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<document id="FD9E7317E7744C85D3061E61799843B8" ID-DOI="10.1111/j.1096-3642.2009.00526.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5443023" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630454731098" checkinUser="felipe" docAuthor="Liew, Thor-Seng, Schilthuizen, Menno &amp; Vermeulen, Jaap Jan" docDate="2009" docId="03F9878AFF96E81A0D088DD827E4FC4A" docLanguage="en" docName="j.1096-3642.2009.00526.x.pdf" docOrigin="Zoological Journal of the Linnean Society 157 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Everettia safriei Liew &amp; Schilthuizen &amp; Vermeulen 2009, SP. NOV." docType="treatment" docVersion="5" lastPageNumber="540" masterDocId="FFC0FFF2FF8FE8030C238C132365FFCA" masterDocTitle="Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo" masterLastPageNumber="550" masterPageNumber="515" pageNumber="540" updateTime="1699276879878" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="719458E82ECB23AADFFA18326757387C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="570ED3B32F783677F09F8F90CFB4679C">
<mods:title id="1DF0EFFB001969A0745DC5A51F94D1DC">Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo</mods:title>
</mods:titleInfo>
<mods:name id="EA4B89344935FBC1B4046AD7AA9B70AD" type="personal">
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<mods:roleTerm id="00F93B8CF1C8C145CE35DEF6D26057B8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EAD05784B91B6B3AECAAA897D8D91727">Liew, Thor-Seng</mods:namePart>
</mods:name>
<mods:name id="BC3496FF4C840C00E7B3E9285A6797EE" type="personal">
<mods:role id="19EF78FAFAF85BCB0571726D90AC203B">
<mods:roleTerm id="5722D1D7D0A1BD2941FECD78E9FACF5B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1FBE7FC47BC7BEDD4A4DE8B59B9504DA">Schilthuizen, Menno</mods:namePart>
</mods:name>
<mods:name id="6DA34D6F51BB8BBEFB7B8BA987D8C024" type="personal">
<mods:role id="60AF23227AB5773D011058209E8C27D9">
<mods:roleTerm id="982CCF965C11408B56BA9DB2B1CD4767">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3A75C327306E397DA47329F9D63698B5">Vermeulen, Jaap Jan</mods:namePart>
</mods:name>
<mods:typeOfResource id="EDC2113A5D87399DA6298CCDC9501EDF">text</mods:typeOfResource>
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<mods:titleInfo id="3D189F785468FA31BD202A1B1E207E2D">
<mods:title id="C1ABA7B6A1291C3B07E5C2903E004460">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="0DD94BEEFCFACC11075977DC1FEAF0E1">
<mods:date id="AAC367B8E0222C2E2C98854DADBE8EEA">2009</mods:date>
<mods:detail id="BB09A972B6C1B769A7C6903C7FEFF631" type="pubDate">
<mods:number id="BD10C3EDC6AD75D22D28D34FD6FBBB74">2009-10-26</mods:number>
</mods:detail>
<mods:detail id="3795520E9A9D560A3FD216AF4BAF3F65" type="volume">
<mods:number id="43B74B954CAF30BCA755EBD2DF60D24F">157</mods:number>
</mods:detail>
<mods:detail id="4E05F35A7A99472D3FFE00EB499BA1F8" type="issue">
<mods:number id="56501898E61C80FEE79F252FDCED8E82">3</mods:number>
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<mods:start id="2522FBBB8857A8EFB9DC35954A9120D6">515</mods:start>
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<mods:location id="23165FAAFFD47FEACDAF4BC70D43131F">
<mods:url id="0D2D20F1E36191A9C0EAA5325DA08FFE">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x</mods:url>
</mods:location>
<mods:classification id="BE3E3FD42EF952FE6FEE24FC32A0C0AA">journal article</mods:classification>
<mods:identifier id="853C0A43310300BCDC730A2F682ED094" type="DOI">10.1111/j.1096-3642.2009.00526.x</mods:identifier>
<mods:identifier id="980CD01B0F49949699DF8CC6D20A18EE" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="ECA26662E28D578F8A6D42EF82388888" type="Zenodo-Dep">5443023</mods:identifier>
</mods:mods>
<treatment id="03F9878AFF96E81A0D088DD827E4FC4A" ID-DOI="http://doi.org/10.5281/zenodo.5492199" ID-Zenodo-Dep="5492199" LSID="urn:lsid:plazi:treatment:03F9878AFF96E81A0D088DD827E4FC4A" httpUri="http://treatment.plazi.org/id/03F9878AFF96E81A0D088DD827E4FC4A" lastPageNumber="540" pageId="25" pageNumber="540">
<subSubSection id="C34A6517FF96E81A0D088DD8218BFD75" pageId="25" pageNumber="540" type="nomenclature">
<paragraph id="8BEF369CFF96E81A0D088DD821E6FE2A" blockId="25.[299,643,459,480]" box="[299,643,459,480]" pageId="25" pageNumber="540">
<heading id="D0A781F0FF96E81A0D088DD821E6FE2A" box="[299,643,459,480]" centered="true" fontSize="9" level="2" pageId="25" pageNumber="540" reason="2">
<taxonomicName id="4C504D1FFF96E81A0D088DD82171FE2A" ID-CoL="3DHB3" authority="Liew &amp; Schilthuizen &amp; Vermeulen, 2009" authorityName="Liew &amp; Schilthuizen &amp; Vermeulen" authorityYear="2009" box="[299,532,459,480]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="540" phylum="Mollusca" rank="species" species="safriei" status="sp. nov.">EVERETTIA SAFRIEI</taxonomicName>
<emphasis id="B924EA8EFF96E81A0E3D8DDD21E6FE2A" bold="true" box="[542,643,459,480]" pageId="25" pageNumber="540">
<taxonomicNameLabel id="A21757F5FF96E81A0E3D8DDD21E6FE2A" box="[542,643,459,480]" pageId="25" pageNumber="540" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF369CFF96E81A0C878DE1218BFD75" blockId="25.[164,779,498,703]" pageId="25" pageNumber="540">
<emphasis id="B924EA8EFF96E81A0C878DE1238FFDCD" box="[164,234,498,519]" italics="true" pageId="25" pageNumber="540">Types:</emphasis>
<materialsCitation id="3B383CC1FF96E81A0CD58DE1218FFD75" collectingDate="2005-04-24" collectionCode="T" collectorName="S. Liew &amp; J. Lapidin &amp; Safrie &amp; Jasilin." country="Malaysia" county="State" elevation="2300" latitude="6.05" location="Kotal's" longLatPrecision="1305" longitude="116.6" municipality="Ranau District" pageId="25" pageNumber="540" specimenCode="BMNH 20080631" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<collectingCountry id="F347760CFF96E81A0CD58DE1221AFDCD" box="[246,383,498,519]" name="Malaysia" pageId="25" pageNumber="540">MALAYSIA</collectingCountry>
:
<collectingCounty id="628E4E10FF96E81A0DAC8DE122A9FDCD" box="[399,460,498,519]" pageId="25" pageNumber="540">State</collectingCounty>
of
<collectingRegion id="4994F87EFF96E81A0DD98DE12120FDC2" box="[506,581,498,520]" country="Malaysia" name="Sabah" pageId="25" pageNumber="540">Sabah</collectingRegion>
:
<collectingMunicipality id="6B8BACE6FF96E81A0E768DE1206DFDC2" box="[597,776,498,520]" pageId="25" pageNumber="540">Ranau District</collectingMunicipality>
. Southern part of Kinabalu Park. At
<quantity id="4CA89B79FF96E81A0E678E0321F2FDEC" box="[580,663,528,550]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="25" pageNumber="540" unit="m" value="2300.0">
<elevation id="007DD1AFFF96E81A0E678E0321F2FDEC" box="[580,663,528,550]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="25" pageNumber="540" unit="m" value="2300.0">2300 m</elevation>
</quantity>
alt. along
<location id="8E8F6047FF96E81A0C878E3C2396FD8F" LSID="urn:lsid:plazi:treatment:03F9878AFF96E81A0D088DD827E4FC4A:8E8F6047FF96E81A0C878E3C2396FD8F" box="[164,243,559,581]" country="Malaysia" county="State" latitude="6.05" longLatPrecision="1305" longitude="116.6" municipality="Ranau District" name="Kotal's" pageId="25" pageNumber="540" stateProvince="Sabah">Kotal’s</location>
route from
<location id="8E8F6047FF96E81A0D5C8E3C229BFD8F" LSID="urn:lsid:plazi:treatment:03F9878AFF96E81A0D088DD827E4FC4A:8E8F6047FF96E81A0D5C8E3C229BFD8F" box="[383,510,559,581]" country="Malaysia" county="State" latitude="6.05" longLatPrecision="1305" longitude="116.6" municipality="Ranau District" name="Bukit Babi" pageId="25" pageNumber="540" stateProvince="Sabah">Bukit Babi</location>
to the eastern ridge of
<location id="8E8F6047FF96E81A0C878E5D2207FDA9" LSID="urn:lsid:plazi:treatment:03F9878AFF96E81A0D088DD827E4FC4A:8E8F6047FF96E81A0C878E5D2207FDA9" box="[164,354,589,611]" country="Malaysia" county="State" latitude="6.05" longLatPrecision="1305" longitude="116.6" municipality="Ranau District" name="Mount Kinabalu" pageId="25" pageNumber="540" stateProvince="Sabah">Mount Kinabalu</location>
(
<geoCoordinate id="EE64505BFF96E81A0D578E5E22D0FDA9" box="[372,437,588,611]" degrees="6" direction="north" minutes="3" orientation="latitude" pageId="25" pageNumber="540" precision="925" value="6.05">
6°3
<emphasis id="B924EA8EFF96E81A0DBA8E5F22FAFDA9" box="[409,415,588,611]" italics="true" pageId="25" pageNumber="540">′</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE64505BFF96E81A0DE08E5E214FFDA9" box="[451,554,588,611]" degrees="116" direction="east" minutes="36" orientation="longitude" pageId="25" pageNumber="540" precision="925" value="116.6">
116°36
<emphasis id="B924EA8EFF96E81A0E338E5F2173FDA9" box="[528,534,588,611]" italics="true" pageId="25" pageNumber="540">′</emphasis>
E
</geoCoordinate>
). Collected by T.-
<collectorName id="26A5534AFF96E81A0ED48E5D23B9FD48" pageId="25" pageNumber="540">S. Liew</collectorName>
,
<collectorName id="26A5534AFF96E81A0CCA8E7F2207FD48" box="[233,354,620,642]" pageId="25" pageNumber="540">J. Lapidin</collectorName>
,
<collectorName id="26A5534AFF96E81A0D4D8E7F22D0FD48" box="[366,437,620,642]" pageId="25" pageNumber="540">Safrie</collectorName>
, and
<collectorName id="26A5534AFF96E81A0DD68E7F212CFD48" box="[501,585,620,642]" pageId="25" pageNumber="540">Jasilin.</collectorName>
<date id="FFEE105CFF96E81A0E718E7F21A2FD48" box="[594,711,620,642]" pageId="25" pageNumber="540" value="2005-04-24">
<collectingDate id="EFAAE9B4FF96E81A0E718E7F21A2FD48" box="[594,711,620,642]" pageId="25" pageNumber="540" value="2005-04-24">24.iv.2005</collectingDate>
</date>
,
<typeStatus id="54EB883EFF96E81A0EF08E7F23BDFD6A" pageId="25" pageNumber="540" type="holotype">holotype</typeStatus>
, BOR/MOL 5307,
<figureCitation id="136B2A19FF96E81A0DE48E98212FFD6A" box="[455,586,651,673]" captionStart="Figure 12" captionStartId="24.[144,223,696,715]" captionTargetBox="[146,1423,199,666]" captionTargetId="figure-241@24.[144,1425,196,666]" captionTargetPageId="24" captionText="Figure 12. Shells. A, Everettia jasilini sp. nov.; B, Everettia monticola sp. nov.; C, Everettia dominiki sp. nov.; D, Everettia safriei sp. nov.; E, Everettia planispira sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443047" httpUri="https://zenodo.org/record/5443047/files/figure.png" pageId="25" pageNumber="540">Figure 12D</figureCitation>
;
<specimenCount id="9D56FD15FF96E81A0E438E982062FD6A" box="[608,775,651,673]" pageId="25" pageNumber="540" type="generic" typeStatus="paratypes">six paratypes</specimenCount>
, three in
<specimenCode id="DBF69EE7FF96E81A0D298EB922B6FD75" box="[266,467,681,703]" collectionCode="BMNH" pageId="25" pageNumber="540">BMNH 20080631</specimenCode>
and three in SP 13063
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6517FF96E81A0C878EFB27E4FC4A" pageId="25" pageNumber="540" type="etymology">
<paragraph id="8BEF369CFF96E81A0C878EFB218FFC90" blockId="25.[164,779,744,858]" pageId="25" pageNumber="540">
<emphasis id="B924EA8EFF96E81A0C878EFB2247FD37" box="[164,290,744,765]" italics="true" pageId="25" pageNumber="540">Etymology:</emphasis>
This species is named after Safrie Hatimin, staff member at Kinabalu Park, who dedicatedly assisted the first author during the intensive malacological survey on Mount Kinabalu in 2005.
</paragraph>
<paragraph id="8BEF369CFF96E81A0C878F91215EFA8F" blockId="25.[164,779,898,1349]" pageId="25" pageNumber="540">
<emphasis id="B924EA8EFF96E81A0C878F9122E7FC5D" box="[164,386,898,919]" italics="true" pageId="25" pageNumber="540">Material examined:</emphasis>
<collectingRegion id="4994F87EFF96E81A0DA98F902286FC52" box="[394,483,899,920]" country="Malaysia" name="Sabah" pageId="25" pageNumber="540">SABAH</collectingRegion>
– Mount Kinabalu: BOR/ MOL 2671, southern slope,
<quantity id="4CA89B79FF96E81A0DC08FB22153FC7D" box="[483,566,929,951]" metricMagnitude="3" metricUnit="m" metricValue="3.3299999999999996" pageId="25" pageNumber="540" unit="m" value="3330.0">3330 m</quantity>
alt.,
<date id="FFEE105CFF96E81A0E508FB221A5FC7C" box="[627,704,929,950]" pageId="25" pageNumber="540" value="2003-10">x.2003</date>
; BOR/ MOL 2672, southern slope,
<quantity id="4CA89B79FF96E81A0E328FD32100FC1F" box="[529,613,960,981]" metricMagnitude="3" metricUnit="m" metricValue="3.3299999999999996" pageId="25" pageNumber="540" unit="m" value="3330.0">3330 m</quantity>
alt.,
<date id="FFEE105CFF96E81A0E9A8FD32063FC1F" box="[697,774,960,981]" pageId="25" pageNumber="540" value="2003-10">x.2003</date>
; BOR/MOL 2682, southern slope,
<quantity id="4CA89B79FF96E81A0E058FCD211CFC3E" box="[550,633,990,1012]" metricMagnitude="3" metricUnit="m" metricValue="3.08" pageId="25" pageNumber="540" unit="m" value="3080.0">3080 m</quantity>
alt.,
<date id="FFEE105CFF96E81A0E9A8FCD2063FC39" box="[697,774,990,1011]" pageId="25" pageNumber="540" value="2003-10">x.2003</date>
; BOR/MOL 2683, southern slope,
<quantity id="4CA89B79FF96E81A0E4C8FEE21A7FBD8" box="[623,706,1021,1042]" metricMagnitude="3" metricUnit="m" metricValue="3.08" pageId="25" pageNumber="540" unit="m" value="3080.0">3080 m</quantity>
alt.,
<date id="FFEE105CFF96E81A0C87880F2394FBFB" box="[164,241,1052,1073]" pageId="25" pageNumber="540" value="2003-10">x.2003</date>
; BOR/MOL 2687, southern slope,
<quantity id="4CA89B79FF96E81A0EA2880F21B1FBFB" box="[641,724,1052,1073]" metricMagnitude="3" metricUnit="m" metricValue="3.08" pageId="25" pageNumber="540" unit="m" value="3080.0">3080 m</quantity>
alt.,
<date id="FFEE105CFF96E81A0C8788292394FB85" box="[164,241,1082,1103]" pageId="25" pageNumber="540" value="2003-10">x.2003</date>
; BOR/MOL 5302, southern slope,
<quantity id="4CA89B79FF96E81A0E948829206EFB9A" box="[695,779,1082,1104]" metricMagnitude="3" metricUnit="m" metricValue="3.024" pageId="25" pageNumber="540" unit="m" value="3024.0">3024 m</quantity>
alt.,
<date id="FFEE105CFF96E81A0CFA884A224BFBA5" box="[217,302,1113,1135]" pageId="25" pageNumber="540" value="2005-09">ix.2005</date>
; BOR/MOL 5306, southern slope,
<quantity id="4CA89B79FF96E81A0E94884A206EFBA4" box="[695,779,1113,1134]" metricMagnitude="3" metricUnit="m" metricValue="3.0889999999999995" pageId="25" pageNumber="540" unit="m" value="3089.0">3089 m</quantity>
alt.,
<date id="FFEE105CFF96E81A0CCE8864222FFB47" box="[237,330,1143,1165]" pageId="25" pageNumber="540" value="2004-12">xii.2004</date>
; BOR/MOL 5303, eastern slope,
<quantity id="4CA89B79FF96E81A0C8788852392FB61" box="[164,247,1174,1196]" metricMagnitude="3" metricUnit="m" metricValue="3.264" pageId="25" pageNumber="540" unit="m" value="3264.0">3264 m</quantity>
alt.,
<date id="FFEE105CFF96E81A0D19888522EAFB66" box="[314,399,1174,1196]" pageId="25" pageNumber="540" value="2005-06">vi.2005</date>
; BOR/MOL 5304, eastern slope,
<quantity id="4CA89B79FF96E81A0C8788A72392FB00" box="[164,247,1204,1226]" metricMagnitude="3" metricUnit="m" metricValue="3.3519999999999994" pageId="25" pageNumber="540" unit="m" value="3352.0">3352 m</quantity>
alt.,
<date id="FFEE105CFF96E81A0D7B88A722C8FB00" box="[344,429,1204,1226]" pageId="25" pageNumber="540" value="2005-06">vi.2005</date>
; BOR/MOL 5305, eastern slope,
<quantity id="4CA89B79FF96E81A0CCC88C02226FB23" box="[239,323,1235,1257]" metricMagnitude="3" metricUnit="m" metricValue="3.432" pageId="25" pageNumber="540" unit="m" value="3432.0">3432 m</quantity>
alt.,
<date id="FFEE105CFF96E81A0DA588C022BFFB23" box="[390,474,1235,1257]" pageId="25" pageNumber="540" value="2005-06">vi.2005</date>
; BOR/MOL 5308, eastern slope,
<quantity id="4CA89B79FF96E81A0D2088E12233FACD" box="[259,342,1266,1288]" metricMagnitude="3" metricUnit="m" metricValue="3.416" pageId="25" pageNumber="540" unit="m" value="3416.0">3416 m</quantity>
alt.,
<date id="FFEE105CFF96E81A0D9C88E22171FACD" box="[447,532,1265,1287]" pageId="25" pageNumber="540" value="2005-06">vi.2005</date>
; BOR/MOL 5309, eastern slope,
<quantity id="4CA89B79FF96E81A0D72890322C1FAEC" box="[337,420,1296,1318]" metricMagnitude="3" metricUnit="m" metricValue="2.924" pageId="25" pageNumber="540" unit="m" value="2924.0">2924 m</quantity>
alt.,
<date id="FFEE105CFF96E81A0DCB8903215FFAEC" box="[488,570,1296,1318]" pageId="25" pageNumber="540" value="2005-04">iv.2005</date>
; BOR/MOL 5310, eastern slope,
<quantity id="4CA89B79FF96E81A0D6E893C22C4FA8E" box="[333,417,1327,1349]" metricMagnitude="3" metricUnit="m" metricValue="3.336" pageId="25" pageNumber="540" unit="m" value="3336.0">3336 m</quantity>
alt.,
<date id="FFEE105CFF96E81A0DC2893C2152FA8F" box="[481,567,1327,1349]" pageId="25" pageNumber="540" value="2005-06">vi.2005</date>
.
</paragraph>
<paragraph id="8BEF369CFF96E81A0C87897D22F2FA34" blockId="25.[164,779,1390,1534]" pageId="25" pageNumber="540">
<emphasis id="B924EA8EFF96E81A0C87897D22B1FA49" box="[164,468,1390,1411]" italics="true" pageId="25" pageNumber="540">Diagnostic characteristics:</emphasis>
The numerous, irregular, very fine, and discontinuous spiral threads on the upper surface of the shell differentiate this species from individuals of
<taxonomicName id="4C504D1FFF96E81A0DB089D92108FA14" authorityName="Liew &amp; Schilthuizen &amp; Vermeulen" authorityYear="2009" box="[403,621,1481,1503]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="540" phylum="Mollusca" rank="species" species="dominiki">
<emphasis id="B924EA8EFF96E81A0DB089D92108FA14" box="[403,621,1481,1503]" italics="true" pageId="25" pageNumber="540">Everettia dominiki</emphasis>
</taxonomicName>
with similar shell shape and size.
</paragraph>
<paragraph id="8BEF369CFF96E81A0C878A3420CBFD81" blockId="25.[164,779,1575,1903]" lastBlockId="25.[826,1441,198,587]" pageId="25" pageNumber="540">
<emphasis id="B924EA8EFF96E81A0C878A342249F9F6" box="[164,300,1575,1596]" italics="true" pageId="25" pageNumber="540">Description:</emphasis>
Shell (
<figureCitation id="136B2A19FF96E81A0D5D8A342283F9F6" box="[382,486,1575,1597]" captionStart="Figure 10" captionStartId="17.[164,243,1552,1571]" captionTargetBox="[325,1283,195,1522]" captionTargetId="figure-92@17.[323,1283,195,1522]" captionTargetPageId="17" captionText="Figure 10. Shell surface sculptures (scanning electron micrographs). A, upper surface of Everettia jasilini sp. nov.; B, underside of E. jasilini sp. nov.; C, upper surface of Everettia safriei sp. nov.; D; underside of Everettia paulbasintali sp. nov.; E, upper surface of E. paulbasintali sp. nov.; F, underside of Everettia klemmantanica; G, upper surface on the fourth whorl of E. klemmantanica; H, upper surface on the second whorl of E. klemmantanica. Scale bar = 100 Mm." figureDoi="http://doi.org/10.5281/zenodo.5443043" httpUri="https://zenodo.org/record/5443043/files/figure.png" pageId="25" pageNumber="540">Figs 10C</figureCitation>
,
<figureCitation id="136B2A19FF96E81A0DD18A342141F9F7" box="[498,548,1575,1597]" captionStart="Figure 12" captionStartId="24.[144,223,696,715]" captionTargetBox="[146,1423,199,666]" captionTargetId="figure-241@24.[144,1425,196,666]" captionTargetPageId="24" captionText="Figure 12. Shells. A, Everettia jasilini sp. nov.; B, Everettia monticola sp. nov.; C, Everettia dominiki sp. nov.; D, Everettia safriei sp. nov.; E, Everettia planispira sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443047" httpUri="https://zenodo.org/record/5443047/files/figure.png" pageId="25" pageNumber="540">12D</figureCitation>
): small, rather thin, greenish brown, moderately elevated, outer whorls rounded. Periphery rounded; spire almost flat. Above the periphery, protoconch smooth (sometimes with inconspicuous, densely placed spiral threads); shell with distinct oblique wrinkling; there are very fine, irregular and discontinuous spiral threads above the wrinkling. Below the periphery, shell is almost smooth. Height up to
<quantity id="4CA89B79FF96E81A0D8C8B0F2163F8FB" box="[431,518,1820,1841]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="25" pageNumber="540" unit="mm" value="4.8">4.8 mm</quantity>
; width up to
<quantity id="4CA89B79FF96E81A0E8C8B0F2063F8FB" box="[687,774,1820,1841]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="25" pageNumber="540" unit="mm" value="8.3">8.3 mm</quantity>
; diameter of the first three whorls 0.75–0.8, 0.55–0.60, and 0.95–1.00 mm, respectively; number of whorls up to five and one-quarter; height aperture up to
<quantity id="4CA89B79FF96E81A09668CD526F9FF11" box="[1349,1436,198,220]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="25" pageNumber="540" unit="mm" value="3.6">3.6 mm</quantity>
; width aperture up to
<quantity id="4CA89B79FF96E81A086B8CF727C5FF30" box="[1096,1184,228,250]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="25" pageNumber="540" unit="mm" value="4.5">4.5 mm</quantity>
. Genitalia (
<figureCitation id="136B2A19FF96E81A09118CF726F1FF31" box="[1330,1428,228,251]" captionStart="Figure 13" captionStartId="24.[144,223,1326,1345]" captionTargetBox="[148,1422,795,1294]" captionTargetId="figure-279@24.[144,1424,792,1296]" captionTargetPageId="24" captionText="Figure 13. Genitalia. A, Everettia jasilini sp. nov.; B, Everettia dominiki sp. nov.; C, Everettia safriei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5443049" httpUri="https://zenodo.org/record/5443049/files/figure.png" pageId="25" pageNumber="540">Fig. 13C</figureCitation>
): maximum length from genital opening to the end of dart-sac (before the visible gland tubules) up to
<quantity id="4CA89B79FF96E81A0F198D5320F7FE9C" box="[826,914,320,342]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="25" pageNumber="540" unit="mm" value="4.5">4.5 mm</quantity>
. The penis, dart-sac, and vagina with very thin transparent walls. P, V, GO, BC, and DS arranged around the short genital atrium, with limited space between the openings of each of the genital structures. BC short, about half of total DS length. Animal (
<figureCitation id="136B2A19FF96E81A0FDB8DCA272BFE25" box="[1016,1102,473,495]" captionStart="Figure 8" captionStartId="14.[144,223,1760,1779]" captionTargetBox="[147,1419,200,1728]" captionTargetId="figure-0@14.[144,1424,195,1730]" captionTargetPageId="14" captionText="Figure 8. Animal and mantle pigmentation. A, Everettia subconsul west coast lowland; B, Everettia layanglayang sp. nov.; C, Everettia monticola sp. nov.; D, Everettia safriei sp. nov.; E, Everettia interior sp. nov.; F, Everettia subconsul east coast lowland; G, Everettia corrugata corrugata; H, Everettia jucundior sp. nov.; I, Everettia klemmantanica; J, Everettia lapidini sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5443039" httpUri="https://zenodo.org/record/5443039/files/figure.png" pageId="25" pageNumber="540">Fig. 8D</figureCitation>
): the head of animal head is uniformly black. The mantle is brownish and in the last half whorl has sparse irregular black and bright markings.
</paragraph>
<paragraph id="8BEF369CFF96E81A0F198E6020C1FD2F" blockId="25.[826,1441,627,741]" pageId="25" pageNumber="540">
<emphasis id="B924EA8EFF96E81A0F198E60270FFD42" box="[826,1130,627,648]" italics="true" pageId="25" pageNumber="540">Distribution and habitat:</emphasis>
Upper montane primary and coniferous forest,
<quantity id="4CA89B79FF96E81A08168E8127A8FD62" box="[1077,1229,658,680]" metricMagnitude="3" metricUnit="m" metricValue="3.1" metricValueMax="3.5" metricValueMin="2.7" pageId="25" pageNumber="540" unit="m" value="3100.0" valueMax="3500.0" valueMin="2700.0">2700–3500 m</quantity>
alt.
<collectingRegion id="4994F87EFF96E81A09228E812629FD62" box="[1281,1356,658,680]" country="Malaysia" name="Sabah" pageId="25" pageNumber="540">Sabah</collectingRegion>
: Mount Kinabalu (eastern and southern ridges only), endemic (
<figureCitation id="136B2A19FF96E81A0F618EDC20FCFD2F" box="[834,921,719,741]" captionStart="Figure 9" captionStartId="15.[164,243,1622,1641]" captionTargetBox="[325,1281,196,1590]" captionTargetId="figure-58@15.[323,1283,194,1592]" captionTargetPageId="15" captionText="Figure 9. Distribution maps (Mount Kinabalu and Mount Tambuyukon). A, Everettia monticola sp. nov. and Everettia safriei sp. nov.; B, Everettia planispira sp. nov., Everettia jasilini sp. nov., and Everettia corrugata corrugata; C, Everettia dominiki sp. nov.; D, Everettia themis, Everettia subconsul, Everettia corrugata williamsi ssp. nov. and Everettia layanglayang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5443041" httpUri="https://zenodo.org/record/5443041/files/figure.png" pageId="25" pageNumber="540">Fig. 9A</figureCitation>
).
</paragraph>
<paragraph id="8BEF369CFF96E81A0F198F1D27E4FC4A" blockId="25.[826,1442,782,896]" pageId="25" pageNumber="540">
<emphasis id="B924EA8EFF96E81A0F198F1D20C3FCE9" box="[826,934,782,803]" italics="true" pageId="25" pageNumber="540">Remarks:</emphasis>
This species occurs sympatrically with
<taxonomicName id="4C504D1FFF96E81A09AB8F1C20E4FC8B" authorityName="Liew &amp; Schilthuizen &amp; Vermeulen" authorityYear="2009" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="540" phylum="Mollusca" rank="species" species="safriei">
<emphasis id="B924EA8EFF96E81A09AB8F1C20E4FC8B" italics="true" pageId="25" pageNumber="540">E. safriei</emphasis>
</taxonomicName>
, to which it is also phylogenetically close. However, both species maintain their distinct and unique shell characteristics.
</paragraph>
</subSubSection>
</treatment>
</document>
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