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<document id="E47BB447A6D76A62E1BE551F7A37A1EB" ID-CLB-Dataset="171013" ID-DOI="10.5852/ejt.2022.843.1947" ID-GBIF-Dataset="044b60a6-0e4e-44be-b75a-614b58434dcc" ID-ISSN="2118-9773" ID-Zenodo-Dep="7222308" ID-ZooBank="C03BE897-EFE2-4CCD-913A-723792CDF050" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666126358981" checkinUser="felipe" docAuthor="Wood, Thomas J. &amp; Monfared, Alireza" docDate="2022" docId="03AB87A2FFB95119DA41D8E3FD92F85C" docLanguage="en" docName="ejt-843_1-136.pdf" docOrigin="European Journal of Taxonomy 843" docSource="http://dx.doi.org/10.5852/ejt.2022.843.1947" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Andrena (Ulandrena) bulbosa Wood &amp; Monfared 2022, sp. nov." docType="treatment" docUuid="86FAABC5-56AB-4758-AC10-8E045BD9AC28" docUuidSource="ZooBank" docVersion="11" lastPageNumber="49" masterDocId="FF92FFDAFF945129D84DD914FFB0FFB4" masterDocTitle="A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species" masterLastPageNumber="136" masterPageNumber="1" pageNumber="46" updateTime="1699408809500" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B873201236C789BDA7DD11F7877B826A">A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species</mods:title>
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<mods:namePart id="D48D8926D19526CAF98A8B0FC675E5CD">Wood, Thomas J.</mods:namePart>
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<mods:affiliation id="9C2E919D7412C126B5442C2257E98808">Laboratory of Zoology, University of Mons, 7000 Mons, Belgium.</mods:affiliation>
<mods:nameIdentifier id="4322A5B6050DD144C18A16453FA032DE" type="email">thomasjames.wood@umons.ac.be</mods:nameIdentifier>
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<mods:namePart id="686762DF5A225F9FA8743FEFEA3257C5">Monfared, Alireza</mods:namePart>
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<mods:affiliation id="B1995CCAA8AB0236C06C1A97C8BB807B">Department of Plant Protection, Faculty of Agriculture, Yasouj University, Yasouj, Iran.</mods:affiliation>
<mods:nameIdentifier id="96A2D1AF709CED75D994683EC91DC8A7" type="email">amonfared@yu.ac.ir</mods:nameIdentifier>
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<treatment id="03AB87A2FFB95119DA41D8E3FD92F85C" ID-DOI="http://doi.org/10.5281/zenodo.7222631" ID-GBIF-Taxon="203124997" ID-Zenodo-Dep="7222631" LSID="urn:lsid:zoobank.org:act:86FAABC5-56AB-4758-AC10-8E045BD9AC28" httpUri="http://treatment.plazi.org/id/03AB87A2FFB95119DA41D8E3FD92F85C" lastPageId="48" lastPageNumber="49" pageId="45" pageNumber="46">
<subSubSection id="C318653FFFB95104DA41D8E3FB98FDA6" box="[524,1064,503,530]" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="8BBD36B4FFB95104DA41D8E3FB98FDA6" blockId="45.[357,1230,503,602]" box="[524,1064,503,530]" pageId="45" pageNumber="46">
<heading id="D0F581D8FFB95104DA41D8E3FB98FDA6" box="[524,1064,503,530]" centered="true" fontSize="11" level="2" pageId="45" pageNumber="46" reason="2">
<taxonomicName id="4C024D37FFB95104DA41D8E3FC75FDA6" authority="Wood" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[524,965,503,530]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="bulbosa" status="sp. nov." subGenus="Ulandrena">
<emphasis id="B976EAA6FFB95104DA41D8E3FCCAFDA6" bold="true" box="[524,890,503,530]" pageId="45" pageNumber="46">
<emphasis id="B976EAA6FFB95104DA41D8E3FDC6FDA6" bold="true" box="[524,630,503,530]" italics="true" pageId="45" pageNumber="46">Andrena</emphasis>
(
<emphasis id="B976EAA6FFB95104DACAD8E3FCB9FDA6" bold="true" box="[647,777,503,530]" italics="true" pageId="45" pageNumber="46">Ulandrena</emphasis>
)
<emphasis id="B976EAA6FFB95104DB56D8E3FCCAFDA6" bold="true" box="[795,890,503,530]" italics="true" pageId="45" pageNumber="46">bulbosa</emphasis>
</emphasis>
Wood
</taxonomicName>
<taxonomicNameLabel id="A24557DDFFB95104DB80D8ECFB98FDA6" box="[973,1064,504,530]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C318653FFFB95104D928DB08FCDAFDEE" pageId="45" pageNumber="46" type="description">
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="8BBD36B4FFB95104DA84DB54FCDAFDEE" blockId="45.[357,1230,503,602]" box="[713,874,576,602]" pageId="45" pageNumber="46">
<figureCitation id="13392A31FFB95104DA84DB54FCDAFDEE" box="[713,874,576,602]" captionStart-0="Figs 109114" captionStart-1="Figs 115120" captionStartId-0="46.[191,243,1965,1991]" captionStartId-1="47.[189,241,1961,1987]" captionTargetBox-0="[189,1398,557,1918]" captionTargetBox-1="[189,1398,558,1919]" captionTargetId-0="figure-119@46.[189,1398,557,1918]" captionTargetId-1="figure-123@47.[189,1398,558,1919]" captionTargetPageId-0="46" captionTargetPageId-1="47" captionText-0="Figs 109114. Andrena bulbosa Wood sp. nov., ♀ (OÖLM). 109. Profile. 110. Face. 111. Process of labrum, detail. 112. Scutum. 113. Terga. 114. Terga 45, detail." captionText-1="Figs 115120. Andrena bulbosa Wood sp. nov., ♂ (OÖLM). 115. Profile. 116. Face. 117. Terga. 118. Genitalia dorsal view. 119. Genitalia lateral view. 120. Genitalia ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.7222388" figureDoi-1="http://doi.org/10.5281/zenodo.7222392" httpUri-0="https://zenodo.org/record/7222388/files/figure.png" httpUri-1="https://zenodo.org/record/7222392/files/figure.png" pageId="45" pageNumber="46">Figs 109120</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C318653FFFB95104D8F0DB96FE93FA0B" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="8BBD36B4FFB95104D8F0DB96FE86FD28" blockId="45.[189,310,642,668]" box="[189,310,642,668]" pageId="45" pageNumber="46">
<heading id="D0F581D8FFB95104D8F0DB96FE86FD28" bold="true" box="[189,310,642,668]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="B976EAA6FFB95104D8F0DB96FE86FD28" bold="true" box="[189,310,642,668]" pageId="45" pageNumber="46">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FFB95104D8F0DBA6FD46FBEF" blockId="45.[187,1399,690,1115]" pageId="45" pageNumber="46">
<taxonomicName id="4C024D37FFB95104D8F0DBA6FE3AFD78" authority="Wood &amp; Monfared, 2022" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[189,394,690,716]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="bulbosa" status="sp. nov.">
<emphasis id="B976EAA6FFB95104D8F0DBA6FE3AFD78" box="[189,394,690,716]" italics="true" pageId="45" pageNumber="46">Andrena bulbosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557DDFFB95104D9DBDBA7FE45FD79" box="[406,501,691,717]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
can swiftly be placed in the subgenus
<taxonomicName id="4C024D37FFB95104DB93DBA6FAAEFD78" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" box="[990,1310,690,716]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ulandrena">
<emphasis id="B976EAA6FFB95104DB93DBA6FBECFD78" box="[990,1116,690,716]" italics="true" pageId="45" pageNumber="46">Ulandrena</emphasis>
<bibRefCitation id="EF934B45FFB95104DC25DBA6FAAEFD78" author="Warncke" box="[1128,1310,690,716]" firstAuthor="Warncke" pageId="45" pageNumber="46" pagination="1 - 110" refId="ref70132" refString="Warncke K. 1968. Die Untergattungen der westpalaarktischen Bienengattung Andrena F. Memorias e Estudos do Museu Zoologico da Universidade de Coimbra 307: 1 - 110." type="journal article" year="1968">Warncke, 1968</bibRefCitation>
</taxonomicName>
by the combination of the inner hind tibial spur that is broadened slightly before its base, the yellow-marked male clypeus (
<figureCitation id="13392A31FFB95104D92EDBEFFE74FCA1" box="[355,452,763,789]" captionStart="Figs 115120" captionStartId="47.[189,241,1961,1987]" captionTargetBox="[189,1398,558,1919]" captionTargetId="figure-123@47.[189,1398,558,1919]" captionTargetPageId="47" captionText="Figs 115120. Andrena bulbosa Wood sp. nov., ♂ (OÖLM). 115. Profile. 116. Face. 117. Terga. 118. Genitalia dorsal view. 119. Genitalia lateral view. 120. Genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.7222392" httpUri="https://zenodo.org/record/7222392/files/figure.png" pageId="45" pageNumber="46">Fig.116</figureCitation>
), and generally enlarged genital capsule (
<figureCitation id="13392A31FFB95104DBD7DBEFFC4DFCA1" box="[922,1021,763,789]" captionStart="Figs 115120" captionStartId="47.[189,241,1961,1987]" captionTargetBox="[189,1398,558,1919]" captionTargetId="figure-123@47.[189,1398,558,1919]" captionTargetPageId="47" captionText="Figs 115120. Andrena bulbosa Wood sp. nov., ♂ (OÖLM). 115. Profile. 116. Face. 117. Terga. 118. Genitalia dorsal view. 119. Genitalia lateral view. 120. Genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.7222392" httpUri="https://zenodo.org/record/7222392/files/figure.png" pageId="45" pageNumber="46">Fig. 118</figureCitation>
). It belongs to the
<taxonomicName id="4C024D37FFB95104DC8ADBEFFAC3FCA1" authorityName="Smith" authorityYear="1847" box="[1223,1395,763,789]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="polita">
<emphasis id="B976EAA6FFB95104DC8ADBEFFAC3FCA1" box="[1223,1395,763,789]" italics="true" pageId="45" pageNumber="46">Andrena polita</emphasis>
</taxonomicName>
species group and is closest to the larger species with red-marked terga, specifically
<taxonomicName id="4C024D37FFB95104DC32DA0BFF48FCE9" authority="Morawitz, 1886" authorityName="Morawitz" authorityYear="1886" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="caspica">
<emphasis id="B976EAA6FFB95104DC32DA0BFB48FC8D" box="[1151,1272,799,825]" italics="true" pageId="45" pageNumber="46">A. caspica</emphasis>
Morawitz, 1886
</taxonomicName>
,
<taxonomicName id="4C024D37FFB95104D948DA50FD9BFCE9" authority="Friese, 1899" authorityName="Friese" authorityYear="1899" box="[261,555,835,862]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="unicincta">
<emphasis id="B976EAA6FFB95104D948DA50FE22FCE9" box="[261,402,836,862]" italics="true" pageId="45" pageNumber="46">A. unicincta</emphasis>
Friese, 1899
</taxonomicName>
, and
<taxonomicName id="4C024D37FFB95104DA2ADA50FC14FCE9" authority="Friese, 1899" authorityName="Friese" authorityYear="1899" box="[615,932,835,862]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="unifasciata">
<emphasis id="B976EAA6FFB95104DA2ADA50FCB9FCE9" box="[615,777,835,862]" italics="true" pageId="45" pageNumber="46">A. unifasciata</emphasis>
Friese, 1899
</taxonomicName>
. Males can instantly be separated by the unique genitalia, which lack gonocoxal teeth (
<figureCitation id="13392A31FFB95104DA9BDA73FC89FC36" box="[726,825,871,898]" captionStart="Figs 115120" captionStartId="47.[189,241,1961,1987]" captionTargetBox="[189,1398,558,1919]" captionTargetId="figure-123@47.[189,1398,558,1919]" captionTargetPageId="47" captionText="Figs 115120. Andrena bulbosa Wood sp. nov., ♂ (OÖLM). 115. Profile. 116. Face. 117. Terga. 118. Genitalia dorsal view. 119. Genitalia lateral view. 120. Genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.7222392" httpUri="https://zenodo.org/record/7222392/files/figure.png" pageId="45" pageNumber="46">Fig. 118</figureCitation>
) (vs strongly produced and pointed in
<taxonomicName id="4C024D37FFB95104DCB1DA7CFAC6FC35" authorityName="Morawitz" authorityYear="1886" box="[1276,1398,872,898]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="caspica">
<emphasis id="B976EAA6FFB95104DCB1DA7CFAC6FC35" box="[1276,1398,872,898]" italics="true" pageId="45" pageNumber="46">A. caspica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C024D37FFB95104D8A2DA98FE22FC12" authorityName="Friese" authorityYear="1899" box="[239,402,908,934]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="unifasciata">
<emphasis id="B976EAA6FFB95104D8A2DA98FE22FC12" box="[239,402,908,934]" italics="true" pageId="45" pageNumber="46">A. unifasciata</emphasis>
</taxonomicName>
, weakly produced and laterally diverging in
<taxonomicName id="4C024D37FFB95104DBE9DA98FB82FC11" authorityName="Friese" authorityYear="1899" box="[932,1074,908,934]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="unicincta">
<emphasis id="B976EAA6FFB95104DBE9DA98FB82FC11" box="[932,1074,908,934]" italics="true" pageId="45" pageNumber="46">A. unicincta</emphasis>
</taxonomicName>
). Females can be separated from
<taxonomicName id="4C024D37FFB95104D94CDAA4FE23FC7E" authorityName="Friese" authorityYear="1899" box="[257,403,944,970]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="unicincta">
<emphasis id="B976EAA6FFB95104D94CDAA4FE23FC7E" box="[257,403,944,970]" italics="true" pageId="45" pageNumber="46">A. unicincta</emphasis>
</taxonomicName>
by the shiny scutum (
<figureCitation id="13392A31FFB95104DAE6DAA4FCA2FC7E" box="[683,786,944,970]" captionStart="Figs 109114" captionStartId="46.[191,243,1965,1991]" captionTargetBox="[189,1398,557,1918]" captionTargetId="figure-119@46.[189,1398,557,1918]" captionTargetPageId="46" captionText="Figs 109114. Andrena bulbosa Wood sp. nov., ♀ (OÖLM). 109. Profile. 110. Face. 111. Process of labrum, detail. 112. Scutum. 113. Terga. 114. Terga 45, detail." figureDoi="http://doi.org/10.5281/zenodo.7222388" httpUri="https://zenodo.org/record/7222388/files/figure.png" pageId="45" pageNumber="46">Fig. 112</figureCitation>
) (vs dull) and interstitial nervulus (vs postfurcal). Separation from
<taxonomicName id="4C024D37FFB95104D9CEDAC0FE4EFC5A" authorityName="Morawitz" authorityYear="1886" box="[387,510,980,1006]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="caspica">
<emphasis id="B976EAA6FFB95104D9CEDAC0FE4EFC5A" box="[387,510,980,1006]" italics="true" pageId="45" pageNumber="46">A. caspica</emphasis>
</taxonomicName>
can be made by the smaller body size of
<quantity id="4CFA9B51FFB95104DBA8DAC0FB88FC5B" box="[997,1080,980,1007]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="45" pageNumber="46" unit="mm" value="10.0">10 mm</quantity>
(
<quantity id="4CFA9B51FFB95104DC05DAC0FB7AFC5A" box="[1096,1226,980,1006]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="45" pageNumber="46" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">1213 mm</quantity>
) as well as the comparatively shinier scutum (vs only slightly shiny in
<taxonomicName id="4C024D37FFB95104DB05DAEDFC72FBA6" authorityName="Morawitz" authorityYear="1886" box="[840,962,1017,1043]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="caspica">
<emphasis id="B976EAA6FFB95104DB05DAEDFC72FBA6" box="[840,962,1017,1043]" italics="true" pageId="45" pageNumber="46">A. caspica</emphasis>
</taxonomicName>
, comparative material required). It is therefore very similar to
<taxonomicName id="4C024D37FFB95104D9A8DD09FD38FB83" authorityName="Friese" authorityYear="1899" box="[485,648,1053,1079]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="unifasciata">
<emphasis id="B976EAA6FFB95104D9A8DD09FD38FB83" box="[485,648,1053,1079]" italics="true" pageId="45" pageNumber="46">A. unifasciata</emphasis>
</taxonomicName>
, and based on the limited female material, separation from this taxon in the female sex is not currently possible.
</paragraph>
<paragraph id="8BBD36B4FFB95104D8F0DD97FE93FA0B" blockId="45.[189,1401,1155,1471]" pageId="45" pageNumber="46">
Similar red-marked female material of
<taxonomicName id="4C024D37FFB95104DAC1DD97FCBAFB29" authorityName="Warncke" authorityYear="1968" box="[652,778,1155,1181]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ulandrena">
<emphasis id="B976EAA6FFB95104DAC1DD97FCBAFB29" box="[652,778,1155,1181]" italics="true" pageId="45" pageNumber="46">Ulandrena</emphasis>
</taxonomicName>
collected from Uludere in
<collectingRegion id="49C6F856FFB95104DC07DD97FAB7FB29" box="[1098,1287,1155,1181]" country="Turkey" name="Sirnak" pageId="45" pageNumber="46">Şırnak Province</collectingRegion>
(
<collectingCountry id="F3157624FFB95104DD5ADD97FADFFB29" box="[1303,1391,1155,1181]" name="Turkey" pageId="45" pageNumber="46">Turkey</collectingCountry>
) by Klaus Warncke (
<date id="FFBC1074FFB95104D9F0DDB3FDD6FB76" box="[445,614,1191,1218]" pageId="45" pageNumber="46" value="1981-06-16">16 Jun. 1981</date>
) is morphologically inseparable from Levantine specimens of
<taxonomicName id="4C024D37FFB95104D8F0DDD8FED2FB52" authorityName="Friese" authorityYear="1899" box="[189,354,1228,1254]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="unifasciata">
<emphasis id="B976EAA6FFB95104D8F0DDD8FED2FB52" box="[189,354,1228,1254]" italics="true" pageId="45" pageNumber="46">A. unifasciata</emphasis>
</taxonomicName>
and the Iranian specimen of
<taxonomicName id="4C024D37FFB95104DA8BDDD8FCF6FB52" authority="Wood &amp; Monfared, 2022" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[710,838,1228,1254]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="bulbosa" status="sp. nov.">
<emphasis id="B976EAA6FFB95104DA8BDDD8FCF6FB52" box="[710,838,1228,1254]" italics="true" pageId="45" pageNumber="46">A. bulbosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557DDFFB95104DB00DDD8FC1AFB52" box="[845,938,1228,1254]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
Males must be found that can resolve the identity of specimens from south-eastern
<collectingCountry id="F3157624FFB95104DA82DDE4FC94FABE" box="[719,804,1264,1290]" name="Turkey" pageId="45" pageNumber="46">Turkey</collectingCountry>
, which has biogeographical affinities with both the Levant and
<collectingCountry id="F3157624FFB95104D901DC01FECFFA9B" box="[332,383,1301,1327]" name="Iran" pageId="45" pageNumber="46">Iran</collectingCountry>
. Molecular data (G. Pisanty pers. comm.) suggests that there may be cryptic species within this group that require clarification and delineation. Based on morphological knowledge, both
<taxonomicName id="4C024D37FFB95104D8F0DC49FEFBFAC2" authorityName="Friese" authorityYear="1899" box="[189,331,1373,1399]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="unicincta">
<emphasis id="B976EAA6FFB95104D8F0DC49FEFBFAC2" box="[189,331,1373,1399]" italics="true" pageId="45" pageNumber="46">A. unicincta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C024D37FFB95104D9C8DC49FE4FFAC2" authorityName="Morawitz" authorityYear="1886" box="[389,511,1373,1399]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="caspica">
<emphasis id="B976EAA6FFB95104D9C8DC49FE4FFAC2" box="[389,511,1373,1399]" italics="true" pageId="45" pageNumber="46">A. caspica</emphasis>
</taxonomicName>
range from the Levant to
<collectingCountry id="F3157624FFB95104DB7EDC49FCD5FAC3" box="[819,869,1373,1399]" name="Iran" pageId="45" pageNumber="46">Iran</collectingCountry>
, but male
<taxonomicName id="4C024D37FFB95104DB92DC49FB32FAC3" authorityName="Friese" authorityYear="1899" box="[991,1154,1373,1399]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="unifasciata">
<emphasis id="B976EAA6FFB95104DB92DC49FB32FAC3" box="[991,1154,1373,1399]" italics="true" pageId="45" pageNumber="46">A. unifasciata</emphasis>
</taxonomicName>
are currently known only from the Levant. The nomenclatural and distributional situation may therefore change pending revision.
</paragraph>
</subSubSection>
<subSubSection id="C318653FFFB95104D8F0DCFCFD55F9E2" pageId="45" pageNumber="46" type="etymology">
<paragraph id="8BBD36B4FFB95104D8F0DCFCFEF4F9B6" blockId="45.[189,324,1512,1538]" box="[189,324,1512,1538]" pageId="45" pageNumber="46">
<heading id="D0F581D8FFB95104D8F0DCFCFEF4F9B6" bold="true" box="[189,324,1512,1538]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="B976EAA6FFB95104D8F0DCFCFEF4F9B6" bold="true" box="[189,324,1512,1538]" pageId="45" pageNumber="46">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FFB95104D8F0DF03FD55F9E2" blockId="45.[189,1398,1559,1622]" pageId="45" pageNumber="46">
From the Latin meaning bulbous in reference to the swollen genital capsule that separates this taxon from similar red-marked species of
<taxonomicName id="4C024D37FFB95104DA2DDF2FFD6EF9E1" authorityName="Warncke" authorityYear="1968" box="[608,734,1595,1621]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ulandrena">
<emphasis id="B976EAA6FFB95104DA2DDF2FFD6EF9E1" box="[608,734,1595,1621]" italics="true" pageId="45" pageNumber="46">Ulandrena</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C318653FFFB95104D8F0DF69FC1FF8E1" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="8BBD36B4FFB95104D8F0DF69FE1AF923" blockId="45.[189,426,1661,1687]" box="[189,426,1661,1687]" pageId="45" pageNumber="46">
<heading id="D0F581D8FFB95104D8F0DF69FE1AF923" bold="true" box="[189,426,1661,1687]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="B976EAA6FFB95104D8F0DF69FE1AF923" bold="true" box="[189,426,1661,1687]" pageId="45" pageNumber="46">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FFB95104D8A7DFB9FBC2F95A" blockId="45.[189,1138,1709,1774]" pageId="45" pageNumber="46">
<materialsCitation id="3B6A3CE9FFB95104D8A7DFB9FBC2F95A" ID-GBIF-Occurrence="3944043612" collectionCode="OOLM" collectorName="K. Warncke" country="Iran" elevation="1800" location="Sarvestan" pageId="45" pageNumber="46" specimenCount="1" specimenCount-male="1" stateProvince="Fars" typeStatus="holotype">
<emphasis id="B976EAA6FFB95104D8A7DFB9FEECF973" bold="true" box="[234,348,1709,1735]" pageId="45" pageNumber="46">
<typeStatus id="54B98816FFB95104D8A7DFB9FEECF973" box="[234,348,1709,1735]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3157624FFB95104D8F0DFC6FEB5F958" box="[189,261,1746,1772]" name="Iran" pageId="45" pageNumber="46">IRAN</collectingCountry>
<specimenCount id="9D04FD3DFFB95104D953DFC6FE87F958" box="[286,311,1746,1772]" pageId="45" pageNumber="46" type="male"></specimenCount>
;
<collectingRegion id="49C6F856FFB95104D90EDFC6FEC8F958" box="[323,376,1746,1772]" country="Iran" name="Fars" pageId="45" pageNumber="46">Fars</collectingRegion>
,
<quantity id="4CFA9B51FFB95104D9C9DFC5FE7EF95F" box="[388,462,1745,1772]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="45" pageNumber="46" unit="km" value="15.0">
<locationDeviation id="9A0F2F96FFB95104D9C9DFC5FE7EF95F" box="[388,462,1745,1772]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="45" pageNumber="46" unit="km" value="15.0">15 km</locationDeviation>
</quantity>
SE of
<location id="8EDD606FFFB95104DA52DFC6FD25F958" LSID="urn:lsid:plazi:treatment:03AB87A2FFB95119DA41D8E3FD92F85C:8EDD606FFFB95104DA52DFC6FD25F958" box="[543,661,1746,1772]" country="Iran" name="Sarvestan" pageId="45" pageNumber="46" stateProvince="Fars">Sarvestan</location>
;
<elevation id="002FD187FFB95104DAECDFC6FC89F95F" box="[673,825,1745,1772]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="45" pageNumber="46" unit="m" value="1800.0">
<quantity id="4CFA9B51FFB95104DAECDFC6FD4AF958" box="[673,762,1746,1772]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="45" pageNumber="46" unit="m" value="1800.0">1800 m</quantity>
a.s.l.
</elevation>
;
<collectorName id="26F75362FFB95104DB0ADFC6FC62F95F" box="[839,978,1745,1772]" pageId="45" pageNumber="46">K. Warncke</collectorName>
leg.;
<collectionCode id="ED13AE71FFB95104DC59DFC6FBDEF95A" box="[1044,1134,1746,1774]" collectionName="OOLM" pageId="45" pageNumber="46">OÖLM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BBD36B4FFB95104D8A7DE00FC1FF8E1" blockId="45.[189,943,1812,1877]" pageId="45" pageNumber="46">
<materialsCitation id="3B6A3CE9FFB95104D8A7DE00FC1BF8E1" ID-GBIF-Occurrence="3944043693" collectionCode="OOLM" collectorName="K. Warncke" country="Iran" elevation="1800" location="Fars" pageId="45" pageNumber="46" specimenCount="1" specimenCount-female="1" stateProvince="Fars" typeStatus="paratype">
<emphasis id="B976EAA6FFB95104D8A7DE00FEECF89A" bold="true" box="[234,348,1812,1838]" pageId="45" pageNumber="46">
<typeStatus id="54B98816FFB95104D8A7DE00FEECF89A" box="[234,348,1812,1838]" pageId="45" pageNumber="46" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="F3157624FFB95104D8F0DE2CFEB5F8E6" box="[189,261,1848,1874]" name="Iran" pageId="45" pageNumber="46">IRAN</collectingCountry>
<specimenCount id="9D04FD3DFFB95104D953DE2CFEFBF8E7" box="[286,331,1848,1875]" pageId="45" pageNumber="46" type="female">1 ♀</specimenCount>
;
<collectingRegion id="49C6F856FFB95104D914DE2DFE3EF8E7" box="[345,398,1849,1875]" country="Iran" name="Fars" pageId="45" pageNumber="46">Fars</collectingRegion>
, same collection data as for holotype;
<collectionCode id="ED13AE71FFB95104DB1CDE2DFC1BF8E1" box="[849,939,1849,1877]" collectionName="OOLM" pageId="45" pageNumber="46">OÖLM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C318653FFFB95119D8F0DE6EFE8BF8C0" lastPageId="48" lastPageNumber="49" pageId="45" pageNumber="46" type="description">
<paragraph id="8BBD36B4FFB95104D8F0DE6EFEFEF820" blockId="45.[189,334,1914,1940]" box="[189,334,1914,1940]" pageId="45" pageNumber="46">
<heading id="D0F581D8FFB95104D8F0DE6EFEFEF820" bold="true" box="[189,334,1914,1940]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="B976EAA6FFB95104D8F0DE6EFEFEF820" bold="true" box="[189,334,1914,1940]" pageId="45" pageNumber="46">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FFB95104D8A7DEBEFEF5F870" blockId="45.[189,745,1962,2025]" box="[234,325,1962,1988]" pageId="45" pageNumber="46">
<emphasis id="B976EAA6FFB95104D8A7DEBEFEF5F870" bold="true" box="[234,325,1962,1988]" pageId="45" pageNumber="46">Female</emphasis>
</paragraph>
<paragraph id="8BBD36B4FFB95104D8F0DEDBFD59F85C" blockId="45.[189,745,1962,2025]" box="[189,745,1998,2025]" pageId="45" pageNumber="46">
MEASUREMENTS. Body length
<quantity id="4CFA9B51FFB95104DA5EDEDAFDD5F85D" box="[531,613,1998,2025]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="45" pageNumber="46" unit="mm" value="10.0">10 mm</quantity>
(
<figureCitation id="13392A31FFB95104DA38DEDAFD6CF85C" box="[629,732,1998,2024]" captionStart="Figs 109114" captionStartId="46.[191,243,1965,1991]" captionTargetBox="[189,1398,557,1918]" captionTargetId="figure-119@46.[189,1398,557,1918]" captionTargetPageId="46" captionText="Figs 109114. Andrena bulbosa Wood sp. nov., ♀ (OÖLM). 109. Profile. 110. Face. 111. Process of labrum, detail. 112. Scutum. 113. Terga. 114. Terga 45, detail." figureDoi="http://doi.org/10.5281/zenodo.7222388" httpUri="https://zenodo.org/record/7222388/files/figure.png" pageId="45" pageNumber="46">Fig. 109</figureCitation>
).
</paragraph>
<paragraph id="8BBD36B4FFBA5106D8F0D81DFAC6FEF2" blockId="46.[189,1399,265,467]" lastBlockId="47.[189,1398,265,327]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
HEAD. 1.3 times as wide as long. Clypeus dark, weakly domed, regularly punctate with exception of narrow impunctate longitudinal midline, punctures otherwise separated by 0.51 puncture diameter; underlying surface faintly shagreened, shining (
<figureCitation id="13392A31FFBA5107DAA6D85BFCE1FEDD" box="[747,849,335,361]" captionStart="Figs 109114" captionStartId="46.[191,243,1965,1991]" captionTargetBox="[189,1398,557,1918]" captionTargetId="figure-119@46.[189,1398,557,1918]" captionTargetPageId="46" captionText="Figs 109114. Andrena bulbosa Wood sp. nov., ♀ (OÖLM). 109. Profile. 110. Face. 111. Process of labrum, detail. 112. Scutum. 113. Terga. 114. Terga 45, detail." figureDoi="http://doi.org/10.5281/zenodo.7222388" httpUri="https://zenodo.org/record/7222388/files/figure.png" pageId="46" pageNumber="47">Fig. 110</figureCitation>
). Process of labrum broadly trapezoidal, apical margin with hint of emargination medially, essentially straight (
<figureCitation id="13392A31FFBA5107DBEAD866FBBBFE39" box="[935,1035,370,397]" captionStart="Figs 109114" captionStartId="46.[191,243,1965,1991]" captionTargetBox="[189,1398,557,1918]" captionTargetId="figure-119@46.[189,1398,557,1918]" captionTargetPageId="46" captionText="Figs 109114. Andrena bulbosa Wood sp. nov., ♀ (OÖLM). 109. Profile. 110. Face. 111. Process of labrum, detail. 112. Scutum. 113. Terga. 114. Terga 45, detail." figureDoi="http://doi.org/10.5281/zenodo.7222388" httpUri="https://zenodo.org/record/7222388/files/figure.png" pageId="46" pageNumber="47">Fig. 111</figureCitation>
). Face, gena, vertex, and scape with whitish hairs. Gena slightly exceeding width of compound eye; ocelloccipital distance subequal to diameter of lateral ocellus. Foveae dorsally occupying ½ distance between compound eye and lateral ocellus, slightly narrowed below at level of antennal insertions; foveae filled with whitish hairs.Antennae dark, A(4)512 slightly lightened dark orange below; A3 slightly exceeding A4+5, shorter than A4+5+6.
</paragraph>
<caption id="DF7D663CFFBA5107D8F2DEB9FC2EF85E" ID-DOI="http://doi.org/10.5281/zenodo.7222388" ID-Zenodo-Dep="7222388" httpUri="https://zenodo.org/record/7222388/files/figure.png" pageId="46" pageNumber="47" startId="46.[191,243,1965,1991]" targetBox="[189,1398,557,1918]" targetPageId="46">
<paragraph id="8BBD36B4FFBA5107D8F2DEB9FC2EF85E" blockId="46.[189,1398,1965,2027]" pageId="46" pageNumber="47">
<emphasis id="B976EAA6FFBA5107D8F2DEB9FED9F873" bold="true" box="[191,361,1965,1991]" pageId="46" pageNumber="47">Figs 109114.</emphasis>
<taxonomicName id="4C024D37FFBA5107D93FDEB9FD3BF873" authority="Wood" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[370,651,1965,1991]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="bulbosa" status="sp. nov.">
<emphasis id="B976EAA6FFBA5107D93FDEB9FD8DF873" box="[370,573,1965,1991]" italics="true" pageId="46" pageNumber="47">Andrena bulbosa</emphasis>
Wood
</taxonomicName>
<taxonomicNameLabel id="A24557DDFFBA5107DAD8DEBAFD45F87C" box="[661,757,1966,1992]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
, ♀ (OÖLM).
<emphasis id="B976EAA6FFBA5107DBD2DEB9FC7BF873" bold="true" box="[927,971,1965,1991]" pageId="46" pageNumber="47">109</emphasis>
. Profile.
<emphasis id="B976EAA6FFBA5107DC71DEB9FBD6F873" bold="true" box="[1084,1126,1965,1991]" pageId="46" pageNumber="47">110</emphasis>
. Face.
<emphasis id="B976EAA6FFBA5107DC8DDEB9FB59F873" bold="true" box="[1216,1257,1965,1991]" pageId="46" pageNumber="47">111</emphasis>
. Process of labrum, detail.
<emphasis id="B976EAA6FFBA5107D922DEC4FE29F85E" bold="true" box="[367,409,2000,2026]" pageId="46" pageNumber="47">112</emphasis>
. Scutum.
<emphasis id="B976EAA6FFBA5107DA5DDEC4FD8AF85E" bold="true" box="[528,570,2000,2026]" pageId="46" pageNumber="47">113</emphasis>
. Terga.
<emphasis id="B976EAA6FFBA5107DAD5DEC4FD72F85E" bold="true" box="[664,706,2000,2026]" pageId="46" pageNumber="47">114</emphasis>
. Terga 45, detail.
</paragraph>
</caption>
<paragraph id="8BBD36B4FFBB5119D8F0D87AFE27FD9E" blockId="47.[186,1399,366,498]" lastBlockId="48.[188,1399,265,554]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
MESOSOMA. Scutum and scutellum with dense and even punctation, punctures almost contiguous to separated by 0.5 puncture diameter; underlying surface generally smooth and shining, scutum shagreened anteriorly (
<figureCitation id="13392A31FFBB5106D974D8A0FE2CFE7A" box="[313,412,436,462]" captionStart="Figs 109114" captionStartId="46.[191,243,1965,1991]" captionTargetBox="[189,1398,557,1918]" captionTargetId="figure-119@46.[189,1398,557,1918]" captionTargetPageId="46" captionText="Figs 109114. Andrena bulbosa Wood sp. nov., ♀ (OÖLM). 109. Profile. 110. Face. 111. Process of labrum, detail. 112. Scutum. 113. Terga. 114. Terga 45, detail." figureDoi="http://doi.org/10.5281/zenodo.7222388" httpUri="https://zenodo.org/record/7222388/files/figure.png" pageId="47" pageNumber="48">Fig. 112</figureCitation>
). Pronotum without humeral angle. Mesepisternum and lateral face of propodeum with fine shagreen, weakly shining, mesepisternum with fine raised pattern of reticulation. Propodeal triangle weakly delineated laterally by fine carinae, internal surface with weak pattern of raised rugae basally, disappearing centrally, here with reticulation, dull. Propodeal triangle therefore weakly contrasting with dorsolateral faces of propodeum, these covered in fine but dense raised reticulation, dull. Mesosoma with whitish hairs, longest on mesepisternum; propodeal corbicula dorsally composed of white plumose hairs. Legs dark, apical tarsal segments lightened orange; pubescence white, tibial and femoral scopae and flocculus unicolourous white. Hind tarsal claws with strong inner tooth. Wings hyaline, stigma and venation dark orange, nervulus interstitial, first recurrent vein enters second submarginal cell slightly beyond its middle.
</paragraph>
<caption id="DF7D663CFFBB5106D8F0DEBDFB35F852" ID-DOI="http://doi.org/10.5281/zenodo.7222392" ID-Zenodo-Dep="7222392" httpUri="https://zenodo.org/record/7222392/files/figure.png" pageId="47" pageNumber="48" startId="47.[189,241,1961,1987]" targetBox="[189,1398,558,1919]" targetPageId="47">
<paragraph id="8BBD36B4FFBB5106D8F0DEBDFB35F852" blockId="47.[189,1399,1961,2023]" pageId="47" pageNumber="48">
<emphasis id="B976EAA6FFBB5106D8F0DEBDFEDBF877" bold="true" box="[189,363,1961,1987]" pageId="47" pageNumber="48">Figs 115120.</emphasis>
<taxonomicName id="4C024D37FFBB5106D934DEBDFD2AF877" authority="Wood" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[377,666,1961,1987]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bulbosa" status="sp. nov.">
<emphasis id="B976EAA6FFBB5106D934DEBDFDF8F877" box="[377,584,1961,1987]" italics="true" pageId="47" pageNumber="48">Andrena bulbosa</emphasis>
Wood
</taxonomicName>
<taxonomicNameLabel id="A24557DDFFBB5106DAE5DEBEFCBBF870" box="[680,779,1962,1988]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ (OÖLM).
<emphasis id="B976EAA6FFBB5106DB8DDEBDFC5AF877" bold="true" box="[960,1002,1961,1987]" pageId="47" pageNumber="48">115</emphasis>
. Profile.
<emphasis id="B976EAA6FFBB5106DC2EDEBDFB3DF877" bold="true" box="[1123,1165,1961,1987]" pageId="47" pageNumber="48">116</emphasis>
. Face.
<emphasis id="B976EAA6FFBB5106DCA3DEBDFAA8F877" bold="true" box="[1262,1304,1961,1987]" pageId="47" pageNumber="48">117</emphasis>
. Terga.
<emphasis id="B976EAA6FFBB5106D8F0DED8FF57F852" bold="true" box="[189,231,1996,2022]" pageId="47" pageNumber="48">118</emphasis>
. Genitalia dorsal view.
<emphasis id="B976EAA6FFBB5106D9B3DED8FD98F852" bold="true" box="[510,552,1996,2022]" pageId="47" pageNumber="48">119</emphasis>
. Genitalia lateral view.
<emphasis id="B976EAA6FFBB5106DB0DDED8FCDCF852" bold="true" box="[832,876,1996,2022]" pageId="47" pageNumber="48">120</emphasis>
. Genitalia ventral view.
</paragraph>
</caption>
<paragraph id="8BBD36B4FFA45119D8F0DB40FD64FCFB" blockId="48.[189,1400,596,847]" pageId="48" pageNumber="49">
METASOMA. Terga dark with extensive red maculations; red maculations
<taxonomicName id="4C024D37FFA45119DC4ADB40FB2FFDDA" box="[1031,1183,596,622]" form="patches" pageId="48" pageNumber="49" rank="form">form patches</taxonomicName>
on T1 basally and apically, T23 almost entirely red-marked, T4 basally and apically; tergal margins lightened yellowhyaline (
<figureCitation id="13392A31FFA45119D96BDB8BFE3EFD0D" box="[294,398,671,697]" captionStart="Figs 109114" captionStartId="46.[191,243,1965,1991]" captionTargetBox="[189,1398,557,1918]" captionTargetId="figure-119@46.[189,1398,557,1918]" captionTargetPageId="46" captionText="Figs 109114. Andrena bulbosa Wood sp. nov., ♀ (OÖLM). 109. Profile. 110. Face. 111. Process of labrum, detail. 112. Scutum. 113. Terga. 114. Terga 45, detail." figureDoi="http://doi.org/10.5281/zenodo.7222388" httpUri="https://zenodo.org/record/7222388/files/figure.png" pageId="48" pageNumber="49">Fig. 113</figureCitation>
). Terga T14 weakly shagreened, shining; T5 dull; all terga densely and uniformly punctate, punctures separated by 1 puncture diameter (
<figureCitation id="13392A31FFA45119DB0CDBD0FC17FD6B" box="[833,935,708,735]" captionStart="Figs 109114" captionStartId="46.[191,243,1965,1991]" captionTargetBox="[189,1398,557,1918]" captionTargetId="figure-119@46.[189,1398,557,1918]" captionTargetPageId="46" captionText="Figs 109114. Andrena bulbosa Wood sp. nov., ♀ (OÖLM). 109. Profile. 110. Face. 111. Process of labrum, detail. 112. Scutum. 113. Terga. 114. Terga 45, detail." figureDoi="http://doi.org/10.5281/zenodo.7222388" httpUri="https://zenodo.org/record/7222388/files/figure.png" pageId="48" pageNumber="49">Fig. 114</figureCitation>
). Tergal discs with very fine, short, and sparse whitish hairs, T24 with weak white apical hair fringes that do not obscure underlying surface, widely interrupted. Apical fringe of T5 and hairs flanking pygidial plate white to golden; pygidial plate rounded triangular, without surface sculpture.
</paragraph>
<paragraph id="8BBD36B4FFA45119D8A7DA6DFE9AFC27" blockId="48.[189,744,889,953]" box="[234,298,889,915]" pageId="48" pageNumber="49">
<emphasis id="B976EAA6FFA45119D8A7DA6DFE9AFC27" bold="true" box="[234,298,889,915]" pageId="48" pageNumber="49">Male</emphasis>
</paragraph>
<paragraph id="8BBD36B4FFA45119D8F0DA8BFD58FC0D" blockId="48.[189,744,889,953]" box="[189,744,926,953]" pageId="48" pageNumber="49">
MEASUREMENTS. Body length
<quantity id="4CFA9B51FFA45119DA5EDA8BFDD5FC0D" box="[531,613,927,953]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="48" pageNumber="49" unit="mm" value="12.0">12 mm</quantity>
(
<figureCitation id="13392A31FFA45119DA38DA8BFD6BFC0D" box="[629,731,926,953]" captionStart="Figs 115120" captionStartId="47.[189,241,1961,1987]" captionTargetBox="[189,1398,558,1919]" captionTargetId="figure-123@47.[189,1398,558,1919]" captionTargetPageId="47" captionText="Figs 115120. Andrena bulbosa Wood sp. nov., ♂ (OÖLM). 115. Profile. 116. Face. 117. Terga. 118. Genitalia dorsal view. 119. Genitalia lateral view. 120. Genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.7222392" httpUri="https://zenodo.org/record/7222392/files/figure.png" pageId="48" pageNumber="49">Fig. 115</figureCitation>
).
</paragraph>
<paragraph id="8BBD36B4FFA45119D8F0DAF7FE98FB6B" blockId="48.[187,1401,994,1247]" pageId="48" pageNumber="49">
HEAD. 1.2 times as wide as long. Clypeus domed, entirely yellow-marked, yellow colouration extending onto lower paraocular areas; clypeus evenly punctate, punctures separated by 1 puncture diameter; underlying surface weakly shining (
<figureCitation id="13392A31FFA45119DA15DD3AFD0CFBF3" box="[600,700,1069,1096]" captionStart="Figs 115120" captionStartId="47.[189,241,1961,1987]" captionTargetBox="[189,1398,558,1919]" captionTargetId="figure-123@47.[189,1398,558,1919]" captionTargetPageId="47" captionText="Figs 115120. Andrena bulbosa Wood sp. nov., ♂ (OÖLM). 115. Profile. 116. Face. 117. Terga. 118. Genitalia dorsal view. 119. Genitalia lateral view. 120. Genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.7222392" httpUri="https://zenodo.org/record/7222392/files/figure.png" pageId="48" pageNumber="49">Fig. 116</figureCitation>
). Process of labrum rounded rectangular, broadly emarginate medially. Lower face and gena with white hairs, becoming golden-brown on vertex, scape, and frons. Gena exceeding width of compound eye; ocelloccipital distance 1½ times diameter of lateral ocellus. Antennae dark, A413 slightly lightened dark brown below; A3 slightly exceeding A4+5, shorter than A4+5+6.
</paragraph>
<paragraph id="8BBD36B4FFA45119D8F0DC1CFD08FA0C" blockId="48.[189,1399,1288,1464]" pageId="48" pageNumber="49">MESOSOMA. Mesosoma structurally as in female. Mesepisternum and propodeum with moderately thick long whitish hairs, becoming golden-brown dorsally on scutum and scutellum. Legs dark, apical tarsal segments lightened orange; pubescence whitish to brownish. Hind tarsal claws with inner tooth. Wings hyaline, stigma and venation orange; nervulus very slightly antefurcal, first recurrent vein enters second submarginal cell clearly beyond its middle.</paragraph>
<paragraph id="8BBD36B4FFA45119D8F0DCF6FE8BF8C0" blockId="48.[186,1399,1506,1908]" pageId="48" pageNumber="49">
METASOMA. Terga dark, red maculations cover entirety of T23, apical margins of T1 and T4; tergal margins otherwise lightened yellow-hyaline (
<figureCitation id="13392A31FFA45119DA8BDF13FC9AF996" box="[710,810,1543,1570]" captionStart="Figs 115120" captionStartId="47.[189,241,1961,1987]" captionTargetBox="[189,1398,558,1919]" captionTargetId="figure-123@47.[189,1398,558,1919]" captionTargetPageId="47" captionText="Figs 115120. Andrena bulbosa Wood sp. nov., ♂ (OÖLM). 115. Profile. 116. Face. 117. Terga. 118. Genitalia dorsal view. 119. Genitalia lateral view. 120. Genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.7222392" httpUri="https://zenodo.org/record/7222392/files/figure.png" pageId="48" pageNumber="49">Fig. 117</figureCitation>
). Tergal discs weakly shagreened, strongly shining; terga densely and evenly punctate, punctures separated by 11.5 puncture diameters. Tergal discs with fine golden pubescence, hairs longest on T1, T24 with complete apical hairbands composed of golden hairs, weakest on T2, becoming progressively thicker to T4, here obscuring underlying surface. S8 short, triangular, ventral surface covered by short golden hairs that obscure underlying shape. Genital capsule large, gonocoxae with inner margin evenly rounded, lacking apical teeth (
<figureCitation id="13392A31FFA45119DCDCDFD7FB47F969" box="[1169,1271,1731,1757]" captionStart="Figs 115120" captionStartId="47.[189,241,1961,1987]" captionTargetBox="[189,1398,558,1919]" captionTargetId="figure-123@47.[189,1398,558,1919]" captionTargetPageId="47" captionText="Figs 115120. Andrena bulbosa Wood sp. nov., ♂ (OÖLM). 115. Profile. 116. Face. 117. Terga. 118. Genitalia dorsal view. 119. Genitalia lateral view. 120. Genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.7222392" httpUri="https://zenodo.org/record/7222392/files/figure.png" pageId="48" pageNumber="49">Fig. 118</figureCitation>
); in lateral view gonocoxae forming rounded 90° angle (
<figureCitation id="13392A31FFA45119DA95DFFDFC8DF8B7" box="[728,829,1769,1795]" captionStart="Figs 115120" captionStartId="47.[189,241,1961,1987]" captionTargetBox="[189,1398,558,1919]" captionTargetId="figure-123@47.[189,1398,558,1919]" captionTargetPageId="47" captionText="Figs 115120. Andrena bulbosa Wood sp. nov., ♂ (OÖLM). 115. Profile. 116. Face. 117. Terga. 118. Genitalia dorsal view. 119. Genitalia lateral view. 120. Genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.7222392" httpUri="https://zenodo.org/record/7222392/files/figure.png" pageId="48" pageNumber="49">Fig. 119</figureCitation>
). Gonostyli with apical blades flattened, sharply right-angled basally, and produced into ventrally projecting points. Penis valves mediolaterally with short hyaline extensions, apically produced into long narrow projection, very subtly bifurcate apically (
<figureCitation id="13392A31FFA45119D88ADE4EFE9EF8C0" box="[199,302,1882,1908]" captionStart="Figs 115120" captionStartId="47.[189,241,1961,1987]" captionTargetBox="[189,1398,558,1919]" captionTargetId="figure-123@47.[189,1398,558,1919]" captionTargetPageId="47" captionText="Figs 115120. Andrena bulbosa Wood sp. nov., ♂ (OÖLM). 115. Profile. 116. Face. 117. Terga. 118. Genitalia dorsal view. 119. Genitalia lateral view. 120. Genitalia ventral view." figureDoi="http://doi.org/10.5281/zenodo.7222392" httpUri="https://zenodo.org/record/7222392/files/figure.png" pageId="48" pageNumber="49">Fig. 120</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C318653FFFA45119D8F0DE89FD92F85C" box="[189,546,1949,2025]" pageId="48" pageNumber="49" type="distribution">
<paragraph id="8BBD36B4FFA45119D8F0DE89FEE6F803" blockId="48.[189,342,1949,1975]" box="[189,342,1949,1975]" pageId="48" pageNumber="49">
<heading id="D0F581D8FFA45119D8F0DE89FEE6F803" bold="true" box="[189,342,1949,1975]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="B976EAA6FFA45119D8F0DE89FEE6F803" bold="true" box="[189,342,1949,1975]" pageId="48" pageNumber="49">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FFA45119D8F0DEDAFD92F85C" blockId="48.[189,546,1998,2025]" box="[189,546,1998,2025]" pageId="48" pageNumber="49">
Southern
<collectingCountry id="F3157624FFA45119D963DEDBFEEDF85D" box="[302,349,1999,2025]" name="Iran" pageId="48" pageNumber="49">Iran</collectingCountry>
(
<collectingRegion id="49C6F856FFA45119D920DEDAFDA6F85C" box="[365,534,1998,2024]" country="Iran" name="Fars" pageId="48" pageNumber="49">Fars Province</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>