1219 lines
160 KiB
XML
1219 lines
160 KiB
XML
<document id="E6257E43B5FB219A049C26F85F1E4BCD" ID-CLB-Dataset="276847" ID-DOI="10.11646/phytotaxa.584.3.4" ID-GBIF-Dataset="51e9548a-d257-49f6-a33c-e34e5215ea6c" ID-ISSN="1179-3163" ID-Zenodo-Dep="7645651" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676538678178" checkinUser="plazi" docAuthor="Zografidis, Aris, Koutroumpa, Konstantina, Liveri, Eleni & Dimopoulos, Panayotis" docDate="2023" docId="03F98790FFFBFFECD1AA8CF0FD50E73A" docLanguage="en" docName="phytotaxa.584.3.4.pdf" docOrigin="Phytotaxa 584 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.584.3.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Centaurea dolopica Zograf., Koutroumpa, Liveri & Dimop. 2023, sp. nov." docType="treatment" docVersion="6" lastPageNumber="183" masterDocId="FFC0FFE8FFF8FFE6D1228C04FF90E202" masterDocTitle="Centaurea dolopica (Asteraceae, Cardueae), a new species of section Acrocentron from Greece" masterLastPageNumber="185" masterPageNumber="173" pageNumber="176" updateTime="1699477740960" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4D05AF29F5EC669C7819AA453933387E">Centaurea dolopica (Asteraceae, Cardueae), a new species of section Acrocentron from Greece</mods:title>
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<mods:namePart id="DE0964CB1BB4820503BC5E9221162D38">Zografidis, Aris</mods:namePart>
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<mods:affiliation id="956CF9AC62BB58F0875C641B3AF6FB0B">Department of Biology, University of Patras, University Campus, GR 26504 Rion, Greece</mods:affiliation>
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<mods:namePart id="E4E6FC706E7A2549BFB54261319CB2AB">Koutroumpa, Konstantina</mods:namePart>
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<mods:affiliation id="F976A9C4DCFAE12E4B81F904DCA601BB">Botanischer Garten und Botanisches Museum Berlin, Freie Universität Berlin, Königin-Luise-Str. 6 - 8, 14195 Berlin, Germany k. koutroumpa @ bo. berlin; https: // orcid. org / 0000 - 0001 - 9391 - 874 X</mods:affiliation>
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<mods:nameIdentifier id="5659767BD6E4FF74E2CAEB4692B6C319" type="email">k.koutroumpa@bo.berlin</mods:nameIdentifier>
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<mods:namePart id="4B35EABE7BCF434117C3BCB15F798E17">Liveri, Eleni</mods:namePart>
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<mods:affiliation id="84E55D6823198165A40275CFED1BC8FD">Department of Biology, University of Patras, University Campus, GR 26504 Rion, Greece & Department of Agriculture, University of Patras, GR 30200 Messolonghi, Greece</mods:affiliation>
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<mods:namePart id="80FE362099643617DCF7A89691EBCB1A">Dimopoulos, Panayotis</mods:namePart>
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<mods:affiliation id="3CC0AF871C93E9CF6EE6E47704EF9FF0">Department of Biology, University of Patras, University Campus, GR 26504 Rion, Greece</mods:affiliation>
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<mods:title id="FCA533CFA879D342F71DA6E01716FA3D">Phytotaxa</mods:title>
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<mods:date id="6301ACE6E69C9A5B83C76204B28D407C">2023</mods:date>
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<mods:number id="E82B38C71A374390B14E3812B1E936C3">2023-02-16</mods:number>
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<mods:number id="8D0A8B8500DE50154FFE3F1E1B7F7B82">584</mods:number>
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<mods:url id="FF54C855D2D2C5A37B7065C42D9FB0B6">http://dx.doi.org/10.11646/phytotaxa.584.3.4</mods:url>
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<treatment id="03F98790FFFBFFECD1AA8CF0FD50E73A" ID-DOI="http://doi.org/10.5281/zenodo.7647113" ID-GBIF-Taxon="217351084" ID-Zenodo-Dep="7647113" LSID="urn:lsid:plazi:treatment:03F98790FFFBFFECD1AA8CF0FD50E73A" httpUri="http://treatment.plazi.org/id/03F98790FFFBFFECD1AA8CF0FD50E73A" lastPageId="10" lastPageNumber="183" pageId="3" pageNumber="176">
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<heading id="D0A781EAFFFBFFE5D1AA8CF0FC03E30C" box="[136,915,244,271]" fontSize="11" level="2" pageId="3" pageNumber="176" reason="3">
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<taxonomicName id="4C504D05FFFBFFE5D1AA8CF0FCBEE30C" ID-CoL="B6KD6" authority="Zograf., Koutroumpa, Liveri & Dimop." authorityName="Zograf., Koutroumpa, Liveri & Dimop." authorityYear="2023" box="[136,814,244,271]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="176" phylum="Tracheophyta" rank="species" species="dolopica" status="sp. nov.">
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<emphasis id="B924EA94FFFBFFE5D1AA8CF0FEFAE30D" bold="true" box="[136,362,244,271]" italics="true" pageId="3" pageNumber="176">Centaurea dolopica</emphasis>
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Zograf., Koutroumpa, Liveri & Dimop.
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</taxonomicName>
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,
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<emphasis id="B924EA94FFFBFFE5D21B8CF0FC03E30C" bold="true" box="[825,915,244,270]" pageId="3" pageNumber="176">
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<taxonomicNameLabel id="A21757EFFFFBFFE5D21B8CF0FC03E30C" box="[825,915,244,270]" pageId="3" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A650DFFFBFFE2D1AA8D18FDE4E3FF" lastPageId="4" lastPageNumber="177" pageId="3" pageNumber="176" type="diagnosis">
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<emphasis id="B924EA94FFFBFFE5D1AA8D18FB1EE37B" italics="true" pageId="3" pageNumber="176">
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Diagnosis:—A species belonging to the
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<taxonomicName id="4C504D05FFFBFFE5D3338D18FD2DE333" box="[529,701,284,305]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="176" phylum="Tracheophyta" rank="species" species="achaia">Centaurea achaia</taxonomicName>
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group (including
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<taxonomicName id="4C504D05FFFBFFE5D24C8D18FC5FE333" box="[878,975,284,305]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="176" phylum="Tracheophyta" rank="species" species="achaia">C. achaia</taxonomicName>
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,
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<taxonomicName id="4C504D05FFFBFFE5D2F88D18FBD7E333" box="[986,1095,284,305]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="176" phylum="Tracheophyta" rank="species" species="aetolica">C. aetolica</taxonomicName>
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,
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<taxonomicName id="4C504D05FFFBFFE5D5738D18FB4DE333" box="[1105,1245,284,305]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="176" phylum="Tracheophyta" rank="species" species="corinthiaca">C. corinthiaca</taxonomicName>
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and
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<taxonomicName id="4C504D05FFFBFFE5D42D8D18FAE9E333" box="[1295,1401,284,305]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="176" phylum="Tracheophyta" rank="species" species="euboica">C. euboica</taxonomicName>
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) and distinguished from the other members by its narrower involucral bracts and by its fewer central florets. Robust individuals of
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||
<taxonomicName id="4C504D05FFFBFFE5D4B78D44FE81E37B" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="176" phylum="Tracheophyta" rank="species" species="dolopica">C. dolopica</taxonomicName>
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also exhibit larger and more branched flowering stems bearing a larger number of capitula.
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</emphasis>
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</paragraph>
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:—
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.
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,
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: ca.
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of
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, scrub on serpentine cut slope and roadsides,
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,
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,
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,
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,
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<emphasis id="B924EA94FFFBFFE5D3008DA8FD20E3C3" box="[546,688,428,449]" italics="true" pageId="3" pageNumber="176">Zografidis 745</emphasis>
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(
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:
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<collectionCode id="ED41AE43FFFBFFE5D23D8DA8FCDFE3C0" box="[799,847,428,450]" country="Greece" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14588" lsid="urn:lsid:biocol.org:col:14588" name="University of Patras" pageId="3" pageNumber="176" type="Herbarium">UPA</collectionCode>
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(39986 & 39987)! mounted on two sheets; isotypes:
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<collectionCode id="ED41AE43FFFBFFE5D46A8DA8FAEEE3C3" box="[1352,1406,428,449]" country="Greece" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13129" lsid="urn:lsid:biocol.org:col:13129" name="Goulandris Natural History Museum" pageId="3" pageNumber="176" type="Herbarium">ATH</collectionCode>
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!,
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<collectionCode id="ED41AE43FFFBFFE5D4A98DA8FA0FE3C3" box="[1419,1439,428,449]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="3" pageNumber="176" type="Herbarium">B</collectionCode>
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!) (
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<figureCitation id="136B2A03FFFBFFE5D1E78DD4FE91E3E7" box="[197,257,464,486]" captionStart="FIGURE 1" captionStartId="3.[136,229,1600,1622]" captionTargetBox="[162,1428,517,1565]" captionTargetId="figure-145@3.[154,1433,508,1573]" captionTargetPageId="3" captionText="FIGURE 1. Centaurea dolopica. A, habit; B, capitulum; C, marginal (left) and central (right) floret, scale bar = 1 cm; D, two marginal (top left) and three central achenes, scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7645653" httpUri="https://zenodo.org/record/7645653/files/figure.png" pageId="3" pageNumber="176">Figs 1</figureCitation>
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,
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<figureCitation id="136B2A03FFFBFFE5D02F8DD4FE8BE3E7" box="[269,283,464,485]" captionStart="FIGURE 2" captionStartId="4.[136,229,1483,1505]" captionTargetBox="[152,1432,543,1452]" captionTargetId="figure-200@4.[146,1441,533,1459]" captionTargetPageId="4" captionText="FIGURE 2. Holotype of Centaurea dolopica (UPA) mounted on two sheets." figureDoi="http://doi.org/10.5281/zenodo.7645655" httpUri="https://zenodo.org/record/7645655/files/figure.png" pageId="3" pageNumber="176">2</figureCitation>
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,
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<figureCitation id="136B2A03FFFBFFE5D0078DD4FEA1E3E7" box="[293,305,464,485]" captionStart="FIGURE 3" captionStartId="5.[136,229,1620,1642]" captionTargetBox="[152,1435,755,1586]" captionTargetId="figure-317@5.[143,1444,749,1595]" captionTargetPageId="5" captionText="FIGURE 3. Centaurea dolopica plants in their natural habitat: (A) several large plants along a roadside and (B) on a cut slope. (C) Eryngium serbicum in the same habitat." figureDoi="http://doi.org/10.5281/zenodo.7645657" httpUri="https://zenodo.org/record/7645657/files/figure.png" pageId="3" pageNumber="176">3</figureCitation>
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(
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<collectionCode id="ED41AE43FFFBFFE5D01A8DD4FEDDE3E7" box="[312,333,464,485]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="3" pageNumber="176" type="Herbarium">A</collectionCode>
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,
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<collectionCode id="ED41AE43FFFBFFE5D0738DD4FEF2E3E7" box="[337,354,464,485]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="3" pageNumber="176" type="Herbarium">B</collectionCode>
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), 4(e))
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</materialsCitation>
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.
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</paragraph>
|
||
<caption id="DF2F660EFFFBFFE5D1AA8A44FD1DE478" ID-DOI="http://doi.org/10.5281/zenodo.7645653" ID-Zenodo-Dep="7645653" httpUri="https://zenodo.org/record/7645653/files/figure.png" pageId="3" pageNumber="176" startId="3.[136,229,1600,1622]" targetBox="[162,1428,517,1565]" targetPageId="3">
|
||
<paragraph id="8BEF3686FFFBFFE5D1AA8A44FD1DE478" blockId="3.[136,1451,1600,1658]" pageId="3" pageNumber="176">
|
||
<emphasis id="B924EA94FFFBFFE5D1AA8A44FF6FE457" bold="true" box="[136,255,1600,1622]" pageId="3" pageNumber="176">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C504D05FFFBFFE5D0278A45FE54E457" box="[261,452,1600,1622]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="176" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFBFFE5D0278A45FE54E457" box="[261,452,1600,1622]" italics="true" pageId="3" pageNumber="176">Centaurea dolopica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B924EA94FFFBFFE5D0F38A44FE72E457" bold="true" box="[465,482,1600,1621]" pageId="3" pageNumber="176">A</emphasis>
|
||
, habit;
|
||
<emphasis id="B924EA94FFFBFFE5D30F8A45FDADE454" bold="true" box="[557,573,1601,1622]" pageId="3" pageNumber="176">B</emphasis>
|
||
, capitulum;
|
||
<emphasis id="B924EA94FFFBFFE5D3958A44FD58E454" bold="true" box="[695,712,1600,1622]" pageId="3" pageNumber="176">C</emphasis>
|
||
, marginal (left) and central (right) floret, scale bar = 1 cm;
|
||
<emphasis id="B924EA94FFFBFFE5D42F8A45FA8EE454" bold="true" box="[1293,1310,1601,1622]" pageId="3" pageNumber="176">D</emphasis>
|
||
, two marginal (top left) and three central achenes, scale bar = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEF3686FFFBFFE2D1AA8AADFDE4E3FF" blockId="3.[136,1452,1705,2092]" lastBlockId="4.[136,1452,158,509]" lastPageId="4" lastPageNumber="177" pageId="3" pageNumber="176">
|
||
Robust, densely arachnoid, perennial herb with one to several leaf rosettes and one to several (40 counted in a large individual) erect flowering stems 40–100(–120) cm in length, often bent in maturity by the weight of their capitula. Flowering stems branched from below middle (in robust, much-branched stems) or above middle (in sparingly branched stems) with 1–11 branches, forming pseudopaniculate-subcorymbose synflorescences, each flowering stem bearing 3–29 capitula. Basal, late successional rosette leaves and lower cauline leaves 1- to 2-pinnatisect to pinnate, ovate-elliptical to oblong-lanceolate in outline, 10–55 ×
|
||
<quantity id="4CA89B63FFFBFFE5D3D58B59FCC7E575" box="[759,855,1885,1912]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="15.0" metricValueMin="3.0" pageId="3" pageNumber="176" unit="cm" value="9.0" valueMax="15.0" valueMin="3.0">3–15 cm</quantity>
|
||
with a petiole up to
|
||
<quantity id="4CA89B63FFFBFFE5D5158B59FBEEE575" box="[1079,1150,1885,1911]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="176" unit="cm" value="18.0">18 cm</quantity>
|
||
; segments ca. 8–12, simple or 1-pinnatisect, attenuate, frequently subtended or intermitted by smaller, simple segments. Simple segments of basal leaves elliptical to narrowly oblong-lanceolate or oblanceolate, up to 10 ×
|
||
<quantity id="4CA89B63FFFBFFE5D2E38BA1FC6AE5BD" box="[961,1018,1957,1983]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="176" unit="cm" value="1.0">1 cm</quantity>
|
||
, subacute-mucronate to acute; margins subentire to irregularly denticulate, rarely lobed. Middle and upper cauline leaves gradually reduced in size and lobe number, the uppermost simple. Capitula 3–3.8 ×
|
||
<quantity id="4CA89B63FFFBFFE5D39E8BE9FCAFEA05" box="[700,831,2029,2056]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.4" metricValueMin="2.5" pageId="3" pageNumber="176" unit="cm" value="2.95" valueMax="3.4" valueMin="2.5">2.5–3.4 cm</quantity>
|
||
; involucre ovoid to ellipsoid-ovoid, 1.8–2.6 ×
|
||
<quantity id="4CA89B63FFFBFFE5D4758BE9FF3EEA29" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="3" pageNumber="176" unit="cm" value="1.2" valueMax="1.5" valueMin="0.9">0.9–1.5 cm</quantity>
|
||
; receptacle
|
||
<quantity id="4CA89B63FFFBFFE5D0108415FE1FEA29" box="[306,399,2065,2092]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="3" pageNumber="176" unit="mm" value="7.0" valueMax="9.0" valueMin="5.0">5–9 mm</quantity>
|
||
in diameter, densely setose; setae
|
||
<quantity id="4CA89B63FFFBFFE5D2328415FCEAEA29" box="[784,890,2065,2092]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="11.0" metricValueMin="5.0" pageId="3" pageNumber="176" unit="mm" value="8.0" valueMax="11.0" valueMin="5.0">5–11 mm</quantity>
|
||
long, caducous in fruiting stage; involucral bracts in several rows, imbricate, coriaceous, with appendages ± covering the bracts; middle bracts 0.8–1.8 ×
|
||
<quantity id="4CA89B63FFFCFFE2D40E8C9AFA3BE2BB" box="[1324,1451,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="4" pageNumber="177" unit="cm" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 cm</quantity>
|
||
(incl. appendage); appendage of middle bracts slightly convex, spiny, fimbriate, decurrent, straw-colored to brownish, 0.47–1.32 ×
|
||
<quantity id="4CA89B63FFFCFFE2D03B8CE2FE26E303" box="[281,438,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" metricValueMax="13.5" metricValueMin="5.699999999999999" pageId="4" pageNumber="177" unit="cm" value="0.96" valueMax="1.35" valueMin="0.57">0.57–1.35 cm</quantity>
|
||
, with (5-)8–14 cilia
|
||
<quantity id="4CA89B63FFFCFFE2D3828CE2FCABE303" box="[672,827,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="5.1" metricValueMin="1.6" pageId="4" pageNumber="177" unit="cm" value="0.335" valueMax="0.51" valueMin="0.16">0.16–0.51 cm</quantity>
|
||
in length on each side and a spine
|
||
<quantity id="4CA89B63FFFCFFE2D5ED8CE2FAC2E303" box="[1231,1362,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="8.0" metricValueMin="1.0" pageId="4" pageNumber="177" unit="cm" value="0.45" valueMax="0.8" valueMin="0.1">0.1–0.8 cm</quantity>
|
||
. Florets purplish; marginal, sterile florets 10–18,
|
||
<quantity id="4CA89B63FFFCFFE2D3778D0EFD43E327" box="[597,723,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.2" metricValueMin="2.2" pageId="4" pageNumber="177" unit="cm" value="2.7" valueMax="3.2" valueMin="2.2">2.2–3.2 cm</quantity>
|
||
in length, radiant, with 3–5(–6) lobes ca.
|
||
<quantity id="4CA89B63FFFCFFE2D58F8D0EFB74E327" box="[1197,1252,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="177" unit="cm" value="1.0">1 cm</quantity>
|
||
in length; central florets 23–39, hermaphrodite, tubular, expanded into 5-lobed corolla,
|
||
<quantity id="4CA89B63FFFCFFE2D2928D2AFBA1E34B" box="[944,1073,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="3.4" metricValueMin="2.3" pageId="4" pageNumber="177" unit="cm" value="2.8499999999999996" valueMax="3.4" valueMin="2.3">2.3–3.4 cm</quantity>
|
||
in length; anthers syngenesious, oblong, ca.
|
||
<quantity id="4CA89B63FFFCFFE2D0328D56FEC3E36F" box="[272,339,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="177" unit="mm" value="4.0">4 mm</quantity>
|
||
in length. Achenes straw-colored to dark greyish with caducous hairs and a lateral hilum. Outer achenes oblong to narrowly oblong 3.2–4.9 ×
|
||
<quantity id="4CA89B63FFFCFFE2D3AB8D72FC82E393" box="[649,786,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.6" metricValueMin="0.9" pageId="4" pageNumber="177" unit="mm" value="1.25" valueMax="1.6" valueMin="0.9">0.9–1.6 mm</quantity>
|
||
; pappus of outer achenes consisting of one series of smooth hairs,
|
||
<quantity id="4CA89B63FFFCFFE2D1EF8D9EFEAFE3B7" box="[205,319,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="4.0" metricValueMin="1.2" pageId="4" pageNumber="177" unit="mm" value="2.6" valueMax="4.0" valueMin="1.2">1.2–4 mm</quantity>
|
||
in length. Central (fertile) achenes ovoid-oblong, 4.1–5.3 ×
|
||
<quantity id="4CA89B63FFFCFFE2D2C78D9EFBFFE3B7" box="[997,1135,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.8" metricValueMin="1.9" pageId="4" pageNumber="177" unit="mm" value="2.3499999999999996" valueMax="2.8" valueMin="1.9">1.9–2.8 mm</quantity>
|
||
, with a pappus of two rows, the inner row consisting of smooth hairs, the outer row consisting of scabrous hairs; larger hairs of inner row
|
||
<quantity id="4CA89B63FFFCFFE2D4758DBAFF27E3FF" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="4" pageNumber="177" unit="mm" value="3.5" valueMax="4.5" valueMin="2.5">2.5–4.5 mm</quantity>
|
||
; larger hairs of outer row
|
||
<quantity id="4CA89B63FFFCFFE2D0FB8DE6FDE0E3FF" box="[473,624,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.5" metricValueMin="7.5" pageId="4" pageNumber="177" unit="mm" value="9.5" valueMax="11.5" valueMin="7.5">7.5–11.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2F660EFFFCFFE2D1AA89CFFCF9E7E0" ID-DOI="http://doi.org/10.5281/zenodo.7645655" ID-Zenodo-Dep="7645655" box="[136,873,1483,1506]" httpUri="https://zenodo.org/record/7645655/files/figure.png" pageId="4" pageNumber="177" startId="4.[136,229,1483,1505]" targetBox="[152,1432,543,1452]" targetPageId="4">
|
||
<paragraph id="8BEF3686FFFCFFE2D1AA89CFFCF9E7E0" blockId="4.[136,873,1483,1506]" box="[136,873,1483,1506]" pageId="4" pageNumber="177">
|
||
<emphasis id="B924EA94FFFCFFE2D1AA89CFFF6EE7E2" bold="true" box="[136,254,1483,1505]" pageId="4" pageNumber="177">FIGURE 2.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C504D05FFFCFFE2D05F89C8FDEFE7E3" authority="(UPA)" baseAuthorityName="UPA" box="[381,639,1484,1505]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="177" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFCFFE2D05F89C8FDABE7E3" box="[381,571,1484,1505]" italics="true" pageId="4" pageNumber="177">Centaurea dolopica</emphasis>
|
||
(UPA)
|
||
</taxonomicName>
|
||
mounted on two sheets.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34A650DFFFCFFE2D19F8A14FB9BE56C" pageId="4" pageNumber="177" type="etymology">
|
||
<paragraph id="8BEF3686FFFCFFE2D19F8A14FEC8E44C" blockId="4.[136,1452,1552,2083]" pageId="4" pageNumber="177">
|
||
<emphasis id="B924EA94FFFCFFE2D19F8A14FED8E428" bold="true" box="[189,328,1552,1578]" pageId="4" pageNumber="177">Etymology:</emphasis>
|
||
—The specific epithet of the new species refers to Dolopia, the ancient name of the region where the plants were found.
|
||
</paragraph>
|
||
<paragraph id="8BEF3686FFFCFFE2D19F8A5CFE4AE494" blockId="4.[136,1452,1552,2083]" pageId="4" pageNumber="177">
|
||
<emphasis id="B924EA94FFFCFFE2D19F8A5CFED3E470" bold="true" box="[189,323,1624,1650]" pageId="4" pageNumber="177">Phenology:</emphasis>
|
||
—The first capitula open in early July and anthesis peaks in mid-July followed by fruiting of the plants in late July and August.
|
||
</paragraph>
|
||
<paragraph id="8BEF3686FFFCFFE2D19F8AA4FB9BE56C" blockId="4.[136,1452,1552,2083]" pageId="4" pageNumber="177">
|
||
<emphasis id="B924EA94FFFCFFE2D19F8AA4FED5E4B8" bold="true" box="[189,325,1696,1722]" pageId="4" pageNumber="177">Karyology:</emphasis>
|
||
—The karyotype formula of
|
||
<taxonomicName id="4C504D05FFFCFFE2D38C8AA4FC05E4B8" box="[686,917,1696,1722]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="177" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFCFFE2D38C8AA4FC05E4B8" box="[686,917,1696,1722]" italics="true" pageId="4" pageNumber="177">Centaurea dolopica</emphasis>
|
||
</taxonomicName>
|
||
is 2
|
||
<emphasis id="B924EA94FFFCFFE2D2F48AA5FC74E4B8" box="[982,996,1697,1722]" italics="true" pageId="4" pageNumber="177">n</emphasis>
|
||
=
|
||
<quantity id="4CA89B63FFFCFFE2D5378AA4FBA6E4B8" box="[1045,1078,1696,1722]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="177" unit="m" value="8.0">8m</quantity>
|
||
+2m-SAT+2sm+2sm-SAT = 22 chromosomes. The karyotype of the new species is diploid (2
|
||
<emphasis id="B924EA94FFFCFFE2D21A8AC1FCD6E4DC" box="[824,838,1733,1758]" italics="true" pageId="4" pageNumber="177">n</emphasis>
|
||
= 2x = 22) and symmetrical. It consists of metacentric (m) and submetacentric (sm) chromosomes (
|
||
<figureCitation id="136B2A03FFFCFFE2D3548AECFD27E501" box="[630,695,1768,1795]" captionStart="FIGURE 5" captionStartId="8.[136,229,586,608]" captionTargetBox="[276,1312,190,552]" captionTargetId="figure-29@8.[276,1312,190,552]" captionTargetPageId="8" captionText="FIGURE 5. A) Karyotype and B) idiogram of Centaurea dolopica. Arrows indicate satellites." figureDoi="http://doi.org/10.5281/zenodo.7645665" httpUri="https://zenodo.org/record/7645665/files/figure.png" pageId="4" pageNumber="177">Fig. 5</figureCitation>
|
||
). The 3rd largest in size chromosome pair is submetacentric. The 8th and 9th chromosome pairs are submetacentric and metacentric, respectively, whereas both are bearing small spherical satellites. The size of the chromosomes varies between 1.51–3.49 μm and the average chromosome length (ACL) equals to 2.43 μm. THL and TCL equal to 26.77 μm and 53.54 μm, respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34A650DFFFCFFECD19F8B7CFD50E73A" lastPageId="10" lastPageNumber="183" pageId="4" pageNumber="177" type="distribution">
|
||
<paragraph id="8BEF3686FFFCFFE3D19F8B7CFE63E0D7" blockId="4.[136,1452,1552,2083]" lastBlockId="5.[136,1452,158,725]" lastPageId="5" lastPageNumber="178" pageId="4" pageNumber="177">
|
||
<emphasis id="B924EA94FFFCFFE2D19F8B7CFE7AE590" bold="true" box="[189,490,1912,1938]" pageId="4" pageNumber="177">Distribution and habitat:</emphasis>
|
||
—Based on the current knowledge,
|
||
<taxonomicName id="4C504D05FFFCFFE2D25C8B7CFBCCE590" box="[894,1116,1912,1938]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="177" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFCFFE2D25C8B7CFBCCE590" box="[894,1116,1912,1938]" italics="true" pageId="4" pageNumber="177">Centaurea dolopica</emphasis>
|
||
</taxonomicName>
|
||
is restricted to a single, small population located along a provincial road ca.
|
||
<quantity id="4CA89B63FFFCFFE2D3AF8B98FD44E5B4" box="[653,724,1948,1974]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="177" unit="km" value="10.0">10 km</quantity>
|
||
west of the settlement of Omvriaki in Nomos Fthiotidos,
|
||
<collectingRegion id="4994F864FFFCFFE2D47B8B98FF4BE5D8" country="Greece" name="Sterea Ellada" pageId="4" pageNumber="177">Central Greece</collectingRegion>
|
||
. The substrate of the gently hilly landscape is ophiolitic and the vegetation is characterized by a
|
||
<taxonomicName id="4C504D05FFFCFFE2D46C8BC4FE61E5FD" authority="Linnaeus (1753: 995)" authorityName="Linnaeus" authorityPageNumber="995" authorityYear="1753" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="177" phylum="Tracheophyta" rank="species" species="coccifera">
|
||
<emphasis id="B924EA94FFFCFFE2D46C8BC4FF60E5FC" italics="true" pageId="4" pageNumber="177">Quercus coccifera</emphasis>
|
||
<bibRefCitation id="EFC14B77FFFCFFE2D1D88BE0FE61E5FD" author="Linnaeus, C." box="[250,497,2020,2047]" pageId="4" pageNumber="177" refId="ref7954" refString="Linnaeus, C. (1753) Species plantarum. L. Salvius, Holmia [Stockholm], 1200 pp." type="book" year="1753">Linnaeus (1753: 995)</bibRefCitation>
|
||
</taxonomicName>
|
||
partly evergreen scrub (pseudomaquis). The
|
||
<taxonomicName id="4C504D05FFFCFFE2D2DF8BE0FBE2E5FC" authorityName="Zografidis & Koutroumpa & Liveri & Dimopoulos" authorityYear="2023" box="[1021,1138,2020,2046]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="177" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B924EA94FFFCFFE2D2DF8BE0FBE2E5FC" box="[1021,1138,2020,2046]" italics="true" pageId="4" pageNumber="177">Centaurea</emphasis>
|
||
</taxonomicName>
|
||
plants grow mainly on the roadsides and the rocky cut slope (
|
||
<figureCitation id="136B2A03FFFCFFE2D329840CFDECEA20" box="[523,636,2056,2082]" captionStart="FIGURE 3" captionStartId="5.[136,229,1620,1642]" captionTargetBox="[152,1435,755,1586]" captionTargetId="figure-317@5.[143,1444,749,1595]" captionTargetPageId="5" captionText="FIGURE 3. Centaurea dolopica plants in their natural habitat: (A) several large plants along a roadside and (B) on a cut slope. (C) Eryngium serbicum in the same habitat." figureDoi="http://doi.org/10.5281/zenodo.7645657" httpUri="https://zenodo.org/record/7645657/files/figure.png" pageId="4" pageNumber="177">Fig. 3A,B</figureCitation>
|
||
), whereas only a few plants were observed inside the surrounding denser scrub. We note that cut slopes—resembling natural steep slopes—and roadsides are frequently inhabited by this photophilous and often chasmophytic genus, and the plants may be more sporadic or even absent from the surrounding, undisturbed vegetation (e.g.,
|
||
<bibRefCitation id="EFC14B77FFFDFFE3D0EE8CE2FD00E302" author="Phitos, D. & Constantinidis, Th. & Kamari, G." box="[460,656,230,257]" pageId="5" pageNumber="178" refId="ref8273" refString="Phitos, D., Constantinidis, Th. & Kamari, G. (Eds.) (2009) The Red Data Book of rare and threatened plants of Greece, vol. 1. Hellenic Botanical Society, Patras, 405 pp. [In Greek]" type="book" year="2009">
|
||
Phitos
|
||
<emphasis id="B924EA94FFFDFFE3D33A8CE3FDD8E302" box="[536,584,230,256]" italics="true" pageId="5" pageNumber="178">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFFDFFE3D3B88CE2FC1EE302" author="Zografidis, A. & Bancheva, S. & Tan, K." box="[666,910,230,256]" pageId="5" pageNumber="178" pagination="57 - 63" refId="ref9201" refString="Zografidis, A., Bancheva, S. & Tan, K. (2014) A new yellow-flowered species of Centaurea (Asteraceae) from Mt Imittos, Attica, Greece. Phytologia Balcanica 20 (1): 57 - 63." type="journal article" year="2014">
|
||
Zografidis
|
||
<emphasis id="B924EA94FFFDFFE3D2368CE3FCDCE302" box="[788,844,230,256]" italics="true" pageId="5" pageNumber="178">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
). Interestingly, among the angiosperms observed and collected in the habitat of
|
||
<taxonomicName id="4C504D05FFFDFFE3D0F98D0FFDCEE326" box="[475,606,266,293]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFDFFE3D0F98D0FFDCEE326" box="[475,606,266,293]" italics="true" pageId="5" pageNumber="178">C. dolopica</emphasis>
|
||
</taxonomicName>
|
||
, was
|
||
<taxonomicName id="4C504D05FFFDFFE3D3BD8D0FFBC7E327" authority="Pancic (1856: 520)" authorityName="Pancic" authorityPageNumber="520" authorityYear="1856" box="[671,1111,266,293]" class="Magnoliopsida" family="Apiaceae" genus="Eryngium" kingdom="Plantae" order="Apiales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="serbicum">
|
||
<emphasis id="B924EA94FFFDFFE3D3BD8D0FFCEBE326" box="[671,891,266,292]" italics="true" pageId="5" pageNumber="178">Eryngium serbicum</emphasis>
|
||
<bibRefCitation id="EFC14B77FFFDFFE3D2A08D0EFBC7E327" author="Pancic, J." box="[898,1111,266,293]" pageId="5" pageNumber="178" pagination="475 - 598" refId="ref8107" refString="Pancic, J. (1856) Verzeichniss der in Serbien wildwachsenden Phanerogamen. Verhandlungen des Zoologisch-Botanischen Vereins in Wien 6: 475 - 598." type="journal article" year="1856">Pančić (1856: 520)</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136B2A03FFFDFFE3D5458D0EFB2CE326" box="[1127,1212,266,292]" captionStart="FIGURE 3" captionStartId="5.[136,229,1620,1642]" captionTargetBox="[152,1435,755,1586]" captionTargetId="figure-317@5.[143,1444,749,1595]" captionTargetPageId="5" captionText="FIGURE 3. Centaurea dolopica plants in their natural habitat: (A) several large plants along a roadside and (B) on a cut slope. (C) Eryngium serbicum in the same habitat." figureDoi="http://doi.org/10.5281/zenodo.7645657" httpUri="https://zenodo.org/record/7645657/files/figure.png" pageId="5" pageNumber="178">Fig. 3C</figureCitation>
|
||
) (voucher
|
||
<emphasis id="B924EA94FFFDFFE3D4158D0EFF22E34A" italics="true" pageId="5" pageNumber="178">Zografidis 743</emphasis>
|
||
UPA!), which represents the first confirmed report from
|
||
<collectingCountry id="F3477616FFFDFFE3D2688D2AFC09E34A" box="[842,921,302,328]" name="Greece" pageId="5" pageNumber="178">Greece</collectingCountry>
|
||
of this conspicuous and characteristic Balkan endemic species. Since the new
|
||
<taxonomicName id="4C504D05FFFDFFE3D0D28D57FDF5E36F" authorityName="Zografidis & Koutroumpa & Liveri & Dimopoulos" authorityYear="2023" box="[496,613,339,365]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B924EA94FFFDFFE3D0D28D57FDF5E36F" box="[496,613,339,365]" italics="true" pageId="5" pageNumber="178">Centaurea</emphasis>
|
||
</taxonomicName>
|
||
and the eryngo species could easily be observed by any botanist traversing this location west of Omvriaki during mid to late summer, the fact that these two species were only currently recorded indicate that the region must have long remained floristically unexplored. A collection by
|
||
<emphasis id="B924EA94FFFDFFE3D5B18D9FFAEDE3B6" box="[1171,1405,410,436]" italics="true" pageId="5" pageNumber="178">Hartvig et al. 10541</emphasis>
|
||
(C) from north of Trilofon, assigned to
|
||
<taxonomicName id="4C504D05FFFDFFE3D3318DBBFD10E3DA" box="[531,640,446,473]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFFDFFE3D3318DBBFD10E3DA" box="[531,640,446,473]" italics="true" pageId="5" pageNumber="178">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFC14B77FFFDFFE3D3888DBAFB8AE3DB" author="Wagenitz, G. & Gamal-Eldin, E." box="[682,1050,446,473]" pageId="5" pageNumber="178" pagination="95 - 127" refId="ref9164" refString="Wagenitz, G. & Gamal-Eldin, E. (1985) Zur Kenntnis der griechischen Centaurea - Arten der sektion Acrokentron. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 107: 95 - 127." type="journal article" year="1985">Wagenitz & Gamal-Eldin (1985)</bibRefCitation>
|
||
, was of interest to be studied due to its geographical proximity to
|
||
<taxonomicName id="4C504D05FFFDFFE3D0FB8DE7FDCDE3FE" box="[473,605,482,509]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFDFFE3D0FB8DE7FDCDE3FE" box="[473,605,482,509]" italics="true" pageId="5" pageNumber="178">C. dolopica</emphasis>
|
||
</taxonomicName>
|
||
, namely just ca.
|
||
<quantity id="4CA89B63FFFDFFE3D2038DE6FCF9E3FE" box="[801,873,482,508]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="178" unit="km" value="10.0">10 km</quantity>
|
||
south of the
|
||
<emphasis id="B924EA94FFFDFFE3D5238DE6FB3AE3FE" box="[1025,1194,482,508]" italics="true" pageId="5" pageNumber="178">locus classicus</emphasis>
|
||
. However, despite our exhaustive search in herbarium C, we could not trace this material and neither did we succeed in finding the respective population of plants in the field. The examination of a duplicate specimen of the aforementioned collection kept in a private herbarium indicated that the taxonomic identity of the specimen is unsettled due to the fact that it was collected late in the season, perhaps representing a regrowth after the plant had been cut or bitten down, and is in poor condition (Arne Strid, pers. comm.). A more detailed field exploration is needed to locate this population of
|
||
<taxonomicName id="4C504D05FFFDFFE3D5C58E93FACCE0B3" authorityName="Zografidis & Koutroumpa & Liveri & Dimopoulos" authorityYear="2023" box="[1255,1372,663,689]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B924EA94FFFDFFE3D5C58E93FACCE0B3" box="[1255,1372,663,689]" italics="true" pageId="5" pageNumber="178">Centaurea</emphasis>
|
||
</taxonomicName>
|
||
and to re-examine its taxonomic status.
|
||
</paragraph>
|
||
<caption id="DF2F660EFFFDFFE3D1AA8A50FD97E48C" ID-DOI="http://doi.org/10.5281/zenodo.7645657" ID-Zenodo-Dep="7645657" httpUri="https://zenodo.org/record/7645657/files/figure.png" pageId="5" pageNumber="178" startId="5.[136,229,1620,1642]" targetBox="[152,1435,755,1586]" targetPageId="5">
|
||
<paragraph id="8BEF3686FFFDFFE3D1AA8A50FD97E48C" blockId="5.[136,1451,1620,1678]" pageId="5" pageNumber="178">
|
||
<emphasis id="B924EA94FFFDFFE3D1AA8A50FE90E46B" bold="true" box="[136,256,1620,1642]" pageId="5" pageNumber="178">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C504D05FFFDFFE3D02B8A51FE5AE46B" box="[265,458,1620,1642]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFDFFE3D02B8A51FE5AE46B" box="[265,458,1620,1642]" italics="true" pageId="5" pageNumber="178">Centaurea dolopica</emphasis>
|
||
</taxonomicName>
|
||
plants in their natural habitat: (
|
||
<emphasis id="B924EA94FFFDFFE3D22A8A50FC89E46B" bold="true" box="[776,793,1620,1641]" pageId="5" pageNumber="178">A</emphasis>
|
||
) several large plants along a roadside and (
|
||
<emphasis id="B924EA94FFFDFFE3D5EC8A50FB4EE46B" bold="true" box="[1230,1246,1620,1641]" pageId="5" pageNumber="178">B</emphasis>
|
||
) on a cut slope. (
|
||
<emphasis id="B924EA94FFFDFFE3D4B08A50FA33E468" bold="true" box="[1426,1443,1620,1642]" pageId="5" pageNumber="178">C</emphasis>
|
||
)
|
||
<taxonomicName id="4C504D05FFFDFFE3D1AA8A7DFED5E48F" authorityName="Pancic" authorityYear="1856" box="[136,325,1656,1678]" class="Magnoliopsida" family="Apiaceae" genus="Eryngium" kingdom="Plantae" order="Apiales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="serbicum">
|
||
<emphasis id="B924EA94FFFDFFE3D1AA8A7DFED5E48F" box="[136,325,1656,1678]" italics="true" pageId="5" pageNumber="178">Eryngium serbicum</emphasis>
|
||
</taxonomicName>
|
||
in the same habitat.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEF3686FFFDFFE0D19F8AB8FEF6E326" blockId="5.[136,1452,1724,2075]" lastBlockId="6.[136,1451,158,292]" lastPageId="6" lastPageNumber="179" pageId="5" pageNumber="178">
|
||
<emphasis id="B924EA94FFFDFFE3D19F8AB8FE64E4D5" bold="true" box="[189,500,1724,1751]" pageId="5" pageNumber="178">Conservation assessment:</emphasis>
|
||
—The sole known population of the new species is very small and restricted. The plants were found along ca.
|
||
<quantity id="4CA89B63FFFDFFE3D0EF8AE5FD86E4F9" box="[461,534,1761,1787]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="178" unit="m" value="300.0">300 m</quantity>
|
||
of the provincial road (see above), accounting for an area of occupancy of well below
|
||
<quantity id="4CA89B63FFFDFFE3D1F68B01FE9DE51D" box="[212,269,1797,1823]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="178" unit="km" value="1.0">1 km</quantity>
|
||
<superScript id="7C259BCEFFFDFFE3D02F8B00FE85E510" attach="left" box="[269,277,1796,1810]" fontSize="6" pageId="5" pageNumber="178">2</superScript>
|
||
. Approximately 70 mature and
|
||
<specimenCount id="9D56FD0FFFFDFFE3D3598B01FC99E51D" box="[635,777,1797,1823]" pageId="5" pageNumber="178" type="immature">40 immature</specimenCount>
|
||
plants were counted.
|
||
<taxonomicName id="4C504D05FFFDFFE3D2DD8B01FB4EE51D" box="[1023,1246,1797,1823]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFDFFE3D2DD8B01FB4EE51D" box="[1023,1246,1797,1823]" italics="true" pageId="5" pageNumber="178">Centaurea dolopica</emphasis>
|
||
</taxonomicName>
|
||
was only recently observed for the first time and no estimation on the size of its population in the past can be drawn. However, given that
|
||
<taxonomicName id="4C504D05FFFDFFE3D1AA8B49FEA1E565" box="[136,305,1869,1895]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="B924EA94FFFDFFE3D1AA8B49FF6DE565" box="[136,253,1869,1895]" italics="true" pageId="5" pageNumber="178">Centaurea</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
of section
|
||
<taxonomicName id="4C504D05FFFDFFE3D08B8B49FDA3E565" box="[425,563,1869,1895]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="section" section="Acrocentron">
|
||
<emphasis id="B924EA94FFFDFFE3D08B8B49FDA3E565" box="[425,563,1869,1895]" italics="true" pageId="5" pageNumber="178">Acrocentron</emphasis>
|
||
</taxonomicName>
|
||
are impressive herbs and sought for by professional and amateur botanists alike, it can be assumed that the new species has been a very rare plant throughout the last century at least.
|
||
<taxonomicName id="4C504D05FFFDFFE3D5FD8B75FA3BE589" authorityName="Linnaeus" authorityYear="1753" box="[1247,1451,1905,1931]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="178" phylum="Tracheophyta" rank="species" species="coccifera">
|
||
<emphasis id="B924EA94FFFDFFE3D5FD8B75FA3BE589" box="[1247,1451,1905,1931]" italics="true" pageId="5" pageNumber="178">Quercus coccifera</emphasis>
|
||
</taxonomicName>
|
||
pseudomaquis in
|
||
<collectingCountry id="F3477616FFFDFFE3D0708B91FE31E5AD" box="[338,417,1941,1967]" name="Greece" pageId="5" pageNumber="178">Greece</collectingCountry>
|
||
has generally been considered secondary vegetation (
|
||
<bibRefCitation id="EFC14B77FFFDFFE3D52D8B91FB6CE5AD" author="Chasapis, M. & Karagiannakidou, V. & Theodoropoulos, K." box="[1039,1276,1941,1967]" pageId="5" pageNumber="178" pagination="357 - 381" refId="ref7008" refString="Chasapis, M., Karagiannakidou, V. & Theodoropoulos, K. (2004) Phytosociological research of Quercus coccifera L. pseudomaquis on Mount Chortiatis, northern Greece. Israel Journal of Plant Sciences 52: 357 - 381. https: // doi. org / 10.1560 / N 0 YU-G 77 C-TLKX-HW 5 K" type="journal article" year="2004">
|
||
Chasapis
|
||
<emphasis id="B924EA94FFFDFFE3D5A28B91FB2BE5AD" box="[1152,1211,1941,1967]" italics="true" pageId="5" pageNumber="178">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
and references therein), but no indication of ongoing decline of the quality of habitat (including grazing) was observed or estimated. The widening of the provincial road would certainly be detrimental for the plants that grow along roadsides, but road upgrading construction works are considered unlikely at this infrequently visited location. However, solely on account of its very small population, the new species should be listed as Endangered: En D (
|
||
<bibRefCitation id="EFC14B77FFFEFFE0D50C8C9AFB28E2BA" author="IUCN" box="[1070,1208,158,184]" pageId="6" pageNumber="179" refId="ref7725" refString="IUCN (2022) The IUCN Red List of Threatened Species. Version 2021 - 3. Available from: https: // www. iucnredlist. org (accessed 23 September 2022)" type="url" year="2022">IUCN 2022</bibRefCitation>
|
||
). It is noteworthy that
|
||
<taxonomicName id="4C504D05FFFEFFE0D1AA8CC7FE95E2DE" box="[136,261,194,221]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="aetolica">
|
||
<emphasis id="B924EA94FFFEFFE0D1AA8CC7FE95E2DE" box="[136,261,194,221]" italics="true" pageId="6" pageNumber="179">C. aetolica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C504D05FFFEFFE0D01D8CC7FE73E2DE" box="[319,483,194,221]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="corinthiaca">
|
||
<emphasis id="B924EA94FFFEFFE0D01D8CC7FE73E2DE" box="[319,483,194,221]" italics="true" pageId="6" pageNumber="179">C. corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
(as
|
||
<taxonomicName id="4C504D05FFFEFFE0D3378CC7FCCEE2DE" authorityName="Phitos & Georgiadis" authorityYear="1981" box="[533,862,194,221]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="subSpecies" species="achaia" subSpecies="corinthiaca">
|
||
<emphasis id="B924EA94FFFEFFE0D3378CC7FD14E2DE" box="[533,644,194,221]" italics="true" pageId="6" pageNumber="179">C. achaia</emphasis>
|
||
subsp.
|
||
<emphasis id="B924EA94FFFEFFE0D3FF8CC6FCCEE2DE" box="[733,862,194,220]" italics="true" pageId="6" pageNumber="179">corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
) from the
|
||
<taxonomicName id="4C504D05FFFEFFE0D2F98CC7FBD9E2DE" box="[987,1097,194,221]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFFEFFE0D2F98CC7FBD9E2DE" box="[987,1097,194,221]" italics="true" pageId="6" pageNumber="179">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
group as well as additional 16 species and 3 subspecies of
|
||
<taxonomicName id="4C504D05FFFEFFE0D0EB8CE3FDAEE303" authorityName="Zografidis & Koutroumpa & Liveri & Dimopoulos" authorityYear="2023" box="[457,574,231,257]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B924EA94FFFEFFE0D0EB8CE3FDAEE303" box="[457,574,231,257]" italics="true" pageId="6" pageNumber="179">Centaurea</emphasis>
|
||
</taxonomicName>
|
||
are included in the Red Data Book of Rare and Threatened Plants of
|
||
<collectingCountry id="F3477616FFFEFFE0D47E8CE2FA3BE302" box="[1372,1451,230,256]" name="Greece" pageId="6" pageNumber="179">Greece</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFC14B77FFFEFFE0D1B28D0EFECAE326" author="Phitos, D. & Constantinidis, Th. & Kamari, G." box="[144,346,266,292]" pageId="6" pageNumber="179" refId="ref8273" refString="Phitos, D., Constantinidis, Th. & Kamari, G. (Eds.) (2009) The Red Data Book of rare and threatened plants of Greece, vol. 1. Hellenic Botanical Society, Patras, 405 pp. [In Greek]" type="book" year="2009">
|
||
Phitos
|
||
<emphasis id="B924EA94FFFEFFE0D1FC8D0FFE87E326" box="[222,279,266,292]" italics="true" pageId="6" pageNumber="179">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF2F660EFFFEFFE0D1AA880FFCE3E66A" ID-DOI="http://doi.org/10.5281/zenodo.7645661" ID-Zenodo-Dep="7645661" httpUri="https://zenodo.org/record/7645661/files/figure.png" pageId="6" pageNumber="179" startId="6.[136,229,1035,1057]" targetBox="[155,1433,317,1010]" targetPageId="6">
|
||
<paragraph id="8BEF3686FFFEFFE0D1AA880FFCE3E66A" blockId="6.[136,1451,1035,1129]" pageId="6" pageNumber="179">
|
||
<emphasis id="B924EA94FFFEFFE0D1AA880FFF6EE622" bold="true" box="[136,254,1035,1057]" pageId="6" pageNumber="179">FIGURE 4.</emphasis>
|
||
Habits (upper-case letters) and flower-heads on approximate same scale (lower-case letters) of
|
||
<taxonomicName id="4C504D05FFFEFFE0D5BB880FFA3BE622" box="[1177,1451,1035,1056]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="section" section="Acrocentron">
|
||
<emphasis id="B924EA94FFFEFFE0D5BB880FFB6DE622" box="[1177,1277,1035,1056]" italics="true" pageId="6" pageNumber="179">Centaurea</emphasis>
|
||
sect.
|
||
<emphasis id="B924EA94FFFEFFE0D417880FFA3BE622" box="[1333,1451,1035,1056]" italics="true" pageId="6" pageNumber="179">Acrocentron</emphasis>
|
||
</taxonomicName>
|
||
representatives from Greece belonging to the
|
||
<taxonomicName id="4C504D05FFFEFFE0D367882BFD34E646" box="[581,676,1071,1092]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFFEFFE0D367882BFD34E646" box="[581,676,1071,1092]" italics="true" pageId="6" pageNumber="179">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
group;
|
||
<emphasis id="B924EA94FFFEFFE0D3D3882BFC84E646" bold="true" box="[753,788,1071,1092]" pageId="6" pageNumber="179">A,a</emphasis>
|
||
:
|
||
<taxonomicName id="4C504D05FFFEFFE0D200882BFC6CE646" box="[802,1020,1071,1092]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="corinthiaca">
|
||
<emphasis id="B924EA94FFFEFFE0D200882BFC6CE646" box="[802,1020,1071,1092]" italics="true" pageId="6" pageNumber="179">Centaurea corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
(photos by Katerina Goula);
|
||
<emphasis id="B924EA94FFFEFFE0D43E882BFAAFE646" bold="true" box="[1308,1343,1071,1092]" pageId="6" pageNumber="179">B,b</emphasis>
|
||
:
|
||
<taxonomicName id="4C504D05FFFEFFE0D46F882BFA3BE646" box="[1357,1451,1071,1092]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFFEFFE0D46F882BFA3BE646" box="[1357,1451,1071,1092]" italics="true" pageId="6" pageNumber="179">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
(photos by Katerina Goula);
|
||
<emphasis id="B924EA94FFFEFFE0D0BE8857FE2EE66B" bold="true" box="[412,446,1107,1129]" pageId="6" pageNumber="179">C,c</emphasis>
|
||
:
|
||
<taxonomicName id="4C504D05FFFEFFE0D0E98857FDA3E66A" box="[459,563,1107,1128]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="euboica">
|
||
<emphasis id="B924EA94FFFEFFE0D0E98857FDA3E66A" box="[459,563,1107,1128]" italics="true" pageId="6" pageNumber="179">C. euboica</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B924EA94FFFEFFE0D31D8857FDF4E66A" bold="true" box="[575,612,1107,1128]" pageId="6" pageNumber="179">D,d</emphasis>
|
||
,
|
||
<taxonomicName id="4C504D05FFFEFFE0D3528857FD49E66A" box="[624,729,1107,1128]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="aetolica">
|
||
<emphasis id="B924EA94FFFEFFE0D3528857FD49E66A" box="[624,729,1107,1128]" italics="true" pageId="6" pageNumber="179">C. aetolica</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B924EA94FFFEFFE0D3C48857FD61E66A" bold="true" box="[742,753,1107,1128]" pageId="6" pageNumber="179">e</emphasis>
|
||
:
|
||
<taxonomicName id="4C504D05FFFEFFE0D3DF8857FCFDE66A" box="[765,877,1107,1128]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFEFFE0D3DF8857FCFDE66A" box="[765,877,1107,1128]" italics="true" pageId="6" pageNumber="179">C. dolopica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEF3686FFFEFFE1D19F8893FAB3E34A" blockId="6.[136,1452,1175,2066]" lastBlockId="7.[136,1451,158,329]" lastPageId="7" lastPageNumber="180" pageId="6" pageNumber="179">
|
||
<emphasis id="B924EA94FFFEFFE0D19F8893FE5BE6B3" bold="true" box="[189,459,1175,1201]" pageId="6" pageNumber="179">Taxonomic comments:</emphasis>
|
||
—Based on its overall morphology, the new species belongs to a range-restricted, Greek endemic group including
|
||
<taxonomicName id="4C504D05FFFEFFE0D08488B8FDFEE6D7" box="[422,622,1211,1238]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFFEFFE0D08488B8FDFEE6D7" box="[422,622,1211,1238]" italics="true" pageId="6" pageNumber="179">Centaurea achaia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C504D05FFFEFFE0D35E88B8FD67E6D7" box="[636,759,1211,1238]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="aetolica">
|
||
<emphasis id="B924EA94FFFEFFE0D35E88B8FD67E6D7" box="[636,759,1211,1238]" italics="true" pageId="6" pageNumber="179">C. aetolica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C504D05FFFEFFE0D22788B8FC36E6D7" box="[773,934,1211,1238]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="corinthiaca">
|
||
<emphasis id="B924EA94FFFEFFE0D22788B8FC36E6D7" box="[773,934,1211,1238]" italics="true" pageId="6" pageNumber="179">C. corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C504D05FFFEFFE0D2FE88B8FACBE6D7" authority="Rechinger (1956: 102)" authorityName="Rechinger" authorityPageNumber="102" authorityYear="1956" box="[988,1371,1211,1238]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="euboica">
|
||
<emphasis id="B924EA94FFFEFFE0D2FE88B8FBC6E6D7" box="[988,1110,1211,1238]" italics="true" pageId="6" pageNumber="179">C. euboica</emphasis>
|
||
<bibRefCitation id="EFC14B77FFFEFFE0D57E88BFFACBE6D7" author="Rechinger, K. H." box="[1116,1371,1211,1238]" pageId="6" pageNumber="179" pagination="95 - 103" refId="ref8517" refString="Rechinger, K. H. (1956) Plantae novae ex insula Euboea. Anzeiger der Osterreichische Akademie der Wissenschaften, Mathematisch- Naturwissenschatliche Klasse 9: 95 - 103." type="journal article" year="1956">Rechinger (1956: 102)</bibRefCitation>
|
||
</taxonomicName>
|
||
, which has been recognized at subsectional rank by some authors viz.
|
||
<emphasis id="B924EA94FFFEFFE0D27088E4FCF5E6F8" box="[850,869,1248,1274]" italics="true" pageId="6" pageNumber="179">C</emphasis>
|
||
. subsect.
|
||
<taxonomicName id="4C504D05FFFEFFE0D2F788DBFAACE6FB" authority="Routsi & Th. Georgiadis" authorityName="Routsi & Th. Georgiadis" box="[981,1340,1247,1273]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFFEFFE0D2F788DBFBB4E6FB" box="[981,1060,1247,1273]" italics="true" pageId="6" pageNumber="179">Achaia</emphasis>
|
||
Routsi & Th.Georgiadis
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="EFC14B77FFFEFFE0D44088DBFF6EE71F" author="Routsi, E." pageId="6" pageNumber="179" refId="ref8730" refString="Routsi, E. (1993) Biosystematic study of the section Acrocentron (Cass.) DC. of the genus Centaurea L. in Greece. PhD Thesis, University of Patras, Greece." type="book" year="1993">Routsi (1993: 14)</bibRefCitation>
|
||
. Plants of this group are characterized by their pinnatisect to pinnate leaves with lanceolate to linear segments, whitish to purplish florets and convex, stramineous appendages of middle bracts that largely cover the nails of their neighboring bracts on the involucre and are usually armed with stout spines (
|
||
<bibRefCitation id="EFC14B77FFFEFFE0D2D1894FFAB0E767" author="Routsi, E. & Georgiadis, T." box="[1011,1312,1355,1381]" pageId="6" pageNumber="179" pagination="139 - 151" refId="ref8875" refString="Routsi, E. & Georgiadis, T. (1999) Cytogeographical study of Centaurea L. sect. Acrocentron (Cass.) DC. (Asteraceae) in Greece. Botanica Helvetica 109 (2): 139 - 151." type="journal article" year="1999">Routsi & Georgiadis 1999</bibRefCitation>
|
||
). The newly described species,
|
||
<taxonomicName id="4C504D05FFFEFFE0D07F8974FDAEE78B" box="[349,574,1391,1418]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFEFFE0D07F8974FDAEE78B" box="[349,574,1391,1418]" italics="true" pageId="6" pageNumber="179">Centaurea dolopica</emphasis>
|
||
</taxonomicName>
|
||
, is among the most distinct species in the group, owing to its tall habit with copious and characteristic capitula, which have fewer central florets and smaller (shorter and narrower) involucral bracts than any of the other species. Furthermore, the appendages of the bracts in the new species are not bearing stout spines but rather thin and short ones. Compared to
|
||
<taxonomicName id="4C504D05FFFEFFE0D35789D8FD71E7F7" box="[629,737,1499,1526]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFFEFFE0D35789D8FD71E7F7" box="[629,737,1499,1526]" italics="true" pageId="6" pageNumber="179">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136B2A03FFFEFFE0D3D389DFFCCDE7F7" box="[753,861,1499,1525]" captionStart="FIGURE 4" captionStartId="6.[136,229,1035,1057]" captionTargetBox="[155,1433,317,1010]" captionTargetId="figure-102@6.[155,1433,317,1010]" captionTargetPageId="6" captionText="FIGURE 4. Habits (upper-case letters) and flower-heads on approximate same scale (lower-case letters) of Centaurea sect. Acrocentron representatives from Greece belonging to the C. achaia group; A,a: Centaurea corinthiaca (photos by Katerina Goula); B,b: C. achaia (photos by Katerina Goula); C,c: C. euboica; D,d, C. aetolica; e: C. dolopica." figureDoi="http://doi.org/10.5281/zenodo.7645661" httpUri="https://zenodo.org/record/7645661/files/figure.png" pageId="6" pageNumber="179">Fig. 4B,b</figureCitation>
|
||
), which is the most widespread and morphologically variable species,
|
||
<taxonomicName id="4C504D05FFFEFFE0D0648A04FE57E41B" box="[326,455,1535,1562]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFEFFE0D0648A04FE57E41B" box="[326,455,1535,1562]" italics="true" pageId="6" pageNumber="179">C. dolopica</emphasis>
|
||
</taxonomicName>
|
||
additionally differs in its denser indumentum—that renders the plants ±greyish,—smaller capitula and shorter achenes.
|
||
<taxonomicName id="4C504D05FFFEFFE0D0FB8A20FD20E43F" box="[473,688,1571,1598]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="aetolica">
|
||
<emphasis id="B924EA94FFFEFFE0D0FB8A20FD20E43F" box="[473,688,1571,1598]" italics="true" pageId="6" pageNumber="179">Centaurea aetolica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136B2A03FFFEFFE0D3E08A27FCBEE43F" box="[706,814,1571,1598]" captionStart="FIGURE 4" captionStartId="6.[136,229,1035,1057]" captionTargetBox="[155,1433,317,1010]" captionTargetId="figure-102@6.[155,1433,317,1010]" captionTargetPageId="6" captionText="FIGURE 4. Habits (upper-case letters) and flower-heads on approximate same scale (lower-case letters) of Centaurea sect. Acrocentron representatives from Greece belonging to the C. achaia group; A,a: Centaurea corinthiaca (photos by Katerina Goula); B,b: C. achaia (photos by Katerina Goula); C,c: C. euboica; D,d, C. aetolica; e: C. dolopica." figureDoi="http://doi.org/10.5281/zenodo.7645661" httpUri="https://zenodo.org/record/7645661/files/figure.png" pageId="6" pageNumber="179">Fig. 4D,d</figureCitation>
|
||
) is sharply distinguished from all other members of the group by its uniformly cream-colored appendages of involucral bracts. The new species differs from
|
||
<taxonomicName id="4C504D05FFFEFFE0D4328A4CFA1CE463" box="[1296,1420,1607,1634]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="aetolica">
|
||
<emphasis id="B924EA94FFFEFFE0D4328A4CFA1CE463" box="[1296,1420,1607,1634]" italics="true" pageId="6" pageNumber="179">C. aetolica</emphasis>
|
||
</taxonomicName>
|
||
in having smaller capitula and lanceolate leaf segments (vs. ±linear).
|
||
<emphasis id="B924EA94FFFEFFE0D24C8A68FB6EE487" box="[878,1278,1643,1670]" italics="true" pageId="6" pageNumber="179">
|
||
<taxonomicName id="4C504D05FFFEFFE0D24C8A68FBFBE487" box="[878,1131,1643,1670]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" sensu="stricto" species="corinthiaca">Centaurea corinthiaca</taxonomicName>
|
||
<taxonomicNameLabel id="A21757EFFFFEFFE0D5538A68FB6EE487" box="[1137,1278,1644,1669]" pageId="6" pageNumber="179" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="136B2A03FFFEFFE0D42C8A6FFAE7E487" box="[1294,1399,1643,1669]" captionStart="FIGURE 4" captionStartId="6.[136,229,1035,1057]" captionTargetBox="[155,1433,317,1010]" captionTargetId="figure-102@6.[155,1433,317,1010]" captionTargetPageId="6" captionText="FIGURE 4. Habits (upper-case letters) and flower-heads on approximate same scale (lower-case letters) of Centaurea sect. Acrocentron representatives from Greece belonging to the C. achaia group; A,a: Centaurea corinthiaca (photos by Katerina Goula); B,b: C. achaia (photos by Katerina Goula); C,c: C. euboica; D,d, C. aetolica; e: C. dolopica." figureDoi="http://doi.org/10.5281/zenodo.7645661" httpUri="https://zenodo.org/record/7645661/files/figure.png" pageId="6" pageNumber="179">Fig. 4A,a</figureCitation>
|
||
) is a very distinct taxon with short stature and capitula with whitish corollas (see
|
||
<bibRefCitation id="EFC14B77FFFEFFE0D2C18A8BFAC6E4A8" author="Wagenitz, G. & Gamal-Eldin, E." box="[995,1366,1679,1706]" pageId="6" pageNumber="179" pagination="95 - 127" refId="ref9164" refString="Wagenitz, G. & Gamal-Eldin, E. (1985) Zur Kenntnis der griechischen Centaurea - Arten der sektion Acrokentron. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 107: 95 - 127." type="journal article" year="1985">Wagenitz & Gamal-Eldin (1985)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFFEFFE0D4408A8BFF4FE4CF" author="Routsi, E." pageId="6" pageNumber="179" refId="ref8730" refString="Routsi, E. (1993) Biosystematic study of the section Acrocentron (Cass.) DC. of the genus Centaurea L. in Greece. PhD Thesis, University of Patras, Greece." type="book" year="1993">Routsi (1993))</bibRefCitation>
|
||
. The involucres of this species are comparable in size to those of
|
||
<taxonomicName id="4C504D05FFFEFFE0D28F8AB0FBBFE4CF" box="[941,1071,1715,1742]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFEFFE0D28F8AB0FBBFE4CF" box="[941,1071,1715,1742]" italics="true" pageId="6" pageNumber="179">C. dolopica</emphasis>
|
||
</taxonomicName>
|
||
; however, the ratio of the width of bract appendages to the width of the bracts is clearly lower in
|
||
<taxonomicName id="4C504D05FFFEFFE0D21B8ADCFC4AE4F3" box="[825,986,1751,1778]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="corinthiaca">
|
||
<emphasis id="B924EA94FFFEFFE0D21B8ADCFC4AE4F3" box="[825,986,1751,1778]" italics="true" pageId="6" pageNumber="179">C. corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
—and the lowest observed in the group— because the bracts are wide with relatively narrow appendages bearing short cilia that cover less the neighboring bracts.
|
||
<taxonomicName id="4C504D05FFFEFFE0D1AA8B24FEF5E53B" box="[136,357,1823,1850]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFEFFE0D1AA8B24FEF5E53B" box="[136,357,1823,1850]" italics="true" pageId="6" pageNumber="179">Centaurea dolopica</emphasis>
|
||
</taxonomicName>
|
||
is additionally distinguished from
|
||
<taxonomicName id="4C504D05FFFEFFE0D3C78B24FC16E53B" box="[741,902,1823,1850]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="corinthiaca">
|
||
<emphasis id="B924EA94FFFEFFE0D3C78B24FC16E53B" box="[741,902,1823,1850]" italics="true" pageId="6" pageNumber="179">C. corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
by its considerably denser indumentum and more branched habit. We, additionally, note that some populations with intermediate characters between
|
||
<taxonomicName id="4C504D05FFFEFFE0D5F88B40FAECE55F" box="[1242,1404,1859,1886]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="corinthiaca">
|
||
<emphasis id="B924EA94FFFEFFE0D5F88B40FAECE55F" box="[1242,1404,1859,1886]" italics="true" pageId="6" pageNumber="179">C. corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C504D05FFFEFFE0D1AA8B6CFF65E583" box="[136,245,1895,1922]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFFEFFE0D1AA8B6CFF65E583" box="[136,245,1895,1922]" italics="true" pageId="6" pageNumber="179">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
occur near the
|
||
<emphasis id="B924EA94FFFEFFE0D0868B63FDDCE583" box="[420,588,1895,1921]" italics="true" pageId="6" pageNumber="179">locus classicus</emphasis>
|
||
of the former, and that probably led to the reduction of the Corinthian taxon to the rank of subspecies, viz.
|
||
<taxonomicName id="4C504D05FFFEFFE0D0F58B88FB1BE5A7" authority="Phitos & Georgiadis (1981: 102)" authorityName="Phitos & Georgiadis" authorityPageNumber="102" authorityYear="1981" box="[471,1163,1931,1958]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="subSpecies" species="achaia" subSpecies="corinthiaca">
|
||
<emphasis id="B924EA94FFFEFFE0D0F58B88FDD3E5A7" box="[471,579,1931,1958]" italics="true" pageId="6" pageNumber="179">C. achaia</emphasis>
|
||
subsp.
|
||
<emphasis id="B924EA94FFFEFFE0D3B48B8FFC87E5A7" box="[662,791,1931,1957]" italics="true" pageId="6" pageNumber="179">corinthiaca</emphasis>
|
||
<bibRefCitation id="EFC14B77FFFEFFE0D23F8B8FFB1BE5A7" author="Phitos, D. & Georgiadis, Th." box="[797,1163,1931,1957]" pageId="6" pageNumber="179" pagination="99 - 114" refId="ref8228" refString="Phitos, D. & Georgiadis, Th. (1981) A propos du grupe de Centaurea achaia Boiss. et Heldr. de la section Acrocentron (Cass.) DC. Botanika Chronika 1 (2): 99 - 114." type="journal article" year="1981">Phitos & Georgiadis (1981: 102)</bibRefCitation>
|
||
</taxonomicName>
|
||
, which has been followed to date. However, we suggest that the two species are to be retained separate as discussed below. Lastly,
|
||
<taxonomicName id="4C504D05FFFEFFE0D4138BB4FA3CE5CB" box="[1329,1452,1967,1994]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="euboica">
|
||
<emphasis id="B924EA94FFFEFFE0D4138BB4FA3CE5CB" box="[1329,1452,1967,1994]" italics="true" pageId="6" pageNumber="179">C. euboica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136B2A03FFFEFFE0D1B38BD7FF6CE5EF" box="[145,252,2003,2029]" captionStart="FIGURE 4" captionStartId="6.[136,229,1035,1057]" captionTargetBox="[155,1433,317,1010]" captionTargetId="figure-102@6.[155,1433,317,1010]" captionTargetPageId="6" captionText="FIGURE 4. Habits (upper-case letters) and flower-heads on approximate same scale (lower-case letters) of Centaurea sect. Acrocentron representatives from Greece belonging to the C. achaia group; A,a: Centaurea corinthiaca (photos by Katerina Goula); B,b: C. achaia (photos by Katerina Goula); C,c: C. euboica; D,d, C. aetolica; e: C. dolopica." figureDoi="http://doi.org/10.5281/zenodo.7645661" httpUri="https://zenodo.org/record/7645661/files/figure.png" pageId="6" pageNumber="179">Fig. 4C,c</figureCitation>
|
||
), an Euboean endemic species, differs from
|
||
<taxonomicName id="4C504D05FFFEFFE0D3D28BD0FCE3E5EF" box="[752,883,2003,2030]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="179" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFEFFE0D3D28BD0FCE3E5EF" box="[752,883,2003,2030]" italics="true" pageId="6" pageNumber="179">C. dolopica</emphasis>
|
||
</taxonomicName>
|
||
in being only sparsely hairy and in having ± more attenuate and narrower leaf segments, a less branched habit and larger capitula. The aforementioned morphological differences between the new species and the other members of the group are summarized in
|
||
<tableCitation id="C6D2033DFFFFFFE1D5B68C9AFB7AE2BA" box="[1172,1258,158,184]" captionStart="TABLE 2" captionStartId="7.[137,229,376,400]" captionTargetPageId="7" captionText="TABLE 2. Diagnostic morphological characters among the Centaurea achaia group. Measurements follow Routsi (1993)." httpUri="http://table.plazi.org/id/DF2F660EFFFFFFE1D1AB8D7CFA36E392" pageId="7" pageNumber="180" tableUuid="DF2F660EFFFFFFE1D1AB8D7CFA36E392">Table 2</tableCitation>
|
||
. We note that the depicted plants of
|
||
<taxonomicName id="4C504D05FFFFFFE1D0768CC7FE45E2DE" box="[340,469,194,221]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="180" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFFFFE1D0768CC7FE45E2DE" box="[340,469,194,221]" italics="true" pageId="7" pageNumber="180">C. dolopica</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<figureCitation id="136B2A03FFFFFFE1D0D58CC6FDD9E2DE" box="[503,585,194,220]" captionStart="FIGURE 1" captionStartId="3.[136,229,1600,1622]" captionTargetBox="[162,1428,517,1565]" captionTargetId="figure-145@3.[154,1433,508,1573]" captionTargetPageId="3" captionText="FIGURE 1. Centaurea dolopica. A, habit; B, capitulum; C, marginal (left) and central (right) floret, scale bar = 1 cm; D, two marginal (top left) and three central achenes, scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7645653" httpUri="https://zenodo.org/record/7645653/files/figure.png" pageId="7" pageNumber="180">Figs 1a</figureCitation>
|
||
and
|
||
<figureCitation id="136B2A03FFFFFFE1D35F8CC6FD1BE2DE" box="[637,651,194,220]" captionStart="FIGURE 2" captionStartId="4.[136,229,1483,1505]" captionTargetBox="[152,1432,543,1452]" captionTargetId="figure-200@4.[146,1441,533,1459]" captionTargetPageId="4" captionText="FIGURE 2. Holotype of Centaurea dolopica (UPA) mounted on two sheets." figureDoi="http://doi.org/10.5281/zenodo.7645655" httpUri="https://zenodo.org/record/7645655/files/figure.png" pageId="7" pageNumber="180">2</figureCitation>
|
||
represent medium-sized individuals in terms of plant height. The height of medium-sized
|
||
<taxonomicName id="4C504D05FFFFFFE1D06F8CE3FE41E302" box="[333,465,230,257]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="180" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFFFFE1D06F8CE3FE41E302" box="[333,465,230,257]" italics="true" pageId="7" pageNumber="180">C. dolopica</emphasis>
|
||
</taxonomicName>
|
||
plants is comparable to large- or very large-sized plants in the rest of the species of the
|
||
<taxonomicName id="4C504D05FFFFFFE1D1AA8D0FFF65E326" box="[136,245,266,293]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="180" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFFFFFE1D1AA8D0FFF65E326" box="[136,245,266,293]" italics="true" pageId="7" pageNumber="180">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
group. In the field, we observed some impressive individuals of
|
||
<taxonomicName id="4C504D05FFFFFFE1D2EE8D0FFBDEE326" box="[972,1102,266,293]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="180" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFFFFE1D2EE8D0FFBDEE326" box="[972,1102,266,293]" italics="true" pageId="7" pageNumber="180">C. dolopica</emphasis>
|
||
</taxonomicName>
|
||
with up to 40 flowering stems, the largest of those exceeding
|
||
<quantity id="4CA89B63FFFFFFE1D0F98D2AFD96E34B" box="[475,518,302,329]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="180" unit="m" value="1.0">1 m</quantity>
|
||
in length and each of the stems bearing up to ca. 30 capitula (
|
||
<figureCitation id="136B2A03FFFFFFE1D59F8D2AFA86E34A" box="[1213,1302,302,328]" captionStart="FIGURE 3" captionStartId="5.[136,229,1620,1642]" captionTargetBox="[152,1435,755,1586]" captionTargetId="figure-317@5.[143,1444,749,1595]" captionTargetPageId="5" captionText="FIGURE 3. Centaurea dolopica plants in their natural habitat: (A) several large plants along a roadside and (B) on a cut slope. (C) Eryngium serbicum in the same habitat." figureDoi="http://doi.org/10.5281/zenodo.7645657" httpUri="https://zenodo.org/record/7645657/files/figure.png" pageId="7" pageNumber="180">Fig. 3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF2F660EFFFFFFE1D1AB8D7CFA36E392" ID-Table-UUID="DF2F660EFFFFFFE1D1AB8D7CFA36E392" box="[137,1446,376,400]" httpUri="http://table.plazi.org/id/DF2F660EFFFFFFE1D1AB8D7CFA36E392" pageId="7" pageNumber="180" startId="7.[137,229,376,400]" targetBox="[144,1432,424,1910]" targetIsTable="true" targetPageId="7">
|
||
<paragraph id="8BEF3686FFFFFFE1D1AB8D7CFA36E392" blockId="7.[137,1446,376,400]" box="[137,1446,376,400]" pageId="7" pageNumber="180">
|
||
<emphasis id="B924EA94FFFFFFE1D1AB8D7CFF6FE392" bold="true" box="[137,255,376,400]" pageId="7" pageNumber="180">TABLE 2.</emphasis>
|
||
Diagnostic morphological characters among the
|
||
<taxonomicName id="4C504D05FFFFFFE1D22E8D7CFC5BE392" box="[780,971,376,400]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="180" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFFFFFE1D22E8D7CFC5BE392" box="[780,971,376,400]" italics="true" pageId="7" pageNumber="180">Centaurea achaia</emphasis>
|
||
</taxonomicName>
|
||
group. Measurements follow
|
||
<bibRefCitation id="EFC14B77FFFFFFE1D42E8D7CFA32E392" author="Routsi, E." box="[1292,1442,376,400]" pageId="7" pageNumber="180" refId="ref8730" refString="Routsi, E. (1993) Biosystematic study of the section Acrocentron (Cass.) DC. of the genus Centaurea L. in Greece. PhD Thesis, University of Patras, Greece." type="book" year="1993">Routsi (1993)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEF3686FFFFFFE1D0F88DACFAE5E56F" pageId="7" pageNumber="180">
|
||
<table id="F950C426FFFF0019D1B28DACFA08E574" box="[144,1432,424,1910]" gridcols="6" gridrows="16" pageId="7" pageNumber="180">
|
||
<tr id="356034C4FFFF0019D1B28DACFA08E3BC" box="[144,1432,424,446]" gridrow="0" pageId="7" pageNumber="180" rowspan-0="1">
|
||
<th id="76B15DB8FFFF0019D0F88DACFD0CE3BC" box="[474,668,424,446]" gridcol="1" gridrow="0" pageId="7" pageNumber="180">
|
||
<taxonomicName id="4C504D05FFFFFFE1D0F88DACFD0CE3BC" box="[474,668,424,446]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="180" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFFFFFE1D0F88DACFD0CE3BC" bold="true" box="[474,668,424,446]" italics="true" pageId="7" pageNumber="180">Centaurea dolopica</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76B15DB8FFFF0019D3918DACFCFCE3BC" box="[691,876,424,446]" gridcol="2" gridrow="0" pageId="7" pageNumber="180">
|
||
<taxonomicName id="4C504D05FFFFFFE1D3918DACFC82E3BC" box="[691,786,424,446]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="180" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFFFFFE1D3918DACFC82E3BC" bold="true" box="[691,786,424,446]" italics="true" pageId="7" pageNumber="180">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76B15DB8FFFF0019D25E8DACFBB6E3BC" box="[892,1062,424,446]" gridcol="3" gridrow="0" pageId="7" pageNumber="180">
|
||
<taxonomicName id="4C504D05FFFFFFE1D25E8DACFC75E3BC" box="[892,997,424,446]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="180" phylum="Tracheophyta" rank="species" species="aetolica">
|
||
<emphasis id="B924EA94FFFFFFE1D25E8DACFC75E3BC" bold="true" box="[892,997,424,446]" italics="true" pageId="7" pageNumber="180">C. aetolica</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76B15DB8FFFF0019D51A8DACFB4EE3BC" box="[1080,1246,424,446]" gridcol="4" gridrow="0" pageId="7" pageNumber="180">
|
||
<taxonomicName id="4C504D05FFFFFFE1D51A8DACFB55E3BC" box="[1080,1221,424,446]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="180" phylum="Tracheophyta" rank="species" species="corinthiaca">
|
||
<emphasis id="B924EA94FFFFFFE1D51A8DACFB55E3BC" bold="true" box="[1080,1221,424,446]" italics="true" pageId="7" pageNumber="180">C. corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76B15DB8FFFF0019D5CC8DACFA08E3BC" box="[1262,1432,424,446]" gridcol="5" gridrow="0" pageId="7" pageNumber="180">
|
||
<emphasis id="B924EA94FFFFFFE1D5CC8DACFAC7E3BC" bold="true" box="[1262,1367,424,446]" italics="true" pageId="7" pageNumber="180">C. euboica</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B28DDBFA08E03F" box="[144,1432,479,573]" gridrow="1" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B28DDBFE58E03F" box="[144,456,479,573]" gridcol="0" gridrow="1" pageId="7" pageNumber="180">Indumentum</th>
|
||
<td id="76B15DB8FFFF0019D0F88DDBFD0CE03F" box="[474,668,479,573]" gridcol="1" gridrow="1" pageId="7" pageNumber="180">Densely arachnoid (plants ± greyish)</td>
|
||
<td id="76B15DB8FFFF0019D3918DDBFCFCE03F" box="[691,876,479,573]" gridcol="2" gridrow="1" pageId="7" pageNumber="180">Moderately arachnoid (plants ± green)</td>
|
||
<td id="76B15DB8FFFF0019D25E8DDBFBB6E03F" box="[892,1062,479,573]" gridcol="3" gridrow="1" pageId="7" pageNumber="180">Densely arachnoid (plants ± greyish)</td>
|
||
<td id="76B15DB8FFFF0019D51A8DDBFB4EE03F" box="[1080,1246,479,573]" gridcol="4" gridrow="1" pageId="7" pageNumber="180">Moderately arachnoid (plants ± green)</td>
|
||
<td id="76B15DB8FFFF0019D5CC8DDBFA08E03F" box="[1262,1432,479,573]" gridcol="5" gridrow="1" pageId="7" pageNumber="180">Sparsely arachnoid (plants green)</td>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B28E59FA08E076" box="[144,1432,605,628]" gridrow="2" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B28E59FE58E076" box="[144,456,605,628]" gridcol="0" gridrow="2" pageId="7" pageNumber="180">Length of stems (cm)</th>
|
||
<td id="76B15DB8FFFF0019D0F88E59FD0CE076" box="[474,668,605,628]" gridcol="1" gridrow="2" pageId="7" pageNumber="180">40–100(–120)</td>
|
||
<td id="76B15DB8FFFF0019D3918E59FCFCE076" box="[691,876,605,628]" gridcol="2" gridrow="2" pageId="7" pageNumber="180">30–60(–80)</td>
|
||
<td id="76B15DB8FFFF0019D25E8E59FBB6E076" box="[892,1062,605,628]" gridcol="3" gridrow="2" pageId="7" pageNumber="180">30–70</td>
|
||
<td id="76B15DB8FFFF0019D51A8E59FB4EE076" box="[1080,1246,605,628]" gridcol="4" gridrow="2" pageId="7" pageNumber="180">20–50</td>
|
||
<td id="76B15DB8FFFF0019D5CC8E59FA08E076" box="[1262,1432,605,628]" gridcol="5" gridrow="2" pageId="7" pageNumber="180">30–70</td>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B28E97FA08E0A8" box="[144,1432,659,682]" gridrow="3" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B28E97FE58E0A8" box="[144,456,659,682]" gridcol="0" gridrow="3" pageId="7" pageNumber="180">Branches per stem1</th>
|
||
<td id="76B15DB8FFFF0019D0F88E97FD0CE0A8" box="[474,668,659,682]" gridcol="1" gridrow="3" pageId="7" pageNumber="180">1–11</td>
|
||
<td id="76B15DB8FFFF0019D3918E97FCFCE0A8" box="[691,876,659,682]" gridcol="2" gridrow="3" pageId="7" pageNumber="180">(0–)1–6(–8)</td>
|
||
<td id="76B15DB8FFFF0019D25E8E97FBB6E0A8" box="[892,1062,659,682]" gridcol="3" gridrow="3" pageId="7" pageNumber="180">1–9</td>
|
||
<td id="76B15DB8FFFF0019D51A8E97FB4EE0A8" box="[1080,1246,659,682]" gridcol="4" gridrow="3" pageId="7" pageNumber="180">0–6</td>
|
||
<td id="76B15DB8FFFF0019D5CC8E97FA08E0A8" box="[1262,1432,659,682]" gridcol="5" gridrow="3" pageId="7" pageNumber="180">(0–)1–5</td>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B28ECEFA08E106" box="[144,1432,714,772]" gridrow="4" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B28ECEFE58E106" box="[144,456,714,772]" gridcol="0" gridrow="4" pageId="7" pageNumber="180">Terminal leaflet</th>
|
||
<td id="76B15DB8FFFF0019D0F88ECEFD0CE106" box="[474,668,714,772]" gridcol="1" gridrow="4" pageId="7" pageNumber="180">± Lanceolate</td>
|
||
<td id="76B15DB8FFFF0019D3918ECEFCFCE106" box="[691,876,714,772]" gridcol="2" gridrow="4" pageId="7" pageNumber="180">± Lanceolate</td>
|
||
<td id="76B15DB8FFFF0019D25E8ECEFBB6E106" box="[892,1062,714,772]" gridcol="3" gridrow="4" pageId="7" pageNumber="180">± Linear</td>
|
||
<td id="76B15DB8FFFF0019D51A8ECEFB4EE106" box="[1080,1246,714,772]" gridcol="4" gridrow="4" pageId="7" pageNumber="180">± Lanceolate</td>
|
||
<td id="76B15DB8FFFF0019D5CC8ECEFA08E106" box="[1262,1432,714,772]" gridcol="5" gridrow="4" pageId="7" pageNumber="180">± Narrowly oblanceolate</td>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B28F20FA08E139" box="[144,1432,804,827]" gridrow="5" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B28F20FE58E139" box="[144,456,804,827]" gridcol="0" gridrow="5" pageId="7" pageNumber="180">Capitula per stem2</th>
|
||
<td id="76B15DB8FFFF0019D0F88F20FD0CE139" box="[474,668,804,827]" gridcol="1" gridrow="5" pageId="7" pageNumber="180">3–29</td>
|
||
<td id="76B15DB8FFFF0019D3918F20FCFCE139" box="[691,876,804,827]" gridcol="2" gridrow="5" pageId="7" pageNumber="180">1–12</td>
|
||
<td id="76B15DB8FFFF0019D25E8F20FBB6E139" box="[892,1062,804,827]" gridcol="3" gridrow="5" pageId="7" pageNumber="180">2–20</td>
|
||
<td id="76B15DB8FFFF0019D51A8F20FB4EE139" box="[1080,1246,804,827]" gridcol="4" gridrow="5" pageId="7" pageNumber="180">1–10</td>
|
||
<td id="76B15DB8FFFF0019D5CC8F20FA08E139" box="[1262,1432,804,827]" gridcol="5" gridrow="5" pageId="7" pageNumber="180">1–10</td>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B28F5FFA08E197" box="[144,1432,859,917]" gridrow="6" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B28F5FFE58E197" box="[144,456,859,917]" gridcol="0" gridrow="6" pageId="7" pageNumber="180">Involucre (cm × cm)</th>
|
||
<td id="76B15DB8FFFF0019D0F88F5FFD0CE197" box="[474,668,859,917]" gridcol="1" gridrow="6" pageId="7" pageNumber="180">1.8–2.6 × 0.9–1.5</td>
|
||
<td id="76B15DB8FFFF0019D3918F5FFCFCE197" box="[691,876,859,917]" gridcol="2" gridrow="6" pageId="7" pageNumber="180">2–3 × 1.6–2.8</td>
|
||
<td id="76B15DB8FFFF0019D25E8F5FFBB6E197" box="[892,1062,859,917]" gridcol="3" gridrow="6" pageId="7" pageNumber="180">2.3–2.7 × 1.6–2.2</td>
|
||
<td id="76B15DB8FFFF0019D51A8F5FFB4EE197" box="[1080,1246,859,917]" gridcol="4" gridrow="6" pageId="7" pageNumber="180">1.9–2.4 × 1.2–1.5</td>
|
||
<td id="76B15DB8FFFF0019D5CC8F5FFA08E197" box="[1262,1432,859,917]" gridcol="5" gridrow="6" pageId="7" pageNumber="180">2.1–2.9 × 1.9–2.9</td>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B28FDEFA08E616" box="[144,1432,986,1044]" gridrow="7" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B28FDEFE58E616" box="[144,456,986,1044]" gridcol="0" gridrow="7" pageId="7" pageNumber="180">Length of middle involucral bracts (cm)</th>
|
||
<td id="76B15DB8FFFF0019D0F88FDEFD0CE616" box="[474,668,986,1044]" gridcol="1" gridrow="7" pageId="7" pageNumber="180">0.8–1.8</td>
|
||
<td id="76B15DB8FFFF0019D3918FDEFCFCE616" box="[691,876,986,1044]" gridcol="2" gridrow="7" pageId="7" pageNumber="180">1.8–4.5(–5.8)</td>
|
||
<td id="76B15DB8FFFF0019D25E8FDEFBB6E616" box="[892,1062,986,1044]" gridcol="3" gridrow="7" pageId="7" pageNumber="180">1.4–2.5</td>
|
||
<td id="76B15DB8FFFF0019D51A8FDEFB4EE616" box="[1080,1246,986,1044]" gridcol="4" gridrow="7" pageId="7" pageNumber="180">1.7–3.4</td>
|
||
<td id="76B15DB8FFFF0019D5CC8FDEFA08E616" box="[1262,1432,986,1044]" gridcol="5" gridrow="7" pageId="7" pageNumber="180">1.3–3.2</td>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B28830FA08E66C" box="[144,1432,1076,1134]" gridrow="8" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B28830FE58E66C" box="[144,456,1076,1134]" gridcol="0" gridrow="8" pageId="7" pageNumber="180">Width of middle involucral bracts (cm)</th>
|
||
<td id="76B15DB8FFFF0019D0F88830FD0CE66C" box="[474,668,1076,1134]" gridcol="1" gridrow="8" pageId="7" pageNumber="180">0.3–0.5</td>
|
||
<td id="76B15DB8FFFF0019D3918830FCFCE66C" box="[691,876,1076,1134]" gridcol="2" gridrow="8" pageId="7" pageNumber="180">(0.5–)0.6–0.8</td>
|
||
<td id="76B15DB8FFFF0019D25E8830FBB6E66C" box="[892,1062,1076,1134]" gridcol="3" gridrow="8" pageId="7" pageNumber="180">(0.55–)0.65–0.7</td>
|
||
<td id="76B15DB8FFFF0019D51A8830FB4EE66C" box="[1080,1246,1076,1134]" gridcol="4" gridrow="8" pageId="7" pageNumber="180">(0.5–)0.55–0.65</td>
|
||
<td id="76B15DB8FFFF0019D5CC8830FA08E66C" box="[1262,1432,1076,1134]" gridcol="5" gridrow="8" pageId="7" pageNumber="180">(0.5–)0.55–0.75</td>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B2888BFA08E6EE" box="[144,1432,1167,1260]" gridrow="9" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B2888BFE58E6EE" box="[144,456,1167,1260]" gridcol="0" gridrow="9" pageId="7" pageNumber="180">
|
||
Width of appendage of middle involucral bracts /Width of bract (mean ±
|
||
<taxonomicName id="4C504D05FFFFFFE1D1C788D3FEA1E6EE" box="[229,305,1239,1260]" pageId="7" pageNumber="180" rank="subSpecies" subSpecies="error">st. error</taxonomicName>
|
||
)
|
||
</th>
|
||
<td id="76B15DB8FFFF0019D0F8888BFD0CE6EE" box="[474,668,1167,1260]" gridcol="1" gridrow="9" pageId="7" pageNumber="180">2.18 ± 0.07</td>
|
||
<td id="76B15DB8FFFF0019D391888BFCFCE6EE" box="[691,876,1167,1260]" gridcol="2" gridrow="9" pageId="7" pageNumber="180">1.93–2.28 ± 0.04–0.1</td>
|
||
<td id="76B15DB8FFFF0019D25E888BFBB6E6EE" box="[892,1062,1167,1260]" gridcol="3" gridrow="9" pageId="7" pageNumber="180">1.84 ± 0.09</td>
|
||
<td id="76B15DB8FFFF0019D51A888BFB4EE6EE" box="[1080,1246,1167,1260]" gridcol="4" gridrow="9" pageId="7" pageNumber="180">1.41 ± 0.04</td>
|
||
<td id="76B15DB8FFFF0019D5CC888BFA08E6EE" box="[1262,1432,1167,1260]" gridcol="5" gridrow="9" pageId="7" pageNumber="180">1.98–2 ± 0.04– 0.07</td>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B28909FA08E769" box="[144,1432,1293,1387]" gridrow="10" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B28909FE58E769" box="[144,456,1293,1387]" gridcol="0" gridrow="10" pageId="7" pageNumber="180">
|
||
Length of middle involucral bracts/Length of appendage (mean ±
|
||
<taxonomicName id="4C504D05FFFFFFE1D1C78952FEA1E768" box="[229,305,1365,1387]" pageId="7" pageNumber="180" rank="subSpecies" subSpecies="error">st. error</taxonomicName>
|
||
)
|
||
</th>
|
||
<td id="76B15DB8FFFF0019D0F88909FD0CE769" box="[474,668,1293,1387]" gridcol="1" gridrow="10" pageId="7" pageNumber="180">1.49 ± 0.03</td>
|
||
<td id="76B15DB8FFFF0019D3918909FCFCE769" box="[691,876,1293,1387]" gridcol="2" gridrow="10" pageId="7" pageNumber="180">1.11–1.19 ± 0.01–0.04</td>
|
||
<td id="76B15DB8FFFF0019D25E8909FBB6E769" box="[892,1062,1293,1387]" gridcol="3" gridrow="10" pageId="7" pageNumber="180">1.24 ± 0.02</td>
|
||
<td id="76B15DB8FFFF0019D51A8909FB4EE769" box="[1080,1246,1293,1387]" gridcol="4" gridrow="10" pageId="7" pageNumber="180">1.22 ± 0.03</td>
|
||
<td id="76B15DB8FFFF0019D5CC8909FA08E769" box="[1262,1432,1293,1387]" gridcol="5" gridrow="10" pageId="7" pageNumber="180">1.18–1.28 ± 0.04–0.07</td>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B2898FFA08E40F" box="[144,1432,1419,1549]" gridrow="11" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B2898FFE58E40F" box="[144,456,1419,1549]" gridcol="0" gridrow="11" pageId="7" pageNumber="180">Stripe of colored part of the appendage (incl. spine)</th>
|
||
<td id="76B15DB8FFFF0019D0F8898FFD0CE40F" box="[474,668,1419,1549]" gridcol="1" gridrow="11" pageId="7" pageNumber="180">Brownish with darker veins, contrasting the cilia</td>
|
||
<td id="76B15DB8FFFF0019D391898FFCFCE40F" box="[691,876,1419,1549]" gridcol="2" gridrow="11" pageId="7" pageNumber="180">Brownish with darker veins, contrasting the cilia</td>
|
||
<td id="76B15DB8FFFF0019D25E898FFBB6E40F" box="[892,1062,1419,1549]" gridcol="3" gridrow="11" pageId="7" pageNumber="180">Whitish with indistinct veins, not contrasting the cilia</td>
|
||
<td id="76B15DB8FFFF0019D51A898FFB4EE40F" box="[1080,1246,1419,1549]" gridcol="4" gridrow="11" pageId="7" pageNumber="180">Brownish with darker veins, contrasting the cilia</td>
|
||
<td id="76B15DB8FFFF0019D5CC898FFA08E40F" box="[1262,1432,1419,1549]" gridcol="5" gridrow="11" pageId="7" pageNumber="180">Brownish with darker veins, contrasting the cilia</td>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B28A2AFA08E46A" box="[144,1432,1582,1640]" gridrow="12" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B28A2AFE58E46A" box="[144,456,1582,1640]" gridcol="0" gridrow="12" pageId="7" pageNumber="180">Spine of middle involucral bracts (cm)</th>
|
||
<td id="76B15DB8FFFF0019D0F88A2AFD0CE46A" box="[474,668,1582,1640]" gridcol="1" gridrow="12" pageId="7" pageNumber="180">0.1–0.8</td>
|
||
<td id="76B15DB8FFFF0019D3918A2AFCFCE46A" box="[691,876,1582,1640]" gridcol="2" gridrow="12" pageId="7" pageNumber="180">0.9–3.2(–4.4)</td>
|
||
<td id="76B15DB8FFFF0019D25E8A2AFBB6E46A" box="[892,1062,1582,1640]" gridcol="3" gridrow="12" pageId="7" pageNumber="180">0.3–1.2</td>
|
||
<td id="76B15DB8FFFF0019D51A8A2AFB4EE46A" box="[1080,1246,1582,1640]" gridcol="4" gridrow="12" pageId="7" pageNumber="180">0.9–2.2</td>
|
||
<td id="76B15DB8FFFF0019D5CC8A2AFA08E46A" box="[1262,1432,1582,1640]" gridcol="5" gridrow="12" pageId="7" pageNumber="180">0.4–2</td>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B28A8CFA08E49C" box="[144,1432,1672,1694]" gridrow="13" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B28A8CFE58E49C" box="[144,456,1672,1694]" gridcol="0" gridrow="13" pageId="7" pageNumber="180">Number of central florets</th>
|
||
<td id="76B15DB8FFFF0019D0F88A8CFD0CE49C" box="[474,668,1672,1694]" gridcol="1" gridrow="13" pageId="7" pageNumber="180">23–39</td>
|
||
<td id="76B15DB8FFFF0019D3918A8CFCFCE49C" box="[691,876,1672,1694]" gridcol="2" gridrow="13" pageId="7" pageNumber="180">(43–)52–71(–88)</td>
|
||
<td id="76B15DB8FFFF0019D25E8A8CFBB6E49C" box="[892,1062,1672,1694]" gridcol="3" gridrow="13" pageId="7" pageNumber="180">(54–)62–96</td>
|
||
<td id="76B15DB8FFFF0019D51A8A8CFB4EE49C" box="[1080,1246,1672,1694]" gridcol="4" gridrow="13" pageId="7" pageNumber="180">36–49(–52)</td>
|
||
<td id="76B15DB8FFFF0019D5CC8A8CFA08E49C" box="[1262,1432,1672,1694]" gridcol="5" gridrow="13" pageId="7" pageNumber="180">28–56</td>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B28ABBFA08E4FB" box="[144,1432,1727,1785]" gridrow="14" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B28ABBFE58E4FB" box="[144,456,1727,1785]" gridcol="0" gridrow="14" pageId="7" pageNumber="180">Corolla</th>
|
||
<td id="76B15DB8FFFF0019D0F88ABBFD0CE4FB" box="[474,668,1727,1785]" gridcol="1" gridrow="14" pageId="7" pageNumber="180">Purplish</td>
|
||
<td id="76B15DB8FFFF0019D3918ABBFCFCE4FB" box="[691,876,1727,1785]" gridcol="2" gridrow="14" pageId="7" pageNumber="180">Purplish</td>
|
||
<td id="76B15DB8FFFF0019D25E8ABBFBB6E4FB" box="[892,1062,1727,1785]" gridcol="3" gridrow="14" pageId="7" pageNumber="180">Purplish</td>
|
||
<td id="76B15DB8FFFF0019D51A8ABBFB4EE4FB" box="[1080,1246,1727,1785]" gridcol="4" gridrow="14" pageId="7" pageNumber="180">Whitish with orange veins</td>
|
||
<td id="76B15DB8FFFF0019D5CC8ABBFA08E4FB" box="[1262,1432,1727,1785]" gridcol="5" gridrow="14" pageId="7" pageNumber="180">Purplish</td>
|
||
</tr>
|
||
<tr id="356034C4FFFF0019D1B28B1DFA08E574" box="[144,1432,1817,1910]" gridrow="15" pageId="7" pageNumber="180">
|
||
<th id="76B15DB8FFFF0019D1B28B1DFE58E574" box="[144,456,1817,1910]" gridcol="0" gridrow="15" pageId="7" pageNumber="180">
|
||
Length of achenes of central florets in cm (mean ±
|
||
<taxonomicName id="4C504D05FFFFFFE1D0468B3AFE20E550" box="[356,432,1853,1875]" pageId="7" pageNumber="180" rank="subSpecies" subSpecies="error">st. error</taxonomicName>
|
||
)
|
||
</th>
|
||
<td id="76B15DB8FFFF0019D0F88B1DFD0CE574" box="[474,668,1817,1910]" gridcol="1" gridrow="15" pageId="7" pageNumber="180">0.41–0.53 (0.47 ± 0.008)</td>
|
||
<td id="76B15DB8FFFF0019D3918B1DFCFCE574" box="[691,876,1817,1910]" gridcol="2" gridrow="15" pageId="7" pageNumber="180">0.5–0.6 (0.5–0.551 ± 0.001–0.014)3</td>
|
||
<td id="76B15DB8FFFF0019D25E8B1DFBB6E574" box="[892,1062,1817,1910]" gridcol="3" gridrow="15" pageId="7" pageNumber="180">0.42–0.46 (0.439 ± 0.006)</td>
|
||
<td id="76B15DB8FFFF0019D51A8B1DFB4EE574" box="[1080,1246,1817,1910]" gridcol="4" gridrow="15" pageId="7" pageNumber="180">0.52–0.56 (0.529 ± 0.005)</td>
|
||
<td id="76B15DB8FFFF0019D5CC8B1DFA08E574" box="[1262,1432,1817,1910]" gridcol="5" gridrow="15" pageId="7" pageNumber="180">0.5–0.55 (0.512–0.522 ± 0.008–0.009)4</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<tableNote id="76B63708FFFFFFE1D1AA8B8FFC3BEA12" pageId="7" pageNumber="180" targetBox="[144,1432,424,1910]" targetPageId="7">
|
||
<paragraph id="8BEF3686FFFFFFE1D1AA8B8FFC2AE5A6" blockId="7.[136,954,1931,2065]" box="[136,954,1931,1956]" pageId="7" pageNumber="180">
|
||
<superScript id="7C259BCEFFFFFFE1D1AA8B8FFF00E59B" attach="left" box="[136,144,1931,1945]" fontSize="6" pageId="7" pageNumber="180">1</superScript>
|
||
Only the branches that produce non-abortive, fertile capitula are accounted.
|
||
</paragraph>
|
||
<paragraph id="8BEF3686FFFFFFE1D1AA8BABFD5CE5CA" blockId="7.[136,954,1931,2065]" box="[136,716,1967,1992]" pageId="7" pageNumber="180">
|
||
<superScript id="7C259BCEFFFFFFE1D1AA8BABFF00E5BF" attach="left" box="[136,144,1967,1981]" fontSize="6" pageId="7" pageNumber="180">2</superScript>
|
||
Only the non-abortive, fertile capitula are accounted.
|
||
</paragraph>
|
||
<paragraph id="8BEF3686FFFFFFE1D1AA8BD7FC3BE5EE" blockId="7.[136,954,1931,2065]" box="[136,939,2003,2029]" pageId="7" pageNumber="180">
|
||
<superScript id="7C259BCEFFFFFFE1D1AA8BD7FF00E5E3" attach="left" box="[136,144,2003,2017]" fontSize="6" pageId="7" pageNumber="180">3</superScript>
|
||
Range of mean values and standard errors of 4 populations (
|
||
<bibRefCitation id="EFC14B77FFFFFFE1D23B8BD0FC0FE5EE" author="Routsi, E." box="[793,927,2004,2028]" pageId="7" pageNumber="180" refId="ref8730" refString="Routsi, E. (1993) Biosystematic study of the section Acrocentron (Cass.) DC. of the genus Centaurea L. in Greece. PhD Thesis, University of Patras, Greece." type="book" year="1993">Routsi 1993</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEF3686FFFFFFE1D1AA8BF3FC3BEA12" blockId="7.[136,954,1931,2065]" box="[136,939,2039,2065]" pageId="7" pageNumber="180">
|
||
<superScript id="7C259BCEFFFFFFE1D1AA8BF3FF00EA07" attach="left" box="[136,144,2039,2053]" fontSize="6" pageId="7" pageNumber="180">4</superScript>
|
||
Range of mean values and standard errors of 2 populations (
|
||
<bibRefCitation id="EFC14B77FFFFFFE1D23B8BFCFC0FEA12" author="Routsi, E." box="[793,927,2040,2064]" pageId="7" pageNumber="180" refId="ref8730" refString="Routsi, E. (1993) Biosystematic study of the section Acrocentron (Cass.) DC. of the genus Centaurea L. in Greece. PhD Thesis, University of Patras, Greece." type="book" year="1993">Routsi 1993</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</tableNote>
|
||
<caption id="DF2F660EFFF0FFEED1AA8E4EFB9EE062" ID-DOI="http://doi.org/10.5281/zenodo.7645665" ID-Zenodo-Dep="7645665" box="[136,1038,585,608]" httpUri="https://zenodo.org/record/7645665/files/figure.png" pageId="8" pageNumber="181" startId="8.[136,229,586,608]" targetBox="[276,1312,190,552]" targetPageId="8">
|
||
<paragraph id="8BEF3686FFF0FFEED1AA8E4EFB9EE062" blockId="8.[136,1038,585,608]" box="[136,1038,585,608]" pageId="8" pageNumber="181">
|
||
<emphasis id="B924EA94FFF0FFEED1AA8E4EFE8BE05D" bold="true" box="[136,283,585,608]" pageId="8" pageNumber="181">FIGURE 5. A)</emphasis>
|
||
Karyotype and
|
||
<emphasis id="B924EA94FFF0FFEED0978E4EFE5DE05D" bold="true" box="[437,461,585,607]" pageId="8" pageNumber="181">B)</emphasis>
|
||
idiogram of
|
||
<taxonomicName id="4C504D05FFF0FFEED3688E4EFC98E05D" box="[586,776,586,607]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="181" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFF0FFEED3688E4EFC98E05D" box="[586,776,586,607]" italics="true" pageId="8" pageNumber="181">Centaurea dolopica</emphasis>
|
||
</taxonomicName>
|
||
. Arrows indicate satellites.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2F660EFFF0FFEED1AA8A9EFE5AE542" ID-DOI="http://doi.org/10.5281/zenodo.7645667" ID-Zenodo-Dep="7645667" httpUri="https://zenodo.org/record/7645667/files/figure.png" pageId="8" pageNumber="181" startId="8.[136,229,1690,1712]" targetBox="[153,1436,722,1660]" targetPageId="8">
|
||
<paragraph id="8BEF3686FFF0FFEED1AA8A9EFE5AE542" blockId="8.[136,1452,1690,1856]" pageId="8" pageNumber="181">
|
||
<emphasis id="B924EA94FFF0FFEED1AA8A9EFF6FE4AD" bold="true" box="[136,255,1690,1712]" pageId="8" pageNumber="181">FIGURE 6.</emphasis>
|
||
Phylogeny of
|
||
<taxonomicName id="4C504D05FFF0FFEED0AD8A9FFCC2E4B2" box="[399,850,1690,1712]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="181" phylum="Tracheophyta" rank="section" section="Acrocentron" subGenus="Lopholoma">
|
||
<emphasis id="B924EA94FFF0FFEED0AD8A9FFE63E4B2" box="[399,499,1691,1712]" italics="true" pageId="8" pageNumber="181">Centaurea</emphasis>
|
||
subg.
|
||
<emphasis id="B924EA94FFF0FFEED3178A9EFD32E4AD" box="[565,674,1690,1711]" italics="true" pageId="8" pageNumber="181">Lopholoma</emphasis>
|
||
sect.
|
||
<emphasis id="B924EA94FFF0FFEED3FE8A9FFCC2E4B2" box="[732,850,1691,1712]" italics="true" pageId="8" pageNumber="181">Acrocentron</emphasis>
|
||
</taxonomicName>
|
||
, including the newly described
|
||
<taxonomicName id="4C504D05FFF0FFEED5A88A9FFADAE4AD" box="[1162,1354,1690,1712]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="181" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFF0FFEED5A88A9FFADAE4AD" box="[1162,1354,1690,1712]" italics="true" pageId="8" pageNumber="181">Centaurea dolopica</emphasis>
|
||
</taxonomicName>
|
||
(in bold), with
|
||
<taxonomicName id="4C504D05FFF0FFEED19F8ABBFE21E4D6" box="[189,433,1726,1748]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="181" phylum="Tracheophyta" rank="subGenus" subGenus="Cyanus">
|
||
<emphasis id="B924EA94FFF0FFEED19F8ABBFEB1E4D6" box="[189,289,1727,1748]" italics="true" pageId="8" pageNumber="181">Centaurea</emphasis>
|
||
subg.
|
||
<emphasis id="B924EA94FFF0FFEED04B8ABBFE21E4D6" box="[361,433,1727,1748]" italics="true" pageId="8" pageNumber="181">Cyanus</emphasis>
|
||
</taxonomicName>
|
||
as outgroup, inferred from Bayesian analysis of the concatenated ITS and ETS sequences. Posterior probabilities (pp) higher than 0.7 and bootstrap values (bs) higher than 50% estimated from MrBayes and RAxML analyses are reported above and below the branches, respectively.At the right, the names of the major clades in
|
||
<emphasis id="B924EA94FFF0FFEED2FD8B03FC66E51E" box="[991,1014,1799,1820]" italics="true" pageId="8" pageNumber="181">C.</emphasis>
|
||
subg.
|
||
<taxonomicName id="4C504D05FFF0FFEED5118B02FB30E519" box="[1075,1184,1798,1819]" class="Magnoliopsida" family="Asteraceae" genus="Lopholoma" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="181" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B924EA94FFF0FFEED5118B02FB30E519" box="[1075,1184,1798,1819]" italics="true" pageId="8" pageNumber="181">Lopholoma</emphasis>
|
||
</taxonomicName>
|
||
correspond to those of Font
|
||
<emphasis id="B924EA94FFF0FFEED1AA8B2FFF22E53D" box="[136,178,1834,1856]" italics="true" pageId="8" pageNumber="181">et al</emphasis>
|
||
.’s (2009) phylogenetic trees.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEF3686FFF0FFEFD19F8B6BFC70E0B2" blockId="8.[136,1452,1902,2073]" lastBlockId="9.[136,1452,158,689]" lastPageId="9" lastPageNumber="182" pageId="8" pageNumber="181">
|
||
The karyological studies for the genus
|
||
<taxonomicName id="4C504D05FFF0FFEED3588B6BFD7FE58B" authorityName="Zografidis & Koutroumpa & Liveri & Dimopoulos" authorityYear="2023" box="[634,751,1903,1929]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="181" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B924EA94FFF0FFEED3588B6BFD7FE58B" box="[634,751,1903,1929]" italics="true" pageId="8" pageNumber="181">Centaurea</emphasis>
|
||
</taxonomicName>
|
||
are numerous (see
|
||
<bibRefCitation id="EFC14B77FFF0FFEED2ED8B6BFB17E58B" author="Rice, A. & Glick, L. & Abadi, S. & Einhorn, M. & Kopelman, N. M. & Salman-Minkov, A. & Mayzel, J. & Chay, O. & Mayrose, I." box="[975,1159,1902,1929]" pageId="8" pageNumber="181" pagination="19 - 26" refId="ref8548" refString="Rice, A., Glick, L., Abadi, S., Einhorn, M., Kopelman, N. M., Salman-Minkov, A., Mayzel, J., Chay, O. & Mayrose, I. (2015) The Chromosome Counts Database (CCDB) - a community resource of plant chromosome numbers. New Phytologist 206: 19 - 26. https: // doi. org / 10.1111 / nph. 13191" type="journal article" year="2015">
|
||
Rice
|
||
<emphasis id="B924EA94FFF0FFEED5298B6BFBD5E58B" box="[1035,1093,1903,1929]" italics="true" pageId="8" pageNumber="181">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
) and have revealed many basic chromosome numbers (x = 8, 9, 10, 11, 12, 13, 14, 15, 16) and several ploidy levels (2x, 3x, 4x, up to 11x) (
|
||
<bibRefCitation id="EFC14B77FFF0FFEED4408B97FF50E5D3" author="Routsi, E." pageId="8" pageNumber="181" refId="ref8730" refString="Routsi, E. (1993) Biosystematic study of the section Acrocentron (Cass.) DC. of the genus Centaurea L. in Greece. PhD Thesis, University of Patras, Greece." type="book" year="1993">Routsi 1993</bibRefCitation>
|
||
and references therein). The taxa of
|
||
<emphasis id="B924EA94FFF0FFEED34B8BB3FD13E5D3" box="[617,643,1975,2001]" italics="true" pageId="8" pageNumber="181">C.</emphasis>
|
||
sect.
|
||
<taxonomicName id="4C504D05FFF0FFEED3E58BB3FCC1E5D3" box="[711,849,1975,2001]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="185" phylum="Tracheophyta" rank="section" section="Acrocentron">
|
||
<emphasis id="B924EA94FFF0FFEED3E58BB3FCC1E5D3" box="[711,849,1975,2001]" italics="true" pageId="8" pageNumber="181">Acrocentron</emphasis>
|
||
</taxonomicName>
|
||
have also been extensively examined karyologically (e.g.,
|
||
<bibRefCitation id="EFC14B77FFF0FFEED1EA8BDFFEEEE5F6" author="Runemark, H." box="[200,382,2010,2037]" pageId="8" pageNumber="181" pagination="161 - 176" refId="ref8919" refString="Runemark, H. (1967) Studies in the Aegean Flora XII. Cytologic and morphologic investigations in Centaurea. Botaniska Notiser 120: 161 - 176." type="journal article" year="1967">Runemark 1967</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF0FFEED0AB8BDFFD8FE5F6" author="Gardou, C." box="[393,543,2010,2037]" pageId="8" pageNumber="181" pagination="29 - 38" refId="ref7421" refString="Gardou, C. (1969) Caryosystematique des Centaurees de la section Acrocentron Cass. Bulletin de la Societe botanique de France 116: 29 - 38. https: // doi. org / 10.1080 / 00378941.1969.10838638" type="journal article" year="1969">Gardou 1969</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF0FFEED30B8BDFFDF4E5F7" author="Gardou, C." box="[553,612,2011,2037]" pageId="8" pageNumber="181" pagination="537 - 547" refId="ref7460" refString="Gardou, C. (1974) Quelques vues synthetiques sur les Centaurees de la section Acrocentron (Cass.) O. Hoffm. dans la flore Mediterraneenne. Colloques internationales du CNRS Montpellier. Flora Mediterranea 235: 537 - 547." type="journal article" year="1974">1974</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF0FFEED34D8BDFFD67E5F7" author="Phitos, D." box="[623,759,2011,2037]" pageId="8" pageNumber="181" pagination="69 - 73" refId="ref8133" refString="Phitos, D. (1970) Zur Polyploidie in der Gattung Centaurea L. Sektion Acrocentron (Compositae). Berichte der Deutschen Botanischen Gesellschaft 83: 69 - 73" type="journal article" year="1970">Phitos 1970</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF0FFEED2208BDFFCACE5F7" author="Phitos, D." box="[770,828,2011,2037]" pageId="8" pageNumber="181" pagination="255 - 259" refId="ref8164" refString="Phitos, D. (1971) Cytotaxonomische Studien der griechischen Centaurea - Arten, Sektion Acrocentron (Compositae). Berichte der Deutschen Botanischen Gesellschaft 84: 255 - 259." type="journal article" year="1971">1971</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF0FFEED26A8BDFFBE7E5F7" author="Damboldt, J. & Matthas, U." box="[840,1143,2010,2037]" pageId="8" pageNumber="181" pagination="103 - 115" refId="ref7090" refString="Damboldt, J. & Matthas, U. (1975) Chromosomenzahlen einiger mediterraner und mitteleuropaischer Centaurea - Arten (Asteraceae). PIant Systematics and Evolution 123: 103 - 115. https: // doi. org / 10.1007 / BF 00981769" type="journal article" year="1975">Damboldt & Matthäs 1975</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF0FFEED5A08BDFFA36E5F7" author="Phitos, D. & Georgiadis, Th." box="[1154,1446,2011,2037]" pageId="8" pageNumber="181" pagination="99 - 114" refId="ref8228" refString="Phitos, D. & Georgiadis, Th. (1981) A propos du grupe de Centaurea achaia Boiss. et Heldr. de la section Acrocentron (Cass.) DC. Botanika Chronika 1 (2): 99 - 114." type="journal article" year="1981">Phitos & Georgiadis 1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF0FFEED1AA8BFBFD87EA1B" author="Georgiadis, Th. & Christodoulakis, D." box="[136,535,2047,2073]" pageId="8" pageNumber="181" pagination="307 - 318" refId="ref7504" refString="Georgiadis, Th. & Christodoulakis, D. (1984) Contribution a l'etude cytogeographique des Centaurees de I'ile de Samos. Candollea 39: 307 - 318." type="journal article" year="1984">Georgiadis & Christodoulakis 1984</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF0FFEED3038BFBFCD8EA1B" author="Routsi, E. & Georgiadis, T." box="[545,840,2047,2073]" pageId="8" pageNumber="181" pagination="396 - 399" refId="ref8767" refString="Routsi, E. & Georgiadis, T. (1988) In: IOPB chromosome number reports XCIX. Taxon 37: 396 - 399." type="journal article" year="1988">Routsi & Georgiadis 1988</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF0FFEED2708BFBFC1DEA1B" author="Routsi, E. & Georgiadis, T." box="[850,909,2047,2073]" pageId="8" pageNumber="181" pagination="139 - 151" refId="ref8875" refString="Routsi, E. & Georgiadis, T. (1999) Cytogeographical study of Centaurea L. sect. Acrocentron (Cass.) DC. (Asteraceae) in Greece. Botanica Helvetica 109 (2): 139 - 151." type="journal article" year="1999">1999</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF0FFEED2B58BFBFBB1EA1B" author="Routsi, E." box="[919,1057,2047,2073]" pageId="8" pageNumber="181" refId="ref8730" refString="Routsi, E. (1993) Biosystematic study of the section Acrocentron (Cass.) DC. of the genus Centaurea L. in Greece. PhD Thesis, University of Patras, Greece." type="book" year="1993">Routsi 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF0FFEED50E8BFBFB4DEA1B" author="Font, M. & Valles, J. & Susanna, A. & Garcia-Jacas, N." box="[1068,1245,2047,2073]" pageId="8" pageNumber="181" pagination="7 - 18" refId="ref7236" refString="Font, M., Valles, J., Susanna, A. & Garcia-Jacas, N. (2008) Auto- and allopolyploidy in Centaurea sect. Acrocentron s. l. (Asteraceae, Cardueae): karyotype and fluorochrome banding pattern analyses. Collectanea Botanica (Barcelona) 27: 7 - 18. https: // doi. org / 10.3989 / collectbot. 2008. v 27.1" type="journal article" year="2008">
|
||
Font
|
||
<emphasis id="B924EA94FFF0FFEED5478BFBFB0DEA1B" box="[1125,1181,2047,2073]" italics="true" pageId="8" pageNumber="181">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF0FFEED5CA8BFBFA37EA1B" author="Uysal, T. & Ertugrul, K. & Susanna, A. & Garcia-Jacas, N." box="[1256,1447,2047,2073]" pageId="8" pageNumber="181" pagination="280 - 286" refId="ref9018" refString="Uysal, T., Ertugrul, K., Susanna, A. & Garcia-Jacas, N. (2009) New chromosome counts in the genus Centaurea (Asteraceae) from Turkey. Botanical Journal of the Linnean Society 159: 280 - 286. https: // doi. org / 10.1111 / j. 1095 - 8339.2008.00939. x" type="journal article" year="2009">
|
||
Uysal
|
||
<emphasis id="B924EA94FFF0FFEED40D8BFBFAF6EA1B" box="[1327,1382,2047,2073]" italics="true" pageId="8" pageNumber="181">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD1AA8C9AFE21E2BA" author="Ranjbar, M. & Negaresh, K." box="[136,433,158,184]" pageId="9" pageNumber="182" pagination="50 - 60" refId="ref8470" refString="Ranjbar, M. & Negaresh, K. (2013) A contribution to Centaurea sect. Acrocentron (Asteraceae) from Iran. Phytotaxa 149 (1): 50 - 60. https: // doi. org / 10.11646 / phytotaxa. 149.1.6" type="journal article" year="2013">Ranjbar & Negaresh 2013</bibRefCitation>
|
||
). These taxa show two basic chromosome numbers, x = 10 and x = 11 (
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD5EF8C9AFAC7E2BA" author="Routsi, E." box="[1229,1367,158,184]" pageId="9" pageNumber="182" refId="ref8730" refString="Routsi, E. (1993) Biosystematic study of the section Acrocentron (Cass.) DC. of the genus Centaurea L. in Greece. PhD Thesis, University of Patras, Greece." type="book" year="1993">Routsi 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD4408C9AFEF5E2DE" author="Routsi, E. & Georgiadis, T." pageId="9" pageNumber="182" pagination="139 - 151" refId="ref8875" refString="Routsi, E. & Georgiadis, T. (1999) Cytogeographical study of Centaurea L. sect. Acrocentron (Cass.) DC. (Asteraceae) in Greece. Botanica Helvetica 109 (2): 139 - 151." type="journal article" year="1999">Routsi & Georgiadis 1999</bibRefCitation>
|
||
). The basic chromosome number x = 11 is considered ancestral and is known only from diploid taxa. Additionally, it has been noted that the number of chromosome counts with x = 11 is more frequent in the eastern than the western Mediterranean region (
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD36F8D0EFC31E326" author="Garcia-Jacas, N. & Susanna, A." box="[589,929,266,292]" pageId="9" pageNumber="182" pagination="1 - 18" refId="ref7377" refString="Garcia-Jacas, N. & Susanna, A. (1992) Karyological notes on Centaurea sect. Acrocentron (Asteraceae). Plant Systematics and Evolution 179: 1 - 18. https: // doi. org / 10.1007 / BF 00938015" type="journal article" year="1992">Garcia-Jacas & Susanna 1992</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD28F8D0EFBE4E326" author="Uysal, T. & Ertugrul, K. & Susanna, A. & Garcia-Jacas, N." box="[941,1140,266,292]" pageId="9" pageNumber="182" pagination="280 - 286" refId="ref9018" refString="Uysal, T., Ertugrul, K., Susanna, A. & Garcia-Jacas, N. (2009) New chromosome counts in the genus Centaurea (Asteraceae) from Turkey. Botanical Journal of the Linnean Society 159: 280 - 286. https: // doi. org / 10.1111 / j. 1095 - 8339.2008.00939. x" type="journal article" year="2009">
|
||
Uysal
|
||
<emphasis id="B924EA94FFF1FFEFD2D48D0FFBA0E326" box="[1014,1072,266,292]" italics="true" pageId="9" pageNumber="182">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
). On the other hand, x = 10 can be observed in both diploids and polyploids reaching up to 2
|
||
<emphasis id="B924EA94FFF1FFEFD2768D2BFCF2E34A" box="[852,866,303,328]" italics="true" pageId="9" pageNumber="182">n</emphasis>
|
||
= 10x and 2
|
||
<emphasis id="B924EA94FFF1FFEFD2CE8D2BFC6AE34A" box="[1004,1018,303,328]" italics="true" pageId="9" pageNumber="182">n</emphasis>
|
||
= 11x in material of
|
||
<taxonomicName id="4C504D05FFF1FFEFD5C38D2BFF7AE36E" authority="Boiss. & Heldr." authorityName="Boiss. & Heldr." class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="182" phylum="Tracheophyta" rank="species" species="spruneri">
|
||
<emphasis id="B924EA94FFF1FFEFD5C38D2BFAF0E34A" box="[1249,1376,303,329]" italics="true" pageId="9" pageNumber="182">C. spruneri</emphasis>
|
||
Boiss. & Heldr.
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD0298D56FE7FE36E" author="Boissier, E." box="[267,495,338,365]" pageId="9" pageNumber="182" refId="ref6898" refString="Boissier, E. (1845) Diagnoses plantarum orientalium novarum, ser. 1, 6. B. Herrmann, Lipsia, 136 pp." type="book" year="1845">Boissier (1845: 132)</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD0DF8D56FD16E36E" author="Phitos, D." box="[509,646,338,364]" pageId="9" pageNumber="182" pagination="69 - 73" refId="ref8133" refString="Phitos, D. (1970) Zur Polyploidie in der Gattung Centaurea L. Sektion Acrocentron (Compositae). Berichte der Deutschen Botanischen Gesellschaft 83: 69 - 73" type="journal article" year="1970">Phitos 1970</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD3B28D56FD5AE36E" author="Phitos, D." box="[656,714,338,364]" pageId="9" pageNumber="182" pagination="255 - 259" refId="ref8164" refString="Phitos, D. (1971) Cytotaxonomische Studien der griechischen Centaurea - Arten, Sektion Acrocentron (Compositae). Berichte der Deutschen Botanischen Gesellschaft 84: 255 - 259." type="journal article" year="1971">1971</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD3F78D56FC42E36E" author="Phitos, D. & Kamari, G." box="[725,978,338,364]" pageId="9" pageNumber="182" pagination="147 - 151" refId="ref8196" refString="Phitos, D. & Kamari, G. (1973) Cytologische Untersuchungen an einigen griechischen Blutenpflanzen (Centaurea, Aethionema). Biologia Gallo-Hellenica 4: 147 - 151." type="journal article" year="1973">Phitos & Kamari 1973</bibRefCitation>
|
||
). Most of the karyotypes belonging to taxa of section
|
||
<taxonomicName id="4C504D05FFF1FFEFD1DC8D73FE18E393" box="[254,392,375,401]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="182" phylum="Tracheophyta" rank="section" section="Acrocentron">
|
||
<emphasis id="B924EA94FFF1FFEFD1DC8D73FE18E393" box="[254,392,375,401]" italics="true" pageId="9" pageNumber="182">Acrocentron</emphasis>
|
||
</taxonomicName>
|
||
consist of metacentric and submetacentric chromosomes as well as satellited chromosomes in many cases (
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD03B8D9EFE34E3B6" author="Routsi, E." box="[281,420,410,436]" pageId="9" pageNumber="182" refId="ref8730" refString="Routsi, E. (1993) Biosystematic study of the section Acrocentron (Cass.) DC. of the genus Centaurea L. in Greece. PhD Thesis, University of Patras, Greece." type="book" year="1993">Routsi 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD0928D9EFD4AE3B6" author="Routsi, E. & Georgiadis, T." box="[432,730,410,436]" pageId="9" pageNumber="182" pagination="139 - 151" refId="ref8875" refString="Routsi, E. & Georgiadis, T. (1999) Cytogeographical study of Centaurea L. sect. Acrocentron (Cass.) DC. (Asteraceae) in Greece. Botanica Helvetica 109 (2): 139 - 151." type="journal article" year="1999">Routsi & Georgiadis 1999</bibRefCitation>
|
||
). The
|
||
<taxonomicName id="4C504D05FFF1FFEFD2028D9FFC1DE3B6" box="[800,909,410,437]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="182" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF1FFEFD2028D9FFC1DE3B6" box="[800,909,410,437]" italics="true" pageId="9" pageNumber="182">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
group comprises taxa with x = 11 (
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD43E8D9EFA37E3B6" author="Routsi, E." box="[1308,1447,410,436]" pageId="9" pageNumber="182" refId="ref8730" refString="Routsi, E. (1993) Biosystematic study of the section Acrocentron (Cass.) DC. of the genus Centaurea L. in Greece. PhD Thesis, University of Patras, Greece." type="book" year="1993">Routsi 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD1AA8DBAFE21E3DA" author="Routsi, E. & Georgiadis, T." box="[136,433,446,472]" pageId="9" pageNumber="182" pagination="139 - 151" refId="ref8875" refString="Routsi, E. & Georgiadis, T. (1999) Cytogeographical study of Centaurea L. sect. Acrocentron (Cass.) DC. (Asteraceae) in Greece. Botanica Helvetica 109 (2): 139 - 151." type="journal article" year="1999">Routsi & Georgiadis 1999</bibRefCitation>
|
||
) including
|
||
<taxonomicName id="4C504D05FFF1FFEFD3108DBBFC81E3DA" box="[562,785,446,473]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="182" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFF1FFEFD3108DBBFC81E3DA" box="[562,785,446,473]" italics="true" pageId="9" pageNumber="182">Centaurea dolopica</emphasis>
|
||
</taxonomicName>
|
||
. The karyotype of the new species (
|
||
<figureCitation id="136B2A03FFF1FFEFD5888DBAFB7BE3DB" box="[1194,1259,446,473]" captionStart="FIGURE 5" captionStartId="8.[136,229,586,608]" captionTargetBox="[276,1312,190,552]" captionTargetId="figure-29@8.[276,1312,190,552]" captionTargetPageId="8" captionText="FIGURE 5. A) Karyotype and B) idiogram of Centaurea dolopica. Arrows indicate satellites." figureDoi="http://doi.org/10.5281/zenodo.7645665" httpUri="https://zenodo.org/record/7645665/files/figure.png" pageId="9" pageNumber="182">Fig. 5</figureCitation>
|
||
) is similar to the karyotypes of the other taxa of
|
||
<taxonomicName id="4C504D05FFF1FFEFD0C28DE7FDDCE3FE" box="[480,588,482,509]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="182" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF1FFEFD0C28DE7FDDCE3FE" box="[480,588,482,509]" italics="true" pageId="9" pageNumber="182">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD3818DE6FCBDE3FE" author="Routsi, E." box="[675,813,482,508]" pageId="9" pageNumber="182" refId="ref8730" refString="Routsi, E. (1993) Biosystematic study of the section Acrocentron (Cass.) DC. of the genus Centaurea L. in Greece. PhD Thesis, University of Patras, Greece." type="book" year="1993">Routsi 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD2158DE6FBCFE3FE" author="Routsi, E. & Georgiadis, T." box="[823,1119,482,508]" pageId="9" pageNumber="182" pagination="139 - 151" refId="ref8875" refString="Routsi, E. & Georgiadis, T. (1999) Cytogeographical study of Centaurea L. sect. Acrocentron (Cass.) DC. (Asteraceae) in Greece. Botanica Helvetica 109 (2): 139 - 151." type="journal article" year="1999">Routsi & Georgiadis 1999</bibRefCitation>
|
||
). The presence of two pairs of SAT-chromosomes in
|
||
<taxonomicName id="4C504D05FFF1FFEFD0A28E03FD93E022" box="[384,515,518,545]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="182" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFF1FFEFD0A28E03FD93E022" box="[384,515,518,545]" italics="true" pageId="9" pageNumber="182">C. dolopica</emphasis>
|
||
</taxonomicName>
|
||
’s karyotype has been also observed in the material of
|
||
<taxonomicName id="4C504D05FFF1FFEFD5478E03FB42E022" box="[1125,1234,518,545]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="182" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF1FFEFD5478E03FB42E022" box="[1125,1234,518,545]" italics="true" pageId="9" pageNumber="182">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C504D05FFF1FFEFD42B8E03FA3BE022" box="[1289,1451,518,545]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="182" phylum="Tracheophyta" rank="species" species="corinthiaca">
|
||
<emphasis id="B924EA94FFF1FFEFD42B8E03FA3BE022" box="[1289,1451,518,545]" italics="true" pageId="9" pageNumber="182">C. corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="4994F864FFF1FFEFD1E68E2EFEC2E046" box="[196,338,554,580]" country="Greece" name="Peloponnisos" pageId="9" pageNumber="182">Peloponnese</collectingRegion>
|
||
and Sterea
|
||
<collectingCountry id="F3477616FFF1FFEFD0F08E2EFD8CE046" box="[466,540,554,580]" name="Greece" pageId="9" pageNumber="182">Hellas</collectingCountry>
|
||
, respectively (
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD39E8E2EFC70E046" author="Phitos, D. & Georgiadis, Th." box="[700,992,554,580]" pageId="9" pageNumber="182" pagination="99 - 114" refId="ref8228" refString="Phitos, D. & Georgiadis, Th. (1981) A propos du grupe de Centaurea achaia Boiss. et Heldr. de la section Acrocentron (Cass.) DC. Botanika Chronika 1 (2): 99 - 114." type="journal article" year="1981">Phitos & Georgiadis 1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD2C98E2EFBE5E046" author="Routsi, E." box="[1003,1141,554,580]" pageId="9" pageNumber="182" refId="ref8730" refString="Routsi, E. (1993) Biosystematic study of the section Acrocentron (Cass.) DC. of the genus Centaurea L. in Greece. PhD Thesis, University of Patras, Greece." type="book" year="1993">Routsi 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD55D8E2EFA33E046" author="Routsi, E. & Georgiadis, T." box="[1151,1443,554,580]" pageId="9" pageNumber="182" pagination="139 - 151" refId="ref8875" refString="Routsi, E. & Georgiadis, T. (1999) Cytogeographical study of Centaurea L. sect. Acrocentron (Cass.) DC. (Asteraceae) in Greece. Botanica Helvetica 109 (2): 139 - 151." type="journal article" year="1999">Routsi & Georgiadis 1999</bibRefCitation>
|
||
) and also in
|
||
<taxonomicName id="4C504D05FFF1FFEFD02B8E4BFE13E06A" box="[265,387,590,617]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="182" phylum="Tracheophyta" rank="species" species="aetolica">
|
||
<emphasis id="B924EA94FFF1FFEFD02B8E4BFE13E06A" box="[265,387,590,617]" italics="true" pageId="9" pageNumber="182">C. aetolica</emphasis>
|
||
</taxonomicName>
|
||
from Sterea
|
||
<collectingCountry id="F3477616FFF1FFEFD3378E4AFDCDE06A" box="[533,605,590,616]" name="Greece" pageId="9" pageNumber="182">Hellas</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFC14B77FFF1FFEFD3498E4AFC04E06A" author="Phitos, D. & Georgiadis, Th." box="[619,916,590,616]" pageId="9" pageNumber="182" pagination="99 - 114" refId="ref8228" refString="Phitos, D. & Georgiadis, Th. (1981) A propos du grupe de Centaurea achaia Boiss. et Heldr. de la section Acrocentron (Cass.) DC. Botanika Chronika 1 (2): 99 - 114." type="journal article" year="1981">Phitos & Georgiadis 1981</bibRefCitation>
|
||
). These findings corroborate the inclusion of
|
||
<taxonomicName id="4C504D05FFF1FFEFD4B08E4BFF7AE08E" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="182" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFF1FFEFD4B08E4BFF7AE08E" italics="true" pageId="9" pageNumber="182">C. dolopica</emphasis>
|
||
</taxonomicName>
|
||
in the
|
||
<taxonomicName id="4C504D05FFF1FFEFD0148E77FE33E08E" box="[310,419,626,653]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="182" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF1FFEFD0148E77FE33E08E" box="[310,419,626,653]" italics="true" pageId="9" pageNumber="182">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
group and their close affinities. Moreover, a karyomorphometric study for all taxa related to
|
||
<taxonomicName id="4C504D05FFF1FFEFD1AA8E93FE9BE0B2" box="[136,267,662,689]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="182" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFF1FFEFD1AA8E93FE9BE0B2" box="[136,267,662,689]" italics="true" pageId="9" pageNumber="182">C. dolopica</emphasis>
|
||
</taxonomicName>
|
||
could be valuable to give a further insight on their relationships.
|
||
</paragraph>
|
||
<caption id="DF2F660EFFF1FFEFD1AA8BD1FCE6E5E9" ID-DOI="http://doi.org/10.5281/zenodo.7645669" ID-Zenodo-Dep="7645669" box="[136,886,2005,2028]" httpUri="https://zenodo.org/record/7645669/files/figure.png" pageId="9" pageNumber="182" startId="9.[136,229,2005,2027]" targetBox="[147,1439,716,1979]" targetPageId="9">
|
||
<paragraph id="8BEF3686FFF1FFEFD1AA8BD1FCE6E5E9" blockId="9.[136,886,2005,2028]" box="[136,886,2005,2028]" pageId="9" pageNumber="182">
|
||
<emphasis id="B924EA94FFF1FFEFD1AA8BD1FF67E5E9" bold="true" box="[136,247,2005,2027]" pageId="9" pageNumber="182">FIGURE 7</emphasis>
|
||
. Distribution map of the members of the
|
||
<taxonomicName id="4C504D05FFF1FFEFD3A78BD2FCA1E5E9" box="[645,817,2006,2027]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="182" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF1FFEFD3A78BD2FCA1E5E9" box="[645,817,2006,2027]" italics="true" pageId="9" pageNumber="182">Centaurea achaia</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEF3686FFF2FFECD19F8C9AFD50E73A" blockId="10.[136,1452,158,1336]" pageId="10" pageNumber="183">
|
||
Phylogenetic analyses revealed the placement of
|
||
<taxonomicName id="4C504D05FFF2FFECD3CE8C9BFC5CE2BA" box="[748,972,158,185]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFF2FFECD3CE8C9BFC5CE2BA" box="[748,972,158,185]" italics="true" pageId="10" pageNumber="183">Centaurea dolopica</emphasis>
|
||
</taxonomicName>
|
||
in the Greek-Aegean clade of the
|
||
<emphasis id="B924EA94FFF2FFECD4758C9BFAE1E2BB" box="[1367,1393,159,185]" italics="true" pageId="10" pageNumber="183">C.</emphasis>
|
||
sect.
|
||
<taxonomicName id="4C504D05FFF2FFECD1AA8CC7FE82E2DF" box="[136,274,195,221]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="185" phylum="Tracheophyta" rank="section" section="Acrocentron">
|
||
<emphasis id="B924EA94FFF2FFECD1AA8CC7FE82E2DF" box="[136,274,195,221]" italics="true" pageId="10" pageNumber="183">Acrocentron</emphasis>
|
||
</taxonomicName>
|
||
in close evolutionary relationship with representatives of the
|
||
<taxonomicName id="4C504D05FFF2FFECD2EB8CC7FBA6E2DE" box="[969,1078,194,221]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD2EB8CC7FBA6E2DE" box="[969,1078,194,221]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<figureCitation id="136B2A03FFF2FFECD5B28CC6FB44E2DE" box="[1168,1236,194,220]" captionStart="FIGURE 6" captionStartId="8.[136,229,1690,1712]" captionTargetBox="[153,1436,722,1660]" captionTargetId="figure-48@8.[151,1436,633,1665]" captionTargetPageId="8" captionText="FIGURE 6. Phylogeny of Centaurea subg. Lopholoma sect. Acrocentron, including the newly described Centaurea dolopica (in bold), with Centaurea subg. Cyanus as outgroup, inferred from Bayesian analysis of the concatenated ITS and ETS sequences. Posterior probabilities (pp) higher than 0.7 and bootstrap values (bs) higher than 50% estimated from MrBayes and RAxML analyses are reported above and below the branches, respectively.At the right, the names of the major clades in C. subg. Lopholoma correspond to those of Font et al.’s (2009) phylogenetic trees." figureDoi="http://doi.org/10.5281/zenodo.7645667" httpUri="https://zenodo.org/record/7645667/files/figure.png" pageId="10" pageNumber="183">Fig. 6</figureCitation>
|
||
), corroborating the morphological data. Specifically,
|
||
<taxonomicName id="4C504D05FFF2FFECD3248CE3FD1BE302" box="[518,651,230,257]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFF2FFECD3248CE3FD1BE302" box="[518,651,230,257]" italics="true" pageId="10" pageNumber="183">C. dolopica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C504D05FFF2FFECD3E68CE3FCA3E302" box="[708,819,230,257]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD3E68CE3FCA3E302" box="[708,819,230,257]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
from its
|
||
<emphasis id="B924EA94FFF2FFECD2BC8CE2FBD7E302" box="[926,1095,230,256]" italics="true" pageId="10" pageNumber="183">locus classicus</emphasis>
|
||
in Mt. Chelmos (
|
||
<collectingRegion id="4994F864FFF2FFECD4318CE2FA34E302" box="[1299,1444,230,256]" country="Greece" name="Peloponnisos" pageId="10" pageNumber="183">Peloponnese</collectingRegion>
|
||
) form a well-supported clade (bs=82 and pp=0.98;
|
||
<figureCitation id="136B2A03FFF2FFECD3E98D0EFC80E326" box="[715,784,266,292]" captionStart="FIGURE 6" captionStartId="8.[136,229,1690,1712]" captionTargetBox="[153,1436,722,1660]" captionTargetId="figure-48@8.[151,1436,633,1665]" captionTargetPageId="8" captionText="FIGURE 6. Phylogeny of Centaurea subg. Lopholoma sect. Acrocentron, including the newly described Centaurea dolopica (in bold), with Centaurea subg. Cyanus as outgroup, inferred from Bayesian analysis of the concatenated ITS and ETS sequences. Posterior probabilities (pp) higher than 0.7 and bootstrap values (bs) higher than 50% estimated from MrBayes and RAxML analyses are reported above and below the branches, respectively.At the right, the names of the major clades in C. subg. Lopholoma correspond to those of Font et al.’s (2009) phylogenetic trees." figureDoi="http://doi.org/10.5281/zenodo.7645667" httpUri="https://zenodo.org/record/7645667/files/figure.png" pageId="10" pageNumber="183">Fig. 6</figureCitation>
|
||
) sister to
|
||
<taxonomicName id="4C504D05FFF2FFECD2A78D0FFB92E326" box="[901,1026,266,293]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="aetolica">
|
||
<emphasis id="B924EA94FFF2FFECD2A78D0FFB92E326" box="[901,1026,266,293]" italics="true" pageId="10" pageNumber="183">C. aetolica</emphasis>
|
||
</taxonomicName>
|
||
(bs=95 and pp=1;
|
||
<figureCitation id="136B2A03FFF2FFECD5C28D0EFAB6E326" box="[1248,1318,266,292]" captionStart="FIGURE 6" captionStartId="8.[136,229,1690,1712]" captionTargetBox="[153,1436,722,1660]" captionTargetId="figure-48@8.[151,1436,633,1665]" captionTargetPageId="8" captionText="FIGURE 6. Phylogeny of Centaurea subg. Lopholoma sect. Acrocentron, including the newly described Centaurea dolopica (in bold), with Centaurea subg. Cyanus as outgroup, inferred from Bayesian analysis of the concatenated ITS and ETS sequences. Posterior probabilities (pp) higher than 0.7 and bootstrap values (bs) higher than 50% estimated from MrBayes and RAxML analyses are reported above and below the branches, respectively.At the right, the names of the major clades in C. subg. Lopholoma correspond to those of Font et al.’s (2009) phylogenetic trees." figureDoi="http://doi.org/10.5281/zenodo.7645667" httpUri="https://zenodo.org/record/7645667/files/figure.png" pageId="10" pageNumber="183">Fig. 6</figureCitation>
|
||
). The other species of the
|
||
<taxonomicName id="4C504D05FFF2FFECD00B8D2BFE06E34A" box="[297,406,302,329]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD00B8D2BFE06E34A" box="[297,406,302,329]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
group, namely
|
||
<taxonomicName id="4C504D05FFF2FFECD36A8D2BFD7AE34A" box="[584,746,302,329]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="corinthiaca">
|
||
<emphasis id="B924EA94FFF2FFECD36A8D2BFD7AE34A" box="[584,746,302,329]" italics="true" pageId="10" pageNumber="183">C. corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C504D05FFF2FFECD2038D2BFC0CE34A" box="[801,924,302,329]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="euboica">
|
||
<emphasis id="B924EA94FFF2FFECD2038D2BFC0CE34A" box="[801,924,302,329]" italics="true" pageId="10" pageNumber="183">C. euboica</emphasis>
|
||
</taxonomicName>
|
||
, together with an individual of
|
||
<taxonomicName id="4C504D05FFF2FFECD4228D2BFAFDE34A" box="[1280,1389,302,329]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD4228D2BFAFDE34A" box="[1280,1389,302,329]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
from the Vilia area in
|
||
<collectingRegion id="4994F864FFF2FFECD01A8D56FEEFE36E" box="[312,383,338,364]" country="Greece" name="Attiki" pageId="10" pageNumber="183">Attica</collectingRegion>
|
||
,
|
||
<taxonomicName id="4C504D05FFF2FFECD0AB8D56FD89E36E" box="[393,537,338,364]" form="another" pageId="10" pageNumber="183" rank="form">form another</taxonomicName>
|
||
well-supported clade (bs=99 and pp=1;
|
||
<figureCitation id="136B2A03FFF2FFECD2F68D56FB85E36E" box="[980,1045,338,364]" captionStart="FIGURE 6" captionStartId="8.[136,229,1690,1712]" captionTargetBox="[153,1436,722,1660]" captionTargetId="figure-48@8.[151,1436,633,1665]" captionTargetPageId="8" captionText="FIGURE 6. Phylogeny of Centaurea subg. Lopholoma sect. Acrocentron, including the newly described Centaurea dolopica (in bold), with Centaurea subg. Cyanus as outgroup, inferred from Bayesian analysis of the concatenated ITS and ETS sequences. Posterior probabilities (pp) higher than 0.7 and bootstrap values (bs) higher than 50% estimated from MrBayes and RAxML analyses are reported above and below the branches, respectively.At the right, the names of the major clades in C. subg. Lopholoma correspond to those of Font et al.’s (2009) phylogenetic trees." figureDoi="http://doi.org/10.5281/zenodo.7645667" httpUri="https://zenodo.org/record/7645667/files/figure.png" pageId="10" pageNumber="183">Fig. 6</figureCitation>
|
||
) sister to
|
||
<taxonomicName id="4C504D05FFF2FFECD55C8D57FF2FE392" authority="Moore (1878: 133)" authorityName="Moore" authorityPageNumber="133" authorityYear="1878" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="ebenoides">
|
||
<emphasis id="B924EA94FFF2FFECD55C8D57FA9DE36E" box="[1150,1293,338,365]" italics="true" pageId="10" pageNumber="183">C. ebenoides</emphasis>
|
||
<bibRefCitation id="EFC14B77FFF2FFECD4308D57FF2FE392" author="Moore, S. L. M." pageId="10" pageNumber="183" pagination="129 - 138" refId="ref8036" refString="Moore, S. L. M. (1878) Alabastra diversa. Journal of Botany 16: 129 - 138." type="journal article" year="1878">Moore (1878: 133)</bibRefCitation>
|
||
</taxonomicName>
|
||
; yet this sister relationship is only moderately supported (bs=76 and pp=0.89;
|
||
<figureCitation id="136B2A03FFF2FFECD5148D72FBEAE392" box="[1078,1146,374,400]" captionStart="FIGURE 6" captionStartId="8.[136,229,1690,1712]" captionTargetBox="[153,1436,722,1660]" captionTargetId="figure-48@8.[151,1436,633,1665]" captionTargetPageId="8" captionText="FIGURE 6. Phylogeny of Centaurea subg. Lopholoma sect. Acrocentron, including the newly described Centaurea dolopica (in bold), with Centaurea subg. Cyanus as outgroup, inferred from Bayesian analysis of the concatenated ITS and ETS sequences. Posterior probabilities (pp) higher than 0.7 and bootstrap values (bs) higher than 50% estimated from MrBayes and RAxML analyses are reported above and below the branches, respectively.At the right, the names of the major clades in C. subg. Lopholoma correspond to those of Font et al.’s (2009) phylogenetic trees." figureDoi="http://doi.org/10.5281/zenodo.7645667" httpUri="https://zenodo.org/record/7645667/files/figure.png" pageId="10" pageNumber="183">Fig. 6</figureCitation>
|
||
).
|
||
<taxonomicName id="4C504D05FFF2FFECD5B28D73FAEDE392" authorityName="Moore" authorityYear="1878" box="[1168,1405,374,401]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="ebenoides">
|
||
<emphasis id="B924EA94FFF2FFECD5B28D73FAEDE392" box="[1168,1405,374,401]" italics="true" pageId="10" pageNumber="183">Centaurea ebenoides</emphasis>
|
||
</taxonomicName>
|
||
is a Greek endemic species with a narrow distribution in Euboea and
|
||
<collectingRegion id="4994F864FFF2FFECD2568D9EFC28E3B6" box="[884,952,410,436]" country="Greece" name="Attiki" pageId="10" pageNumber="183">Attica</collectingRegion>
|
||
that is morphologically very different from the
|
||
<taxonomicName id="4C504D05FFF2FFECD1908DBBFE8FE3DA" box="[178,287,446,473]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD1908DBBFE8FE3DA" box="[178,287,446,473]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
group as it has a short, procumbent habit and very small, inconspicuous bract appendages. Although in the current phylogeny, the
|
||
<taxonomicName id="4C504D05FFF2FFECD0978DE7FDB2E3FE" box="[437,546,482,509]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD0978DE7FDB2E3FE" box="[437,546,482,509]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
group is not monophyletic due to the phylogenetic position of
|
||
<taxonomicName id="4C504D05FFF2FFECD5CB8DE7FAEBE3FE" authorityName="Moore" authorityYear="1878" box="[1257,1403,482,509]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="ebenoides">
|
||
<emphasis id="B924EA94FFF2FFECD5CB8DE7FAEBE3FE" box="[1257,1403,482,509]" italics="true" pageId="10" pageNumber="183">C. ebenoides</emphasis>
|
||
</taxonomicName>
|
||
, the incomplete taxon sampling of this study, especially regarding other related species to
|
||
<taxonomicName id="4C504D05FFF2FFECD5698E03FB4CE022" authorityName="Moore" authorityYear="1878" box="[1099,1244,518,545]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="ebenoides">
|
||
<emphasis id="B924EA94FFF2FFECD5698E03FB4CE022" box="[1099,1244,518,545]" italics="true" pageId="10" pageNumber="183">C. ebenoides</emphasis>
|
||
</taxonomicName>
|
||
, and the moderate support of the sister relationship of
|
||
<taxonomicName id="4C504D05FFF2FFECD30A8E2FFD2CE046" authorityName="Moore" authorityYear="1878" box="[552,700,554,581]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="ebenoides">
|
||
<emphasis id="B924EA94FFF2FFECD30A8E2FFD2CE046" box="[552,700,554,581]" italics="true" pageId="10" pageNumber="183">C. ebenoides</emphasis>
|
||
</taxonomicName>
|
||
to one of the two well-supported clades of the
|
||
<taxonomicName id="4C504D05FFF2FFECD5CB8E2FFAC9E046" box="[1257,1369,554,581]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD5CB8E2FFAC9E046" box="[1257,1369,554,581]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
group, cannot confidently reject the monophyly of the
|
||
<taxonomicName id="4C504D05FFF2FFECD3818E4BFC81E06A" box="[675,785,590,617]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD3818E4BFC81E06A" box="[675,785,590,617]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
group. The division of the clade comprising species of the
|
||
<taxonomicName id="4C504D05FFF2FFECD1AA8E77FF65E08E" box="[136,245,626,653]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD1AA8E77FF65E08E" box="[136,245,626,653]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
group and
|
||
<taxonomicName id="4C504D05FFF2FFECD0518E77FD93E08E" authorityName="Moore" authorityYear="1878" box="[371,515,626,653]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="ebenoides">
|
||
<emphasis id="B924EA94FFF2FFECD0518E77FD93E08E" box="[371,515,626,653]" italics="true" pageId="10" pageNumber="183">C. ebenoides</emphasis>
|
||
</taxonomicName>
|
||
into two subclades demonstrates an interesting biogeographic pattern of speciation. Specifically, species diversification in the subclade of
|
||
<taxonomicName id="4C504D05FFF2FFECD3D68E93FCE9E0B2" box="[756,889,662,689]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="dolopica">
|
||
<emphasis id="B924EA94FFF2FFECD3D68E93FCE9E0B2" box="[756,889,662,689]" italics="true" pageId="10" pageNumber="183">C. dolopica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C504D05FFF2FFECD2AB8E93FBE7E0B2" authority="(Chelmos)" baseAuthorityName="Chelmos" box="[905,1143,662,689]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD2AB8E93FC67E0B2" box="[905,1015,662,689]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
(Chelmos)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C504D05FFF2FFECD5928E93FABDE0B2" box="[1200,1325,662,689]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="aetolica">
|
||
<emphasis id="B924EA94FFF2FFECD5928E93FABDE0B2" box="[1200,1325,662,689]" italics="true" pageId="10" pageNumber="183">C. aetolica</emphasis>
|
||
</taxonomicName>
|
||
took place in north
|
||
<collectingRegion id="4994F864FFF2FFECD1CF8EBEFEEDE0D6" box="[237,381,698,724]" country="Greece" name="Peloponnisos" pageId="10" pageNumber="183">Peloponnese</collectingRegion>
|
||
,
|
||
<collectingRegion id="4994F864FFF2FFECD0A98EBFFD8BE0D6" box="[395,539,698,725]" country="Greece" name="Dytiki Ellada" pageId="10" pageNumber="183">West Greece</collectingRegion>
|
||
and East
|
||
<collectingRegion id="4994F864FFF2FFECD3B08EBEFCD0E0D6" box="[658,832,698,724]" country="Greece" name="Sterea Ellada" pageId="10" pageNumber="183">Central Greece</collectingRegion>
|
||
, whereas species diversification in the other subclade took place further eastwards, in the areas of
|
||
<collectingRegion id="4994F864FFF2FFECD3598EDAFD2FE0FA" box="[635,703,734,760]" country="Greece" name="Attiki" pageId="10" pageNumber="183">Attica</collectingRegion>
|
||
and Euboea (
|
||
<figureCitation id="136B2A03FFF2FFECD27E8EDAFC30E0FA" box="[860,928,734,760]" captionStart="FIGURE 7" captionStartId="9.[136,229,2005,2027]" captionTargetBox="[147,1439,716,1979]" captionTargetId="figure-349@9.[145,1442,713,1981]" captionTargetPageId="9" captionText="FIGURE 7. Distribution map of the members of the Centaurea achaia group." figureDoi="http://doi.org/10.5281/zenodo.7645669" httpUri="https://zenodo.org/record/7645669/files/figure.png" pageId="10" pageNumber="183">Fig. 7</figureCitation>
|
||
). This pattern could suggest the existence of a western and an eastern ancestor in
|
||
<collectingRegion id="4994F864FFF2FFECD3328F06FD22E11E" box="[528,690,770,796]" country="Greece" name="Sterea Ellada" pageId="10" pageNumber="183">central Greece</collectingRegion>
|
||
that diversified allopatrically giving rise to the two aforementioned lineages. Additional taxon sampling and ancestral range estimation analysis in a phylogenetic framework would be required to clarify the biogeographic origins of this clade. Another remarkable finding in this phylogenetic study is the non-monophyletic
|
||
<taxonomicName id="4C504D05FFF2FFECD0AA8F6BFE66E18A" box="[392,502,878,905]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD0AA8F6BFE66E18A" box="[392,502,878,905]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
, which raises questions about the species limits. In the most recent circumscription (FoG Web 2022),
|
||
<taxonomicName id="4C504D05FFF2FFECD0738F97FE2DE1AE" box="[337,445,914,941]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD0738F97FE2DE1AE" box="[337,445,914,941]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
comprises two subspecies, i.e.,
|
||
<taxonomicName id="4C504D05FFF2FFECD2038F97FBBEE1AE" box="[801,1070,914,941]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="subSpecies" species="achaia" subSpecies="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD2038F97FC1EE1AE" box="[801,910,914,941]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
subsp.
|
||
<emphasis id="B924EA94FFF2FFECD2C08F96FBBEE1AE" box="[994,1070,914,940]" italics="true" pageId="10" pageNumber="183">achaia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C504D05FFF2FFECD5418F97FA34E1AE" authorityName="Phitos & Georgiadis" authorityYear="1981" box="[1123,1444,914,941]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="subSpecies" species="achaia" subSpecies="corinthiaca">
|
||
<emphasis id="B924EA94FFF2FFECD5418F97FB5FE1AE" box="[1123,1231,914,941]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
subsp.
|
||
<emphasis id="B924EA94FFF2FFECD4018F96FA34E1AE" box="[1315,1444,914,940]" italics="true" pageId="10" pageNumber="183">corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
. Based on our results, the sample of
|
||
<taxonomicName id="4C504D05FFF2FFECD3018FB3FD01E1D2" box="[547,657,950,977]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD3018FB3FD01E1D2" box="[547,657,950,977]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<emphasis id="B924EA94FFF2FFECD2268FB2FC3CE1D2" box="[772,940,950,976]" italics="true" pageId="10" pageNumber="183">locus classicus</emphasis>
|
||
(Chelmos) and that of
|
||
<taxonomicName id="4C504D05FFF2FFECD5958FB3FACAE1D2" box="[1207,1370,950,977]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="corinthiaca">
|
||
<emphasis id="B924EA94FFF2FFECD5958FB3FACAE1D2" box="[1207,1370,950,977]" italics="true" pageId="10" pageNumber="183">C. corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
do not form a monophyletic group but are placed in two different well-supported clades (
|
||
<figureCitation id="136B2A03FFF2FFECD5118FDEFBE8E1F6" box="[1075,1144,986,1012]" captionStart="FIGURE 6" captionStartId="8.[136,229,1690,1712]" captionTargetBox="[153,1436,722,1660]" captionTargetId="figure-48@8.[151,1436,633,1665]" captionTargetPageId="8" captionText="FIGURE 6. Phylogeny of Centaurea subg. Lopholoma sect. Acrocentron, including the newly described Centaurea dolopica (in bold), with Centaurea subg. Cyanus as outgroup, inferred from Bayesian analysis of the concatenated ITS and ETS sequences. Posterior probabilities (pp) higher than 0.7 and bootstrap values (bs) higher than 50% estimated from MrBayes and RAxML analyses are reported above and below the branches, respectively.At the right, the names of the major clades in C. subg. Lopholoma correspond to those of Font et al.’s (2009) phylogenetic trees." figureDoi="http://doi.org/10.5281/zenodo.7645667" httpUri="https://zenodo.org/record/7645667/files/figure.png" pageId="10" pageNumber="183">Fig. 6</figureCitation>
|
||
), thus rejecting the current classification of
|
||
<taxonomicName id="4C504D05FFF2FFECD0638FFBFE72E61A" box="[321,482,1022,1049]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="corinthiaca">
|
||
<emphasis id="B924EA94FFF2FFECD0638FFBFE72E61A" box="[321,482,1022,1049]" italics="true" pageId="10" pageNumber="183">C. corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
as a subspecies of
|
||
<taxonomicName id="4C504D05FFF2FFECD3948FFBFCB3E61A" box="[694,803,1022,1049]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD3948FFBFCB3E61A" box="[694,803,1022,1049]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
. Furthermore, the sample of
|
||
<taxonomicName id="4C504D05FFF2FFECD5458FFBFB44E61A" box="[1127,1236,1022,1049]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD5458FFBFB44E61A" box="[1127,1236,1022,1049]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
from Vilia (
|
||
<collectingRegion id="4994F864FFF2FFECD47E8FFAFA34E61A" box="[1372,1444,1022,1048]" country="Greece" name="Attiki" pageId="10" pageNumber="183">Attica</collectingRegion>
|
||
) is not sister to
|
||
<taxonomicName id="4C504D05FFF2FFECD0088827FE07E63E" box="[298,407,1058,1085]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD0088827FE07E63E" box="[298,407,1058,1085]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
from Chelmos, indicating that this species, even after the exclusion of
|
||
<taxonomicName id="4C504D05FFF2FFECD58D8827FAC1E63E" box="[1199,1361,1058,1085]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="corinthiaca">
|
||
<emphasis id="B924EA94FFF2FFECD58D8827FAC1E63E" box="[1199,1361,1058,1085]" italics="true" pageId="10" pageNumber="183">C. corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
, is nonmonophyletic.
|
||
<taxonomicName id="4C504D05FFF2FFECD0138843FE6AE662" box="[305,506,1094,1121]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD0138843FE6AE662" box="[305,506,1094,1121]" italics="true" pageId="10" pageNumber="183">Centaurea achaia</emphasis>
|
||
</taxonomicName>
|
||
is the most widely distributed species of the group (
|
||
<figureCitation id="136B2A03FFF2FFECD56E8842FB00E662" box="[1100,1168,1094,1120]" captionStart="FIGURE 7" captionStartId="9.[136,229,2005,2027]" captionTargetBox="[147,1439,716,1979]" captionTargetId="figure-349@9.[145,1442,713,1981]" captionTargetPageId="9" captionText="FIGURE 7. Distribution map of the members of the Centaurea achaia group." figureDoi="http://doi.org/10.5281/zenodo.7645669" httpUri="https://zenodo.org/record/7645669/files/figure.png" pageId="10" pageNumber="183">Fig. 7</figureCitation>
|
||
) and exhibits the highest morphological variability (
|
||
<bibRefCitation id="EFC14B77FFF2FFECD09A886EFDD4E686" author="Routsi, E." box="[440,580,1130,1156]" pageId="10" pageNumber="183" refId="ref8730" refString="Routsi, E. (1993) Biosystematic study of the section Acrocentron (Cass.) DC. of the genus Centaurea L. in Greece. PhD Thesis, University of Patras, Greece." type="book" year="1993">Routsi 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC14B77FFF2FFECD36D886EFC23E687" author="Wagenitz, G. & Gamal-Eldin, E." box="[591,947,1130,1157]" pageId="10" pageNumber="183" pagination="95 - 127" refId="ref9164" refString="Wagenitz, G. & Gamal-Eldin, E. (1985) Zur Kenntnis der griechischen Centaurea - Arten der sektion Acrokentron. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 107: 95 - 127." type="journal article" year="1985">Wagenitz & Gamal-Eldin 1985</bibRefCitation>
|
||
). A more detailed morphological study of
|
||
<taxonomicName id="4C504D05FFF2FFECD4B0886FFF44E6AA" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD4B0886FFF44E6AA" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
across its distributional range and an enhanced sampling of different populations of the species in a phylogenetic framework could clarify the species limits of
|
||
<taxonomicName id="4C504D05FFF2FFECD3A188B7FD7FE6CE" box="[643,751,1202,1229]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="achaia">
|
||
<emphasis id="B924EA94FFF2FFECD3A188B7FD7FE6CE" box="[643,751,1202,1229]" italics="true" pageId="10" pageNumber="183">C. achaia</emphasis>
|
||
</taxonomicName>
|
||
and further guide the decision of whether the individuals from
|
||
<collectingRegion id="4994F864FFF2FFECD1AA88D2FF5CE6F2" box="[136,204,1238,1264]" country="Greece" name="Attiki" pageId="10" pageNumber="183">Attica</collectingRegion>
|
||
should be included in
|
||
<taxonomicName id="4C504D05FFF2FFECD0E588D3FDF8E6F2" box="[455,616,1238,1265]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="species" species="corinthiaca">
|
||
<emphasis id="B924EA94FFF2FFECD0E588D3FDF8E6F2" box="[455,616,1238,1265]" italics="true" pageId="10" pageNumber="183">C. corinthiaca</emphasis>
|
||
</taxonomicName>
|
||
or they constitute a different taxon. Species delimitations and relationships in
|
||
<taxonomicName id="4C504D05FFF2FFECD18488FFFE8BE717" authorityName="Zografidis & Koutroumpa & Liveri & Dimopoulos" authorityYear="2023" box="[166,283,1275,1301]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="183" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B924EA94FFF2FFECD18488FFFE8BE717" box="[166,283,1275,1301]" italics="true" pageId="10" pageNumber="183">Centaurea</emphasis>
|
||
</taxonomicName>
|
||
are being addressed in an on-going integrative taxonomic study of the genus by the second author that employs morphometric and phylogenomic studies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |