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<document id="87F18E85BDC9AAFBC70CC19F21A18C2E" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629706805705" checkinUser="felipe" docAuthor="Hara, Marcos Ryotaro &amp; Pinto-Da-Rocha, Ricardo" docDate="2010" docId="039387E5953F460EFF65F9E089E3F9FD" docLanguage="en" docName="zt02698p136.pdf" docOrigin="Zootaxa 2698" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Eusarcus signatus Hara &amp; Pinto-Da-Rocha 2010, comb. n." docType="treatment" docVersion="4" lastPageNumber="103" masterDocId="FFAAFF9D955B4668FFF2FFA78A56FF9F" masterDocTitle="Systematic review and cladistic analysis of the genus Eusarcus Perty 1833 (Arachnida, Opiliones, Gonyleptidae) 2698" masterLastPageNumber="136" masterPageNumber="1" pageNumber="101" updateTime="1699098874055" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="BE2725AFACC40134BFC721CEEDBAE6CE">Systematic review and cladistic analysis of the genus Eusarcus Perty 1833 (Arachnida, Opiliones, Gonyleptidae) 2698</mods:title>
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<mods:namePart id="53F30052B31C06C0FF32469EDF5AF2A5">Hara, Marcos Ryotaro</mods:namePart>
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<mods:namePart id="DF067BDCDFA6C8D86BBE799E3A45908D">Pinto-Da-Rocha, Ricardo</mods:namePart>
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<mods:date id="9228BA1DA353F42A821D0645D7910C9B">2010</mods:date>
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<mods:number id="1D2FCCB490E1B0A095803CCBEE11FC13">2010-12-03</mods:number>
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<treatment id="039387E5953F460EFF65F9E089E3F9FD" ID-DOI="http://doi.org/10.5281/zenodo.5295615" ID-Zenodo-Dep="5295615" LSID="urn:lsid:plazi:treatment:039387E5953F460EFF65F9E089E3F9FD" httpUri="http://treatment.plazi.org/id/039387E5953F460EFF65F9E089E3F9FD" lastPageId="102" lastPageNumber="103" pageId="100" pageNumber="101">
<subSubSection id="C3206578953F460CFF65F9E08834F9FE" box="[151,610,1607,1633]" pageId="100" pageNumber="101" type="nomenclature">
<paragraph id="8B8536F3953F460CFF65F9E08834F9FE" blockId="100.[151,610,1607,1668]" box="[151,610,1607,1633]" pageId="100" pageNumber="101">
<heading id="D0CD819F953F460CFF65F9E08834F9FE" bold="true" box="[151,610,1607,1633]" fontSize="11" level="1" pageId="100" pageNumber="101" reason="1">
<emphasis id="B94EEAE1953F460CFF65F9E08834F9FE" bold="true" box="[151,610,1607,1633]" pageId="100" pageNumber="101">
<taxonomicName id="4C3A4D70953F460CFF65F9E08BA6F9FE" ID-CoL="7W5S6" authority="(Roewer)" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" baseAuthorityName="Roewer" box="[151,496,1607,1633]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="signatus" status="comb. nov.">
<emphasis id="B94EEAE1953F460CFF65F9E08B23F9FE" bold="true" box="[151,373,1607,1633]" italics="true" pageId="100" pageNumber="101">Eusarcus signatus</emphasis>
(Roewer)
</taxonomicName>
<taxonomicNameLabel id="A27D579A953F460CFE05F9E08834F9FE" box="[503,610,1607,1633]" pageId="100" pageNumber="101" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3206578953F460CFF65F9CD8AABF91B" box="[151,253,1642,1668]" pageId="100" pageNumber="101" type="description">
<paragraph id="8B8536F3953F460CFF65F9CD8AABF91B" blockId="100.[151,610,1607,1668]" box="[151,253,1642,1668]" pageId="100" pageNumber="101">
(
<figureCitation id="13012A76953F460CFF6DF9CD8AA2F91B" box="[159,244,1642,1668]" captionStart="FIGURE 32" captionStartId="101.[151,256,1950,1974]" captionTargetBox="[197,1390,262,1925]" captionTargetId="figure-93@101.[188,1399,247,1933]" captionTargetPageId="101" captionText="FIGURE 32. Eusarcus signatus (Roewer) comb. n. Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view (in part); D, right trochanter and femur IV, dorsal view; E, idem, ventral view; F, idem, prolateral view. Scale bars, 1 mm." pageId="100" pageNumber="101">Fig. 32</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3206578953F460CFF65F9128EA5F973" pageId="100" pageNumber="101" type="reference_group">
<paragraph id="8B8536F3953F460CFF65F9128EA5F973" blockId="100.[151,1436,1717,1772]" pageId="100" pageNumber="101">
<treatmentCitationGroup id="AB2A11DD953F460CFF65F9128EA5F973" pageId="100" pageNumber="101">
<treatmentCitation id="0A9B10E2953F460CFF65F9128816F953" author="Roewer, C. F." box="[151,576,1717,1740]" page="57" pageId="100" pageNumber="101" year="1949">
<taxonomicName id="4C3A4D70953F460CFF65F9128816F953" ID-CoL="9X4TN" authority="Roewer 1949: 57" authorityName="Roewer" authorityPageNumber="57" authorityYear="1949" box="[151,576,1717,1740]" class="Arachnida" family="Gonyleptidae" genus="Antetriceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="signatus">
<emphasis id="B94EEAE1953F460CFF65F9128B2AF953" box="[151,380,1717,1740]" italics="true" pageId="100" pageNumber="101">Antetriceras signatus</emphasis>
<bibRefCitation id="EFAB4B02953F460CFE71F9128816F953" author="Roewer, C. F." box="[387,576,1717,1740]" pageId="100" pageNumber="101" pagination="11 - 61" refId="ref86525" refString="Roewer, C. F. (1949) Weitere Weberknechte XIII. Uber Phalangodiden I. (Subfam. Phalangodinae, Tricommatinae, Samoinae). Senckenbergiana, 30 (1 - 3), 11 - 61." type="journal article" year="1949">Roewer 1949: 57</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 111a–d;
<treatmentCitation id="0A9B10E2953F460CFD26F91289DDF953" author="Kury, A. B." box="[724,907,1717,1740]" page="156" pageId="100" pageNumber="101" year="2003">
<bibRefCitation id="EFAB4B02953F460CFD26F91289DDF953" author="Kury, A. B." box="[724,907,1717,1740]" pageId="100" pageNumber="101" pagination="5 - 337" refId="ref83946" refString="Kury, A. B. (2003 a) Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. esp. monografico n o 1, 5 - 337." type="journal article" year="2003">Kury 2003a: 156</bibRefCitation>
</treatmentCitation>
(cat); 2003b: 6 (syst); (ma
<typeStatus id="54818851953F460CFB45F9128F4EF953" box="[1207,1304,1717,1740]" pageId="100" pageNumber="101" type="holotype">holotype</typeStatus>
; &quot;Terra Vermelha,
<collectingRegion id="49FEF811953F460CFEE7F9728BE4F973" box="[277,434,1749,1772]" country="Brazil" name="Rio de Janeiro" pageId="100" pageNumber="101">Rio de Janeiro</collectingRegion>
; SMF RII 5078/12&quot;; examined, without genitalia, spine of ocularium broken).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3206578953F460CFF65F8BD88DAF8C5" pageId="100" pageNumber="101" type="materials_examined">
<paragraph id="8B8536F3953F460CFF65F8BD88DAF8C5" blockId="100.[151,1436,1818,2031]" pageId="100" pageNumber="101">
<emphasis id="B94EEAE1953F460CFF65F8BD8BD0F8AB" bold="true" box="[151,390,1818,1844]" pageId="100" pageNumber="101">Material examined</emphasis>
:
<materialsCitation id="3B523CAE953F460CFE64F8BD88EBF8AB" box="[406,701,1818,1844]" country="Brazil" location="Rio de Janeiro" pageId="100" pageNumber="101" specimenCount="1" stateProvince="Rio de Janeiro">
<collectingCountry id="F32D7663953F460CFE64F8BD8852F8AB" box="[406,516,1818,1844]" name="Brazil" pageId="100" pageNumber="101">BRAZIL</collectingCountry>
.
<collectingRegion id="49FEF811953F460CFDE3F8BD88EBF8AB" box="[529,701,1818,1844]" country="Brazil" name="Rio de Janeiro" pageId="100" pageNumber="101">Rio de Janeiro</collectingRegion>
</materialsCitation>
:
<materialsCitation id="3B523CAE953F460CFD3FF8BD88DEF8C5" collectionCode="SMF" country="Brazil" location="Rio de Janeiro" pageId="100" pageNumber="101" specimenCode="RII 5078" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<collectingRegion id="49FEF811953F460CFD3FF8BD892DF8AB" box="[717,891,1818,1844]" country="Brazil" name="Rio de Janeiro" pageId="100" pageNumber="101">Rio de Janeiro</collectingRegion>
? (Terra Vermelha), without name of collector and date, ma
<typeStatus id="54818851953F460CFEC1F8E78BCFF8C5" box="[307,409,1856,1882]" pageId="100" pageNumber="101" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED2BAE36953F460CFE5BF8E78BB3F8C5" box="[425,485,1856,1882]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="100" pageNumber="101">SMF</collectionCode>
<specimenCode id="DB9C9E88953F460CFE1EF8E7880CF8C5" box="[492,602,1856,1882]" collectionCode="RII" pageId="100" pageNumber="101">RII 5078</specimenCode>
/12)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3206578953F460DFF34F8C28820FF48" lastPageId="101" lastPageNumber="102" pageId="100" pageNumber="101" type="diagnosis">
<paragraph id="8B8536F3953F460DFF34F8C28820FF48" blockId="100.[151,1436,1818,2031]" lastBlockId="101.[151,1436,152,215]" lastPageId="101" lastPageNumber="102" pageId="100" pageNumber="101">
<emphasis id="B94EEAE1953F460CFF34F8C28B1EF8E0" bold="true" box="[198,328,1893,1919]" pageId="100" pageNumber="101">Diagnosis:</emphasis>
<taxonomicName id="4C3A4D70953F460CFEABF8C28BB2F8E1" authorityName="Roewer" authorityYear="1949" box="[345,484,1893,1919]" class="Arachnida" family="Gonyleptidae" genus="Antetriceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="signatus">
<emphasis id="B94EEAE1953F460CFEABF8C28B3DF8E1" box="[345,363,1893,1918]" italics="true" pageId="100" pageNumber="101">E</emphasis>
.
<emphasis id="B94EEAE1953F460CFE71F8C28BB2F8E1" box="[387,484,1893,1918]" italics="true" pageId="100" pageNumber="101">signatus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C3A4D70953F460CFD88F8C28953F8E1" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1942" box="[634,773,1893,1919]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="aduncus">
<emphasis id="B94EEAE1953F460CFD88F8C288DAF8E1" box="[634,652,1893,1918]" italics="true" pageId="100" pageNumber="101">E</emphasis>
.
<emphasis id="B94EEAE1953F460CFD56F8C28953F8E1" box="[676,773,1893,1918]" italics="true" pageId="100" pageNumber="101">aduncus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D70953F460CFCEFF8C289C4F8E1" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" baseAuthorityName="Hara &amp; Pinto-Da-Rocha" baseAuthorityYear="2010" box="[797,914,1893,1919]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="berlae">
<emphasis id="B94EEAE1953F460CFCEFF8C28979F8E1" box="[797,815,1893,1918]" italics="true" pageId="100" pageNumber="101">E</emphasis>
.
<emphasis id="B94EEAE1953F460CFCB5F8C289C4F8E1" box="[839,914,1893,1918]" italics="true" pageId="100" pageNumber="101">berlae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D70953F460CFC5BF8C28E4CF8E1" authorityName="Kury" authorityYear="2008" box="[937,1050,1893,1919]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="elinae">
<emphasis id="B94EEAE1953F460CFC5BF8C289EDF8E1" box="[937,955,1893,1918]" italics="true" pageId="100" pageNumber="101">E</emphasis>
.
<emphasis id="B94EEAE1953F460CFC21F8C28E4CF8E1" box="[979,1050,1893,1918]" italics="true" pageId="100" pageNumber="101">elinae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D70953F460CFBC0F8C28EF4F8E1" box="[1074,1186,1893,1919]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="fulvus">
<emphasis id="B94EEAE1953F460CFBC0F8C28E12F8E1" box="[1074,1092,1893,1918]" italics="true" pageId="100" pageNumber="101">E</emphasis>
.
<emphasis id="B94EEAE1953F460CFBAEF8C28EF4F8E1" box="[1116,1186,1893,1918]" italics="true" pageId="100" pageNumber="101">fulvus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D70953F460CFB48F8C28F3CF8E1" authorityName="Soares &amp; Soares" authorityYear="1946" box="[1210,1386,1893,1919]" class="Arachnida" family="Gonyleptidae" genus="Metagraphinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="sooretamae">
<emphasis id="B94EEAE1953F460CFB48F8C28E9AF8E1" box="[1210,1228,1893,1918]" italics="true" pageId="100" pageNumber="101">E</emphasis>
.
<emphasis id="B94EEAE1953F460CFB11F8C28F3CF8E1" box="[1251,1386,1893,1918]" italics="true" pageId="100" pageNumber="101">sooretamae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D70953F460CFA71F8C28B75F83B" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="B94EEAE1953F460CFA71F8C28FC3F8E1" box="[1411,1429,1893,1918]" italics="true" pageId="100" pageNumber="101">E</emphasis>
.
<emphasis id="B94EEAE1953F460CFF65F82C8B75F83B" box="[151,291,1931,1956]" italics="true" pageId="100" pageNumber="101">cavernicola</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D70953F460CFEC1F82C8BE9F83B" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[307,447,1930,1956]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="B94EEAE1953F460CFEC1F82C8B13F83B" box="[307,325,1931,1956]" italics="true" pageId="100" pageNumber="101">E</emphasis>
.
<emphasis id="B94EEAE1953F460CFEA6F82C8BE9F83B" box="[340,447,1931,1956]" italics="true" pageId="100" pageNumber="101">mirabilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D70953F460CFE09F82C88C8F83B" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" box="[507,670,1930,1956]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="sergipanus">
<emphasis id="B94EEAE1953F460CFE09F82C885BF83B" box="[507,525,1931,1956]" italics="true" pageId="100" pageNumber="101">E</emphasis>
.
<emphasis id="B94EEAE1953F460CFDEFF82C88C8F83B" box="[541,670,1931,1956]" italics="true" pageId="100" pageNumber="101">sergipanus</emphasis>
</taxonomicName>
because of the conical prolateral median apophysis on the male trochanter IV, and can be distinguished by: The presence of PAM; median spine of scutal area III of medium size; basitarsus I not elongated; prolateral apical apophysis of the male coxa IV with triangular ventral projection; femur IV not longer than 1.5 times dorsal scutum length, with 1 PVS and a large retrolateral apical tubercle; ocularium with a median spine.
</paragraph>
</subSubSection>
<caption id="DF45667B953E460DFF65F8398962F86A" pageId="101" pageNumber="102" startId="101.[151,256,1950,1974]" targetBox="[197,1390,262,1925]" targetPageId="101">
<paragraph id="8B8536F3953E460DFF65F8398962F86A" blockId="101.[151,1436,1950,2037]" pageId="101" pageNumber="102">
<emphasis id="B94EEAE1953E460DFF65F8398B74F829" bold="true" box="[151,290,1950,1974]" pageId="101" pageNumber="102">FIGURE 32</emphasis>
.
<taxonomicName id="4C3A4D70953E460DFEC3F8398835F82A" authority="(Roewer)" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" baseAuthorityName="Hara &amp; Pinto-Da-Rocha" baseAuthorityYear="2010" box="[305,611,1950,1973]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="101" pageNumber="102" phylum="Arthropoda" rank="species" species="signatus" status="comb. nov.">
<emphasis id="B94EEAE1953E460DFEC3F8398BA0F82A" box="[305,502,1950,1973]" italics="true" pageId="101" pageNumber="102">Eusarcus signatus</emphasis>
(Roewer)
</taxonomicName>
<emphasis id="B94EEAE1953E460DFD9EF8398898F829" bold="true" box="[620,718,1950,1974]" pageId="101" pageNumber="102">
<taxonomicNameLabel id="A27D579A953E460DFD9EF8398898F829" box="[620,718,1950,1974]" pageId="101" pageNumber="102" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view (in part); D, right trochanter and femur IV, dorsal view; E, idem, ventral view; F, idem, prolateral view. Scale bars, 1 mm.
</paragraph>
</caption>
<subSubSection id="C3206578953D460EFF37FF308B14FAC3" pageId="102" pageNumber="103" type="description">
<paragraph id="8B8536F3953D460EFF37FF308B5EFE0D" blockId="102.[151,1436,151,1634]" pageId="102" pageNumber="103">
<emphasis id="B94EEAE1953D460EFF37FF308BD4FF2E" bold="true" box="[197,386,151,177]" pageId="102" pageNumber="103">Redescription:</emphasis>
Male (
<typeStatus id="54818851953D460EFE29FF3F881BFF2D" box="[475,589,152,178]" pageId="102" pageNumber="103" type="holotype">holotype</typeStatus>
): Dorsum (
<figureCitation id="13012A76953D460EFD26FF3F8934FF2D" box="[724,866,152,178]" captionStart="FIGURE 32" captionStartId="101.[151,256,1950,1974]" captionTargetBox="[197,1390,262,1925]" captionTargetId="figure-93@101.[188,1399,247,1933]" captionTargetPageId="101" captionText="FIGURE 32. Eusarcus signatus (Roewer) comb. n. Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view (in part); D, right trochanter and femur IV, dorsal view; E, idem, ventral view; F, idem, prolateral view. Scale bars, 1 mm." pageId="102" pageNumber="103">Fig. 32A,B</figureCitation>
): Measurements: SL 2.80; SMW 2.50; femur I 1.50; II 3.00; III 2.20; IV 2.70. Median paracheliceral projection of similar-size as PAM. Ocularium far from anterior scutal margin; with a median spine, 6 tubercles. Carapace with 23 scattered tubercles. Scutal area I with 13–14 scattered tubercles on each side; II with 28; III with 27 and a median spine of medium size, reaching groove V; IV with 21 tubercles. Posterior margin of dorsal scutum with 7 scattered tubercles and a row of 13 tubercles. Free tergites I–III with a row of 9, 8, 9 tubercles, respectively. Anal operculum with 14 tubercles.
</paragraph>
<paragraph id="8B8536F3953D460EFF37FE3A89CBFE43" blockId="102.[151,1436,151,1634]" pageId="102" pageNumber="103">Venter: Coxa I with 20 tubercles, median and distal ones largest; II–III with 31–37 and 26–36, respectively, increasing in size distally; IV irregularly tuberculate.</paragraph>
<paragraph id="8B8536F3953D460EFF37FE4F8874FD9D" blockId="102.[151,1436,151,1634]" box="[197,546,488,514]" pageId="102" pageNumber="103">Chelicera: Segment I smooth.</paragraph>
<paragraph id="8B8536F3953D460EFF37FDAA8BCBFDED" blockId="102.[151,1436,151,1634]" pageId="102" pageNumber="103">Pedipalpus: Trochanter dorsally inflated, with 2 tubercles. Femur dorsally with small, scattered tubercles; ventrally with 2 scattered tubercles. Tibial setation: Prolateral and retrolateral IiIi. Tarsal setation: Prolateral iiIi/iIi, retrolateral IiIi.</paragraph>
<paragraph id="8B8536F3953D460EFF37FDDA8908FC78" blockId="102.[151,1436,151,1634]" pageId="102" pageNumber="103">
Legs (
<figureCitation id="13012A76953D460EFEE3FDDA8BF5FD08" box="[273,419,637,663]" captionStart="FIGURE 32" captionStartId="101.[151,256,1950,1974]" captionTargetBox="[197,1390,262,1925]" captionTargetId="figure-93@101.[188,1399,247,1933]" captionTargetPageId="101" captionText="FIGURE 32. Eusarcus signatus (Roewer) comb. n. Male (holotype): A, habitus, dorsal view; B, dorsal scutum, right lateral view; C, right coxa and trochanter IV, posterior view (in part); D, right trochanter and femur IV, dorsal view; E, idem, ventral view; F, idem, prolateral view. Scale bars, 1 mm." pageId="102" pageNumber="103">Fig. 32C–F</figureCitation>
): Coxa II with 1–2 tubercles on retrolateral margin; IV densely tuberculate, ending between grooves III and IV of dorsal scutum, surpassing dorsal scutum in dorsal view only apically, with 1 transversal prolateral apical apophysis, this conical, short, blunt, with 1 triangular ventral projection and another one, transversal, in the middle. Trochanters I–IV with scattered tubercles; I with 1 large ventro-basal tubercle; III with 1 large retrolateral apical tubercle; IV prolaterally with 1 median apophysis, this conical, curved anteriad; retrolaterally with 1 short, pointed apical apophysis. Femur I with reduced PDS, RDS; II–III with reduced PDS, medium RDS; III sinuous, ventrally with a prolateral row of tubercles slightly increasing in size distally, 1 large prolateral apical tubercle; IV sinuous, with medium PDS, RDS; ventrally with 2 rows of tubercles increasing in size distally, prolateral ones enlarged, 1 prolateral apical apophysis, 1 large retrolateral apical tubercle. Tarsal segmentation: 6, 8–9, 6, 6.
</paragraph>
<paragraph id="8B8536F3953D460EFF37FC558919FB93" blockId="102.[151,1436,151,1634]" box="[197,847,1010,1036]" pageId="102" pageNumber="103">Coloration: Brown; dark brown on prosoma and leg IV.</paragraph>
<paragraph id="8B8536F3953D460EFF37FBBF8BF0FBAD" blockId="102.[151,1436,151,1634]" box="[197,422,1048,1074]" pageId="102" pageNumber="103">Female: Unknown.</paragraph>
<paragraph id="8B8536F3953D460EFF37FB9A8B14FAC3" blockId="102.[151,1436,151,1634]" pageId="102" pageNumber="103">
<emphasis id="B94EEAE1953D460EFF37FB9A8BFEFBC8" bold="true" box="[197,424,1085,1111]" pageId="102" pageNumber="103">Taxonomical note</emphasis>
:
<bibRefCitation id="EFAB4B02953D460EFE4BFB9A880BFBC8" author="Kury, A. B." box="[441,605,1085,1111]" pageId="102" pageNumber="103" pagination="1 - 8" refId="ref83989" refString="Kury, A. B. (2003 b) A new species of Pherania Strand, 1942 from southern Brazil (Arachnida: Opiliones: Gonyleptidae). Zootaxa, 363, 1 - 8." type="journal article" year="2003">Kury (2003b)</bibRefCitation>
suggested an affinity between the genera
<taxonomicName id="4C3A4D70953D460EFBAFFB9A8FCDFBC8" authority="Roewer 1949" authorityName="Roewer" authorityYear="1949" box="[1117,1435,1085,1111]" class="Arachnida" family="Gonyleptidae" genus="Antetriceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="102" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAE1953D460EFBAFFB9A8EA7FBC9" box="[1117,1265,1085,1110]" italics="true" pageId="102" pageNumber="103">Antetriceras</emphasis>
<bibRefCitation id="EFAB4B02953D460EFB09FB9A8FCDFBC8" author="Roewer, C. F." box="[1275,1435,1085,1111]" pageId="102" pageNumber="103" pagination="11 - 61" refId="ref86525" refString="Roewer, C. F. (1949) Weitere Weberknechte XIII. Uber Phalangodiden I. (Subfam. Phalangodinae, Tricommatinae, Samoinae). Senckenbergiana, 30 (1 - 3), 11 - 61." type="journal article" year="1949">Roewer 1949</bibRefCitation>
</taxonomicName>
(here placed in the synonymy of
<taxonomicName id="4C3A4D70953D460EFDC4FBC488F1FBE3" authorityName="Perty" authorityYear="1833" box="[566,679,1123,1148]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="102" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAE1953D460EFDC4FBC488F1FBE3" box="[566,679,1123,1148]" italics="true" pageId="102" pageNumber="103">Eusarcus</emphasis>
</taxonomicName>
) and
<taxonomicName id="4C3A4D70953D460EFD00FBC48E53FBE3" authority="Strand 1942" authorityName="Strand" authorityYear="1942" box="[754,1029,1122,1148]" class="Arachnida" family="Gonyleptidae" genus="Pherania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="102" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAE1953D460EFD00FBC48933FBE3" box="[754,869,1123,1148]" italics="true" pageId="102" pageNumber="103">Pherania</emphasis>
<bibRefCitation id="EFAB4B02953D460EFC9CFBC58E53FBE3" author="Strand, E." box="[878,1029,1122,1148]" pageId="102" pageNumber="103" pagination="386 - 402" refId="ref88546" refString="Strand, E. (1942) Miscellanea nomenclatoria zoologica et paleontologica. X. Folia zoologica et hydrobiologica, 11 (1), 386 - 402." type="journal article" year="1942">Strand 1942</bibRefCitation>
</taxonomicName>
on the basis of descriptions and illustrations of
<taxonomicName id="4C3A4D70953D460EFEBBFB2F8B9DFB3E" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" baseAuthorityName="Hara &amp; Pinto-Da-Rocha" baseAuthorityYear="2010" box="[329,459,1160,1185]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="signatus">
<emphasis id="B94EEAE1953D460EFEBBFB2F8B9DFB3E" box="[329,459,1160,1185]" italics="true" pageId="102" pageNumber="103">A. signatus</emphasis>
</taxonomicName>
. Unfortunately, illustrations of apophyses as well as the width of coxa IV provided by Roewer were inaccurate. Thus
<bibRefCitation id="EFAB4B02953D460EFDDFFB0A8898FB58" author="Kury, A. B." box="[557,718,1197,1223]" pageId="102" pageNumber="103" pagination="1 - 8" refId="ref83989" refString="Kury, A. B. (2003 b) A new species of Pherania Strand, 1942 from southern Brazil (Arachnida: Opiliones: Gonyleptidae). Zootaxa, 363, 1 - 8." type="journal article" year="2003">Kury (2003b)</bibRefCitation>
was misled to propose an unlikely association. Although no member of
<taxonomicName id="4C3A4D70953D460EFEEDFB748BDAFB73" authorityName="Strand" authorityYear="1942" box="[287,396,1235,1260]" class="Arachnida" family="Gonyleptidae" genus="Pherania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="102" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAE1953D460EFEEDFB748BDAFB73" box="[287,396,1235,1260]" italics="true" pageId="102" pageNumber="103">Pherania</emphasis>
</taxonomicName>
was included in any cladistic analysis, it is improbable that this genus is closely related to
<taxonomicName id="4C3A4D70953D460EFF44FB5F8B1FFA8E" authorityName="Roewer" authorityYear="1949" box="[182,329,1272,1297]" class="Arachnida" family="Gonyleptidae" genus="Antetriceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="102" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAE1953D460EFF44FB5F8B1FFA8E" box="[182,329,1272,1297]" italics="true" pageId="102" pageNumber="103">Antetriceras</emphasis>
</taxonomicName>
, since the former does not have the synapomorphies of
<taxonomicName id="4C3A4D70953D460EFC19FB5F8E0EFA8E" authorityName="Perty" authorityYear="1833" box="[1003,1112,1272,1297]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="102" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAE1953D460EFC19FB5F8E0EFA8E" box="[1003,1112,1272,1297]" italics="true" pageId="102" pageNumber="103">Eusarcus</emphasis>
</taxonomicName>
, such as a median spine in scutal area III or a paramedian pair of apophyses placed ectally to the chelicera on the anterior margin of the dorsal scutum.
</paragraph>
</subSubSection>
<subSubSection id="C3206578953D460EFF37FAC089E3F9FD" pageId="102" pageNumber="103" type="materials_examined">
<paragraph id="8B8536F3953D460EFF37FAC08F30F9A3" blockId="102.[151,1436,151,1634]" pageId="102" pageNumber="103">
<emphasis id="B94EEAE1953D460EFF37FAC08B38FA1E" bold="true" box="[197,366,1383,1409]" pageId="102" pageNumber="103">Type locality:</emphasis>
<materialsCitation id="3B523CAE953D460EFE87FACF8BA3FA53" country="Brazil" location="Rio de Janeiro" pageId="102" pageNumber="103" specimenCount="1" stateProvince="Rio de Janeiro">
<collectingCountry id="F32D7663953D460EFE87FACF8BE9FA1D" box="[373,447,1384,1410]" name="Brazil" pageId="102" pageNumber="103">Brazil</collectingCountry>
,
<collectingRegion id="49FEF811953D460EFE39FACF8821FA1D" box="[459,631,1384,1410]" country="Brazil" name="Rio de Janeiro" pageId="102" pageNumber="103">Rio de Janeiro</collectingRegion>
,
<collectingRegion id="49FEF811953D460EFD70FACF897CFA1D" box="[642,810,1384,1410]" country="Brazil" name="Rio de Janeiro" pageId="102" pageNumber="103">Rio de Janeiro</collectingRegion>
(?). Precising the exact type locality of this species is a difficult task. When Roewer described the species, he only stated “Terra Vermelha”, a very common name that occurs in many places of
</materialsCitation>
<materialsCitation id="3B523CAE953D460EFE0CFA158F34F9A3" country="Brazil" location="However" municipality="On" pageId="102" pageNumber="103" specimenCount="1" stateProvince="Santa Catarina" typeStatus="holotype">
<collectingCountry id="F32D7663953D460EFE0CFA15881CFA53" box="[510,586,1458,1484]" name="Brazil" pageId="102" pageNumber="103">Brazil</collectingCountry>
.
<collectingMunicipality id="6BE1AC89953D460EFDA4FA15882CFA53" box="[598,634,1458,1484]" pageId="102" pageNumber="103">On</collectingMunicipality>
the basis of the localities and information from the collector,
<bibRefCitation id="EFAB4B02953D460EFAADFA158AA2FA6D" author="Kury, A. B." pageId="102" pageNumber="103" pagination="5 - 337" refId="ref83946" refString="Kury, A. B. (2003 a) Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. esp. monografico n o 1, 5 - 337." type="journal article" year="2003">Kury (2003a)</bibRefCitation>
gave “
<collectingRegion id="49FEF811953D460EFEBBFA7F8854FA6D" box="[329,514,1496,1522]" country="Brazil" name="Santa Catarina" pageId="102" pageNumber="103">Santa Catarina</collectingRegion>
,
<collectingRegion id="49FEF811953D460EFDFCFA7F8802FA6D" box="[526,596,1496,1522]" country="Brazil" name="Ceara" pageId="102" pageNumber="103">Seara</collectingRegion>
, Terra Vermelha” as the type locality.
<location id="8EE56028953D460EFBDBFA7F8ECFFA6D" LSID="urn:lsid:plazi:treatment:039387E5953F460EFF65F9E089E3F9FD:8EE56028953D460EFBDBFA7F8ECFFA6D" box="[1065,1177,1496,1522]" country="Brazil" municipality="On" name="However" pageId="102" pageNumber="103" stateProvince="Santa Catarina">However</location>
, we verified that it is clearly written “
<collectingRegion id="49FEF811953D460EFEA4FA5A885FF988" box="[342,521,1533,1559]" country="Brazil" name="Rio de Janeiro" pageId="102" pageNumber="103">Rio de Janeiro</collectingRegion>
” on the label of the
<typeStatus id="54818851953D460EFD0EFA5A8931F988" box="[764,871,1533,1559]" pageId="102" pageNumber="103" type="holotype">holotype</typeStatus>
. Although risky, considering the distribution of species closely related to
<taxonomicName id="4C3A4D70953D460EFE30F9848812F9A3" authorityName="Hara &amp; Pinto-Da-Rocha" authorityYear="2010" baseAuthorityName="Hara &amp; Pinto-Da-Rocha" baseAuthorityYear="2010" box="[450,580,1571,1596]" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" order="Opiliones" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="signatus">
<emphasis id="B94EEAE1953D460EFE30F9848812F9A3" box="[450,580,1571,1596]" italics="true" pageId="102" pageNumber="103">A. signatus</emphasis>
</taxonomicName>
, we suggest the city of
<collectingRegion id="49FEF811953D460EFCA8F9858E55F9A3" box="[858,1027,1570,1596]" country="Brazil" name="Rio de Janeiro" pageId="102" pageNumber="103">Rio de Janeiro</collectingRegion>
as the probable exact locality
</materialsCitation>
.
</paragraph>
<paragraph id="8B8536F3953D460EFF37F9E089E3F9FD" blockId="102.[151,1436,151,1634]" box="[197,949,1607,1634]" pageId="102" pageNumber="103">
<emphasis id="B94EEAE1953D460EFF37F9E0885FF9FE" bold="true" box="[197,521,1607,1633]" pageId="102" pageNumber="103">Geographical distribution</emphasis>
: Known only from the
<typeStatus id="54818851953D460EFCEFF9EF8919F9FD" box="[797,847,1608,1634]" pageId="102" pageNumber="103">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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