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<document id="EA24BAAE9D814F6B6C2FF7F546848FBC" ID-DOI="10.5281/zenodo.5170031" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170031" ID-ZooBank="A5348E25-CC3E-476B-9AD2-0A6C3A37A61A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628280835588" checkinUser="felipe" docAuthor="Gordon, Robert D. &amp; Hanley, Guy A." docDate="2017" docId="038E8B10D849FFDEBF96E2E6290BFC11" docLanguage="en" docName="InsectaMundi.0601.1-158.pdf" docOrigin="Insecta Mundi 2017 (601)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Prodilis maryann Gordon and Hanley 2017, new species" docType="treatment" docVersion="6" lastPageNumber="93" masterDocId="FFB7F368D814FF80BF56E4652816FFCA" masterDocTitle="South American Coccinellidae (Coleoptera), Part XVII: systematic revision of Western Hemisphere Cephaloscymnini (Coccinellinae) with description of a cryptic new genus and species of Coccidulini (Coccinellinae)" masterLastPageNumber="158" masterPageNumber="1" pageNumber="92" updateTime="1699039770044" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="4919422F97BBD7B8AF690F32CED762A5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="64B58F6479026592ABCF639769F65642">
<mods:title id="CDA299A13D243BFF9B57CB0BD89A41E8">South American Coccinellidae (Coleoptera), Part XVII: systematic revision of Western Hemisphere Cephaloscymnini (Coccinellinae) with description of a cryptic new genus and species of Coccidulini (Coccinellinae)</mods:title>
</mods:titleInfo>
<mods:name id="D2B983DC90927F689A8299084DC979E1" type="personal">
<mods:role id="CAFB146CE0E79279137641A737E5788E">
<mods:roleTerm id="65710203C865ED113675503558F600ED">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E8BD1D009C4760FEFC23E67459A5D97A">Gordon, Robert D.</mods:namePart>
</mods:name>
<mods:name id="FB5D7A544704A8AE9141BBDB37BA6AA2" type="personal">
<mods:role id="E4F66B70E0CECC1896C75E8C8DF10028">
<mods:roleTerm id="382C3D0FAA02607B548B49CFB8882948">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7BCD6CD62444E7B0D7D7404546E1B771">Hanley, Guy A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B961440A087D9C43FAA13A23000627D8">text</mods:typeOfResource>
<mods:relatedItem id="CE638BE0C100C77C7B48222BFCFBD852" type="host">
<mods:titleInfo id="FCAD5E2EE07A570FA0058E3201254424">
<mods:title id="7DE7EB76C78AC82E5C769A5F4632A9BB">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="CCF96D954C0FA61892EF108C664BE686">
<mods:date id="EFD86490BE94CA72D67E0F55EC7A0A59">2017</mods:date>
<mods:detail id="4B26A94B1B66460405C376CF58902012" type="pubDate">
<mods:number id="7083D8238E4058A232107AFA10F0ED6F">2017-12-29</mods:number>
</mods:detail>
<mods:detail id="E78413B4B090275D8977DE58D20FF849" type="volume">
<mods:number id="28B0264113C5F04665AE8140A894F3A8">2017</mods:number>
</mods:detail>
<mods:detail id="E0F89696BA2F535734F624E26851D356" type="issue">
<mods:number id="1B855B4D1A5BE859AD6A7B07DF9D28F8">601</mods:number>
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<mods:start id="A19AE463B688AD4CBF0A567030583079">1</mods:start>
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<mods:classification id="1647DBD640B84BD69FAD6D923AA3674E">journal article</mods:classification>
<mods:identifier id="CD90F165E39F94147DD9D6E71A9AE22D" type="DOI">10.5281/zenodo.5170031</mods:identifier>
<mods:identifier id="AC1FBEDA6C2B837A5FDBB493A8F8C78C" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="59E430E65427FCD97FADD510D3B1BA1E" type="Zenodo-Dep">5170031</mods:identifier>
<mods:identifier id="9DA31D513738D9E4430DE024B00C9424" type="ZooBank">A5348E25-CC3E-476B-9AD2-0A6C3A37A61A</mods:identifier>
</mods:mods>
<treatment id="038E8B10D849FFDEBF96E2E6290BFC11" ID-DOI="http://doi.org/10.5281/zenodo.5186246" ID-Zenodo-Dep="5186246" LSID="urn:lsid:plazi:treatment:038E8B10D849FFDEBF96E2E6290BFC11" httpUri="http://treatment.plazi.org/id/038E8B10D849FFDEBF96E2E6290BFC11" lastPageId="94" lastPageNumber="93" pageId="93" pageNumber="92">
<subSubSection id="C33D698DD849FFDDBF96E2E62BBFF951" box="[192,937,1667,1691]" pageId="93" pageNumber="92" type="nomenclature">
<paragraph id="8B983A06D849FFDDBF96E2E62BBFF951" blockId="93.[192,937,1667,1691]" box="[192,937,1667,1691]" pageId="93" pageNumber="92">
<heading id="D0D08D6AD849FFDDBF96E2E62BBFF951" bold="true" box="[192,937,1667,1691]" fontSize="10" level="2" pageId="93" pageNumber="92" reason="3">
<emphasis id="B953E614D849FFDDBF96E2E62BBFF951" bold="true" box="[192,937,1667,1691]" pageId="93" pageNumber="92">
51.
<taxonomicName id="4C274185D849FFDDBFB8E2E62AEFF951" authority="Gordon and Hanley" authorityName="Gordon and Hanley" authorityYear="2017" box="[238,761,1667,1691]" class="Insecta" family="Coccinellidae" genus="Prodilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="92" phylum="Arthropoda" rank="species" species="maryann" status="sp. nov.">
<emphasis id="B953E614D849FFDDBFB8E2E629F4F950" bold="true" box="[238,482,1667,1691]" italics="true" pageId="93" pageNumber="92">Prodilis maryann</emphasis>
Gordon and Hanley
</taxonomicName>
,
<taxonomicNameLabel id="A2605B6FD849FFDDBC51E2E62BBFF951" box="[775,937,1667,1691]" pageId="93" pageNumber="92" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D698DD849FFDEBF96E2A629C7FD31" lastPageId="94" lastPageNumber="93" pageId="93" pageNumber="92" type="description">
<paragraph id="8B983A06D849FFDEBF96E2A62B74FDB1" blockId="93.[192,1441,1731,1915]" lastBlockId="94.[192,1441,195,635]" lastPageId="94" lastPageNumber="93" pageId="93" pageNumber="92">
<emphasis id="B953E614D849FFDDBF96E2A629A1F911" bold="true" box="[192,439,1731,1755]" pageId="93" pageNumber="92">Description. Male</emphasis>
<typeStatus id="549C84A4D849FFDDBE97E2A62A38F911" box="[449,558,1731,1755]" pageId="93" pageNumber="92" type="holotype">holotype</typeStatus>
. Length
<quantity id="4CDF97E3D849FFDDBDCAE2A62AEDF911" box="[668,763,1731,1755]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="93" pageNumber="92" unit="mm" value="2.4">2.4 mm</quantity>
, width
<quantity id="4CDF97E3D849FFDDBC0FE2A62BACF911" box="[857,954,1731,1755]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="93" pageNumber="92" unit="mm" value="1.9">1.9 mm</quantity>
; body oval, somewhat rounded, elytron with side rounded, wider than pronotal base, widest at middle of elytra. Dorsal surface entirely shiny. Color yellowish red; head greenish black, apical ½ yellow (
<figureCitation id="131C2683D849FFDDBC2AE3662BF5F8D1" box="[892,995,1795,1819]" captionStart="Figures 591–607" captionStartId="151.[192,287,1617,1639]" captionTargetBox="[206,1424,212,1590]" captionTargetId="figure-16@151.[197,1444,197,1628]" captionTargetPageId="151" captionText="Figures 591–607. Prodilis spp. 591–596) Prodilis eunice 591) Habitus. 592) Basal abdominal ventrites. 593) Frons. 594) Phallobase ventral. 595) Phallobase lateral. 596) Sipho, apex lost. 597–601) Prodilis angie 597) Habitus. 598) Basal abdominal ventrite. 599) Frons. 600) Phallobase ventral. 601) Phallobase lateral. 602–607) Prodilis maryann 602) Habitus. 603) Basal abdominal ventrites. 604) Frons. 605) Phallobase ventral. 606) Phallobase lateral. 607) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170103" httpUri="https://zenodo.org/record/5170103/files/figure.png" pageId="93" pageNumber="92">Fig. 604</figureCitation>
); pronotum greenish black, lateral 1/ 5 brown; elytron with apical 1/10 obscurely darkened, lateral reflexed border yellow (
<figureCitation id="131C2683D849FFDDBBEDE3462D36F8F1" box="[1211,1312,1827,1851]" captionStart="Figures 591–607" captionStartId="151.[192,287,1617,1639]" captionTargetBox="[206,1424,212,1590]" captionTargetId="figure-16@151.[197,1444,197,1628]" captionTargetPageId="151" captionText="Figures 591–607. Prodilis spp. 591–596) Prodilis eunice 591) Habitus. 592) Basal abdominal ventrites. 593) Frons. 594) Phallobase ventral. 595) Phallobase lateral. 596) Sipho, apex lost. 597–601) Prodilis angie 597) Habitus. 598) Basal abdominal ventrite. 599) Frons. 600) Phallobase ventral. 601) Phallobase lateral. 602–607) Prodilis maryann 602) Habitus. 603) Basal abdominal ventrites. 604) Frons. 605) Phallobase ventral. 606) Phallobase lateral. 607) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170103" httpUri="https://zenodo.org/record/5170103/files/figure.png" pageId="93" pageNumber="92">Fig. 602</figureCitation>
); antenna, mouthparts, epipleuron, legs yellow; abdomen yellow. Head punctures small, separated by a diameter or less; pronotal punctures as large as on head, separated by a diameter or less; elytral punctures larger than on pronotum, separated by less than twice a diameter; prosternal, mesosternal punctures large, separated by about a diameter; metasternal punctures as large as on mesosternum, separated by less than a diameter except more widely spaced medially; abdomen with punctures on ventrites 1–3 large, separated by less than twice a diameter, punctures on remaining ventrites smaller, separated by about a diameter. Head with frons not widened from vertex to clypeus, lateral margins parallel, 1.4 times width of eye measured at vertex; eye canthus short; apical maxillary palpomere short, widened from base to apex. Pronotum widest at middle, reflexed lateral margin wide, widest in anterior 1/3. Epipleuron nearly flat, wide in basal ½, as wide as pronotal hypomeron, with feeble depressions for reception of femoral apices. Prosternum slightly longer than wide, longer than mesosternum, base strongly arcuate medially, lateral carina slender, extended anterior to apex of procoxa. Postcoxal line on ventrite 1 long, slightly angulate, extended 2/3 distance to apex of ventrite (
<figureCitation id="131C2683D84AFFDEBC66E6662B80FDD1" box="[816,918,515,539]" captionStart="Figures 591–607" captionStartId="151.[192,287,1617,1639]" captionTargetBox="[206,1424,212,1590]" captionTargetId="figure-16@151.[197,1444,197,1628]" captionTargetPageId="151" captionText="Figures 591–607. Prodilis spp. 591–596) Prodilis eunice 591) Habitus. 592) Basal abdominal ventrites. 593) Frons. 594) Phallobase ventral. 595) Phallobase lateral. 596) Sipho, apex lost. 597–601) Prodilis angie 597) Habitus. 598) Basal abdominal ventrite. 599) Frons. 600) Phallobase ventral. 601) Phallobase lateral. 602–607) Prodilis maryann 602) Habitus. 603) Basal abdominal ventrites. 604) Frons. 605) Phallobase ventral. 606) Phallobase lateral. 607) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170103" httpUri="https://zenodo.org/record/5170103/files/figure.png" pageId="94" pageNumber="93">Fig. 603</figureCitation>
). Apex of ventrite 5 arcuate. Genitalia with slender phallobase, basal lobe longer than paramere, slender, narrowed from base to acute apex; paramere short, widest at middle, abruptly narrowed to abruptly rounded apex in apical ½, without marginal serrations (
<figureCitation id="131C2683D84AFFDEBE1AE60629F8FDB1" box="[332,494,611,635]" captionStart="Figures 591–607" captionStartId="151.[192,287,1617,1639]" captionTargetBox="[206,1424,212,1590]" captionTargetId="figure-16@151.[197,1444,197,1628]" captionTargetPageId="151" captionText="Figures 591–607. Prodilis spp. 591–596) Prodilis eunice 591) Habitus. 592) Basal abdominal ventrites. 593) Frons. 594) Phallobase ventral. 595) Phallobase lateral. 596) Sipho, apex lost. 597–601) Prodilis angie 597) Habitus. 598) Basal abdominal ventrite. 599) Frons. 600) Phallobase ventral. 601) Phallobase lateral. 602–607) Prodilis maryann 602) Habitus. 603) Basal abdominal ventrites. 604) Frons. 605) Phallobase ventral. 606) Phallobase lateral. 607) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170103" httpUri="https://zenodo.org/record/5170103/files/figure.png" pageId="94" pageNumber="93">Fig. 605, 606</figureCitation>
); sipho long, robust (
<figureCitation id="131C2683D84AFFDEBDB8E6062B43FDB1" box="[750,853,611,635]" captionStart="Figures 591–607" captionStartId="151.[192,287,1617,1639]" captionTargetBox="[206,1424,212,1590]" captionTargetId="figure-16@151.[197,1444,197,1628]" captionTargetPageId="151" captionText="Figures 591–607. Prodilis spp. 591–596) Prodilis eunice 591) Habitus. 592) Basal abdominal ventrites. 593) Frons. 594) Phallobase ventral. 595) Phallobase lateral. 596) Sipho, apex lost. 597–601) Prodilis angie 597) Habitus. 598) Basal abdominal ventrite. 599) Frons. 600) Phallobase ventral. 601) Phallobase lateral. 602–607) Prodilis maryann 602) Habitus. 603) Basal abdominal ventrites. 604) Frons. 605) Phallobase ventral. 606) Phallobase lateral. 607) Sipho." figureDoi="http://doi.org/10.5281/zenodo.5170103" httpUri="https://zenodo.org/record/5170103/files/figure.png" pageId="94" pageNumber="93">Fig. 607</figureCitation>
).
</paragraph>
<paragraph id="8B983A06D84AFFDEBF96E6C629A5FD71" blockId="94.[192,435,675,699]" box="[192,435,675,699]" pageId="94" pageNumber="93">
<emphasis id="B953E614D84AFFDEBF96E6C6293AFD71" bold="true" box="[192,300,675,699]" pageId="94" pageNumber="93">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B983A06D84AFFDEBF96E68629C7FD31" blockId="94.[192,465,739,763]" box="[192,465,739,763]" pageId="94" pageNumber="93">
<emphasis id="B953E614D84AFFDEBF96E6862954FD31" bold="true" box="[192,322,739,763]" pageId="94" pageNumber="93">Variation</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33D698DD84AFFDEBF96E7462C72FC91" pageId="94" pageNumber="93" type="materials_examined">
<paragraph id="8B983A06D84AFFDEBF96E7462C72FC91" blockId="94.[192,1440,802,860]" pageId="94" pageNumber="93">
<emphasis id="B953E614D84AFFDEBF96E746299FFCF1" bold="true" box="[192,393,803,827]" pageId="94" pageNumber="93">Type material.</emphasis>
<materialsCitation id="3B4F305BD84AFFDEBEC6E7462C72FC91" collectingDate="2000-01-05" collectionCode="USNM" country="Costa Rica" elevation="100" latitude="10.433333" location="Eugena" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Biol. La Selva" pageId="94" pageNumber="93" specimenCount="1" stateProvince="Heredia" typeStatus="holotype">
<typeStatus id="549C84A4D84AFFDEBEC6E74629E9FCF1" box="[400,511,803,827]" pageId="94" pageNumber="93" type="holotype">Holotype</typeStatus>
male;
<collectingCountry id="F3307A96D84AFFDEBD18E7462AEFFCF1" box="[590,761,803,827]" name="Costa Rica" pageId="94" pageNumber="93">COSTA RICA</collectingCountry>
:
<collectingRegion id="49E3F4E4D84AFFDEBC5FE7462B78FCF1" box="[777,878,803,827]" country="Costa Rica" name="Heredia" pageId="94" pageNumber="93">Heredia</collectingRegion>
,
<collectingMunicipality id="6BFCA07CD84AFFDEBC2EE7412C4CFCF1" box="[888,1114,803,827]" pageId="94" pageNumber="93">Est. Biol. La Selva</collectingMunicipality>
,
<quantity id="4CDF97E3D84AFFDEBB32E7462CC5FCF1" box="[1124,1235,803,827]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="94" pageNumber="93" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">
<elevation id="000ADD35D84AFFDEBB32E7462CC5FCF1" box="[1124,1235,803,827]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="94" pageNumber="93" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50–150m</elevation>
</quantity>
<geoCoordinate id="EE135CC1D84AFFDEBB8FE7462D21FCF1" box="[1241,1335,802,827]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="94" pageNumber="93" precision="925" value="10.433333">
10
<superScript id="7C52974ED84AFFDEBBA0E7472CE8FCFA" attach="none" box="[1270,1278,802,816]" fontSize="6" pageId="94" pageNumber="93">o</superScript>
26'N
</geoCoordinate>
<geoCoordinate id="EE135CC1D84AFFDEBA6BE7462D8DFCF1" box="[1341,1435,802,827]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="94" pageNumber="93" precision="925" value="-84.01667">
84
<superScript id="7C52974ED84AFFDEBA0CE7472D74FCFA" attach="none" box="[1370,1378,802,816]" fontSize="6" pageId="94" pageNumber="93">o</superScript>
01'W
</geoCoordinate>
, Prov. ALAS, INBio–OET, FT/46/01–40,
<location id="8EF86CDDD84AFFDEBDF6E7212AEBFC96" LSID="urn:lsid:plazi:treatment:038E8B10D849FFDEBF96E2E6290BFC11:8EF86CDDD84AFFDEBDF6E7212AEBFC96" box="[672,765,836,860]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Biol. La Selva" name="Eugena" pageId="94" pageNumber="93" stateProvince="Heredia">Eugena</location>
sp.,
<date id="FF991CC6D84AFFDEBC63E7262BFCFC91" box="[821,1002,835,859]" pageId="94" pageNumber="93" value="2000-01-05">
<collectingDate id="EFDDE52ED84AFFDEBC63E7262BFCFC91" box="[821,1002,835,859]" pageId="94" pageNumber="93" value="2000-01-05">05 Enero 2000</collectingDate>
</date>
. (
<collectionCode id="ED36A2C3D84AFFDEBCABE7262C4EFC91" box="[1021,1112,835,859]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="94" pageNumber="93" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D698DD84AFFDEBF96E7E6290BFC11" pageId="94" pageNumber="93" type="discussion">
<paragraph id="8B983A06D84AFFDEBF96E7E6290BFC11" blockId="94.[192,1440,899,987]" pageId="94" pageNumber="93">
<emphasis id="B953E614D84AFFDEBF96E7E62952FC51" bold="true" box="[192,324,899,923]" pageId="94" pageNumber="93">Remarks.</emphasis>
One of several species with dorsal coloration similar to
<taxonomicName id="4C274185D84AFFDEBCB8E7E12C88FC51" authorityName="Gorham" authorityYear="1897" box="[1006,1182,899,923]" class="Insecta" family="Coccinellidae" genus="Neaporia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="93" phylum="Arthropoda" rank="species" species="chiriquensis">
<emphasis id="B953E614D84AFFDEBCB8E7E12C88FC51" box="[1006,1182,899,923]" italics="true" pageId="94" pageNumber="93">P. chiriquensis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C274185D84AFFDEBBFAE7E62D93FC51" authorityName="Gordon and Hanley" authorityYear="2017" box="[1196,1413,899,923]" class="Insecta" family="Coccinellidae" genus="Prodilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="93" phylum="Arthropoda" rank="species" species="maryann">
<emphasis id="B953E614D84AFFDEBBFAE7E62D93FC51" box="[1196,1413,899,923]" italics="true" pageId="94" pageNumber="93">Prodilis maryann</emphasis>
</taxonomicName>
is distinguished from those species by a nearly total lack of dark coloration on the elytral apex. See key to species.
</paragraph>
</subSubSection>
</treatment>
</document>
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