treatments-xml/data/03/E4/43/03E4436DFFD9FFECFF0CFC69FD514F8C.xml
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<document id="406DE877EF45D3545E82DEEEC454C5E5" ID-CLB-Dataset="280865" ID-DOI="10.11646/zootaxa.5382.1.8" ID-GBIF-Dataset="7b2fa3e0-92a3-4862-9ff0-a70a838a4978" ID-ISSN="1175-5326" ID-Zenodo-Dep="10279897" ID-ZooBank="8DEA006E-732F-4ECF-97A5-FC9468E3FF9D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701938063791" checkinUser="plazi" docAuthor="Yin, Jia-Dong, Bu, Wen-Jun &amp; Xie, Qiang" docDate="2023" docId="03E4436DFFD9FFECFF0CFC69FD514F8C" docLanguage="en" docName="zootaxa.5382.1.8.pdf" docOrigin="Zootaxa 5382 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5382.1.8/52424" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heissophila macrotheleae Schuh 2006" docType="treatment" docVersion="2" lastPageNumber="56" masterDocId="FFDD3B15FFD8FFEDFF9BFFA4FFD04823" masterDocTitle="Heissophila macrotheleae Schuh, 2006, the first record of the subfamily Heissophilinae from China (Hemiptera: Heteroptera: Plokiophilidae)" masterLastPageNumber="61" masterPageNumber="55" pageNumber="56" updateTime="1701938722411" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="355AE68667FBA313D27302E129A23BCB">Heissophila macrotheleae Schuh, 2006, the first record of the subfamily Heissophilinae from China (Hemiptera: Heteroptera: Plokiophilidae)</mods:title>
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<mods:namePart id="9F6ECC85834EEDBBFA07E4C487684A8D">Yin, Jia-Dong</mods:namePart>
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<mods:affiliation id="6A164CA3526123F94FC955E52AF3779C">School of Life Sciences, State Key Laboratory of Biocontrol, 135 Xingangxi Road, Sun Yat-sen University, Guangzhou 510275, Guangdong, China. &amp; yinjd 3 @ mail 2. sysu. edu. cn; https: // orcid. org / 0000 - 0002 - 5756 - 7011</mods:affiliation>
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<mods:namePart id="8F86567AA275AEFB1FB695ED98D90A81">Bu, Wen-Jun</mods:namePart>
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<mods:namePart id="CAFC829A0ECF4282B9C12D0DCF267A73">Xie, Qiang</mods:namePart>
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<mods:affiliation id="FF6E923C8836A403CF54DC5E018154F5">School of Life Sciences, State Key Laboratory of Biocontrol, 135 Xingangxi Road, Sun Yat-sen University, Guangzhou 510275, Guangdong, China. &amp; xieq 8 @ mail. sysu. edu. cn; https: // orcid. org / 0000 - 0001 - 6376 - 8808</mods:affiliation>
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<paragraph id="8BF2F27BFFD9FFECFF0CFC69FDA04BCB" blockId="1.[151,624,973,1000]" box="[151,624,973,1000]" pageId="1" pageNumber="56">
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<taxonomicName id="4C4D89F8FFD9FFECFF0CFC69FDA04BCB" authority="Schuh, 2006" authorityName="Schuh" authorityYear="2006" box="[151,624,973,1000]" class="Insecta" family="Plokiophilidae" genus="Heissophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="macrotheleae">
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<emphasis id="B9392E69FFD9FFECFF0CFC69FE014BCB" bold="true" box="[151,465,973,1000]" italics="true" pageId="1" pageNumber="56">Heissophila macrotheleae</emphasis>
<bibRefCitation id="EFDC8F8AFFD9FFECFE43FC69FDA04BCB" author="Schuh, R. T." box="[472,624,973,1000]" pageId="1" pageNumber="56" pagination="637 - 645" refId="ref2879" refString="Schuh, R. T. (2006) Heissophila macrotheleae, a new genus and new species of Plokiophilidae from Thailand (Hemiptera, Heteroptera), with comments on the family diagnosis. Denisia, 19, 637 - 645." type="journal article" year="2006">Schuh, 2006</bibRefCitation>
</emphasis>
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<treatmentCitation id="0AECD46AFFD9FFECFF0CFBB6FD8B4C09" author="Schuh, R. T." box="[151,603,1042,1066]" page="644" pageId="1" pageNumber="56" year="2006">
<taxonomicName id="4C4D89F8FFD9FFECFF0CFBB6FD8B4C09" authority="Schuh, 2006: 644" authorityName="Schuh" authorityPageNumber="644" authorityYear="2006" box="[151,603,1042,1066]" class="Insecta" family="Plokiophilidae" genus="Heissophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="macrotheleae">
<emphasis id="B9392E69FFD9FFECFF0CFBB6FE704C09" box="[151,416,1042,1066]" italics="true" pageId="1" pageNumber="56">Heissophila macrotheleae</emphasis>
<materialsCitation id="3B25F826FFD9FFECFE3DFBB6FD8B4C09" box="[422,603,1042,1066]" country="Thailand" location="Chang Rai Province" pageId="1" pageNumber="56" specimenCount="1">
<bibRefCitation id="EFDC8F8AFFD9FFECFE3DFBB6FD8B4C09" author="Schuh, R. T." box="[422,603,1042,1066]" pageId="1" pageNumber="56" pagination="637 - 645" refId="ref2879" refString="Schuh, R. T. (2006) Heissophila macrotheleae, a new genus and new species of Plokiophilidae from Thailand (Hemiptera, Heteroptera), with comments on the family diagnosis. Denisia, 19, 637 - 645." type="journal article" year="2006">Schuh, 2006: 644</bibRefCitation>
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.
<typeStatus id="54F64CD9FFD9FFECFDFFFBB7FD464C08" box="[612,662,1043,1067]" pageId="1" pageNumber="56">Type</typeStatus>
locality:
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(
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).
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</paragraph>
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<emphasis id="B9392E69FFD9FFECFF0CFBFCFE954C51" bold="true" box="[151,325,1112,1138]" pageId="1" pageNumber="56">Redescription.</emphasis>
The Chinese samples conform mostly in all diagnostic features to the species description by
<bibRefCitation id="EFDC8F8AFFD9FFECFAD4FBFCFF1F4CB5" author="Schuh, R. T." pageId="1" pageNumber="56" pagination="637 - 645" refId="ref2879" refString="Schuh, R. T. (2006) Heissophila macrotheleae, a new genus and new species of Plokiophilidae from Thailand (Hemiptera, Heteroptera), with comments on the family diagnosis. Denisia, 19, 637 - 645." type="journal article" year="2006">Schuh, 2006</bibRefCitation>
from
<collectingCountry id="F35AB2EBFFD9FFECFE88FBD8FEA94CB5" box="[275,377,1148,1174]" name="Thailand" pageId="1" pageNumber="56">Thailand</collectingCountry>
, except for the following new supplement.
</paragraph>
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Body length (
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). Ground colour and head dark brown, ventral surface of head paler. Antennal segment I dark reddish-brown; segment II light brown, distal pale yellow; segment III yellow, distal 1/3 pale brown; segment IV yellow, distal 1/4 pale yellow. Pronotum, propleura and scutellum brown. Membrane of forewing light brown, with four conspicuous longitudinal veins; area around costal fracture whitish. Coxae dark reddish-brown; trochanters basal pale yellow, distal reddish-brown; femora dark reddish-brown and distal part paler; tibiae basal 1/2 brown and distal 1/2 part paler; tarsi paler (
<figureCitation id="1376EEFEFFD9FFECFDE0FAF0FD294D4C" box="[635,761,1364,1391]" captionStart="FIGURE 1" captionStartId="2.[151,250,1730,1754]" captionTargetBox="[152,1435,179,1703]" captionTargetId="figure-24@2.[151,1435,179,1704]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Heissophila macrotheleae. A. male in dorsal view; B. male in ventral view; C. female in dorsal view; D. female in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10279899" httpUri="https://zenodo.org/record/10279899/files/figure.png" pageId="1" pageNumber="56">Fig. 1AD</figureCitation>
).
</paragraph>
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<emphasis id="B9392E69FFD9FFECFF5CFADCFE604DB1" bold="true" box="[199,432,1400,1426]" pageId="1" pageNumber="56">Material examined.</emphasis>
<materialsCitation id="3B25F826FFD9FFECFE2CFADCFB1D4D94" collectingDate="2021-05-15" collectionCode="XTBG" collectorName="Qiang Xie &amp; Yu Men" country="China" county="Mengla County" elevation="924" latitude="21.978031" location="Menglun Town" longLatPrecision="1" longitude="101.17531" municipality="Xishuangbanna" pageId="1" pageNumber="56" specimenCount="10" specimenCount-female="5" specimenCount-male="5" stateProvince="Yunnan">
<specimenCount id="9D4B39F2FFD9FFECFE2CFADCFE204DB0" box="[439,496,1400,1427]" count="5" pageId="1" pageNumber="56" type="male">5♂♂</specimenCount>
,
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,
<collectingCountry id="F35AB2EBFFD9FFECFDD9FADDFD4E4DB0" box="[578,670,1401,1427]" name="China" pageId="1" pageNumber="56">CHINA</collectingCountry>
,
<collectingRegion id="49893C99FFD9FFECFD3CFADDFCB94DB1" box="[679,873,1400,1427]" country="China" name="Yunnan" pageId="1" pageNumber="56">Yunnan Province</collectingRegion>
,
<collectingMunicipality id="6B966801FFD9FFECFCEFFADCFBF54DB1" box="[884,1061,1400,1426]" pageId="1" pageNumber="56">Xishuangbanna</collectingMunicipality>
,
<collectingCounty id="62938AF7FFD9FFECFBABFADCFB304DB0" box="[1072,1248,1400,1427]" pageId="1" pageNumber="56">Mengla County</collectingCounty>
,
<location id="8E92A4A0FFD9FFECFB70FADCFA484DB0" LSID="urn:lsid:plazi:treatment:03E4436DFFD9FFECFF0CFC69FD514F8C:8E92A4A0FFD9FFECFB70FADCFA484DB0" box="[1259,1432,1400,1427]" country="China" county="Mengla County" latitude="21.978031" longLatPrecision="1" longitude="101.17531" municipality="Xishuangbanna" name="Menglun Town" pageId="1" pageNumber="56" stateProvince="Yunnan">Menglun Town</location>
,
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:
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,
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; ca.
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</quantity>
elev.; leg.
<collectorName id="26B897ADFFD9FFECFCCEFA38FC184D95" box="[853,968,1436,1462]" pageId="1" pageNumber="56">Qiang Xie</collectorName>
&amp;
<collectorName id="26B897ADFFD9FFECFC71FA39FB974D94" box="[1002,1095,1437,1463]" pageId="1" pageNumber="56">Yu Men</collectorName>
;
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<collectingDate id="EFB72D53FFD9FFECFBC9FA38FB1D4D94" box="[1106,1229,1436,1463]" pageId="1" pageNumber="56" value="2021-05-15">2021-V-15</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B25F826FFD9FFECFB42FA39FEED4DF9" collectingDate="2021-05-15" collectionCode="XTBG" collectorName="Hua-yan Chen" country="China" county="Mengla County" elevation="924" latitude="21.978031" location="Menglun Town" longLatPrecision="1" longitude="101.17531" municipality="Xishuangbanna" pageId="1" pageNumber="56" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D4B39F2FFD9FFECFB42FA39FB2D4D94" box="[1241,1277,1437,1463]" count="1" pageId="1" pageNumber="56" type="male">1♂</specimenCount>
,
<specimenCount id="9D4B39F2FFD9FFECFA92FA39FAFC4D94" box="[1289,1324,1437,1463]" count="1" pageId="1" pageNumber="56" type="female">1♀</specimenCount>
, ibid, leg.
<collectorName id="26B897ADFFD9FFECFF0CFA65FEED4DF9" box="[151,317,1472,1499]" pageId="1" pageNumber="56">Hua-yan Chen</collectorName>
</materialsCitation>
,
<date id="FFF3D4BBFFD9FFECFED3FA65FE014DF8" box="[328,465,1473,1499]" pageId="1" pageNumber="56" value="2021-04-14">2021-IV-14.</date>
</paragraph>
<paragraph id="8BF2F27BFFD9FFECFF5CFA41FC5F4E01" blockId="1.[151,1437,1112,1967]" pageId="1" pageNumber="56">
<emphasis id="B9392E69FFD9FFECFF5CFA41FEC14DDC" bold="true" box="[199,273,1509,1535]" pageId="1" pageNumber="56">Notes.</emphasis>
The Chinese samples do not differ significantly from the Thai samples, and all the morphological characters fall well within the original description by
<bibRefCitation id="EFDC8F8AFFD9FFECFD6BF9ACFC5A4E01" author="Schuh, R. T." box="[752,906,1544,1570]" pageId="1" pageNumber="56" pagination="637 - 645" refId="ref2879" refString="Schuh, R. T. (2006) Heissophila macrotheleae, a new genus and new species of Plokiophilidae from Thailand (Hemiptera, Heteroptera), with comments on the family diagnosis. Denisia, 19, 637 - 645." type="journal article" year="2006">Schuh (2006)</bibRefCitation>
.
</paragraph>
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<emphasis id="B9392E69FFD9FFECFF5CF988FEA04E64" bold="true" box="[199,368,1580,1607]" pageId="1" pageNumber="56">DNA barcode.</emphasis>
The
<emphasis id="B9392E69FFD9FFECFE33F989FE084E64" box="[424,472,1581,1607]" italics="true" pageId="1" pageNumber="56">COI</emphasis>
barcode of
<taxonomicName id="4C4D89F8FFD9FFECFDC0F989FCC34E65" authorityName="Schuh" authorityYear="2006" box="[603,787,1580,1606]" class="Insecta" family="Plokiophilidae" genus="Heissophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="macrotheleae">
<emphasis id="B9392E69FFD9FFECFDC0F989FCC34E65" box="[603,787,1580,1606]" italics="true" pageId="1" pageNumber="56">H. macrotheleae</emphasis>
</taxonomicName>
was deposited in GenBank (accession
<collectionCode id="ED5C6ABEFFD9FFECFB5EF989FB3F4E64" box="[1221,1263,1581,1607]" pageId="1" pageNumber="56">OQ</collectionCode>
197447), which was further searched against the
<collectionCode id="ED5C6ABEFFD9FFECFD9DF9F5FD994E48" box="[518,585,1617,1643]" pageId="1" pageNumber="56">NCBI</collectionCode>
Nucleotide database: the closest result was
<taxonomicName id="4C4D89F8FFD9FFECFBACF9F4FB0A4E49" authorityName="China" authorityYear="1953" box="[1079,1242,1616,1642]" class="Insecta" family="Plokiophilidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="family">Plokiophilidae</taxonomicName>
sp. (
<collectionCode id="ED5C6ABEFFD9FFECFA8BF9F4FAEE4E49" box="[1296,1342,1616,1642]" country="New Zealand" name="Otago Museum" pageId="1" pageNumber="56">OM</collectionCode>
323306) with an identity no lower than 98%. Moreover, the
<emphasis id="B9392E69FFD9FFECFD42F9D1FCD94EAC" box="[729,777,1653,1679]" italics="true" pageId="1" pageNumber="56">COI</emphasis>
barcode of the host spider was also deposited in GenBank (accession
<collectionCode id="ED5C6ABEFFD9FFECFE88F93CFEED4E91" box="[275,317,1688,1714]" pageId="1" pageNumber="56">OQ</collectionCode>
361630).
</paragraph>
</subSubSection>
<subSubSection id="C357A1F0FFD9FFECFF5CF918FD6F4F45" pageId="1" pageNumber="56" type="biology_ecology">
<paragraph id="8BF2F27BFFD9FFECFF5CF918FD6F4F45" blockId="1.[151,1437,1112,1967]" pageId="1" pageNumber="56">
<emphasis id="B9392E69FFD9FFECFF5CF918FEF74EF5" bold="true" box="[199,295,1724,1750]" pageId="1" pageNumber="56">Biology.</emphasis>
Habitat (
<figureCitation id="1376EEFEFFD9FFECFE0AF918FDDD4EF4" box="[401,525,1724,1751]" captionStart="FIGURE 2" captionStartId="3.[151,250,1275,1299]" captionTargetBox="[151,1435,181,1251]" captionTargetId="figure-20@3.[151,1435,181,1251]" captionTargetPageId="3" captionText="FIGURE 2. Habitus images of live individuals for Heissophila macrotheleae. A. an adult on the web in side view; B. an adult on the web in dorsal view; C. habitat of collected site. (Photographs were authorized by Dr. Hua-Yan Chen)" figureDoi="http://doi.org/10.5281/zenodo.10279901" httpUri="https://zenodo.org/record/10279901/files/figure.png" pageId="1" pageNumber="56">Fig. 2AC</figureCitation>
). In total, six to eight spider webs were overlapping each other in about 1 square meter on the afternoon of
<date id="FFF3D4BBFFD9FFECFE5BF944FDBD4ED8" box="[448,621,1760,1787]" pageId="1" pageNumber="56" value="2021-05-15">15th May 2021</date>
under the canopy shade of
<taxonomicName id="4C4D89F8FFD9FFECFC3CF945FBDF4ED8" box="[935,1039,1761,1787]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="1" pageNumber="56" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B9392E69FFD9FFECFC3CF945FC354ED9" box="[935,997,1761,1786]" italics="true" pageId="1" pageNumber="56">Ficus</emphasis>
sp.
</taxonomicName>
, of which only four webs had both spiders and bugs. There were about 15 individuals of them on each web. Based on our intuitive observations, the movement of
<taxonomicName id="4C4D89F8FFD9FFECFEFFF88DFDF04F61" authorityName="Schuh" authorityYear="2006" box="[356,544,1832,1858]" class="Insecta" family="Plokiophilidae" genus="Heissophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="macrotheleae">
<emphasis id="B9392E69FFD9FFECFEFFF88DFDF04F61" box="[356,544,1832,1858]" italics="true" pageId="1" pageNumber="56">H. macrotheleae</emphasis>
</taxonomicName>
appears to be less nimble than that of
<taxonomicName id="4C4D89F8FFD9FFECFC7CF88DFB534F61" authorityName="Luo, Peng &amp; Xie" authorityYear="2021" box="[999,1155,1832,1858]" class="Insecta" family="Plokiophilidae" genus="Plokiophiloides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="bannaensis">
<emphasis id="B9392E69FFD9FFECFC7CF88DFB534F61" box="[999,1155,1832,1858]" italics="true" pageId="1" pageNumber="56">P. bannaensis</emphasis>
</taxonomicName>
. We examined ten adult specimens, but nymphs and eggs were not found.
</paragraph>
</subSubSection>
<subSubSection id="C357A1F0FFD9FFECFF5CF8D4FD514F8C" pageId="1" pageNumber="56" type="distribution">
<paragraph id="8BF2F27BFFD9FFECFF5CF8D4FD514F8C" blockId="1.[151,1437,1112,1967]" pageId="1" pageNumber="56">
<emphasis id="B9392E69FFD9FFECFF5CF8D4FE894FA9" bold="true" box="[199,345,1904,1930]" pageId="1" pageNumber="56">Distribution</emphasis>
:
<collectingCountry id="F35AB2EBFFD9FFECFEF2F8D5FE104FA8" box="[361,448,1905,1931]" name="China" pageId="1" pageNumber="56">CHINA</collectingCountry>
(
<collectingRegion id="49893C99FFD9FFECFE54F8D5FD464FA9" box="[463,662,1904,1931]" country="China" name="Yunnan" pageId="1" pageNumber="56">Yunnan Province</collectingRegion>
),
<collectingCountry id="F35AB2EBFFD9FFECFD31F8D5FC914FA8" box="[682,833,1905,1931]" name="Indonesia" pageId="1" pageNumber="56">INDONESIA</collectingCountry>
(
<collectingRegion id="49893C99FFD9FFECFCCBF8D5FBC04FA9" box="[848,1040,1904,1931]" country="Indonesia" name="Kalimantan Timur" pageId="1" pageNumber="56">East Kalimantan</collectingRegion>
),
<collectingCountry id="F35AB2EBFFD9FFECFBB8F8D5FB644FA8" box="[1059,1204,1905,1931]" name="Thailand" pageId="1" pageNumber="56">THAILAND</collectingCountry>
(Chang Rai,
<collectingRegion id="49893C99FFD9FFECFAD0F8D4FF144F8D" country="Thailand" name="Chiang Mai" pageId="1" pageNumber="56">Chiang Mai</collectingRegion>
and
<collectingRegion id="49893C99FFD9FFECFF60F830FE714F8D" box="[251,417,1940,1966]" country="Thailand" name="Sakon Nakhon" pageId="1" pageNumber="56">Sakon Nakhon</collectingRegion>
Provinces) (
<figureCitation id="1376EEFEFFD9FFECFDB5F830FDA34F8C" box="[558,627,1940,1967]" captionStart="FIGURE 3" captionStartId="4.[151,250,1392,1416]" captionTargetBox="[151,1435,181,1368]" captionTargetId="figure-24@4.[151,1435,181,1368]" captionTargetPageId="4" captionText="FIGURE 3. Distribution of all known specimens of Heissophila Schuh, 2006. Blue dot = distribution in Thailand; yellow dot = distribution in Indonesia; red dot = distribution in China." figureDoi="http://doi.org/10.5281/zenodo.10279903" httpUri="https://zenodo.org/record/10279903/files/figure.png" pageId="1" pageNumber="56">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>