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<document id="1F1D8570659E1EBA3D0A1C617BA87002" ID-DOI="10.3853/j.0067-1975.62.2010.1554" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629574225023" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2010" docId="03F987C5FF9EF74D02BB15A64898CFCD" docLanguage="en" docName="RecAustMus.62.2.217-284.pdf" docOrigin="Records of the Australian Museum 62 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada montesquieuana Köhler 2010, n.sp." docType="treatment" docUuid="AE449D6D-B9AE-44E6-905B-A6258CB686E0" docUuidSource="ZooBank" docVersion="3" lastPageNumber="233" masterDocId="FFC0FFBDFF91F75C027510454D15CE74" masterDocTitle="Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)" masterLastPageNumber="284" masterPageNumber="217" pageNumber="232" updateTime="1699078283413" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="56C44B1710DB6CAD3A31CAFB9C930FBA">Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)</mods:title>
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<mods:namePart id="D5B38037890971DC8A3FA596C1BB01E4">Köhler, Frank</mods:namePart>
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<mods:title id="3390D1DA7302FC74859A3E75A489B62E">Records of the Australian Museum</mods:title>
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<mods:date id="7006DA2093AE65A8ADB8785781D33A49">2010</mods:date>
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<mods:number id="8CC22DF55DB5BAC7AE63A1A6D42A4EDF">2010-11-24</mods:number>
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<mods:url id="C479DFC43CEB2C15CD796C0C9451A47D">http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554</mods:url>
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<mods:classification id="301348B63B1DE6A40037900F77CF9970">journal article</mods:classification>
<mods:identifier id="18952C6377020B63A036BB4615087ABB" type="DOI">10.3853/j.0067-1975.62.2010.1554</mods:identifier>
<mods:identifier id="947F6ECA6F3D234B356DC347AFE44324" type="ISSN">2201-4349</mods:identifier>
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<mods:identifier id="E9E732DCC6EB66BDB96873B81C0EB90C" type="ZooBank">749A271A-6942-4D4C-B0DF-B968462BFF0C</mods:identifier>
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<treatment id="03F987C5FF9EF74D02BB15A64898CFCD" LSID="urn:lsid:zoobank.org:act:AE449D6D-B9AE-44E6-905B-A6258CB686E0" httpUri="http://treatment.plazi.org/id/03F987C5FF9EF74D02BB15A64898CFCD" lastPageId="17" lastPageNumber="233" pageId="15" pageNumber="232">
<subSubSection id="C34A6558FF9EF75302BB15A64F8DCB89" box="[206,664,1507,1533]" pageId="15" pageNumber="232" type="nomenclature">
<paragraph id="8BEF36D3FF9EF75302BB15A64F8DCB89" blockId="15.[206,664,1507,1533]" box="[206,664,1507,1533]" pageId="15" pageNumber="232">
<heading id="D0A781BFFF9EF75302BB15A64F8DCB89" box="[206,664,1507,1533]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="232" reason="2">
<emphasis id="B924EAC1FF9EF75302BB15A64F8DCB89" bold="true" box="[206,664,1507,1533]" pageId="15" pageNumber="232">
<taxonomicName id="4C504D50FF9EF75302BB15A64F46CB89" ID-CoL="D7RM" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[206,595,1507,1533]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="232" phylum="Mollusca" rank="species" species="montesquieuana" status="sp. nov.">
<emphasis id="B924EAC1FF9EF75302BB15A64F46CB89" bold="true" box="[206,595,1507,1533]" italics="true" pageId="15" pageNumber="232">Amplirhagada montesquieuana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF9EF753002915A64F8DCB89" box="[604,664,1507,1533]" pageId="15" pageNumber="232" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C34A6558FF9EF753020C16524DF7C890" pageId="15" pageNumber="232" type="materials_examined">
<paragraph id="8BEF36D3FF9EF753020C16524C14C8F3" blockId="15.[121,749,1559,1671]" pageId="15" pageNumber="232">
<materialsCitation id="3B383C8EFF9EF753020C16524DE8C8F3" collectingDate="2002-08-28" collectionCode="FERT" collectorName="Harvey" county="northwestern Kimberley" latitude="-14.137777" location="Admiralty Gulf" longLatPrecision="21" longitude="125.69861" municipality="Bonaparte Archipelago" pageId="15" pageNumber="232" specimenCode="FERT 04" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B924EAC1FF9EF753020C16524C06C85B" bold="true" box="[121,275,1559,1583]" pageId="15" pageNumber="232">
<typeStatus id="54EB8871FF9EF753020C16524DA6C85B" box="[121,179,1559,1583]" pageId="15" pageNumber="232">Type</typeStatus>
locality
</emphasis>
(
<figureCitation id="136B2A56FF9EF7530353165D4C7FC85B" box="[294,362,1560,1583]" captionStart="Figure 1" captionStartId="1.[121,184,1433,1454]" captionTargetBox="[121,1421,142,1408]" captionTargetId="figure-129@1.[121,1421,138,1414]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species in the Kimberley region, northwestern Australia. (1) A. euroa n.sp., Adolphus Island. (2) A. solemiana n.sp., Middle Osborn Island. (3) A. indistincta n.sp., Southwest Osborn Island, Kidney Island. (4) A. combeana, Cassini Island. (5) A. mckenziei n.sp., Oliver Island. (6) A. ponderi n.sp., Kingsmill Island. (7) A. montesquieuana n.sp., Fenelon Island. (8) A. descartesana n.sp., Descartes Island. (9) A. katerana, Katers Island. (10) A. puescheli n.sp., unnamed island near Prudhoe Island. (11) A. decora n.sp., A. kessneri n.sp., Bigge Island. (12) A. berthierana n.sp., Berthier Island. (13) A. lamarckiana n.sp., Lamarck Island.(14) A. anderdonensis n.sp., unnamed island, Anderdon Islands. (15) A. tricenaria n.sp., Prince Frederick Harbour, north of Hunter River mouth. (16) A. regia n.sp., A. boongareensis n.sp., A. gibsoni n.sp., and Amplirhagada sp., Boongaree Island. (17) A. yorkensis n.sp., Coronation Island. (18) A. buffonensis n.sp., Buffon Island. (19) A. uwinsensis n.sp., Uwins Island. (20) A. sphaeroidea n.sp., St. Andrews Island. (21) A. basilica n.sp., and A. camdenensis n.sp., Augustus Island. (22) A. kimberleyana n.sp., NW of Wilson Point. (23) A. gemina n.sp., 1.5 km SE of Hall Point. (24) A. dubitabile n.sp., Steep Island. (25) A. napierana, north Napier ranges. (26) A. percita, Napier ranges. (27) A. castra, A. mitchelliana, and A. varia, Mitchell Plateau." figureDoi="http://doi.org/10.5281/zenodo.5239088" httpUri="https://zenodo.org/record/5239088/files/figure.png" pageId="15" pageNumber="232">Fig. 1</figureCitation>
).
<collectingRegion id="4994F831FF9EF75303F0165D4F5AC85B" box="[389,591,1560,1583]" country="Australia" name="Western Australia" pageId="15" pageNumber="232">Western Australia</collectingRegion>
,
<collectingCounty id="628E4E5FFF9EF7530028165D4DFFC838" pageId="15" pageNumber="232">northwestern Kimberley</collectingCounty>
,
<collectingMunicipality id="6B8BACA9FF9EF753028716704CF0C838" box="[242,485,1589,1612]" pageId="15" pageNumber="232">Bonaparte Archipelago</collectingMunicipality>
,
<location id="8E8F6008FF9EF753039916704F85C838" LSID="urn:lsid:plazi:treatment:03F987C5FF9EF74D02BB15A64898CFCD:8E8F6008FF9EF753039916704F85C838" box="[492,656,1589,1612]" county="northwestern Kimberley" latitude="-14.137777" longLatPrecision="21" longitude="125.69861" municipality="Bonaparte Archipelago" name="Admiralty Gulf" pageId="15" pageNumber="232" stateProvince="Western Australia">Admiralty Gulf</location>
,
<location id="8E8F6008FF9EF75300ED16704DABC81D" LSID="urn:lsid:plazi:treatment:03F987C5FF9EF74D02BB15A64898CFCD:8E8F6008FF9EF75300ED16704DABC81D" county="northwestern Kimberley" latitude="-14.137777" longLatPrecision="21" longitude="125.69861" municipality="Bonaparte Archipelago" name="Fenelon Island" pageId="15" pageNumber="232" stateProvince="Western Australia">Fenelon Island</location>
,
<geoCoordinate id="EE645014FF9EF75302B916174C5DC81D" box="[204,328,1618,1641]" degrees="14" direction="south" minutes="08" orientation="latitude" pageId="15" pageNumber="232" precision="15" seconds="16" value="-14.137777">14°08'16&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FF9EF753032716174CCAC81D" box="[338,479,1618,1641]" degrees="125" direction="east" minutes="41" orientation="longitude" pageId="15" pageNumber="232" precision="15" seconds="55" value="125.69861">125°41'55&quot;E</geoCoordinate>
;
<specimenCode id="DBF69EA8FF9EF753039816174F46C81D" box="[493,595,1618,1641]" collectionCode="FERT" pageId="15" pageNumber="232">FERT 04</specimenCode>
(leg.
<collectorName id="26A55305FF9EF75300ED16174FFDC81D" box="[664,744,1618,1641]" pageId="15" pageNumber="232">Harvey</collectorName>
,
<date id="FFEE1013FF9EF753020C16354DE3C8F3" box="[121,246,1648,1671]" pageId="15" pageNumber="232" value="2002-08-28">
<collectingDate id="EFAAE9FBFF9EF753020C16354DE3C8F3" box="[121,246,1648,1671]" pageId="15" pageNumber="232" value="2002-08-28">28.08.2002</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF36D3FF9EF753020C16E54DF7C890" blockId="15.[121,749,1696,1764]" pageId="15" pageNumber="232">
<emphasis id="B924EAC1FF9EF753020C16E54C17C8C7" bold="true" box="[121,258,1696,1715]" pageId="15" pageNumber="232">Type material</emphasis>
.
<materialsCitation id="3B383C8EFF9EF753036416E44F47C8C0" accessionNumber="S34606" box="[273,594,1697,1716]" collectionCode="WAM" pageId="15" pageNumber="232" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EB8871FF9EF753036416E44C70C8C0" box="[273,357,1697,1716]" pageId="15" pageNumber="232" type="holotype">Holotype</typeStatus>
<collectionCode id="ED41AE16FF9EF753031916E44CB7C8C0" box="[364,418,1697,1716]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="15" pageNumber="232" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF9EF75303DE16E44CFAC8C0" box="[427,495,1697,1716]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34606" pageId="15" pageNumber="232" type="EnaNcbi">S34606</accessionNumber>
(Pl. 1.11)
</materialsCitation>
.
<materialsCitation id="3B383C8EFF9EF753002B16E44DCBC890" accessionNumber="S41453" collectionCode="WAM" pageId="15" pageNumber="232" specimenCount="1" typeStatus="paratype">
<typeStatus id="54EB8871FF9EF753002B16E44FA0C8C0" box="[606,693,1697,1716]" pageId="15" pageNumber="232" type="paratype">Paratypes</typeStatus>
AMS C463748 (9 preserved specimens),
<collectionCode id="ED41AE16FF9EF75303A316FC4F18C8B8" box="[470,525,1721,1740]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="15" pageNumber="232" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF9EF753006316FC4F4AC8B8" box="[534,607,1721,1740]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41453" pageId="15" pageNumber="232" type="EnaNcbi">S41453</accessionNumber>
(15 preserved specimens)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FF9EF74D020C16BA4898CFCD" lastPageId="17" lastPageNumber="234" pageId="15" pageNumber="232" type="etymology">
<paragraph id="8BEF36D3FF9EF753020C16BA4C6DC940" blockId="15.[121,748,1791,1844]" pageId="15" pageNumber="232">
<emphasis id="B924EAC1FF9EF753020C16BA4DE1C963" bold="true" box="[121,244,1791,1815]" pageId="15" pageNumber="232">Etymology</emphasis>
. In reference to Montesquieu Islands, to which Fenelon Island belongs.
</paragraph>
<paragraph id="8BEF36D3FF9EF753020C17154CA7C913" blockId="15.[121,434,1872,1895]" box="[121,434,1872,1895]" pageId="15" pageNumber="232">Sealing strategy. Rock sealer.</paragraph>
<paragraph id="8BEF36D3FF9EF753020C17C7490FC84E" blockId="15.[121,749,1922,2004]" lastBlockId="15.[794,1422,1189,1594]" pageId="15" pageNumber="232">
<emphasis id="B924EAC1FF9EF753020C17C74DA4C9EE" bold="true" box="[121,177,1922,1946]" pageId="15" pageNumber="232">Shell</emphasis>
(
<figureCitation id="136B2A56FF9EF75302C817C74C2EC9ED" box="[189,315,1922,1945]" captionStart="Figure 21" captionStartId="16.[166,229,1539,1560]" captionTargetBox="[166,1466,151,1526]" captionTargetId="figure-8@16.[166,1466,151,1526]" captionTargetPageId="16" captionText="Figure 21. SEM photographs of Amplirhagada montesquieuana n.sp., paratypes AMS C463748. (AB) Shell: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture on last two whorls viewed from above (scale 200 µm). (CF) Radula: (C) central and lateral teeth (Scale 20 µm); (D) close-up of central and inner lateral teeth (scale 10 µm); (E) outer lateral and inner marginal teeth (scale 10 µm); (F) close-up of middle marginal teeth (scale 10 µm). (G) Jaw (scale 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239131" httpUri="https://zenodo.org/record/5239131/files/figure.png" pageId="15" pageNumber="232">Fig. 21AB</figureCitation>
, Pl. 1.11). Broadly conical with low spire; thin (translucent). Whorls evenly rounded in cross-section. Umbilicus open, forming a chink to narrowly winding opening, 8095 percent concealed by columellar reflection. Background colour whitish or yellowish to golden brown; with diffuse, thin, brown sub-sutural and mid-whorl bands most conspicuous on last whorl; ventral colour whitish; outer lip colour same as shell or lighter; inner lip whitish and translucent. Protoconch c.
<quantity id="4CA89B36FF9EF753061C157D49BDCB3B" box="[1129,1192,1336,1359]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="232" unit="mm" value="2.0">2 mm</quantity>
in diameter with 1.5 whorls, sculptured by fine but distinct axial lirae. Teleoconch with coarse axial lirae, rounded in cross-section with regular spacing, spaces equal to thickness of lirae. Sculpture evenly distributed across shell and whorl diameter, height reduced underneath suture. Angle of aperture 30°, outer lip rounded, sharp to moderately thick, slightly expanded and reflected; basal node of lip weak, palatal node absent. Parietal wall of inner lip inconspicuous.
</paragraph>
<paragraph id="8BEF36D3FF9EF753016F161849E8C8DB" blockId="15.[794,1422,1629,1711]" pageId="15" pageNumber="232">
<emphasis id="B924EAC1FF9EF753016F16184912C801" bold="true" box="[794,1031,1629,1653]" pageId="15" pageNumber="232">Pallial morphology</emphasis>
. Pallial cavity moderately deep, extending
<emphasis id="B924EAC1FF9EF75301F1163E4E99C8FD" bold="true" box="[900,908,1659,1673]" pageId="15" pageNumber="232">
<superScript id="7C259B9BFF9EF75301F1163E4E99C8FD" attach="right" box="[900,908,1659,1673]" fontSize="6" pageId="15" pageNumber="232">3</superScript>
</emphasis>
<emphasis id="B924EAC1FF9EF75301E416C44E8CC8FB" bold="true" box="[913,921,1665,1679]" pageId="15" pageNumber="232">4</emphasis>
whorl. Mantle with mottled, black pigmentation. Kidney extending about half of pallial cavity.
</paragraph>
<paragraph id="8BEF36D3FF9EF74C016F169748AFC8DB" blockId="15.[794,1422,1746,2004]" lastBlockId="16.[839,1466,1688,1711]" lastPageId="16" lastPageNumber="233" pageId="15" pageNumber="232">
<emphasis id="B924EAC1FF9EF753016F16974EE9C89E" bold="true" box="[794,1020,1746,1770]" pageId="15" pageNumber="232">Genital morphology</emphasis>
(
<figureCitation id="136B2A56FF9EF753067E16964999C89E" box="[1035,1164,1747,1770]" captionStart-0="Figure 22" captionStart-1="Figure 23" captionStartId-0="17.[121,186,632,653]" captionStartId-1="17.[121,184,1629,1650]" captionTargetBox-0="[126,736,149,618]" captionTargetBox-1="[121,728,734,1757]" captionTargetId-0="figure-9@17.[121,748,141,627]" captionTargetId-1="figure-34@17.[121,748,728,1723]" captionTargetPageId-0="17" captionTargetPageId-1="17" captionText-0="Figure 22. Genitalia of Amplirhagada montesquieuana n.sp., paratypeAMS C463748 (28August, scale 10 mm). Compare with Fig. 3 for labelling of structures." captionText-1="Figure 23. Interior of penial chamber" figureDoi-0="http://doi.org/10.5281/zenodo.5239133" figureDoi-1="http://doi.org/10.5281/zenodo.5239135" httpUri-0="https://zenodo.org/record/5239133/files/figure.png" httpUri-1="https://zenodo.org/record/5239135/files/figure.png" pageId="15" pageNumber="232">Figs. 2223</figureCitation>
). Penis straight, of more or less same length as anterior part of oviduct to slightly longer. Vas deferens coils once before entering penis. Penial retractor muscle clearly shorter than penis. Penial verge very short, slender to spatulate, with pointed tip. Penial wall pustules of normal size, slightly elongated, arranged in sparsely distributed, distinct rows over entire length of inner penial wall. Towards base of penial chamber, pustules fuse to form corrugated, narrow pilasters. Main stimulatory pilaster differentiated, forming cone-shaped, prominent ridge covered by enlarged pustules at apical to median portion of penial chamber. Pilaster ridges carry little toe-shaped extensions that support hooks. Vas deferens entering penial sheath in upper third. Vagina moderately long, tubular. Inner vaginal wall with smooth longitudinal pilasters. Spermathecal duct wide; inner wall with transversely structured, longitudinal pilasters. Spermathecal head globular to elongate, connected with oviduct by connective tissue; wall delicate. Free oviduct comprising±half of anterior part of oviduct, rather straight. Spermoviduct longer than anterior part of oviduct. Talon embedded in albumen gland at junction with spermoviduct.
</paragraph>
<caption id="DF2F665BFF81F74C02D316464E9EC81C" ID-DOI="http://doi.org/10.5281/zenodo.5239131" ID-Zenodo-Dep="5239131" httpUri="https://zenodo.org/record/5239131/files/figure.png" pageId="16" pageNumber="233" startId="16.[166,229,1539,1560]" targetBox="[166,1466,151,1526]" targetPageId="16">
<paragraph id="8BEF36D3FF81F74C02D316464E9EC81C" blockId="16.[166,1467,1539,1640]" pageId="16" pageNumber="233">
Figure 21. SEM photographs of
<taxonomicName id="4C504D50FF81F74C039416464E1AC86C" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[481,783,1539,1560]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="233" phylum="Mollusca" rank="species" species="montesquieuana" status="sp. nov.">
<emphasis id="B924EAC1FF81F74C039416464E1AC86C" box="[481,783,1539,1560]" italics="true" pageId="16" pageNumber="233">Amplirhagada montesquieuana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF81F74C016216464E53C86C" box="[791,838,1539,1560]" pageId="16" pageNumber="233" rank="species">n.sp.</taxonomicNameLabel>
, paratypes AMS C463748. (
<emphasis id="B924EAC1FF81F74C062F16464996C86C" box="[1114,1155,1539,1560]" italics="true" pageId="16" pageNumber="233">AB</emphasis>
) Shell: (
<emphasis id="B924EAC1FF81F74C06AC164649FDC86C" box="[1241,1256,1539,1560]" italics="true" pageId="16" pageNumber="233">A</emphasis>
) apical whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FF81F74C03B4165B4CC5C847" box="[449,464,1566,1587]" italics="true" pageId="16" pageNumber="233">B</emphasis>
) sculpture on last two whorls viewed from above (scale 200 µm). (
<emphasis id="B924EAC1FF81F74C062E165B4993C847" box="[1115,1158,1566,1587]" italics="true" pageId="16" pageNumber="233">CF</emphasis>
) Radula: (
<emphasis id="B924EAC1FF81F74C069B165B49EBC847" box="[1262,1278,1566,1587]" italics="true" pageId="16" pageNumber="233">C</emphasis>
) central and lateral teeth (Scale 20 µm); (
<emphasis id="B924EAC1FF81F74C0306167D4C91C839" box="[371,388,1592,1613]" italics="true" pageId="16" pageNumber="233">D</emphasis>
) close-up of central and inner lateral teeth (scale 10 µm); (
<emphasis id="B924EAC1FF81F74C01C7167D4ED4C839" box="[946,961,1592,1613]" italics="true" pageId="16" pageNumber="233">E</emphasis>
) outer lateral and inner marginal teeth (scale 10 µm); (
<emphasis id="B924EAC1FF81F74C02DB16164DA8C81C" box="[174,189,1619,1640]" italics="true" pageId="16" pageNumber="233">F</emphasis>
) close-up of middle marginal teeth (scale 10 µm). (
<emphasis id="B924EAC1FF81F74C00DE16164FA9C81C" box="[683,700,1619,1640]" italics="true" pageId="16" pageNumber="233">G</emphasis>
) Jaw (scale 100 µm).
</paragraph>
</caption>
<paragraph id="8BEF36D3FF81F74D013216974898CFCD" blockId="16.[839,1467,1746,2004]" lastBlockId="17.[794,1422,154,441]" lastPageId="17" lastPageNumber="234" pageId="16" pageNumber="233">
<emphasis id="B924EAC1FF81F74C013216974928C89E" bold="true" box="[839,1085,1746,1770]" pageId="16" pageNumber="233">Radular morphology</emphasis>
(
<figureCitation id="136B2A56FF81F74C0625169649C1C89E" box="[1104,1236,1747,1770]" captionStart="Figure 21" captionStartId="16.[166,229,1539,1560]" captionTargetBox="[166,1466,151,1526]" captionTargetId="figure-8@16.[166,1466,151,1526]" captionTargetPageId="16" captionText="Figure 21. SEM photographs of Amplirhagada montesquieuana n.sp., paratypes AMS C463748. (AB) Shell: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture on last two whorls viewed from above (scale 200 µm). (CF) Radula: (C) central and lateral teeth (Scale 20 µm); (D) close-up of central and inner lateral teeth (scale 10 µm); (E) outer lateral and inner marginal teeth (scale 10 µm); (F) close-up of middle marginal teeth (scale 10 µm). (G) Jaw (scale 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239131" httpUri="https://zenodo.org/record/5239131/files/figure.png" pageId="16" pageNumber="233">Fig. 21CF</figureCitation>
). Rectangular. Tooth formula C+11+4+14. Average number of rows of teeth 150±32.5 with 38.1±0.3 rows per mm (n = 2). Central teeth with sharply pointed, triangular mesocones, shorter than base of tooth; ectocones small. Lateral teeth with bluntly pointed, triangular mesocones, length equal to base of tooth; ectocones well developed, endocones vestigial. Marginal teeth with elongate mesocones; ectocones shorter and narrower than mesocones, split into two denticles; endocones reduced in size.
<emphasis id="B924EAC1FF80F74D020C176C4C61C935" bold="true" box="[121,372,1833,1857]" pageId="17" pageNumber="234">Comparative remarks</emphasis>
. The shell of this species is smaller than that of other species from the Montesquieu Islands but otherwise difficult to differentiate by means of colour, sculpture and shape (
<tableCitation id="C6D20368FF80F74D031417C44CA1C9EC" box="[353,436,1921,1944]" captionStart="Table 1" captionStartId="61.[120,176,474,495]" captionText="Table 1. Shell parameters of Amplirhagada species, given in the form maximumminimum (mean±standard deviation) measures of shells for n specimens." pageId="17" pageNumber="234">Table 1</tableCitation>
). Its axial sculpture is more regular than that of
<taxonomicName id="4C504D50FF80F74D032117DA4CF7C9C2" box="[340,482,1951,1974]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="234" phylum="Mollusca" rank="species" species="combeana">
<emphasis id="B924EAC1FF80F74D032117DA4CF7C9C2" box="[340,482,1951,1974]" italics="true" pageId="17" pageNumber="234">A. combeana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C504D50FF80F74D006E17DA4FB6C9C2" authorityName="Köhler" authorityYear="2010" box="[539,675,1951,1974]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="234" phylum="Mollusca" rank="species" species="mckenziei">
<emphasis id="B924EAC1FF80F74D006E17DA4FB6C9C2" box="[539,675,1951,1974]" italics="true" pageId="17" pageNumber="234">A. mckenziei</emphasis>
</taxonomicName>
. From
<taxonomicName id="4C504D50FF80F74D020C17F94DEAC9A7" authorityName="Köhler" authorityYear="2010" box="[121,255,1980,2003]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="234" phylum="Mollusca" rank="species" species="mckenziei">
<emphasis id="B924EAC1FF80F74D020C17F94DEAC9A7" box="[121,255,1980,2003]" italics="true" pageId="17" pageNumber="234">A. mckenziei</emphasis>
</taxonomicName>
it also differs by a smaller and comparatively flatter shell; a one-way ANOVA revealed that both species differ significantly in the shell parameters H, D, FW and H/D. The inner penial wall is characteristic by its rather sparsely distributed rows of pustules that give rise to corrugated, narrow, longitudinal pilasters towards the base of the penial chamber, and the shape and development of the main stimulatory pilaster being shorter and broader than in
<taxonomicName id="4C504D50FF80F74D016F112D4E9ECF0B" authorityName="Köhler" authorityYear="2010" box="[794,907,360,383]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="234" phylum="Mollusca" rank="species" species="ponderi">
<emphasis id="B924EAC1FF80F74D016F112D4E9ECF0B" box="[794,907,360,383]" italics="true" pageId="17" pageNumber="234">A. ponderi</emphasis>
</taxonomicName>
and larger than in
<taxonomicName id="4C504D50FF80F74D0623112D49C9CF0B" authorityName="Köhler" authorityYear="2010" box="[1110,1244,360,383]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="234" phylum="Mollusca" rank="species" species="mckenziei">
<emphasis id="B924EAC1FF80F74D0623112D49C9CF0B" box="[1110,1244,360,383]" italics="true" pageId="17" pageNumber="234">A. mckenziei</emphasis>
</taxonomicName>
. The possession of well-developed “toe-shaped” extensions of the pilaster ridges is unique amongst species from Montesquieu Islands.
</paragraph>
</subSubSection>
</treatment>
</document>