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<document id="2B264DAA8AE60A825681D2FDF4C7D0B6" ID-DOI="10.15407/zoo2021.02.143" ID-GBIF-Dataset="38e28475-92b4-45cc-b0e2-bbefb71cbca5" ID-ISSN="2707-7268" ID-Zenodo-Dep="6455840" IM.bibliography_approvedBy="admin" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="admin" IM.tables_requiresApprovalFor="existingObjects,operationResults" IM.taxonomicNames_approvedBy="admin" IM.treatments_approvedBy="admin" checkinTime="1649789298792" checkinUser="felipe" docAuthor="Koretsky, I. A. &amp; Rahmat, S. J." docDate="2021" docId="039CDB4195554C62FF5FFCEDFF73FEC0" docLanguage="en" docName="Zoodiversity.55.2.143-154.pdf" docOrigin="Zoodiversity 55 (2)" docSource="http://dx.doi.org/10.15407/zoo2021.02.143" docStyle="DocumentStyle:65E6F982F72656C606C1639D3996B172.2:Zoodiversity.2020-.journal_article" docStyleId="65E6F982F72656C606C1639D3996B172" docStyleName="Zoodiversity.2020-.journal_article" docStyleVersion="2" docTitle="Phocinae Gray 1821" docType="treatment" docVersion="6" lastPageNumber="150" masterDocId="FFA5A33995534C65FFF9FFC2FF99FFEA" masterDocTitle="Unique Short-Faced Miocene Seal Discovered In Grytsiv (Ukraine)" masterLastPageNumber="154" masterPageNumber="143" pageNumber="149" updateTime="1695856864423" updateUser="admin" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="1313F0B066DFF3B9F793E2F0A8BA230C">Unique Short-Faced Miocene Seal Discovered In Grytsiv (Ukraine)</mods:title>
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<mods:namePart id="F565018DC490BB3F7868BAE92901AB2D">Koretsky, I. A.</mods:namePart>
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<mods:namePart id="07735B3D5CC76162C1269C97122EEE27">Rahmat, S. J.</mods:namePart>
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<treatment id="039CDB4195554C62FF5FFCEDFF73FEC0" ID-DOI="http://doi.org/10.5281/zenodo.6462356" ID-GBIF-Taxon="194602225" ID-Zenodo-Dep="6462356" LSID="urn:lsid:plazi:treatment:039CDB4195554C62FF5FFCEDFF73FEC0" httpUri="http://treatment.plazi.org/id/039CDB4195554C62FF5FFCEDFF73FEC0" lastPageId="7" lastPageNumber="150" pageId="6" pageNumber="149">
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<paragraph id="8B8A6A5795554C63FF5FFCEDFE5CFCA3" blockId="6.[166,453,815,841]" box="[166,453,815,841]" pageId="6" pageNumber="149">
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<emphasis id="B941B64595554C63FF5FFCEDFE5CFCA3" bold="true" box="[166,453,815,841]" pageId="6" pageNumber="149">
Subfamily
<taxonomicName id="4C3511D495554C63FED3FCEDFE5CFCA3" authorityName="Gray" authorityYear="1821" box="[298,453,815,841]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="subFamily" subFamily="Phocinae">PHOCINAE</taxonomicName>
</emphasis>
</heading>
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<subSubSection id="C32F39DC95554C62FF22FCB2FF73FEC0" lastPageId="7" lastPageNumber="150" pageId="6" pageNumber="149" type="discussion">
<paragraph id="8B8A6A5795554C63FF22FCB2FE26FBC3" blockId="6.[166,1225,879,1065]" pageId="6" pageNumber="149">
Due to the lack of preservation and fragility of fossil seal skulls, less than
<specimenCount id="9D33A1DE95554C63FBD5FCADFB51FC63" box="[1068,1224,879,905]" pageId="6" pageNumber="149" type="generic">20 specimens</specimenCount>
have been described so far, with only a few belonging to representatives of the subfamily
<taxonomicName id="4C3511D495554C63FF5FFC6DFE8FFC23" authorityName="Gray" authorityYear="1821" box="[166,278,943,969]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="subFamily" subFamily="Phocinae">Phocinae</taxonomicName>
. This new partial seal skull of
<taxonomicName id="4C3511D495554C63FD74FC6DFCE5FC23" authorityName="Koretsky &amp; Rahmat" authorityYear="2021" box="[653,892,943,969]" class="Mammalia" family="Phocidae" genus="Planopusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="semenovi">
<emphasis id="B941B64595554C63FD74FC6DFCE5FC23" box="[653,892,943,969]" italics="true" pageId="6" pageNumber="149">Planopusa semenovi</emphasis>
</taxonomicName>
is extremely small and has a short face (based on the size of the rostral part and teeth). Based on teeth formula,
<taxonomicName id="4C3511D495554C63FF5FFC2DFEAFFBE3" authority="Koretsky &amp; Rahmat, 2021" authorityName="Koretsky &amp; Rahmat" authorityYear="2021" box="[166,310,1007,1033]" class="Mammalia" family="Phocidae" genus="Planopusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="semenovi" status="sp. n.">
<emphasis id="B941B64595554C63FF5FFC2DFEAFFBE3" box="[166,310,1007,1033]" italics="true" pageId="6" pageNumber="149">Pl. semenovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2720B3E95554C63FEC4FC32FEE3FBE0" box="[317,378,1008,1034]" pageId="6" pageNumber="149" rank="species">sp. n.</taxonomicNameLabel>
belongs to the subfamily
<taxonomicName id="4C3511D495554C63FD5FFC2DFC8AFBE3" authorityName="Gray" authorityYear="1821" box="[678,787,1007,1033]" class="Mammalia" family="Phocidae" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="subFamily" subFamily="Phocinae">Phocinae</taxonomicName>
and will be compared to other recent and fossil phocine seals.
</paragraph>
<paragraph id="8B8A6A5795554C63FF5FFB8DFDB3FB83" blockId="6.[166,1225,1103,1834]" box="[166,554,1103,1129]" pageId="6" pageNumber="149">
<taxonomicName id="4C3511D495554C63FF5FFB8DFE08FB83" authorityName="Koretsky &amp; Rahmat" authorityYear="2021" box="[166,401,1103,1129]" class="Mammalia" family="Phocidae" genus="Planopusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="semenovi">
<emphasis id="B941B64595554C63FF5FFB8DFE08FB83" box="[166,401,1103,1129]" italics="true" pageId="6" pageNumber="149">Planopusa semenovi</emphasis>
</taxonomicName>
differs from:
</paragraph>
<paragraph id="8B8A6A5795554C63FF22FBB2FE15FB23" blockId="6.[166,1225,1103,1834]" pageId="6" pageNumber="149">
Recent
<taxonomicName id="4C3511D495554C63FECDFBB2FE51FB63" box="[308,456,1136,1161]" class="Mammalia" family="Phocidae" genus="Pusa" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="capsica">
<emphasis id="B941B64595554C63FECDFBB2FE51FB63" box="[308,456,1136,1161]" italics="true" pageId="6" pageNumber="149">Pusa capsica</emphasis>
</taxonomicName>
by: being smaller, having well-developed, round anterior palatal foramen; the presence of palatal groove; the absence of diastema; the shape of the proximal edge of hard palate.
</paragraph>
<paragraph id="8B8A6A5795554C63FF22FB0DFECAFA80" blockId="6.[166,1225,1103,1834]" pageId="6" pageNumber="149">
<taxonomicName id="4C3511D495554C63FF22FB0DFE1FFB00" box="[219,390,1231,1258]" class="Mammalia" family="Phocidae" genus="Phocanella" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B941B64595554C63FF22FB0DFEC3FB03" box="[219,346,1231,1257]" italics="true" pageId="6" pageNumber="149">Phocanella</emphasis>
sp.
</taxonomicName>
, cf.
<taxonomicName id="4C3511D495554C63FE42FB12FDA9FB03" baseAuthorityName="Van Beneden" baseAuthorityYear="1876" box="[443,560,1231,1257]" class="Mammalia" family="Phocidae" genus="Phocanella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="pumila">
<emphasis id="B941B64595554C63FE42FB12FDA9FB03" box="[443,560,1231,1257]" italics="true" pageId="6" pageNumber="149">P. pumila</emphasis>
</taxonomicName>
(Pliocene, Yorktown Formation 5.33.6 Ma) from Lee Creek Mine,
<collectingCountry id="F3222AC795554C63FEC6FB2DFEEEFAE3" box="[319,375,1263,1289]" name="United States of America" pageId="6" pageNumber="149">USA</collectingCountry>
(
<bibRefCitation id="EFA417A695554C63FE7CFB2DFDA2FAE0" author="Ray" box="[389,571,1263,1290]" etAl="et al." firstAuthor="Ray" pageId="6" pageNumber="149" pagination="81 - 140" refId="ref7321" refString="Ray, C. E., Bohaska, D. J., Koretsky I. A., Ward, L. W., Barnes, L. G. 2008. Geology and Paleontology of the Lee Creek Mine, North Carolina, IV. Virginia Museum of Natural History Special Publication, 14, 81 - 140." type="journal article" year="2008">Ray et al., 2008</bibRefCitation>
) by: the lack of diastema between teeth; short concavity over P
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preorbital part of the maxilla; the flattened palatal process of maxilla; the reverse ratios between P4 and M1; wider rostrum; rounded anterior palatal foramen, and deeper palatal groove.
</paragraph>
<paragraph id="8B8A6A5795554C63FF22FAADFD2CFA00" blockId="6.[166,1225,1103,1834]" pageId="6" pageNumber="149">
<taxonomicName id="4C3511D495554C63FF22FAADFE45FA63" authorityName="Koretsky" authorityYear="2001" box="[219,476,1391,1417]" class="Mammalia" family="Phocidae" genus="Histriophoca" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="alekseevi">
<emphasis id="B941B64595554C63FF22FAADFE45FA63" box="[219,476,1391,1417]" italics="true" pageId="6" pageNumber="149">Histriophoca alekseevi</emphasis>
</taxonomicName>
(Middle Miocene, middle Sarmatian [~11.69.9 Ma] of the Eastern Paratethys) by: the absence of diastemata; the flattened palatal process of the maxilla; the reverse ratios between P4 and M1; wider rostrum; round and deep anterior palatal foramen; uneven depth of the palatal groove.
</paragraph>
<paragraph id="8B8A6A5795554C63FF22FA2DFC3EF940" blockId="6.[166,1225,1103,1834]" pageId="6" pageNumber="149">
<taxonomicName id="4C3511D495554C63FF22FA2DFE56F9E3" baseAuthorityName="Eichwald" baseAuthorityYear="1850" box="[219,463,1519,1545]" class="Mammalia" family="Phocidae" genus="Monachopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="pontica">
<emphasis id="B941B64595554C63FF22FA2DFE56F9E3" box="[219,463,1519,1545]" italics="true" pageId="6" pageNumber="149">Monachopsis pontica</emphasis>
</taxonomicName>
(Middle Miocene, middle Sarmatian [~11.69.9 Ma] of the Eastern Paratethys) by: the smaller width of the palatal process; narrowest infraorbital foramen of any known fossil or modern seal, but larger than in other seals from the same region (such as
<taxonomicName id="4C3511D495554C63FEE9F98DFD8DF983" authorityName="Koretsky" authorityYear="2001" box="[272,532,1615,1641]" class="Mammalia" family="Phocidae" genus="Histriophoca" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="alekseevi">
<emphasis id="B941B64595554C63FEE9F98DFD8DF983" box="[272,532,1615,1641]" italics="true" pageId="6" pageNumber="149">Histriophoca alekseevi</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C3511D495554C63FDA1F992FCF6F983" baseAuthorityName="Toula" baseAuthorityYear="1897" box="[600,879,1615,1641]" class="Mammalia" family="Phocidae" genus="Praepusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="vindobonensis">
<emphasis id="B941B64595554C63FDA1F992FCF6F983" box="[600,879,1615,1641]" italics="true" pageId="6" pageNumber="149">Praepusa vindobonensis</emphasis>
</taxonomicName>
); longer and narrower teeth; width of the rostrum part of the skull (
<tableCitation id="C6B75FEC95554C63FD8BF9ADFD5AF963" box="[626,707,1647,1673]" captionStart="Table 1" captionStartId="4.[166,237,175,199]" captionTargetBox="[170,1218,240,1455]" captionTargetPageId="4" captionText="Table 1. Cranial measurements, mm" httpUri="http://table.plazi.org/id/DF4A3ADF95574C61FF5FFF6DFDDCFF2D" pageId="6" pageNumber="149" tableUuid="DF4A3ADF95574C61FF5FFF6DFDDCFF2D">table 1</tableCitation>
; which is similar in size to
<taxonomicName id="4C3511D495554C63FBFEF9ADFB50F963" baseAuthorityName="True" baseAuthorityYear="1906" box="[1031,1225,1647,1673]" class="Mammalia" family="Phocidae" genus="Leptophoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="lenis">
<emphasis id="B941B64595554C63FBFEF9ADFB50F963" box="[1031,1225,1647,1673]" italics="true" pageId="6" pageNumber="149">Leptophoca lenis</emphasis>
</taxonomicName>
despite the smaller size, but wider than in
<taxonomicName id="4C3511D495554C63FD6DF94DFC0FF943" authorityName="Koretsky" authorityYear="2001" box="[660,918,1679,1705]" class="Mammalia" family="Phocidae" genus="Histriophoca" kingdom="Animalia" order="Carnivora" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="alekseevi">
<emphasis id="B941B64595554C63FD6DF94DFC0FF943" box="[660,918,1679,1705]" italics="true" pageId="6" pageNumber="149">Histriophoca alekseevi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B8A6A5795554C63FF22F972FB8AF8C0" blockId="6.[166,1225,1103,1834]" pageId="6" pageNumber="149">
<taxonomicName id="4C3511D495554C63FF22F972FE70F923" box="[219,489,1711,1737]" class="Magnoliopsida" family="Gentianaceae" genus="Prepusa" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="149" phylum="Tracheophyta" rank="species" species="vindobonensis">
<emphasis id="B941B64595554C63FF22F972FE70F923" box="[219,489,1711,1737]" italics="true" pageId="6" pageNumber="149">Prepusa vindobonensis</emphasis>
</taxonomicName>
(middle Miocene, early Sarmatian [~12.811.6 Ma] of the Central and Eastern Paratethys) by: upper incisors forming a straight line; the absence of diastemata; the short concavity over P
<quantity id="4CCDC7B295554C63FD92F92DFD0DF8E3" box="[619,660,1775,1801]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="149" unit="in" value="1.0">1 in</quantity>
preorbital part of the maxilla; rounded anterior palatal foramen; the similar width of rostrum (despite overall smaller size).
</paragraph>
<paragraph id="8B8A6A5795544C62FF1AFF6DFF73FEC0" blockId="7.[174,1232,175,298]" pageId="7" pageNumber="150">
<taxonomicName id="4C3511D495544C62FF1AFF6DFE3AFF23" baseAuthorityName="True" baseAuthorityYear="1906" box="[227,419,175,201]" class="Mammalia" family="Phocidae" genus="Leptophoca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="150" phylum="Chordata" rank="species" species="lenis">
<emphasis id="B941B64595544C62FF1AFF6DFE3AFF23" box="[227,419,175,201]" italics="true" pageId="7" pageNumber="150">Leptophoca lenis</emphasis>
</taxonomicName>
(Early Miocene [~16 Ma],
<collectingCountry id="F3222AC795544C62FD27FF6DFC81FF20" box="[734,792,175,202]" name="United States of America" pageId="7" pageNumber="150">USA</collectingCountry>
) by: upper incisors forming a straight line; the absence of diastema; short concavity over P1 of the preorbital part of the maxilla; rounded anterior palatal foramen; similar width of the rostrum (despite overall smaller size).
</paragraph>
</subSubSection>
</treatment>
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