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<document id="19E137E2F819EB636D698123344D2B5E" ID-CLB-Dataset="2886" ID-DOI="10.1206/0003-0090.454.1.1" ID-GBIF-Dataset="3a7cdaa4-c2f2-4370-9301-c2ab339b2e0d" ID-ISSN="0003-0090" ID-Zenodo-Dep="5788835" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639747398861" checkinUser="felipe" docAuthor="Esselstyn, Jacob A., Achmadi, Anang S., Handika, Heru, Swanson, Mark T., Giarla, Thomas C. &amp; Rowe, Kevin C." docDate="2021" docId="038AB318014BE9594F89FAE8FCEFB0FF" docLanguage="en" docName="BulAmeMusNatHist.454.1-108.pdf" docOrigin="Bulletin of the American Museum of Natural History 2021 (454)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-454/issue-1/0003-0090.454.1.1/Fourteen-New-Endemic-Species-of-Shrew-Genus-Crocidura-from-Sulawesi/10.1206/0003-0090.454.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Crocidura ordinaria Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe 2021, new species" docType="treatment" docUuid="62BBDEF5-DF9F-45B8-BA4F-0CCDDB65B8AD" docUuidSource="ZooBank" docVersion="13" lastPageNumber="86" masterDocId="FFB3CB600119E90F4F33FF82FF9DB404" masterDocTitle="Fourteen New, Endemic Species Of Shrew (Genus Crocidura) From Sulawesi Reveal A Spectacular Island Radiation" masterLastPageNumber="109" masterPageNumber="1" pageNumber="82" updateTime="1699472789874" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7939AD5AE28F9605C39398FBE3AB7EBC">Fourteen New, Endemic Species Of Shrew (Genus Crocidura) From Sulawesi Reveal A Spectacular Island Radiation</mods:title>
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<mods:namePart id="A0FAD4191C2A31C39F99C5C0E46230B2">Esselstyn, Jacob A.</mods:namePart>
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<mods:namePart id="A467A0822EF758DA241EE70C3F8FECEA">Achmadi, Anang S.</mods:namePart>
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<mods:namePart id="D5B827EC2327FE677593DC30B642A7E8">Handika, Heru</mods:namePart>
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<mods:namePart id="CF15FE2B85A5352B01AF787E4F81DDF7">Swanson, Mark T.</mods:namePart>
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<mods:date id="B6CA3D2F02232B8AC6E2DF2B808FA064">2021</mods:date>
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<taxonomicName id="4C23798D014BE95D4F89FAE8FE02B186" authority="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe, 2021" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[186,415,1386,1410]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="82" pageNumber="82" phylum="Chordata" rank="species" species="ordinaria" status="sp. nov.">
<emphasis id="B957DE1C014BE95D4F89FAE8FE02B186" bold="true" box="[186,415,1386,1410]" italics="true" pageId="82" pageNumber="82">Crocidura ordinaria</emphasis>
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,
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<taxonomicNameLabel id="A2646367014BE95D4E9EFAE9FDB3B187" box="[429,558,1387,1411]" pageId="82" pageNumber="82" rank="species">new species</taxonomicNameLabel>
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LSID:
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urn:lsid:zoobank. org:act:
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<typeStatus id="5498BCAC014BE95D4FBFF988FE9EB226" box="[140,259,1546,1570]" pageId="82" pageNumber="82" type="holotype">HOLOTYPE</typeStatus>
:
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43011 (=
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<collectionCode id="ED329ACB014BE95D4EF7F989FD89B226" box="[452,532,1547,1570]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="82" pageNumber="82" type="Museum">FMNH</collectionCode>
218726
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), an adult male, collected on
<date id="FF9D24CE014BE95D4E96F9A9FDB5B247" box="[421,552,1579,1604]" pageId="82" pageNumber="82" value="2012-05-05">
<collectingDate id="EFD9DD26014BE95D4E96F9A9FDB5B247" box="[421,552,1579,1604]" pageId="82" pageNumber="82" value="2012-05-05">5 May 2012</collectingDate>
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by
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specimen comprises a skull (
<figureCitation id="13181E8B014BE95D4D6BF9CEFF07B281" captionStart="FIG" captionStartId="85.[104,146,1438,1462]" captionTargetBox="[130,1585,465,1409]" captionTargetId="figure-10@85.[130,1585,465,1409]" captionTargetPageId="85" captionText="FIG. 41. Images showing dorsal, ventral, and lateral views of the skull and lateral and occlusal views of the dentary of three species belonging to the Ordinary Group: A, Crocidura normalis, FMNH 213457; B, C. ordinaria, FMNH 218726; and C, C. solita, FMNH 213044." figureDoi="http://doi.org/10.5281/zenodo.5788907" httpUri="https://zenodo.org/record/5788907/files/figure.png" pageId="82" pageNumber="82">fig. 41B</figureCitation>
), formalin-fixed body, and frozen tissues.
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measurements from the
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are
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×
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×
<quantity id="4CDBAFEB014BE95D4E65F92DFE38B2C3" box="[342,421,1711,1735]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="82" pageNumber="82" unit="mm" value="16.0">16 mm</quantity>
×
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= 11.0 g.
<collectorName id="26D667D8014BE95D4D9DFD70FD4AB70E" box="[686,727,754,778]" pageId="82" pageNumber="82">The</collectorName>
voucher specimen and a tissue sample will be permanently curated at
<collectionCode id="ED329ACB014BE95D4CD7FC96FB80B72F" box="[996,1053,788,811]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="82" pageNumber="82" type="Museum">MZB</collectionCode>
and an additional tissue sample will be retained at
<collectionCode id="ED329ACB014BE95D4B64FCB7FB37B748" box="[1111,1194,821,844]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="82" pageNumber="82" type="Museum">FMNH</collectionCode>
.
</paragraph>
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<collectionCode id="ED329ACB014BE95D4DFFFCD4FCE3B769" box="[716,894,853,877]" pageId="82" pageNumber="82">TYPE LOCALITY</collectionCode>
:
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,
<collectingRegion id="49E7CCEC014BE95D4B23FCD7FB25B769" box="[1040,1208,853,877]" country="Indonesia" name="Sulawesi Barat" pageId="82" pageNumber="82">Sulawesi Barat</collectingRegion>
,
<collectingCounty id="62FD7A82014BE95D4D9FFCF5FC93B78A" box="[684,782,887,910]" pageId="82" pageNumber="82">Mamasa</collectingCounty>
,
<collectingMunicipality id="6BF89874014BE95D4C17FCF5FC1BB78A" box="[804,902,887,910]" pageId="82" pageNumber="82">Mamasa</collectingMunicipality>
,
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,
<location id="8EFC54D5014BE95D4B43FCF5FC9FB7AB" LSID="urn:lsid:plazi:treatment:038AB318014BE9594F89FAE8FCEFB0FF:8EFC54D5014BE95D4B43FCF5FC9FB7AB" country="Indonesia" county="Mamasa" municipality="Mamasa" name="Rantepangko" pageId="82" pageNumber="82" stateProvince="Sulawesi Barat">Rantepangko</location>
,
<location id="8EFC54D5014BE95D4C29FC1AFBB4B7AB" LSID="urn:lsid:plazi:treatment:038AB318014BE9594F89FAE8FCEFB0FF:8EFC54D5014BE95D4C29FC1AFBB4B7AB" box="[794,1065,919,943]" country="Indonesia" county="Mamasa" municipality="Mamasa" name="Mt. Gandang Dewata" pageId="82" pageNumber="82" stateProvince="Sulawesi Barat">Mt. Gandang Dewata</location>
;
<geoCoordinate id="EE1764C9014BE95D4B73FC1AFB2AB7B4" box="[1088,1207,920,944]" degrees="2.8181" direction="south" orientation="latitude" pageId="82" pageNumber="82" precision="5" value="-2.8181">2.8181° S</geoCoordinate>
,
<geoCoordinate id="EE1764C9014BE95D4D9FFC3BFCB2B7D4" box="[684,815,953,977]" degrees="119.3823" direction="east" orientation="longitude" pageId="82" pageNumber="82" precision="5" value="119.3823">119.3823° E</geoCoordinate>
,
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elevation.
</paragraph>
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</subSubSection>
<subSubSection id="C3395185014BE95D4DFFFC58FBF0B050" pageId="82" pageNumber="82" type="etymology">
<paragraph id="8B9C020E014BE95D4DFFFC58FBF0B050" blockId="82.[684,1214,754,1737]" pageId="82" pageNumber="82">
ETYMOLOGY:
<emphasis id="B957DE1C014BE95D4C6CFC58FC54B7F6" box="[863,969,986,1010]" italics="true" pageId="82" pageNumber="82">Ordinaria</emphasis>
is Latin for “ordinary,” used in recognition that this is yet another species of shrew with no striking or unique phenotypic traits worthy of a descriptive name.
</paragraph>
</subSubSection>
<subSubSection id="C3395185014BE95D4DFFFBDCFB98B138" pageId="82" pageNumber="82" type="distribution">
<paragraph id="8B9C020E014BE95D4DFFFBDCFB98B138" blockId="82.[684,1214,754,1737]" pageId="82" pageNumber="82">
GEOGRAPHIC DISTRIBUTION:
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<emphasis id="B957DE1C014BE95D4B20FBDCFD7BB093" italics="true" pageId="82" pageNumber="82">Crocidura ordinaria</emphasis>
</taxonomicName>
is found in the west-central area of endemism (Mt. Gandang Dewata,
<collectingRegion id="49E7CCEC014BE95D4B24FB23FC91B0DD" country="Indonesia" name="Sulawesi Barat" pageId="82" pageNumber="82">West Sulawesi Province</collectingRegion>
and Mt. Torompupu,
<collectingRegion id="49E7CCEC014BE95D4B36FB43FC93B0FE" country="Indonesia" name="Sulawesi Tengah" pageId="82" pageNumber="82">Central Sulawesi Province</collectingRegion>
) (
<figureCitation id="13181E8B014BE95D4C16FB60FCF2B0FE" box="[805,879,1250,1274]" captionStart="FIG" captionStartId="83.[108,150,1052,1073]" captionTargetId="graphics-267@83.[275,1045,225,1023]" captionTargetPageId="83" captionText="FIG. 39. Map of Sulawesi showing localities sampled for shrews. Colored areas enclose localities with known records of members of the Ordinary Group. To maintain clarity of presentation, the widespread Crocidura nigripes is not included." figureDoi="http://doi.org/10.5281/zenodo.5789164" httpUri="https://zenodo.org/record/5789164/files/figure.png" pageId="82" pageNumber="82">fig. 39</figureCitation>
). The species spans an unusually broad elevational range from approximately
<quantity id="4CDBAFEB014BE95D4D9DFAA6FCD6B138" box="[686,843,1316,1340]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="2.6" metricValueMin="0.2" pageId="82" pageNumber="82" unit="m" value="1400.0" valueMax="2600.0" valueMin="200.0">200 to 2600 m</quantity>
(
<figureCitation id="13181E8B014BE95D4C68FAA6FC3CB138" box="[859,929,1316,1340]" captionStart="FIG" captionStartId="31.[104,146,1395,1419]" captionTargetBox="[104,1603,469,1356]" captionTargetId="graphics-9@31.[188,1530,466,1251]" captionTargetPageId="31" captionText="FIG. 13. Elevational records of all species of Crocidura known from Sulawesi. Each point represents a specimen. For specimens associated with a minimum and maximum elevation, we used the center of the given elevational range. Sample sizes are given above the x-axis. Species are grouped according to the species groups used in the text (Thick = Thick-Tailed Group)." figureDoi="http://doi.org/10.5281/zenodo.5788859" httpUri="https://zenodo.org/record/5788859/files/figure.png" pageId="82" pageNumber="82">fig. 13</figureCitation>
;
<tableCitation id="C6A137B5014BE95D4C9EFAA6FC65B138" box="[941,1016,1316,1340]" captionStart="TABLE 3" captionStartId="27.[611,691,226,250]" captionTargetBox="[117,1200,329,1675]" captionTargetPageId="27" captionText="TABLE 3 Elevational Ranges (m) and Species Richness of Crocidura on Mountains of Sulawesia" httpUri="http://table.plazi.org/id/DF5C52860102E9144D50FF60FBDEB51A" pageId="82" pageNumber="82" tableUuid="DF5C52860102E9144D50FF60FBDEB51A">table 3</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3395185014BE95C4DFFFAC7FDC7B206" lastPageId="83" lastPageNumber="83" pageId="82" pageNumber="82" type="diagnosis">
<paragraph id="8B9C020E014BE95C4DFFFAC7FDC7B206" blockId="82.[684,1214,754,1737]" lastBlockId="83.[108,638,1183,1736]" lastPageId="83" lastPageNumber="83" pageId="82" pageNumber="82">
DIAGNOSIS: A moderately sized shrew (
<tableCitation id="C6A137B5014BE95D4B4FFAC7FD21B17B" captionStart="TABLE 2" captionStartId="25.[610,690,226,250]" captionTargetBox="[119,1184,329,1682]" captionTargetPageId="25" captionText="TABLE 2 Descriptive Statisticsa for External Measurements (mm) and Mass (g) for Species of Sulawesi Crocidura" httpUri="http://table.plazi.org/id/DF5C52860100E9164D51FF60FBE5B523" pageId="82" pageNumber="82" tableUuid="DF5C52860100E9164D51FF60FBE5B523">tables 2</tableCitation>
,
<tableCitation id="C6A137B5014BE95D4DFBFAE5FD7EB17B" box="[712,739,1383,1407]" captionStart="TABLE 14" captionStartId="77.[603,683,226,250]" captionTargetBox="[119,1190,358,1139]" captionTargetPageId="77" captionText="TABLE 14 Descriptive Statisticsa for Craniodental Measurements (mm) for Species of the Ordinary Group of Sulawesi Crocidura" httpUri="http://table.plazi.org/id/DF5C52860154E9424D68FF60FCC5B540" pageId="82" pageNumber="82" tableUuid="DF5C52860154E9424D68FF60FCC5B540">14</tableCitation>
) with a somewhat stocky build. The pelage is medium to dark gray-brown and thick, with hairs at the middorsum typically
<quantity id="4CDBAFEB014BE95D4B13FA2AFBE3B1C4" box="[1056,1150,1448,1472]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="82" pageNumber="82" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">67 mm</quantity>
long. The venter is paler, with hairs that are pale gray at the tip, but dark gray at the base. In some specimens, the tips of some ventral hairs are reddish brown, giving the belly and chest pale cinnamon highlights. The mystacial vibrissae are short and darkly pigmented for at least half their lengths. Dorsally, the feet are nearly as dark as the pelage, but some specimens have much paler digits (
<figureCitation id="13181E8B014AE95C4F86FB1DFE90B0B3" box="[181,269,1183,1207]" captionStart="FIG" captionStartId="84.[104,146,1285,1309]" captionTargetBox="[104,1612,588,1258]" captionTargetId="figure-10@84.[104,1612,588,1258]" captionTargetPageId="84" captionText="FIG. 40. Images showing the ventral surface of the hind foot and dorsal surfaces of the tail base (approximately 1 cm from rump) and tail tip from three members of the Ordinary Group: A, Crocidura normalis, FMNH 213174 (left hind foot); B, C. ordinaria, FMNH 218726 (right hind foot); and C, C. solita, FMNH 213015 (left hind foot). Upper scale bar for foot, lower bar corresponds to both tail images." figureDoi="http://doi.org/10.5281/zenodo.5788905" httpUri="https://zenodo.org/record/5788905/files/figure.png" pageId="83" pageNumber="83">fig. 40B</figureCitation>
). The tail is shorter than head and body (
<figureCitation id="13181E8B014AE95C4F87FB42FF6DB0DC" box="[180,240,1216,1240]" captionStart="FIG" captionStartId="24.[108,150,1587,1608]" captionTargetBox="[137,1192,224,1561]" captionTargetId="graphics-9@24.[203,1191,224,1433]" captionTargetPageId="24" captionText="FIG. 9. Box plots showing variation in external measurements from all species of Sulawesi shrew. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are grouped according to the species groups used in the text (Thick = Thick- Tailed Group). All measurements in mm. HBL = head-and-body length." figureDoi="http://doi.org/10.5281/zenodo.5788853" httpUri="https://zenodo.org/record/5788853/files/figure.png" pageId="83" pageNumber="83">fig. 9</figureCitation>
) and the abundance of tail bristles and applied hairs is variable. The skull of this species is typical in its length (relative to body size) for a
<collectingRegion id="49E7CCEC014AE95C4FC3FAA1FED2B13F" box="[240,335,1315,1339]" country="Indonesia" name="Sulawesi Utara" pageId="83" pageNumber="83">Sulawesi</collectingRegion>
shrew but is wide at the braincase and interorbital region relative to skull length (
<figureCitation id="13181E8B014AE95C4FF1FAE7FE97B179" box="[194,266,1381,1405]" captionStart="FIG" captionStartId="26.[108,150,1414,1435]" captionTargetId="graphics-12@26.[213,1196,224,1265]" captionTargetPageId="26" captionText="FIG. 10. Box plots of relative skull measures showing braincase breadth (BB), interorbital width (IOW), and rostral length (RL) divided by condyloincisive length (CIL) and BB divided by IOW for all species of Sulawesi shrew. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are ordered according to the species groups used in the text (Thick = Thick-Tailed Group)." figureDoi="http://doi.org/10.5281/zenodo.5789149" httpUri="https://zenodo.org/record/5789149/files/figure.png" pageId="83" pageNumber="83">fig. 10</figureCitation>
;
<tableCitation id="C6A137B5014AE95C4E24FAE7FEECB179" box="[279,369,1381,1405]" captionStart="TABLE 14" captionStartId="77.[603,683,226,250]" captionTargetBox="[119,1190,358,1139]" captionTargetPageId="77" captionText="TABLE 14 Descriptive Statisticsa for Craniodental Measurements (mm) for Species of the Ordinary Group of Sulawesi Crocidura" httpUri="http://table.plazi.org/id/DF5C52860154E9424D68FF60FCC5B540" pageId="83" pageNumber="83" tableUuid="DF5C52860154E9424D68FF60FCC5B540">table 14</tableCitation>
). The braincase, though broad, is not especially inflated vertically. The dentition is more prominent than expected given the palatal width (
<figureCitation id="13181E8B014AE95C4E09FA4AFE08B1E5" box="[314,405,1480,1505]" captionStart="FIG" captionStartId="85.[104,146,1438,1462]" captionTargetBox="[130,1585,465,1409]" captionTargetId="figure-10@85.[130,1585,465,1409]" captionTargetPageId="85" captionText="FIG. 41. Images showing dorsal, ventral, and lateral views of the skull and lateral and occlusal views of the dentary of three species belonging to the Ordinary Group: A, Crocidura normalis, FMNH 213457; B, C. ordinaria, FMNH 218726; and C, C. solita, FMNH 213044." figureDoi="http://doi.org/10.5281/zenodo.5788907" httpUri="https://zenodo.org/record/5788907/files/figure.png" pageId="83" pageNumber="83">fig. 41B</figureCitation>
), but, otherwise, the skull of
<taxonomicName id="4C23798D014AE95C4FF6FA68FE00B206" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[197,413,1514,1538]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014AE95C4FF6FA68FE00B206" box="[197,413,1514,1538]" italics="true" pageId="83" pageNumber="83">Crocidura ordinaria</emphasis>
</taxonomicName>
is unremarkable.
</paragraph>
</subSubSection>
<paragraph id="8B9C020E014AE95C4E83FF7FFC7EB508" blockId="83.[300,997,253,997]" box="[432,995,253,268]" pageId="83" pageNumber="83">
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</paragraph>
<paragraph id="8B9C020E014AE95C4E1FFEBCFEC9B549" blockId="83.[300,997,253,997]" box="[300,340,318,333]" pageId="83" pageNumber="83">
<geoCoordinate id="EE1764C9014AE95C4E1FFEBCFEC9B549" box="[300,340,318,333]" degrees="1.5" direction="north" orientation="latitude" pageId="83" pageNumber="83" precision="5555" value="1.5">1.5°N</geoCoordinate>
</paragraph>
<paragraph id="8B9C020E014AE95C4E1FFE40FEC9B650" blockId="83.[300,997,253,997]" pageId="83" pageNumber="83">
<emphasis id="B957DE1C014AE95C4C19FE52FCEAB637" italics="true" pageId="83" pageNumber="83">
<taxonomicName id="4C23798D014AE95C4C19FE52FC0DB5E0" box="[810,912,464,484]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="musseri">C. musseri</taxonomicName>
<taxonomicName id="4C23798D014AE95C4C19FE69FC0BB604" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[810,918,491,512]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="normalis">C. normalis</taxonomicName>
<taxonomicName id="4C23798D014AE95C4C19FD84FC05B61E" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[810,920,518,538]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="ordinaria">C. ordinaria</taxonomicName>
<taxonomicName id="4C23798D014AE95C4C19FD9CFCEAB637" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[810,887,542,563]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="solita">C. solita</taxonomicName>
</emphasis>
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</paragraph>
<paragraph id="8B9C020E014AE95C4DD4FD32FCFAB6FF" blockId="83.[300,997,253,997]" pageId="83" pageNumber="83">
Recent sample sites
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Miller and Hollister (1921)
<typeStatus id="5498BCAC014AE95C4DD4FD65FC92B6FF" box="[743,783,743,763]" pageId="83" pageNumber="83">type</typeStatus>
localities
</paragraph>
<paragraph id="8B9C020E014AE95C4C7CFCB4FECEB753" blockId="83.[300,997,253,997]" pageId="83" pageNumber="83">
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<geoCoordinate id="EE1764C9014AE95C4E1FFCCAFECEB753" box="[300,339,840,855]" degrees="4.5" direction="south" orientation="latitude" pageId="83" pageNumber="83" precision="5555" value="-4.5">4.5°S</geoCoordinate>
</paragraph>
<paragraph id="8B9C020E014AE95C4C51FC07FC2BB7E1" blockId="83.[300,997,253,997]" pageId="83" pageNumber="83">
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<quantity id="4CDBAFEB014AE95C4C51FC28FC78B7BA" box="[866,997,938,958]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="83" pageNumber="83" unit="m" value="1500.0" valueMax="2000.0" valueMin="1000.0">10002000 m</quantity>
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&gt;
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</paragraph>
<caption id="DF5C5286014AE95C4F5FFB9EFEC2B068" ID-DOI="http://doi.org/10.5281/zenodo.5789164" ID-Zenodo-Dep="5789164" httpUri="https://zenodo.org/record/5789164/files/figure.png" pageId="83" pageNumber="83" startId="83.[108,150,1052,1073]" targetBox="[300,997,253,997]" targetPageId="83">
<paragraph id="8B9C020E014AE95C4F5FFB9EFEC2B068" blockId="83.[108,1212,1050,1132]" pageId="83" pageNumber="83">
FIG. 39. Map of Sulawesi showing localities sampled for shrews. Colored areas enclose localities with known records of members of the Ordinary Group. To maintain clarity of presentation, the widespread
<taxonomicName id="4C23798D014AE95C4B6BFBB5FF24B068" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="nigripes">
<emphasis id="B957DE1C014AE95C4B6BFBB5FF24B068" italics="true" pageId="83" pageNumber="83">Crocidura nigripes</emphasis>
</taxonomicName>
is not included.
</paragraph>
</caption>
<subSubSection id="C3395185014AE9594FBFF989FC3AB78B" lastPageId="86" lastPageNumber="86" pageId="83" pageNumber="83" type="description">
<paragraph id="8B9C020E014AE95C4FBFF989FB21B2CC" blockId="83.[108,638,1183,1736]" lastBlockId="83.[684,1212,1183,1736]" pageId="83" pageNumber="83">
COMPARISONS:
<taxonomicName id="4C23798D014AE95C4E79F989FDA6B227" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[330,571,1547,1571]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014AE95C4E79F989FDA6B227" box="[330,571,1547,1571]" italics="true" pageId="83" pageNumber="83">Crocidura ordinaria</emphasis>
</taxonomicName>
is a moderately sized member of the Ordinary Group. The tail is shorter than head-and-body length (
<figureCitation id="13181E8B014AE95C4FF4F9ECFE99B282" box="[199,260,1646,1670]" captionStart="FIG" captionStartId="24.[108,150,1587,1608]" captionTargetBox="[137,1192,224,1561]" captionTargetId="graphics-9@24.[203,1191,224,1433]" captionTargetPageId="24" captionText="FIG. 9. Box plots showing variation in external measurements from all species of Sulawesi shrew. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are grouped according to the species groups used in the text (Thick = Thick- Tailed Group). All measurements in mm. HBL = head-and-body length." figureDoi="http://doi.org/10.5281/zenodo.5788853" httpUri="https://zenodo.org/record/5788853/files/figure.png" pageId="83" pageNumber="83">fig. 9</figureCitation>
;
<tableCitation id="C6A137B5014AE95C4E20F9ECFEFEB282" box="[275,355,1646,1670]" captionStart="TABLE 2" captionStartId="25.[610,690,226,250]" captionTargetBox="[119,1184,329,1682]" captionTargetPageId="25" captionText="TABLE 2 Descriptive Statisticsa for External Measurements (mm) and Mass (g) for Species of Sulawesi Crocidura" httpUri="http://table.plazi.org/id/DF5C52860100E9164D51FF60FBE5B523" pageId="83" pageNumber="83" tableUuid="DF5C52860100E9164D51FF60FBE5B523">table 2</tableCitation>
). Members of the Long- Tailed Group are larger and have much longer tails. Rhoditis Group members
<taxonomicName id="4C23798D014AE95C4EE7F932FDD4B2CC" authorityName="Esselstyn et al., 2019:" authorityYear="1716" box="[468,585,1712,1736]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="rhoditis">
<emphasis id="B957DE1C014AE95C4EE7F932FDD4B2CC" box="[468,585,1712,1736]" italics="true" pageId="83" pageNumber="83">C. rhoditis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C23798D014AE95C4D9FFB1DFCF2B0B3" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[684,879,1183,1207]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="pseudorhoditis">
<emphasis id="B957DE1C014AE95C4D9FFB1DFCF2B0B3" box="[684,879,1183,1207]" italics="true" pageId="83" pageNumber="83">C. pseudorhoditis</emphasis>
</taxonomicName>
are larger.
<taxonomicName id="4C23798D014AE95C4CC5FB1DFB21B0B3" box="[1014,1212,1183,1207]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="pallida">
<emphasis id="B957DE1C014AE95C4CC5FB1DFB21B0B3" box="[1014,1212,1183,1207]" italics="true" pageId="83" pageNumber="83">Crocidura pallida</emphasis>
</taxonomicName>
and
<taxonomicName id="4C23798D014AE95C4DEDFB42FCC1B0DC" box="[734,860,1216,1240]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="australis">
<emphasis id="B957DE1C014AE95C4DEDFB42FCC1B0DC" box="[734,860,1216,1240]" italics="true" pageId="83" pageNumber="83">C. australis</emphasis>
</taxonomicName>
, also members of the Rhoditis Group, are only slightly larger than
<taxonomicName id="4C23798D014AE95C4B7AFB63FD51B11E" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014AE95C4B7AFB63FD51B11E" italics="true" pageId="83" pageNumber="83">C. ordinaria</emphasis>
</taxonomicName>
in head-and-body length. Compared to
<taxonomicName id="4C23798D014AE95C4B97FA81FC88B13F" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014AE95C4B97FA81FC88B13F" italics="true" pageId="83" pageNumber="83">C. ordinaria</emphasis>
</taxonomicName>
,
<taxonomicName id="4C23798D014AE95C4C15FAA6FC0BB13F" box="[806,918,1315,1339]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="pallida">
<emphasis id="B957DE1C014AE95C4C15FAA6FC0BB13F" box="[806,918,1315,1339]" italics="true" pageId="83" pageNumber="83">C. pallida</emphasis>
</taxonomicName>
has paler feet dorsally, a narrower relative braincase breadth (BB/CIL) and a narrower relative interorbital width (IOW/CIL) whereas
<taxonomicName id="4C23798D014AE95C4CA7FA05FB8DB19A" box="[916,1040,1414,1438]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="australis">
<emphasis id="B957DE1C014AE95C4CA7FA05FB8DB19A" box="[916,1040,1414,1438]" italics="true" pageId="83" pageNumber="83">C. australis</emphasis>
</taxonomicName>
has a narrower relative interorbital width, but wider relative braincase breadth than
<taxonomicName id="4C23798D014AE95C4CF1FA4BFBD3B1E4" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[962,1102,1480,1504]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014AE95C4CF1FA4BFBD3B1E4" box="[962,1102,1480,1504]" italics="true" pageId="83" pageNumber="83">C. ordinaria</emphasis>
</taxonomicName>
(
<figureCitation id="13181E8B014AE95C4B52FA4AFB32B1E5" box="[1121,1199,1480,1505]" captionStart="FIG" captionStartId="26.[108,150,1414,1435]" captionTargetId="graphics-12@26.[213,1196,224,1265]" captionTargetPageId="26" captionText="FIG. 10. Box plots of relative skull measures showing braincase breadth (BB), interorbital width (IOW), and rostral length (RL) divided by condyloincisive length (CIL) and BB divided by IOW for all species of Sulawesi shrew. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are ordered according to the species groups used in the text (Thick = Thick-Tailed Group)." figureDoi="http://doi.org/10.5281/zenodo.5789149" httpUri="https://zenodo.org/record/5789149/files/figure.png" pageId="83" pageNumber="83">fig. 10</figureCitation>
). All members of the Small-Bodied Group are much smaller than
<taxonomicName id="4C23798D014AE95C4CAAF989FBB5B227" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[921,1064,1547,1571]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014AE95C4CAAF989FBB5B227" box="[921,1064,1547,1571]" italics="true" pageId="83" pageNumber="83">C. ordinaria</emphasis>
</taxonomicName>
. Within the Ordinary Group,
<taxonomicName id="4C23798D014AE95C4C5CF9AEFC7EB247" box="[879,995,1580,1603]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="nigripes">
<emphasis id="B957DE1C014AE95C4C5CF9AEFC7EB247" box="[879,995,1580,1603]" italics="true" pageId="83" pageNumber="83">C. nigripes</emphasis>
</taxonomicName>
has darker feet and a relatively much narrower interorbital region and braincase than
<taxonomicName id="4C23798D014AE95C4CBEF9ECFB8AB282" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[909,1047,1646,1670]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014AE95C4CBEF9ECFB8AB282" box="[909,1047,1646,1670]" italics="true" pageId="83" pageNumber="83">C. ordinaria</emphasis>
</taxonomicName>
. Compared to
<taxonomicName id="4C23798D014AE95C4D9FF90DFCA5B2A3" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[684,824,1679,1703]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014AE95C4D9FF90DFCA5B2A3" box="[684,824,1679,1703]" italics="true" pageId="83" pageNumber="83">C. ordinaria</emphasis>
</taxonomicName>
,
<taxonomicName id="4C23798D014AE95C4C7AF90DFC5CB2A3" box="[841,961,1679,1703]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="musseri">
<emphasis id="B957DE1C014AE95C4C7AF90DFC5CB2A3" box="[841,961,1679,1703]" italics="true" pageId="83" pageNumber="83">C. musseri</emphasis>
</taxonomicName>
has a thicker, darker pelage, and darker feet.
<taxonomicName id="4C23798D014AE95C4CF6F932FB3DB2CC" authorityName="Esselstyn &amp; Achmadi &amp; Handika &amp; Swanson &amp; Giarla &amp; Rowe" authorityYear="2021" box="[965,1184,1712,1736]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="83" pageNumber="83" phylum="Chordata" rank="species" species="normalis">
<emphasis id="B957DE1C014AE95C4CF6F932FB3DB2CC" box="[965,1184,1712,1736]" italics="true" pageId="83" pageNumber="83">Crocidura normalis</emphasis>
</taxonomicName>
is
</paragraph>
<caption id="DF5C5286014DE95B4F5BFA87FB46B15D" ID-DOI="http://doi.org/10.5281/zenodo.5788905" ID-Zenodo-Dep="5788905" httpUri="https://zenodo.org/record/5788905/files/figure.png" pageId="84" pageNumber="84" startId="84.[104,146,1285,1309]" targetBox="[104,1612,588,1258]" targetPageId="84">
<paragraph id="8B9C020E014DE95B4F5BFA87FB46B15D" blockId="84.[104,1612,1284,1369]" pageId="84" pageNumber="84">
FIG. 40. Images showing the ventral surface of the hind foot and dorsal surfaces of the tail base (approximately 1 cm from rump) and tail tip from three members of the Ordinary Group:
<emphasis id="B957DE1C014DE95B4ECDFAA3FD8BB13F" bold="true" box="[510,534,1313,1339]" pageId="84" pageNumber="84">A,</emphasis>
<taxonomicName id="4C23798D014DE95B4D2FFAA3FCA9B13F" authority=", FMNH" authorityName="FMNH" box="[540,820,1313,1339]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="84" pageNumber="84" phylum="Chordata" rank="species" species="normalis">
<emphasis id="B957DE1C014DE95B4D2FFAA3FD43B13F" box="[540,734,1313,1339]" italics="true" pageId="84" pageNumber="84">Crocidura normalis</emphasis>
, FMNH
</taxonomicName>
213174 (left hind foot);
<emphasis id="B957DE1C014DE95B4B1CFAA3FBD8B13F" bold="true" box="[1071,1093,1313,1339]" pageId="84" pageNumber="84">B,</emphasis>
<taxonomicName id="4C23798D014DE95B4B78FAA3FA81B13F" authority=", FMNH" authorityName="FMNH" box="[1099,1308,1313,1339]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="84" pageNumber="84" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014DE95B4B78FAA3FB5BB13F" box="[1099,1222,1313,1339]" italics="true" pageId="84" pageNumber="84">C. ordinaria</emphasis>
, FMNH
</taxonomicName>
218726 (right hind foot); and
<emphasis id="B957DE1C014DE95B4F5AFABDFF1DB15D" bold="true" box="[105,128,1343,1369]" pageId="84" pageNumber="84">C,</emphasis>
<taxonomicName id="4C23798D014DE95B4FB4FABDFEB2B15D" authority=", FMNH" authorityName="FMNH" box="[135,303,1343,1369]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="84" pageNumber="84" phylum="Chordata" rank="species" species="solita">
<emphasis id="B957DE1C014DE95B4FB4FABDFF44B15D" box="[135,217,1343,1369]" italics="true" pageId="84" pageNumber="84">C. solita</emphasis>
, FMNH
</taxonomicName>
213015 (left hind foot). Upper scale bar for foot, lower bar corresponds to both tail images.
</paragraph>
</caption>
<caption id="DF5C5286014CE95A4F5BFA1CFADAB1D0" ID-DOI="http://doi.org/10.5281/zenodo.5788907" ID-Zenodo-Dep="5788907" httpUri="https://zenodo.org/record/5788907/files/figure.png" pageId="85" pageNumber="85" startId="85.[104,146,1438,1462]" targetBox="[130,1585,465,1409]" targetPageId="85">
<paragraph id="8B9C020E014CE95A4F5BFA1CFADAB1D0" blockId="85.[104,1612,1437,1492]" pageId="85" pageNumber="85">
FIG. 41. Images showing dorsal, ventral, and lateral views of the skull and lateral and occlusal views of the dentary of three species belonging to the Ordinary Group:
<emphasis id="B957DE1C014CE95A4E2EFA38FEA8B1D0" bold="true" box="[285,309,1466,1492]" pageId="85" pageNumber="85">A,</emphasis>
<taxonomicName id="4C23798D014CE95A4E08FA38FDC8B1D0" authority=", FMNH" authorityName="FMNH" box="[315,597,1466,1492]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="85" pageNumber="85" phylum="Chordata" rank="species" species="normalis">
<emphasis id="B957DE1C014CE95A4E08FA38FE63B1D0" box="[315,510,1466,1492]" italics="true" pageId="85" pageNumber="85">Crocidura normalis</emphasis>
, FMNH
</taxonomicName>
213457;
<emphasis id="B957DE1C014CE95A4D80FA38FD54B1D0" bold="true" box="[691,713,1466,1492]" pageId="85" pageNumber="85">B,</emphasis>
<taxonomicName id="4C23798D014CE95A4DFCFA38FC3DB1D0" authority=", FMNH" authorityName="FMNH" box="[719,928,1466,1492]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="85" pageNumber="85" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014CE95A4DFCFA38FCD7B1D0" box="[719,842,1466,1492]" italics="true" pageId="85" pageNumber="85">C. ordinaria</emphasis>
, FMNH
</taxonomicName>
218726; and
<emphasis id="B957DE1C014CE95A4B18FA38FBDFB1D0" bold="true" box="[1067,1090,1466,1492]" pageId="85" pageNumber="85">C,</emphasis>
<taxonomicName id="4C23798D014CE95A4B7BFA38FB6DB1D0" authority=", FMNH" authorityName="FMNH" box="[1096,1264,1466,1492]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="85" pageNumber="85" phylum="Chordata" rank="species" species="solita">
<emphasis id="B957DE1C014CE95A4B7BFA38FB07B1D0" box="[1096,1178,1466,1492]" italics="true" pageId="85" pageNumber="85">C. solita</emphasis>
, FMNH
</taxonomicName>
213044.
</paragraph>
</caption>
<paragraph id="8B9C020E014FE9594D9FFF60FC3AB78B" blockId="86.[684,1213,226,1275]" pageId="86" pageNumber="86">
darker in color, smaller in body size, has more bristles on its darkly colored tail, and its skull is narrower with a shorter relative rostral length (RL/CIL) than
<taxonomicName id="4C23798D014FE9594C6CFEC7FC71B559" box="[863,1004,325,349]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="86" pageNumber="86" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014FE9594C6CFEC7FC71B559" box="[863,1004,325,349]" italics="true" pageId="86" pageNumber="86">C. ordinaria</emphasis>
</taxonomicName>
.
<taxonomicName id="4C23798D014FE9594CCDFEC7FB2BB559" box="[1022,1206,325,349]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="86" pageNumber="86" phylum="Chordata" rank="species" species="solita">
<emphasis id="B957DE1C014FE9594CCDFEC7FB2BB559" box="[1022,1206,325,349]" italics="true" pageId="86" pageNumber="86">Crocidura solita</emphasis>
</taxonomicName>
, another member of the Ordinary Group, is difficult to distinguish morphologically from
<taxonomicName id="4C23798D014FE9594B97FE05FC8BB5C4" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="86" pageNumber="86" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014FE9594B97FE05FC8BB5C4" italics="true" pageId="86" pageNumber="86">C. ordinaria</emphasis>
</taxonomicName>
. External measurements are nearly identical between these two species (
<figureCitation id="13181E8B014FE9594B74FE4BFB1FB5E5" box="[1095,1154,457,481]" captionStart="FIG" captionStartId="24.[108,150,1587,1608]" captionTargetBox="[137,1192,224,1561]" captionTargetId="graphics-9@24.[203,1191,224,1433]" captionTargetPageId="24" captionText="FIG. 9. Box plots showing variation in external measurements from all species of Sulawesi shrew. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are grouped according to the species groups used in the text (Thick = Thick- Tailed Group). All measurements in mm. HBL = head-and-body length." figureDoi="http://doi.org/10.5281/zenodo.5788853" httpUri="https://zenodo.org/record/5788853/files/figure.png" pageId="86" pageNumber="86">fig. 9</figureCitation>
), but
<taxonomicName id="4C23798D014FE9594D9FFE69FCABB606" box="[684,822,490,514]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="86" pageNumber="86" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014FE9594D9FFE69FCABB606" box="[684,822,490,514]" italics="true" pageId="86" pageNumber="86">C. ordinaria</emphasis>
</taxonomicName>
has a higher mass-to-length ratio (
<figureCitation id="13181E8B014FE9594D87FD89FC9FB627" box="[692,770,523,547]" captionStart="FIG" captionStartId="37.[107,149,993,1014]" captionTargetBox="[134,1190,224,965]" captionTargetId="graphics-291@37.[206,1189,224,845]" captionTargetPageId="37" captionText="FIG. 17. Box plots showing the length of the hind foot (HF) relative to the head-and-body length (HBL) and the ratio of mass to HBL. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are ordered along the x-axis according to the species groups used in the text (Thick = Thick-Tailed Group). Lengths are in mm and mass is in grams." figureDoi="http://doi.org/10.5281/zenodo.5788865" httpUri="https://zenodo.org/record/5788865/files/figure.png" pageId="86" pageNumber="86">fig. 17</figureCitation>
). The pelage and feet of
<taxonomicName id="4C23798D014FE9594B1CFD8EFB21B627" box="[1071,1212,523,547]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="86" pageNumber="86" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014FE9594B1CFD8EFB21B627" box="[1071,1212,523,547]" italics="true" pageId="86" pageNumber="86">C. ordinaria</emphasis>
</taxonomicName>
are darker and have a smaller hypothenar (
<figureCitation id="13181E8B014FE9594BA4FDAEFD51B661" captionStart="FIG" captionStartId="84.[104,146,1285,1309]" captionTargetBox="[104,1612,588,1258]" captionTargetId="figure-10@84.[104,1612,588,1258]" captionTargetPageId="84" captionText="FIG. 40. Images showing the ventral surface of the hind foot and dorsal surfaces of the tail base (approximately 1 cm from rump) and tail tip from three members of the Ordinary Group: A, Crocidura normalis, FMNH 213174 (left hind foot); B, C. ordinaria, FMNH 218726 (right hind foot); and C, C. solita, FMNH 213015 (left hind foot). Upper scale bar for foot, lower bar corresponds to both tail images." figureDoi="http://doi.org/10.5281/zenodo.5788905" httpUri="https://zenodo.org/record/5788905/files/figure.png" pageId="86" pageNumber="86">fig. 40</figureCitation>
), on average, than in
<taxonomicName id="4C23798D014FE9594CF7FDCCFBBDB661" box="[964,1056,589,613]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="86" pageNumber="86" phylum="Chordata" rank="species" species="solita">
<emphasis id="B957DE1C014FE9594CF7FDCCFBBDB661" box="[964,1056,589,613]" italics="true" pageId="86" pageNumber="86">C. solita</emphasis>
</taxonomicName>
. Cranially,
<taxonomicName id="4C23798D014FE9594B97FDCCFC89B682" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="86" pageNumber="86" phylum="Chordata" rank="species" species="ordinaria">
<emphasis id="B957DE1C014FE9594B97FDCCFC89B682" italics="true" pageId="86" pageNumber="86">C. ordinaria</emphasis>
</taxonomicName>
has a wider skull, observable in the absolute and relative breadths at the rostrum, interorbital region, and braincase (
<figureCitation id="13181E8B014FE9594B68FD32FB2AB6CD" box="[1115,1207,688,713]" captionStart="FIG" captionStartId="26.[108,150,1414,1435]" captionTargetId="graphics-12@26.[213,1196,224,1265]" captionTargetPageId="26" captionText="FIG. 10. Box plots of relative skull measures showing braincase breadth (BB), interorbital width (IOW), and rostral length (RL) divided by condyloincisive length (CIL) and BB divided by IOW for all species of Sulawesi shrew. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are ordered according to the species groups used in the text (Thick = Thick-Tailed Group)." figureDoi="http://doi.org/10.5281/zenodo.5789149" httpUri="https://zenodo.org/record/5789149/files/figure.png" pageId="86" pageNumber="86">figs. 10</figureCitation>
,
<figureCitation id="13181E8B014FE9594D9FFD50FD54B6EE" box="[684,713,722,746]" captionStart="FIG" captionStartId="86.[107,149,1242,1263]" captionTargetBox="[125,556,238,1216]" captionTargetPageId="86" captionText="FIG. 42. Box plots showing subtle differences in cranial measurements between Crocidura solita and C. ordinaria, two members of the Ordinary Group. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. All measurements in mm." figureDoi="http://doi.org/10.5281/zenodo.5788911" httpUri="https://zenodo.org/record/5788911/files/figure.png" pageId="86" pageNumber="86">42</figureCitation>
;
<tableCitation id="C6A137B5014FE9594DEAFD50FCA6B6EE" box="[729,827,722,746]" captionStart="TABLE 14" captionStartId="77.[603,683,226,250]" captionTargetBox="[119,1190,358,1139]" captionTargetPageId="77" captionText="TABLE 14 Descriptive Statisticsa for Craniodental Measurements (mm) for Species of the Ordinary Group of Sulawesi Crocidura" httpUri="http://table.plazi.org/id/DF5C52860154E9424D68FF60FCC5B540" pageId="86" pageNumber="86" tableUuid="DF5C52860154E9424D68FF60FCC5B540">table 14</tableCitation>
). Principal components analyses of external and cranial dimensions show that these two species overlap broadly in multivariate morphometric space, more so with external measurements than with cranial measurements (
<figureCitation id="13181E8B014FE9594D87FCF5FD63B78B" box="[692,766,887,911]" captionStart="FIG" captionStartId="87.[109,151,828,849]" captionTargetBox="[167,1154,227,794]" captionTargetId="graphics-435@87.[252,668,233,736]" captionTargetPageId="87" captionText="FIG. 43. Bivariate plots showing the first two axes from principal components analyses of A, five external and B, 12 cranial measurements from Crocidura solita and C. ordinaria, two members of the Ordinary Group. Loadings and variance explained are given in tables 17 and 18, respectively." figureDoi="http://doi.org/10.5281/zenodo.5788913" httpUri="https://zenodo.org/record/5788913/files/figure.png" pageId="86" pageNumber="86">fig. 43</figureCitation>
;
<tableCitation id="C6A137B5014FE9594C39FCF5FCF2B78B" box="[778,879,887,911]" captionStart="TABLE 17" captionStartId="88.[604,684,226,250]" captionTargetBox="[283,1007,329,900]" captionTargetPageId="88" captionText="TABLE 17 Results of Principal Components Analysis of Craniodental Measurements of Crocidura ordinaria and C. solita" pageId="86" pageNumber="86">tables 17</tableCitation>
,
<tableCitation id="C6A137B5014FE9594C4FFCF5FC07B78B" box="[892,922,887,911]" captionStart="TABLE 18" captionStartId="89.[603,683,226,250]" captionTargetBox="[308,981,329,642]" captionTargetPageId="89" captionText="TABLE 18 Results of Principal Components Analysis of External Measurements of Crocidura ordinaria and C. solita" pageId="86" pageNumber="86">18</tableCitation>
).
</paragraph>
</subSubSection>
<paragraph id="8B9C020E014FE9594F4EFDC9FF5FB668" blockId="86.[125,547,587,825]" box="[125,194,587,666]" pageId="86" pageNumber="86">
<collectionCode id="ED329ACB014FE9594F4EFDC9FF07B69E" box="[125,154,587,666]" pageId="86" pageNumber="86">WIDTH</collectionCode>
3
</paragraph>
<paragraph id="8B9C020E014FE9594F4EFD22FF5FB6F5" blockId="86.[125,547,587,825]" box="[125,195,666,785]" pageId="86" pageNumber="86">ROSTRAL 2.75 2.5</paragraph>
<paragraph id="8B9C020E014FE9594FAAFC9DFDBEB73D" blockId="86.[125,547,587,825]" box="[153,547,799,825]" pageId="86" pageNumber="86">
<geoCoordinate id="EE1764C9014FE9594FAAFC9DFEA5B73D" box="[153,312,799,825]" degrees="2.25" direction="north" orientation="latitude" pageId="86" pageNumber="86" precision="555" value="2.25">2.25 N</geoCoordinate>
= 16 N = 43
</paragraph>
<paragraph id="8B9C020E014FE9594F97FCDFFF5FB775" blockId="86.[125,556,861,1216]" box="[164,194,861,881]" pageId="86" pageNumber="86">5.7</paragraph>
<subSubSection id="C3395185014FE9594DFFFC1AFB10B7F6" pageId="86" pageNumber="86" type="discussion">
<paragraph id="8B9C020E014FE9594DFFFC1AFB10B7F6" blockId="86.[684,1213,226,1275]" pageId="86" pageNumber="86">
COMMENTS: See extensive comments detailing our decision to distinguish this species from
<taxonomicName id="4C23798D014FE9594D9EFC58FCC4B7F6" box="[685,857,986,1010]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="86" pageNumber="86" phylum="Chordata" rank="species" species="solita">
<emphasis id="B957DE1C014FE9594D9EFC58FCC4B7F6" box="[685,857,986,1010]" italics="true" pageId="86" pageNumber="86">Crocidura solita</emphasis>
</taxonomicName>
in the next species account.
</paragraph>
</subSubSection>
<paragraph id="8B9C020E014FE9594F97FC2AFF5FB7B8" blockId="86.[125,556,861,1216]" box="[164,194,936,956]" pageId="86" pageNumber="86">5.4</paragraph>
<paragraph id="8B9C020E014FE9594F4EFC3AFF07B7C2" blockId="86.[125,556,861,1216]" box="[125,154,952,966]" pageId="86" pageNumber="86">3</paragraph>
<paragraph id="8B9C020E014FE9594F4EFC47FF07B7DD" blockId="86.[125,556,861,1216]" box="[125,154,965,985]" pageId="86" pageNumber="86">M</paragraph>
<paragraph id="8B9C020E014FE9594F4EFC62FF59B002" blockId="86.[125,556,861,1216]" box="[125,196,992,1030]" pageId="86" pageNumber="86">TO 5.1</paragraph>
<paragraph id="8B9C020E014FE9594F4EFB8AFF07B012" blockId="86.[125,556,861,1216]" box="[125,154,1032,1046]" pageId="86" pageNumber="86">4</paragraph>
<subSubSection id="C3395185014FE9594DFFFC79FCEFB0FF" pageId="86" pageNumber="86" type="materials_examined">
<paragraph id="8B9C020E014FE9594DFFFC79FCEFB0FF" blockId="86.[684,1213,226,1275]" pageId="86" pageNumber="86">
SPECIMENS EXAMINED:
<materialsCitation id="3B4B0853014FE9594CE3FC7EFC44B0BD" ID-GBIF-Occurrence="3421079442" collectionCode="FMNH, MZB, NMV, Z" location="Mt. Gandang Dewata" pageId="86" pageNumber="86" specimenCode="FMNH 218727-218743, 218763-218767, 218769, 218771, 218772, MZB 34806, 34808, 34816, 34820, 34827, 34833, 34856, 34870, 34871, 34876, 34877, 34883, 43011, NMV Z21910, Z21911, Z21938" specimenCount="1">
<location id="8EFC54D5014FE9594CE3FC7EFB21B017" LSID="urn:lsid:plazi:treatment:038AB318014BE9594F89FAE8FCEFB0FF:8EFC54D5014FE9594CE3FC7EFB21B017" box="[976,1212,1019,1043]" name="Mt. Gandang Dewata" pageId="86" pageNumber="86">Mt. Gandang Dewata</location>
(
<specimenCode id="DB85AA75014FE9594D85FB9EFC7CB030" box="[694,993,1052,1076]" pageId="86" pageNumber="86">FMNH 218727218743</specimenCode>
,
<specimenCode id="DB85AA75014FE9594CCCFB9EFB24B030" box="[1023,1209,1052,1076]" pageId="86" pageNumber="86">218763218767</specimenCode>
,
<specimenCode id="DB85AA75014FE9594D9FFBBFFD62B051" box="[684,767,1085,1109]" pageId="86" pageNumber="86">218769</specimenCode>
,
<specimenCode id="DB85AA75014FE9594C3CFBBFFCFCB051" box="[783,865,1085,1109]" pageId="86" pageNumber="86">218771</specimenCode>
,
<specimenCode id="DB85AA75014FE9594C40FBBFFC5BB051" box="[883,966,1085,1109]" pageId="86" pageNumber="86">218772</specimenCode>
;
<specimenCode id="DB85AA75014FE9594CE5FBBCFBFFB051" box="[982,1122,1085,1109]" pageId="86" pageNumber="86">MZB 34806</specimenCode>
,
<specimenCode id="DB85AA75014FE9594B40FBBFFB25B051" box="[1139,1208,1085,1109]" pageId="86" pageNumber="86">34808</specimenCode>
,
<specimenCode id="DB85AA75014FE9594D9FFBDCFD6FB072" box="[684,754,1118,1142]" pageId="86" pageNumber="86">34816</specimenCode>
,
<specimenCode id="DB85AA75014FE9594C34FBDDFCD0B073" box="[775,845,1119,1143]" pageId="86" pageNumber="86">34820</specimenCode>
,
<specimenCode id="DB85AA75014FE9594C51FBDDFC35B073" box="[866,936,1119,1143]" pageId="86" pageNumber="86">34827</specimenCode>
,
<specimenCode id="DB85AA75014FE9594C8FFBDDFB9FB073" box="[956,1026,1119,1143]" pageId="86" pageNumber="86">34833</specimenCode>
,
<specimenCode id="DB85AA75014FE9594B24FBDCFBC0B072" box="[1047,1117,1118,1142]" pageId="86" pageNumber="86">34856</specimenCode>
,
<specimenCode id="DB85AA75014FE9594B41FBDDFB25B073" box="[1138,1208,1119,1143]" pageId="86" pageNumber="86">34870</specimenCode>
,
<specimenCode id="DB85AA75014FE9594D9FFB02FD6FB09C" box="[684,754,1152,1176]" pageId="86" pageNumber="86">34871</specimenCode>
,
<specimenCode id="DB85AA75014FE9594C3AFBFDFCCDB093" box="[777,848,1151,1175]" pageId="86" pageNumber="86">34876</specimenCode>
,
<specimenCode id="DB85AA75014FE9594C56FB02FC31B09C" box="[869,940,1152,1176]" pageId="86" pageNumber="86">34877</specimenCode>
,
<specimenCode id="DB85AA75014FE9594CF1FB02FB94B09C" box="[962,1033,1152,1176]" pageId="86" pageNumber="86">34883</specimenCode>
,
<specimenCode id="DB85AA75014FE9594B2CFB02FBF8B09C" box="[1055,1125,1152,1176]" pageId="86" pageNumber="86">43011</specimenCode>
;
<specimenCode id="DB85AA75014FE9594B48FB02FC9FB0BD" pageId="86" pageNumber="86">NMV Z21910</specimenCode>
,
<specimenCode id="DB85AA75014FE9594C22FB23FCFBB0BD" box="[785,870,1185,1209]" pageId="86" pageNumber="86">Z21911</specimenCode>
,
<specimenCode id="DB85AA75014FE9594C45FB23FC50B0BD" box="[886,973,1185,1209]" pageId="86" pageNumber="86">Z21938</specimenCode>
),
</materialsCitation>
<materialsCitation id="3B4B0853014FE9594CD7FB22FC97B0DE" ID-GBIF-Occurrence="3421079400" collectionCode="FMNH" location="Salu Tiwo" pageId="86" pageNumber="86" specimenCode="FMNH 218744" specimenCount="1">
<location id="8EFC54D5014FE9594CD7FB22FBCBB0BC" LSID="urn:lsid:plazi:treatment:038AB318014BE9594F89FAE8FCEFB0FF:8EFC54D5014FE9594CD7FB22FBCBB0BC" box="[996,1110,1184,1208]" name="Salu Tiwo" pageId="86" pageNumber="86">Salu Tiwo</location>
(
<specimenCode id="DB85AA75014FE9594B59FB23FD62B0DE" pageId="86" pageNumber="86">FMNH 218744</specimenCode>
),
</materialsCitation>
<materialsCitation id="3B4B0853014FE9594C22FB40FCEFB0FF" ID-GBIF-Occurrence="3421079407" collectionCode="LSUMZ, NMV, C" location="Mt. Torompupu" pageId="86" pageNumber="86" specimenCode="LSUMZ 39480, NMV C40252, C40273" specimenCount="1">
<location id="8EFC54D5014FE9594C22FB40FC22B0DD" LSID="urn:lsid:plazi:treatment:038AB318014BE9594F89FAE8FCEFB0FF:8EFC54D5014FE9594C22FB40FC22B0DD" box="[785,959,1218,1241]" name="Mt. Torompupu" pageId="86" pageNumber="86">Mt. Torompupu</location>
(
<specimenCode id="DB85AA75014FE9594CFEFB40FBEFB0DE" box="[973,1138,1218,1242]" pageId="86" pageNumber="86">LSUMZ 39480</specimenCode>
;
<specimenCode id="DB85AA75014FE9594B4FFB40FC9EB0FF" pageId="86" pageNumber="86">NMV C40252</specimenCode>
,
<specimenCode id="DB85AA75014FE9594C3DFB61FCFAB0FF" box="[782,871,1251,1275]" pageId="86" pageNumber="86">C40273</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>