197 lines
20 KiB
XML
197 lines
20 KiB
XML
<document id="9586055BBCEB2AF8F99C41FBC918E69B" ID-DOI="10.1111/j.1096-3642.2007.00306.x" ID-ISSN="0024-4082" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630428936140" checkinUser="felipe" docAuthor="De Pinna, Mário C. C., Ferraris, Carl J. & Vari, Richard P." docDate="2007" docId="03F987EDCD22FFACFCA12B8B1A0CEB67" docLanguage="en" docName="j.1096-3642.2007.00306.x.pdf" docOrigin="Zoological Journal of the Linnean Society 150 (4)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2007.00306.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Cetopsinae Bleeker 1858" docType="treatment" docVersion="2" lastPageNumber="802" masterDocId="FFC0FF95CD0CFF83FFD02876187CE81B" masterDocTitle="A phylogenetic study of the neotropical catfish family Cetopsidae (Osteichthyes, Ostariophysi, Siluriformes), with a new classification" masterLastPageNumber="813" masterPageNumber="755" pageNumber="801" updateTime="1699266303805" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo id="59909855E3704F8BCAD3699F03C38D3A">
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<mods:title id="B8BE1614725B680140568A0904A743C4">A phylogenetic study of the neotropical catfish family Cetopsidae (Osteichthyes, Ostariophysi, Siluriformes), with a new classification</mods:title>
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<mods:namePart id="7462AA8DBA716FF0337515B34F9F0DA2">De Pinna, Mário C. C.</mods:namePart>
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<mods:namePart id="79222D07CEB7129E7EF608D3C1809631">Ferraris, Carl J.</mods:namePart>
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<mods:namePart id="270FA019B63AD56A22669776D52B9FC0">Vari, Richard P.</mods:namePart>
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<mods:typeOfResource id="D8246EEA5DAE2D02293F9D73D191FA7A">text</mods:typeOfResource>
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<mods:title id="7DAFACC6EAA9FF8D152983658EF1E4F0">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="B186ADCA3947F4191E5249403855A136">
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<mods:date id="61C94AEBB36ADFB52AA652BF7536A5C7">2007</mods:date>
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<mods:number id="81FEDD4C432D4184CA9BEA9F02878CBD">2007-08-31</mods:number>
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<mods:number id="8ADEA05D3EE09296AC791C7F6946346C">150</mods:number>
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<mods:detail id="D4B7DC4840C54CF4B8F23A4BB9DED3D8" type="issue">
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<mods:url id="21536A2D7B8E2C6761DF59B6E135F371">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2007.00306.x</mods:url>
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<mods:classification id="D0D92C289F32E299B085F11D7E774FDE">journal article</mods:classification>
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<mods:identifier id="F7F2F95161321388F6BCF97E08472553" type="DOI">10.1111/j.1096-3642.2007.00306.x</mods:identifier>
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<mods:identifier id="FEF67F2925EB0E0B530A1329FCE3A3E9" type="ISSN">0024-4082</mods:identifier>
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</mods:mods>
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<treatment id="03F987EDCD22FFACFCA12B8B1A0CEB67" LSID="urn:lsid:plazi:treatment:03F987EDCD22FFACFCA12B8B1A0CEB67" httpUri="http://treatment.plazi.org/id/03F987EDCD22FFACFCA12B8B1A0CEB67" lastPageId="47" lastPageNumber="802" pageId="46" pageNumber="801">
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<subSubSection id="C34A6570CD22FFADFCA12B8B1D3FEC0E" box="[881,1347,1021,1045]" pageId="46" pageNumber="801" type="nomenclature">
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<paragraph id="8BEF36FBCD22FFADFCA12B8B1D3FEC0E" blockId="46.[881,1347,1021,1045]" box="[881,1347,1021,1045]" pageId="46" pageNumber="801">
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<heading id="D0A78197CD22FFADFCA12B8B1D3FEC0E" box="[881,1347,1021,1045]" centered="true" fontSize="9" level="2" pageId="46" pageNumber="801" reason="5">
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SUBFAMILY
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<taxonomicName id="4C504D78CD22FFADFC2B2B8B1D3FEC0E" authority="BLEEKER, 1858" authorityName="Bleeker" authorityYear="1858" box="[1019,1347,1021,1045]" class="Actinopterygii" family="Cetopsidae" kingdom="Animalia" order="Siluriformes" pageId="46" pageNumber="801" phylum="Chordata" rank="subFamily" subFamily="Cetopsinae">CETOPSINAE BLEEKER, 1858</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A6570CD22FFACFCF62C581A0CEB67" lastPageId="47" lastPageNumber="802" pageId="46" pageNumber="801" type="materials_examined">
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<paragraph id="8BEF36FBCD22FFADFCF62C581D58EC59" blockId="46.[806,1316,1069,1091]" box="[806,1316,1069,1091]" pageId="46" pageNumber="801">
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<emphasis id="B924EAE9CD22FFADFCF62C581BD8EC58" box="[806,932,1070,1091]" italics="true" pageId="46" pageNumber="801">
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<typeStatus id="54EB8859CD22FFADFCF62C581B27EC58" box="[806,859,1070,1091]" pageId="46" pageNumber="801">Type</typeStatus>
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genus
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</emphasis>
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:
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<taxonomicName id="4C504D78CD22FFADFC632C581D58EC59" authority="Spix and Agassiz, 1829" authorityName="Spix and Agassiz" authorityYear="1829" box="[947,1316,1069,1091]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="46" pageNumber="801" phylum="Chordata" rank="genus">
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<emphasis id="B924EAE9CD22FFADFC632C581C6DEC58" box="[947,1041,1070,1091]" italics="true" pageId="46" pageNumber="801">Cetopsis</emphasis>
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Spix and Agassiz, 1829
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</taxonomicName>
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</paragraph>
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<paragraph id="8BEF36FBCD22FFADFCF62C2A1CD9EC6A" blockId="46.[806,1422,1115,1904]" box="[806,1189,1115,1137]" pageId="46" pageNumber="801">
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<emphasis id="B924EAE9CD22FFADFCF62C2A1BEAEC6A" box="[806,918,1116,1137]" italics="true" pageId="46" pageNumber="801">Diagnosis</emphasis>
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(Br:> 5, jk: 99, bs: 99):
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</paragraph>
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<paragraph id="8BEF36FBCD22FFADFCEF2C0C1CD8ECF7" blockId="46.[806,1422,1115,1904]" pageId="46" pageNumber="801">Single row of dentary teeth, or more than one row of teeth present only along approximately one-half of length of tooth-bearing surface, plus some additional teeth located near symphysis (4);</paragraph>
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<paragraph id="8BEF36FBCD22FFADFCEF2C831D17ED32" blockId="46.[806,1422,1115,1904]" pageId="46" pageNumber="801">Adductor mandibulae muscle hypertrophied, extending dorsally to midline of neurocranium (7);</paragraph>
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<paragraph id="8BEF36FBCD22FFADFCEF2D451BB4ED9E" blockId="46.[806,1422,1115,1904]" pageId="46" pageNumber="801">
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<emphasis id="B924EAE9CD22FFADFCEF2D451C38ED5C" box="[831,1092,1330,1352]" italics="true" pageId="46" pageNumber="801">Levator arcus palatini</emphasis>
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muscle very developed and clearly visible in dorsal view (reversed in
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<taxonomicName id="4C504D78CD22FFADFAE02D271BF8ED9F" baseAuthorityName="Lundberg & Rapp Py-Daniel" baseAuthorityYear="1994" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="46" pageNumber="801" phylum="Chordata" rank="species" species="oliveirai">
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<emphasis id="B924EAE9CD22FFADFAE02D271BF8ED9F" italics="true" pageId="46" pageNumber="801">Cetopsis oliveirai</emphasis>
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</taxonomicName>
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) (16);
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</paragraph>
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<paragraph id="8BEF36FBCD22FFADFCEF2DF81C93EDFA" blockId="46.[806,1422,1115,1904]" pageId="46" pageNumber="801">Dorsal portion of hyomandibula with distinct anterior process forming long, direct articulation with ventrolateral margin of neurocranium (17);</paragraph>
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<paragraph id="8BEF36FBCD22FFADFCEF2D9C1BC3EE04" blockId="46.[806,1422,1115,1904]" pageId="46" pageNumber="801">Metapterygoid extending posteromedial of entopterygoid (21);</paragraph>
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<paragraph id="8BEF36FBCD22FFADFCEF2E511D6AEE26" blockId="46.[806,1422,1115,1904]" box="[831,1302,1575,1597]" pageId="46" pageNumber="801">Vomerine teeth disposed in one row (39);</paragraph>
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<paragraph id="8BEF36FBCD22FFADFCEF2E301BCAEEA3" blockId="46.[806,1422,1115,1904]" pageId="46" pageNumber="801">Rigid layer of fibrous cartilage associated with anterolateral margins of mesethmoid, forming protective wall surrounding anteromedial portions of olfactory capsule (52);</paragraph>
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<paragraph id="8BEF36FBCD22FFADFCEF2EB71CE9EEEE" blockId="46.[806,1422,1115,1904]" pageId="46" pageNumber="801">
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Distal portion of posterior ceratohyal broadened (78, reversed in
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<taxonomicName id="4C504D78CD22FFADFC7F2E961CFAEEEF" authorityName="Ferraris, ANSP" authorityYear="1996" box="[943,1158,1759,1781]" class="Actinopterygii" family="Cetopsidae" genus="Denticetopsis" kingdom="Animalia" order="Siluriformes" pageId="46" pageNumber="801" phylum="Chordata" rank="species" species="sauli">
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<emphasis id="B924EAE9CD22FFADFC7F2E961CFAEEEF" box="[943,1158,1759,1781]" italics="true" pageId="46" pageNumber="801">Denticetopsis sauli</emphasis>
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</taxonomicName>
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);
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</paragraph>
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<paragraph id="8BEF36FBCD22FFADFCEF2E881C13EF28" blockId="46.[806,1422,1115,1904]" pageId="46" pageNumber="801">Third and fourth branchiostegal rays distinctly shorter than other rays (82);</paragraph>
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<paragraph id="8BEF36FBCD22FFADFCEF2F4D1B86EF6B" blockId="46.[806,1422,1115,1904]" pageId="46" pageNumber="801">Coracoids extending for two-thirds or less of length of cleithrum (102);</paragraph>
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<paragraph id="8BEF36FBCD23FFACFF6B28B51A4AE90C" blockId="47.[162,778,195,892]" pageId="47" pageNumber="802">Contralateral posterior basipterygial cartilages fused medially at base, forming single stem for posterior cartilaginous process(es) (110);</paragraph>
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<paragraph id="8BEF36FBCD23FFACFF6B29691A09E969" blockId="47.[162,778,195,892]" pageId="47" pageNumber="802">
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Lateral branch of basal bifurcation of posterior basipterygial cartilage ax-head shaped (112, State 2, reversed to state 0 in
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<taxonomicName id="4C504D78CD23FFACFE71292B1A19E969" baseAuthorityName="Lundberg & Rapp Py-Daniel" baseAuthorityYear="1994" box="[417,613,349,370]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="oliveirai">
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<emphasis id="B924EAE9CD23FFACFE71292B1A19E969" box="[417,613,349,370]" italics="true" pageId="47" pageNumber="802">Cetopsis oliveirai</emphasis>
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</taxonomicName>
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);
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</paragraph>
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<paragraph id="8BEF36FBCD23FFACFF6B290D1AB8E9D5" blockId="47.[162,778,195,892]" pageId="47" pageNumber="802">
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First pelvic-fin ray base segments orientated obliquely, with distinct laminar expansion (115, State 2, reversed to State
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<quantity id="4CA89B1ECD23FFACFE5B29CF19C4E9D4" box="[395,440,441,463]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="47" pageNumber="802" unit="in" value="1.0">1 in</quantity>
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<taxonomicName id="4C504D78CD23FFACFE1129CF1AC9E9D5" authorityName="INHS" baseAuthorityName="Fernandez-Yepez" baseAuthorityYear="1972" box="[449,693,441,462]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="morenoi">
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<emphasis id="B924EAE9CD23FFACFE1129CF1AC9E9D5" box="[449,693,441,462]" italics="true" pageId="47" pageNumber="802">Cetopsidium morenoi</emphasis>
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</taxonomicName>
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);
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</paragraph>
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<caption id="DF2F6673CD23FFACFCEA29D41C09E9C9" pageId="47" pageNumber="802" startId="47.[826,904,418,437]" targetBox="[832,1435,199,380]" targetPageId="47">
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<paragraph id="8BEF36FBCD23FFACFCEA29D41C09E9C9" blockId="47.[826,1442,418,466]" pageId="47" pageNumber="802">
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<emphasis id="B924EAE9CD23FFACFCEA29D41BD3E9AE" bold="true" box="[826,943,418,437]" pageId="47" pageNumber="802">Figure 35.</emphasis>
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<taxonomicName id="4C504D78CD23FFACFC6B29D41C8BE9AE" authority=", ANSP" authorityName="ANSP" box="[955,1271,418,437]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="minutum">
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<emphasis id="B924EAE9CD23FFACFC6B29D41CD4E9AF" box="[955,1192,418,437]" italics="true" pageId="47" pageNumber="802">Cetopsidium minutum</emphasis>
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, ANSP
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</taxonomicName>
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175839, 21 mm SL; Guyana, Essequibo River.
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</paragraph>
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</caption>
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<paragraph id="8BEF36FBCD23FFACFF6B29A11A08EA31" blockId="47.[162,778,195,892]" pageId="47" pageNumber="802">
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Pelvic fins with medial membrane joining contralateral medial rays and abdomen (117, State 1 or 2, reversed to State 0 in
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<taxonomicName id="4C504D78CD23FFACFE752A631A19EA31" baseAuthorityName="Spix and Agassiz" baseAuthorityYear="1829" box="[421,613,533,554]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="candiru">
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<emphasis id="B924EAE9CD23FFACFE752A631A19EA31" box="[421,613,533,554]" italics="true" pageId="47" pageNumber="802">Cetopsis candiru</emphasis>
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</taxonomicName>
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);
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</paragraph>
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<paragraph id="8BEF36FBCD23FFACFF6B2A421B74EB24" blockId="47.[162,778,195,892]" pageId="47" pageNumber="802">
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Last anal-fin ray and caudal peduncle united by membranous connection extending at least to basal one-half of ray and may extend along entire length of ray (119, reversed to State 0 in species of
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<taxonomicName id="4C504D78CD23FFACFDA52AE61B7FEABE" authorityName="Bleeker" authorityYear="1862" box="[629,771,656,677]" class="Actinopterygii" family="Cetopsidae" genus="Paracetopsis" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="genus">
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<emphasis id="B924EAE9CD23FFACFDA52AE61B7FEABE" box="[629,771,656,677]" italics="true" pageId="47" pageNumber="802">Paracetopsis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C504D78CD23FFACFF722AD819DEEADF" baseAuthorityName="Schultz" baseAuthorityYear="1944" box="[162,418,686,708]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="motatanensis">
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<emphasis id="B924EAE9CD23FFACFF722AD819DEEADF" box="[162,418,686,708]" italics="true" pageId="47" pageNumber="802">Cetopsis motatanensis</emphasis>
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</taxonomicName>
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, the clade formed by
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<taxonomicName id="4C504D78CD23FFACFD7C2AD8197CEAF9" baseAuthorityName="Lundberg & Rapp Py-Daniel" baseAuthorityYear="1994" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="oliveirai">
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<emphasis id="B924EAE9CD23FFACFD7C2AD8197CEAF9" italics="true" pageId="47" pageNumber="802">Cetopsis oliveirai</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C504D78CD23FFACFEDC2ABB19E0EAF9" box="[268,412,717,738]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="coecutiens">
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<emphasis id="B924EAE9CD23FFACFEDC2ABB19E0EAF9" box="[268,412,717,738]" italics="true" pageId="47" pageNumber="802">C. coecutiens</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C504D78CD23FFACFE032ABB1A37EAF9" baseAuthorityName="Spix and Agassiz" baseAuthorityYear="1829" box="[467,587,717,738]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="candiru">
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<emphasis id="B924EAE9CD23FFACFE032ABB1A37EAF9" box="[467,587,717,738]" italics="true" pageId="47" pageNumber="802">C. candiru</emphasis>
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</taxonomicName>
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, and in the clade formed by
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<taxonomicName id="4C504D78CD23FFACFEC92A9A19E4EB1A" box="[281,408,748,769]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="pearsoni">
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<emphasis id="B924EAE9CD23FFACFEC92A9A19E4EB1A" box="[281,408,748,769]" italics="true" pageId="47" pageNumber="802">C. pearsoni</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C504D78CD23FFACFE732A9A1A5FEB1B" box="[419,547,747,769]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="plumbea">
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<emphasis id="B924EAE9CD23FFACFE732A9A1A5FEB1B" box="[419,547,747,769]" italics="true" pageId="47" pageNumber="802">C. plumbea</emphasis>
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</taxonomicName>
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,
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|
<taxonomicName id="4C504D78CD23FFACFDFF2A9A1ADBEB1B" box="[559,679,747,769]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="sandrae">
|
|
<emphasis id="B924EAE9CD23FFACFDFF2A9A1ADBEB1B" box="[559,679,747,769]" italics="true" pageId="47" pageNumber="802">C. sandrae</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C504D78CD23FFACFD642A9A18AAEB3B" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="montana">
|
|
<emphasis id="B924EAE9CD23FFACFD642A9A18AAEB3B" italics="true" pageId="47" pageNumber="802">C. montana</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C504D78CD23FFACFF3A2B7C1925EB3B" baseAuthorityName="Schultz" baseAuthorityYear="1944" box="[234,345,778,800]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="orinoco">
|
|
<emphasis id="B924EAE9CD23FFACFF3A2B7C1925EB3B" box="[234,345,778,800]" italics="true" pageId="47" pageNumber="802">C. orinoco</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C504D78CD23FFACFEBD2B7C1988EB04" authorityName="Kner" authorityYear="1857" box="[365,500,778,799]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="gobioides">
|
|
<emphasis id="B924EAE9CD23FFACFEBD2B7C1988EB04" box="[365,500,778,799]" italics="true" pageId="47" pageNumber="802">C. gobioides</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C504D78CD23FFACFDEA2B7C1ADBEB3B" box="[570,679,778,800]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="arcana">
|
|
<emphasis id="B924EAE9CD23FFACFDEA2B7C1ADBEB3B" box="[570,679,778,800]" italics="true" pageId="47" pageNumber="802">C. arcana</emphasis>
|
|
</taxonomicName>
|
|
, with a subsequent transition to State
|
|
<quantity id="4CA89B1ECD23FFACFDD42B5F1A53EB24" box="[516,559,809,831]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="47" pageNumber="802" unit="in" value="1.0">1 in</quantity>
|
|
the latter species);
|
|
</paragraph>
|
|
<paragraph id="8BEF36FBCD23FFACFF6B2B311A0CEB67" blockId="47.[162,778,195,892]" pageId="47" pageNumber="802">
|
|
Adipose fin absent in adults (122, convergent in the outgroup species
|
|
<taxonomicName id="4C504D78CD23FFACFEBA2B111A23EB67" baseAuthorityName="Guichenot" baseAuthorityYear="1848" box="[362,607,871,892]" class="Actinopterygii" family="Nematogenyidae" genus="Nematogenys" kingdom="Animalia" order="Siluriformes" pageId="47" pageNumber="802" phylum="Chordata" rank="species" species="inermis">
|
|
<emphasis id="B924EAE9CD23FFACFEBA2B111A23EB67" box="[362,607,871,892]" italics="true" pageId="47" pageNumber="802">Nematogenys inermis</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |