">Sulawesi</collectingRegion>
(including the islands of Muna, Buton, Kabaena and Rubia) (cf. 
<bibRefCitation id="EFF83399FFBD585BFD7E00BBFCBFF801" author="Elbert, J." box="[642,777,1939,1965]" pageId="4" pageNumber="318" refId="ref4549" refString="Elbert, J. (1911) Die Sunda-Expedition des Vereins fur Geographie und Statistik zu Frankfurt am Main. Festschrift zur Feier des 75 jahrigen Bestehens des Vereins. Band 1. H. Minjon, Frankfurt am Main, 274 pp., 29 pls." type="book" year="1911">Elbert 1911</bibRefCitation>
, 
<bibRefCitation id="EFF83399FFBD585BFCE400BBFCE2F801" author="Elbert, J." box="[792,852,1939,1965]" pageId="4" pageNumber="318" refId="ref4597" refString="Elbert, J. (1912) Die Sunda-Expedition des Vereins fur Geographie und Statistik zu Frankfurt am Main. Festschrift zur Feier des 75 jahrigen Bestehens des Vereins. Band 2. H. Minjon, Frankfurt am Main, 373 pp., 32 pls., 7 maps." type="book" year="1912">1912</bibRefCitation>
). While the expedition focussed on geography and geology, there were many interesting cultural reports as well, and a fair amount of botanical and zoological material was also collected. While 
<bibRefCitation id="EFF83399FFBD585BFE4600F3FD09F859" author="Elbert, J." box="[442,703,2011,2037]" pageId="4" pageNumber="318" refId="ref4549" refString="Elbert, J. (1911) Die Sunda-Expedition des Vereins fur Geographie und Statistik zu Frankfurt am Main. Festschrift zur Feier des 75 jahrigen Bestehens des Vereins. Band 1. H. Minjon, Frankfurt am Main, 274 pp., 29 pls." type="book" year="1911">Elbert (1911: 143–174)</bibRefCitation>
dedicated an entire chapter to his study of the island of Muna and its occupants, he did not mention the animals he collected on the island. The large number of specimens obtained suggests they may have been collected by traps; normally, they are not easily collected by hand in cave habitats (see 
<bibRefCitation id="EFF83399FFBC585AFF6B07F7FE8AFF55" author="Ng, P. K. L. &amp; Guinot, D. &amp; Iliffe, T. M." box="[151,316,223,249]" pageId="5" pageNumber="319" pagination="109 - 134" refId="ref5235" refString="Ng, P. K. L., Guinot, D. &amp; Iliffe, T. M. (1996) Revision of the anchialine varunine crabs of the genus Orcovita Ng &amp; Tomascik, 1994 (Crustacea: Decapoda: Brachyura: Grapsidae), with descriptions of four new species. Raffles Bulletin of Zoology, 44 (1), 109 - 134." type="journal article" year="1996">
Ng 
<emphasis id="B91D927AFFBC585AFF3D07F7FF4FFF55" box="[193,249,223,249]" italics="true" pageId="5" pageNumber="319">et al.</emphasis>
1996
</bibRefCitation>
; 
<bibRefCitation id="EFF83399FFBC585AFEB407F7FDA0FF55" author="Davie, P. J. F. &amp; Ng, P. K. L." box="[328,534,223,249]" pageId="5" pageNumber="319" pagination="57 - 70" refId="ref4490" refString="Davie, P. J. F. &amp; Ng, P. K. L. (2012) Two new species of Orcovita (Crustacea: Decapoda: Brachyura: Varunidae) from anchialine caves on Christmas Island, eastern Indian Ocean. Raffles Bulletin of Zoology, 60 (1), 57 - 70." type="journal article" year="2012">Davie &amp; Ng 2012</bibRefCitation>
). The most recent specimen (ZRC 2023.0015) was collected inside a submerged anchialine cave (see also 
<bibRefCitation id="EFF83399FFBC585AFE50062BFDCEFEB1" author="Anonymous" box="[428,632,259,285]" pageId="5" pageNumber="319" pagination="1 - 6" refId="ref4457" refString="Anonymous (2022) Seeing the Enchantment of Paradise Cave Diving - ' Oasis of Buton Island' in Southeast Sulawesi. Kendarinfo, Travel, 18 September 2022, 1 - 6." type="book chapter" year="2022">Anonymous 2022</bibRefCitation>
). The cave-adapted hymenosomatid crab 
<taxonomicName id="4C6935EBFFBC585AFBBC062BFEC7FEED" authority="Naruse, P. K. L. Ng &amp; Guinot, 2006" authorityName="Naruse, P. K. L. Ng &amp; Guinot" authorityYear="2006" class="Malacostraca" family="Hymenosomatidae" genus="Sulaplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="319" phylum="Arthropoda" rank="species" species="ensifer">
<emphasis id="B91D927AFFBC585AFBBC062BFB42FEB1" box="[1088,1268,259,285]" italics="true" pageId="5" pageNumber="319">Sulaplax ensifer</emphasis>
Naruse, P.K.L. Ng &amp; Guinot, 2006
</taxonomicName>
is also known from the caves of island (see 
<bibRefCitation id="EFF83399FFBC585AFC9D060FFB85FEED" author="Naruse, T. &amp; Tamura, H." box="[865,1075,295,321]" pageId="5" pageNumber="319" pagination="147 - 151" refId="ref5035" refString="Naruse, T. &amp; Tamura, H. (2006) A first record of anchialine crab of the genus Orcovita Ng and Tomascik, 1994 (Decapoda: Brachyura: Varunidae) from Japan, with description of the species. Limnology, 7, 147 - 151, figs. 1 + 2, table 1. https: // doi. org / 10.1007 / s 10201 - 006 - 0176 - y" type="journal article" year="2006">
Naruse 
<emphasis id="B91D927AFFBC585AFC4B060FFC46FEED" box="[951,1008,295,321]" italics="true" pageId="5" pageNumber="319">et al.</emphasis>
2006
</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF161EE0FFBC585AFF6B004DFEB8F845" ID-DOI="http://doi.org/10.5281/zenodo.12681800" ID-Zenodo-Dep="12681800" httpUri="https://zenodo.org/record/12681800/files/figure.png" pageId="5" pageNumber="319" startId="5.[151,250,1893,1917]" targetBox="[151,1435,345,1870]" targetPageId="5" targetType="figure">
<paragraph id="8BD64E68FFBC585AFF6B004DFEB8F845" blockId="5.[151,1436,1893,2025]" pageId="5" pageNumber="319">
<emphasis id="B91D927AFFBC585AFF6B004DFEB8F8D1" bold="true" box="[151,270,1893,1917]" pageId="5" pageNumber="319">FIGURE 4</emphasis>
. 
<taxonomicName id="4C6935EBFFBC585AFEE0004EFE68F8D1" authorityName="Ng &amp; Naruse" authorityYear="2024" box="[284,478,1894,1917]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="319" phylum="Arthropoda" rank="species" species="varungan" status="sp. nov.">
<emphasis id="B91D927AFFBC585AFEE0004EFE68F8D1" box="[284,478,1894,1917]" italics="true" pageId="5" pageNumber="319">Orcovita varungan</emphasis>
</taxonomicName>
<emphasis id="B91D927AFFBC585AFE1A004EFD96F8D2" bold="true" box="[486,544,1894,1918]" pageId="5" pageNumber="319">
<taxonomicNameLabel id="A22E2F01FFBC585AFE1A004EFD96F8D2" box="[486,544,1894,1918]" pageId="5" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, D, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; E, F, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, B, male anterior thoracic sternum and pleon; C–E, male thoracic sternum and pleon; F, male pleon.
</paragraph>
</caption>
<caption id="DF161EE0FFBF5859FF6B0265FB2CFA02" ID-DOI="http://doi.org/10.5281/zenodo.12681804" ID-Zenodo-Dep="12681804" httpUri="https://zenodo.org/record/12681804/files/figure.png" pageId="6" pageNumber="320" startId="6.[151,250,1357,1381]" targetBox="[151,1435,181,1333]" targetPageId="6" targetType="figure">
<paragraph id="8BD64E68FFBF5859FF6B0265FB2CFA02" blockId="6.[151,1436,1357,1454]" pageId="6" pageNumber="320">
<emphasis id="B91D927AFFBF5859FF6B0265FEBAFAC9" bold="true" box="[151,268,1357,1381]" pageId="6" pageNumber="320">FIGURE 5</emphasis>
. 
<taxonomicName id="4C6935EBFFBF5859FEE40266FE6EFAC9" authorityName="Ng &amp; Naruse" authorityYear="2024" box="[280,472,1358,1381]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="320" phylum="Arthropoda" rank="species" species="varungan" status="sp. nov.">
<emphasis id="B91D927AFFBF5859FEE40266FE6EFAC9" box="[280,472,1358,1381]" italics="true" pageId="6" pageNumber="320">Orcovita varungan</emphasis>
</taxonomicName>
<emphasis id="B91D927AFFBF5859FE220266FDA0FAC9" bold="true" box="[478,534,1358,1381]" pageId="6" pageNumber="320">
<taxonomicNameLabel id="A22E2F01FFBF5859FE220266FDA0FAC9" box="[478,534,1358,1381]" pageId="6" pageNumber="320" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, A, B, E, F, paratype female (17.6 × 13.9 mm), ZRC 2023.0014, Sulawesi; C, D, paratype female (19.8 × 15.6 mm), SMF 5257b, Sulawesi. A, C, dorsal view of carapace; B, outer view of left chela; D, outer view of right chela; E, female anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae.
</paragraph>
</caption>
<subSubSection id="C3731DE3FFBF5856FF3B02F4FD89FB7A" lastPageId="9" lastPageNumber="323" pageId="6" pageNumber="320" type="discussion">
<paragraph id="8BD64E68FFBF5859FF3B02F4FCD3F906" blockId="6.[151,1437,1500,2030]" pageId="6" pageNumber="320">
<emphasis id="B91D927AFFBF5859FF3B02F4FE82FA5A" bold="true" box="[199,308,1500,1526]" pageId="6" pageNumber="320">Remarks</emphasis>
. Davie &amp; P.K.L. Ng (2012: 67) divided 
<taxonomicName id="4C6935EBFFBF5859FD0602F4FCEBFA5A" authorityName="P. K. L. Ng &amp; Tomascik" authorityYear="1994" box="[762,861,1500,1526]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B91D927AFFBF5859FD0602F4FCEBFA5A" box="[762,861,1500,1526]" italics="true" pageId="6" pageNumber="320">Orcovita</emphasis>
</taxonomicName>
into two groups based on the carapace and relative proportions of the ambulatory legs. In their second group, the species have only one epibranchial tooth (excluding the external orbital tooth), the postfrontal cristae are low to undiscernible, the ambulatory legs are relatively longer, and the junction between the male cheliped fingers are usually not setose. In this group, only 
<taxonomicName id="4C6935EBFFBF5859FB390160FAEFF9CE" authorityName="P. K. L. Ng, Guinot &amp; Iliffe" authorityYear="1996" box="[1221,1369,1608,1634]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="320" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B91D927AFFBF5859FB390160FAEFF9CE" box="[1221,1369,1608,1634]" italics="true" pageId="6" pageNumber="320">O. gracilipes</emphasis>
</taxonomicName>
(both male and female) and 
<taxonomicName id="4C6935EBFFBF5859FE6C0144FD9EF92A" authorityName="N. K. Ng &amp; P. K. L. Ng" authorityYear="2002" box="[400,552,1644,1670]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="320" phylum="Arthropoda" rank="species" species="mcneiceae">
<emphasis id="B91D927AFFBF5859FE6C0144FD9EF92A" box="[400,552,1644,1670]" italics="true" pageId="6" pageNumber="320">O. mcneiceae</emphasis>
</taxonomicName>
(male only) have a distinct tuft of setae between the cheliped fingers. 
<taxonomicName id="4C6935EBFFBF5859FAC60144FEB4F906" authorityName="Ng &amp; Naruse" authorityYear="2024" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="320" phylum="Arthropoda" rank="species" species="varungan" status="sp. nov.">
<emphasis id="B91D927AFFBF5859FAC60144FEB4F906" italics="true" pageId="6" pageNumber="320">Orcovita varungan</emphasis>
</taxonomicName>
<emphasis id="B91D927AFFBF5859FEF601B8FEFEF906" bold="true" box="[266,328,1680,1706]" pageId="6" pageNumber="320">
<taxonomicNameLabel id="A22E2F01FFBF5859FEF601B8FEFEF906" box="[266,328,1680,1706]" pageId="6" pageNumber="320" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
belongs in the same group as these two species.
</paragraph>
<paragraph id="8BD64E68FFBF5858FF3B019CFD19FF79" blockId="6.[151,1437,1500,2030]" lastBlockId="7.[151,1437,151,465]" lastPageId="7" lastPageNumber="321" pageId="6" pageNumber="320">
<taxonomicName id="4C6935EBFFBF5859FF3B019CFE2BF962" authorityName="Ng &amp; Naruse" authorityYear="2024" box="[199,413,1716,1742]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="320" phylum="Arthropoda" rank="species" species="varungan" status="sp. nov.">
<emphasis id="B91D927AFFBF5859FF3B019CFE2BF962" box="[199,413,1716,1742]" italics="true" pageId="6" pageNumber="320">Orcovita varungan</emphasis>
</taxonomicName>
<emphasis id="B91D927AFFBF5859FE59019CFE52F962" bold="true" box="[421,484,1716,1742]" pageId="6" pageNumber="320">
<taxonomicNameLabel id="A22E2F01FFBF5859FE59019CFE52F962" box="[421,484,1716,1742]" pageId="6" pageNumber="320" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can easily be separated from 
<taxonomicName id="4C6935EBFFBF5859FCCB019CFC7CF962" authorityName="P. K. L. Ng, Guinot &amp; Iliffe" authorityYear="1996" box="[823,970,1716,1742]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="320" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B91D927AFFBF5859FCCB019CFC7CF962" box="[823,970,1716,1742]" italics="true" pageId="6" pageNumber="320">O. gracilipes</emphasis>
</taxonomicName>
(from 
<collectingCountry id="F37E0EF8FFBF5859FBE6019CFB28F962" box="[1050,1182,1716,1742]" name="Niue Island" pageId="6" pageNumber="320">Niue Island</collectingCountry>
) by its carapace being proportionally narrower (carapace width 1.27–1.36 times length) (
<figureCitation id="135252EDFFBF5859FC7A01F0FBB7F95E" box="[902,1025,1752,1778]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="6" pageNumber="320">Fig. 2A–D</figureCitation>
) (versus carapace wider, width 1.40 times length) (
<figureCitation id="135252EDFFBF5859FEBF01D4FE14F8BA" box="[323,418,1788,1814]" captionStart="FIGURE 7" captionStartId="8.[151,250,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-17@8.[151,1435,181,1746]" captionTargetPageId="8" captionText="FIGURE 7. A, C, Orcovita gracilipes P.K.L. Ng, Guinot &amp; Iliffe, 1996, paratype male (15.6 × 11.5 mm), ZRC 1996.0096, Niue; B, D, O. mcneiceae N.K. Ng &amp; P.K.L. Ng, 2002, paratype female (14.1 × 11.1 mm), ZRC 2001.1149, New Caledonia. A, B, overall dorsal habitus; C, D, right side of carapace." figureDoi="http://doi.org/10.5281/zenodo.12681808" httpUri="https://zenodo.org/record/12681808/files/figure.png" pageId="6" pageNumber="320">Fig. 7A</figureCitation>
); proportionally less slender ambulatory legs (
<figureCitation id="135252EDFFBF5859FC3B01D4FB94F8BA" box="[967,1058,1788,1814]" captionStart="FIGURE 3" captionStartId="4.[151,250,1658,1682]" captionTargetBox="[158,1430,489,1634]" captionTargetId="figure-182@4.[158,1430,489,1634]" captionTargetPageId="4" captionText="FIGURE 3. Orcovita varungan n. sp., A, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; B, C, D, E, G, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, outer view of right chela; B, outer view of left chela; C, view of pulvinus at base of fingers of left chela; D, dorsal view of left cheliped; E, right P4 and P5; F, G, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.12681798" httpUri="https://zenodo.org/record/12681798/files/figure.png" pageId="6" pageNumber="320">Fig. 3E</figureCitation>
; 
<tableCitation id="C6EB7BD3FFBF5859FBCE01D4FB3EF8BA" box="[1074,1160,1788,1814]" captionStart="TABLE 1" captionStartId="9.[151,239,151,175]" captionTargetPageId="9" captionText="TABLE 1. Length to width ratios of P4 and P5 merus, propodus and dactylus of Orcovita varungan n. sp." httpUri="http://table.plazi.org/id/DF161EE0FFB05856FF6B07BFFAB5FF1C" pageId="6" pageNumber="320" tableUuid="DF161EE0FFB05856FF6B07BFFAB5FF1C">Table 1</tableCitation>
) (versus proportionally slender; cf. 
<bibRefCitation id="EFF83399FFBF5859FEDD0008FE78F896" author="Ng, P. K. L. &amp; Guinot, D. &amp; Iliffe, T. M." box="[289,462,1824,1850]" pageId="6" pageNumber="320" pagination="109 - 134" refId="ref5235" refString="Ng, P. K. L., Guinot, D. &amp; Iliffe, T. M. (1996) Revision of the anchialine varunine crabs of the genus Orcovita Ng &amp; Tomascik, 1994 (Crustacea: Decapoda: Brachyura: Grapsidae), with descriptions of four new species. Raffles Bulletin of Zoology, 44 (1), 109 - 134." type="journal article" year="1996">
Ng 
<emphasis id="B91D927AFFBF5859FEB10009FE3EF896" box="[333,392,1824,1850]" italics="true" pageId="6" pageNumber="320">et al.</emphasis>
1996
</bibRefCitation>
: fig. 1B, table 1; 
<figureCitation id="135252EDFFBF5859FD610008FD4FF896" box="[669,761,1824,1850]" captionStart="FIGURE 7" captionStartId="8.[151,250,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-17@8.[151,1435,181,1746]" captionTargetPageId="8" captionText="FIGURE 7. A, C, Orcovita gracilipes P.K.L. Ng, Guinot &amp; Iliffe, 1996, paratype male (15.6 × 11.5 mm), ZRC 1996.0096, Niue; B, D, O. mcneiceae N.K. Ng &amp; P.K.L. Ng, 2002, paratype female (14.1 × 11.1 mm), ZRC 2001.1149, New Caledonia. A, B, overall dorsal habitus; C, D, right side of carapace." figureDoi="http://doi.org/10.5281/zenodo.12681808" httpUri="https://zenodo.org/record/12681808/files/figure.png" pageId="6" pageNumber="320">Fig. 7A</figureCitation>
); the posterolateral margin is less convergent (
<figureCitation id="135252EDFFBF5859FAEB0008FA22F896" box="[1303,1428,1824,1850]" captionStart="FIGURE 2" captionStartId="3.[151,250,1706,1730]" captionTargetBox="[151,1435,181,1682]" captionTargetId="figure-20@3.[151,1435,181,1682]" captionTargetPageId="3" captionText="FIGURE 2. Orcovita varungan n. sp., A, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi; B, paratype male (22.9 × 17.7 mm), SMF 5257b, Sulawesi; C, E, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; D, G, H, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A–D, dorsal view of carapace; E–H, frontal view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.12681796" httpUri="https://zenodo.org/record/12681796/files/figure.png" pageId="6" pageNumber="320">Fig. 2A–D</figureCitation>
) (versus distinctly converging towards posterior carapace margin; cf. 
<bibRefCitation id="EFF83399FFBF5859FC65006CFB88F8F2" author="Ng, P. K. L. &amp; Guinot, D. &amp; Iliffe, T. M." box="[921,1086,1860,1886]" pageId="6" pageNumber="320" pagination="109 - 134" refId="ref5235" refString="Ng, P. K. L., Guinot, D. &amp; Iliffe, T. M. (1996) Revision of the anchialine varunine crabs of the genus Orcovita Ng &amp; Tomascik, 1994 (Crustacea: Decapoda: Brachyura: Grapsidae), with descriptions of four new species. Raffles Bulletin of Zoology, 44 (1), 109 - 134." type="journal article" year="1996">
Ng 
<emphasis id="B91D927AFFBF5859FC3E006DFC4DF8F2" box="[962,1019,1860,1886]" italics="true" pageId="6" pageNumber="320">et al.</emphasis>
1996
</bibRefCitation>
: figs. 1b, 7A; 
<figureCitation id="135252EDFFBF5859FB20006CFA84F8F2" box="[1244,1330,1860,1886]" captionStart="FIGURE 7" captionStartId="8.[151,250,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-17@8.[151,1435,181,1746]" captionTargetPageId="8" captionText="FIGURE 7. A, C, Orcovita gracilipes P.K.L. Ng, Guinot &amp; Iliffe, 1996, paratype male (15.6 × 11.5 mm), ZRC 1996.0096, Niue; B, D, O. mcneiceae N.K. Ng &amp; P.K.L. Ng, 2002, paratype female (14.1 × 11.1 mm), ZRC 2001.1149, New Caledonia. A, B, overall dorsal habitus; C, D, right side of carapace." figureDoi="http://doi.org/10.5281/zenodo.12681808" httpUri="https://zenodo.org/record/12681808/files/figure.png" pageId="6" pageNumber="320">Fig. 7A</figureCitation>
); and the distal corneous beak of the G1 is directed obliquely dorsolaterally (
<figureCitation id="135252EDFFBF5859FC640040FB6EF82E" box="[920,1240,1896,1922]" captionStart="FIGURE 6" captionStartId="7.[151,250,1851,1875]" captionTargetBox="[177,1411,501,1810]" captionTargetId="figure-164@7.[151,1435,489,1827]" captionTargetPageId="7" captionText="FIGURE 6. Gonopods of Orcovita varungan n. sp., holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi. A–D, with setae, before denuding; E–O, setae denuded. A–G, left G1; H–M, distal part of left G1 with corneous distal part; N, O. left G2. A, E, H, N, ventral view; B, I, subventral view; C, J, outer mesial view; D, G, K, O, dorsal view; L, subdorsal view; F, M, inner mesial view. Scales: E–G, N, O = 0.5 mm; H–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12681806" httpUri="https://zenodo.org/record/12681806/files/figure.png" pageId="6" pageNumber="320">Fig. 6A, B, D, E, G–I, K, L</figureCitation>
) (versus directed more vertically, cf. 
<bibRefCitation id="EFF83399FFBF5859FE8900A4FDABF80A" author="Ng, P. K. L. &amp; Guinot, D. &amp; Iliffe, T. M." box="[373,541,1932,1958]" pageId="6" pageNumber="320" pagination="109 - 134" refId="ref5235" refString="Ng, P. K. L., Guinot, D. &amp; Iliffe, T. M. (1996) Revision of the anchialine varunine crabs of the genus Orcovita Ng &amp; Tomascik, 1994 (Crustacea: Decapoda: Brachyura: Grapsidae), with descriptions of four new species. Raffles Bulletin of Zoology, 44 (1), 109 - 134." type="journal article" year="1996">
Ng 
<emphasis id="B91D927AFFBF5859FE6300A5FE6FF80A" box="[415,473,1932,1958]" italics="true" pageId="6" pageNumber="320">et al.</emphasis>
1996
</bibRefCitation>
: fig. 8B–F). 
<taxonomicName id="4C6935EBFFBF5859FD5200A4FC32F80A" authorityName="Ng &amp; Naruse" authorityYear="2024" box="[686,900,1932,1958]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="320" phylum="Arthropoda" rank="species" species="varungan" status="sp. nov.">
<emphasis id="B91D927AFFBF5859FD5200A4FC32F80A" box="[686,900,1932,1958]" italics="true" pageId="6" pageNumber="320">Orcovita varungan</emphasis>
</taxonomicName>
<emphasis id="B91D927AFFBF5859FC7000A4FC7AF80A" bold="true" box="[908,972,1932,1958]" pageId="6" pageNumber="320">
<taxonomicNameLabel id="A22E2F01FFBF5859FC7000A4FC7AF80A" box="[908,972,1932,1958]" pageId="6" pageNumber="320" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be distinguished from 
<taxonomicName id="4C6935EBFFBF5859FAFF00A4FA2AF80A" authorityName="N. K. Ng &amp; P. K. L. Ng" authorityYear="2002" box="[1283,1436,1932,1958]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="320" phylum="Arthropoda" rank="species" species="mcneiceae">
<emphasis id="B91D927AFFBF5859FAFF00A4FA2AF80A" box="[1283,1436,1932,1958]" italics="true" pageId="6" pageNumber="320">O. mcneiceae</emphasis>
</taxonomicName>
(from 
<collectingRegion id="49AD808AFFBF5859FF220098FE39F866" box="[222,399,1968,1994]" country="New Caledonia" name="Province des Iles" pageId="6" pageNumber="320">Loyalty Islands</collectingRegion>
, 
<collectingCountry id="F37E0EF8FFBF5859FE670098FDFCF866" box="[411,586,1968,1994]" name="New Caledonia" pageId="6" pageNumber="320">New Caledonia</collectingCountry>
) by its relatively longer P5 dactylus and shorter P4 propodus and P5 merus and propodus (
<figureCitation id="135252EDFFBF5859FEBD00FCFE2EF842" box="[321,408,2004,2030]" captionStart="FIGURE 3" captionStartId="4.[151,250,1658,1682]" captionTargetBox="[158,1430,489,1634]" captionTargetId="figure-182@4.[158,1430,489,1634]" captionTargetPageId="4" captionText="FIGURE 3. Orcovita varungan n. sp., A, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; B, C, D, E, G, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, outer view of right chela; B, outer view of left chela; C, view of pulvinus at base of fingers of left chela; D, dorsal view of left cheliped; E, right P4 and P5; F, G, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.12681798" httpUri="https://zenodo.org/record/12681798/files/figure.png" pageId="6" pageNumber="320">Fig. 3E</figureCitation>
; 
<tableCitation id="C6EB7BD3FFBF5859FE5900FCFE41F842" box="[421,503,2004,2030]" captionStart="TABLE 1" captionStartId="9.[151,239,151,175]" captionTargetPageId="9" captionText="TABLE 1. Length to width ratios of P4 and P5 merus, propodus and dactylus of Orcovita varungan n. sp." httpUri="http://table.plazi.org/id/DF161EE0FFB05856FF6B07BFFAB5FF1C" pageId="6" pageNumber="320" tableUuid="DF161EE0FFB05856FF6B07BFFAB5FF1C">Table 1</tableCitation>
) (versus distinctly shorter; N.K. Ng &amp; P.K.L. 
<bibRefCitation id="EFF83399FFBF5859FBFB00FCFBD8F842" author="Ng, N. K. &amp; Ng, P. K. L." box="[1031,1134,2004,2030]" pageId="6" pageNumber="320" pagination="663 - 670" refId="ref5175" refString="Ng, N. K. &amp; Ng, P. K. L. (2002) Orcovita mcneiceae, a new species of anchialine crab (Brachyura, Grapsidae, Varuninae) from the Loyalty Islands. Crustaceana, 75 (5), 663 - 670. https: // doi. org / 10.1163 / 156854002760202660" type="journal article" year="2002">Ng 2002</bibRefCitation>
: fig. 2A, table 1; 
<figureCitation id="135252EDFFBF5859FACA00FCFA38F842" box="[1334,1422,2004,2030]" captionStart="FIGURE 7" captionStartId="8.[151,250,1770,1794]" captionTargetBox="[151,1435,181,1746]" captionTargetId="figure-17@8.[151,1435,181,1746]" captionTargetPageId="8" captionText="FIGURE 7. A, C, Orcovita gracilipes P.K.L. Ng, Guinot &amp; Iliffe, 1996, paratype male (15.6 × 11.5 mm), ZRC 1996.0096, Niue; B, D, O. mcneiceae N.K. Ng &amp; P.K.L. Ng, 2002, paratype female (14.1 × 11.1 mm), ZRC 2001.1149, New Caledonia. A, B, overall dorsal habitus; C, D, right side of carapace." figureDoi="http://doi.org/10.5281/zenodo.12681808" httpUri="https://zenodo.org/record/12681808/files/figure.png" pageId="6" pageNumber="320">Fig. 7B</figureCitation>
); and distal corneous beak of the G1 gently bent proximally (
<figureCitation id="135252EDFFBE5858FCBF07BFFB32FF1D" box="[835,1156,151,177]" captionStart="FIGURE 6" captionStartId="7.[151,250,1851,1875]" captionTargetBox="[177,1411,501,1810]" captionTargetId="figure-164@7.[151,1435,489,1827]" captionTargetPageId="7" captionText="FIGURE 6. Gonopods of Orcovita varungan n. sp., holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi. A–D, with setae, before denuding; E–O, setae denuded. A–G, left G1; H–M, distal part of left G1 with corneous distal part; N, O. left G2. A, E, H, N, ventral view; B, I, subventral view; C, J, outer mesial view; D, G, K, O, dorsal view; L, subdorsal view; F, M, inner mesial view. Scales: E–G, N, O = 0.5 mm; H–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12681806" httpUri="https://zenodo.org/record/12681806/files/figure.png" pageId="7" pageNumber="321">Fig. 6A, B, D, E, G–I, K, L</figureCitation>
) (versus directed almost vertically; N.K. Ng &amp; P.K.L. 
<bibRefCitation id="EFF83399FFBE5858FE1D0793FDF0FF79" author="Ng, N. K. &amp; Ng, P. K. L." box="[481,582,187,213]" pageId="7" pageNumber="321" pagination="663 - 670" refId="ref5175" refString="Ng, N. K. &amp; Ng, P. K. L. (2002) Orcovita mcneiceae, a new species of anchialine crab (Brachyura, Grapsidae, Varuninae) from the Loyalty Islands. Crustaceana, 75 (5), 663 - 670. https: // doi. org / 10.1163 / 156854002760202660" type="journal article" year="2002">Ng 2002</bibRefCitation>
: fig. 3C).
</paragraph>
<paragraph id="8BD64E68FFBE5858FF3B07F7FF09FE7D" blockId="7.[151,1437,151,465]" pageId="7" pageNumber="321">
The subhexagonal carapace shape of 
<taxonomicName id="4C6935EBFFBE5858FD8907F7FCB2FF54" authorityName="Ng &amp; Naruse" authorityYear="2024" box="[629,772,223,249]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="321" phylum="Arthropoda" rank="species" species="varungan" status="n. sp.">
<emphasis id="B91D927AFFBE5858FD8907F7FCB2FF54" box="[629,772,223,249]" italics="true" pageId="7" pageNumber="321">O. varungan</emphasis>
</taxonomicName>
<emphasis id="B91D927AFFBE5858FCF207F7FCF9FF55" bold="true" box="[782,847,223,249]" pageId="7" pageNumber="321">
<taxonomicNameLabel id="A22E2F01FFBE5858FCF207F7FCF9FF55" box="[782,847,223,249]" pageId="7" pageNumber="321" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
most closely resembles that of 
<taxonomicName id="4C6935EBFFBE5858FB4307F7FAE5FF55" authorityName="Stasolla &amp; Innocenti" authorityYear="2014" box="[1215,1363,223,249]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="321" phylum="Arthropoda" rank="species" species="tabiacoud">
<emphasis id="B91D927AFFBE5858FB4307F7FAE5FF55" box="[1215,1363,223,249]" italics="true" pageId="7" pageNumber="321">O. tabiacoud</emphasis>
</taxonomicName>
(from 
<collectingRegion id="49AD808AFFBE5858FF6B062BFF4FFEB1" box="[151,249,259,285]" country="Philippines" name="Palawan" pageId="7" pageNumber="321">Palawan</collectingRegion>
, 
<collectingCountry id="F37E0EF8FFBE5858FEF9062BFE3EFEB1" box="[261,392,259,285]" name="Philippines" pageId="7" pageNumber="321">Philippines</collectingCountry>
), but the former species can easily be separated by the presence of a prominent tuft of setae between the fingers of the adult male chela (
<figureCitation id="135252EDFFBE5858FD7C060FFD4EFEED" box="[640,760,295,321]" captionStart="FIGURE 3" captionStartId="4.[151,250,1658,1682]" captionTargetBox="[158,1430,489,1634]" captionTargetId="figure-182@4.[158,1430,489,1634]" captionTargetPageId="4" captionText="FIGURE 3. Orcovita varungan n. sp., A, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; B, C, D, E, G, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, outer view of right chela; B, outer view of left chela; C, view of pulvinus at base of fingers of left chela; D, dorsal view of left cheliped; E, right P4 and P5; F, G, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.12681798" httpUri="https://zenodo.org/record/12681798/files/figure.png" pageId="7" pageNumber="321">Fig. 3A–C</figureCitation>
) (absent in 
<taxonomicName id="4C6935EBFFBE5858FC85060FFBBCFEED" authorityName="Stasolla &amp; Innocenti" authorityYear="2014" box="[889,1034,295,321]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="321" phylum="Arthropoda" rank="species" species="tabiacoud">
<emphasis id="B91D927AFFBE5858FC85060FFBBCFEED" box="[889,1034,295,321]" italics="true" pageId="7" pageNumber="321">O. tabiacoud</emphasis>
</taxonomicName>
; cf. 
<bibRefCitation id="EFF83399FFBE5858FBC7060FFAD7FEED" author="Stasolla, G. &amp; Innocenti, G." box="[1083,1377,295,321]" pageId="7" pageNumber="321" pagination="591 - 599" refId="ref5361" refString="Stasolla, G. &amp; Innocenti, G. (2014) A new species of cavernicolous crab from Coron Island, Palawan, the Philippines (Decapoda: Brachyura: Varunidae). Raffles Bulletin of Zoology, 62, 591 - 599." type="journal article" year="2014">Stasolla &amp; Innocenti 2014</bibRefCitation>
: figs. 6, 7D), the ischium of the third maxilliped is proportionately longer (
<figureCitation id="135252EDFFBE5858FC6B0663FBBFFEC9" box="[919,1033,331,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1658,1682]" captionTargetBox="[158,1430,489,1634]" captionTargetId="figure-182@4.[158,1430,489,1634]" captionTargetPageId="4" captionText="FIGURE 3. Orcovita varungan n. sp., A, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; B, C, D, E, G, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, outer view of right chela; B, outer view of left chela; C, view of pulvinus at base of fingers of left chela; D, dorsal view of left cheliped; E, right P4 and P5; F, G, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.12681798" httpUri="https://zenodo.org/record/12681798/files/figure.png" pageId="7" pageNumber="321">Fig. 3F, G</figureCitation>
) (versus relatively shorter and more quadrate; 
<bibRefCitation id="EFF83399FFBE5858FEF40647FD83FE25" author="Stasolla, G. &amp; Innocenti, G." box="[264,565,367,393]" pageId="7" pageNumber="321" pagination="591 - 599" refId="ref5361" refString="Stasolla, G. &amp; Innocenti, G. (2014) A new species of cavernicolous crab from Coron Island, Palawan, the Philippines (Decapoda: Brachyura: Varunidae). Raffles Bulletin of Zoology, 62, 591 - 599." type="journal article" year="2014">Stasolla &amp; Innocenti 2014</bibRefCitation>
: figs. 5B, 7F); and the P4 dactylus is proportionately longer, length to width 9.46–9.83 (
<figureCitation id="135252EDFFBE5858FEEB06BBFE2BFE01" box="[279,413,403,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1935,1959]" captionTargetBox="[151,1435,181,1911]" captionTargetId="figure-17@2.[151,1435,181,1911]" captionTargetPageId="2" captionText="FIGURE 1. Orcovita varungan n. sp., overall dorsal habitus. A, holotype male (22.9 × 17.6 mm) (SMF 5257a), Sulawesi; B, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; C, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi." figureDoi="http://doi.org/10.5281/zenodo.12681794" httpUri="https://zenodo.org/record/12681794/files/figure.png" pageId="7" pageNumber="321">Figs. 1A, C</figureCitation>
, 
<figureCitation id="135252EDFFBE5858FE5606BBFE7DFE01" box="[426,459,403,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1658,1682]" captionTargetBox="[158,1430,489,1634]" captionTargetId="figure-182@4.[158,1430,489,1634]" captionTargetPageId="4" captionText="FIGURE 3. Orcovita varungan n. sp., A, F, paratype male (19.5 × 14.9 mm), ZRC 2023.0014, Sulawesi; B, C, D, E, G, male (22.0 × 16.2 mm), ZRC 2023.0015, Sulawesi. A, outer view of right chela; B, outer view of left chela; C, view of pulvinus at base of fingers of left chela; D, dorsal view of left cheliped; E, right P4 and P5; F, G, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.12681798" httpUri="https://zenodo.org/record/12681798/files/figure.png" pageId="7" pageNumber="321">3E</figureCitation>
; 
<tableCitation id="C6EB7BD3FFBE5858FE2406BBFD9FFE01" box="[472,553,403,429]" captionStart="TABLE 1" captionStartId="9.[151,239,151,175]" captionTargetPageId="9" captionText="TABLE 1. Length to width ratios of P4 and P5 merus, propodus and dactylus of Orcovita varungan n. sp." httpUri="http://table.plazi.org/id/DF161EE0FFB05856FF6B07BFFAB5FF1C" pageId="7" pageNumber="321" tableUuid="DF161EE0FFB05856FF6B07BFFAB5FF1C">Table 1</tableCitation>
) (proportionally shorter, ratio 6.69 cf. 
<bibRefCitation id="EFF83399FFBE5858FC1E06BBFABBFE01" author="Stasolla, G. &amp; Innocenti, G." box="[994,1293,403,429]" pageId="7" pageNumber="321" pagination="591 - 599" refId="ref5361" refString="Stasolla, G. &amp; Innocenti, G. (2014) A new species of cavernicolous crab from Coron Island, Palawan, the Philippines (Decapoda: Brachyura: Varunidae). Raffles Bulletin of Zoology, 62, 591 - 599." type="journal article" year="2014">Stasolla &amp; Innocenti 2014</bibRefCitation>
: figs. 5A, B, 7I).
</paragraph>
<caption id="DF161EE0FFBE5858FF6B0013FD1FF813" ID-DOI="http://doi.org/10.5281/zenodo.12681806" ID-Zenodo-Dep="12681806" httpUri="https://zenodo.org/record/12681806/files/figure.png" pageId="7" pageNumber="321" startId="7.[151,250,1851,1875]" targetBox="[177,1411,501,1810]" targetPageId="7" targetType="figure">
<paragraph id="8BD64E68FFBE5858FF6B0013FD1FF813" blockId="7.[151,1436,1851,1983]" pageId="7" pageNumber="321">
<emphasis id="B91D927AFFBE5858FF6B0013FEBBF8FF" bold="true" box="[151,269,1851,1875]" pageId="7" pageNumber="321">FIGURE 6</emphasis>
. Gonopods of 
<taxonomicName id="4C6935EBFFBE5858FE580014FDD2F8FF" authorityName="Ng &amp; Naruse" authorityYear="2024" box="[420,612,1852,1875]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="321" phylum="Arthropoda" rank="species" species="varungan" status="sp. nov.">
<emphasis id="B91D927AFFBE5858FE580014FDD2F8FF" box="[420,612,1852,1875]" italics="true" pageId="7" pageNumber="321">Orcovita varungan</emphasis>
</taxonomicName>
<emphasis id="B91D927AFFBE5858FD970013FD12F8FF" bold="true" box="[619,676,1851,1875]" pageId="7" pageNumber="321">
<taxonomicNameLabel id="A22E2F01FFBE5858FD970013FD12F8FF" box="[619,676,1851,1875]" pageId="7" pageNumber="321" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male (22.9 × 17.6 mm), SMF 5257a, Sulawesi. A–D, with setae, before denuding; E–O, setae denuded. A–G, left G1; H–M, distal part of left G1 with corneous distal part; N, O. left G2. A, E, H, N, ventral view; B, I, subventral view; C, J, outer mesial view; D, G, K, O, dorsal view; L, subdorsal view; F, M, inner mesial view. Scales: E–G, N, O = 0.5 mm; H–M = 0.2 mm.
</paragraph>
</caption>
<caption id="DF161EE0FFB15857FF6B01C2FD2AF8E6" ID-DOI="http://doi.org/10.5281/zenodo.12681808" ID-Zenodo-Dep="12681808" httpUri="https://zenodo.org/record/12681808/files/figure.png" pageId="8" pageNumber="322" startId="8.[151,250,1770,1794]" targetBox="[151,1435,181,1746]" targetPageId="8" targetType="figure">
<paragraph id="8BD64E68FFB15857FF6B01C2FD2AF8E6" blockId="8.[151,1437,1770,1866]" pageId="8" pageNumber="322">
<emphasis id="B91D927AFFB15857FF6B01C2FEBAF8AE" bold="true" box="[151,268,1770,1794]" pageId="8" pageNumber="322">FIGURE 7</emphasis>
. A, C, 
<taxonomicName id="4C6935EBFFB15857FEAC01C3FCD6F8AE" authority="P. K. L. Ng, Guinot &amp; Iliffe, 1996" authorityName="P. K. L. Ng, Guinot &amp; Iliffe" authorityYear="1996" box="[336,864,1770,1795]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="322" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B91D927AFFB15857FEAC01C3FDA2F8AE" box="[336,532,1770,1794]" italics="true" pageId="8" pageNumber="322">Orcovita gracilipes</emphasis>
P.K.L. 
<bibRefCitation id="EFF83399FFB15857FDA201C3FCD6F8AE" author="Ng, P. K. L. &amp; Guinot, D. &amp; Iliffe, T. M." box="[606,864,1770,1795]" pageId="8" pageNumber="322" pagination="109 - 134" refId="ref5235" refString="Ng, P. K. L., Guinot, D. &amp; Iliffe, T. M. (1996) Revision of the anchialine varunine crabs of the genus Orcovita Ng &amp; Tomascik, 1994 (Crustacea: Decapoda: Brachyura: Grapsidae), with descriptions of four new species. Raffles Bulletin of Zoology, 44 (1), 109 - 134." type="journal article" year="1996">Ng, Guinot &amp; Iliffe, 1996</bibRefCitation>
</taxonomicName>
, paratype male (15.6 × 11.5 mm), ZRC 1996.0096, Niue; B, D, 
<taxonomicName id="4C6935EBFFB15857FF2A0027FD38F88A" authority="N. K. Ng &amp; P. K. L. Ng, 2002" authorityName="N. K. Ng &amp; P. K. L. Ng" authorityYear="2002" box="[214,654,1806,1831]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="322" phylum="Arthropoda" rank="species" species="mcneiceae">
<emphasis id="B91D927AFFB15857FF2A0027FED4F88A" box="[214,354,1807,1830]" italics="true" pageId="8" pageNumber="322">O. mcneiceae</emphasis>
N.K. Ng &amp; P.K.L. 
<bibRefCitation id="EFF83399FFB15857FDD00027FD38F88A" author="Ng, N. K. &amp; Ng, P. K. L." box="[556,654,1806,1831]" pageId="8" pageNumber="322" pagination="663 - 670" refId="ref5175" refString="Ng, N. K. &amp; Ng, P. K. L. (2002) Orcovita mcneiceae, a new species of anchialine crab (Brachyura, Grapsidae, Varuninae) from the Loyalty Islands. Crustaceana, 75 (5), 663 - 670. https: // doi. org / 10.1163 / 156854002760202660" type="journal article" year="2002">Ng, 2002</bibRefCitation>
</taxonomicName>
, paratype female (14.1 × 11.1 mm), ZRC 2001.1149, New Caledonia. A, B, overall dorsal habitus; C, D, right side of carapace.
</paragraph>
</caption>
<paragraph id="8BD64E68FFB15857FF3B0051FAE8F877" blockId="8.[151,1437,1912,2011]" pageId="8" pageNumber="322">
<taxonomicName id="4C6935EBFFB15857FF3B0051FE2DF83E" authorityName="Ng &amp; Naruse" authorityYear="2024" box="[199,411,1913,1939]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="322" phylum="Arthropoda" rank="species" species="varungan" status="sp. nov.">
<emphasis id="B91D927AFFB15857FF3B0051FE2DF83E" box="[199,411,1913,1939]" italics="true" pageId="8" pageNumber="322">Orcovita varungan</emphasis>
</taxonomicName>
<emphasis id="B91D927AFFB15857FE5E0051FE56F83F" bold="true" box="[418,480,1913,1939]" pageId="8" pageNumber="322">
<taxonomicNameLabel id="A22E2F01FFB15857FE5E0051FE56F83F" box="[418,480,1913,1939]" pageId="8" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is only the second species of the genus known from 
<collectingCountry id="F37E0EF8FFB15857FBD50050FB2FF83E" box="[1065,1177,1912,1938]" name="Indonesia" pageId="8" pageNumber="322">Indonesia</collectingCountry>
, the first being the 
<typeStatus id="54D2F0CAFFB15857FA900051FA2AF83F" box="[1388,1436,1913,1939]" pageId="8" pageNumber="322">type</typeStatus>
species, 
<taxonomicName id="4C6935EBFFB15857FF0400B5FECCF81A" authorityName="P. K. L. Ng &amp; Tomascik" authorityYear="1994" box="[248,378,1948,1975]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="322" phylum="Arthropoda" rank="species" species="saltatrix">
<emphasis id="B91D927AFFB15857FF0400B5FECCF81A" box="[248,378,1948,1975]" italics="true" pageId="8" pageNumber="322">O. saltatrix</emphasis>
</taxonomicName>
, from Kakaban Island, which is between Borneo and 
<collectingRegion id="49AD808AFFB15857FC1500B4FBF8F81A" box="[1001,1102,1948,1974]" country="Indonesia" name="Sulawesi Utara" pageId="8" pageNumber="322">Sulawesi</collectingRegion>
(
<bibRefCitation id="EFF83399FFB15857FBA300B5FAEAF81B" author="Ng, P. K. L. &amp; Tomascik, T." box="[1119,1372,1948,1975]" pageId="8" pageNumber="322" pagination="937 - 948" refId="ref5304" refString="Ng, P. K. L. &amp; Tomascik, T. (1994) Orcovita saltatrix, a new genus and species of anchialine varunine crab (Crustacea: Decapoda: Brachyura: Grapsidae) from Kakaban Island, Indonesia. Raffles Bulletin of Zoology, 42 (4), 937 - 948." type="journal article" year="1994">Ng &amp; Tomascik 1994</bibRefCitation>
). The highest diversity of species is from the 
<collectingCountry id="F37E0EF8FFB15857FDB300E8FD66F876" box="[591,720,1984,2010]" name="Philippines" pageId="8" pageNumber="322">Philippines</collectingCountry>
, with four known species (see 
<bibRefCitation id="EFF83399FFB15857FBD400E8FAE4F877" author="Stasolla, G. &amp; Innocenti, G." box="[1064,1362,1984,2011]" pageId="8" pageNumber="322" pagination="591 - 599" refId="ref5361" refString="Stasolla, G. &amp; Innocenti, G. (2014) A new species of cavernicolous crab from Coron Island, Palawan, the Philippines (Decapoda: Brachyura: Varunidae). Raffles Bulletin of Zoology, 62, 591 - 599." type="journal article" year="2014">Stasolla &amp; Innocenti 2014</bibRefCitation>
).
</paragraph>
<caption id="DF161EE0FFB05856FF6B07BFFAB5FF1C" ID-Table-UUID="DF161EE0FFB05856FF6B07BFFAB5FF1C" box="[151,1283,151,176]" httpUri="http://table.plazi.org/id/DF161EE0FFB05856FF6B07BFFAB5FF1C" pageId="9" pageNumber="323" startId="9.[151,239,151,175]" targetBox="[167,1397,192,444]" targetIsTable="true" targetPageId="9" targetType="table">
<paragraph id="8BD64E68FFB05856FF6B07BFFAB5FF1C" blockId="9.[151,1283,151,176]" box="[151,1283,151,176]" pageId="9" pageNumber="323">
<emphasis id="B91D927AFFB05856FF6B07BFFEB5FF03" bold="true" box="[151,259,151,175]" pageId="9" pageNumber="323">TABLE 1</emphasis>
. Length to width ratios of P4 and P5 merus, propodus and dactylus of 
<taxonomicName id="4C6935EBFFB05856FC0A07B0FB76FF03" authorityName="Ng &amp; Naruse" authorityYear="2024" box="[1014,1216,152,175]" class="Malacostraca" family="Varunidae" genus="Orcovita" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="323" phylum="Arthropoda" rank="species" species="varungan" status="sp. nov.">
<emphasis id="B91D927AFFB05856FC0A07B0FB76FF03" box="[1014,1216,152,175]" italics="true" pageId="9" pageNumber="323">Orcovita varungan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E2F01FFB05856FB3B07B0FAB5FF1C" box="[1223,1283,152,176]" pageId="9" pageNumber="323" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BD64E68FFB05856FE8507E8FAFDFE07" pageId="9" pageNumber="323">
<table id="F969BCC8FFB0A7A0FF5B07E8FAC3FE10" box="[167,1397,192,444]" gridcols="7" gridrows="4" pageId="9" pageNumber="323">
<tr id="35594C2AFFB0A7A0FF5B07E8FAC3FF50" box="[167,1397,192,252]" gridrow="0" pageId="9" pageNumber="323" rowspan-0="1">
<th id="76882556FFB0A7A0FEAD07E8FE73FF50" box="[337,453,192,252]" gridcol="1" gridrow="0" pageId="9" pageNumber="323">3M = P4 merus</th>
<th id="76882556FFB0A7A0FE0807E8FD3EFF50" box="[500,648,192,252]" gridcol="2" gridrow="0" pageId="9" pageNumber="323">3P = P4 propodus</th>
<th id="76882556FFB0A7A0FD4507E8FCF5FF50" box="[697,835,192,252]" gridcol="3" gridrow="0" pageId="9" pageNumber="323">3D = P4 dactylus</th>
<th id="76882556FFB0A7A0FC8A07E8FC5FFF50" box="[886,1001,192,252]" gridcol="4" gridrow="0" pageId="9" pageNumber="323">4M = P5 merus</th>
<th id="76882556FFB0A7A0FBE107E8FB07FF50" box="[1053,1201,192,252]" gridcol="5" gridrow="0" pageId="9" pageNumber="323">4P = P5 propodus</th>
<th id="76882556FFB0A7A0FB1007E8FAC3FF50" box="[1260,1397,192,252]" gridcol="6" gridrow="0" pageId="9" pageNumber="323">4D = P5 dactylus</th>
</tr>
<tr id="35594C2AFFB0A7A0FF5B0624FAC3FEE4" box="[167,1397,268,328]" gridrow="1" pageId="9" pageNumber="323">
<th id="76882556FFB0A7A0FF5B0624FE9CFEE4" box="[167,298,268,328]" gridcol="0" gridrow="1" pageId="9" pageNumber="323">Ratios (min– max)</th>
<td id="76882556FFB0A7A0FEAD0624FE73FEE4" box="[337,453,268,328]" gridcol="1" gridrow="1" pageId="9" pageNumber="323">4.58–5.42</td>
<td id="76882556FFB0A7A0FE080624FD3EFEE4" box="[500,648,268,328]" gridcol="2" gridrow="1" pageId="9" pageNumber="323">4.00–4.37</td>
<td id="76882556FFB0A7A0FD450624FCF5FEE4" box="[697,835,268,328]" gridcol="3" gridrow="1" pageId="9" pageNumber="323">9.46–9.83</td>
<td id="76882556FFB0A7A0FC8A0624FC5FFEE4" box="[886,1001,268,328]" gridcol="4" gridrow="1" pageId="9" pageNumber="323">3.91–4.37</td>
<td id="76882556FFB0A7A0FBE10624FB07FEE4" box="[1053,1201,268,328]" gridcol="5" gridrow="1" pageId="9" pageNumber="323">2.58–2.74</td>
<td id="76882556FFB0A7A0FB100624FAC3FEE4" box="[1260,1397,268,328]" gridcol="6" gridrow="1" pageId="9" pageNumber="323">5.58–5.93</td>
</tr>
<tr id="35594C2AFFB0A7A0FF5B0670FAC3FEDC" box="[167,1397,344,368]" gridrow="2" pageId="9" pageNumber="323">
<th id="76882556FFB0A7A0FF5B0670FE9CFEDC" box="[167,298,344,368]" gridcol="0" gridrow="2" pageId="9" pageNumber="323">Mean</th>
<td id="76882556FFB0A7A0FEAD0670FE73FEDC" box="[337,453,344,368]" gridcol="1" gridrow="2" pageId="9" pageNumber="323">5.06</td>
<td id="76882556FFB0A7A0FE080670FD3EFEDC" box="[500,648,344,368]" gridcol="2" gridrow="2" pageId="9" pageNumber="323">4.21</td>
<td id="76882556FFB0A7A0FD450670FCF5FEDC" box="[697,835,344,368]" gridcol="3" gridrow="2" pageId="9" pageNumber="323">9.65</td>
<td id="76882556FFB0A7A0FC8A0670FC5FFEDC" box="[886,1001,344,368]" gridcol="4" gridrow="2" pageId="9" pageNumber="323">4.17</td>
<td id="76882556FFB0A7A0FBE10670FB07FEDC" box="[1053,1201,344,368]" gridcol="5" gridrow="2" pageId="9" pageNumber="323">2.68</td>
<td id="76882556FFB0A7A0FB100670FAC3FEDC" box="[1260,1397,344,368]" gridcol="6" gridrow="2" pageId="9" pageNumber="323">5.72</td>
</tr>
<tr id="35594C2AFFB0A7A0FF5B06A8FAC3FE10" box="[167,1397,384,444]" gridrow="3" pageId="9" pageNumber="323">
<th id="76882556FFB0A7A0FF5B06A8FE9CFE10" box="[167,298,384,444]" gridcol="0" gridrow="3" pageId="9" pageNumber="323">Number of individuals</th>
<td id="76882556FFB0A7A0FEAD06A8FE73FE10" box="[337,453,384,444]" gridcol="1" gridrow="3" pageId="9" pageNumber="323">n = 6</td>
<td id="76882556FFB0A7A0FE0806A8FD3EFE10" box="[500,648,384,444]" gridcol="2" gridrow="3" pageId="9" pageNumber="323">n = 3</td>
<td id="76882556FFB0A7A0FD4506A8FCF5FE10" box="[697,835,384,444]" gridcol="3" gridrow="3" pageId="9" pageNumber="323">n = 2</td>
<td id="76882556FFB0A7A0FC8A06A8FC5FFE10" box="[886,1001,384,444]" gridcol="4" gridrow="3" pageId="9" pageNumber="323">n = 6</td>
<td id="76882556FFB0A7A0FBE106A8FB07FE10" box="[1053,1201,384,444]" gridcol="5" gridrow="3" pageId="9" pageNumber="323">n = 4</td>
<td id="76882556FFB0A7A0FB1006A8FAC3FE10" box="[1260,1397,384,444]" gridcol="6" gridrow="3" pageId="9" pageNumber="323">n = 3</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD64E68FFB05856FF3B06C4FD89FB7A" blockId="9.[151,1437,492,1239]" pageId="9" pageNumber="323">
A note on the specimens is necessary. 
<collectorName id="269C2BBEFFB05856FD8606C5FD27FDAB" box="[634,657,493,519]" pageId="9" pageNumber="323">In</collectorName>
the tray set aside by 
<location id="8EB618B3FFB05856FC7806C5FC40FDAB" LSID="urn:lsid:plazi:treatment:03C0FF7EFFB85856FF6B05A7FD89FB7A:8EB618B3FFB05856FC7806C5FC40FDAB" box="[900,1014,493,519]" name="Ngan Kee" pageId="9" pageNumber="323">Ngan Kee</location>
in the 
<collectionCode id="ED78D6ADFFB05856FBBA06C4FBC9FDAA" box="[1094,1151,492,518]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="323">ZRC</collectionCode>
, there were three bottles. 
<collectorName id="269C2BBEFFB05856FF6B0538FF70FD86" box="[151,198,528,554]" pageId="9" pageNumber="323">One</collectorName>
bottle was the most recent male specimen collected by 
<collectorName id="269C2BBEFFB05856FCB60538FC4DFD86" box="[842,1019,528,554]" pageId="9" pageNumber="323">Franck Brehner</collectorName>
in 2007, which was passed to 
<collectorName id="269C2BBEFFB05856FA9C0539FF72FDE3" pageId="9" pageNumber="323">Ngan Kee</collectorName>
by the first author. 
<collectorName id="269C2BBEFFB05856FE60051CFE7EFDE2" box="[412,456,564,590]" pageId="9" pageNumber="323">The</collectorName>
second bottle had 
<specimenCount id="9D6F85E1FFB05856FD6A051DFC8AFDE2" box="[662,828,564,591]" count="2" pageId="9" pageNumber="323" type="generic">two specimens</specimenCount>
(
<specimenCount id="9D6F85E1FFB05856FCB5051CFC05FDE2" box="[841,947,564,590]" count="1" pageId="9" pageNumber="323" type="male">one male</specimenCount>
, 
<specimenCount id="9D6F85E1FFB05856FC41051DFB8CFDE2" box="[957,1082,564,591]" count="1" pageId="9" pageNumber="323" type="female">one female</specimenCount>
) with the label “ex 
<specimenCode id="DBCFE613FFB05856FAEE051CFA38FDE3" box="[1298,1422,564,591]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF 5257</specimenCode>
”. 
<location id="8EB618B3FFB05856FF6B0570FF75FDDE" LSID="urn:lsid:plazi:treatment:03C0FF7EFFB85856FF6B05A7FD89FB7A:8EB618B3FFB05856FF6B0570FF75FDDE" box="[151,195,600,626]" municipality="The" name="The" pageId="9" pageNumber="323">The</location>
third bottle, without any labels, had 
<specimenCount id="9D6F85E1FFB05856FDA30570FCBCFDDE" box="[607,778,600,626]" count="4" pageId="9" pageNumber="323" type="generic">four specimens</specimenCount>
(
<specimenCount id="9D6F85E1FFB05856FCE50570FC3AFDDE" box="[793,908,600,626]" count="2" pageId="9" pageNumber="323" type="male">two males</specimenCount>
and 
<specimenCount id="9D6F85E1FFB05856FC3F0571FBE6FDDE" box="[963,1104,600,627]" count="2" pageId="9" pageNumber="323" type="female">two females</specimenCount>
); these were in the same state of preservation as the one labelled as “ex 
<specimenCode id="DBCFE613FFB05856FD8A0554FD42FD3B" box="[630,756,636,663]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF 5257</specimenCode>
”. 
<collectingMunicipality id="6BB2D412FFB05856FCF70554FC81FD3A" box="[779,823,636,662]" pageId="9" pageNumber="323">The</collectingMunicipality>
<collectionCode id="ED78D6ADFFB05856FCC30554FCC1FD3A" box="[831,887,636,662]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF</collectionCode>
online catalogue (
<collectorName id="269C2BBEFFB05856FBB60554FB2DFD3A" box="[1098,1179,636,662]" pageId="9" pageNumber="323">SeSam</collectorName>
) lists 
<specimenCount id="9D6F85E1FFB05856FB1D0554FAF0FD3A" box="[1249,1350,636,663]" count="15" pageId="9" pageNumber="323" type="male">15 males</specimenCount>
and 
<specimenCount id="9D6F85E1FFB05856FA7C0554FF58FD16" count="21" pageId="9" pageNumber="323" type="female">21 females</specimenCount>
under 
<specimenCode id="DBCFE613FFB05856FEC70588FE03FD17" box="[315,437,672,699]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF 5257</specimenCode>
. 
<collectorName id="269C2BBEFFB05856FE420588FDE7FD16" box="[446,593,672,698]" pageId="9" pageNumber="323">Torben Riehl</collectorName>
kindly sent us all the specimens in the lot listed as 
<specimenCode id="DBCFE613FFB05856FB740588FB48FD17" box="[1160,1278,672,699]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF 5257</specimenCode>
to the authors to examine. 
<collectorName id="269C2BBEFFB05856FEDF05EDFED2FD73" box="[291,356,709,735]" pageId="9" pageNumber="323">Many</collectorName>
of these have lost their ambulatory legs and chelipeds and are in various states of preservation, but all closely resemble the condition of the specimens in the 
<collectionCode id="ED78D6ADFFB05856FCC105C0FCC5FCAE" box="[829,883,744,770]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="323">ZRC</collectionCode>
labelled as “ex 
<specimenCode id="DBCFE613FFB05856FBDE05C0FB2CFCAF" box="[1058,1178,744,771]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF 5257</specimenCode>
” and those in the bottle without any labels. 
<collectorName id="269C2BBEFFB05856FE920424FE37FC8A" box="[366,385,780,806]" pageId="9" pageNumber="323">If</collectorName>
we add the 
<specimenCount id="9D6F85E1FFB05856FDFF0424FDD3FC8A" box="[515,613,780,806]" count="13" pageId="9" pageNumber="323" type="male">13 males</specimenCount>
and 
<specimenCount id="9D6F85E1FFB05856FD650424FCA4FC8A" box="[665,786,780,806]" count="19" pageId="9" pageNumber="323" type="female">19 females</specimenCount>
in the lot labelled as 
<specimenCode id="DBCFE613FFB05856FC050424FBD9FC8B" box="[1017,1135,780,807]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF 5257</specimenCode>
sent to us, to the 
<specimenCount id="9D6F85E1FFB05856FAD70425FA2AFC8A" box="[1323,1436,780,807]" count="2" pageId="9" pageNumber="323" type="male">two males</specimenCount>
and 
<specimenCount id="9D6F85E1FFB05856FF3A0419FEF8FCE6" box="[198,334,816,843]" count="2" pageId="9" pageNumber="323" type="female">two females</specimenCount>
from the bottle in the 
<collectionCode id="ED78D6ADFFB05856FDBA0418FDCAFCE6" box="[582,636,816,842]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="323">ZRC</collectionCode>
without any labels, we have a total of 
<specimenCount id="9D6F85E1FFB05856FBD60418FB38FCE6" box="[1066,1166,816,843]" count="15" pageId="9" pageNumber="323" type="male">15 males</specimenCount>
and 
<specimenCount id="9D6F85E1FFB05856FB380418FAF7FCE6" box="[1220,1345,816,842]" count="21" pageId="9" pageNumber="323" type="female">21 females</specimenCount>
, exactly as stated in the 
<collectionCode id="ED78D6ADFFB05856FEBD047CFECFFCC2" box="[321,377,852,878]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF</collectionCode>
online catalogue. 
<collectorName id="269C2BBEFFB05856FDBE047CFDD7FCC2" box="[578,609,852,878]" pageId="9" pageNumber="323">As</collectorName>
such we are very confident the unlabelled bottle in the 
<collectionCode id="ED78D6ADFFB05856FB3A047CFB4AFCC2" box="[1222,1276,852,878]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="323">ZRC</collectionCode>
are part of the 
<specimenCode id="DBCFE613FFB05856FF6B0450FEA4FC3F" box="[151,274,888,915]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF 5257</specimenCode>
. 
<collectorName id="269C2BBEFFB05856FEE00451FEF7FC3F" box="[284,321,889,915]" pageId="9" pageNumber="323">We</collectorName>
believe the 
<specimenCount id="9D6F85E1FFB05856FE350451FD99FC3E" box="[457,559,888,915]" count="1" pageId="9" pageNumber="323" type="male">one male</specimenCount>
and 
<specimenCount id="9D6F85E1FFB05856FD9A0451FD54FC3E" box="[614,738,888,915]" count="1" pageId="9" pageNumber="323" type="female">one female</specimenCount>
in the bottle labelled as “ex 
<specimenCode id="DBCFE613FFB05856FBDF0450FB2AFC3F" box="[1059,1180,888,915]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF 5257</specimenCode>
” were either separated out early and given to 
<location id="8EB618B3FFB05856FE5904B5FDACFC1B" LSID="urn:lsid:plazi:treatment:03C0FF7EFFB85856FF6B05A7FD89FB7A:8EB618B3FFB05856FE5904B5FDACFC1B" box="[421,538,925,951]" municipality="Michael Turkay" name="Ngan Kee" pageId="9" pageNumber="323">Ngan Kee</location>
by 
<collectingMunicipality id="6BB2D412FFB05856FDB704B4FCB4FC1A" box="[587,770,924,950]" pageId="9" pageNumber="323">Michael Türkay</collectingMunicipality>
or already excluded from the online catalogue by recent update, and therefore the given specimens were not included in the 
<collectionCode id="ED78D6ADFFB05856FC7504E8FC77FC76" box="[905,961,960,986]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF</collectionCode>
catalogue. 
<collectorName id="269C2BBEFFB05856FBBD04E9FBEEFC77" box="[1089,1112,961,987]" pageId="9" pageNumber="323">In</collectorName>
any case, we have examined all the specimens and they are conspecific. 
<collectorName id="269C2BBEFFB05856FD7B04CDFD12FC53" box="[647,676,997,1023]" pageId="9" pageNumber="323">To</collectorName>
ensure the history of these specimens is properly documented and separated, we have labelled the specimens in 
<specimenCode id="DBCFE613FFB05856FD6E0320FCBFFB8F" box="[658,777,1032,1059]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF 5257</specimenCode>
separately even though they are all from the same location and collected at the same time. 
<location id="8EB618B3FFB05856FDF80304FD86FBEA" LSID="urn:lsid:plazi:treatment:03C0FF7EFFB85856FF6B05A7FD89FB7A:8EB618B3FFB05856FDF80304FD86FBEA" box="[516,560,1068,1094]" name="The" pageId="9" pageNumber="323">The</location>
<typeStatus id="54D2F0CAFFB05856FDC50304FD2DFBEA" box="[569,667,1068,1094]" pageId="9" pageNumber="323" type="holotype">holotype</typeStatus>
male was selected from the unlabelled 
<collectionCode id="ED78D6ADFFB05856FB980304FB2CFBEA" box="[1124,1178,1068,1094]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="323">ZRC</collectionCode>
bottle with 
<specimenCount id="9D6F85E1FFB05856FAD40305FA2AFBEA" box="[1320,1436,1068,1095]" count="2" pageId="9" pageNumber="323" type="male">two males</specimenCount>
and 
<specimenCount id="9D6F85E1FFB05856FF3A0379FEF8FBC6" box="[198,334,1104,1131]" count="2" pageId="9" pageNumber="323" type="female">two females</specimenCount>
as it was the most complete specimen and is here labelled as 
<specimenCode id="DBCFE613FFB05856FC060378FBC2FBC7" box="[1018,1140,1104,1131]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF 5257</specimenCode>
a, with the remaining 
<specimenCount id="9D6F85E1FFB05856FA980378FEBAFB22" count="3" pageId="9" pageNumber="323" type="generic">three specimens</specimenCount>
labelled as 
<specimenCode id="DBCFE613FFB05856FE69035CFDA7FB23" box="[405,529,1140,1167]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF 5257</specimenCode>
b. 
<collectorName id="269C2BBEFFB05856FDD6035CFDE0FB22" box="[554,598,1140,1166]" pageId="9" pageNumber="323">The</collectorName>
<specimenCount id="9D6F85E1FFB05856FDA2035CFD72FB22" box="[606,708,1140,1166]" count="13" pageId="9" pageNumber="323" type="male">13 males</specimenCount>
and 
<specimenCount id="9D6F85E1FFB05856FD01035CFCCFFB22" box="[765,889,1140,1166]" count="19" pageId="9" pageNumber="323" type="female">19 females</specimenCount>
most recently sent to us from 
<collectionCode id="ED78D6ADFFB05856FB2B035CFAB9FB22" box="[1239,1295,1140,1166]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF</collectionCode>
to complete this study are labelled as 
<specimenCode id="DBCFE613FFB05856FE5303B0FD9CFB1F" box="[431,554,1176,1203]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF 5257</specimenCode>
c. 
<collectorName id="269C2BBEFFB05856FDBC03B0FDDAFB1E" box="[576,620,1176,1202]" pageId="9" pageNumber="323">The</collectorName>
<specimenCount id="9D6F85E1FFB05856FD8E03B1FCA1FB1E" box="[626,791,1176,1203]" count="2" pageId="9" pageNumber="323" type="generic">two specimens</specimenCount>
given to 
<collectorName id="269C2BBEFFB05856FC8303B1FC59FB1F" box="[895,1007,1177,1203]" pageId="9" pageNumber="323">Ngan Kee</collectorName>
and with the label “ex 
<specimenCode id="DBCFE613FFB05856FB0C03B0FADFFB1F" box="[1264,1385,1176,1203]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="323">SMF 5257</specimenCode>
” are here recatalogued as 
<specimenCode id="DBCFE613FFB05856FE7F0394FD8DFB7A" box="[387,571,1212,1238]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="323">ZRC 2023.0014</specimenCode>
.
</paragraph>
</subSubSection>
</treatment>
</document>