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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.167.1833" ID-GBIF-Dataset="ff2e62f9-b478-406a-9812-d87c5b0f1275" ID-PMC="PMC3272638" ID-Pensoft-Pub="1313-2970-167-1" ID-PubMed="22363201" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-167-1" ModsDocOrigin="ZooKeys 167" ModsDocTitle="Systematics, conservation and morphology of the spider genus Tayshaneta (Araneae, Leptonetidae) in Central Texas Caves" checkinTime="1451249361926" checkinUser="pensoft" docAuthor="Ledford, Joel, Paquin, Pierre, Cokendolpher, James, Campbell, Josh &amp; Griswold, Charles" docDate="2012" docId="EED39BB8F8548DB6C05FB8340D69022B" docLanguage="en" docName="ZooKeys 167: 1-102" docOrigin="ZooKeys 167" docSource="http://dx.doi.org/10.3897/zookeys.167.1833" docTitle="Tayshaneta archambaulti Ledford, Paquin, Cokendolpher, Campbell &amp; Griswold, 2012, sp. n." docType="treatment" docVersion="4" lastPageNumber="20" masterDocId="FFABFF9EDA67666EBE2E632CFFF08176" masterDocTitle="Systematics, conservation and morphology of the spider genus Tayshaneta (Araneae, Leptonetidae) in Central Texas Caves" masterLastPageNumber="102" masterPageNumber="1" pageNumber="19" updateTime="1668153202787" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systematics, conservation and morphology of the spider genus Tayshaneta (Araneae, Leptonetidae) in Central Texas Caves</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ledford, Joel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Paquin, Pierre</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cokendolpher, James</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Campbell, Josh</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Griswold, Charles</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>167</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>102</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.167.1833</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.167.1833</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-167-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152033156" LSID="urn:lsid:zoobank.org:act:0A3E7B18-3AB5-44F9-AF73-4623ABDD215C" httpUri="http://treatment.plazi.org/id/EED39BB8F8548DB6C05FB8340D69022B" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection pageId="18" pageNumber="19" type="nomenclature">
<paragraph pageId="18" pageNumber="19">
<taxonomicName LSID="urn:lsid:zoobank.org:act:0A3E7B18-3AB5-44F9-AF73-4623ABDD215C" class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta archambaulti" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="archambaulti">Tayshaneta archambaulti</taxonomicName>
<taxonomicNameLabel pageId="18" pageNumber="19">sp. n.</taxonomicNameLabel>
Figs 13
<normalizedToken originalValue="AF34AF">A-F34A-F</normalizedToken>
52C58
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="type data">
<paragraph pageId="18" pageNumber="19">Type data.</paragraph>
<paragraph pageId="18" pageNumber="19">
Male holotype from Grapevine Cave, 7 miles west of Wimberly, Hays County, Texas, 18-Nov-2009, J. Ledford, K.
<normalizedToken originalValue="OConnor">O'Connor</normalizedToken>
,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="30.04">30.04N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="555" value="-98.22">98.22W</geoCoordinate>
, (CASC).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="etymology">
<paragraph pageId="18" pageNumber="19">Etymology.</paragraph>
<paragraph pageId="18" pageNumber="19">This species is named in honor of Martin Archambault, fellow caver and friend who helped collect many leptonetids in Texas and Mexico.</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="other material examined">
<paragraph pageId="18" pageNumber="19">Other material examined.</paragraph>
<paragraph pageId="18" pageNumber="19">
USA: Hays County: Burnett Ranch Cave, 7mi. W. of Wimberly, 1982, A. Grubbs,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="30.02">30.02N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="555" value="-98.21">98.21W</geoCoordinate>
, 1♂, 2 ♀, 3 juvs (AMNH); Grapevine Cave, 7mi. W. of Wimberly, 26-May-1989, A. Grubbs,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="30.04">30.04N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="555" value="-98.22">98.22W</geoCoordinate>
, 1♂ (AMNH); Grapevine Cave, 7mi. W. of Wimberly, 23-April-1995, A. Grubbs, Vreeland,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="30.04">30.04N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="555" value="-98.22">98.22W</geoCoordinate>
, 1♂, 1 ♀, 5 juvs, (TMM);Grapevine Cave, 7mi. W. of Wimberly, 18-November-2009, J. Ledford, K.
<normalizedToken originalValue="OConnor">O'Connor</normalizedToken>
,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="30.04">30.04N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="555" value="-98.22">98.22W</geoCoordinate>
, 4♂, 5 ♀, 7 juvs, (TMM).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="diagnosis">
<paragraph pageId="18" pageNumber="19">Diagnosis.</paragraph>
<paragraph pageId="18" pageNumber="19">
<taxonomicName class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta archambaulti" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="archambaulti">Tayshaneta archambaulti</taxonomicName>
can be separated from all
<taxonomicName class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Tayshaneta</taxonomicName>
species that lack a ventral sclerite, except
<taxonomicName class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta coeca" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="coeca">Tayshaneta coeca</taxonomicName>
and
<taxonomicName class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta devia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="devia">Tayshaneta devia</taxonomicName>
, by the following combination of characters: embolus oval to quadrate, lacking sculpture along its margin (E, Fig. 34D); retrolateral tibial spine short, occupying less than 0.50
<normalizedToken originalValue="×">x</normalizedToken>
the length of the palpal tarsus (RTS, Fig. 34A). Separated from
<taxonomicName class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta devia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="devia">Tayshaneta devia</taxonomicName>
by having a retrolateral tibial spine with sculpture along its entire length and from
<taxonomicName class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta coeca" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="coeca">Tayshaneta coeca</taxonomicName>
by having the embolus curved distally and extending beyond the apical portion of the bulb (E, Fig. 34E).
</paragraph>
</subSubSection>
<subSubSection lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph pageId="18" pageNumber="19">Description.</paragraph>
<paragraph pageId="18" pageNumber="19">
Male (holotype). Body length 1.35, carapace 0.64 long, 0.56 wide, length 1.13
<normalizedToken originalValue="×">x</normalizedToken>
width. Carapace light brown, eyes reduced, sparsely setose (Figs 13
<normalizedToken originalValue="AF">A-F</normalizedToken>
). Legs elongate and thin, femur I 1.5
<normalizedToken originalValue="×">x</normalizedToken>
carapace length, covered in fine setae with few scattered spines.Palpal tarsus entire, tapering apically; retrolateral tibial spine weakly recurved, on weakly elevated base, sculptured throughout, length 0.32
<normalizedToken originalValue="×">x</normalizedToken>
tarsus length (RTS, Fig. 34A). Bulb suboval, length 1.71
<normalizedToken originalValue="×">x</normalizedToken>
width; embolus oval to quadrate (E, Fig. 34D), length 2.0
<normalizedToken originalValue="×">x</normalizedToken>
width. Abdomen pale to yellow-brown, without pattern, 0.70 long, 0.54 wide, covered in fine setae.
</paragraph>
<caption pageId="19" pageNumber="20">
<paragraph pageId="19" pageNumber="20">
<pageBreakToken pageId="19" pageNumber="20" start="start">Figure</pageBreakToken>
13.
<taxonomicName class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta archambaulti" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="archambaulti">Tayshaneta archambaulti</taxonomicName>
sp. n., Burnett Ranch Cave, Hays County, Texas (AMNH), habitus. A
<taxonomicName class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta archambaulti" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="archambaulti">Tayshaneta archambaulti</taxonomicName>
male, dorsal B
<taxonomicName class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta archambaulti" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="archambaulti">Tayshaneta archambaulti</taxonomicName>
male, ventral C
<taxonomicName class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta archambaulti" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="archambaulti">Tayshaneta archambaulti</taxonomicName>
male, lateral D
<taxonomicName class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta archambaulti" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="archambaulti">Tayshaneta archambaulti</taxonomicName>
female, dorsal E
<taxonomicName class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta archambaulti" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="archambaulti">Tayshaneta archambaulti</taxonomicName>
female, ventral F
<taxonomicName class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta archambaulti" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="archambaulti">Tayshaneta archambaulti</taxonomicName>
female, lateral.
</paragraph>
</caption>
<paragraph pageId="19" pageNumber="20">
Variation(n = 4). Total length 1.34-1.63; carapace length 0.88-1.36
<normalizedToken originalValue="×">x</normalizedToken>
carapace width; length femur I 1.35-1.65
<normalizedToken originalValue="×">x</normalizedToken>
carapace width.
</paragraph>
<paragraph pageId="19" pageNumber="20">
Female (Grapevine Cave). Body length 1.32, carapace 0.63 long, 0.51 wide, length 1.25
<normalizedToken originalValue="×">x</normalizedToken>
width. Pigmentation and setation same as for male (Figs 13
<normalizedToken originalValue="DF">D-F</normalizedToken>
). Legs elongate and thin, femur I 1.34
<normalizedToken originalValue="×">x</normalizedToken>
carapace length, covered in fine setae with few scattered spines.Atrium oval, length 0.51
<normalizedToken originalValue="×">x</normalizedToken>
width, spermathecae with twisted stalks and large, circular heads (Fig. 52C). Abdomen pale to yellow-brown, without pattern, 0.69 long, 0.54 wide, covered in fine setae.
</paragraph>
<paragraph pageId="19" pageNumber="20">
Variation (n = 4). Total length 1.32-1.72; carapace length 1.20-1.29
<normalizedToken originalValue="×">x</normalizedToken>
carapace width; length femur I 1.34-1.75
<normalizedToken originalValue="×">x</normalizedToken>
carapace width.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="distribution">
<paragraph pageId="19" pageNumber="20">Distribution.</paragraph>
<paragraph pageId="19" pageNumber="20">This species is known only from Burnett Ranch Cave and Grapevine Cave in southwestern Hays County (Fig. 58).</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="natural history">
<paragraph pageId="19" pageNumber="20">Natural History.</paragraph>
<paragraph pageId="19" pageNumber="20">
Individuals for this species were collected throughout Grapevine Cave, however, most specimens were encountered at the base of the
<normalizedToken originalValue="caves">cave's</normalizedToken>
vertical entrance in the twilight area under stones. They were collected in fine sheet webs similar to other
<taxonomicName class="Arachnida" family="Leptonetidae" genus="Tayshaneta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tayshaneta" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Tayshaneta</taxonomicName>
species.
</paragraph>
</subSubSection>
</treatment>
</document>