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<document id="A5B2CD04EE44A61600E7A035724CDB5B" ID-CLB-Dataset="6279" ID-DOI="10.5852/ejt.2021.751.1381" ID-GBIF-Dataset="4e3af683-d8a6-40de-991b-c4584d66b8cc" ID-ISSN="2118-9773" ID-Zenodo-Dep="4906007" ID-ZooBank="1D27EA41-2396-463A-B1B2-0CCF79C9F3D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623068023717" checkinUser="felipe" docAuthor="Zamani, Alireza, Chatzaki, Maria, Esyunin, Sergei L. &amp; Marusik, Yuri M." docDate="2021" docId="03F987850367EF7B1C507919FCFF6E07" docLanguage="en" docName="ejt-751_68-114.pdf" docOrigin="European Journal of Taxonomy 751 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.751.1381" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Marjanus isfahanicus Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="9CB7FC88-C04C-4670-A98D-7A3ACC436976" docUuidSource="ZooBank" docVersion="8" lastPageNumber="94" masterDocId="FFC0FFFD037FEF611E347F73FFC16D41" masterDocTitle="One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations" masterLastPageNumber="114" masterPageNumber="68" pageNumber="92" updateTime="1699000119853" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="04C19E16EBD46D47E55937B5C43123D5">One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations</mods:title>
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<mods:namePart id="1D5E74FBF09C4DBA7319CAB940C94531">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="DCD7D443EF2B576195A6CE74A557E1CA">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland. Department of Molecular Biology and Genetics, Democritus University of Thrace, Dragana, 68100 Alexandroupolis, Greece. Perm State University, Bukireva Street 15, Perm, 614600, Russia. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
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<mods:namePart id="0BFBB4592714A02CF492A30B3E82F3FC">Chatzaki, Maria</mods:namePart>
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<mods:namePart id="B0E8E5873E45BCF8E2A621F73ACB909D">Esyunin, Sergei L.</mods:namePart>
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<mods:namePart id="8ADAE281791890F9054D83A76E75CD47">Marusik, Yuri M.</mods:namePart>
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<treatment id="03F987850367EF7B1C507919FCFF6E07" ID-DOI="http://doi.org/10.5281/zenodo.4906483" ID-GBIF-Taxon="182597847" ID-Zenodo-Dep="4906483" LSID="urn:lsid:zoobank.org:act:9CB7FC88-C04C-4670-A98D-7A3ACC436976" httpUri="http://treatment.plazi.org/id/03F987850367EF7B1C507919FCFF6E07" lastPageId="26" lastPageNumber="94" pageId="24" pageNumber="92">
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<taxonomicName id="4C504D100367EF791C507919FCAF6BC5" ID-CoL="98YVP" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[612,878,1641,1668]" class="Arachnida" family="Gnaphosidae" genus="Marjanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="92" phylum="Arthropoda" rank="species" species="isfahanicus" status="sp. nov.">
<emphasis id="B924EA810367EF791C507919FCAF6BC5" bold="true" box="[612,878,1641,1668]" italics="true" pageId="24" pageNumber="92">Marjanus isfahanicus</emphasis>
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<taxonomicNameLabel id="A21757FA0367EF791D427918FC116BC4" box="[886,976,1643,1669]" pageId="24" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<figureCitation id="136B2A160367EF791CC479CBFC826B93" box="[752,835,1720,1746]" captionStart="Fig" captionStartId="25.[191,234,1964,1990]" captionTargetBox="[189,1398,1028,1918]" captionTargetId="figure-221@25.[189,1398,1028,1925]" captionTargetPageId="25" captionText="Fig. 16. Marjanus isfahanicus sp. nov., holotype, ♀ (MHNG). A. Habitus, dorsal view. B, D, F. Epigyne, ventral view. C, E, G. Epigyne, dorsal view. Scale bars: A = 0.5 mm; B = 0.2 mm; FG = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4906055" httpUri="https://zenodo.org/record/4906055/files/figure.png" pageId="24" pageNumber="92">Fig. 16</figureCitation>
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<paragraph id="8BEF36930367EF791E89798EFEF76A56" blockId="24.[189,310,1789,1815]" box="[189,310,1789,1815]" pageId="24" pageNumber="92">
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<emphasis id="B924EA810367EF791E89798EFEF76A56" bold="true" box="[189,310,1789,1815]" pageId="24" pageNumber="92">Diagnosis</emphasis>
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<paragraph id="8BEF36930367EF791E897843FACB6A33" blockId="24.[189,1398,1840,1906]" pageId="24" pageNumber="92">The new species differs from the generotype by having interrupted anterior margins (vs uninterrupted), a strongly rebordered W-shaped atrium (vs inconspicuous) and round receptacles (vs tubular).</paragraph>
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<subSubSection id="C34A65180367EF791E8978EFFAB96AA9" box="[189,1400,1948,2025]" pageId="24" pageNumber="92" type="etymology">
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<heading id="D0A781FF0367EF791E8978EFFE856AF7" bold="true" box="[189,324,1948,1974]" fontSize="11" level="3" pageId="24" pageNumber="92" reason="3">
<emphasis id="B924EA810367EF791E8978EFFE856AF7" bold="true" box="[189,324,1948,1974]" pageId="24" pageNumber="92">Etymology</emphasis>
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The specific epithet is an adjective and refers to
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,
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, the
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locality of the new species.
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<emphasis id="B924EA810366EF781E897E7AFEAD6C63" bold="true" box="[189,364,264,291]" pageId="25" pageNumber="93">Type material</emphasis>
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<materialsCitation id="3B383CCE0366EF781EDE7E4BFE986CFC" ID-GBIF-Occurrence="3127434323" collectingDate="2016-05-19" collectionCode="MHNG" collectorName="P. Ponel" country="Iran" county="Qamsar" elevation="2710" latitude="33.616665" location="Barzok Protected Area" longLatPrecision="1204" longitude="51.316666" municipality="Barzok Protected Area" pageId="25" pageNumber="93" specimenCount="1" specimenCount-female="1" stateProvince="Isfahan Province" typeStatus="holotype">
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<typeStatus id="54EB88310366EF781EDE7E4BFE9D6C13" box="[234,348,312,338]" pageId="25" pageNumber="93" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F34776030366EF781E897E28FEC96C34" box="[189,264,347,373]" name="Iran" pageId="25" pageNumber="93">IRAN</collectingCountry>
<specimenCount id="9D56FD1A0366EF781F117E2FFEFD6C37" box="[293,316,348,374]" pageId="25" pageNumber="93" type="female"></specimenCount>
;
<collectingRegion id="4994F8710366EF781F787E28FDD76C34" box="[332,534,347,373]" country="Iran" name="Esfahan" pageId="25" pageNumber="93">Isfahan Province</collectingRegion>
,
<collectingCounty id="628E4E1F0366EF781C107E28FDBE6C34" box="[548,639,347,373]" pageId="25" pageNumber="93">Qamsar</collectingCounty>
&amp;
<collectingMunicipality id="6B8BACE90366EF781C9C7E28FC766C34" box="[680,951,347,373]" pageId="25" pageNumber="93">Barzok Protected Area</collectingMunicipality>
,
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<quantity id="4CA89B760366EF781DF17E28FBD06C34" box="[965,1041,347,374]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="25" pageNumber="93" unit="km" value="55.0">55 km</quantity>
SW of Qamsar
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,
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NE of Kamoo
</locationDeviation>
,
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;
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,
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;
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a.s.l.
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;
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<collectingDate id="EFAAE9BB0366EF781A5B7E0CFACC6CD8" box="[1135,1293,383,410]" pageId="25" pageNumber="93" value="2016-05-19">19 May 2016</collectingDate>
</date>
;
<collectorName id="26A553450366EF781B2D7EF3FAB66CD8" box="[1305,1399,383,410]" pageId="25" pageNumber="93">P. Ponel</collectorName>
leg.;
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.
</materialsCitation>
</paragraph>
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<emphasis id="B924EA810366EF781E897E97FE8F6CBF" bold="true" box="[189,334,484,510]" pageId="25" pageNumber="93">Description</emphasis>
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<emphasis id="B924EA810366EF781EDE7D67FE846F6F" bold="true" box="[234,325,532,558]" pageId="25" pageNumber="93">Female</emphasis>
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Habitus as in
<figureCitation id="136B2A160366EF781F697D44FE096F10" box="[349,456,567,593]" captionStart="Fig" captionStartId="25.[191,234,1964,1990]" captionTargetBox="[189,1398,1028,1918]" captionTargetId="figure-221@25.[189,1398,1028,1925]" captionTargetPageId="25" captionText="Fig. 16. Marjanus isfahanicus sp. nov., holotype, ♀ (MHNG). A. Habitus, dorsal view. B, D, F. Epigyne, ventral view. C, E, G. Epigyne, dorsal view. Scale bars: A = 0.5 mm; B = 0.2 mm; FG = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4906055" httpUri="https://zenodo.org/record/4906055/files/figure.png" pageId="25" pageNumber="93">Fig. 16A</figureCitation>
. Total length 3.04. Carapace 1.40 long, 1.01 wide. Eye sizes and interdistance of PMEs: AME = 0.05, ALE = 0.07, PME = 0.06, PLE = 0.06, PMEPME = 0.05. Carapace and chelicerae dark brown; carapace darker at the margins, with a pattern of light patches. Chelicera with 3 promarginal teeth and 1 retromarginal tooth. Sternum, maxillae and labium light brown. Legs lighter than carapace, without annulations, tarsi lighter. Abdomen dark dorsally, light brown ventrally, without any markings. Spinnerets light gray and uniform in color. Measurements of legs: I: 3.27 (0.95, 0.63, 0.69, 0.54, 0.46), II: 3.14 (0.91, 0.52, 0.69, 0.52, 0.50), III: 2.59 (0.74, 0.37, 0.51, 0.47, 0.50), IV: 3.98 (1.08, 0.56, 0.83, 0.92, 0.59).
</paragraph>
<paragraph id="8BEF36930366EF781E897C07FAB76E94" blockId="25.[189,1398,884,982]" pageId="25" pageNumber="93">
Epigyne as in
<figureCitation id="136B2A160366EF781F5E7C07FE3A6ECF" box="[362,507,884,910]" captionStart="Fig" captionStartId="25.[191,234,1964,1990]" captionTargetBox="[189,1398,1028,1918]" captionTargetId="figure-221@25.[189,1398,1028,1925]" captionTargetPageId="25" captionText="Fig. 16. Marjanus isfahanicus sp. nov., holotype, ♀ (MHNG). A. Habitus, dorsal view. B, D, F. Epigyne, ventral view. C, E, G. Epigyne, dorsal view. Scale bars: A = 0.5 mm; B = 0.2 mm; FG = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4906055" httpUri="https://zenodo.org/record/4906055/files/figure.png" pageId="25" pageNumber="93">Fig. 16BG</figureCitation>
. Atrium with distinct and separate anterior hoods; lateral margins distinct, converging; endogyne with membranous bursae near copulatory openings; copulatory ducts thin, forming 2 screw-like, full lateral coils in mid part, straight posteriorly, with blind glandular heads, connected
</paragraph>
<caption id="DF2F661B0366EF781E8B78DFFAFF6AAB" ID-DOI="http://doi.org/10.5281/zenodo.4906055" ID-Zenodo-Dep="4906055" httpUri="https://zenodo.org/record/4906055/files/figure.png" pageId="25" pageNumber="93" startId="25.[191,234,1964,1990]" targetBox="[189,1398,1028,1918]" targetPageId="25">
<paragraph id="8BEF36930366EF781E8B78DFFAFF6AAB" blockId="25.[188,1398,1964,2026]" pageId="25" pageNumber="93">
<emphasis id="B924EA810366EF781E8B78DFFED96A87" bold="true" box="[191,280,1964,1990]" pageId="25" pageNumber="93">Fig. 16.</emphasis>
<taxonomicName id="4C504D100366EF781F2878DEFDD86A87" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[284,537,1964,1990]" class="Arachnida" family="Gnaphosidae" genus="Marjanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="93" phylum="Arthropoda" rank="species" species="isfahanicus" status="sp. nov.">
<emphasis id="B924EA810366EF781F2878DEFDD86A87" box="[284,537,1964,1990]" italics="true" pageId="25" pageNumber="93">Marjanus isfahanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA0366EF781C2B78DFFDBD6A87" box="[543,636,1964,1990]" pageId="25" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♀ (MHNG).
<emphasis id="B924EA810366EF781DBB78DFFC656A87" bold="true" box="[911,932,1964,1990]" pageId="25" pageNumber="93">A</emphasis>
. Habitus, dorsal view.
<emphasis id="B924EA810366EF781A9E78DFFB3E6A87" bold="true" box="[1194,1279,1964,1990]" pageId="25" pageNumber="93">B, D, F</emphasis>
. Epigyne, ventral view.
<emphasis id="B924EA810366EF781F6878BCFE786AA8" bold="true" box="[348,441,1999,2025]" pageId="25" pageNumber="93">C, E, G</emphasis>
. Epigyne, dorsal view. Scale bars: A = 0.5 mm; B = 0.2 mm; FG = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BEF36930365EF7B1E897E7AFC396C09" blockId="26.[189,1398,265,328]" pageId="26" pageNumber="94">to ventral additional bursae with extra glandular tubes; receptacles globular, 3 times as wide as ducts, touching each other, with fertilization ducts located at their upper part.</paragraph>
<paragraph id="8BEF36930365EF7B1EDE7E1CFEEB6CC8" blockId="26.[189,313,367,429]" box="[234,298,367,393]" pageId="26" pageNumber="94">
<emphasis id="B924EA810365EF7B1EDE7E1CFEEB6CC8" bold="true" box="[234,298,367,393]" pageId="26" pageNumber="94">Male</emphasis>
</paragraph>
<paragraph id="8BEF36930365EF7B1E897EE0FEF86CEC" blockId="26.[189,313,367,429]" box="[189,313,403,429]" pageId="26" pageNumber="94">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C34A65180365EF7B1E897EA6FAB66FF1" pageId="26" pageNumber="94" type="discussion">
<paragraph id="8BEF36930365EF7B1E897EA6FE856CAE" blockId="26.[189,324,469,495]" box="[189,324,469,495]" pageId="26" pageNumber="94">
<heading id="D0A781FF0365EF7B1E897EA6FE856CAE" bold="true" box="[189,324,469,495]" fontSize="11" level="3" pageId="26" pageNumber="94" reason="3">
<emphasis id="B924EA810365EF7B1E897EA6FE856CAE" bold="true" box="[189,324,469,495]" pageId="26" pageNumber="94">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930365EF7B1E897D76FAB66FF1" blockId="26.[188,1400,517,688]" pageId="26" pageNumber="94">
Based on the similarity of the epigynal conformation to that of the
<typeStatus id="54EB88310365EF7B1DF67D75FC346F61" box="[962,1013,518,544]" pageId="26" pageNumber="94">type</typeStatus>
species, we tentatively place this species here in the absence of male specimens. However, only two of the three diagnostic elements of the females of this genus are met in this new species: the long, coiled copulatory ducts and additional anterior copulatory bursae and the receptacles being round rather than tubular. The latter character may have to be omitted from the generic diagnosis when more species are described and included in this genus.
</paragraph>
</subSubSection>
<subSubSection id="C34A65180365EF7B1E897DA4FCFF6E07" pageId="26" pageNumber="94" type="distribution">
<paragraph id="8BEF36930365EF7B1E897DA4FE976FB0" blockId="26.[189,342,727,753]" box="[189,342,727,753]" pageId="26" pageNumber="94">
<heading id="D0A781FF0365EF7B1E897DA4FE976FB0" bold="true" box="[189,342,727,753]" fontSize="11" level="3" pageId="26" pageNumber="94" reason="3">
<emphasis id="B924EA810365EF7B1E897DA4FE976FB0" bold="true" box="[189,342,727,753]" pageId="26" pageNumber="94">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930365EF7B1E897C7BFCFF6E07" blockId="26.[189,1398,775,838]" pageId="26" pageNumber="94">
Known only from the
<typeStatus id="54EB88310365EF7B1FF47C7BFE326E63" box="[448,499,776,802]" pageId="26" pageNumber="94">type</typeStatus>
locality in
<collectingRegion id="4994F8710365EF7B1C427C74FCFE6E60" box="[630,831,775,801]" country="Iran" name="Esfahan" pageId="26" pageNumber="94">Isfahan Province</collectingRegion>
, central
<collectingCountry id="F34776030365EF7B1D957C7BFC156E63" box="[929,980,776,802]" name="Iran" pageId="26" pageNumber="94">Iran</collectingCountry>
. This is the first record of
<taxonomicName id="4C504D100365EF7B1B307C7BFAB76E60" authorityName="Chatzaki" authorityYear="2018" box="[1284,1398,776,801]" class="Arachnida" family="Gnaphosidae" genus="Marjanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B924EA810365EF7B1B307C7BFAB76E60" box="[1284,1398,776,801]" italics="true" pageId="26" pageNumber="94">Marjanus</emphasis>
</taxonomicName>
in
<collectingCountry id="F34776030365EF7B1EEF7C5FFECC6E07" box="[219,269,812,838]" name="Iran" pageId="26" pageNumber="94">Iran</collectingCountry>
, and the second species described in this genus.
</paragraph>
</subSubSection>
</treatment>
</document>