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<document id="8BCE5693797A5BD6523C53399EBA586D" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629706805705" checkinUser="felipe" docAuthor="Hara, Marcos Ryotaro &amp; Pinto-Da-Rocha, Ricardo" docDate="2010" docId="039387E595474676FF65FF3089F5FA95" docLanguage="en" docName="zt02698p136.pdf" docOrigin="Zootaxa 2698" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Eusarcus alpinus Hara &amp; Pinto-Da-Rocha 2010, sp. n." docType="treatment" docVersion="3" lastPageNumber="31" masterDocId="FFAAFF9D955B4668FFF2FFA78A56FF9F" masterDocTitle="Systematic review and cladistic analysis of the genus Eusarcus Perty 1833 (Arachnida, Opiliones, Gonyleptidae) 2698" masterLastPageNumber="136" masterPageNumber="1" pageNumber="29" updateTime="1699098874055" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="2F31012420CCB4DB0EFD6D25D9F21909" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8EEE8988BFFACF0BA9C4F3C1C8DA13A2">
<mods:title id="551573066AFD1E8D8081B6D904898F07">Systematic review and cladistic analysis of the genus Eusarcus Perty 1833 (Arachnida, Opiliones, Gonyleptidae) 2698</mods:title>
</mods:titleInfo>
<mods:name id="95C5AE8DB27E9F3D85DF0F781DDC5D0E" type="personal">
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<mods:namePart id="AB8485D4481DEBC28EA05A3273532510">Hara, Marcos Ryotaro</mods:namePart>
</mods:name>
<mods:name id="45B14720B370BA40A2C42AADF541F320" type="personal">
<mods:role id="3EBFC44A494586D99447670A9F197FCC">
<mods:roleTerm id="4B4E4B1D51D0A55C5C289188F9B34EF0">Author</mods:roleTerm>
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<mods:namePart id="211DDCE98CF77A839D92C01B4B18AC7D">Pinto-Da-Rocha, Ricardo</mods:namePart>
</mods:name>
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<mods:title id="240059D0513012E3359D456DE7C43EFE">Zootaxa</mods:title>
</mods:titleInfo>
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<mods:date id="DC9E6EBBFC90737376A2E446A43B12A8">2010</mods:date>
<mods:detail id="E910EE054F5D706DE193D27E95D86858" type="pubDate">
<mods:number id="7E712377738ABA9C031B070E9C58AA3E">2010-12-03</mods:number>
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<mods:detail id="E7972BD4C40E700ADCB33E320A3D8859" type="volume">
<mods:number id="7456637FD731278CBC1B1044E2783845">2698</mods:number>
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<mods:classification id="1D0ED2A1FAF6DFA71ECD8D28E28CF388">journal article</mods:classification>
<mods:identifier id="F37B606E9F6C3DCE88EC31F2818414E7" type="ISSN">1175­5334</mods:identifier>
</mods:mods>
<treatment id="039387E595474676FF65FF3089F5FA95" ID-DOI="http://doi.org/10.5281/zenodo.5295580" ID-Zenodo-Dep="5295580" LSID="urn:lsid:plazi:treatment:039387E595474676FF65FF3089F5FA95" httpUri="http://treatment.plazi.org/id/039387E595474676FF65FF3089F5FA95" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
<subSubSection id="C320657895474674FF65FF308BE5FF2E" box="[151,435,151,177]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8B8536F395474674FF65FF308BE5FF2E" blockId="28.[151,435,151,212]" box="[151,435,151,177]" pageId="28" pageNumber="29">
<heading id="D0CD819F95474674FF65FF308BE5FF2E" bold="true" box="[151,435,151,177]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<emphasis id="B94EEAE195474674FF65FF308BE5FF2E" bold="true" box="[151,435,151,177]" pageId="28" pageNumber="29">
<taxonomicName id="4C3A4D7095474674FF65FF308B3CFF2E" ID-CoL="7W5R3" authority="Hara &amp; Pinto-Da-Rocha, 2010" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[151,362,151,177]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="alpinus" status="sp. nov.">
<emphasis id="B94EEAE195474674FF65FF308B3CFF2E" bold="true" box="[151,362,151,177]" italics="true" pageId="28" pageNumber="29">Eusarcus alpinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D579A95474674FE83FF308BE5FF2E" box="[369,435,151,177]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320657895474674FF65FF1D8BC1FF4B" box="[151,407,186,212]" pageId="28" pageNumber="29" type="description">
<paragraph id="8B8536F395474674FF65FF1D8BC1FF4B" blockId="28.[151,435,151,212]" box="[151,407,186,212]" pageId="28" pageNumber="29">
(
<figureCitation id="13012A7695474674FF6DFF1D8AA4FF4B" box="[159,242,186,212]" captionStart="FIGURE 4" captionStartId="29.[151,255,1937,1961]" captionTargetBox="[203,1399,198,1908]" captionTargetId="figure-75@29.[197,1401,189,1918]" captionTargetPageId="29" captionText="FIGURE 4. Eusarcus alpinus sp. n. Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view(in part); D, right trochanter and femur IV, prolateral view; E, idem, dorsal view; F, idem, ventral view; G, right femur III, ventral view; H, right tibia III, ventral view. Scale bars, 1 mm." pageId="28" pageNumber="29">Figs. 4</figureCitation>
,
<figureCitation id="13012A7695474674FF0DFF1D8B19FF4B" box="[255,335,186,212]" captionStart="FIGURE 43" captionStartId="119.[151,255,1559,1583]" captionTargetBox="[157,1429,312,1530]" captionTargetId="figure-316@119.[151,1436,293,1538]" captionTargetPageId="119" captionText="FIGURE 43. Distal part of penis in dorsal and lateral view: A–B, Eusarcus alpinus sp. n.; C–D, E. didactylus sp. n.; E– F, E. nigrimaculatus. Scale bars, 0.1 mm." pageId="28" pageNumber="29">43A,B</figureCitation>
,
<figureCitation id="13012A7695474674FEA9FF1D8BD9FF4B" box="[347,399,186,212]" captionStart="FIGURE 49" captionStartId="125.[151,255,2004,2028]" captionTargetBox="[180,1389,206,1966]" captionTargetId="figure-26@125.[175,1399,194,1974]" captionTargetPageId="125" captionText="FIGURE 49. A, B: Localities of Eusarcus spp. in eastern Brazil." pageId="28" pageNumber="29">49A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C320657895474674FF65FEA5885AFE13" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8B8536F395474674FF65FEA5885AFE13" blockId="28.[151,1436,258,2002]" pageId="28" pageNumber="29">
<emphasis id="B94EEAE195474674FF65FEA58B0AFE83" bold="true" box="[151,348,258,284]" pageId="28" pageNumber="29">Type material:</emphasis>
<materialsCitation id="3B523CAE95474674FE97FEA58901FEDD" collectingDate="1998-03-23" collectingDateMax="1998-03-25" collectingDateMin="1998-03-23" collectorName="A. B. Kury &amp; M. Baptista &amp; A. P. L. Giupponi" country="Brazil" location="Santa Maria Madalena" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<collectingCountry id="F32D766395474674FE97FEA58B8EFE83" box="[357,472,258,284]" name="Brazil" pageId="28" pageNumber="29">BRAZIL</collectingCountry>
.
<collectingRegion id="49FEF81195474674FE15FEA588CAFE83" box="[487,668,258,284]" country="Brazil" name="Rio de Janeiro" pageId="28" pageNumber="29">Rio de Janeiro</collectingRegion>
:
<location id="8EE5602895474674FD5CFEA5899CFE83" LSID="urn:lsid:plazi:treatment:039387E595474676FF65FF3089F5FA95:8EE5602895474674FD5CFEA5899CFE83" box="[686,970,258,284]" country="Brazil" name="Santa Maria Madalena" pageId="28" pageNumber="29" stateProvince="Rio de Janeiro">Santa Maria Madalena</location>
(P.E. do Desengano),
<collectorName id="26CF532595474674FB15FEA58F3AFE83" box="[1255,1388,258,284]" pageId="28" pageNumber="29">A.B. Kury</collectorName>
,
<collectorName id="26CF532595474674FA88FEA58AACFEDD" pageId="28" pageNumber="29">M. Baptista</collectorName>
,
<collectorName id="26CF532595474674FEF4FE8F8B9EFEDD" box="[262,456,296,322]" pageId="28" pageNumber="29">A.P.L. Giupponi</collectorName>
leg.,
<date id="FF84103395474674FDF8FE8F88EEFEDD" box="[522,696,296,322]" pageId="28" pageNumber="29" value="1998-03-23" valueMax="1998-03-25" valueMin="1998-03-23">
<collectingDate id="EFC0E9DB95474674FDF8FE8F88EEFEDD" box="[522,696,296,322]" pageId="28" pageNumber="29" value="1998-03-23" valueMax="1998-03-25" valueMin="1998-03-23">23–25.iii.1998</collectingDate>
</date>
, ma
<typeStatus id="5481885195474674FD03FE8F8901FEDD" box="[753,855,296,322]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
</materialsCitation>
&amp;
<materialsCitation id="3B523CAE95474674FC8DFE8F885EFE13" collectingDate="1998-03-23" collectingDateMax="1998-03-25" collectingDateMin="1998-03-23" collectionCode="ZMUC" collectorName="Teresopolis &amp; E. A. Goeldi" country="Brazil" location="Santa Maria Madalena" pageId="28" pageNumber="29" specimenCode="MNRJ 5719, MZSP 28665" specimenCount="3" stateProvince="Rio de Janeiro" typeStatus="paratype">
2 fe
<typeStatus id="5481885195474674FC46FE8F8E72FEDD" box="[948,1060,296,322]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
(
<specimenCode id="DB9C9E8895474674FBC6FE8F8E9AFEDD" box="[1076,1228,296,322]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="28" pageNumber="29">MNRJ 5719</specimenCode>
); idem, 1 ma &amp; 1 fe
<typeStatus id="5481885195474674FF47FEEA8B73FEF8" box="[181,293,333,359]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
(
<specimenCode id="DB9C9E8895474674FEC7FEEA8B8FFEF8" box="[309,473,333,359]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="28" pageNumber="29" type="Museum">MZSP 28665</specimenCode>
);
<collectorName id="26CF532595474674FE1CFEEA8822FEF8" box="[494,628,333,359]" pageId="28" pageNumber="29">Teresópolis</collectorName>
(Colônia Alpina),
<collectorName id="26CF532595474674FCA1FEEA89B6FEF8" box="[851,992,333,359]" pageId="28" pageNumber="29">E.A. Goeldi</collectorName>
leg., without date, 3 ma, 3 beta male, 2 fe &amp; 2 im
<typeStatus id="5481885195474674FED7FED58BC3FE13" box="[293,405,370,396]" pageId="28" pageNumber="29" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED2BAE3695474674FE57FED58856FE13" box="[421,512,370,396]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="28" pageNumber="29">ZMUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C320657895474674FF34FE308F72FD28" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8B8536F395474674FF34FE308F72FD28" blockId="28.[151,1436,258,2002]" pageId="28" pageNumber="29">
<emphasis id="B94EEAE195474674FF34FE308B1EFE2E" bold="true" box="[198,328,407,433]" pageId="28" pageNumber="29">Diagnosis:</emphasis>
<taxonomicName id="4C3A4D7095474674FEA2FE3F8B91FE2E" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[336,455,408,434]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="B94EEAE195474674FEA2FE3F8B34FE2E" box="[336,354,408,433]" italics="true" pageId="28" pageNumber="29">E</emphasis>
.
<emphasis id="B94EEAE195474674FE83FE3F8B91FE2E" box="[369,455,408,433]" italics="true" pageId="28" pageNumber="29">alpinus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C3A4D7095474674FDB9FE3F88FCFE2E" box="[587,682,408,434]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="incus">
<emphasis id="B94EEAE195474674FDB9FE3F880BFE2E" box="[587,605,408,433]" italics="true" pageId="28" pageNumber="29">E</emphasis>
.
<emphasis id="B94EEAE195474674FD9EFE3F88FCFE2E" box="[620,682,408,433]" italics="true" pageId="28" pageNumber="29">incus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D7095474674FD4BFE3F8962FE2E" box="[697,820,408,434]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B94EEAE195474674FD4BFE3F889DFE2E" box="[697,715,408,433]" italics="true" pageId="28" pageNumber="29">E</emphasis>
.
<emphasis id="B94EEAE195474674FD2BFE3F8962FE2E" box="[729,820,408,433]" italics="true" pageId="28" pageNumber="29">pusillus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D7095474674FCB1FE3F8E43FE2E" box="[835,1045,408,434]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="nigrimaculatus">
<emphasis id="B94EEAE195474674FCB1FE3F8903FE2E" box="[835,853,408,433]" italics="true" pageId="28" pageNumber="29">E</emphasis>
.
<emphasis id="B94EEAE195474674FC96FE3F8E43FE2E" box="[868,1045,408,433]" italics="true" pageId="28" pageNumber="29">nigrimaculatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D7095474674FBD6FE3F8EA2FE2E" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[1060,1268,408,434]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="acrophthalmus">
<emphasis id="B94EEAE195474674FBD6FE3F8E60FE2E" box="[1060,1078,408,433]" italics="true" pageId="28" pageNumber="29">E</emphasis>
.
<emphasis id="B94EEAE195474674FBB7FE3F8EA2FE2E" box="[1093,1268,408,433]" italics="true" pageId="28" pageNumber="29">acrophthalmus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D7095474674FAF1FE3F8FCAFE2E" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[1283,1436,408,434]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="didactylus">
<emphasis id="B94EEAE195474674FAF1FE3F8F43FE2E" box="[1283,1301,408,433]" italics="true" pageId="28" pageNumber="29">E</emphasis>
.
<emphasis id="B94EEAE195474674FAD6FE3F8FCAFE2E" box="[1316,1436,408,433]" italics="true" pageId="28" pageNumber="29">didactylus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D7095474674FF3BFE1A8B2CFE49" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[201,378,445,471]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="geometricus">
<emphasis id="B94EEAE195474674FF3BFE1A8A8DFE49" box="[201,219,445,470]" italics="true" pageId="28" pageNumber="29">E</emphasis>
.
<emphasis id="B94EEAE195474674FF19FE1A8B2CFE49" box="[235,378,445,470]" italics="true" pageId="28" pageNumber="29">geometricus</emphasis>
</taxonomicName>
because of the cylindrical prolateral median process on male trochanter IV and generally large body size, and can be distinguished by: The ocularium with 2 large tubercles, the rectangular projection on the retrolateral margin of the prolateral apical apophysis of coxa IV and the shape of cylindrical prolateral median process of male trochanter IV: Prolateral edge of apex inflated, blunt; retrolateral edge of apex with blunt, upwards pointing finger-shaped projection; basally with a small, blunt, slightly conical apophysis. Penis without ventral process of glans (unusual within the genus); stylus curved ventrad, without trichomes and angular medio-ventral projection; ventral plate elliptic; trunk covering base of glans dorsoventrally.
</paragraph>
</subSubSection>
<subSubSection id="C320657895474674FF34FD658BE3FCB8" pageId="28" pageNumber="29" type="etymology">
<paragraph id="8B8536F395474674FF34FD658BE3FCB8" blockId="28.[151,1436,258,2002]" pageId="28" pageNumber="29">
<emphasis id="B94EEAE195474674FF34FD658B01FD43" bold="true" box="[198,343,706,732]" pageId="28" pageNumber="29">Etymology:</emphasis>
The species name, an adjective, refers to the
<typeStatus id="5481885195474674FC8DFD6589E7FD43" box="[895,945,706,732]" pageId="28" pageNumber="29">type</typeStatus>
locality, Colônia Alpina, a Swiss colony founded by Emil August Göldi. He was a remarkable and important Swiss researcher who helped to promote natural history in
<collectingCountry id="F32D766395474674FE94FCAA8BE7FCB8" box="[358,433,781,807]" name="Brazil" pageId="28" pageNumber="29">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C320657895474676FF34FC958AFAFB20" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="description">
<paragraph id="8B8536F395474674FF34FC9589C6FBB3" blockId="28.[151,1436,258,2002]" pageId="28" pageNumber="29">
<emphasis id="B94EEAE195474674FF34FC958B37FCD3" bold="true" box="[198,353,818,844]" pageId="28" pageNumber="29">Description:</emphasis>
<emphasis id="B94EEAE195474674FE9AFC958BF2FCD4" box="[360,420,818,843]" italics="true" pageId="28" pageNumber="29">Male</emphasis>
(
<typeStatus id="5481885195474674FE46FC958874FCD3" box="[436,546,818,844]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
): Dorsum (
<figureCitation id="13012A7695474674FD51FC95894BFCD3" box="[675,797,818,844]" captionStart="FIGURE 4" captionStartId="29.[151,255,1937,1961]" captionTargetBox="[203,1399,198,1908]" captionTargetId="figure-75@29.[197,1401,189,1918]" captionTargetPageId="29" captionText="FIGURE 4. Eusarcus alpinus sp. n. Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view(in part); D, right trochanter and femur IV, prolateral view; E, idem, dorsal view; F, idem, ventral view; G, right femur III, ventral view; H, right tibia III, ventral view. Scale bars, 1 mm." pageId="28" pageNumber="29">Fig. 4A,B</figureCitation>
): Measurements: SL 4.80; SMW 4.30; femur I 2.15; II 4.20; III 3.20; IV 3.80. Median paracheliceral projection of similar-size as PAM. Ocularium far from anterior scutal margin; with 2 large tubercles and 12 smaller ones. Carapace with 35 scattered tubercles. Scutal area I with 43 scattered tubercles on each side; II with 91; III with 100 and a median spine directed backwards, almost straight, curved only apically, surpassing posterior margin of dorsal scutum; IV with 50 tubercles. Posterior margin of dorsal scutum with 31 scattered tubercles. Free tergites I–III with an irregular row of 22, 18, 18 tubercles, respectively. Anal operculum with 39 tubercles.
</paragraph>
<paragraph id="8B8536F395474674FF37FB9F8E79FBE8" blockId="28.[151,1436,258,2002]" pageId="28" pageNumber="29">Venter: Coxa I with 33–34 tubercles, median and distal ones largest; II–III with 56–58, 60–69 tubercles increasing in size distally, respectively; IV densely and irregularly tuberculate.</paragraph>
<paragraph id="8B8536F395474674FF37FB2588D0FB03" blockId="28.[151,1436,258,2002]" box="[197,646,1154,1180]" pageId="28" pageNumber="29">Chelicera: Segment I with 2 tubercles.</paragraph>
<paragraph id="8B8536F395474674FF37FB0F8E0FFA93" blockId="28.[151,1436,258,2002]" pageId="28" pageNumber="29">Pedipalpus: Trochanter dorsally inflated, with 4 tubercles. Femur dorsally with 4 aligned tubercles, 5 apical small ones; ventrally with 1 large pointed basal tubercle, 2 small scattered tubercles. Tibial setation: Prolateral IiIi, retrolateral iiIi/IiIi. Tarsal setation: Prolateral IIi, retrolateral IiIi/Iii.</paragraph>
<paragraph id="8B8536F395474674FF37FABF8E0FF8A3" blockId="28.[151,1436,258,2002]" pageId="28" pageNumber="29">
Legs (
<figureCitation id="13012A7695474674FEE0FABF8BCDFAAD" box="[274,411,1304,1330]" captionStart="FIGURE 4" captionStartId="29.[151,255,1937,1961]" captionTargetBox="[203,1399,198,1908]" captionTargetId="figure-75@29.[197,1401,189,1918]" captionTargetPageId="29" captionText="FIGURE 4. Eusarcus alpinus sp. n. Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view(in part); D, right trochanter and femur IV, prolateral view; E, idem, dorsal view; F, idem, ventral view; G, right femur III, ventral view; H, right tibia III, ventral view. Scale bars, 1 mm." pageId="28" pageNumber="29">Fig. 4C–H</figureCitation>
): Coxa II with 5–7 tubercles on retrolateral margin; IV densely tuberculate, with 1 oblique prolateral apical apophysis, this conical, long, blunt, curved posteriad and slightly ventrad, with 1 rectangular projection on retrolateral margin. Trochanters I–IV with scattered tubercles; I–III with large retrolateral tubercle; I–II with 1 large ventro-basal tubercle; IV prolaterally with 1 cylindrical and flattened median process, this prolateral edge of apex inflated, blunt; retrolateral edge of apex with blunt, upwards pointing finger-shaped projection; basally with a small, blunt, slightly conical apophysis; retrolaterally with 1 large pointed subasal tubercle, 1 median and 1 apical large blunt tubercle; 1 large pointed ventro-apical tubercle. Femur I with reduced PDS, RDS; II–III with reduced PDS, large RDS; III sinuous, ventrally with 2 rows of tubercles increasing in size distally, prolateral ones enlarged, pointed, 1 small, blunt prolateral apical spine; IV sinuous, with medium PDS, large RDS; a retrolateral row of enlarged, slightly pointed tubercles; ventrally with two irregular rows of tubercles increasing in size distally, prolateral ones becoming pointed (5 large), 2 apical spines. Patellae III–IV with enlarged, slightly pointed ventral tubercles. Tibia III ventrally with 2 rows of pointed tubercles increasing in size subapically, retrolateral ones largest (3–4 large); IV ventrally with 2 rows of tubercles increasing in size distally, becoming pointed, 2 apical spines. Metatarsi III–IV with tubercles decreasing in size and number distally. Tarsal segmentation: 6, 8–9, 6, 6.
</paragraph>
<paragraph id="8B8536F395474674FF37F8EF8905F84D" blockId="28.[151,1436,258,2002]" pageId="28" pageNumber="29">
Penis (
<figureCitation id="13012A7695474674FEE4F8EF8BC9F8FD" box="[278,415,1864,1890]" captionStart="FIGURE 43" captionStartId="119.[151,255,1559,1583]" captionTargetBox="[157,1429,312,1530]" captionTargetId="figure-316@119.[151,1436,293,1538]" captionTargetPageId="119" captionText="FIGURE 43. Distal part of penis in dorsal and lateral view: A–B, Eusarcus alpinus sp. n.; C–D, E. didactylus sp. n.; E– F, E. nigrimaculatus. Scale bars, 0.1 mm." pageId="28" pageNumber="29">Fig. 43A,B</figureCitation>
): Stylus smooth, apex swollen, curved ventrad; without trichomes and angular medioventral projection. Ventral process of glans absent. Ventral plate elliptic; with 3 pairs of short distal setae; 1 pair of short, straight, thick median setae; 4 pairs of long, straight basal setae, basalmost pair very close to trunk of penis. Trunk covering base of glans dorsoventrally.
</paragraph>
<caption id="DF45667B95464675FF65F8368F70F86B" pageId="29" pageNumber="30" startId="29.[151,255,1937,1961]" targetBox="[203,1399,198,1908]" targetPageId="29">
<paragraph id="8B8536F395464675FF65F8368F70F86B" blockId="29.[151,1436,1937,2036]" pageId="29" pageNumber="30">
<emphasis id="B94EEAE195464675FF65F8368B4CF836" bold="true" box="[151,282,1937,1961]" pageId="29" pageNumber="30">FIGURE 4.</emphasis>
<taxonomicName id="4C3A4D7095464675FED1F8368B8AF837" authority="Hara &amp; Pinto-Da-Rocha, 2010" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[291,476,1937,1960]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="alpinus" status="sp. nov.">
<emphasis id="B94EEAE195464675FED1F8368B8AF837" box="[291,476,1937,1960]" italics="true" pageId="29" pageNumber="30">Eusarcus alpinus</emphasis>
</taxonomicName>
<emphasis id="B94EEAE195464675FE16F8368874F836" bold="true" box="[484,546,1937,1961]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A27D579A95464675FE16F8368874F836" box="[484,546,1937,1961]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view(in part); D, right trochanter and femur IV, prolateral view; E, idem, dorsal view; F, idem, ventral view; G, right femur III, ventral view; H, right tibia III, ventral view. Scale bars, 1 mm.
</paragraph>
</caption>
<paragraph id="8B8536F395454676FF37FF3F8959FF48" blockId="30.[151,1437,152,1290]" pageId="30" pageNumber="31">Coloration: Brown; dark brown on body margin, spines, apophyses and patellae, basal and apical areas of femora and tibiae. Light brown on metatarsi and tarsi.</paragraph>
<paragraph id="8B8536F395454676FF34FF458F24FD13" blockId="30.[151,1437,152,1290]" pageId="30" pageNumber="31">
<emphasis id="B94EEAE195454676FF34FF458B48FF64" box="[198,286,226,251]" italics="true" pageId="30" pageNumber="31">Female</emphasis>
(
<typeStatus id="5481885195454676FEDDFF458BCFFF63" box="[303,409,226,252]" pageId="30" pageNumber="31" type="paratype">paratype</typeStatus>
; MNRJ 5719): Dorsum: Measurements: SL 4.95; SMW 3.80 femur I 2.05; II 4.00; III 3.00; IV 3.70. Free tergites I–III with rows of 15 and 22, 12 and 24, 12 and 16 tubercles, respectively. Pedipalpus: Tibial setation: Prolateral and retrolateral IiIi. Tarsal setation: Prolateral IIi, retrolateral IiIi. Legs: Coxa II with 1 prolateral tubercle, 4 retrolateral; IV densely tuberculate, with 1 oblique prolateral apical apophysis, this conical, short, pointed, curved backwards, surpassing posterior margin of this segment. Trochanter IV with scattered tubercles, retrolaterally with 1 large median tubercle, 1 short, blunt apical apophysis. Femur I ventrally with 2 rows of slightly enlarged tubercles; III ventrally with 2 rows of tubercles increasing in size distally, prolateral ones enlarged, 1 short, blunt prolateral apical spine, 1 large retrolateral apical tubercle; IV ventrally with two irregular rows of tubercles (prolateral ones enlarged) increasing in size distally, 1 blunt prolateral apical spine, 1 large retrolateral apical tubercle. Tibiae I, IV with two ventral rows of slightly enlarged tubercles, without spines; III with rows of similar-sized tubercles. Tarsal segmentation: 6, 8, 6, 6.
</paragraph>
<paragraph id="8B8536F395454676FF37FD3D8F18FC40" blockId="30.[151,1437,152,1290]" pageId="30" pageNumber="31">
<emphasis id="B94EEAE195454676FF37FD3D8BF2FD2C" box="[197,420,666,691]" italics="true" pageId="30" pageNumber="31">Variation in males</emphasis>
(n=5): Measurements: SL 4.05–4.80; SMW 3.70–4.00; femur I 1.80–2.15; II 3.80– 4.20; III 2.90–3.60; IV 3.50–3.80. Median paracheliceral projection smaller or similar-sized as PAM. Pedipalpus: Tibial setation: Prolateral IiIi/iii, retrolateral IiIii/iiIi/IiIi/IIi. Tarsal setation: Prolateral IIi/iIi, retrolateral IiIi/Iii. Trochanter III with or without large ventro-basal tubercle; IV with 1–2 large retrolateral apical tubercles. Femora I–II ventrally with 2 rows of slightly enlarged or similar-sized tubercles; I with reduced or small PDS, RDS; II–III with reduced or small PDS, medium or large RDS; III with or without 1 large retrolateral ventro-apical tubercle. Tibia I with 2 rows of similar-sized tubercles or increasing in size distally; IV with or without 1 retrolateral apical spine. The beta male form has its coxa IV reaching grooves III–IV and its prolateral median apophysis of trochanter IV conical instead of cylindrical and flattened.
</paragraph>
<paragraph id="8B8536F395454676FF37FC4D8AFAFB20" blockId="30.[151,1437,152,1290]" pageId="30" pageNumber="31">
<emphasis id="B94EEAE195454676FF37FC4D8BE3FB9C" box="[197,437,1002,1027]" italics="true" pageId="30" pageNumber="31">Variation in females</emphasis>
(n=7): Measurements: SL 4.50–4.95; SMW 3.00–3.80; femur I 1.80–2.05; II 3.50– 4.00; III 2.80–3.00; IV 3.50–3.70. Pedipalpus: Tibial setation: Prolateral IiiIi/IiIi/IIi, retrolateral IiIii/IiIi/IIi. Trochanter III with or without large ventro-basal tubercle. Femur I with reduced or small RDS; II with medium or large RDS; III with reduced or small PDS, medium or large RDS; IV with medium or large RDS. Tibia I with two ventral rows of slightly enlarged or similar-sized tubercles. Tarsal segmentation: 6, 7–10, 6, 6.
</paragraph>
</subSubSection>
<subSubSection id="C320657895454676FF37FB6D89F5FA95" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8B8536F395454676FF37FB6D8F59FB7B" blockId="30.[151,1437,152,1290]" box="[197,1295,1226,1252]" pageId="30" pageNumber="31">
<materialsCitation id="3B523CAE95454676FF37FB6D8F5DFB7B" box="[197,1291,1226,1252]" country="Brazil" location="Parque Estadual do Desengano" municipality="Santa Maria Madalena" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<emphasis id="B94EEAE195454676FF37FB6D8B38FB7B" bold="true" box="[197,366,1226,1252]" pageId="30" pageNumber="31">
<typeStatus id="5481885195454676FF37FB6D8B54FB7B" box="[197,258,1226,1252]" pageId="30" pageNumber="31">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F32D766395454676FE84FB6D8B96FB7B" box="[374,448,1226,1252]" name="Brazil" pageId="30" pageNumber="31">Brazil</collectingCountry>
,
<collectingRegion id="49FEF81195454676FE3EFB6D882EFB7B" box="[460,632,1226,1252]" country="Brazil" name="Rio de Janeiro" pageId="30" pageNumber="31">Rio de Janeiro</collectingRegion>
,
<collectingMunicipality id="6BE1AC8995454676FD76FB6D89C6FB7B" box="[644,912,1226,1252]" pageId="30" pageNumber="31">Santa Maria Madalena</collectingMunicipality>
,
<location id="8EE5602895454676FC6EFB6D8F5DFB7B" LSID="urn:lsid:plazi:treatment:039387E595474676FF65FF3089F5FA95:8EE5602895454676FC6EFB6D8F5DFB7B" box="[924,1291,1226,1252]" country="Brazil" municipality="Santa Maria Madalena" name="Parque Estadual do Desengano" pageId="30" pageNumber="31" stateProvince="Rio de Janeiro">Parque Estadual do Desengano</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8536F395454676FF37FB4889F5FA95" blockId="30.[151,1437,152,1290]" box="[197,931,1263,1290]" pageId="30" pageNumber="31">
<emphasis id="B94EEAE195454676FF37FB48885FFA96" bold="true" box="[197,521,1263,1289]" pageId="30" pageNumber="31">Geographical distribution</emphasis>
(
<figureCitation id="13012A7695454676FDE8FB5788DFFA95" box="[538,649,1264,1290]" captionStart="FIGURE 49" captionStartId="125.[151,255,2004,2028]" captionTargetBox="[180,1389,206,1966]" captionTargetId="figure-26@125.[175,1399,194,1974]" captionTargetPageId="125" captionText="FIGURE 49. A, B: Localities of Eusarcus spp. in eastern Brazil." pageId="30" pageNumber="31">Fig. 49A</figureCitation>
):
<collectingCountry id="F32D766395454676FD6EFB5788B1FA95" box="[668,743,1264,1290]" name="Brazil" pageId="30" pageNumber="31">Brazil</collectingCountry>
.
<collectingRegion id="49FEF81195454676FD00FB5789C9FA95" box="[754,927,1264,1290]" country="Brazil" name="Rio de Janeiro" pageId="30" pageNumber="31">Rio de Janeiro</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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