141 lines
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141 lines
14 KiB
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<document ID-CLB-Dataset="275942" ID-DOI="http://dx.doi.org/10.3897/asp.81.e107579" ID-GBIF-Dataset="742ed870-c756-4a81-8324-e95c9808643a" ID-Pensoft-Pub="1864-8312-81-819" ID-Pensoft-UUID="70148F72D847500A930DA76D2196759A" ID-ZooBank="50EE94AAD90A43E1B54E4FADE22F4668" ModsDocID="1864-8312-81-819" checkinTime="1698951166303" checkinUser="pensoft" docAuthor="Ge, Si-Xun, Jiang, Zhuo-Heng, Ren, Li-Li, van Achterberg, Cornelis & Tan, Jiang-Li" docDate="2023" docId="7D4FE0AD08A45A36BE76477444A171BA" docLanguage="en" docName="ArthropodSystPhyl 81: 819-844" docOrigin="Arthropod Systematics & amp; Phylogeny 81" docPubDate="2023-11-02" docSource="http://dx.doi.org/10.3897/asp.81.e107579" docTitle="Aphanostephanus Ge and Tan 2023, gen. nov." docType="treatment" docUuid="10114E82-8813-4751-8F3B-98F4B5BD1E79" docUuidSource="ZooBank" docVersion="2" id="70148F72D847500A930DA76D2196759A" lastPageNumber="819" masterDocId="70148F72D847500A930DA76D2196759A" masterDocTitle="New insights into the phylogeny of Stephanidae (Hymenoptera: Apocrita), with a revision of the fossil species" masterLastPageNumber="844" masterPageNumber="819" pageNumber="819" updateTime="1698998894021" updateUser="ExternalLinkService">
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<mods:titleInfo id="06D0B65699AB3E8C1191FFDADC96C3D9">
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<mods:title id="2FADF8E5B287B4B25BB013DFBC790A25">New insights into the phylogeny of Stephanidae (Hymenoptera: Apocrita), with a revision of the fossil species</mods:title>
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<mods:namePart id="9F38AE04ECAA9B6D8EBB1E140D5CE163">Ge, Si-Xun</mods:namePart>
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<mods:nameIdentifier id="9887D26353765C7FB65DDC0C56860A88" type="ORCID">https://orcid.org/0000-0003-3769-1530</mods:nameIdentifier>
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<mods:affiliation id="574F35E9713E4D0D64773F53D852ECC8">College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="6EF9E2F52FEF915680069EF6C78EFF59">Author</mods:roleTerm>
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<mods:namePart id="5976FE382A4F87D53F361E77DA0F50EF">Jiang, Zhuo-Heng</mods:namePart>
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<mods:affiliation id="15F18466E593305C9B531A626328E5A4">School of Science, Westlake University, Hangzhou, China</mods:affiliation>
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</mods:name>
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<mods:name id="6B9FF7C41698953526A293A1F868703F" type="personal">
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<mods:roleTerm id="2E93661D9BBF422142360F07B4ABF5DA">Author</mods:roleTerm>
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<mods:namePart id="D74D74FA89C59D8D257B7A36B8E2662C">Ren, Li-Li</mods:namePart>
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<mods:affiliation id="10627E1E17E2B5505C6AFCACD3323D4D">College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
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<mods:nameIdentifier id="2AA2546D00B85B5AE83037CAD71777BE" type="email">lily_ren@bjfu.edu.cn</mods:nameIdentifier>
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<mods:name id="0F6381A1ABA5E631A20D9EB2011BB7A9" type="personal">
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</mods:role>
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<mods:namePart id="901A8E5352B3495616326BE17F2016EF">van Achterberg, Cornelis</mods:namePart>
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<mods:nameIdentifier id="52D7DE232AF387CECC4FD88431072ABA" type="ORCID">https://orcid.org/0000-0002-6495-4853</mods:nameIdentifier>
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<mods:affiliation id="654A1B5FC754D843C130E872D9CF6687">Shaanxi Key Laboratory for Animal Conservation / Key Laboratory of Resource Biology and Biotechnology in Western China, College of Life Sciences, Northwest University, 229 North Taibai Road, Xi' an, Shaanxi 710069, China</mods:affiliation>
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<mods:nameIdentifier id="27AD461882AE24DF25FB59B241A8B862" type="email">kees@vanachterberg.org</mods:nameIdentifier>
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<mods:namePart id="4B165A3BB87A9166106C10673FEF9015">Tan, Jiang-Li</mods:namePart>
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<mods:affiliation id="747CC5B68AF76898C269F76DE1B6FF0A">Shaanxi Key Laboratory for Animal Conservation / Key Laboratory of Resource Biology and Biotechnology in Western China, College of Life Sciences, Northwest University, 229 North Taibai Road, Xi' an, Shaanxi 710069, China</mods:affiliation>
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<mods:nameIdentifier id="8FAEEEE5555713FBE662E7065ED2EB84" type="email">tanjiangli@sina.com</mods:nameIdentifier>
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<mods:typeOfResource id="6CFA1126280D009200EC91ECCBBC856A">text</mods:typeOfResource>
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<mods:titleInfo id="A89AD8DB01AA672906864CE4F3E94CB7">
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<mods:title id="44515CE9476A0B7B1052F82A4F1008FB">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:part id="360CB74DB062344C62F453980E1992C6">
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<mods:date id="DDA1A2232BF109401F7B620BE5F9F0CB">2023</mods:date>
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<mods:number id="6D2B7EFB9A160F8A3142201FE9FD1A93">2023-11-02</mods:number>
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<mods:detail id="1AE784C33FBC3EB341F88269179D7576" type="volume">
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<mods:number id="5B4F71D9568414F02C70D39E9165FB3C">81</mods:number>
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<mods:start id="3FC7800CED03F8BA3EB76B9757AC45D6">819</mods:start>
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<mods:end id="958E89B225ED91E4475EEAA856953961">844</mods:end>
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<mods:url id="8E9977B243C8E425BFF06E7C536CF6E5">http://dx.doi.org/10.3897/asp.81.e107579</mods:url>
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<mods:classification id="E12002EFF44F289490353FF1404052BB">journal article</mods:classification>
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<mods:identifier id="CA002A1EB30916DE4736844C03941D10" type="DOI">http://dx.doi.org/10.3897/asp.81.e107579</mods:identifier>
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<mods:identifier id="B87D943CB406092CC7FA5E43B8AA74D3" type="Pensoft-Pub">1864-8312-81-819</mods:identifier>
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<treatment id="7D4FE0AD08A45A36BE76477444A171BA" ID-GBIF-Taxon="216248472" LSID="urn:lsid:zoobank.org:act:10114E82-8813-4751-8F3B-98F4B5BD1E79" httpUri="http://treatment.plazi.org/id/7D4FE0AD08A45A36BE76477444A171BA" lastPageNumber="819" pageId="0" pageNumber="819">
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<subSubSection id="8BB59167C6A4A7C51369928C1A49D787" pageId="0" pageNumber="819" type="nomenclature">
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<paragraph id="FFDDD3C75F8082182951F4ED47BF8A49" pageId="0" pageNumber="819">
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Genus †
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<taxonomicName id="7F51FC65339C0D66FE598EF6ABC3DCA4" LSID="https://zoobank.org/10114E82-8813-4751-8F3B-98F4B5BD1E79" authority="Ge and Tan" authorityName="Ge and Tan" authorityYear="2023" class="Magnoliopsida" family="Stephanidae" genus="Aphanostephanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aphanostephanus" order="Hymenoptera" pageId="0" pageNumber="819" phylum="Tracheophyta" rank="genus" status="gen. nov.">Aphanostephanus Ge and Tan</taxonomicName>
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<taxonomicNameLabel id="B352FC60523ABBD3B271C759417F46C5" pageId="0" pageNumber="819">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="E50C304E9E989998FC8AD6ABAE34180A" pageId="0" pageNumber="819" type="type species">
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<paragraph id="6316AFF8F853576F1935D7B6BC95D2B0" pageId="0" pageNumber="819">Type species.</paragraph>
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<paragraph id="8ECF66F2A72C3FADE3A30CE1E9D33C82" pageId="0" pageNumber="819">
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†
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<taxonomicName id="13D6185853252D145F6392978DA92D8E" authorityName="Engel" authorityYear="2005" class="Insecta" family="Stephanidae" genus="Electrostephanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Electrostephanus janzeni" order="Hymenoptera" pageId="0" pageNumber="819" phylum="Arthropoda" rank="species" species="janzeni">
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<emphasis id="08ECB58CCA1467E15B49BB4110F120A2" italics="true" pageId="0" pageNumber="819">Electrostephanus janzeni</emphasis>
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</taxonomicName>
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Engel, 2005.
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</paragraph>
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</subSubSection>
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<subSubSection id="F037E9AD954C5B9F9136B5818AF5ED7C" pageId="0" pageNumber="819" type="diagnosis">
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<paragraph id="A7CA3AFA793DADDD74AFE5CA449D5322" pageId="0" pageNumber="819">Diagnosis.</paragraph>
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<paragraph id="6A5CADD015C0888E3546740D42E2AA66" pageId="0" pageNumber="819">Head globose, compound eyes comparatively small, occupying less than half portion of lateral surface. Pronotum elongated with neck differentiated. Forewing with vein 2Rs+M rather short, directly connecting to veins 2-Rs and apical abscissa of vein M without incisions; vein 2Cua pigmented and vein 2Cub absent. Metasoma with tergum I and sternum I not fused laterally, tergite I short and robust, about as long as tergite II. Ovipositor almost as long as body length.</paragraph>
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</subSubSection>
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<subSubSection id="85DC8AA61CDD5DF1B153B0003DE44BD0" pageId="0" pageNumber="819" type="etymology">
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<paragraph id="583410016C3348DCEF54F93A27D0600A" pageId="0" pageNumber="819">Etymology.</paragraph>
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<paragraph id="23E34C2A9408C56FDB78417CA3ECC2BE" pageId="0" pageNumber="819">
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From
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<normalizedToken id="4CF12B740F966868617D75B02E970B48" originalValue="“Aphanes”">"Aphanes"</normalizedToken>
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(Greek for
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<normalizedToken id="C1A7167D68D8577C6CB9FD349D2FA2D7" originalValue="“vague”">"vague"</normalizedToken>
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) and the generic name
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<taxonomicName id="1020476B42D8F8B3DAECF12296D1EB45" class="Insecta" family="Stephanidae" genus="Stephanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stephanus" order="Hymenoptera" pageId="0" pageNumber="819" phylum="Arthropoda" rank="genus">
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<emphasis id="E7F9CECA90FC1FFC331B99417C70928C" italics="true" pageId="0" pageNumber="819">Stephanus</emphasis>
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</taxonomicName>
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Jurine. The name refers to the puzzling position of the genus within a family. The gender of the name is masculine.
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</paragraph>
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</subSubSection>
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<subSubSection id="D3A2C89942564DCB983FBE8E4453C455" pageId="0" pageNumber="819" type="remarks">
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<paragraph id="8EE967F74F4858587051041AC8AF10D6" pageId="0" pageNumber="819">Remarks.</paragraph>
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<paragraph id="03D52F573F937848BC4FDDA12F59FEF4" pageId="0" pageNumber="819">
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In the original description by
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<bibRefCitation id="1566CC5995550A84F7C49E95DF6DCFB6" author="Engel, MS" journalOrPublisher="Polskie Pismo Entomologiczne" pageId="0" pageNumber="819" pagination="317 - 332" refId="B17" refString="Engel, MS, 2005. The crown wasp Electrostephanus (Hymenoptera: Stephanidae): discovery of the female and a new species. Polskie Pismo Entomologiczne 74 (3): 317 - 332" title="The crown wasp Electrostephanus (Hymenoptera: Stephanidae): discovery of the female and a new species." volume="74" year="2005">Engel (2005)</bibRefCitation>
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, the type species was indicated to be similar to †
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<taxonomicName id="249B73FF0153BBABD1A99D482FF40FDB" family="Stephanidae" genus="Neurastephanus" kingdom="Animalia" lsidName="Neurastephanus neovenatus" order="Hymenoptera" pageId="0" pageNumber="819" rank="species" species="neovenatus">
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<emphasis id="1A78CE70A1D279C3B87F87EBAAC1D0BC" italics="true" pageId="0" pageNumber="819">Neurastephanus neovenatus</emphasis>
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</taxonomicName>
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<emphasis id="1C20DB053332EA43DF2F79DFFDAA7199" bold="true" pageId="0" pageNumber="819">comb. nov.</emphasis>
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which is a species here considered to belong to †
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<taxonomicName id="55CB5AF1BF0B377197EC29FC1F176B86" authorityName="Ge and Tan" authorityYear="2023" family="Stephanidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="819" rank="subFamily" subFamily="Lagenostephaninae">Lagenostephaninae</taxonomicName>
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<emphasis id="3BDDE8C50C4512B5C2FAFAA5BB2ED5F1" bold="true" pageId="0" pageNumber="819">subf. nov.</emphasis>
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However, it differs from †
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<taxonomicName id="4224F23EA112DE26C3A71180E600013B" family="Stephanidae" kingdom="Animalia" lsidName="N. neovenatus" order="Hymenoptera" pageId="0" pageNumber="819" rank="species" species="neovenatus">
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<emphasis id="3E05ED9CBF6A309C3A417671CB555E44" italics="true" pageId="0" pageNumber="819">N. neovenatus</emphasis>
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</taxonomicName>
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in having vein 1-M straight; vein 2Rs + M short, directly connected to vein 2-Rs, and the apical abscissa of vein M. In addition, the paratype female also showed its ovipositor sheath about as long as the body length (in †
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<taxonomicName id="01A28F1BBDDE3DCF6128EEBF613455BD" authorityName="Ge and Tan" authorityYear="2023" family="Stephanidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="819" rank="subFamily" subFamily="Lagenostephaninae">Lagenostephaninae</taxonomicName>
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as long as the metasoma). Combining our phylogenetic results, this taxa showed its lineage sister to †
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<taxonomicName id="ECDB6C3C3400DCDD295B04D798F2AC9F" authorityName="Ge and Tan" authorityYear="2023" family="Stephanidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="819" rank="subFamily" subFamily="Lagenostephaninae">Lagenostephaninae</taxonomicName>
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+
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<taxonomicName id="B7E83EC7BB923ED29C5921553431358C" authorityName="Orfila" authorityYear="1949" lsidName="" pageId="0" pageNumber="819" rank="subFamily" subFamily="Schlettereriinae">Schlettereriinae</taxonomicName>
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, however, there is no synapomorphies or characteristics with significant evolutionary significance that can define it as a distinct group; thus, we prudently treated the genus as
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<emphasis id="46F60BAC68509DD1FA7194308D42ADDE" italics="true" pageId="0" pageNumber="819">incertae sedis</emphasis>
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in the
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<taxonomicName id="F2B24C4A39157CDB5E13F54A35F75D15" class="Insecta" family="Stephanidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="819" phylum="Arthropoda" rank="family">Stephanidae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |