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<document id="ADD4A322321CCC200205D2D01A6A1383" ID-CLB-Dataset="28065" ID-DOI="10.11646/zootaxa.4504.4.5" ID-GBIF-Dataset="343a6a67-e8bf-467c-9818-de9621bbe955" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606614" ID-ZooBank="08E1A79F-BAA4-4EAB-9636-FE44BF7EB7F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585132792" checkinUser="plazi" docAuthor="Jäger, Peter" docDate="2018" docId="03AB87D4E7666F6912F9F952C8137A59" docLanguage="en" docName="zootaxa.4504.4.5.pdf" docOrigin="Zootaxa 4504 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systaria bregibec Jäger 2018, spec. nov." docType="treatment" docVersion="8" lastPageNumber="535" masterDocId="FF92FFACE76F6F62126EFFC4C9517912" masterDocTitle="On the genus Systaria (Araneae: Clubionidae) in Southeast Asia: new species from caves and forests" masterLastPageNumber="544" masterPageNumber="524" pageNumber="533" updateTime="1698712839155" updateUser="ExternalLinkService">
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<mods:title id="4A62F07C90DFC155ED1D28DDFF58884D">On the genus Systaria (Araneae: Clubionidae) in Southeast Asia: new species from caves and forests</mods:title>
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<mods:namePart id="0181F49BB17090395F8604E74327A493">Jäger, Peter</mods:namePart>
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<mods:title id="E61775EE41F31D9E8EE1FC6B14EC906C">Zootaxa</mods:title>
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<mods:date id="F0DD9D19D1C74DB0694E8DDFC2C706BD">2018</mods:date>
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<mods:number id="B3135D0129C3174FF49A30A85977657C">2018-10-26</mods:number>
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<mods:number id="3D15FB47FC3C98E2E71DC59C2F3C796B">4504</mods:number>
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<mods:classification id="B58BA3873BCD178D0059964CB39E19BA">journal article</mods:classification>
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<mods:identifier id="52A59761991749EC25B6DE61FCFAF6D4" type="DOI">10.11646/zootaxa.4504.4.5</mods:identifier>
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<treatment id="03AB87D4E7666F6912F9F952C8137A59" ID-DOI="http://doi.org/10.5281/zenodo.5988010" ID-GBIF-Taxon="154495200" ID-Zenodo-Dep="5988010" LSID="urn:lsid:plazi:treatment:03AB87D4E7666F6912F9F952C8137A59" httpUri="http://treatment.plazi.org/id/03AB87D4E7666F6912F9F952C8137A59" lastPageId="11" lastPageNumber="535" pageId="9" pageNumber="533">
<subSubSection id="C3186549E7666F6B12F9F952C8B47FA2" box="[151,485,1686,1712]" pageId="9" pageNumber="533" type="nomenclature">
<paragraph id="8BBD36C2E7666F6B12F9F952C8B47FA2" blockId="9.[151,485,1686,1746]" box="[151,485,1686,1712]" pageId="9" pageNumber="533">
<heading id="D0F581AEE7666F6B12F9F952C8B47FA2" bold="true" box="[151,485,1686,1712]" fontSize="11" level="1" pageId="9" pageNumber="533" reason="1">
<emphasis id="B976EAD0E7666F6B12F9F952C8B47FA2" bold="true" box="[151,485,1686,1712]" pageId="9" pageNumber="533">
<taxonomicName id="4C024D41E7666F6B12F9F952C8347FA2" authority="Jäger, 2018" authorityName="Jäger" authorityYear="2018" box="[151,357,1686,1712]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="533" phylum="Arthropoda" rank="species" species="bregibec" status="sp. nov.">
<emphasis id="B976EAD0E7666F6B12F9F952C8347FA2" bold="true" box="[151,357,1686,1712]" italics="true" pageId="9" pageNumber="533">Systaria bregibec</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557ABE7666F6B1303F952C8B47FA2" box="[365,485,1686,1712]" pageId="9" pageNumber="533" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7666F6B12F9F97DC8397FC0" box="[151,360,1721,1746]" pageId="9" pageNumber="533" type="description">
<paragraph id="8BBD36C2E7666F6B12F9F97DC8397FC0" blockId="9.[151,485,1686,1746]" box="[151,360,1721,1746]" pageId="9" pageNumber="533">
<figureCitation id="13392A47E7666F6B12F9F97DC8477FC0" box="[151,278,1721,1746]" captionStart="FIGURES 27–32" captionStartId="12.[151,263,1874,1897]" captionTargetBox="[151,1412,217,1843]" captionTargetId="figure@12.[151,1436,213,1843]" captionTargetPageId="12" captionText="FIGURES 27–32. Systaria spp., copulatory organs. 27–28 Systaria bregibec spec. nov., holotype male from Cambodia, Rabbit Island. 29–30 Systaria bifidops spec. nov., holotype male from Malaysia, Bukit Fraser. 31–32 Systaria panay spec. nov., holotype female from Philippines, Panay Island. 27–30 Left palp (27, 29 ventral, 28, 30 retrolateral). 31 Epigyne, ventral. 32 Vulva, dorsal. Abbreviations: BP—bifid process of RTA, CO—copulatory opening, LO—loop of copulatory ducts, TPproximal projection of tegulum, VH—ventral hump of palpal tibia." httpUri="https://zenodo.org/record/2606652/files/figure.png" pageId="9" pageNumber="533">Figs 27–28</figureCitation>
,
<figureCitation id="13392A47E7666F6B134CF97DC8397FC0" box="[290,360,1721,1746]" captionStart-0="FIGURE 45" captionStart-1="FIGURE 46" captionStartId-0="18.[151,250,1002,1024]" captionStartId-1="18.[151,250,1733,1755]" captionTargetBox-0="[150,1436,194,981]" captionTargetBox-1="[150,1436,1106,1712]" captionTargetId-0="figure@18.[150,1436,193,981]" captionTargetId-1="figure@18.[150,1436,1105,1712]" captionTargetPageId-0="18" captionTargetPageId-1="18" captionText-0="FIGURE 45. Records of Systaria spp. in SE Asia described in this paper. Triangles—species recorded from caves, circlesspecies recorded from forest." captionText-1="FIGURE 46. All known records of the genus Systaria (World Spider Catalog 2018) including undescribed species mentioned in Deeleman-Reinhold (2001) according to habitats. Triangles—species recorded from caves, circles—species recorded from forest. See discussion." httpUri-0="https://zenodo.org/record/2606658/files/figure.png" httpUri-1="https://zenodo.org/record/2606660/files/figure.png" pageId="9" pageNumber="533">45–46</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7666F6B12F9F8C5CC2E7E2C" pageId="9" pageNumber="533" type="materials_examined">
<paragraph id="8BBD36C2E7666F6B12F9F8C5CC2E7E2C" blockId="9.[151,1436,1793,2034]" pageId="9" pageNumber="533">
<emphasis id="B976EAD0E7666F6B12F9F8C5C8197E08" bold="true" box="[151,328,1793,1818]" pageId="9" pageNumber="533">Type material:</emphasis>
<materialsCitation id="3B6A3C9FE7666F6B1321F8C6CC2E7E2C" ID-GBIF-Occurrence="2028138111" collectingDate="2017-07-09" collectionCode="SMF" collectorName="P. Jager" country="Cambodia" elevation="21" latitude="10.436514" location="Rabbit Island" longLatPrecision="1" longitude="104.320274" pageId="9" pageNumber="533" specimenCount="1" stateProvince="Kep" typeStatus="holotype">
<collectingCountry id="F3157652E7666F6B1321F8C6C8BD7E08" box="[335,492,1794,1818]" name="Cambodia" pageId="9" pageNumber="533">CAMBODIA</collectingCountry>
:
<emphasis id="B976EAD0E7666F6B1399F8C6CBC87E0B" box="[503,665,1794,1817]" italics="true" pageId="9" pageNumber="533">
<collectingRegion id="49C6F820E7666F6B1399F8C6CBC27E0B" box="[503,659,1794,1817]" country="Cambodia" name="Kep" pageId="9" pageNumber="533">Kep Province</collectingRegion>
:
</emphasis>
<emphasis id="B976EAD0E7666F6B10CCF8C5CA5E7E08" bold="true" box="[674,783,1793,1818]" pageId="9" pageNumber="533">
<typeStatus id="54B98860E7666F6B10CCF8C5CA5E7E08" box="[674,783,1793,1818]" pageId="9" pageNumber="533" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<location id="8EDD6019E7666F6B110DF8C5CAAC7E08" LSID="urn:lsid:plazi:treatment:03AB87D4E7666F6912F9F952C8137A59:8EDD6019E7666F6B110DF8C5CAAC7E08" box="[867,1021,1793,1818]" country="Cambodia" latitude="10.436514" longLatPrecision="1" longitude="104.320274" name="Rabbit Island" pageId="9" pageNumber="533" stateProvince="Kep">Rabbit Island</location>
,
<geoCoordinate id="EE365005E7666F6B1667F8C5CDE47E08" box="[1033,1205,1793,1818]" direction="north" orientation="latitude" pageId="9" pageNumber="533" precision="1" value="10.436514">10°26’11.45''N</geoCoordinate>
,
<geoCoordinate id="EE365005E7666F6B16AFF8C6CC257E08" box="[1217,1396,1794,1818]" direction="east" orientation="longitude" pageId="9" pageNumber="533" precision="1" value="104.320274">104°19'12.98''E</geoCoordinate>
,
<quantity id="4CFA9B27E7666F6B17EEF8C6C9FC7E2F" metricMagnitude="1" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="533" unit="m" value="21.0">
<elevation id="002FD1F1E7666F6B17EEF8C6C9FC7E2F" metricMagnitude="1" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="533" unit="m" value="21.0">21 m</elevation>
</quantity>
elevation, disturbed secondary forest, forest floor and foliage, at night,
<collectorName id="26F75314E7666F6B11BEF8E2CD747E2C" box="[976,1061,1830,1854]" pageId="9" pageNumber="533">P. Jäger</collectorName>
leg. by hand,
<date id="FFBC1002E7666F6B16AAF8E2CC777E2C" box="[1220,1318,1830,1854]" pageId="9" pageNumber="533" value="2017-07-09">
<collectingDate id="EFF8E9EAE7666F6B16AAF8E2CC777E2C" box="[1220,1318,1830,1854]" pageId="9" pageNumber="533" value="2017-07-09">9.7.2017</collectingDate>
</date>
(
<collectionCode id="ED13AE07E7666F6B175BF8E2CC227E2C" box="[1333,1395,1830,1854]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="533">SMF</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7666F6B12A9F88DCB767EE0" pageId="9" pageNumber="533" type="etymology">
<paragraph id="8BBD36C2E7666F6B12A9F88DCB767EE0" blockId="9.[151,1436,1793,2034]" pageId="9" pageNumber="533">
<emphasis id="B976EAD0E7666F6B12A9F88DC81F7E70" bold="true" box="[199,334,1865,1890]" pageId="9" pageNumber="533">Etymology.</emphasis>
The species name is a combination of three Latin words: “brevis”, meaning “short”, referring to the short embolus, “gibber”, meaning “hump”, referring to the distinct ventral hump on the male palpal tibia, and “beccus”, meaning “beak”, referring to the beak-shaped distal part of the RTA in retrolateral view (
<figureCitation id="13392A47E7666F6B168DF855CC377EB8" box="[1251,1382,1937,1962]" captionStart="FIGURES 27–32" captionStartId="12.[151,263,1874,1897]" captionTargetBox="[151,1412,217,1843]" captionTargetId="figure@12.[151,1436,213,1843]" captionTargetPageId="12" captionText="FIGURES 27–32. Systaria spp., copulatory organs. 27–28 Systaria bregibec spec. nov., holotype male from Cambodia, Rabbit Island. 29–30 Systaria bifidops spec. nov., holotype male from Malaysia, Bukit Fraser. 31–32 Systaria panay spec. nov., holotype female from Philippines, Panay Island. 27–30 Left palp (27, 29 ventral, 28, 30 retrolateral). 31 Epigyne, ventral. 32 Vulva, dorsal. Abbreviations: BP—bifid process of RTA, CO—copulatory opening, LO—loop of copulatory ducts, TPproximal projection of tegulum, VH—ventral hump of palpal tibia." httpUri="https://zenodo.org/record/2606652/files/figure.png" pageId="9" pageNumber="533">Figs 27–28</figureCitation>
). All three character states are characteristic for the species group with currently three species included (see diagnosis and discussion); term in apposition.
</paragraph>
</subSubSection>
<caption id="DF7D664AE7656F6812F9F8D0CA0F7E7A" httpUri="https://zenodo.org/record/2606650/files/figure.png" pageId="10" pageNumber="534" startId="10.[151,263,1812,1835]" targetBox="[165,1431,188,1798]" targetPageId="10">
<paragraph id="8BBD36C2E7656F6812F9F8D0CA0F7E7A" blockId="10.[151,1436,1812,1896]" pageId="10" pageNumber="534">
<emphasis id="B976EAD0E7656F6812F9F8D0C8057E38" bold="true" box="[151,340,1812,1835]" pageId="10" pageNumber="534">FIGURES 22–26.</emphasis>
<taxonomicName id="4C024D41E7656F681334F8D1CB527E39" authority="Jäger, 2018" authorityName="Jäger" authorityYear="2018" box="[346,515,1813,1835]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="534" phylum="Arthropoda" rank="species" species="procera" status="sp. nov.">
<emphasis id="B976EAD0E7656F681334F8D1CB527E39" box="[346,515,1813,1835]" italics="true" pageId="10" pageNumber="534">Systaria procera</emphasis>
</taxonomicName>
<emphasis id="B976EAD0E7656F681064F8D1CB207E38" bold="true" box="[522,625,1813,1834]" pageId="10" pageNumber="534">
<taxonomicNameLabel id="A24557ABE7656F681064F8D1CB207E38" box="[522,625,1813,1834]" pageId="10" pageNumber="534" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, females from NW Cambodia (22–23 La Ang Pracheu, 24–25 Roung Onkham, 26 Roung Pnomh Chakrey 8). 22, 24, 26 Epigyne, ventral (26 not dissected). 23, 25 Vulva, dorsal. Abbreviations: COcopulatory opening, MP—median protrusion of anterior atrial margin.
</paragraph>
</caption>
<subSubSection id="C3186549E7656F6912A9F850CBA479C3" lastPageId="11" lastPageNumber="535" pageId="10" pageNumber="534" type="diagnosis">
<paragraph id="8BBD36C2E7656F6912A9F850CBA479C3" blockId="10.[151,1437,1940,2032]" lastBlockId="11.[151,1437,151,843]" lastPageId="11" lastPageNumber="535" pageId="10" pageNumber="534">
<emphasis id="B976EAD0E7656F6812A9F850C8107EBF" bold="true" box="[199,321,1940,1965]" pageId="10" pageNumber="534">Diagnosis.</emphasis>
Small spiders with body length of males: 9.1. Male copulatory organs similar to
<taxonomicName id="4C024D41E7656F68174BF851CB417EDC" authority="Dankittipakul &amp; Singtripop 2011" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2011" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="534" phylum="Arthropoda" rank="species" species="insolita">
<emphasis id="B976EAD0E7656F68174BF851CCCD7EBF" box="[1317,1436,1940,1965]" italics="true" pageId="10" pageNumber="534">S. insolita</emphasis>
<bibRefCitation id="EF934B33E7656F6812F9F871CB417EDC" author="Dankittipakul, P. &amp; Singtripop, T." box="[151,528,1973,1998]" pageId="10" pageNumber="534" pagination="16 - 32" refId="ref11494" refString="Dankittipakul, P. &amp; Singtripop, T. (2011) Seven new species of Systaria Simon, 1897 from southeast Asia (Araneae, Clubionidae, Systariinae). Zootaxa, 2905, 16 - 32." type="journal article" year="2011">Dankittipakul &amp; Singtripop 2011</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C024D41E7656F681025F872CD557EDC" authority="Zhang, Fu &amp; Zhu 2009" authorityName="Zhang, Fu &amp; Zhu" authorityYear="2009" box="[587,1028,1973,1998]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="534" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B976EAD0E7656F681025F872CBBE7EDC" box="[587,751,1973,1998]" italics="true" pageId="10" pageNumber="534">S. hainanensis</emphasis>
<bibRefCitation id="EF934B33E7656F681096F871CD557EDC" author="Zhang, F. &amp; Fu, J. Y. &amp; Zhu, M. S." box="[760,1028,1973,1998]" pageId="10" pageNumber="534" pagination="51 - 60" refId="ref12597" refString="Zhang, F., Fu, J. Y. &amp; Zhu, M. S. (2009) First report of the sac spider genus Systaria Simon (Araneae: Clubionidae) from China: Systaria hainanensis sp. nov. and Systaria mengla (Song &amp; Zhu, 1994) comb. nov. Zootaxa, 2305, 51 - 60." type="journal article" year="2009">
Zhang, Fu &amp; Zhu
<quantity id="4CFA9B27E7656F6811A2F872CD557EDC" box="[972,1028,1974,1998]" metricMagnitude="1" metricUnit="m" metricValue="5.10286" pageId="10" pageNumber="534" unit="in" value="2009.0">2009</quantity>
</bibRefCitation>
</taxonomicName>
in having a short embolus, a beakshaped (double-headed) RTA tip and a distinct ventral hump at the palpal tibia, but can be distinguished by both processes of distal RTA pointed in retrolateral view (blunt in
<taxonomicName id="4C024D41E7646F691132FF5CCD5179A2" authorityName="Zhang, Fu &amp; Zhu" authorityYear="2009" box="[860,1024,151,176]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="535" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B976EAD0E7646F691132FF5CCD5179A2" box="[860,1024,151,176]" italics="true" pageId="11" pageNumber="535">S. hainanensis</emphasis>
</taxonomicName>
) and of similar size (dorsal process distinctly more prominent in
<taxonomicName id="4C024D41E7646F6913B3FF7ECB1C79C0" authorityName="Dankittipakul &amp; Singtripop" authorityYear="2011" box="[477,589,185,210]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="535" phylum="Arthropoda" rank="species" species="insolita">
<emphasis id="B976EAD0E7646F6913B3FF7ECB1C79C0" box="[477,589,185,210]" italics="true" pageId="11" pageNumber="535">S. insolita</emphasis>
</taxonomicName>
) (
<figureCitation id="13392A47E7646F691008FF7DCBB879C3" box="[614,745,185,210]" captionStart="FIGURES 27–32" captionStartId="12.[151,263,1874,1897]" captionTargetBox="[151,1412,217,1843]" captionTargetId="figure@12.[151,1436,213,1843]" captionTargetPageId="12" captionText="FIGURES 27–32. Systaria spp., copulatory organs. 27–28 Systaria bregibec spec. nov., holotype male from Cambodia, Rabbit Island. 29–30 Systaria bifidops spec. nov., holotype male from Malaysia, Bukit Fraser. 31–32 Systaria panay spec. nov., holotype female from Philippines, Panay Island. 27–30 Left palp (27, 29 ventral, 28, 30 retrolateral). 31 Epigyne, ventral. 32 Vulva, dorsal. Abbreviations: BP—bifid process of RTA, CO—copulatory opening, LO—loop of copulatory ducts, TPproximal projection of tegulum, VH—ventral hump of palpal tibia." httpUri="https://zenodo.org/record/2606652/files/figure.png" pageId="11" pageNumber="535">Figs 27–28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7646F6912A9FF1EC8C87BF5" pageId="11" pageNumber="535" type="description">
<paragraph id="8BBD36C2E7646F6912A9FF1ECDD47BD4" blockId="11.[151,1437,151,843]" pageId="11" pageNumber="535">
<emphasis id="B976EAD0E7646F6912A9FF1EC80879E1" bold="true" box="[199,345,218,243]" pageId="11" pageNumber="535">Description.</emphasis>
<emphasis id="B976EAD0E7646F691308FF1EC8F179E1" box="[358,416,218,243]" italics="true" pageId="11" pageNumber="535">Male</emphasis>
(
<typeStatus id="54B98860E7646F6913D8FF1ECB4E79E1" box="[438,543,218,243]" pageId="11" pageNumber="535" type="holotype">holotype</typeStatus>
): prosoma length 4.5, prosoma width 3.1, anterior width of prosoma 2.1, opisthosoma length 4.6, opisthosoma width 2.1. Eyes: AME 0.29, ALE 0.25, PME 0.20, PLE 0.26, AME–AME 0.16, AME–ALE 0.06, PME–PME 0.36, PME–PLE 0.24, AME–PME 0.18, ALE–PLE 0.02. Cheliceral furrow with 3 promarginal and 2 retromarginal teeth. Spination: Palp: femur p001, d012, patella–tarsus with bristles, not counted in spination; femur I p0011, d111, r110 (small), II p0111, d111, r110 (small), III p111, d111, r111, IV p1101, d111, r111; patella I–IV 000; tibia I p01, v 220, II p01, v220(2120), III–IV p11, d010, r11, v222; metatarsus II v100, III p112, r112, v202, IV p112, r122, v222. Leg formula: 4123. Measurements of palp and legs: Palp 4.4 (1.8, 0.7, 0.7, -, 1.2), I 13.9 (3.9, 1.7, 3.6, 3.0, 1.7), II 13.3 (3.7, 1.7, 3.4, 2.9, 1.6), III 11.6 (3.3, 1.4, 2.6, 2.9, 1.4), IV 16.6 (4.5, 1.7, 3.9, 4.6, 1.9). Palp as in diagnosis (
<figureCitation id="13392A47E7646F6910A7FE21CA1D78EC" box="[713,844,485,510]" captionStart="FIGURES 27–32" captionStartId="12.[151,263,1874,1897]" captionTargetBox="[151,1412,217,1843]" captionTargetId="figure@12.[151,1436,213,1843]" captionTargetPageId="12" captionText="FIGURES 27–32. Systaria spp., copulatory organs. 27–28 Systaria bregibec spec. nov., holotype male from Cambodia, Rabbit Island. 29–30 Systaria bifidops spec. nov., holotype male from Malaysia, Bukit Fraser. 31–32 Systaria panay spec. nov., holotype female from Philippines, Panay Island. 27–30 Left palp (27, 29 ventral, 28, 30 retrolateral). 31 Epigyne, ventral. 32 Vulva, dorsal. Abbreviations: BP—bifid process of RTA, CO—copulatory opening, LO—loop of copulatory ducts, TPproximal projection of tegulum, VH—ventral hump of palpal tibia." httpUri="https://zenodo.org/record/2606652/files/figure.png" pageId="11" pageNumber="535">Figs 27–28</figureCitation>
). Cymbium two times longer than tibia. RTA arising distally from tibia with its disto-dorsal margin straight, disto-ventrad. Embolus arising in 9-o’clock-position from tegulum, slightly bent, with broad base, its tip situated close to tip of membranous conductor in 12-o’clockposition. Broad part of spermophor visible retrolaterally and proximally, S-shaped narrow part visible proximally to embolus base. Colouration. Deep yellowish- to reddish-brown without pattern. Fovea black, chelicerae, labium, gnathocoxae deep reddish-brown, sternum marginally darker brown. Opisthosoma dorsally with one pair of black spots and some black transverse striae in front of spinnerets. Dark setae around spinnerets
</paragraph>
<paragraph id="8BBD36C2E7646F6912A9FD0AC8C87BF5" blockId="11.[151,1437,151,843]" box="[199,409,718,743]" pageId="11" pageNumber="535">
<emphasis id="B976EAD0E7646F6912A9FD0AC84A7BF5" box="[199,283,718,743]" italics="true" pageId="11" pageNumber="535">Female</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3186549E7646F6912A9FD2BC8137A59" pageId="11" pageNumber="535" type="distribution">
<paragraph id="8BBD36C2E7646F6912A9FD2BCADB7A1A" blockId="11.[151,1437,151,843]" box="[199,906,751,776]" pageId="11" pageNumber="535">
<emphasis id="B976EAD0E7646F6912A9FD2BC8317A1A" bold="true" box="[199,352,751,776]" pageId="11" pageNumber="535">Distribution.</emphasis>
Known only from the
<typeStatus id="54B98860E7646F69100EFD34CBC17A1A" box="[608,656,752,776]" pageId="11" pageNumber="535">type</typeStatus>
locality (
<figureCitation id="13392A47E7646F691094FD2BCA2C7A1A" box="[762,893,751,776]" captionStart-0="FIGURE 45" captionStart-1="FIGURE 46" captionStartId-0="18.[151,250,1002,1024]" captionStartId-1="18.[151,250,1733,1755]" captionTargetBox-0="[150,1436,194,981]" captionTargetBox-1="[150,1436,1106,1712]" captionTargetId-0="figure@18.[150,1436,193,981]" captionTargetId-1="figure@18.[150,1436,1105,1712]" captionTargetPageId-0="18" captionTargetPageId-1="18" captionText-0="FIGURE 45. Records of Systaria spp. in SE Asia described in this paper. Triangles—species recorded from caves, circlesspecies recorded from forest." captionText-1="FIGURE 46. All known records of the genus Systaria (World Spider Catalog 2018) including undescribed species mentioned in Deeleman-Reinhold (2001) according to habitats. Triangles—species recorded from caves, circles—species recorded from forest. See discussion." httpUri-0="https://zenodo.org/record/2606658/files/figure.png" httpUri-1="https://zenodo.org/record/2606660/files/figure.png" pageId="11" pageNumber="535">Figs 45–46</figureCitation>
).
</paragraph>
<paragraph id="8BBD36C2E7646F6912A9FCD5C8137A59" blockId="11.[151,1437,151,843]" pageId="11" pageNumber="535">
<emphasis id="B976EAD0E7646F6912A9FCD5C8D77A38" bold="true" box="[199,390,785,810]" pageId="11" pageNumber="535">Natural history.</emphasis>
The specimen has been found on a slope in a degraded secondary forest, with dry and weak leaf litter layer.
</paragraph>
</subSubSection>
</treatment>
</document>
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