354 lines
46 KiB
XML
354 lines
46 KiB
XML
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<mods:title id="9F74DBE92F704D5B167AEF7B6DC252A1">Diversity and systematics of Haminoeidae gastropods (Heterobranchia: Cephalaspidea) in the tropical West Pacific Ocean: new data on the genera Aliculastrum, Atys, Diniatys and Liloa</mods:title>
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<mods:namePart id="CE206F80109B75D0C8ADEC9535AF909A">Too, Chin Chin</mods:namePart>
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<mods:affiliation id="6567297A6D6A4AF8517DDF43B2A8EC3F">Affiliated to: Marine Laboratory, University of Guam, Mangilao, GU 96923</mods:affiliation>
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<mods:affiliation id="50D8822D9BE1062D31F00116C5EE5AD5">Affiliated to: Marine Laboratory, University of Guam, Mangilao, GU 96923</mods:affiliation>
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<mods:namePart id="C16799DDD111C1DC313E909901C1EF4E">Malaquias, Manuel António E.</mods:namePart>
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<emphasis id="B94EEACCFF94A274FF78FE59FEB368FC" bold="true" box="[151,351,476,502]" italics="true" pageId="30" pageNumber="385">Liloa porcellana</emphasis>
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(
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)
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</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph id="8B8536DEFF94A274FF78FDA7FB946B32" blockId="30.[151,1322,546,660]" box="[151,1144,546,568]" pageId="30" pageNumber="385">
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<treatmentCitation id="0A9B10CFFF94A274FF78FDA7FE1D6B32" author="Gould, A. A." box="[151,497,546,568]" page="138" pageId="30" pageNumber="385" year="1859">
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<taxonomicName id="4C3A4D5DFF94A274FF78FDA7FE1D6B32" ID-CoL="JP4C" authority="Gould, 1859: 138" authorityName="Gould" authorityPageNumber="138" authorityYear="1859" box="[151,497,546,568]" class="Gastropoda" family="Haminoeidae" genus="Atys" kingdom="Animalia" order="Cephalaspidea" pageId="30" pageNumber="385" phylum="Mollusca" rank="species" species="porcellana">
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<emphasis id="B94EEACCFF94A274FF78FDA7FEDA6B32" box="[151,310,546,568]" italics="true" pageId="30" pageNumber="385">Atys porcellana</emphasis>
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<bibRefCitation id="EFAB4B2FFF94A274FED3FDA7FE1D6B32" author="Gould, A. A." box="[316,497,546,568]" pageId="30" pageNumber="385" pagination="138 - 143" refId="ref18907" refString="Gould, A. A. (1859) Description of new species of shells brought home by the North Pacific Exploring Expedition. Proceedings of the Boston Society of Natural History, 7, 138 - 143." type="journal article" year="1859">Gould, 1859: 138</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<bibRefCitation id="EFAB4B2FFF94A274FE15FDA7FD7D6B32" author="Sowerby, G. B." box="[506,657,546,568]" pageId="30" pageNumber="385" pagination="1843 - 1878" refId="ref21194" refString="Sowerby, G. B. (1870) Monograph of the genus Atys. In: Reeve, L. A. (Ed.), Conchologica Iconica, or Illustrations of the shells of molluscous animals 1843 - 1878. Vol. 17. L. Reeve & Co, London. [pages not numbered]" type="book chapter" year="1870">Sowerby 1870</bibRefCitation>
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: pl. 5, sp. 30.
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<treatmentCitation id="0A9B10CFFF94A274FCF2FDA7FC336B32" author="Johnson, R. I." box="[797,991,546,568]" page="130" pageId="30" pageNumber="385" year="1964">
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<bibRefCitation id="EFAB4B2FFF94A274FCF2FDA7FC336B32" author="Johnson, R. I." box="[797,991,546,568]" pageId="30" pageNumber="385" pagination="1 - 182" refId="ref19203" refString="Johnson, R. I. (1964) The recent mollusc of Augustus Addison Gould: Illustrations of the types described by Gould with a bibliography and catalogue of his species. Smithsonian Institution, Washington, D. C, pp. 1 - 182." type="book chapter" year="1964">Johnson 1964: 130</bibRefCitation>
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</treatmentCitation>
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, pl. 12, fig. 13.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B8536DEFF94A274FF78FDC4FBDB6B5D" blockId="30.[151,1322,546,660]" box="[151,1079,577,599]" pageId="30" pageNumber="385">
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<treatmentCitationGroup id="AB2A11F0FF94A274FF78FDC4FBDB6B5D" box="[151,1079,577,599]" pageId="30" pageNumber="385">
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<taxonomicName id="4C3A4D5DFF94A274FF78FDC4FE7A6B5D" ID-CoL="JP4C" authorityName="Gould" authorityYear="1859" box="[151,406,577,599]" class="Gastropoda" family="Haminoeidae" genus="Atys" kingdom="Animalia" order="Cephalaspidea" pageId="30" pageNumber="385" phylum="Mollusca" rank="species" species="porcellana" subGenus="Alicula">
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<emphasis id="B94EEACCFF94A274FF78FDC4FE7A6B5D" box="[151,406,577,599]" italics="true" pageId="30" pageNumber="385">Atys (Alicula) porcellana</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="0A9B10CFFF94A274FE40FDC4FD896B5D" author="Pilsbry, H. A." box="[431,613,577,599]" page="268" pageId="30" pageNumber="385" year="1895">
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<bibRefCitation id="EFAB4B2FFF94A274FE40FDC4FD896B5D" author="Pilsbry, H. A." box="[431,613,577,599]" pageId="30" pageNumber="385" refId="ref20498" refString="Pilsbry, H. A. (1895) Manual of Conchology, structural and systematic. Vol. 15. Academy of Natural Sciences of Philadelphia, Philadelphia, 436 pp." type="book" year="1895">Pilsbry 1895: 268</bibRefCitation>
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</treatmentCitation>
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, pl. 28, fig. 23.
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||
<treatmentCitation id="0A9B10CFFF94A274FCEAFDC4FC406B5D" author="Kobelt, W." box="[773,940,577,599]" page="26" pageId="30" pageNumber="385" year="1896">
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<bibRefCitation id="EFAB4B2FFF94A274FCEAFDC4FC406B5D" author="Kobelt, W." box="[773,940,577,599]" pageId="30" pageNumber="385" pagination="1 - 32" refId="ref19292" refString="Kobelt, W. (1896) Die Familie Bullidae. In: Systematisches Conchylien-Cabinet. Verlag von Bauer & Raspe, Nurenberg, pp. 1 - 32." type="book chapter" year="1896">Kobelt 1896: 26</bibRefCitation>
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</treatmentCitation>
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, pl. 8, fig. 11.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B8536DEFF94A274FF78FDE4FD826B7C" blockId="30.[151,1322,546,660]" box="[151,622,608,630]" pageId="30" pageNumber="385">
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<treatmentCitationGroup id="AB2A11F0FF94A274FF78FDE4FD826B7C" box="[151,622,608,630]" pageId="30" pageNumber="385">
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<treatmentCitation id="0A9B10CFFF94A274FF78FDE4FDF76B7C" author="Pilsbry, H. A." box="[151,539,608,630]" page="369" pageId="30" pageNumber="385" year="1921">
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<taxonomicName id="4C3A4D5DFF94A274FF78FDE4FDF76B7C" ID-CoL="6LBQH" authority="Pilsbry, 1921: 369" authorityName="Pilsbry" authorityPageNumber="369" authorityYear="1921" box="[151,539,608,630]" class="Gastropoda" family="Haminoeidae" genus="Haminoea" kingdom="Animalia" order="Cephalaspidea" pageId="30" pageNumber="385" phylum="Mollusca" rank="species" species="olopana">
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<emphasis id="B94EEACCFF94A274FF78FDE4FEB56B7C" box="[151,345,608,630]" italics="true" pageId="30" pageNumber="385">Haminoea olopana</emphasis>
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<bibRefCitation id="EFAB4B2FFF94A274FE8FFDE5FDF76B7C" author="Pilsbry, H. A." box="[352,539,608,630]" pageId="30" pageNumber="385" pagination="360 - 382" refId="ref20603" refString="Pilsbry, H. A. (1921) Marine mollusks of Hawaii: XIV, XV. Proceedings of the Academy of Natural Sciences of Philadelphia, 72 (3), 360 - 382." type="journal article" year="1921">Pilsbry, 1921: 369</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, fig. 8b.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B8536DEFF94A274FF78FDFBFAC66B9E" blockId="30.[151,1322,546,660]" box="[151,1322,638,660]" pageId="30" pageNumber="385">
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<treatmentCitationGroup id="AB2A11F0FF94A274FF78FDFBFAC66B9E" box="[151,1322,638,660]" pageId="30" pageNumber="385">
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<taxonomicName id="4C3A4D5DFF94A274FF78FDFBFEAC6B9E" baseAuthorityName="Gould" baseAuthorityYear="1859" box="[151,320,638,660]" class="Malacostraca" family="Atyidae" genus="Liloa" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="385" phylum="Arthropoda" rank="species" species="porcellana">
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<emphasis id="B94EEACCFF94A274FF78FDFBFEAC6B9E" box="[151,320,638,660]" italics="true" pageId="30" pageNumber="385">Liloa porcellana</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="0A9B10CFFF94A274FEB6FDFBFE106B9E" author="Habe, T." box="[345,508,638,660]" page="151" pageId="30" pageNumber="385" year="1952">
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<bibRefCitation id="EFAB4B2FFF94A274FEB6FDFBFE106B9E" author="Habe, T." box="[345,508,638,660]" pageId="30" pageNumber="385" pagination="137 - 152" refId="ref18981" refString="Habe, T. (1952) Atyidae in Japan. In: Kuroda, T., (Ed.), Illustrated Catalogue of Japanese shells, 20, pp. 137 - 152." type="journal article" year="1952">Habe 1952: 151</bibRefCitation>
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</treatmentCitation>
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, pl. 21, figs 20, 22, 25.
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<treatmentCitation id="0A9B10CFFF94A274FD04FDFBFC916B9E" author="Lin, G. Y." box="[747,893,638,660]" page="106" pageId="30" pageNumber="385" year="1997">
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<bibRefCitation id="EFAB4B2FFF94A274FD04FDFBFC916B9E" author="Lin, G. Y." box="[747,893,638,660]" pageId="30" pageNumber="385" refId="ref19471" refString="Lin, G. Y. (1997) Fauna Sinica: Phylum Mollusca, Class Gastropoda, Sub-Class Opisthobranchia, Order Cephalaspidea. Science Presss, Beijing, 246 pp." type="book" year="1997">Lin 1997: 106</bibRefCitation>
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</treatmentCitation>
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, pl. 10, fig. 3.
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<treatmentCitation id="0A9B10CFFF94A274FBFFFDFBFB746B9E" author="Qi, Z. Y." box="[1040,1176,638,660]" page="147" pageId="30" pageNumber="385" year="2004">
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<bibRefCitation id="EFAB4B2FFF94A274FBFFFDFBFB746B9E" author="Qi, Z. Y." box="[1040,1176,638,660]" pageId="30" pageNumber="385" refId="ref20697" refString="Qi, Z. Y. (2004) Seashells of China. China Ocean Press, Beijing, 418 pp." type="book" year="2004">Qi 2004: 147</bibRefCitation>
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</treatmentCitation>
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, pl. 83, fig. O.
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</paragraph>
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<emphasis id="B94EEACCFF94A274FF78FD44FED96BD0" bold="true" box="[151,309,705,730]" inLineHeading="true" pageId="30" pageNumber="385" reason="1">
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<typeStatus id="5481887CFF94A274FF78FD44FF3E6BD0" box="[151,210,705,730]" pageId="30" pageNumber="385">Type</typeStatus>
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locality.
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</emphasis>
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<collectingRegion id="49FEF83CFF94A274FED3FD44FE506BD0" box="[316,444,705,730]" country="Japan" name="Kagoshima" pageId="30" pageNumber="385">Kagoshima</collectingRegion>
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<location id="8EE56005FF94A274FE2CFD47FE1D6BD0" LSID="urn:lsid:plazi:treatment:039387C8FF94A275FF78FE59FEBB6B12:8EE56005FF94A274FE2CFD47FE1D6BD0" box="[451,497,706,730]" country="Japan" name="Bay" pageId="30" pageNumber="385" stateProvince="Kagoshima">Bay</location>
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,
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<collectingCountry id="F32D764EFF94A274FE13FD47FDD36BD3" box="[508,575,706,729]" name="Japan" pageId="30" pageNumber="385">Japan</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B8536DEFF94A274FF28FD60FE3E6A63" blockId="30.[151,1437,705,2026]" pageId="30" pageNumber="385">
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<emphasis id="B94EEACCFF94A274FF28FD60FE586BF4" bold="true" box="[199,436,741,766]" inLineHeading="true" pageId="30" pageNumber="385" reason="1">Material examined.</emphasis>
|
||
<materialsCitation id="3B523C83FF94A274FE50FD60FADC6BF7" ID-GBIF-Occurrence="3319782303" box="[447,1328,741,766]" collectionCode="USNM" country="Japan" location="Kagoshima" pageId="30" pageNumber="385" specimenCode="USNM 1357" specimenCount="1" stateProvince="Kagoshima" typeStatus="holotype">
|
||
<collectingRegion id="49FEF83CFF94A274FE50FD60FDAE6BF4" box="[447,578,741,766]" country="Japan" name="Kagoshima" pageId="30" pageNumber="385">Kagoshima</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32D764EFF94A274FDBEFD63FD786BF7" box="[593,660,742,765]" name="Japan" pageId="30" pageNumber="385">Japan</collectingCountry>
|
||
, 1 shell examined (
|
||
<typeStatus id="5481887CFF94A274FC94FD60FC0F6BF4" box="[891,995,741,766]" pageId="30" pageNumber="385" type="holotype">holotype</typeStatus>
|
||
),
|
||
<specimenCode id="DB9C9EA5FF94A274FC15FD60FB7D6BF4" box="[1018,1169,741,766]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="30" pageNumber="385" type="Museum">USNM 1357</specimenCode>
|
||
, H =
|
||
<quantity id="4CC29B3BFF94A274FB35FD60FADC6BF7" box="[1242,1328,741,766]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="30" pageNumber="385" unit="mm" value="12.0">12 mm</quantity>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B523C83FF94A274FAD0FD60FC956A28" ID-GBIF-Occurrence="3319782314" accessionNumber="UF 299895" collectionCode="UF" country="Northern Mariana Islands" location="Mariana Islands" pageId="30" pageNumber="385" specimenCount="1" stateProvince="Kagoshima" typeStatus="holotype">
|
||
<collectingCountry id="F32D764EFF94A274FAD0FD60FF056A28" name="Northern Mariana Islands" pageId="30" pageNumber="385">Mariana Islands</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32D764EFF94A274FF1BFC8CFED76A28" box="[244,315,777,802]" name="Guam" pageId="30" pageNumber="385">Guam</collectingCountry>
|
||
, 3 spcs dissected,
|
||
<collectionCode id="ED2BAE1BFF94A274FDE6FC8FFDC16A2B" box="[521,557,778,801]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="30" pageNumber="385" type="Museum">UF</collectionCode>
|
||
299895, H =
|
||
<quantity id="4CC29B3BFF94A274FD27FC8CFCA06A2B" box="[712,844,777,802]" metricMagnitude="-3" metricUnit="m" metricValue="9.85" metricValueMax="10.0" metricValueMin="9.7" pageId="30" pageNumber="385" unit="mm" value="9.85" valueMax="10.0" valueMin="9.7">9.7–10 mm</quantity>
|
||
; the
|
||
</materialsCitation>
|
||
<materialsCitation id="3B523C83FF94A274FC6EFC8CFE226A63" ID-GBIF-Occurrence="3319782302" collectionCode="MNHN" country="Philippines" location="Paris" pageId="30" pageNumber="385" specimenCode="OT926, ZMBN 89712" specimenCount="1" stateProvince="Kagoshima" typeStatus="holotype">
|
||
<collectingCountry id="F32D764EFF94A274FC6EFC8CFBEE6A28" box="[897,1026,777,802]" name="Philippines" pageId="30" pageNumber="385">Philippines</collectingCountry>
|
||
, 1 spc. dissected,
|
||
<collectionCode id="ED2BAE1BFF94A274FB25FC8FFACE6A2B" box="[1226,1314,778,801]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="30" pageNumber="385">MNHN</collectionCode>
|
||
,
|
||
<location id="8EE56005FF94A274FAC1FC8CFA8A6A28" LSID="urn:lsid:plazi:treatment:039387C8FF94A275FF78FE59FEBB6B12:8EE56005FF94A274FAC1FC8CFA8A6A28" box="[1326,1382,777,802]" country="Philippines" name="Paris" pageId="30" pageNumber="385" stateProvince="Kagoshima">Paris</location>
|
||
(S6/
|
||
<specimenCode id="DB9C9EA5FF94A274FF78FCA8FF066A4C" box="[151,234,813,838]" collectionCode="OT" pageId="30" pageNumber="385">OT926</specimenCode>
|
||
), H =
|
||
<quantity id="4CC29B3BFF94A274FEC0FCA8FE646A4F" box="[303,392,813,838]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="30" pageNumber="385" unit="mm" value="7.8">7.8 mm</quantity>
|
||
. Maui, Hawaii, 1 spc dissected,
|
||
<specimenCode id="DB9C9EA5FF94A274FD1FFCABFC7C6A4C" box="[752,912,813,838]" collectionCode="ZMBN" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="30" pageNumber="385">ZMBN 89712</specimenCode>
|
||
, H =
|
||
<quantity id="4CC29B3BFF94A274FC22FCA8FBCA6A4F" box="[973,1062,813,838]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="30" pageNumber="385" unit="mm" value="4.2">4.2 mm</quantity>
|
||
. Maui, Hawaii, 1 shell examined,
|
||
<specimenCode id="DB9C9EA5FF94A274FF78FCD7FED46A60" box="[151,312,849,874]" collectionCode="ZMBN" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="30" pageNumber="385">ZMBN 89712</specimenCode>
|
||
, H =
|
||
<quantity id="4CC29B3BFF94A274FE9AFCD4FE226A63" box="[373,462,849,874]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="30" pageNumber="385" unit="mm" value="2.8">2.8 mm</quantity>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8536DEFF94A274FF28FCF0FE986AB8" blockId="30.[151,1437,705,2026]" pageId="30" pageNumber="385">
|
||
<emphasis id="B94EEACCFF94A274FF28FCF0FE2A6A84" bold="true" box="[199,454,885,910]" pageId="30" pageNumber="385">
|
||
Animal (
|
||
<figureCitation id="13012A5BFF94A274FEDFFCF0FE5B6A84" box="[304,439,885,910]" captionStart="FIGURE 1" captionStartId="3.[151,250,1820,1842]" captionTargetBox="[218,1352,193,1797]" captionTargetId="figure-175@3.[218,1352,193,1797]" captionTargetPageId="3" captionText="FIGURE 1. Live images. A. Aliculastrum cylindricum (Koumac, northern New Caledonia; H = 28 mm, MNHN, Paris), B. Aliculastrum debilis (Maui, Hawaii; ZMBN 81658; H = 9 mm), C. Aliculastrum parallelum (Guam; UF 374138; H = 19 mm), D. Atys multistriatus (Guam; UF 374136; H = 7 mm), E. Atys pittmani nov. sp. (Maui, Hawaii; ZMBN 81673; H = 10 mm), F. Atys semistriatus (Maui, Hawaii; ZMBN 81656; H = 6 mm), G. Atys ukulele nov. sp. (Maui, Hawaii; ZMBN 89710; H = 5.4 mm), H. Diniatys dentifer (Guam; UF 374130; H = 9 mm), I. Diniatys dubia (Guam; UF 299907; H = 12.7 mm), J. Liloa curta (Guam; UF 374131; H = 15 mm), K. Liloa porcellana (Maui, Hawaii; ZMBN 89712; H = 4.2 mm); L. Liloa porcellana (Maui, Hawaii; ZMBN 89712; H = 2.8 mm). Image A, courtesy of W. B. Rudman; Images B, E–G, K, L, courtesy of C. Pittman." figureDoi="http://doi.org/10.5281/zenodo.4914929" httpUri="https://zenodo.org/record/4914929/files/figure.png" pageId="30" pageNumber="385">Figs 1K, L</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Body whitish-translucent, white dots scattered over the body, more dense at cephalic shield; eyes visible.
|
||
</paragraph>
|
||
<paragraph id="8B8536DEFF94A274FF28FC38FDF36D48" blockId="30.[151,1437,705,2026]" pageId="30" pageNumber="385">
|
||
<emphasis id="B94EEACCFF94A274FF28FC38FE096ADC" bold="true" box="[199,485,957,982]" pageId="30" pageNumber="385">
|
||
Shell (
|
||
<figureCitation id="13012A5BFF94A274FEFAFC38FE7D6ADC" box="[277,401,957,982]" captionStart="FIGURE 3" captionStartId="5.[151,250,1881,1903]" captionTargetBox="[171,1407,193,1851]" captionTargetId="figure-110@5.[171,1407,193,1851]" captionTargetPageId="5" captionText="FIGURE 3. Shells. A. Atys ukulele nov. sp. (Maui, Hawaii; ZMBN 89711; H = 4.6 mm), B. Diniatys dentifer (the Philippines, MNHN, Paris (B17/ OT793); H = 3.5 mm), C. Diniatys dubia (Guam; UF 299907; H = 7.5 mm), D. Liloa curta (the Philippines; MNHN, Paris (S12/ OT575); H = 3.1 mm), E. Liloa porcellana (Guam; UF 299895; H = 6.7 mm), F. Liloa porcellana (Maui, Hawaii; ZMBN 89712; H = 4.2 mm), G. Liloa porcellana (Maui, Hawaii; ZMBN 89712; H = 2.8 mm). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4914937" httpUri="https://zenodo.org/record/4914937/files/figure.png" pageId="30" pageNumber="385">Fig. 3E–G</figureCitation>
|
||
,
|
||
<figureCitation id="13012A5BFF94A274FE4DFC38FE396ADC" box="[418,469,957,982]" captionStart="FIGURE 18" captionStartId="31.[151,250,1890,1912]" captionTargetBox="[197,1389,599,1867]" captionTargetId="figure-302@31.[191,1396,591,1868]" captionTargetPageId="31" captionText="FIGURE 18. Shell and anatomy of Liloa porcellana. A. Detail of middle part of shell. B. Detail of jaw. C. Front view of anterior part of radula. D. Detail of median tooth at the middle part of radula. E. Denticulation on inner lateral tooth at right side of radula. F. Detail of outer lateral teeth at left side of radula. Scale bars: A = 100 µm; B = 2 µm; C, F = 20 µm; D, E = 20 µm. A, B: UF 299895, H = 9.8 mm; C–F: UF 299895, H = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4914973" httpUri="https://zenodo.org/record/4914973/files/figure.png" pageId="30" pageNumber="385">18A</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Maximum height
|
||
<quantity id="4CC29B3BFF94A274FD50FC38FCF96ADF" box="[703,789,957,982]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="30" pageNumber="385" unit="mm" value="19.0">19 mm</quantity>
|
||
; whitish translucent; fragile, thin, elongated-cylindrical, both anterior and posterior ends truncated, anterior end narrower; spire sunken, outer lip thin; spiral grooves covering entire shell in adults (which become closer towards the ends) and only visible at shell ends in juveniles; faint axial lines present irregularly.
|
||
</paragraph>
|
||
<paragraph id="8B8536DEFF94A274FF28FBC8FBED6D6C" blockId="30.[151,1437,705,2026]" box="[199,1025,1101,1126]" pageId="30" pageNumber="385">
|
||
<emphasis id="B94EEACCFF94A274FF28FBC8FE656D6C" bold="true" box="[199,393,1101,1126]" pageId="30" pageNumber="385">
|
||
Jaws (
|
||
<figureCitation id="13012A5BFF94A274FEFEFBC8FE966D6C" box="[273,378,1101,1126]" captionStart="FIGURE 18" captionStartId="31.[151,250,1890,1912]" captionTargetBox="[197,1389,599,1867]" captionTargetId="figure-302@31.[191,1396,591,1868]" captionTargetPageId="31" captionText="FIGURE 18. Shell and anatomy of Liloa porcellana. A. Detail of middle part of shell. B. Detail of jaw. C. Front view of anterior part of radula. D. Detail of median tooth at the middle part of radula. E. Denticulation on inner lateral tooth at right side of radula. F. Detail of outer lateral teeth at left side of radula. Scale bars: A = 100 µm; B = 2 µm; C, F = 20 µm; D, E = 20 µm. A, B: UF 299895, H = 9.8 mm; C–F: UF 299895, H = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4914973" httpUri="https://zenodo.org/record/4914973/files/figure.png" pageId="30" pageNumber="385">Fig. 18B</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Present, crescent shape, jaw rodlets with 6-10 denticles.
|
||
</paragraph>
|
||
<paragraph id="8B8536DEFF94A274FF28FBF4FEE26C10" blockId="30.[151,1437,705,2026]" pageId="30" pageNumber="385">
|
||
<emphasis id="B94EEACCFF94A274FF28FBF4FE216D80" bold="true" box="[199,461,1137,1162]" pageId="30" pageNumber="385">
|
||
Radula (
|
||
<figureCitation id="13012A5BFF94A274FEC0FBF4FE536D80" box="[303,447,1137,1162]" captionStart="FIGURE 18" captionStartId="31.[151,250,1890,1912]" captionTargetBox="[197,1389,599,1867]" captionTargetId="figure-302@31.[191,1396,591,1868]" captionTargetPageId="31" captionText="FIGURE 18. Shell and anatomy of Liloa porcellana. A. Detail of middle part of shell. B. Detail of jaw. C. Front view of anterior part of radula. D. Detail of median tooth at the middle part of radula. E. Denticulation on inner lateral tooth at right side of radula. F. Detail of outer lateral teeth at left side of radula. Scale bars: A = 100 µm; B = 2 µm; C, F = 20 µm; D, E = 20 µm. A, B: UF 299895, H = 9.8 mm; C–F: UF 299895, H = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4914973" httpUri="https://zenodo.org/record/4914973/files/figure.png" pageId="30" pageNumber="385">Figs 18C–F</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Radular formula at mid-point 18–20 x 7–4.1.1.1.4–7; median tooth with broad base, triangular central cusp prominent, small triangular lateral cusp; inner lateral tooth hook-shaped, base with semicircular projection outwardly, groove along outer margin, weak denticulation along groove; outer lateral teeth hook-shaped, base with semi-circular projection outwardly, weak groove along outer margin, size decreases outwardly.
|
||
</paragraph>
|
||
<paragraph id="8B8536DEFF94A274FF28FAA0FD196C8C" blockId="30.[151,1437,705,2026]" pageId="30" pageNumber="385">
|
||
<emphasis id="B94EEACCFF94A274FF28FAA0FDF36C34" bold="true" box="[199,543,1317,1342]" pageId="30" pageNumber="385">
|
||
Gizzard plates (
|
||
<figureCitation id="13012A5BFF94A274FE6DFAA0FDFC6C34" box="[386,528,1317,1342]" captionStart="FIGURE 19" captionStartId="33.[151,250,858,880]" captionTargetBox="[151,1436,194,833]" captionTargetId="figure-486@33.[151,1436,193,837]" captionTargetPageId="33" captionText="FIGURE 19. Anatomy of Liloa porcellana. A. Top view of gizzard plate. B. Rods on middle ridges of gizzard plate. C. Male reproductive system. go, genital opening; (?) gd, (?) glandular duct; pr, prostate; ps, penial sheath; sd 1, first seminal duct; sd 2, second seminal duct. Scale bars: A = 100 µm; B = 20 µm; C = 1 mm. A–C: UF 299895, H = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4914975" httpUri="https://zenodo.org/record/4914975/files/figure.png" pageId="30" pageNumber="385">Figs 19A, B</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Three gizzard plates; almost equal size from anterior to posterior, 10–11 ridges, covered by rods with pointed tips along top edge of ridges, dense tiny rods with pointed tips on both anterior and posterior sides of ridges, gaps between ridges smooth.
|
||
</paragraph>
|
||
<paragraph id="8B8536DEFF94A274FF28FA14FCAB6F54" blockId="30.[151,1437,705,2026]" pageId="30" pageNumber="385">
|
||
<emphasis id="B94EEACCFF94A274FF28FA14FD676CA0" bold="true" box="[199,651,1425,1450]" pageId="30" pageNumber="385">
|
||
Male reproductive system (
|
||
<figureCitation id="13012A5BFF94A274FDFFFA14FD916CA0" box="[528,637,1425,1450]" captionStart="FIGURE 19" captionStartId="33.[151,250,858,880]" captionTargetBox="[151,1436,194,833]" captionTargetId="figure-486@33.[151,1436,193,837]" captionTargetPageId="33" captionText="FIGURE 19. Anatomy of Liloa porcellana. A. Top view of gizzard plate. B. Rods on middle ridges of gizzard plate. C. Male reproductive system. go, genital opening; (?) gd, (?) glandular duct; pr, prostate; ps, penial sheath; sd 1, first seminal duct; sd 2, second seminal duct. Scale bars: A = 100 µm; B = 20 µm; C = 1 mm. A–C: UF 299895, H = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4914975" httpUri="https://zenodo.org/record/4914975/files/figure.png" pageId="30" pageNumber="385">Fig. 19C</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Total length
|
||
<quantity id="4CC29B3BFF94A274FCCAFA14FC916CA3" box="[805,893,1425,1450]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="30" pageNumber="385" unit="mm" value="4.2">4.2 mm</quantity>
|
||
(H =
|
||
<quantity id="4CC29B3BFF94A274FC2AFA14FBF66CA0" box="[965,1050,1425,1450]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="385" unit="mm" value="10.0">10 mm</quantity>
|
||
). Formed by three parts: prostate, seminal ducts and penial region; prostate oval, bilobed, proximal lobe smaller, opaque-brownish, distal lobe lightyellowish; translucent duct connects prostate to creamy-yellowish striated (?)glandular region, which opens distally to first seminal duct that possesses hook-like posterior end, and proximally opens to second seminal duct, translucent, coils twice before connecting to penial region; penial region oval, translucent, with convoluted opaqueyellowish inner glandular duct connecting to genital opening.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3206555FF94A274FF28F9ECFD216FC0" pageId="30" pageNumber="385" type="biology_ecology">
|
||
<paragraph id="8B8536DEFF94A274FF28F9ECFC2B6F88" blockId="30.[151,1437,705,2026]" box="[199,967,1641,1666]" pageId="30" pageNumber="385">
|
||
<emphasis id="B94EEACCFF94A274FF28F9ECFEC76F88" bold="true" box="[199,299,1641,1666]" inLineHeading="true" pageId="30" pageNumber="385" reason="1">Ecology.</emphasis>
|
||
Sandy bottom below
|
||
<quantity id="4CC29B3BFF94A274FDCEF9ECFDB66F8B" box="[545,602,1641,1666]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="30" pageNumber="385" unit="m" value="29.0">29 m</quantity>
|
||
deep (
|
||
<bibRefCitation id="EFAB4B2FFF94A274FD4AF9ECFCFB6F88" author="Lin, G. Y." box="[677,791,1641,1666]" pageId="30" pageNumber="385" refId="ref19471" refString="Lin, G. Y. (1997) Fauna Sinica: Phylum Mollusca, Class Gastropoda, Sub-Class Opisthobranchia, Order Cephalaspidea. Science Presss, Beijing, 246 pp." type="book" year="1997">Lin, 1997</bibRefCitation>
|
||
; present study).
|
||
</paragraph>
|
||
<paragraph id="8B8536DEFF94A274FF28F908FD216FC0" blockId="30.[151,1437,705,2026]" pageId="30" pageNumber="385">
|
||
<emphasis id="B94EEACCFF94A274FF28F908FDE86FAC" bold="true" box="[199,516,1677,1702]" inLineHeading="true" pageId="30" pageNumber="385" reason="1">Geographical distribution.</emphasis>
|
||
<collectingCountry id="F32D764EFF94A274FDE4F90BFDA26FAF" box="[523,590,1678,1701]" name="Japan" pageId="30" pageNumber="385">Japan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32D764EFF94A274FDB6F908FD736FAC" box="[601,671,1677,1702]" name="China" pageId="30" pageNumber="385">China</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32D764EFF94A274FD46F908FD126FAC" box="[681,766,1677,1702]" name="Taiwan" pageId="30" pageNumber="385">Taiwan</collectingCountry>
|
||
, the
|
||
<collectingCountry id="F32D764EFF94A274FCDDF908FC5F6FAC" box="[818,947,1677,1702]" name="Philippines" pageId="30" pageNumber="385">Philippines</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32D764EFF94A274FC50F908FBEA6FAC" box="[959,1030,1677,1702]" name="Guam" pageId="30" pageNumber="385">Guam</collectingCountry>
|
||
, Big Island, Maui, Oahu and French Frigate Shoals of Hawaii (
|
||
<bibRefCitation id="EFAB4B2FFF94A274FE52F934FDF06FC0" author="Qi, Z. Y." box="[445,540,1713,1738]" pageId="30" pageNumber="385" refId="ref20697" refString="Qi, Z. Y. (2004) Seashells of China. China Ocean Press, Beijing, 418 pp." type="book" year="2004">Qi 2004</bibRefCitation>
|
||
; present study).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3206555FF94A275FF28F950FEBB6B12" lastPageId="31" lastPageNumber="386" pageId="30" pageNumber="385" type="discussion">
|
||
<paragraph id="8B8536DEFF94A275FF28F950FC1A69F2" blockId="30.[151,1437,705,2026]" lastBlockId="31.[151,1437,151,536]" lastPageId="31" lastPageNumber="386" pageId="30" pageNumber="385">
|
||
<emphasis id="B94EEACCFF94A274FF28F950FED76FE4" bold="true" box="[199,315,1749,1774]" inLineHeading="true" pageId="30" pageNumber="385" reason="1">Remarks.</emphasis>
|
||
There are a few conflicting features about the description of this species. Some authors referred to
|
||
<taxonomicName id="4C3A4D5DFF94A274FF78F97CFEBD6E1B" baseAuthorityName="Gould" baseAuthorityYear="1859" box="[151,337,1785,1809]" class="Malacostraca" family="Atyidae" genus="Liloa" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="385" phylum="Arthropoda" rank="species" species="porcellana">
|
||
<emphasis id="B94EEACCFF94A274FF78F97CFEBD6E1B" box="[151,337,1785,1809]" italics="true" pageId="30" pageNumber="385">Liloa porcellana</emphasis>
|
||
</taxonomicName>
|
||
with spiral grooves only at both ends of shell (
|
||
<bibRefCitation id="EFAB4B2FFF94A274FC8FF97CFC186E18" author="Pilsbry, H. A." box="[864,1012,1785,1810]" pageId="30" pageNumber="385" refId="ref20498" refString="Pilsbry, H. A. (1895) Manual of Conchology, structural and systematic. Vol. 15. Academy of Natural Sciences of Philadelphia, Philadelphia, 436 pp." type="book" year="1895">Pilsbry 1895</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAB4B2FFF94A274FC10F97CFB916E18" author="Habe, T." box="[1023,1149,1785,1810]" pageId="30" pageNumber="385" pagination="137 - 152" refId="ref18981" refString="Habe, T. (1952) Atyidae in Japan. In: Kuroda, T., (Ed.), Illustrated Catalogue of Japanese shells, 20, pp. 137 - 152." type="journal article" year="1952">Habe 1952</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAB4B2FFF94A274FB66F97CFB1F6E18" author="Lin, G. Y." box="[1161,1267,1785,1810]" pageId="30" pageNumber="385" refId="ref19471" refString="Lin, G. Y. (1997) Fauna Sinica: Phylum Mollusca, Class Gastropoda, Sub-Class Opisthobranchia, Order Cephalaspidea. Science Presss, Beijing, 246 pp." type="book" year="1997">Lin 1997</bibRefCitation>
|
||
). However, the original description states that spiral grooves are more visible at both ends, which does not imply that the middle part of the shell is smooth (
|
||
<bibRefCitation id="EFAB4B2FFF94A274FE28F8C4FDBD6E50" author="Gould, A. A." box="[455,593,1857,1882]" pageId="30" pageNumber="385" pagination="138 - 143" refId="ref18907" refString="Gould, A. A. (1859) Description of new species of shells brought home by the North Pacific Exploring Expedition. Proceedings of the Boston Society of Natural History, 7, 138 - 143." type="journal article" year="1859">Gould 1859</bibRefCitation>
|
||
). The study of the
|
||
<typeStatus id="5481887CFF94A274FCCDF8C4FC686E50" box="[802,900,1857,1882]" pageId="30" pageNumber="385" type="holotype">holotype</typeStatus>
|
||
(USNM 1357) confirmed the presence of spiral grooves throughout the shell but they are in fact faint in the middle part of the shell. Larger specimens that we examined (H ≥
|
||
<quantity id="4CC29B3BFF94A274FEBDF80CFE416EA8" box="[338,429,1929,1954]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="30" pageNumber="385" unit="mm" value="4.2">4.2 mm</quantity>
|
||
) have spiral grooves along the entire shell, with distance between grooves decreasing towards both ends; whereas the small specimen (H =
|
||
<quantity id="4CC29B3BFF94A274FD11F828FCBB6ECC" box="[766,855,1965,1990]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="30" pageNumber="385" unit="mm" value="2.8">2.8 mm</quantity>
|
||
) have spiral grooves only at both ends of the shell (
|
||
<figureCitation id="13012A5BFF94A274FF70F854FEF06EE0" box="[159,284,2001,2026]" captionStart="FIGURE 1" captionStartId="3.[151,250,1820,1842]" captionTargetBox="[218,1352,193,1797]" captionTargetId="figure-175@3.[218,1352,193,1797]" captionTargetPageId="3" captionText="FIGURE 1. Live images. A. Aliculastrum cylindricum (Koumac, northern New Caledonia; H = 28 mm, MNHN, Paris), B. Aliculastrum debilis (Maui, Hawaii; ZMBN 81658; H = 9 mm), C. Aliculastrum parallelum (Guam; UF 374138; H = 19 mm), D. Atys multistriatus (Guam; UF 374136; H = 7 mm), E. Atys pittmani nov. sp. (Maui, Hawaii; ZMBN 81673; H = 10 mm), F. Atys semistriatus (Maui, Hawaii; ZMBN 81656; H = 6 mm), G. Atys ukulele nov. sp. (Maui, Hawaii; ZMBN 89710; H = 5.4 mm), H. Diniatys dentifer (Guam; UF 374130; H = 9 mm), I. Diniatys dubia (Guam; UF 299907; H = 12.7 mm), J. Liloa curta (Guam; UF 374131; H = 15 mm), K. Liloa porcellana (Maui, Hawaii; ZMBN 89712; H = 4.2 mm); L. Liloa porcellana (Maui, Hawaii; ZMBN 89712; H = 2.8 mm). Image A, courtesy of W. B. Rudman; Images B, E–G, K, L, courtesy of C. Pittman." figureDoi="http://doi.org/10.5281/zenodo.4914929" httpUri="https://zenodo.org/record/4914929/files/figure.png" pageId="30" pageNumber="385">Figs 1K–L</figureCitation>
|
||
,
|
||
<figureCitation id="13012A5BFF94A274FEC5F854FE836EE0" box="[298,367,2001,2026]" captionStart="FIGURE 2" captionStartId="4.[151,250,1879,1901]" captionTargetBox="[193,1376,193,1837]" captionTargetId="figure-110@4.[193,1376,193,1837]" captionTargetPageId="4" captionText="FIGURE 2. Shells. A. Aliculastrum cylindricum (the Philippines; MNHN, Paris (S3/ OT236); H = 22 mm), B. Aliculastrum cylindricum (New Caledonia; MNHN, Paris; H = 13.7 mm), C. Aliculastrum debilis (the Philippines; MNHN, Paris (OT834); H = 4.5 mm), D. Aliculastrum parallelum (Guam; UF 374138; H = 10 mm), E. Atys multistriatus (Guam; UF 374136; H = 5.3 mm), F. Atys pittmani nov. sp. (Maui, Hawaii; ZMBN 81673; H = 10 mm); G. Atys semistriatus (Tahiti; ZMBN 87082; H = 4.1 mm). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4914933" httpUri="https://zenodo.org/record/4914933/files/figure.png" pageId="30" pageNumber="385">2E–G</figureCitation>
|
||
). The
|
||
<typeStatus id="5481887CFF94A274FE57F854FDF66EE0" box="[440,538,2001,2026]" pageId="30" pageNumber="385" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C3A4D5DFF94A274FDACF854FD126EE3" baseAuthorityName="Gould" baseAuthorityYear="1859" box="[579,766,2001,2025]" class="Malacostraca" family="Atyidae" genus="Liloa" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="385" phylum="Arthropoda" rank="species" species="porcellana">
|
||
<emphasis id="B94EEACCFF94A274FDACF854FD126EE3" box="[579,766,2001,2025]" italics="true" pageId="30" pageNumber="385">Liloa porcellana</emphasis>
|
||
</taxonomicName>
|
||
has a narrower posterior end compared to its anterior end. However, this was not exactly the case for the shells examined during this study, but differences in shell height may explain this phenotypic difference, since none of our specimens was as large as the
|
||
<typeStatus id="5481887CFF95A275FBB4FF39FB5169DF" box="[1115,1213,188,213]" pageId="31" pageNumber="386" type="holotype">holotype</typeStatus>
|
||
(H =
|
||
<quantity id="4CC29B3BFF95A275FAE9FF39FAB769DF" box="[1286,1371,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="31" pageNumber="386" unit="mm" value="12.0">12 mm</quantity>
|
||
). The anatomical features of our specimens match those described by
|
||
<bibRefCitation id="EFAB4B2FFF95A275FC8EFF5AFC1D69F2" author="Habe, T." box="[865,1009,223,248]" pageId="31" pageNumber="386" pagination="137 - 152" refId="ref18981" refString="Habe, T. (1952) Atyidae in Japan. In: Kuroda, T., (Ed.), Illustrated Catalogue of Japanese shells, 20, pp. 137 - 152." type="journal article" year="1952">Habe (1952)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8536DEFF95A275FF28FE81FEBB6B12" blockId="31.[151,1437,151,536]" pageId="31" pageNumber="386">
|
||
This species can be distinguished from
|
||
<taxonomicName id="4C3A4D5DFF95A275FD68FE81FCEA6816" baseAuthorityName="Adams" baseAuthorityYear="1850" box="[647,774,260,284]" class="Malacostraca" family="Atyidae" genus="Liloa" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="386" phylum="Arthropoda" rank="species" species="curta">
|
||
<emphasis id="B94EEACCFF95A275FD68FE81FCEA6816" box="[647,774,260,284]" italics="true" pageId="31" pageNumber="386">Liloa curta</emphasis>
|
||
</taxonomicName>
|
||
by a slender-elongated shell shape and spiral grooves that become closer towards the ends of the shell, although the latter feature may vary ontogenetically. Also,
|
||
<taxonomicName id="4C3A4D5DFF95A275FA8EFEADFEE3686E" baseAuthorityName="Gould" baseAuthorityYear="1859" class="Malacostraca" family="Atyidae" genus="Liloa" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="386" phylum="Arthropoda" rank="species" species="porcellana">
|
||
<emphasis id="B94EEACCFF95A275FA8EFEADFEE3686E" italics="true" pageId="31" pageNumber="386">Liloa porcellana</emphasis>
|
||
</taxonomicName>
|
||
only has spiral grooves in the mid-region of the shell in adult specimens; in juveniles, those can be faint or absent. Both species have similar radula. The gizzard plates of
|
||
<taxonomicName id="4C3A4D5DFF95A275FC93FEF5FBDA6882" baseAuthorityName="Gould" baseAuthorityYear="1859" box="[892,1078,368,392]" class="Malacostraca" family="Atyidae" genus="Liloa" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="386" phylum="Arthropoda" rank="species" species="porcellana">
|
||
<emphasis id="B94EEACCFF95A275FC93FEF5FBDA6882" box="[892,1078,368,392]" italics="true" pageId="31" pageNumber="386">Liloa porcellana</emphasis>
|
||
</taxonomicName>
|
||
have fewer and far apart ridges than
|
||
<taxonomicName id="4C3A4D5DFF95A275FF22FE11FEA568A6" baseAuthorityName="Adams" baseAuthorityYear="1850" box="[205,329,404,428]" class="Malacostraca" family="Atyidae" genus="Liloa" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="386" phylum="Arthropoda" rank="species" species="curta">
|
||
<emphasis id="B94EEACCFF95A275FF22FE11FEA568A6" box="[205,329,404,428]" italics="true" pageId="31" pageNumber="386">Liloa curta</emphasis>
|
||
</taxonomicName>
|
||
where the ridges are numerous and tight together. Moreover, the male reproductive system of
|
||
<taxonomicName id="4C3A4D5DFF95A275FA8EFE11FEE368DA" baseAuthorityName="Gould" baseAuthorityYear="1859" class="Malacostraca" family="Atyidae" genus="Liloa" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="386" phylum="Arthropoda" rank="species" species="porcellana">
|
||
<emphasis id="B94EEACCFF95A275FA8EFE11FEE368DA" italics="true" pageId="31" pageNumber="386">Liloa porcellana</emphasis>
|
||
</taxonomicName>
|
||
has a longer and twisted secondary seminal duct (
|
||
<figureCitation id="13012A5BFF95A275FCAFFE32FC4868DA" box="[832,932,439,464]" captionStart="FIGURE 19" captionStartId="33.[151,250,858,880]" captionTargetBox="[151,1436,194,833]" captionTargetId="figure-486@33.[151,1436,193,837]" captionTargetPageId="33" captionText="FIGURE 19. Anatomy of Liloa porcellana. A. Top view of gizzard plate. B. Rods on middle ridges of gizzard plate. C. Male reproductive system. go, genital opening; (?) gd, (?) glandular duct; pr, prostate; ps, penial sheath; sd 1, first seminal duct; sd 2, second seminal duct. Scale bars: A = 100 µm; B = 20 µm; C = 1 mm. A–C: UF 299895, H = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4914975" httpUri="https://zenodo.org/record/4914975/files/figure.png" pageId="31" pageNumber="386">Fig. 19C</figureCitation>
|
||
, sd 2) and a conspicuous duct connecting the prostate to a globose glandular region whereas in
|
||
<taxonomicName id="4C3A4D5DFF95A275FD01FE59FC9D68FE" baseAuthorityName="Adams" baseAuthorityYear="1850" box="[750,881,476,500]" class="Malacostraca" family="Atyidae" genus="Liloa" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="386" phylum="Arthropoda" rank="species" species="curta">
|
||
<emphasis id="B94EEACCFF95A275FD01FE59FC9D68FE" box="[750,881,476,500]" italics="true" pageId="31" pageNumber="386">Liloa curta</emphasis>
|
||
</taxonomicName>
|
||
the prostate almost sits on top of the globose glandular region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |