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<document ID-ISSN="2107-7207" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683046568874" checkinUser="felipe" docAuthor="PešićK, Vladimir, K, Harry Smit &amp; K, Mer Man Gurung" docDate="2022" docId="03E087CD5675FFFE4D96B69EFBE4F84F" docLanguage="en" docName="Acarologia.62.3.821-860.pdf" docOrigin="Acarologia 1941 (3)" docStyle="DocumentStyle:0387F17C4AF9F8D6336E46CC5E5664D0.6:Acarologia.2018-.journal_article" docStyleId="0387F17C4AF9F8D6336E46CC5E5664D0" docStyleName="Acarologia.2018-.journal_article" docStyleVersion="6" docTitle="Torrenticola trongsaensis PešićK &amp; K &amp; K 2022, sp. nov." docType="treatment" docVersion="1" lastPageNumber="843" masterDocId="FFD9FFB55663FFE94C7DB12FFFC2FFB2" masterDocTitle="Torrenticolid water mites of Bhutan. Genera Torrenticola Piersig, 1896 and Neoatractides Lundblad, 1941 (Acari: Hydrachnidia: Torrenticolidae)" masterLastPageNumber="860" masterPageNumber="821" pageNumber="842" updateTime="1683046720932" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Torrenticolid water mites of Bhutan. Genera Torrenticola Piersig, 1896 and Neoatractides Lundblad, 1941 (Acari: Hydrachnidia: Torrenticolidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>PešićK, Vladimir</mods:namePart>
<mods:affiliation>University of Montenegro, Cetinjski put bb, 81000 Podgorica, Montenegro.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>K, Harry Smit</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>K, Mer Man Gurung</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Acarologia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1941</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
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<mods:extent unit="page">
<mods:start>821</mods:start>
<mods:end>860</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="ISSN">2107-7207</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03E087CD5675FFFE4D96B69EFBE4F84F" httpUri="http://treatment.plazi.org/id/03E087CD5675FFFE4D96B69EFBE4F84F" lastPageId="23" lastPageNumber="843" pageId="22" pageNumber="842">
<subSubSection pageId="22" pageNumber="842" type="nomenclature">
<paragraph blockId="22.[491,920,1969,1995]" box="[491,920,1969,1995]" pageId="22" pageNumber="842">
<heading bold="true" box="[491,920,1969,1995]" fontSize="11" level="2" pageId="22" pageNumber="842" reason="3">
<emphasis bold="true" box="[491,920,1969,1995]" pageId="22" pageNumber="842">
<taxonomicName authority="PešićK &amp; K &amp; K, 2022" authorityName="PešićK &amp; K &amp; K" authorityYear="2022" box="[491,814,1969,1995]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="22" pageNumber="842" phylum="Arthropoda" rank="species" species="trongsaensis" status="sp. nov.">
<emphasis bold="true" box="[491,814,1969,1995]" italics="true" pageId="22" pageNumber="842">Torrenticola trongsaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[822,920,1969,1995]" pageId="22" pageNumber="842" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="22.[491,998,2025,2044]" box="[491,998,2025,2044]" pageId="22" pageNumber="842">
Zoobank:
<uuid box="[580,998,2025,2044]" pageId="22" pageNumber="842">67C92662-2D1B-4678-8E87-E6C0AFFA9F29</uuid>
</paragraph>
</subSubSection>
<subSubSection box="[531,766,229,253]" pageId="23" pageNumber="843" type="description">
<paragraph blockId="23.[491,1559,229,2045]" box="[531,766,229,253]" pageId="23" pageNumber="843">
<figureCitation box="[531,650,229,253]" captionStart="Figure 15" captionStartId="24.[119,190,2015,2036]" captionTargetBox="[143,1458,227,1952]" captionTargetId="figure-2@24.[137,1460,221,1959]" captionTargetPageId="24" captionText="Figure 15 Torrenticola trongsaensissp. nov., ♂ holotype, MG15 Khabab Chhu: A frontal margin of idiosoma, dorsal view; B dorsal shield; C ventral shield; D palp, lateral view; E palp, medial view; F I-L-5 and -6. Scale bars = 100 μm." pageId="23" pageNumber="843">Figures 15</figureCitation>
,
<figureCitation box="[662,692,229,253]" captionStart="Figure 16" captionStartId="25.[119,190,1901,1922]" captionTargetBox="[105,1490,267,1840]" captionTargetId="figure-2@25.[103,1494,259,1847]" captionTargetPageId="25" captionText="Figure 16 Torrenticola trongsaensissp. nov., ♀, MG14 Chuserbu stream: A dorsal shield; B ventral shield; C palp, medial view; D palp, lateral view; E gnathosoma and chelicera. Scale bars = 100 μm." pageId="23" pageNumber="843">16</figureCitation>
,
<figureCitation box="[703,766,229,253]" captionStart="Figure 19" captionStartId="29.[119,190,1992,2013]" captionTargetBox="[175,1422,221,1938]" captionTargetId="figure-2@29.[175,1422,221,1939]" captionTargetPageId="29" captionText="Figure 19 Photographs of selected structures. A, B, D, H, I, Dorsal shield; C, G, J, Ejaculatory complex; E, F, frontal margin of idiosoma, dorsal view.AG Torrenticola dbgurungisp. nov. (AC, EG ♂, D, F ♀): A, CF MG8 Dangdung Chhu; B, G MG9 Kartigang Chhu. HJ T. trongsaensissp. nov. (H, J ♂, I ♀): H, J MG15 Khabab Chhu; I MG14 Chuserbu stream." pageId="23" pageNumber="843">19HJ</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="843" type="materials_examined">
<paragraph blockId="23.[491,1559,229,2045]" box="[531,1527,264,288]" pageId="23" pageNumber="843">
<emphasis bold="true" box="[531,753,264,288]" pageId="23" pageNumber="843">Material examined</emphasis>
<materialsCitation box="[787,1527,264,288]" collectionCode="MG" country="Bhutan" location="Khabab" pageId="23" pageNumber="843" specimenCode="MG15" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[787,881,264,288]" pageId="23" pageNumber="843">Holotype</typeStatus>
<specimenCount box="[889,911,265,288]" pageId="23" pageNumber="843" type="male"></specimenCount>
, dissected and slide mounted,
<collectingCountry box="[1264,1348,264,288]" name="Bhutan" pageId="23" pageNumber="843">Bhutan</collectingCountry>
,
<specimenCode box="[1359,1429,264,288]" collectionCode="MG" country="China" name="Museum of Zoology" pageId="23" pageNumber="843">MG15</specimenCode>
<location LSID="urn:lsid:plazi:treatment:03E087CD5675FFFE4D96B69EFBE4F84F:8E9660005674FFFE49E1B027FA35FE92" box="[1436,1527,264,288]" country="Bhutan" name="Khabab" pageId="23" pageNumber="843">Khabab</location>
</materialsCitation>
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" pageId="23" pageNumber="843">
Chhu,
<geoCoordinate box="[575,681,298,322]" degrees="27.48492" direction="north" orientation="latitude" pageId="23" pageNumber="843" precision="1" value="27.48492">27.48492°N</geoCoordinate>
,
<geoCoordinate box="[693,801,298,322]" degrees="90.33490" direction="east" orientation="longitude" pageId="23" pageNumber="843" precision="1" value="90.3349">90.33490°E</geoCoordinate>
,
<quantity box="[812,890,298,322]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="23" pageNumber="843" unit="m" value="2500.0">2500 m</quantity>
asl,
<date box="[943,1042,298,322]" pageId="23" pageNumber="843" value="2021-05-14">14.v.2021</date>
.
<typeStatus box="[1059,1162,299,323]" pageId="23" pageNumber="843">Paratypes</typeStatus>
:
<specimenCount box="[1178,1213,298,322]" pageId="23" pageNumber="843" type="male">1♂</specimenCount>
,
<specimenCount box="[1227,1261,298,322]" pageId="23" pageNumber="843" type="female">4♀</specimenCount>
, same data as
<typeStatus box="[491,589,333,357]" pageId="23" pageNumber="843">holotype</typeStatus>
;
<specimenCount box="[603,637,333,357]" pageId="23" pageNumber="843" type="female">1♀</specimenCount>
,
<collectionCode box="[653,698,333,357]" country="China" name="Museum of Zoology" pageId="23" pageNumber="843">MG</collectionCode>
14 Chuserbu stream,
<geoCoordinate box="[933,1052,333,357]" degrees="27.50246" direction="north" orientation="latitude" pageId="23" pageNumber="843" precision="1" value="27.50246">27.50246°N</geoCoordinate>
,
<geoCoordinate box="[1065,1168,333,357]" degrees="90.31782" direction="east" orientation="longitude" pageId="23" pageNumber="843" precision="1" value="90.31782">90.31782°E</geoCoordinate>
,
<quantity box="[1180,1264,333,357]" metricMagnitude="3" metricUnit="m" metricValue="2.666" pageId="23" pageNumber="843" unit="m" value="2666.0">2666 m</quantity>
asl,
<date box="[1318,1412,333,357]" pageId="23" pageNumber="843" value="2021-05-13">13.v.2021</date>
,
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,1466,367,391]" pageId="23" pageNumber="843">
dissected and slide mounted;
<specimenCount box="[836,869,367,391]" pageId="23" pageNumber="843" type="female">1♀</specimenCount>
,
<collectionCode box="[884,926,367,391]" country="China" name="Museum of Zoology" pageId="23" pageNumber="843">MG</collectionCode>
12 Nikka Chhu,
<geoCoordinate box="[1111,1230,367,391]" degrees="27.52601" direction="north" orientation="latitude" pageId="23" pageNumber="843" precision="1" value="27.52601">27.52601°N</geoCoordinate>
,
<geoCoordinate box="[1241,1340,367,391]" degrees="90.29947" direction="east" orientation="longitude" pageId="23" pageNumber="843" precision="1" value="90.29947">90.29947°E</geoCoordinate>
,
<quantity box="[1350,1423,367,391]" metricMagnitude="3" metricUnit="m" metricValue="2.6089999999999995" pageId="23" pageNumber="843" unit="m" value="2609.0">2609 m</quantity>
asl,
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,594,402,426]" pageId="23" pageNumber="843">
<date box="[491,590,402,426]" pageId="23" pageNumber="843" value="2021-05-10">10.v.2021</date>
.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="843" type="diagnosis">
<paragraph blockId="23.[491,1559,229,2045]" box="[531,1479,436,461]" pageId="23" pageNumber="843">
<emphasis bold="true" box="[531,648,437,461]" pageId="23" pageNumber="843">Diagnosis</emphasis>
— Idiosoma large in size,&gt; 1000 µm in both sexes; Cx-I relatively short,
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" pageId="23" pageNumber="843">anteriorly broad; gnathosomal rostrum short, less than width of gnathosoma; in both sexes suture lines of Cx-IV extending posteriorly beyond posterior margin of genital field; ejaculatory complex with well-developed anterior keel and proximal arms.</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="843" type="description">
<paragraph blockId="23.[491,1559,229,2045]" pageId="23" pageNumber="843">
<emphasis bold="true" box="[531,662,574,598]" pageId="23" pageNumber="843">Description</emphasis>
<emphasis box="[707,896,574,598]" italics="true" pageId="23" pageNumber="843">General features</emphasis>
Idiosoma roundish; dorsal shield without a colour pattern (as photographed in
<figureCitation box="[816,973,609,633]" captionStart="Figure 19" captionStartId="29.[119,190,1992,2013]" captionTargetBox="[175,1422,221,1938]" captionTargetId="figure-2@29.[175,1422,221,1939]" captionTargetPageId="29" captionText="Figure 19 Photographs of selected structures. A, B, D, H, I, Dorsal shield; C, G, J, Ejaculatory complex; E, F, frontal margin of idiosoma, dorsal view.AG Torrenticola dbgurungisp. nov. (AC, EG ♂, D, F ♀): A, CF MG8 Dangdung Chhu; B, G MG9 Kartigang Chhu. HJ T. trongsaensissp. nov. (H, J ♂, I ♀): H, J MG15 Khabab Chhu; I MG14 Chuserbu stream." pageId="23" pageNumber="843">Figures 19HI</figureCitation>
); area of primary sclerotization of the dorsal plate with four dorsoglandularia; frontal platelets broad, frontal platelets broad, relatively short; Cx-I relatively short, anteriorly broad (
<figureCitation box="[848,975,677,701]" captionStart="Figure 15" captionStartId="24.[119,190,2015,2036]" captionTargetBox="[143,1458,227,1952]" captionTargetId="figure-2@24.[137,1460,221,1959]" captionTargetPageId="24" captionText="Figure 15 Torrenticola trongsaensissp. nov., ♂ holotype, MG15 Khabab Chhu: A frontal margin of idiosoma, dorsal view; B dorsal shield; C ventral shield; D palp, lateral view; E palp, medial view; F I-L-5 and -6. Scale bars = 100 μm." pageId="23" pageNumber="843">Figure 15C</figureCitation>
); gnathosomal bay U-shaped, proximally rounded;
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" pageId="23" pageNumber="843">Cxgl-4 subapical; medial suture line of Cx-II+III relatively short; suture lines of Cx-IV in both sexes distinctly extending posteriorly beyond posterior margin of genital field, laterally curved;</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" pageId="23" pageNumber="843">
excretory pore and Vgl-2 away from the line of primary sclerotization, excretory pore on the level of Vgl-2; gnathosomal rostrum short, less than depth of gnathosoma (
<figureCitation box="[1347,1477,815,839]" captionStart="Figure 16" captionStartId="25.[119,190,1901,1922]" captionTargetBox="[105,1490,267,1840]" captionTargetId="figure-2@25.[103,1494,259,1847]" captionTargetPageId="25" captionText="Figure 16 Torrenticola trongsaensissp. nov., ♀, MG14 Chuserbu stream: A dorsal shield; B ventral shield; C palp, medial view; D palp, lateral view; E gnathosoma and chelicera. Scale bars = 100 μm." pageId="23" pageNumber="843">Figure 16E</figureCitation>
); P-2
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" pageId="23" pageNumber="843">
ventral margin nearly straight, P-2 and P-3 ventrodistal protrusions bluntly pointed, P-4 with a ventral tubercle bearing one long and three shorter setae (
<figureCitation box="[1130,1284,884,908]" captionStart="Figure 15" captionStartId="24.[119,190,2015,2036]" captionTargetBox="[143,1458,227,1952]" captionTargetId="figure-2@24.[137,1460,221,1959]" captionTargetPageId="24" captionText="Figure 15 Torrenticola trongsaensissp. nov., ♂ holotype, MG15 Khabab Chhu: A frontal margin of idiosoma, dorsal view; B dorsal shield; C ventral shield; D palp, lateral view; E palp, medial view; F I-L-5 and -6. Scale bars = 100 μm." pageId="23" pageNumber="843">Figures 15D, E</figureCitation>
)
<emphasis box="[1283,1342,884,908]" italics="true" pageId="23" pageNumber="843">Male</emphasis>
Genital field elongated, subrectangular; ejaculatory complex conventional in shape (
<figureCitation box="[1267,1380,919,943]" captionStart="Figure 19" captionStartId="29.[119,190,1992,2013]" captionTargetBox="[175,1422,221,1938]" captionTargetId="figure-2@29.[175,1422,221,1939]" captionTargetPageId="29" captionText="Figure 19 Photographs of selected structures. A, B, D, H, I, Dorsal shield; C, G, J, Ejaculatory complex; E, F, frontal margin of idiosoma, dorsal view.AG Torrenticola dbgurungisp. nov. (AC, EG ♂, D, F ♀): A, CF MG8 Dangdung Chhu; B, G MG9 Kartigang Chhu. HJ T. trongsaensissp. nov. (H, J ♂, I ♀): H, J MG15 Khabab Chhu; I MG14 Chuserbu stream." pageId="23" pageNumber="843">Figure 19J</figureCitation>
).
<emphasis box="[1389,1473,919,943]" italics="true" pageId="23" pageNumber="843">Female</emphasis>
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,984,953,977]" pageId="23" pageNumber="843">Genital field large and pentagonal in shape.</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[531,1487,988,1012]" pageId="23" pageNumber="843">
<emphasis box="[531,685,989,1012]" italics="true" pageId="23" pageNumber="843">Measurements</emphasis>
.
<emphasis box="[704,763,988,1012]" italics="true" pageId="23" pageNumber="843">Male</emphasis>
(
<typeStatus box="[774,871,988,1012]" pageId="23" pageNumber="843">holotype</typeStatus>
) Idiosoma (ventral view:
<figureCitation box="[1180,1306,988,1012]" captionStart="Figure 15" captionStartId="24.[119,190,2015,2036]" captionTargetBox="[143,1458,227,1952]" captionTargetId="figure-2@24.[137,1460,221,1959]" captionTargetPageId="24" captionText="Figure 15 Torrenticola trongsaensissp. nov., ♂ holotype, MG15 Khabab Chhu: A frontal margin of idiosoma, dorsal view; B dorsal shield; C ventral shield; D palp, lateral view; E palp, medial view; F I-L-5 and -6. Scale bars = 100 μm." pageId="23" pageNumber="843">Figure 15C</figureCitation>
) L 1065, W 853;
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,1485,1022,1046]" pageId="23" pageNumber="843">
dorsal shield (
<figureCitation box="[659,785,1022,1046]" captionStart="Figure 15" captionStartId="24.[119,190,2015,2036]" captionTargetBox="[143,1458,227,1952]" captionTargetId="figure-2@24.[137,1460,221,1959]" captionTargetPageId="24" captionText="Figure 15 Torrenticola trongsaensissp. nov., ♂ holotype, MG15 Khabab Chhu: A frontal margin of idiosoma, dorsal view; B dorsal shield; C ventral shield; D palp, lateral view; E palp, medial view; F I-L-5 and -6. Scale bars = 100 μm." pageId="23" pageNumber="843">Figure 15B</figureCitation>
) L 909, W 738, L/W ratio 1.23; dorsal plate L 825; shoulder plate L
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,1425,1057,1081]" pageId="23" pageNumber="843">288294, W 125131, L/W ratio 2.22.3; frontal plate L 194, W 113116, L/W ratio 1.681.72;</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,1460,1091,1115]" pageId="23" pageNumber="843">shoulder/frontal plate L 1.481.52. Gnathosomal bay L 206, Cx-I total L 397, Cx-I mL 191,</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,1487,1125,1149]" pageId="23" pageNumber="843">Cx-II+III mL 96; ratio Cx-I L/Cx-II+III mL 4.1; Cx-I mL/Cx-II+III mL 2.0. Genital field L/W</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" pageId="23" pageNumber="843">234/162, ratio 1.45; distance genital field-excretory pore 200, genital field-caudal idiosoma margin 313. Gnathosoma vL 334, chelicera L 384; palp total L 360, dL/H, dL/H ratio: P-1,</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,1394,1229,1253]" pageId="23" pageNumber="843">38/39, 0.96; P-2, 116/64, 1.81; P-3, 69/56, 1.22; P-4, 114/34, 3.32; P-5, 23/14, 1.7; L ratio</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,1386,1263,1287]" pageId="23" pageNumber="843">P-2/P-4 1.01. dL of I-L-26: 94, 109, 134, 153, 144; I-L-6 H 48; dL/H I-L-6 ratio 3.0.</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[531,1501,1298,1322]" pageId="23" pageNumber="843">
<emphasis box="[531,615,1298,1322]" italics="true" pageId="23" pageNumber="843">Female</emphasis>
(
<typeStatus box="[625,714,1298,1322]" pageId="23" pageNumber="843">paratype</typeStatus>
from Chuserbu stream, n = 1) Idiosoma (ventral view:
<figureCitation box="[1342,1469,1298,1322]" captionStart="Figure 16" captionStartId="25.[119,190,1901,1922]" captionTargetBox="[105,1490,267,1840]" captionTargetId="figure-2@25.[103,1494,259,1847]" captionTargetPageId="25" captionText="Figure 16 Torrenticola trongsaensissp. nov., ♀, MG14 Chuserbu stream: A dorsal shield; B ventral shield; C palp, medial view; D palp, lateral view; E gnathosoma and chelicera. Scale bars = 100 μm." pageId="23" pageNumber="843">Figure 16B</figureCitation>
) L
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,1462,1332,1356]" pageId="23" pageNumber="843">
1108, W 906; dorsal shield (
<figureCitation box="[806,937,1332,1356]" captionStart="Figure 16" captionStartId="25.[119,190,1901,1922]" captionTargetBox="[105,1490,267,1840]" captionTargetId="figure-2@25.[103,1494,259,1847]" captionTargetPageId="25" captionText="Figure 16 Torrenticola trongsaensissp. nov., ♀, MG14 Chuserbu stream: A dorsal shield; B ventral shield; C palp, medial view; D palp, lateral view; E gnathosoma and chelicera. Scale bars = 100 μm." pageId="23" pageNumber="843">Figure 16A</figureCitation>
) L 947, W 768, L/W ratio 1.23; dorsal plate L 869;
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,1436,1367,1391]" pageId="23" pageNumber="843">shoulder plate L 294300, W 134138, L/W ratio 2.172.2; frontal plate L 194, W 119, L/W</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,1476,1401,1425]" pageId="23" pageNumber="843">ratio 1.63; shoulder/frontal plate L 1.521.55. Gnathosomal bay L 206, Cx-I total L 397, Cx-I</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" pageId="23" pageNumber="843">mL 191, Cx-II+III mL 40; ratio Cx-I L/Cx-II+III mL 9.9; Cx-I mL/Cx-II+III mL 4.8. Genital field L/W 247/209, ratio 1.18; distance genital field-excretory pore 250, genital field-caudal idiosoma margin 397. Gnathosoma vL 353, chelicera L 366; palp total L 366, dL/H, dL/H ratio:</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,1416,1539,1563]" pageId="23" pageNumber="843">P-1, 38/40, 0.96; P-2, 118/61, 1.94; P-3, 72/58, 1.25; P-4, 117/36, 3.37; P-5, 21/14, 1.5; L ratio</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,615,1573,1597]" pageId="23" pageNumber="843">P-2/P-4 1.0.</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="843" type="etymology">
<paragraph blockId="23.[491,1559,229,2045]" pageId="23" pageNumber="843">
<emphasis bold="true" box="[531,658,1608,1632]" pageId="23" pageNumber="843">Etymology</emphasis>
— The species is named after the district
<collectingRegion box="[1127,1219,1609,1633]" country="Bhutan" name="Tongsa" pageId="23" pageNumber="843">Trongsa</collectingRegion>
from where the new species was collected.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="843" type="discussion">
<paragraph blockId="23.[491,1559,229,2045]" pageId="23" pageNumber="843">
<emphasis bold="true" box="[531,651,1677,1701]" pageId="23" pageNumber="843">Discussion</emphasis>
— In regard to the presence of an anteriorly broad and short Cx-I, an elongated genital field in the male (L/W ratio about 1.4), and a deep gnathosoma, with a short rostrum,
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,1459,1746,1770]" pageId="23" pageNumber="843">
the new species resembles
<taxonomicName authority="(Lundblad, 1941)" baseAuthorityName="Lundblad" baseAuthorityYear="1941" box="[788,1149,1746,1770]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="843" phylum="Arthropoda" rank="species" species="retractipora">
<emphasis box="[788,955,1747,1770]" italics="true" pageId="23" pageNumber="843">T. retractipora</emphasis>
(
<bibRefCitation author="Lundblad, O." box="[963,1142,1746,1770]" pageId="23" pageNumber="843" pagination="97 - 121" refId="ref15588" refString="Lundblad, O. (1941) Neue Wassermilben. Vorlaufige Mitteilung. Entomol. Tidskr., 62, 97 - 121." type="journal article" year="1941">Lundblad, 1941</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName authority="Pesic &amp; Smit, 2009" authorityName="Pesic &amp; Smit" authorityYear="2009" box="[1201,1459,1746,1770]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="843" phylum="Arthropoda" rank="species" species="siamis">
<emphasis box="[1201,1301,1747,1770]" italics="true" pageId="23" pageNumber="843">T. siamis</emphasis>
<bibRefCitation author="Pesic V. &amp; Smit H." box="[1307,1459,1746,1770]" pageId="23" pageNumber="843" pagination="38 - 62" refId="ref15897" refString="Pesic V., Smit H. 2009. Water mites of the family Torrenticolidae Piersig, 1902 (Acari: Hydrachnidia) from Thailand, Part I. The genera Torrenticola Piersig, 1896, Neoatractides Lundblad, 1941 and Pseudotorrenticola Walter, 1906. Zootaxa, 1982: 38 - 62. https: // doi. org / 10.11646 / zootaxa. 1982.1.2" type="journal article" year="2009">Pešić &amp; Smit,</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" pageId="23" pageNumber="843">
2009. The latter two species can be separated from
<taxonomicName authority="PešićK &amp; K &amp; K, 2022" authorityName="PešićK &amp; K &amp; K" authorityYear="2022" box="[1055,1221,1781,1804]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="843" phylum="Arthropoda" rank="species" species="trongsaensis" status="sp. nov.">
<emphasis box="[1055,1221,1781,1804]" italics="true" pageId="23" pageNumber="843">T. trongsaensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1224,1318,1781,1805]" pageId="23" pageNumber="843">
<taxonomicNameLabel box="[1224,1318,1781,1805]" pageId="23" pageNumber="843" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by Cx-IV not extended posterior of the genital field, and a comparatively shorter and stouter gnathosomal rostrum.
<taxonomicName baseAuthorityName="Lundblad" baseAuthorityYear="1941" box="[591,858,1849,1873]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="843" phylum="Arthropoda" rank="species" species="retractipora">
<emphasis box="[591,858,1849,1873]" italics="true" pageId="23" pageNumber="843">Torrenticola retractipora</emphasis>
</taxonomicName>
, known from a single male specimen collected in a stream at Kambaiti pass in
<collectingCountry box="[709,808,1884,1908]" name="Myanmar" pageId="23" pageNumber="843">Myanmar</collectingCountry>
differs by the characteristic shape of the ejaculatory complex
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" pageId="23" pageNumber="843">
(similar to
<taxonomicName authorityName="Pesic &amp; Smit" authorityYear="2007" box="[609,732,1918,1942]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="843" phylum="Arthropoda" rank="species" species="bhutana">
<emphasis box="[609,732,1918,1942]" italics="true" pageId="23" pageNumber="843">T. bhutana</emphasis>
</taxonomicName>
as illustrated in
<figureCitation box="[908,1038,1918,1942]" captionStart="Figure 11" captionStartId="18.[119,190,1435,1456]" captionTargetBox="[118,1473,233,1381]" captionTargetId="figure-2@18.[103,1494,221,1381]" captionTargetPageId="18" captionText="Figure 11 Torrenticola longisetasp. nov., ♀ paratype, MG5 Dakpay Chhu: A ventral shield; B palp, medial view. Scale bars = 100 μm." pageId="23" pageNumber="843">Figure 11G</figureCitation>
: proximal and distal arms short, proximal chamber large, proximal horns reduced, see
<bibRefCitation author="Lundblad O." box="[993,1159,1952,1976]" pageId="23" pageNumber="843" pagination="289 - 443" refId="ref15673" refString="Lundblad O. 1969. Indische Wassermilben, hauptsachlich von Hinterindien. Ark. Zool., 22: 289 - 443." type="journal article" year="1969">Lundblad 1969</bibRefCitation>
, Figure 27f).
<taxonomicName authorityName="Pesic &amp; Smit" authorityYear="2009" box="[1279,1484,1952,1976]" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="843" phylum="Arthropoda" rank="species" species="siamis">
<emphasis box="[1279,1484,1952,1976]" italics="true" pageId="23" pageNumber="843">Torrenticola siamis</emphasis>
</taxonomicName>
,
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,1559,1987,2011]" pageId="23" pageNumber="843">
a species originally described from
<collectingCountry box="[882,991,1987,2011]" name="Thailand" pageId="23" pageNumber="843">Thailand</collectingCountry>
, has a more elongated dorsal shield (L/W ratio 1.4
</paragraph>
<paragraph blockId="23.[491,1559,229,2045]" box="[491,1062,2021,2045]" pageId="23" pageNumber="843">
in
<specimenCount box="[519,539,2022,2045]" pageId="23" pageNumber="843" type="male"></specimenCount>
<typeStatus box="[545,642,2021,2045]" pageId="23" pageNumber="843">holotype</typeStatus>
, data taken from
<bibRefCitation author="Pesic V. &amp; Smit H." box="[832,1051,2021,2045]" pageId="23" pageNumber="843" pagination="38 - 62" refId="ref15897" refString="Pesic V., Smit H. 2009. Water mites of the family Torrenticolidae Piersig, 1902 (Acari: Hydrachnidia) from Thailand, Part I. The genera Torrenticola Piersig, 1896, Neoatractides Lundblad, 1941 and Pseudotorrenticola Walter, 1906. Zootaxa, 1982: 38 - 62. https: // doi. org / 10.11646 / zootaxa. 1982.1.2" type="journal article" year="2009">Pešić and Smit 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>