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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.66.49910" ID-GBIF-Dataset="f465159b-eaf9-4dea-8100-ed11756bff9b" ID-PMC="PMC7148386" ID-Pensoft-Pub="1314-4049-66-83" ID-Pensoft-UUID="9944774B7E5756C9B10B5EC09A81F0DF" ID-PubMed="32296287" ModsDocID="1314-4049-66-83" checkinTime="1585985437246" checkinUser="pensoft" docAuthor="Mu, Yan-Hong, Hu, Ya-Ping, Wei, Yu-Lian &amp; Yuan, Hai-Sheng" docDate="2020" docId="F149B8A970B1509DB706EFAB4952B890" docLanguage="en" docName="MycoKeys 66: 83-103" docOrigin="MycoKeys 66" docSource="http://dx.doi.org/10.3897/mycokeys.66.49910" docTitle="Sarcodon coactus Y. H. Mu &amp; H. S. Yuan 2020, sp. nov." docType="treatment" docVersion="4" id="9944774B7E5756C9B10B5EC09A81F0DF" lastPageNumber="83" masterDocId="9944774B7E5756C9B10B5EC09A81F0DF" masterDocTitle="Hydnaceous fungi of China 8. Morphological and molecular identification of three new species of Sarcodon and a new record from southwest China" masterLastPageNumber="103" masterPageNumber="83" pageNumber="83" updateTime="1668136447372" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Hydnaceous fungi of China 8. Morphological and molecular identification of three new species of Sarcodon and a new record from southwest China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mu, Yan-Hong</mods:namePart>
<mods:affiliation>CAS Key Laboratory of Forest Ecology and Management, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110164, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hu, Ya-Ping</mods:namePart>
<mods:affiliation>University of the Chinese Academy of Sciences, Beijing 100049, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wei, Yu-Lian</mods:namePart>
<mods:affiliation>CAS Key Laboratory of Forest Ecology and Management, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110164, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yuan, Hai-Sheng</mods:namePart>
<mods:affiliation>CAS Key Laboratory of Forest Ecology and Management, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110164, China</mods:affiliation>
<mods:nameIdentifier type="email">hsyuan@iae.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>MycoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>66</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>83</mods:start>
<mods:end>103</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.66.49910</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.66.49910</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-66-83</mods:identifier>
<mods:identifier type="Pensoft-UUID">9944774B7E5756C9B10B5EC09A81F0DF</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="163433665" LSID="urn:lsid:plazi:treatment:F149B8A970B1509DB706EFAB4952B890" httpUri="http://treatment.plazi.org/id/F149B8A970B1509DB706EFAB4952B890" lastPageNumber="83" pageId="0" pageNumber="83">
<subSubSection pageId="0" pageNumber="83" type="nomenclature">
<paragraph pageId="0" pageNumber="83">
<taxonomicName LSID="F149B8A9-70B1-509D-B706-EFAB4952B890" authority="Y. H. Mu &amp; H. S. Yuan" authorityName="Y. H. Mu &amp; H. S. Yuan" authorityYear="2020" class="Agaricomycetes" family="Bankeraceae" genus="Sarcodon" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Sarcodon coactus" order="Thelephorales" pageId="0" pageNumber="83" phylum="Basidiomycota" rank="species" species="coactus" status="sp. nov.">Sarcodon coactus Y.H. Mu &amp; H.S. Yuan</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="83">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A basidiocarp of Sarcodon coactus (holotype: IFP 019351)." figureDoi="10.3897/mycokeys.66.49910.figure2" httpUri="https://binary.pensoft.net/fig/397187" pageId="0" pageNumber="83">Figures 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM of basidiospores of Sarcodon coactus (holotype: IFP 019351)." figureDoi="10.3897/mycokeys.66.49910.figure3" httpUri="https://binary.pensoft.net/fig/397188" pageId="0" pageNumber="83">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Microscopic structures of Sarcodon coactus (drawn from IFP 019351) a basidiospores b section of hymenophoral trama with basidia c hyphae from pileal context." figureDoi="10.3897/mycokeys.66.49910.figure4" httpUri="https://binary.pensoft.net/fig/397189" pageId="0" pageNumber="83">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="83" type="diagnoses">
<paragraph pageId="0" pageNumber="83">Diagnoses.</paragraph>
<paragraph pageId="0" pageNumber="83">
Differs from
<taxonomicName authorityName="Maas Geesteranus" authorityYear="1964" baseAuthorityName="Berk. &amp; Broome" class="Agaricomycetes" family="Bankeraceae" genus="Sarcodon" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Sarcodon thwaitesii" order="Thelephorales" pageId="0" pageNumber="83" phylum="Basidiomycota" rank="species" species="thwaitesii">
<emphasis italics="true" pageId="0" pageNumber="83">Sarcodon thwaitesii</emphasis>
</taxonomicName>
by slightly shorter and decurrent spines, olivaceous tissues in KOH, simple-septate hyphae in all parts of basidiocarp, narrower basidia with shorter sterigmata and smaller basidiospores.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="83" type="materials_examined">
<paragraph pageId="0" pageNumber="83">Type.</paragraph>
<paragraph pageId="0" pageNumber="83">
<emphasis bold="true" pageId="0" pageNumber="83">China.</emphasis>
Yunnan Province, Chuxiong, Zixishan Nat. Res.,
<geoCoordinate degrees="24" direction="north" minutes="58" orientation="latitude" precision="15" seconds="28" value="24.974445">24°58'28&quot;N</geoCoordinate>
,
<geoCoordinate degrees="101" direction="east" minutes="22" orientation="longitude" precision="15" seconds="13" value="101.37028">101°22'13&quot;E</geoCoordinate>
, 2000 m alt., solitary to gregarious, on the ground in
<taxonomicName family="Fagaceae" lsidName="" pageId="0" pageNumber="83" rank="family">Fagaceae</taxonomicName>
forest, 19.07.2018,
<emphasis italics="true" pageId="0" pageNumber="83">Wei 8094</emphasis>
(holotype: IFP 019351).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="83" type="etymology">
<paragraph pageId="0" pageNumber="83">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="83">
<taxonomicName genus="Coactus" lsidName="Coactus" pageId="0" pageNumber="83" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="83">Coactus</emphasis>
</taxonomicName>
(Lat.), refers to the felted pileal surface.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="83" type="description">
<paragraph pageId="0" pageNumber="83">Description.</paragraph>
<paragraph pageId="0" pageNumber="83">Basidiocarps annual, solitary to gregarious, soft and fleshy when fresh, becoming firm and light in weight upon drying; taste none, odour farinaceous when dry. Pileus planar, ellipsoid when young, later round with age, up to 35 mm across and 4-8 mm thick at centre. Pileal surface reddish-brown (8D5) to dark brown (8F8), azonate, pubescent, floccose to felted when fresh, becoming smooth, rugose, scrobiculate when dry; margin white (7A1) when fresh, greyish-brown (7D3) with age, incurved, rarely lobed. Spine surface white (4A1) to yellowish-white (4A2) when fresh, brownish-orange (5C5) to yellowish-brown (5F6) when dry; spines up to 2.1 mm long, base up to 0.3 mm diam., conical, 3-5 per mm, decurrent on stipe, without spines at pileus margin, brittle when dry. Context not duplex, up to 6 mm thick, light brown (5D5), firm; Stipe central, up to 5.5 cm long and 1.3 cm diam., fleshy, greyish-brown (8D3) to violet brown (10F7) when fresh, becoming hollow with age, greyish-orange (5B3) to dark brown (7F7) upon drying, rugous, columniform or attenuate below with bulbous base when old.</paragraph>
<paragraph pageId="0" pageNumber="83">Hyphal structure. Hyphal system monomitic; generative hyphae with simple-septa, CB-, IKI-; tissues olivaceous in KOH.</paragraph>
<paragraph pageId="0" pageNumber="83">
Context. Generative hyphae hyaline, thin-walled, rarely branched, simple-septate, inflated, partly short-celled, interwoven, mostly 4-10
<normalizedToken originalValue="μm">μm</normalizedToken>
diam.
</paragraph>
<paragraph pageId="0" pageNumber="83">
Spines. Tramal hyphae hyaline, thin-walled, frequently branched, more or less parallel along spines, frequently simple-septate, straight, 2-5
<normalizedToken originalValue="μm">μm</normalizedToken>
diam. Cystidia and cystidioles absent. Basidia clavate, thin-walled, with four sterigmata (3.1-5.2
<normalizedToken originalValue="μm">μm</normalizedToken>
long), simple-septate at base, 16.5-50
<normalizedToken originalValue="×">x</normalizedToken>
6.2-9.4
<normalizedToken originalValue="μm">μm</normalizedToken>
; basidioles similar to basidia.
</paragraph>
<paragraph pageId="0" pageNumber="83">
Basidiospores irregular subglobose, brown, thin-walled, tuberculate, CB-, IKI-, (5.1-)5.7-7(-7.1)
<normalizedToken originalValue="×">x</normalizedToken>
(4.6-)4.7-5.9(-6)
<normalizedToken originalValue="μm">μm</normalizedToken>
, Lm = 6.2
<normalizedToken originalValue="μm">μm</normalizedToken>
, Wm = 5.3
<normalizedToken originalValue="μm">μm</normalizedToken>
, Q = 1.17-1.18 (n = 60/2); tuberculi usually isolated or grouped in 2 or more, bi- to trifurcate-like in shape, up to 1.0
<normalizedToken originalValue="μm">μm</normalizedToken>
long.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="83" type="additional specimen examined">
<paragraph pageId="0" pageNumber="83">Additional specimen examined</paragraph>
<paragraph pageId="0" pageNumber="83">
-
<emphasis bold="true" pageId="0" pageNumber="83">China.</emphasis>
Yunnan Province, Maguan County, On the way from Dalishu Township to Damagu Village,
<geoCoordinate degrees="23" direction="north" minutes="4" orientation="latitude" precision="15" seconds="55" value="23.081945">23°4'55&quot;N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="12" orientation="longitude" precision="15" seconds="59" value="104.216385">104°12'59&quot;E</geoCoordinate>
, 1616 m alt., solitary, on the ground in
<taxonomicName family="Fagaceae" lsidName="" pageId="0" pageNumber="83" rank="family">Fagaceae</taxonomicName>
forest, 7.08.2017,
<emphasis italics="true" pageId="0" pageNumber="83">Shi 181</emphasis>
(IFP 019352).
</paragraph>
<caption doi="10.3897/mycokeys.66.49910.figure2" httpUri="https://binary.pensoft.net/fig/397187" pageId="0" pageNumber="83" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="83">
<emphasis bold="true" pageId="0" pageNumber="83">Figure 2.</emphasis>
A basidiocarp of
<taxonomicName authorityName="Y. H. Mu &amp; H. S. Yuan" authorityYear="2020" class="Agaricomycetes" family="Bankeraceae" genus="Sarcodon" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Sarcodon coactus" order="Thelephorales" pageId="0" pageNumber="83" phylum="Basidiomycota" rank="species" species="coactus">
<emphasis italics="true" pageId="0" pageNumber="83">Sarcodon coactus</emphasis>
</taxonomicName>
(holotype: IFP 019351).
</paragraph>
</caption>
<caption doi="10.3897/mycokeys.66.49910.figure3" httpUri="https://binary.pensoft.net/fig/397188" pageId="0" pageNumber="83" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="83">
<emphasis bold="true" pageId="0" pageNumber="83">Figure 3.</emphasis>
SEM of basidiospores of
<taxonomicName authorityName="Y. H. Mu &amp; H. S. Yuan" authorityYear="2020" class="Agaricomycetes" family="Bankeraceae" genus="Sarcodon" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Sarcodon coactus" order="Thelephorales" pageId="0" pageNumber="83" phylum="Basidiomycota" rank="species" species="coactus">
<emphasis italics="true" pageId="0" pageNumber="83">Sarcodon coactus</emphasis>
</taxonomicName>
(holotype: IFP 019351).
</paragraph>
</caption>
<caption doi="10.3897/mycokeys.66.49910.figure4" httpUri="https://binary.pensoft.net/fig/397189" pageId="0" pageNumber="83" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="83">
<emphasis bold="true" pageId="0" pageNumber="83">Figure 4.</emphasis>
Microscopic structures of
<taxonomicName authorityName="Y. H. Mu &amp; H. S. Yuan" authorityYear="2020" class="Agaricomycetes" family="Bankeraceae" genus="Sarcodon" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Sarcodon coactus" order="Thelephorales" pageId="0" pageNumber="83" phylum="Basidiomycota" rank="species" species="coactus">
<emphasis italics="true" pageId="0" pageNumber="83">Sarcodon coactus</emphasis>
</taxonomicName>
(drawn from IFP 019351)
<emphasis bold="true" pageId="0" pageNumber="83">a</emphasis>
basidiospores
<emphasis bold="true" pageId="0" pageNumber="83">b</emphasis>
section of hymenophoral trama with basidia
<emphasis bold="true" pageId="0" pageNumber="83">c</emphasis>
hyphae from pileal context.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>