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<mods:title id="C78F5EC5FD2D406F7BAB53C3EC6915CE">Revision of the genus Porcellionides Miers, 1877 (Isopoda: Porcellionidae) in the Ibero-Balearic region</mods:title>
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<mods:namePart id="6D0A729DE5C2D6ABA8933A23D11790CC">Cifuentes, Julio</mods:namePart>
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<mods:affiliation id="136C46915C06A263D4164BA8E92746EE">Departamento de Biología (Zoología), Facultad de Ciencias, Universidad Autónoma de Madrid, 28049 Cantoblanco, Madrid, Spain.</mods:affiliation>
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<mods:affiliation id="5CAEB5FB8610FE46D03CB82D6FDA681F">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Portugal. &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661 Vairão, Portugal.</mods:affiliation>
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<treatment id="03AB87C5FFB1FF9037A8FC9FFC07F857" ID-DOI="http://doi.org/10.5281/zenodo.12157048" ID-GBIF-Taxon="233240668" ID-Zenodo-Dep="12157048" LSID="urn:lsid:plazi:treatment:03AB87C5FFB1FF9037A8FC9FFC07F857" httpUri="http://treatment.plazi.org/id/03AB87C5FFB1FF9037A8FC9FFC07F857" lastPageNumber="13" pageId="12" pageNumber="13">
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<paragraph id="8BBD36D3FFB1FF9037A8FC9FFBA0FCC5" blockId="12.[470,1117,776,838]" box="[470,1117,776,804]" pageId="12" pageNumber="13">
<heading id="D0F581BFFFB1FF9037A8FC9FFBA0FCC5" box="[470,1117,776,804]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="4C024D50FFB1FF9037A8FC9FFBA0FCC5" ID-CoL="4LWJ6" authority="(Budde-Lund, 1885)" baseAuthorityName="Budde-Lund" baseAuthorityYear="1885" box="[470,1117,776,804]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="fuscomarmoratus">
<emphasis id="B976EAC1FFB1FF9037A8FC9FFC98FCC3" bold="true" box="[470,869,776,803]" italics="true" pageId="12" pageNumber="13">Porcellionides fuscomarmoratus</emphasis>
(
<bibRefCitation id="EF934B22FFB1FF90350AFC9EFBA8FCC5" author="Budde-Lund G." box="[884,1109,777,804]" pageId="12" pageNumber="13" refId="ref23578" refString="Budde-Lund G. 1885. Crustacea Isopoda Terrestria per Familias et Genera et Species Descripta. Nielsen &amp; Lydiche, Copenhagen. https: // doi. org / 10.5962 / bhl. title. 109769" type="book" year="1885">Budde-Lund, 1885</bibRefCitation>
)
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<subSubSection id="C3186558FFB1FF90341EFCBBFC29FCA7" box="[608,980,812,838]" pageId="12" pageNumber="13" type="description">
<paragraph id="8BBD36D3FFB1FF90341EFCBBFC29FCA7" blockId="12.[470,1117,776,838]" box="[608,980,812,838]" pageId="12" pageNumber="13">
<figureCitation id="13392A56FFB1FF90341EFCBBFD43FCA7" box="[608,702,812,838]" captionStart="Fig" captionStartId="6.[189,232,1716,1742]" captionTargetBox="[213,1355,503,1668]" captionTargetId="figure-106@6.[189,1398,496,1681]" captionTargetPageId="6" captionText="Fig. 2. Relation between noduli laterales coordinates (following Vandel 1960) d/c (y-axis) and b/c (x-axis) multiplied by 100 for all studied species. A. Porcellionides cingendus (Kinahan, 1857). B. Porcellionides elegans(Pollo Zorita, 1982). C. Porcellionides fuscomarmoratus(Budde-Lund, 1885). D. Porcellionides glaber (Koch, 1856). E. Porcellionides hispanus (Vandel, 1953). F. Porcellionides lucasioides (Vandel, 1953). G. Porcellionides lusitanus (Vandel, 1946). H. Porcellionides molleri (Verhoeff, 1901). I. Porcellionides pruinosus (Brandt, 1833). J. Porcellionides sexfasciatus (BuddeLund, 1885). K. Porcellionides ibericus sp. nov. L. P. sexfasciatus and its previously considered subspecies: P. glaber (g), P. hispanus (h), P. lucasioides (l), P. lusitanus (lu), P. molleri (m) and P. sexfasciatus (s)." figureDoi="http://doi.org/10.5281/zenodo.12157016" httpUri="https://zenodo.org/record/12157016/files/figure.png" pageId="12" pageNumber="13">Figs 2C</figureCitation>
,
<figureCitation id="13392A56FFB1FF9034B4FCBBFD12FCA7" box="[714,751,812,838]" captionStart="Fig" captionStartId="7.[189,232,1786,1812]" captionTargetBox="[231,1365,1102,1731]" captionTargetId="figure-323@7.[189,1398,1089,1751]" captionTargetPageId="7" captionText="Fig. 3. Scale-setae. A. Porcellionides cingendus (Kinahan, 1857), ♂ (JC104). B. Porcellionides elegans (Pollo Zorita, 1982), ♂ (MNCN 20.04/7306). C. Porcellionides fuscomarmoratus (BuddeLund, 1885), ♀ (MNCN 20.04/8129). D. Porcellionides glaber (Koch, 1856), ♀ (MNCN 20.04/11490). E. Porcellionides hispanus (Vandel, 1953), ♂ (MNCN 20.04/7163). F. Porcellionides lucasioides (Vandel, 1953), ♂ (JC352). G. Porcellionides lusitanus (Vandel, 1946), ♂ (LPS661). H. Porcellionides molleri (Verhoeff, 1901), ♀ (LPS135). I. Porcellionides pruinosus (Brandt, 1833), ♀ (JC318). J. Porcellionides sexfasciatus (Budde-Lund, 1885), ♂ (LPS455a). Scale bar = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.12157020" httpUri="https://zenodo.org/record/12157020/files/figure.png" pageId="12" pageNumber="13">3C</figureCitation>
,
<figureCitation id="13392A56FFB1FF903485FCBBFCCAFCA7" box="[763,823,812,838]" captionStart="Fig" captionStartId="8.[189,232,1704,1730]" captionTargetBox="[323,1265,274,1643]" captionTargetId="figure-17@8.[280,1308,262,1666]" captionText="Fig. 4. Pleopods. AD. Porcellionides cingendus (Kinahan, 1857). AB. ♂ (LPS215). A. Exopod I. B. Exopod II. CD. ♀ (MNCN 20.04/11972). C. Exopod I. D. Exopod II. EH. Porcellionides elegans (Pollo Zorita, 1982). EF. ♂ (JC490). E. Exopod I. F.Exopod II. GH. ♀ (MNCN 20.04/9590). G. Exopod I. H. Exopod II. IL. Porcellionides fuscomarmoratus (Budde-Lund, 1885). IJ. ♂ (UGMSS-2). I. Exopod I. J. Exopod II. KL. ♀ (MNCN 20.04/8129. K. Exopod I. L. Exopod II. MP. Porcellionides glaber (Koch, 1856). ML. ♂ (MNCN 20.04/7886). M. Exopod I. N. Exopod II. O. ♀ (MNCN 20.04/11490), exopod I. P. ♀ (MNCN 20.04/11490), exopod II. QT. Porcellionides hispanus (Vandel, 1953). QR. ♂ (MNCN 20.04/7163). Q. Exopod I. R. Exopod II. ST. ♀ (JC380). S. Exopod I. T. ♀ (JC380), exopod II. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12188976" httpUri="https://zenodo.org/record/12188976/files/figure.png" pageId="12" pageNumber="13">4IL</figureCitation>
, 89,
<tableCitation id="C6800368FFB1FF903503FCBBFC29FCA7" box="[893,980,812,838]" captionStart="Table 1" captionStartId="5.[185,255,713,741]" captionTargetPageId="5" captionText="Table 1. Summary of comparative characters for species within the genus Porcellionides Miers, 1877 found in the Ibero-Balearic region." httpUri="http://table.plazi.org/id/DF7D665BFFB8FF9936C7FD5EF8F9FD04" pageId="12" pageNumber="13" tableUuid="DF7D665BFFB8FF9936C7FD5EF8F9FD04">Table 1</tableCitation>
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<taxonomicName id="4C024D50FFB1FF9036C3FCFAFC90FC69" ID-CoL="42HJB" authority="Budde-Lund, 1885: 189" authorityName="Budde-Lund" authorityPageNumber="189" authorityYear="1885" box="[189,877,877,904]" class="Malacostraca" family="Porcellionidae" genus="Metoponorthus" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="fuscomarmoratus">
<emphasis id="B976EAC1FFB1FF9036C3FCFAFDB4FC69" box="[189,585,877,904]" italics="true" pageId="12" pageNumber="13">Metoponorthus fuscomarmoratus</emphasis>
<bibRefCitation id="EF934B22FFB1FF90342EFCFAFC90FC69" author="Budde-Lund G." box="[592,877,877,904]" pageId="12" pageNumber="13" refId="ref23578" refString="Budde-Lund G. 1885. Crustacea Isopoda Terrestria per Familias et Genera et Species Descripta. Nielsen &amp; Lydiche, Copenhagen. https: // doi. org / 10.5962 / bhl. title. 109769" type="book" year="1885">Budde-Lund, 1885: 189</bibRefCitation>
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.
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<emphasis id="B976EAC1FFB1FF9036C3FC38FE50FC28" bold="true" box="[189,429,943,969]" pageId="12" pageNumber="13">Material examined</emphasis>
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<collectingDate id="EFF8E9FBFFB1FF903400FC4AFCE3FC19" box="[638,798,989,1016]" pageId="12" pageNumber="13" value="2023-03-25">25 Mar. 2023</collectingDate>
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and
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leg.; JC758
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;
<collectingMunicipality id="6BD9ACA9FFB1FF903768FB96FDDEFBFA" box="[278,547,1025,1051]" pageId="12" pageNumber="13">Tajos del Campanario</collectingMunicipality>
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a.s.l.
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;
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<collectingDate id="EFF8E9FBFFB1FF90324AFB96FB4BFBFA" box="[1076,1206,1025,1051]" pageId="12" pageNumber="13" value="2014-08">Aug. 2014</collectingDate>
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;
<collectorName id="26F75305FFB1FF9032BAFB96FAC5FBFA" box="[1220,1336,1025,1051]" pageId="12" pageNumber="13">A. Tinaut</collectorName>
leg.; UGMSS-2
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<specimenCount id="9D04FD5AFFB1FF90372DFBB3FE6BFBDF" box="[339,406,1060,1086]" count="4" pageId="12" pageNumber="13" type="female">4 ♀♀</specimenCount>
;
<location id="8EDD6008FFB1FF9037DBFBB3FDB1FBDF" LSID="urn:lsid:plazi:treatment:03AB87C5FFB1FF9037A8FC9FFC07F857:8EDD6008FFB1FF9037DBFBB3FDB1FBDF" box="[421,588,1060,1086]" country="Spain" name="Sierra Nevada" pageId="12" pageNumber="13" stateProvince="Granada">Sierra Nevada</location>
(Monte caballo);
<date id="FFBC1013FFB1FF903561FBB3FC51FBDE" box="[799,940,1060,1087]" pageId="12" pageNumber="13" value="1965-10-04">
<collectingDate id="EFF8E9FBFFB1FF903561FBB3FC51FBDE" box="[799,940,1060,1087]" pageId="12" pageNumber="13" value="1965-10-04">4 Oct. 1965</collectingDate>
</date>
;
<specimenCode id="DBA49EA8FFB1FF9035C6FBB3FB5DFBDF" box="[952,1184,1060,1086]" pageId="12" pageNumber="13">MNCN 20.04/8129</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BBD36D3FFB1FF9036C3FBF2FE3EFB27" blockId="12.[188,1399,1125,1222]" pageId="12" pageNumber="13">
The latest specimens are cited by
<bibRefCitation id="EF934B22FFB1FF90343AFBF2FCE7FB9E" author="Cifuentes J." box="[580,794,1125,1151]" pageId="12" pageNumber="13" pagination="133" refId="ref23773" refString="Cifuentes J. 2021 a. Los isopodos terrestres de Andalucia, Espana (Crustacea: Isopoda, Oniscidea). Graellsia 77 (1): e 133. https: // doi. org / 10.3989 / graellsia. 2021. v 77.276" type="journal article" year="2021">Cifuentes (2021a)</bibRefCitation>
. Furthermore, 179 other specimens were previously studied, consisting of
<specimenCount id="9D04FD5AFFB1FF9037BAFB1EFDCFFB43" box="[452,562,1160,1187]" count="38" pageId="12" pageNumber="13" type="male">38 males</specimenCount>
,
<specimenCount id="9D04FD5AFFB1FF90343EFB1FFD2CFB43" box="[576,721,1160,1186]" count="136" pageId="12" pageNumber="13" type="female">136 females</specimenCount>
and
<specimenCount id="9D04FD5AFFB1FF903570FB1FFC5FFB43" box="[782,930,1160,1187]" count="5" pageId="12" pageNumber="13" type="immature">5 immatures</specimenCount>
in earlier works (see
<bibRefCitation id="EF934B22FFB1FF9032D4FB1FFA8CFB43" author="Cifuentes J." box="[1194,1393,1160,1186]" pageId="12" pageNumber="13" pagination="143" refId="ref23818" refString="Cifuentes J. 2021 b. Contribucion al conocimiento de los isopodos terrestres de la Comunidad Valenciana y de la Region de Murcia, Espana (Crustacea: Isopoda, Oniscidea). Graellsia 77 (2): e 143. https: // doi. org / 10.3989 / graellsia. 2021. v 77.296" type="journal article" year="2021">Cifuentes 2021b</bibRefCitation>
;
<bibRefCitation id="EF934B22FFB1FF9036C1FB3BFE4BFB27" author="Cifuentes J. &amp; Lopez Martinez S. &amp; Molina Pardo J. L. &amp; Barranco Vega P." box="[191,438,1196,1222]" pageId="12" pageNumber="13" pagination="261 - 276" refId="ref23875" refString="Cifuentes J., Lopez Martinez S., Molina Pardo J. L. &amp; Barranco Vega P. 2021. Isopodos (Crustacea: Oniscidea) presentes en el Medio Subterraneo Superficial (MSS) del Sureste Iberico (Almeria, Espana). Boletin Asociacion espanola de Entomologia 45 (3 - 4): 261 - 276." type="journal article" year="2021">
Cifuentes
<emphasis id="B976EAC1FFB1FF90374BFB3AFE8DFB27" box="[309,368,1196,1222]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2021
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3186558FFB1FF9036C3FB7AFC67F992" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BBD36D3FFB1FF9036C3FB7AFECDFAE6" blockId="12.[189,304,1261,1287]" box="[189,304,1261,1287]" pageId="12" pageNumber="13">
<heading id="D0F581BFFFB1FF9036C3FB7AFECDFAE6" bold="true" box="[189,304,1261,1287]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B976EAC1FFB1FF9036C3FB7AFECDFAE6" bold="true" box="[189,304,1261,1287]" pageId="12" pageNumber="13">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36D3FFB1FF9036C3FA8BFC67F992" blockId="12.[188,1400,1308,1651]" pageId="12" pageNumber="13">
This species is easily recognized by having a highly convex body (
<tableCitation id="C6800368FFB1FF9035B2FA8BFBDEFAD7" box="[972,1059,1308,1334]" captionStart="Table 1" captionStartId="5.[185,255,713,741]" captionTargetPageId="5" captionText="Table 1. Summary of comparative characters for species within the genus Porcellionides Miers, 1877 found in the Ibero-Balearic region." httpUri="http://table.plazi.org/id/DF7D665BFFB8FF9936C7FD5EF8F9FD04" pageId="12" pageNumber="13" tableUuid="DF7D665BFFB8FF9936C7FD5EF8F9FD04">Table 1</tableCitation>
) and by its brown-yellowish coloration, with a distinct marbling pattern (
<figureCitation id="13392A56FFB1FF9034B5FAA8FCD7FAB8" box="[715,810,1343,1369]" captionStart="Fig" captionStartId="13.[189,232,1928,1954]" captionTargetBox="[263,1324,265,1889]" captionTargetId="figure-12@13.[263,1324,265,1889]" captionTargetPageId="13" captionText="Fig. 8. Porcellionides fuscomarmoratus (Budde-Lund, 1885). A. ♀ (JC758), habitus. B. Noduli laterales coordinates. C. ♂ (UGMSS-2), exopod I. D. ♂ (UGMSS-2), exopod II. Scale bars: A = 1 mm; CD = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12157054" httpUri="https://zenodo.org/record/12157054/files/figure.png" pageId="12" pageNumber="13">Fig. 8A</figureCitation>
). It lacks granulations. The position of the noduli laterales shows little variation across tergites, although they follow the general pattern of being situated closer to the lateral and posterior edges in an anterior-posterior direction (
<figureCitation id="13392A56FFB1FF903246FA12FB66FA7E" box="[1080,1179,1413,1439]" captionStart="Fig" captionStartId="6.[189,232,1716,1742]" captionTargetBox="[213,1355,503,1668]" captionTargetId="figure-106@6.[189,1398,496,1681]" captionTargetPageId="6" captionText="Fig. 2. Relation between noduli laterales coordinates (following Vandel 1960) d/c (y-axis) and b/c (x-axis) multiplied by 100 for all studied species. A. Porcellionides cingendus (Kinahan, 1857). B. Porcellionides elegans(Pollo Zorita, 1982). C. Porcellionides fuscomarmoratus(Budde-Lund, 1885). D. Porcellionides glaber (Koch, 1856). E. Porcellionides hispanus (Vandel, 1953). F. Porcellionides lucasioides (Vandel, 1953). G. Porcellionides lusitanus (Vandel, 1946). H. Porcellionides molleri (Verhoeff, 1901). I. Porcellionides pruinosus (Brandt, 1833). J. Porcellionides sexfasciatus (BuddeLund, 1885). K. Porcellionides ibericus sp. nov. L. P. sexfasciatus and its previously considered subspecies: P. glaber (g), P. hispanus (h), P. lucasioides (l), P. lusitanus (lu), P. molleri (m) and P. sexfasciatus (s)." figureDoi="http://doi.org/10.5281/zenodo.12157016" httpUri="https://zenodo.org/record/12157016/files/figure.png" pageId="12" pageNumber="13">Figs 2C</figureCitation>
,
<figureCitation id="13392A56FFB1FF9032D4FA12FB32FA7E" box="[1194,1231,1413,1439]" captionStart="Fig" captionStartId="13.[189,232,1928,1954]" captionTargetBox="[263,1324,265,1889]" captionTargetId="figure-12@13.[263,1324,265,1889]" captionTargetPageId="13" captionText="Fig. 8. Porcellionides fuscomarmoratus (Budde-Lund, 1885). A. ♀ (JC758), habitus. B. Noduli laterales coordinates. C. ♂ (UGMSS-2), exopod I. D. ♂ (UGMSS-2), exopod II. Scale bars: A = 1 mm; CD = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12157054" httpUri="https://zenodo.org/record/12157054/files/figure.png" pageId="12" pageNumber="13">8B</figureCitation>
). Scales-setae are triangular with very long and fine tips (
<figureCitation id="13392A56FFB1FF9034BAFA3FFCDFFA22" box="[708,802,1448,1475]" captionStart="Fig" captionStartId="7.[189,232,1786,1812]" captionTargetBox="[231,1365,1102,1731]" captionTargetId="figure-323@7.[189,1398,1089,1751]" captionTargetPageId="7" captionText="Fig. 3. Scale-setae. A. Porcellionides cingendus (Kinahan, 1857), ♂ (JC104). B. Porcellionides elegans (Pollo Zorita, 1982), ♂ (MNCN 20.04/7306). C. Porcellionides fuscomarmoratus (BuddeLund, 1885), ♀ (MNCN 20.04/8129). D. Porcellionides glaber (Koch, 1856), ♀ (MNCN 20.04/11490). E. Porcellionides hispanus (Vandel, 1953), ♂ (MNCN 20.04/7163). F. Porcellionides lucasioides (Vandel, 1953), ♂ (JC352). G. Porcellionides lusitanus (Vandel, 1946), ♂ (LPS661). H. Porcellionides molleri (Verhoeff, 1901), ♀ (LPS135). I. Porcellionides pruinosus (Brandt, 1833), ♀ (JC318). J. Porcellionides sexfasciatus (Budde-Lund, 1885), ♂ (LPS455a). Scale bar = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.12157020" httpUri="https://zenodo.org/record/12157020/files/figure.png" pageId="12" pageNumber="13">Fig. 3C</figureCitation>
). The lateral lobes of the cephalon are reduced to protrusions. There is no transverse ridge on the pereon (
<figureCitation id="13392A56FFB1FF90351CFA5BFC3CFA07" box="[866,961,1484,1510]" captionStart="Fig" captionStartId="13.[189,232,1928,1954]" captionTargetBox="[263,1324,265,1889]" captionTargetId="figure-12@13.[263,1324,265,1889]" captionTargetPageId="13" captionText="Fig. 8. Porcellionides fuscomarmoratus (Budde-Lund, 1885). A. ♀ (JC758), habitus. B. Noduli laterales coordinates. C. ♂ (UGMSS-2), exopod I. D. ♂ (UGMSS-2), exopod II. Scale bars: A = 1 mm; CD = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12157054" httpUri="https://zenodo.org/record/12157054/files/figure.png" pageId="12" pageNumber="13">Fig. 8A</figureCitation>
,
<tableCitation id="C6800368FFB1FF9035B1FA5BFBD5FA07" box="[975,1064,1484,1510]" captionStart="Table 1" captionStartId="5.[185,255,713,741]" captionTargetPageId="5" captionText="Table 1. Summary of comparative characters for species within the genus Porcellionides Miers, 1877 found in the Ibero-Balearic region." httpUri="http://table.plazi.org/id/DF7D665BFFB8FF9936C7FD5EF8F9FD04" pageId="12" pageNumber="13" tableUuid="DF7D665BFFB8FF9936C7FD5EF8F9FD04">Table 1</tableCitation>
). The pleon is more or less retracted in relation to the pereon. In males, the carpus of the pereopod 1 has a brush of setae, the exopod of the pleopod I has a short, triangular, and pointed posterior inner tip (
<figureCitation id="13392A56FFB1FF90327CF985FBA4F9CD" box="[1026,1113,1554,1580]" captionStart="Fig" captionStartId="8.[189,232,1704,1730]" captionTargetBox="[323,1265,274,1643]" captionTargetId="figure-17@8.[280,1308,262,1666]" captionText="Fig. 4. Pleopods. AD. Porcellionides cingendus (Kinahan, 1857). AB. ♂ (LPS215). A. Exopod I. B. Exopod II. CD. ♀ (MNCN 20.04/11972). C. Exopod I. D. Exopod II. EH. Porcellionides elegans (Pollo Zorita, 1982). EF. ♂ (JC490). E. Exopod I. F.Exopod II. GH. ♀ (MNCN 20.04/9590). G. Exopod I. H. Exopod II. IL. Porcellionides fuscomarmoratus (Budde-Lund, 1885). IJ. ♂ (UGMSS-2). I. Exopod I. J. Exopod II. KL. ♀ (MNCN 20.04/8129. K. Exopod I. L. Exopod II. MP. Porcellionides glaber (Koch, 1856). ML. ♂ (MNCN 20.04/7886). M. Exopod I. N. Exopod II. O. ♀ (MNCN 20.04/11490), exopod I. P. ♀ (MNCN 20.04/11490), exopod II. QT. Porcellionides hispanus (Vandel, 1953). QR. ♂ (MNCN 20.04/7163). Q. Exopod I. R. Exopod II. ST. ♀ (JC380). S. Exopod I. T. ♀ (JC380), exopod II. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12188976" httpUri="https://zenodo.org/record/12188976/files/figure.png" pageId="12" pageNumber="13">Figs 4I</figureCitation>
, 8C), and the exopod of the pleopod II is triangular with a very long posterior inner tip (
<figureCitation id="13392A56FFB1FF9035E2F9A2FC09F9AE" box="[924,1012,1589,1615]" captionStart="Fig" captionStartId="8.[189,232,1704,1730]" captionTargetBox="[323,1265,274,1643]" captionTargetId="figure-17@8.[280,1308,262,1666]" captionText="Fig. 4. Pleopods. AD. Porcellionides cingendus (Kinahan, 1857). AB. ♂ (LPS215). A. Exopod I. B. Exopod II. CD. ♀ (MNCN 20.04/11972). C. Exopod I. D. Exopod II. EH. Porcellionides elegans (Pollo Zorita, 1982). EF. ♂ (JC490). E. Exopod I. F.Exopod II. GH. ♀ (MNCN 20.04/9590). G. Exopod I. H. Exopod II. IL. Porcellionides fuscomarmoratus (Budde-Lund, 1885). IJ. ♂ (UGMSS-2). I. Exopod I. J. Exopod II. KL. ♀ (MNCN 20.04/8129. K. Exopod I. L. Exopod II. MP. Porcellionides glaber (Koch, 1856). ML. ♂ (MNCN 20.04/7886). M. Exopod I. N. Exopod II. O. ♀ (MNCN 20.04/11490), exopod I. P. ♀ (MNCN 20.04/11490), exopod II. QT. Porcellionides hispanus (Vandel, 1953). QR. ♂ (MNCN 20.04/7163). Q. Exopod I. R. Exopod II. ST. ♀ (JC380). S. Exopod I. T. ♀ (JC380), exopod II. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12188976" httpUri="https://zenodo.org/record/12188976/files/figure.png" pageId="12" pageNumber="13">Figs 4J</figureCitation>
, 8D). In females, they are similar to those of other studied species within the genus (
<figureCitation id="13392A56FFB1FF903573F9CFFC70F992" box="[781,909,1624,1651]" captionStart="Fig" captionStartId="8.[189,232,1704,1730]" captionTargetBox="[323,1265,274,1643]" captionTargetId="figure-17@8.[280,1308,262,1666]" captionText="Fig. 4. Pleopods. AD. Porcellionides cingendus (Kinahan, 1857). AB. ♂ (LPS215). A. Exopod I. B. Exopod II. CD. ♀ (MNCN 20.04/11972). C. Exopod I. D. Exopod II. EH. Porcellionides elegans (Pollo Zorita, 1982). EF. ♂ (JC490). E. Exopod I. F.Exopod II. GH. ♀ (MNCN 20.04/9590). G. Exopod I. H. Exopod II. IL. Porcellionides fuscomarmoratus (Budde-Lund, 1885). IJ. ♂ (UGMSS-2). I. Exopod I. J. Exopod II. KL. ♀ (MNCN 20.04/8129. K. Exopod I. L. Exopod II. MP. Porcellionides glaber (Koch, 1856). ML. ♂ (MNCN 20.04/7886). M. Exopod I. N. Exopod II. O. ♀ (MNCN 20.04/11490), exopod I. P. ♀ (MNCN 20.04/11490), exopod II. QT. Porcellionides hispanus (Vandel, 1953). QR. ♂ (MNCN 20.04/7163). Q. Exopod I. R. Exopod II. ST. ♀ (JC380). S. Exopod I. T. ♀ (JC380), exopod II. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12188976" httpUri="https://zenodo.org/record/12188976/files/figure.png" pageId="12" pageNumber="13">Fig. 4KL</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3186558FFB1FF9036C3F90DFC07F857" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8BBD36D3FFB1FF9036C3F90DFEA5F955" blockId="12.[189,344,1690,1716]" box="[189,344,1690,1716]" pageId="12" pageNumber="13">
<heading id="D0F581BFFFB1FF9036C3F90DFEA5F955" bold="true" box="[189,344,1690,1716]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B976EAC1FFB1FF9036C3F90DFEA5F955" bold="true" box="[189,344,1690,1716]" pageId="12" pageNumber="13">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36D3FFB1FF9036C3F95FFC07F857" blockId="12.[188,1399,1736,1974]" pageId="12" pageNumber="13">
Only in the eastern part of the Ibero-Balearic region (
<figureCitation id="13392A56FFB1FF903548F95FFC82F902" box="[822,895,1736,1763]" captionStart="Fig" captionStartId="14.[189,232,1963,1989]" captionTargetBox="[228,1362,927,1918]" captionTargetId="figure-193@14.[189,1398,921,1927]" captionTargetPageId="14" captionText="Fig. 9. Localization of the Ibero-Balearic region in Europe and the distribution of Porcellionides fuscomarmoratus (Budde-Lund, 1885)." figureDoi="http://doi.org/10.5281/zenodo.12157062" httpUri="https://zenodo.org/record/12157062/files/figure.png" pageId="12" pageNumber="13">Fig. 9</figureCitation>
). Spanish providences of
<emphasis id="B976EAC1FFB1FF9032CDF95FFAD9F902" bold="true" box="[1203,1316,1736,1763]" pageId="12" pageNumber="13">Almería:</emphasis>
Dalias (
<bibRefCitation id="EF934B22FFB1FF9036BBF97BFE9DF8E7" author="Vandel A." box="[197,352,1771,1798]" pageId="12" pageNumber="13" pagination="45 - 75" refId="ref25690" refString="Vandel A. 1953. Les isopodes terrestres des provinces d'Almeria et de Granada. Archivos del Instituto de Aclimatacion (Almeria) 1: 45 - 75." type="journal article" year="1953">Vandel 1953</bibRefCitation>
), Felix, Fondón, María and Vélez Rubio (
<bibRefCitation id="EF934B22FFB1FF90352EF97BFBB2F8E7" author="Cifuentes J. &amp; Lopez Martinez S. &amp; Molina Pardo J. L. &amp; Barranco Vega P." box="[848,1103,1772,1798]" pageId="12" pageNumber="13" pagination="261 - 276" refId="ref23875" refString="Cifuentes J., Lopez Martinez S., Molina Pardo J. L. &amp; Barranco Vega P. 2021. Isopodos (Crustacea: Oniscidea) presentes en el Medio Subterraneo Superficial (MSS) del Sureste Iberico (Almeria, Espana). Boletin Asociacion espanola de Entomologia 45 (3 - 4): 261 - 276." type="journal article" year="2021">
Cifuentes
<emphasis id="B976EAC1FFB1FF9035B2F97BFBFAF8E4" box="[972,1031,1772,1798]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2021
</bibRefCitation>
);
<emphasis id="B976EAC1FFB1FF903218F97CFB22F8E7" bold="true" box="[1126,1247,1771,1798]" pageId="12" pageNumber="13">Córdoba:</emphasis>
Santa María de Trassierra (
<bibRefCitation id="EF934B22FFB1FF90371FF898FE05F8C8" author="Garcia L." box="[353,504,1807,1833]" pageId="12" pageNumber="13" pagination="27 - 32" refId="ref24419" refString="Garcia L. 2019. Nuevos registros de Isopodos terrestres (Crustacea: Oniscidea) en Espana meridional (Andalucia y Murcia). Revista Sociedad Gaditana Historia Natural 13: 27 - 32." type="journal article" year="2019">Garcia 2019</bibRefCitation>
);
<emphasis id="B976EAC1FFB1FF903476F898FD7EF8C8" bold="true" box="[520,643,1807,1833]" pageId="12" pageNumber="13">
<collectingCountry id="F3157643FFB1FF903476F898FD83F8C8" box="[520,638,1807,1833]" name="Grenada" pageId="12" pageNumber="13">Granada</collectingCountry>
:
</emphasis>
Sagra (
<bibRefCitation id="EF934B22FFB1FF9034A5F898FC8CF8CB" author="Vandel A." box="[731,881,1807,1834]" pageId="12" pageNumber="13" pagination="45 - 75" refId="ref25690" refString="Vandel A. 1953. Les isopodes terrestres des provinces d'Almeria et de Granada. Archivos del Instituto de Aclimatacion (Almeria) 1: 45 - 75." type="journal article" year="1953">Vandel 1953</bibRefCitation>
) and Sierra Nevada (
<bibRefCitation id="EF934B22FFB1FF90321DF898FB06F8CB" author="Vandel A." box="[1123,1275,1807,1834]" pageId="12" pageNumber="13" pagination="45 - 75" refId="ref25690" refString="Vandel A. 1953. Les isopodes terrestres des provinces d'Almeria et de Granada. Archivos del Instituto de Aclimatacion (Almeria) 1: 45 - 75." type="journal article" year="1953">Vandel 1953</bibRefCitation>
;
<bibRefCitation id="EF934B22FFB1FF903378F898FEF5F8AD" author="Cifuentes J." pageId="12" pageNumber="13" pagination="133" refId="ref23773" refString="Cifuentes J. 2021 a. Los isopodos terrestres de Andalucia, Espana (Crustacea: Isopoda, Oniscidea). Graellsia 77 (1): e 133. https: // doi. org / 10.3989 / graellsia. 2021. v 77.276" type="journal article" year="2021">Cifuentes 2021a</bibRefCitation>
);
<emphasis id="B976EAC1FFB1FF903763F8A5FE2FF8AD" bold="true" box="[285,466,1842,1868]" pageId="12" pageNumber="13">Islas Baleares:</emphasis>
Cabrera (
<bibRefCitation id="EF934B22FFB1FF90343DF8A5FCCEF8AD" author="Garcia L. &amp; Cruz A." box="[579,819,1842,1868]" pageId="12" pageNumber="13" pagination="323 - 332" refId="ref24454" refString="Garcia L. &amp; Cruz A. 1993. Els isopodes terrestres (Crustacea: Isopoda: Oniscidea). In: Alcover J., Ballesteros E. &amp; Fornos J. (eds) Historia Natural de l'Arxipelag de Cabrera. XIX. Monografies de la Societat d'Historia Natural de les Baleares 2: 323 - 332. Editorial Moll, Mallorca." type="journal article" year="1993">Garcia &amp; Cruz 1993</bibRefCitation>
,
<bibRefCitation id="EF934B22FFB1FF903540F8A5FC81F8AD" author="Garcia L. &amp; Cruz A." box="[830,892,1842,1868]" pageId="12" pageNumber="13" pagination="77 - 99" refId="ref24519" refString="Garcia L. &amp; Cruz A. 1996. Els isopodos terrestres (Crustacea: Isopoda: Oniscidea) de les Illes Baleares: cataleg d'especies. Bolleti de la Societat d' Historia natural de les Baleares 39: 77 - 99." type="journal article" year="1996">1996</bibRefCitation>
), Dragonera (
<bibRefCitation id="EF934B22FFB1FF903267F8A5FB53F8AD" author="Garcia L." box="[1049,1198,1842,1868]" pageId="12" pageNumber="13" pagination="203 - 224" refId="ref24254" refString="Garcia L. 2008. Els isopodes terrestres (Crustacea: Isopoda: Oniscidea) del Parc Natural de l'iLa de sa Dragonera (Islas Baleares, Mediterrania occidental). Bolleti de la Societat d'Historia Natural de les Baleares 51: 203 - 224." type="journal article" year="2008">Garcia 2008</bibRefCitation>
), Ibiza (
<bibRefCitation id="EF934B22FFB1FF903374F8A5FECAF88E" author="Garcia L. &amp; Cruz A." pageId="12" pageNumber="13" pagination="77 - 99" refId="ref24519" refString="Garcia L. &amp; Cruz A. 1996. Els isopodos terrestres (Crustacea: Isopoda: Oniscidea) de les Illes Baleares: cataleg d'especies. Bolleti de la Societat d' Historia natural de les Baleares 39: 77 - 99." type="journal article" year="1996">Garcia &amp; Cruz 1996</bibRefCitation>
) and Mallorca (
<bibRefCitation id="EF934B22FFB1FF903791F8C2FD1FF88E" author="Garcia L. &amp; Cruz A." box="[495,738,1877,1903]" pageId="12" pageNumber="13" pagination="77 - 99" refId="ref24519" refString="Garcia L. &amp; Cruz A. 1996. Els isopodos terrestres (Crustacea: Isopoda: Oniscidea) de les Illes Baleares: cataleg d'especies. Bolleti de la Societat d' Historia natural de les Baleares 39: 77 - 99." type="journal article" year="1996">Garcia &amp; Cruz 1996</bibRefCitation>
;
<bibRefCitation id="EF934B22FFB1FF903490F8C2FC7EF88E" author="Garcia L." box="[750,899,1877,1903]" pageId="12" pageNumber="13" pagination="129 - 146" refId="ref24300" refString="Garcia L. 2009. Les &quot; someretes &quot; (Crustacis isopodes terrestres: Isopoda, Oniscidea) de la vall de Soller (Mallorca). Aproximacio a una biodiversitat menystinguda. In: Ajuntament de Soller (eds) III Jornades Estudis Locals a Soller: 129 - 146. Soller, Mallorca." type="book chapter" year="2009">Garcia 2009</bibRefCitation>
);
<emphasis id="B976EAC1FFB1FF9035EBF8C3FC24F88E" bold="true" box="[917,985,1876,1903]" pageId="12" pageNumber="13">Jaén:</emphasis>
Cazorla (
<bibRefCitation id="EF934B22FFB1FF903235F8C2FB1CF88E" author="Garcia L." box="[1099,1249,1877,1903]" pageId="12" pageNumber="13" pagination="78 - 87" refId="ref24356" refString="Garcia L. 2013. Isopodos terrestres (Crustacea: Oniscidea) recolectados en cavidades subterraneas de Jaen. In: Perez Fernandez T. &amp; Perez Ruiz A. (coords) Los Invertebrados de Habitats Subterraneos de Jaen: 78 - 87. Grupo de Espeleologia de Villacarrillo (G. E. V.), Villacarrillo, Spain." type="book chapter" year="2013">Garcia 2013</bibRefCitation>
), Poblado de Vadillo-Castril, Quesada and Sierra de Cazorla (
<bibRefCitation id="EF934B22FFB1FF90348DF8EFFC47F872" author="Cifuentes J." box="[755,954,1912,1939]" pageId="12" pageNumber="13" pagination="133" refId="ref23773" refString="Cifuentes J. 2021 a. Los isopodos terrestres de Andalucia, Espana (Crustacea: Isopoda, Oniscidea). Graellsia 77 (1): e 133. https: // doi. org / 10.3989 / graellsia. 2021. v 77.276" type="journal article" year="2021">Cifuentes 2021a</bibRefCitation>
);
<emphasis id="B976EAC1FFB1FF9035B1F8EFFBC5F872" bold="true" box="[975,1080,1912,1939]" pageId="12" pageNumber="13">Málaga:</emphasis>
Antequera (
<bibRefCitation id="EF934B22FFB1FF9032B6F8EFFA94F872" author="Dollfus A." box="[1224,1385,1912,1939]" pageId="12" pageNumber="13" pagination="161 - 190" refId="ref24087" refString="Dollfus A. 1892. Catalogue raisonne des isopodes terrestres de l'Espagne. Anales de la Sociedad espanola de Historia natural 21: 161 - 190." type="journal article" year="1892">Dollfus 1892</bibRefCitation>
);
<emphasis id="B976EAC1FFB1FF9036C3F80CFED9F854" bold="true" box="[189,292,1947,1973]" pageId="12" pageNumber="13">Murcia:</emphasis>
Sierra de Espuña (
<bibRefCitation id="EF934B22FFB1FF90347FF80BFD3BF857" author="Cifuentes J." box="[513,710,1948,1974]" pageId="12" pageNumber="13" pagination="143" refId="ref23818" refString="Cifuentes J. 2021 b. Contribucion al conocimiento de los isopodos terrestres de la Comunidad Valenciana y de la Region de Murcia, Espana (Crustacea: Isopoda, Oniscidea). Graellsia 77 (2): e 143. https: // doi. org / 10.3989 / graellsia. 2021. v 77.296" type="journal article" year="2021">Cifuentes 2021b</bibRefCitation>
) and Yecla (
<bibRefCitation id="EF934B22FFB1FF903529F80BFC13F857" author="Garcia L." box="[855,1006,1948,1974]" pageId="12" pageNumber="13" pagination="27 - 32" refId="ref24419" refString="Garcia L. 2019. Nuevos registros de Isopodos terrestres (Crustacea: Oniscidea) en Espana meridional (Andalucia y Murcia). Revista Sociedad Gaditana Historia Natural 13: 27 - 32." type="journal article" year="2019">Garcia 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>