261 lines
37 KiB
XML
261 lines
37 KiB
XML
<document id="A885FC901288043DDA5CFFC60F364306" ID-CLB-Dataset="28065" ID-DOI="10.11646/zootaxa.4504.4.5" ID-GBIF-Dataset="343a6a67-e8bf-467c-9818-de9621bbe955" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606614" ID-ZooBank="08E1A79F-BAA4-4EAB-9636-FE44BF7EB7F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585132792" checkinUser="plazi" docAuthor="Jäger, Peter" docDate="2018" docId="03AB87D4E76C6F6712F9FB9ACD977F8A" docLanguage="en" docName="zootaxa.4504.4.5.pdf" docOrigin="Zootaxa 4504 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systaria longinqua Jäger 2018, spec. nov." docType="treatment" docVersion="8" lastPageNumber="529" masterDocId="FF92FFACE76F6F62126EFFC4C9517912" masterDocTitle="On the genus Systaria (Araneae: Clubionidae) in Southeast Asia: new species from caves and forests" masterLastPageNumber="544" masterPageNumber="524" pageNumber="527" updateTime="1698712839155" updateUser="ExternalLinkService">
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<mods:title id="85771B9CB8AB18C0277DF7572B14E73E">On the genus Systaria (Araneae: Clubionidae) in Southeast Asia: new species from caves and forests</mods:title>
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<mods:name id="95F132114023F384F341810D87C19F5A" type="personal">
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<mods:namePart id="5CA4B919048D115C5882188168324F7F">Jäger, Peter</mods:namePart>
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<mods:title id="8F3767AF4D0F21E220B659FAF991D496">Zootaxa</mods:title>
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<mods:part id="5FB58888F7B66CE6F1440A81764C50A5">
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<mods:date id="255871219BB44FE19EDF8C2040DFD35F">2018</mods:date>
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<mods:number id="9DD2BDCDA455D390736501B0A3990D04">2018-10-26</mods:number>
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<mods:number id="325573253F2DDD2C528476D0A7FCC830">4504</mods:number>
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<treatment id="03AB87D4E76C6F6712F9FB9ACD977F8A" ID-DOI="http://doi.org/10.5281/zenodo.5988004" ID-GBIF-Taxon="154495197" ID-Zenodo-Dep="5988004" LSID="urn:lsid:plazi:treatment:03AB87D4E76C6F6712F9FB9ACD977F8A" httpUri="http://treatment.plazi.org/id/03AB87D4E76C6F6712F9FB9ACD977F8A" lastPageId="5" lastPageNumber="529" pageId="3" pageNumber="527">
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<paragraph id="8BBD36C2E76C6F6112F9FB9AC8AB7D6A" blockId="3.[151,506,1118,1178]" box="[151,506,1118,1144]" pageId="3" pageNumber="527">
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<heading id="D0F581AEE76C6F6112F9FB9AC8AB7D6A" bold="true" box="[151,506,1118,1144]" fontSize="11" level="1" pageId="3" pageNumber="527" reason="1">
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<emphasis id="B976EAD0E76C6F6112F9FB9AC8AB7D6A" bold="true" box="[151,506,1118,1144]" pageId="3" pageNumber="527">
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<taxonomicName id="4C024D41E76C6F6112F9FB9AC82A7D6A" authority="Jäger, 2018" authorityName="Jäger" authorityYear="2018" box="[151,379,1118,1144]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="longinqua" status="sp. nov.">
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<emphasis id="B976EAD0E76C6F6112F9FB9AC82A7D6A" bold="true" box="[151,379,1118,1144]" italics="true" pageId="3" pageNumber="527">Systaria longinqua</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24557ABE76C6F6113EDFB9AC8AB7D6A" box="[387,506,1118,1144]" pageId="3" pageNumber="527" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3186549E76C6F6112F9FB45C8D57D88" box="[151,388,1153,1178]" pageId="3" pageNumber="527" type="description">
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<paragraph id="8BBD36C2E76C6F6112F9FB45C8D57D88" blockId="3.[151,506,1118,1178]" box="[151,388,1153,1178]" pageId="3" pageNumber="527">
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<figureCitation id="13392A47E76C6F6112F9FB45C8597D88" box="[151,264,1153,1178]" captionStart-0="FIGURES 6–10" captionStart-1="FIGURES 11–15" captionStartId-0="4.[151,263,1811,1834]" captionStartId-1="5.[151,263,1396,1419]" captionTargetBox-0="[161,1433,593,1765]" captionTargetBox-1="[151,1435,193,1383]" captionTargetId-0="figure@4.[161,1433,593,1799]" captionTargetId-1="figure@5.[151,1435,193,1383]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 6–10. Systaria longinqua spec. nov., male from N Laos, Tham Pathok. 6–8 Left palp (6 ventral, 7 retrolateral, 8 dorsal). 9 Right chelicera, ventral. 10 Eye arrangement, dorsal. Abbreviations: C—Conductor, CT—Cymbial tubercle, E— Embolus, RTA—Retrolateral tibial apophysis, Sp—Spermophor [9–10 not to scale]." captionText-1="FIGURES 11–15. Systaria longinqua spec. nov., holotype female from N Laos, Tham Pathok. 11 Epigyne, ventral. 12 Vulva, dorsal. 13 Left chelicera, ventral. 14 Prosoma, dorsal. 15 Spinnerets, ventral (not all setae drawn). Abbreviation: COcopulatory opening." httpUri-0="https://zenodo.org/record/2606638/files/figure.png" httpUri-1="https://zenodo.org/record/2606640/files/figure.png" pageId="3" pageNumber="527">Figs 6–15</figureCitation>
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,
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<figureCitation id="13392A47E76C6F61137AFB46C8637D88" box="[276,306,1154,1178]" captionStart="FIGURES 18–19" captionStartId="7.[151,263,677,700]" captionTargetBox="[158,1431,208,642]" captionTargetId="figure@7.[151,1436,193,657]" captionTargetPageId="7" captionText="FIGURES 18–19. Systaria spp., live habitus. 18 Systaria longinqua spec. nov., holotype female from N Laos, Tham Pathok. 19 Systaria procera spec. nov., female paratype from NW Cambodia, La Ang Pracheu (19 photo courtesy of H. Steiner)." httpUri="https://zenodo.org/record/2606646/files/figure.png" pageId="3" pageNumber="527">18</figureCitation>
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,
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<figureCitation id="13392A47E76C6F611350FB45C8D57D88" box="[318,388,1153,1178]" captionStart-0="FIGURE 45" captionStart-1="FIGURE 46" captionStartId-0="18.[151,250,1002,1024]" captionStartId-1="18.[151,250,1733,1755]" captionTargetBox-0="[150,1436,194,981]" captionTargetBox-1="[150,1436,1106,1712]" captionTargetId-0="figure@18.[150,1436,193,981]" captionTargetId-1="figure@18.[150,1436,1105,1712]" captionTargetPageId-0="18" captionTargetPageId-1="18" captionText-0="FIGURE 45. Records of Systaria spp. in SE Asia described in this paper. Triangles—species recorded from caves, circlesspecies recorded from forest." captionText-1="FIGURE 46. All known records of the genus Systaria (World Spider Catalog 2018) including undescribed species mentioned in Deeleman-Reinhold (2001) according to habitats. Triangles—species recorded from caves, circles—species recorded from forest. See discussion." httpUri-0="https://zenodo.org/record/2606658/files/figure.png" httpUri-1="https://zenodo.org/record/2606660/files/figure.png" pageId="3" pageNumber="527">45–46</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3186549E76C6F6112F9FB00CBB77C52" pageId="3" pageNumber="527" type="materials_examined">
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<paragraph id="8BBD36C2E76C6F6112F9FB00CDEB7DEC" blockId="3.[151,1437,1220,2011]" pageId="3" pageNumber="527">
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<emphasis id="B976EAD0E76C6F6112F9FB00C81B7DCF" bold="true" box="[151,330,1220,1245]" pageId="3" pageNumber="527">Type material:</emphasis>
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<materialsCitation id="3B6A3C9FE76C6F61133AFB00CDEB7DEC" ID-GBIF-Occurrence="2028138109" collectingDate="2006-10-03" collectionCode="SMF" collectorName="P. Jager & J. Altmann" country="Laos" elevation="373" latitude="20.551367" location="Tham Pathok" longLatPrecision="1" longitude="102.63208" municipality="Nong Khiao" pageId="3" pageNumber="527" specimenCount="1" stateProvince="Louangphabang" typeStatus="holotype">
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<collectingCountry id="F3157652E76C6F61133AFB00C8F37DCE" box="[340,418,1220,1244]" name="Laos" pageId="3" pageNumber="527">LAOS</collectingCountry>
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:
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<emphasis id="B976EAD0E76C6F6113C1FB01CB887DCE" box="[431,729,1220,1245]" italics="true" pageId="3" pageNumber="527">
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<collectingRegion id="49C6F820E76C6F6113C1FB01CB827DCE" box="[431,723,1220,1245]" country="Laos" name="Louangphabang" pageId="3" pageNumber="527">Luang Prabang Province</collectingRegion>
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:
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</emphasis>
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<emphasis id="B976EAD0E76C6F61108AFB00CA007DCF" bold="true" box="[740,849,1220,1245]" pageId="3" pageNumber="527">
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<typeStatus id="54B98860E76C6F61108AFB00CA007DCF" box="[740,849,1220,1245]" pageId="3" pageNumber="527" type="holotype">Holotype</typeStatus>
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</emphasis>
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: female,
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<collectingMunicipality id="6BD9ACB8E76C6F6111AEFB01CD1E7DCF" box="[960,1103,1220,1245]" pageId="3" pageNumber="527">Nong Khiao</collectingMunicipality>
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,
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<location id="8EDD6019E76C6F611630FB00CDAB7DCF" LSID="urn:lsid:plazi:treatment:03AB87D4E76C6F6712F9FB9ACD977F8A:8EDD6019E76C6F611630FB00CDAB7DCF" box="[1118,1274,1220,1245]" country="Laos" latitude="20.551367" longLatPrecision="1" longitude="102.63208" municipality="Nong Khiao" name="Tham Pathok" pageId="3" pageNumber="527" stateProvince="Louangphabang">Tham Pathok</location>
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,
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<geoCoordinate id="EE365005E76C6F611769FB00CCC67DCE" box="[1287,1431,1220,1244]" direction="north" orientation="latitude" pageId="3" pageNumber="527" precision="1" value="20.551367">20°33.082'N</geoCoordinate>
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,
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<geoCoordinate id="EE365005E76C6F6112F9FB21C8607DEC" box="[151,305,1253,1278]" direction="east" orientation="longitude" pageId="3" pageNumber="527" precision="1" value="102.63208">102°37.925'E</geoCoordinate>
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,
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<quantity id="4CFA9B27E76C6F611353FB22C8D57DEF" box="[317,388,1254,1278]" metricMagnitude="2" metricUnit="m" metricValue="3.7299999999999995" pageId="3" pageNumber="527" unit="m" value="373.0">
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<elevation id="002FD1F1E76C6F611353FB22C8D57DEF" box="[317,388,1254,1278]" metricMagnitude="2" metricUnit="m" metricValue="3.7299999999999995" pageId="3" pageNumber="527" unit="m" value="373.0">373 m</elevation>
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</quantity>
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elevation, in cave,
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<collectorName id="26F75314E76C6F611030FB22CBE27DEC" box="[606,691,1254,1278]" pageId="3" pageNumber="527">P. Jäger</collectorName>
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&
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<collectorName id="26F75314E76C6F6110B9FB22CA037DEC" box="[727,850,1253,1278]" pageId="3" pageNumber="527">J. Altmann</collectorName>
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leg. by hand,
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<date id="FFBC1002E76C6F61119FFB21CD307DEC" box="[1009,1121,1253,1278]" pageId="3" pageNumber="527" value="2006-10-03">
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<collectingDate id="EFF8E9EAE76C6F61119FFB21CD307DEC" box="[1009,1121,1253,1278]" pageId="3" pageNumber="527" value="2006-10-03">10.3.2006</collectingDate>
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</date>
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(
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<collectionCode id="ED13AE07E76C6F61161EFB22CDFF7DEC" box="[1136,1198,1254,1278]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="527">SMF</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BBD36C2E76C6F6112A9FAC2CBB77C52" blockId="3.[151,1437,1220,2011]" pageId="3" pageNumber="527">
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<emphasis id="B976EAD0E76C6F6112A9FAC2CB7A7C0D" bold="true" box="[199,555,1286,1311]" pageId="3" pageNumber="527">Additional material examined</emphasis>
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.
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<materialsCitation id="3B6A3C9FE76C6F611057FAC3CBB37C52" ID-GBIF-Occurrence="2028138110" collectingDate="2007-03-18" collectingDateMax="2007-10-20" collectingDateMin="2007-03-18" collectionCode="SMF" collectorName="P. Jager & F. Steinmetz" country="Laos" elevation="373" latitude="20.551367" location="Tham Pathok" longLatPrecision="1" longitude="102.63208" municipality="Nong Khiao" pageId="3" pageNumber="527" specimenCount="1" specimenCount-male="1" stateProvince="Louangphabang">
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<specimenCount id="9D04FD4BE76C6F611057FAC3CBD77C0D" box="[569,646,1286,1311]" pageId="3" pageNumber="527" type="male">1 male</specimenCount>
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, with same data as for holotype except for:
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<collectorName id="26F75314E76C6F611604FAC3CDEE7C0D" box="[1130,1215,1287,1311]" pageId="3" pageNumber="527">P. Jäger</collectorName>
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&
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<collectorName id="26F75314E76C6F61168CFAC3CC3D7C0D" box="[1250,1388,1286,1311]" pageId="3" pageNumber="527">F. Steinmetz</collectorName>
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leg.
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||
<date id="FFBC1002E76C6F6112F9FAECC8567C52" box="[151,263,1320,1344]" pageId="3" pageNumber="527" value="2007-03-18">
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||
<collectingDate id="EFF8E9EAE76C6F6112F9FAECC8567C52" box="[151,263,1320,1344]" pageId="3" pageNumber="527" value="2007-03-18">18.3.2007</collectingDate>
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||
</date>
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as subadult, adult
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<date id="FFBC1002E76C6F6113B6FAECCB077C52" box="[472,598,1320,1344]" pageId="3" pageNumber="527" value="2007-10-20">
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<collectingDate id="EFF8E9EAE76C6F6113B6FAECCB077C52" box="[472,598,1320,1344]" pageId="3" pageNumber="527" value="2007-10-20">20.10.2007</collectingDate>
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||
</date>
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||
(
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<collectionCode id="ED13AE07E76C6F61100BFAECCBF37C52" box="[613,674,1320,1344]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="527">SMF</collectionCode>
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||
; lost)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3186549E76C6F6112A9FA8DC8317CB7" pageId="3" pageNumber="527" type="etymology">
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||
<paragraph id="8BBD36C2E76C6F6112A9FA8DC8317CB7" blockId="3.[151,1437,1220,2011]" pageId="3" pageNumber="527">
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<emphasis id="B976EAD0E76C6F6112A9FA8DC81F7C70" bold="true" box="[199,334,1353,1378]" pageId="3" pageNumber="527">Etymology.</emphasis>
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The species name is derived from the Latin adjective “longinquus”, meaning among others “distant, far away” and referring to the unusually wide distance between the bursae in the female internal duct system; adjective.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3186549E76C6F6112A9FA69CCCD7F9C" pageId="3" pageNumber="527" type="diagnosis">
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||
<paragraph id="8BBD36C2E76C6F6112A9FA69CCCD7F9C" blockId="3.[151,1437,1220,2011]" pageId="3" pageNumber="527">
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||
<emphasis id="B976EAD0E76C6F6112A9FA69C8107CD4" bold="true" box="[199,321,1453,1478]" pageId="3" pageNumber="527">Diagnosis.</emphasis>
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Small spiders with body length of females: 9.9. Males of the new species are similar to those of
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<taxonomicName id="4C024D41E76C6F6117E9FA6ACB1E7CF5" authority="Dankittipakul & Singtripop 2011" authorityName="Dankittipakul & Singtripop" authorityYear="2011" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="lanna">
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||
<emphasis id="B976EAD0E76C6F6117E9FA6AC9867CF5" italics="true" pageId="3" pageNumber="527">S. lanna</emphasis>
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<bibRefCitation id="EF934B33E76C6F6112B3FA0ACB1E7CF5" author="Dankittipakul, P. & Singtripop, T." box="[221,591,1486,1511]" pageId="3" pageNumber="527" pagination="16 - 32" refId="ref11494" refString="Dankittipakul, P. & Singtripop, T. (2011) Seven new species of Systaria Simon, 1897 from southeast Asia (Araneae, Clubionidae, Systariinae). Zootaxa, 2905, 16 - 32." type="journal article" year="2011">
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||
Dankittipakul & Singtripop
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||
<quantity id="4CFA9B27E76C6F611076FA0BCB1E7CF5" box="[536,591,1487,1511]" metricMagnitude="1" metricUnit="m" metricValue="5.10794" pageId="3" pageNumber="527" unit="in" value="2011.0">2011</quantity>
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</bibRefCitation>
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</taxonomicName>
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in having a similar palpal conformation especially a similarly shaped RTA, tegulum and embolus (
|
||
<figureCitation id="13392A47E76C6F6113F0FA2BCB577F1A" box="[414,518,1519,1544]" captionStart="FIGURES 6–10" captionStartId="4.[151,263,1811,1834]" captionTargetBox="[161,1433,593,1765]" captionTargetId="figure@4.[161,1433,593,1799]" captionTargetPageId="4" captionText="FIGURES 6–10. Systaria longinqua spec. nov., male from N Laos, Tham Pathok. 6–8 Left palp (6 ventral, 7 retrolateral, 8 dorsal). 9 Right chelicera, ventral. 10 Eye arrangement, dorsal. Abbreviations: C—Conductor, CT—Cymbial tubercle, E— Embolus, RTA—Retrolateral tibial apophysis, Sp—Spermophor [9–10 not to scale]." httpUri="https://zenodo.org/record/2606638/files/figure.png" pageId="3" pageNumber="527">Figs 6–8</figureCitation>
|
||
), but can be distinguished by disto-ventral process of RTA short and straight in retrolateral view (longer and distinctly hooked in
|
||
<taxonomicName id="4C024D41E76C6F6110ABF9D6CA707F38" authorityName="Dankittipakul & Singtripop" authorityYear="2011" box="[709,801,1553,1578]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="lanna">
|
||
<emphasis id="B976EAD0E76C6F6110ABF9D6CA707F38" box="[709,801,1553,1578]" italics="true" pageId="3" pageNumber="527">S. lanna</emphasis>
|
||
</taxonomicName>
|
||
). Females are similar to those of
|
||
<taxonomicName id="4C024D41E76C6F6116F4F9D6CDA77F38" authorityName="Dankittipakul & Singtripop" authorityYear="2011" box="[1178,1270,1553,1578]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="lanna">
|
||
<emphasis id="B976EAD0E76C6F6116F4F9D6CDA77F38" box="[1178,1270,1553,1578]" italics="true" pageId="3" pageNumber="527">S. lanna</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C024D41E76C6F611743F9D6CCCD7F38" box="[1325,1436,1553,1578]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="mengla">
|
||
<emphasis id="B976EAD0E76C6F611743F9D6CCCD7F38" box="[1325,1436,1553,1578]" italics="true" pageId="3" pageNumber="527">S. mengla</emphasis>
|
||
</taxonomicName>
|
||
(Song & Zhu 1994) in having a similar conformation of the epigyne including course of atrial rims and position of copulatory openings (
|
||
<figureCitation id="13392A47E76C6F6113E7F990CB587F7E" box="[393,521,1620,1645]" captionStart="FIGURES 11–15" captionStartId="5.[151,263,1396,1419]" captionTargetBox="[151,1435,193,1383]" captionTargetId="figure@5.[151,1435,193,1383]" captionTargetPageId="5" captionText="FIGURES 11–15. Systaria longinqua spec. nov., holotype female from N Laos, Tham Pathok. 11 Epigyne, ventral. 12 Vulva, dorsal. 13 Left chelicera, ventral. 14 Prosoma, dorsal. 15 Spinnerets, ventral (not all setae drawn). Abbreviation: COcopulatory opening." httpUri="https://zenodo.org/record/2606640/files/figure.png" pageId="3" pageNumber="527">Figs 11–12</figureCitation>
|
||
), but may be distinguished by the bursae being distinctly shorter and more widely separated from each other over their entire length (almost touching each other anteriorly in
|
||
<taxonomicName id="4C024D41E76C6F6116E3F9B2CDB87F9C" authorityName="Dankittipakul & Singtripop" authorityYear="2011" box="[1165,1257,1653,1678]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="lanna">
|
||
<emphasis id="B976EAD0E76C6F6116E3F9B2CDB87F9C" box="[1165,1257,1653,1678]" italics="true" pageId="3" pageNumber="527">S. lanna</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C024D41E76C6F611770F9B2CCDA7F9C" box="[1310,1419,1653,1678]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="mengla">
|
||
<emphasis id="B976EAD0E76C6F611770F9B2CCDA7F9C" box="[1310,1419,1653,1678]" italics="true" pageId="3" pageNumber="527">S. mengla</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3186549E76C6F6612A9F952CDE27B0D" lastPageId="4" lastPageNumber="528" pageId="3" pageNumber="527" type="description">
|
||
<paragraph id="8BBD36C2E76C6F6612A9F952CCD97824" blockId="3.[151,1437,1220,2011]" lastBlockId="4.[151,1437,151,543]" lastPageId="4" lastPageNumber="528" pageId="3" pageNumber="527">
|
||
<emphasis id="B976EAD0E76C6F6112A9F952C8087FBD" bold="true" box="[199,345,1686,1711]" pageId="3" pageNumber="527">Description.</emphasis>
|
||
<emphasis id="B976EAD0E76C6F61130AF952C8E87FBD" box="[356,441,1686,1711]" italics="true" pageId="3" pageNumber="527">Female</emphasis>
|
||
(
|
||
<typeStatus id="54B98860E76C6F6113A3F952CB677FBD" box="[461,566,1686,1711]" pageId="3" pageNumber="527" type="holotype">holotype</typeStatus>
|
||
): prosoma length 4.8, prosoma width 3.4, anterior width of prosoma 2.2, opisthosoma length 5.1, opisthosoma width 3.0. Eyes (
|
||
<figureCitation id="13392A47E76C6F611175F973CA227FC2" box="[795,883,1719,1744]" captionStart="FIGURES 11–15" captionStartId="5.[151,263,1396,1419]" captionTargetBox="[151,1435,193,1383]" captionTargetId="figure@5.[151,1435,193,1383]" captionTargetPageId="5" captionText="FIGURES 11–15. Systaria longinqua spec. nov., holotype female from N Laos, Tham Pathok. 11 Epigyne, ventral. 12 Vulva, dorsal. 13 Left chelicera, ventral. 14 Prosoma, dorsal. 15 Spinnerets, ventral (not all setae drawn). Abbreviation: COcopulatory opening." httpUri="https://zenodo.org/record/2606640/files/figure.png" pageId="3" pageNumber="527">Fig. 14</figureCitation>
|
||
): AME 0.24, ALE 0.20, PME 0.15, PLE 0.21, AME–AME 0.10, AME–ALE 0.06, PME–PME 0.34, PME–PLE 0.26, AME–PME 0.22, ALE–PLE 0.05. Cheliceral furrow with 5(4) promarginal and 2 retromarginal teeth (
|
||
<figureCitation id="13392A47E76C6F6111F9F93ECABD7E01" box="[919,1004,1786,1811]" captionStart="FIGURES 11–15" captionStartId="5.[151,263,1396,1419]" captionTargetBox="[151,1435,193,1383]" captionTargetId="figure@5.[151,1435,193,1383]" captionTargetPageId="5" captionText="FIGURES 11–15. Systaria longinqua spec. nov., holotype female from N Laos, Tham Pathok. 11 Epigyne, ventral. 12 Vulva, dorsal. 13 Left chelicera, ventral. 14 Prosoma, dorsal. 15 Spinnerets, ventral (not all setae drawn). Abbreviation: COcopulatory opening." httpUri="https://zenodo.org/record/2606640/files/figure.png" pageId="3" pageNumber="527">Fig. 13</figureCitation>
|
||
). Spination: Palp: femur d012, patella p1, tibia p21, d11, metatarsus p21, d10, r10, v022; legs: femur I p011, d111, r110, II p011, d111, r111, III p1111, d111, r1111, IV p111, d111, r001; patella I–IV 000; tibia I v200, II v100, III p11, d10, r11, v212, IV p11, d10, r11, v222; metatarsus I–II v200, III p112, r112, v222, IV p122, r122, v222. Leg formula: 4123. Measurements of palp and legs: Palp 5.2 (1.8, 0.8, 1.1, -, 1.5), I 13.9 (4.0, 1.9, 3.6, 2.9, 1.5), II 12.8 (3.6, 1.7, 3.4, 2.7, 1.4), III 12.4 (3.5, 1.5, 2.9, 3.0, 1.5), IV 16.9 (4.5, 1.9, 4.2, 4.5, 1.8). Copulatory organ as in diagnosis (
|
||
<figureCitation id="13392A47E76C6F61160DF865CDB47EA8" box="[1123,1253,1953,1978]" captionStart="FIGURES 11–15" captionStartId="5.[151,263,1396,1419]" captionTargetBox="[151,1435,193,1383]" captionTargetId="figure@5.[151,1435,193,1383]" captionTargetPageId="5" captionText="FIGURES 11–15. Systaria longinqua spec. nov., holotype female from N Laos, Tham Pathok. 11 Epigyne, ventral. 12 Vulva, dorsal. 13 Left chelicera, ventral. 14 Prosoma, dorsal. 15 Spinnerets, ventral (not all setae drawn). Abbreviation: COcopulatory opening." httpUri="https://zenodo.org/record/2606640/files/figure.png" pageId="3" pageNumber="527">Figs 11–12</figureCitation>
|
||
). Epigynal field slightly wider than long, with 1 pair of slit sensilla, without anterior bands. Sclerotised spheres (~75) tiny, visible through cuticle in ventral view. Anterior margin of atrium arched, indistinct, atrial rims forming two U-shaped structures leading to blind-ending pockets. Copulatory openings located laterally posterior to atrial rims. Internal duct system with moderately large stout oval bursae. Connecting duct narrow, spermathecae separated by more than one of their diameter, fertilisation ducts antero-laterad. Colouration. Yellowish-brown without pattern. Fovea, eye region, chelicerae, labium gnathocoxae and sternum darker. Live specimens with shiny appearance (
|
||
<figureCitation id="13392A47E76B6F661749FED9CC2A7824" box="[1319,1403,285,310]" captionStart="FIGURES 18–19" captionStartId="7.[151,263,677,700]" captionTargetBox="[158,1431,208,642]" captionTargetId="figure@7.[151,1436,193,657]" captionTargetPageId="7" captionText="FIGURES 18–19. Systaria spp., live habitus. 18 Systaria longinqua spec. nov., holotype female from N Laos, Tham Pathok. 19 Systaria procera spec. nov., female paratype from NW Cambodia, La Ang Pracheu (19 photo courtesy of H. Steiner)." httpUri="https://zenodo.org/record/2606646/files/figure.png" pageId="4" pageNumber="528">Fig. 18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD36C2E76B6F6612A9FEFACDE27B0D" blockId="4.[151,1437,151,543]" pageId="4" pageNumber="528">
|
||
<emphasis id="B976EAD0E76B6F6612A9FEFAC8507845" box="[199,257,318,343]" italics="true" pageId="4" pageNumber="528">Male</emphasis>
|
||
(SMF; lost after drawing): cheliceral furrow with 3 promarginal and 2 retromarginal teeth (
|
||
<figureCitation id="13392A47E76B6F66176AFEFACC197845" box="[1284,1352,318,343]" captionStart="FIGURES 6–10" captionStartId="4.[151,263,1811,1834]" captionTargetBox="[161,1433,593,1765]" captionTargetId="figure@4.[161,1433,593,1799]" captionTargetPageId="4" captionText="FIGURES 6–10. Systaria longinqua spec. nov., male from N Laos, Tham Pathok. 6–8 Left palp (6 ventral, 7 retrolateral, 8 dorsal). 9 Right chelicera, ventral. 10 Eye arrangement, dorsal. Abbreviations: C—Conductor, CT—Cymbial tubercle, E— Embolus, RTA—Retrolateral tibial apophysis, Sp—Spermophor [9–10 not to scale]." httpUri="https://zenodo.org/record/2606638/files/figure.png" pageId="4" pageNumber="528">Fig. 9</figureCitation>
|
||
). AME largest, anterior eyes closer to each other than those of posterior row (
|
||
<figureCitation id="13392A47E76B6F6611ADFE9BCD48786A" box="[963,1049,351,376]" captionStart="FIGURES 6–10" captionStartId="4.[151,263,1811,1834]" captionTargetBox="[161,1433,593,1765]" captionTargetId="figure@4.[161,1433,593,1799]" captionTargetPageId="4" captionText="FIGURES 6–10. Systaria longinqua spec. nov., male from N Laos, Tham Pathok. 6–8 Left palp (6 ventral, 7 retrolateral, 8 dorsal). 9 Right chelicera, ventral. 10 Eye arrangement, dorsal. Abbreviations: C—Conductor, CT—Cymbial tubercle, E— Embolus, RTA—Retrolateral tibial apophysis, Sp—Spermophor [9–10 not to scale]." httpUri="https://zenodo.org/record/2606638/files/figure.png" pageId="4" pageNumber="528">Fig. 10</figureCitation>
|
||
). Palp as in diagnosis (
|
||
<figureCitation id="13392A47E76B6F661748FE9BCCDE786A" box="[1318,1423,351,376]" captionStart="FIGURES 6–10" captionStartId="4.[151,263,1811,1834]" captionTargetBox="[161,1433,593,1765]" captionTargetId="figure@4.[161,1433,593,1799]" captionTargetPageId="4" captionText="FIGURES 6–10. Systaria longinqua spec. nov., male from N Laos, Tham Pathok. 6–8 Left palp (6 ventral, 7 retrolateral, 8 dorsal). 9 Right chelicera, ventral. 10 Eye arrangement, dorsal. Abbreviations: C—Conductor, CT—Cymbial tubercle, E— Embolus, RTA—Retrolateral tibial apophysis, Sp—Spermophor [9–10 not to scale]." httpUri="https://zenodo.org/record/2606638/files/figure.png" pageId="4" pageNumber="528">Figs 6–8</figureCitation>
|
||
). Cymbium two times longer than tibia. RTA arising distally from tibia with 4 apices in retrolateral view and 2 apices in dorsal view; two apices long, retrolatero-proximad in ventral view, the other two short, retrolatero-distad. Embolus arising in 7- to 8-o’clock-position from tegulum, almost straight, narrowing suddenly from broad base to long narrow apical part. Tip situated close to membranous conductor in 12-o’clock-position. Moderately broad part of spermophor visible in retrolateral and proximal part of tegulum. Colouration. As in female.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7D664AE76B6F6612F9F8D7CABD7E75" httpUri="https://zenodo.org/record/2606638/files/figure.png" pageId="4" pageNumber="528" startId="4.[151,263,1811,1834]" targetBox="[161,1433,593,1765]" targetPageId="4">
|
||
<paragraph id="8BBD36C2E76B6F6612F9F8D7CABD7E75" blockId="4.[151,1436,1811,1895]" pageId="4" pageNumber="528">
|
||
<emphasis id="B976EAD0E76B6F6612F9F8D7C8197E3B" bold="true" box="[151,328,1811,1834]" pageId="4" pageNumber="528">FIGURES 6–10.</emphasis>
|
||
<taxonomicName id="4C024D41E76B6F66133EF8D0CB437E38" authority="Jäger, 2018" authorityName="Jäger" authorityYear="2018" box="[336,530,1812,1834]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="528" phylum="Arthropoda" rank="species" species="longinqua" status="sp. nov.">
|
||
<emphasis id="B976EAD0E76B6F66133EF8D0CB437E38" box="[336,530,1812,1834]" italics="true" pageId="4" pageNumber="528">Systaria longinqua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B976EAD0E76B6F661074F8D0CBD27E3B" bold="true" box="[538,643,1812,1833]" pageId="4" pageNumber="528">
|
||
<taxonomicNameLabel id="A24557ABE76B6F661074F8D0CBD27E3B" box="[538,643,1812,1833]" pageId="4" pageNumber="528" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male from N Laos, Tham Pathok. 6–8 Left palp (6 ventral, 7 retrolateral, 8 dorsal). 9 Right chelicera, ventral. 10 Eye arrangement, dorsal. Abbreviations: C—Conductor, CT—Cymbial tubercle, E— Embolus, RTA—Retrolateral tibial apophysis, Sp—Spermophor [9–10 not to scale].
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7D664AE76A6F6712F9FAB0C8337CDA" httpUri="https://zenodo.org/record/2606640/files/figure.png" pageId="5" pageNumber="529" startId="5.[151,263,1396,1419]" targetBox="[151,1435,193,1383]" targetPageId="5">
|
||
<paragraph id="8BBD36C2E76A6F6712F9FAB0C8337CDA" blockId="5.[151,1436,1396,1480]" pageId="5" pageNumber="529">
|
||
<emphasis id="B976EAD0E76A6F6712F9FAB0C8037C98" bold="true" box="[151,338,1396,1419]" pageId="5" pageNumber="529">FIGURES 11–15.</emphasis>
|
||
<taxonomicName id="4C024D41E76A6F671337FAB1CB4B7C98" authority="Jäger, 2018" authorityName="Jäger" authorityYear="2018" box="[345,538,1396,1418]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="529" phylum="Arthropoda" rank="species" species="longinqua" status="sp. nov.">
|
||
<emphasis id="B976EAD0E76A6F671337FAB1CB4B7C98" box="[345,538,1396,1418]" italics="true" pageId="5" pageNumber="529">Systaria longinqua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B976EAD0E76A6F67104EFAB1CBD67C98" bold="true" box="[544,647,1397,1418]" pageId="5" pageNumber="529">
|
||
<taxonomicNameLabel id="A24557ABE76A6F67104EFAB1CBD67C98" box="[544,647,1397,1418]" pageId="5" pageNumber="529" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype female from N Laos, Tham Pathok. 11 Epigyne, ventral. 12 Vulva, dorsal. 13 Left chelicera, ventral. 14 Prosoma, dorsal. 15 Spinnerets, ventral (not all setae drawn). Abbreviation: COcopulatory opening.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3186549E76A6F6712A9FA2BCD977F8A" pageId="5" pageNumber="529" type="distribution">
|
||
<paragraph id="8BBD36C2E76A6F6712A9FA2BCADB7F1A" blockId="5.[151,1437,1519,1688]" box="[199,906,1519,1544]" pageId="5" pageNumber="529">
|
||
<emphasis id="B976EAD0E76A6F6712A9FA2BC8317F1A" bold="true" box="[199,352,1519,1544]" pageId="5" pageNumber="529">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54B98860E76A6F67100EFA34CBC17F1A" box="[608,656,1520,1544]" pageId="5" pageNumber="529">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13392A47E76A6F671094FA2BCA2C7F1A" box="[762,893,1519,1544]" captionStart-0="FIGURE 45" captionStart-1="FIGURE 46" captionStartId-0="18.[151,250,1002,1024]" captionStartId-1="18.[151,250,1733,1755]" captionTargetBox-0="[150,1436,194,981]" captionTargetBox-1="[150,1436,1106,1712]" captionTargetId-0="figure@18.[150,1436,193,981]" captionTargetId-1="figure@18.[150,1436,1105,1712]" captionTargetPageId-0="18" captionTargetPageId-1="18" captionText-0="FIGURE 45. Records of Systaria spp. in SE Asia described in this paper. Triangles—species recorded from caves, circlesspecies recorded from forest." captionText-1="FIGURE 46. All known records of the genus Systaria (World Spider Catalog 2018) including undescribed species mentioned in Deeleman-Reinhold (2001) according to habitats. Triangles—species recorded from caves, circles—species recorded from forest. See discussion." httpUri-0="https://zenodo.org/record/2606658/files/figure.png" httpUri-1="https://zenodo.org/record/2606660/files/figure.png" pageId="5" pageNumber="529">Figs 45–46</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD36C2E76A6F6712A9F9D7CD977F8A" blockId="5.[151,1437,1519,1688]" pageId="5" pageNumber="529">
|
||
<emphasis id="B976EAD0E76A6F6712A9F9D7C8D87F3E" bold="true" box="[199,393,1555,1580]" pageId="5" pageNumber="529">Natural history.</emphasis>
|
||
These spiders have been recorded from the dark parts of this relatively small cave. The following organisms have been also found:
|
||
<taxonomicName id="4C024D41E76A6F6710E9F9F3CD177F42" authority="Jager & Ono 2000" authorityName="Jager & Ono" authorityYear="2000" box="[647,1094,1591,1616]" class="Arachnida" family="Sparassidae" genus="Heteropoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="529" phylum="Arthropoda" rank="species" species="simplex">
|
||
<emphasis id="B976EAD0E76A6F6710E9F9F3CA387F42" box="[647,873,1591,1616]" italics="true" pageId="5" pageNumber="529">Heteropoda simplex</emphasis>
|
||
<bibRefCitation id="EF934B33E76A6F67111FF9FCCD177F42" author="Jager, P. & Ono, H." box="[881,1094,1592,1616]" pageId="5" pageNumber="529" pagination="41 - 60" refId="ref11957" refString="Jager, P. & Ono, H. (2000) Sparassidae of Japan. I. New species of Olios, Heteropoda and Sinopoda with remarks on known species (Arachnida: Araneae). Acta Arachnologica, 49 (1), 41 - 60. https: // doi. org / 10.2476 / asjaa. 49.41" type="journal article" year="2000">Jäger & Ono 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C024D41E76A6F67163CF9F3CCC67F42" authority="Huber 2011" authorityName="Huber" authorityYear="2011" box="[1106,1431,1591,1616]" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="529" phylum="Arthropoda" rank="species" species="erawan">
|
||
<emphasis id="B976EAD0E76A6F67163CF9F3CC597F5D" box="[1106,1288,1591,1616]" italics="true" pageId="5" pageNumber="529">Pholcus erawan</emphasis>
|
||
Huber 2011
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C024D41E76A6F6712F9F99FC88D7F66" authority="Huber 2011" authorityName="Huber" authorityYear="2011" box="[151,476,1627,1652]" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="529" phylum="Arthropoda" rank="species" species="namou">
|
||
<emphasis id="B976EAD0E76A6F6712F9F99FC8167F66" box="[151,327,1627,1652]" italics="true" pageId="5" pageNumber="529">Pholcus namou</emphasis>
|
||
Huber 2011
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C024D41E76A6F671384F99FCB157F66" box="[490,580,1627,1652]" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="529" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976EAD0E76A6F671384F99FCB157F66" box="[490,580,1627,1652]" italics="true" pageId="5" pageNumber="529">Pholcus</emphasis>
|
||
</taxonomicName>
|
||
sp. cf.
|
||
<emphasis id="B976EAD0E76A6F6710F1F99FCA577F66" box="[671,774,1627,1652]" italics="true" pageId="5" pageNumber="529">namkhan</emphasis>
|
||
Huber 2011, Telemidae, Opiliones (Laniatores),
|
||
<taxonomicName id="4C024D41E76A6F671751F99FC8AD7F8A" authority=", Diplopoda" authorityName="Diplopoda" class="Insecta" family="Rhaphidophoridae" genus="Ensifera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="529" phylum="Arthropoda" rank="subGenus" subGenus="Rhaphidophoridae">Ensifera (Rhaphidophoridae), Diplopoda</taxonomicName>
|
||
div.,
|
||
<emphasis id="B976EAD0E76A6F671054F9BBCA057F8A" box="[570,852,1663,1688]" italics="true" pageId="5" pageNumber="529">Thereuopoda longicornis</emphasis>
|
||
(Scutigeromorpha) and Isopoda.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |