578 lines
70 KiB
XML
578 lines
70 KiB
XML
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<mods:title id="FEADDD1B3EE4B1EC6355B47EDD817BCE">Fourteen New, Endemic Species Of Shrew (Genus Crocidura) From Sulawesi Reveal A Spectacular Island Radiation</mods:title>
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<mods:namePart id="0B0581204E06CF5921163664EF2C6005">Esselstyn, Jacob A.</mods:namePart>
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<mods:namePart id="0EFB2AA547AF37211BD007B14C46D404">Achmadi, Anang S.</mods:namePart>
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<mods:namePart id="8E6DFA987855BAC4705B6FC3D927275E">Handika, Heru</mods:namePart>
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<mods:namePart id="A33CAD1121ACE531C8908294D5D140AF">Swanson, Mark T.</mods:namePart>
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<mods:namePart id="70455D2CBA79D32CBC69083708E5529D">Giarla, Thomas C.</mods:namePart>
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<mods:namePart id="3D6B3DDE7F6488BB6C27A9524B92DDE9">Rowe, Kevin C.</mods:namePart>
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<mods:title id="BC93C7D6D4CC438DD36EE094F6888072">Bulletin of the American Museum of Natural History</mods:title>
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<mods:date id="2208CE105570A9E533C95DF74295E1BF">2021</mods:date>
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<mods:number id="F7A9DC52F9954A66556F3133AFEC1148">2021-12-15</mods:number>
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<mods:url id="3811FE043130B65592520B178DED2FE7">https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-454/issue-1/0003-0090.454.1.1/Fourteen-New-Endemic-Species-of-Shrew-Genus-Crocidura-from-Sulawesi/10.1206/0003-0090.454.1.1.full</mods:url>
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<treatment id="038AB3180126E94D4DC9FA07FE0DB7D5" ID-DOI="http://doi.org/10.5281/zenodo.5795536" ID-GBIF-Taxon="191690926" ID-Zenodo-Dep="5795536" LSID="urn:lsid:zoobank.org:act:1587FDA2-B27F-47C4-95DF-E44A1081CCA4" httpUri="http://treatment.plazi.org/id/038AB3180126E94D4DC9FA07FE0DB7D5" lastPageId="66" lastPageNumber="66" pageId="63" pageNumber="63">
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<heading id="D0D4B5620126E9304DC9FA07FBF3B19A" box="[762,1134,1413,1438]" centered="true" fontSize="9" level="2" pageId="63" pageNumber="63" reason="2">
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<emphasis id="B957DE1C0126E9304DC9FA07FC7BB19A" bold="true" box="[762,998,1413,1438]" pageId="63" pageNumber="63">
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<taxonomicName id="4C23798D0126E9304DC9FA07FC42B199" authority="Esselstyn & Achmadi & Handika & Swanson & Giarla & Rowe, 2021" authorityName="Esselstyn & Achmadi & Handika & Swanson & Giarla & Rowe" authorityYear="2021" box="[762,991,1413,1437]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="63" pageNumber="63" phylum="Chordata" rank="species" species="mediocris" status="sp. nov.">
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||
<emphasis id="B957DE1C0126E9304DC9FA07FC42B199" bold="true" box="[762,991,1413,1437]" italics="true" pageId="63" pageNumber="63">Crocidura mediocris</emphasis>
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</taxonomicName>
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||
,
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||
</emphasis>
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<taxonomicNameLabel id="A26463670126E9304CDEFA04FBF3B19A" box="[1005,1134,1414,1438]" pageId="63" pageNumber="63" rank="species">new species</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="C33951850126E9304C01FA3AFB92B216" pageId="63" pageNumber="63" type="description">
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<heading id="D0D4B5620126E9304C01FA3AFB92B216" centered="true" fontSize="10" level="2" pageId="63" pageNumber="63" reason="4">
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LSID:
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<uri id="FFB20E0C0126E9304C4BFA35FB92B216" pageId="63" pageNumber="63">
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urn:lsid:zoobank. org:act:
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</uri>
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</heading>
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</paragraph>
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<subSubSection id="C33951850126E94E4DFFF9A9FE5DB5C4" lastPageId="65" lastPageNumber="65" pageId="63" pageNumber="63" type="materials_examined">
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<materialsCitation id="3B4B08530126E94E4DFFF9A9FE5DB5C4" ID-GBIF-Occurrence="3421079365" accessionNumber="MZB 43007" collectingDate="2010-10-22" collectionCode="MZB, FMNH" collectorName="J. A. Esselstyn & The & External" country="Indonesia" county="Luwu Utara" lastPageId="65" lastPageNumber="65" location="Mt. Balease" municipality="Sukamaju" pageId="63" pageNumber="63" specimenCode="FMNH 210603" specimenCount="2" stateProvince="Sulawesi Selatan" typeStatus="holotype">
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<paragraph id="8B9C020E0126E94E4DFFF9A9FDB9B558" blockId="63.[684,1213,1579,1735]" lastBlockId="65.[108,638,226,1738]" lastPageId="65" lastPageNumber="65" pageId="63" pageNumber="63">
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<typeStatus id="5498BCAC0126E9304DFFF9A9FCDEB247" box="[716,835,1579,1603]" pageId="63" pageNumber="63" type="holotype">HOLOTYPE</typeStatus>
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:
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<collectionCode id="ED329ACB0126E9304C60F9AEFC11B247" box="[851,908,1580,1603]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="63" pageNumber="63" type="Museum">MZB</collectionCode>
|
||
43007 (=
|
||
<specimenCode id="DB85AA750126E9304B37F9AEFB2DB247" box="[1028,1200,1579,1603]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="63" pageNumber="63" type="Museum">
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<collectionCode id="ED329ACB0126E9304B37F9AEFBC9B247" box="[1028,1108,1580,1603]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="63" pageNumber="63" type="Museum">FMNH</collectionCode>
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210603
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||
</specimenCode>
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||
), an adult female collected by
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<collectorName id="26D667D80126E9304CD6F9CEFBEAB260" box="[997,1143,1612,1636]" pageId="63" pageNumber="63">J.A. Esselstyn</collectorName>
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||
on
|
||
<date id="FF9D24CE0126E9304B91F9CEFCDBB281" pageId="63" pageNumber="63" value="2010-10-22">
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||
<collectingDate id="EFD9DD260126E9304B91F9CEFCDBB281" pageId="63" pageNumber="63" value="2010-10-22">22 October 2010</collectingDate>
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</date>
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.
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<collectorName id="26D667D80126E9304C61F9EFFCE6B281" box="[850,891,1645,1669]" pageId="63" pageNumber="63">The</collectorName>
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||
specimen consists of a dried skin, cleaned skull and skeleton, and tissue samples.
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||
<collectorName id="26D667D80126E9304DD5F92DFCDFB2C3" box="[742,834,1711,1735]" pageId="63" pageNumber="63">External</collectorName>
|
||
measurements from the
|
||
<typeStatus id="5498BCAC0126E9304B6EF92DFB21B2C3" box="[1117,1212,1711,1735]" pageId="63" pageNumber="63" type="holotype">holotype</typeStatus>
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are:
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<quantity id="4CDBAFEB0158E94E4FA9FF60FF69B4FD" box="[154,244,226,250]" metricMagnitude="-1" metricUnit="m" metricValue="1.02" pageId="65" pageNumber="65" unit="mm" value="102.0">102 mm</quantity>
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×
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<quantity id="4CDBAFEB0158E94E4E22FF60FEC2B4FD" box="[273,351,226,250]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="65" pageNumber="65" unit="mm" value="41.0">41 mm</quantity>
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×
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<quantity id="4CDBAFEB0158E94E4E48FF60FE54B4FD" box="[379,457,226,250]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="65" pageNumber="65" unit="mm" value="11.0">11 mm</quantity>
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×
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<quantity id="4CDBAFEB0158E94E4ED6FF60FDBBB4FD" box="[485,550,226,249]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="65" pageNumber="65" unit="mm" value="7.0">7 mm</quantity>
|
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=
|
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<quantity id="4CDBAFEB0158E94E4D71FF60FDE5B4FE" box="[578,632,226,250]" metricMagnitude="-3" metricUnit="kg" metricValue="3.7" pageId="65" pageNumber="65" unit="g" value="3.7">3.7 g</quantity>
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.
|
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<collectorName id="26D667D80158E94E4F5FFE81FF08B51F" box="[108,149,259,283]" pageId="65" pageNumber="65">The</collectorName>
|
||
voucher specimen and a tissue sample will be permanently curated at
|
||
<collectionCode id="ED329ACB0158E94E4EAFFEA6FE4AB53F" box="[412,471,292,315]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="65" pageNumber="65" type="Museum">MZB</collectionCode>
|
||
, with an additional tissue sample retained by
|
||
<collectionCode id="ED329ACB0158E94E4EFEFEC7FDBDB558" box="[461,544,325,348]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="65" pageNumber="65" type="Museum">FMNH</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF5C52860159E94F4F5BFA18FF6AB1EA" ID-DOI="http://doi.org/10.5281/zenodo.5788891" ID-Zenodo-Dep="5788891" httpUri="https://zenodo.org/record/5788891/files/figure.png" pageId="64" pageNumber="64" startId="64.[104,146,1434,1458]" targetBox="[105,1612,439,1407]" targetPageId="64">
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<paragraph id="8B9C020E0159E94F4F5BFA18FF6AB1EA" blockId="64.[104,1613,1433,1518]" pageId="64" pageNumber="64">
|
||
FIG. 31. Images showing dorsal, ventral, and lateral views of the skull and lateral and occlusal views of the dentary of the three members of the Small- Bodied Group that are not from the northern peninsula:
|
||
<emphasis id="B957DE1C0159E94F4D8EFA35FD48B1D5" bold="true" box="[701,725,1463,1489]" pageId="64" pageNumber="64">A,</emphasis>
|
||
<taxonomicName id="4C23798D0159E94F4DEEFA35FC70B1D5" authority=", FMNH" authorityName="FMNH" box="[733,1005,1463,1489]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="64" pageNumber="64" phylum="Chordata" rank="species" species="levicula">
|
||
<emphasis id="B957DE1C0159E94F4DEEFA35FC0BB1D5" box="[733,918,1463,1489]" italics="true" pageId="64" pageNumber="64">Crocidura levicula</emphasis>
|
||
, FMNH
|
||
</taxonomicName>
|
||
213362;
|
||
<emphasis id="B957DE1C0159E94F4B7EFA35FBFEB1D5" bold="true" box="[1101,1123,1463,1489]" pageId="64" pageNumber="64">B,</emphasis>
|
||
<taxonomicName id="4C23798D0159E94F4B58FA35FADFB1D5" authority=", FMNH" authorityName="FMNH" box="[1131,1346,1463,1489]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="64" pageNumber="64" phylum="Chordata" rank="species" species="mediocris">
|
||
<emphasis id="B957DE1C0159E94F4B58FA35FB77B1D5" box="[1131,1258,1463,1489]" italics="true" pageId="64" pageNumber="64">C. mediocris</emphasis>
|
||
, FMNH
|
||
</taxonomicName>
|
||
210603; and
|
||
<emphasis id="B957DE1C0159E94F4AFCFA35FA7BB1D5" bold="true" box="[1487,1510,1463,1489]" pageId="64" pageNumber="64">C,</emphasis>
|
||
<taxonomicName id="4C23798D0159E94F4ADCFA35FF3DB1EA" authority=", MVZ" authorityName="MVZ" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="64" pageNumber="64" phylum="Chordata" rank="species" species="parva">
|
||
<emphasis id="B957DE1C0159E94F4ADCFA35F9DAB1D5" box="[1519,1607,1463,1489]" italics="true" pageId="64" pageNumber="64">C. parva</emphasis>
|
||
, MVZ
|
||
</taxonomicName>
|
||
237577.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9C020E0158E94E4FBFFEE4FE5DB5C4" blockId="65.[108,638,226,1738]" pageId="65" pageNumber="65">
|
||
<collectionCode id="ED329ACB0158E94E4FBFFEE4FEA5B57A" box="[140,312,358,382]" pageId="65" pageNumber="65">TYPE LOCALITY</collectionCode>
|
||
:
|
||
<collectingCountry id="F334429E0158E94E4E74FEE4FE29B57A" box="[327,436,358,382]" name="Indonesia" pageId="65" pageNumber="65">Indonesia</collectingCountry>
|
||
,
|
||
<collectingRegion id="49E7CCEC0158E94E4EF1FEE4FDE5B57A" box="[450,632,358,382]" country="Indonesia" name="Sulawesi Selatan" pageId="65" pageNumber="65">Sulawesi Selatan</collectingRegion>
|
||
,
|
||
<collectingCounty id="62FD7A820158E94E4F5FFE05FF6DB59A" box="[108,240,391,414]" pageId="65" pageNumber="65">Luwu Utara</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BF898740158E94E4FCEFE05FEF7B59B" box="[253,362,391,415]" pageId="65" pageNumber="65">Sukamaju</collectingMunicipality>
|
||
,
|
||
<location id="8EFC54D50158E94E4E45FE05FE6AB59B" LSID="urn:lsid:plazi:treatment:038AB3180126E94D4DC9FA07FE0DB7D5:8EFC54D50158E94E4E45FE05FE6AB59B" box="[374,503,391,415]" country="Indonesia" county="Luwu Utara" municipality="Sukamaju" name="Mt. Balease" pageId="65" pageNumber="65" stateProvince="Sulawesi Selatan">Mt. Balease</location>
|
||
;
|
||
<geoCoordinate id="EE1764C90158E94E4D30FE05FDE5B59B" box="[515,632,391,415]" degrees="2.50002" direction="south" orientation="latitude" pageId="65" pageNumber="65" precision="1" value="-2.50002">2.50002° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1764C90158E94E4F5DFE2AFF63B5C4" box="[110,254,424,448]" degrees="120.48726" direction="east" orientation="longitude" pageId="65" pageNumber="65" precision="1" value="120.48726">120.48726° E</geoCoordinate>
|
||
,
|
||
<quantity id="4CDBAFEB0158E94E4E39FE2AFED2B5C4" box="[266,335,424,448]" metricMagnitude="2" metricUnit="m" metricValue="8.62" pageId="65" pageNumber="65" unit="m" value="862.0">862 m</quantity>
|
||
elevation.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
</subSubSection>
|
||
<subSubSection id="C33951850158E94E4FBFFE4BFE7FB640" pageId="65" pageNumber="65" type="etymology">
|
||
<paragraph id="8B9C020E0158E94E4FBFFE4BFE7FB640" blockId="65.[108,638,226,1738]" pageId="65" pageNumber="65">
|
||
ETYMOLOGY:
|
||
<emphasis id="B957DE1C0158E94E4E11FE4BFE17B5E5" box="[290,394,457,481]" italics="true" pageId="65" pageNumber="65">Mediocris</emphasis>
|
||
is Latin for “average,” applied in recognition that this is a species of shrew with no striking phenotypic traits worthy of hanging a descriptive name on.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33951850158E94E4FBEFDCFFEF7B7F6" pageId="65" pageNumber="65" type="distribution">
|
||
<paragraph id="8B9C020E0158E94E4FBEFDCFFEF7B7F6" blockId="65.[108,638,226,1738]" pageId="65" pageNumber="65">
|
||
GEOGRAPHIC DISTRIBUTION: This species is found across the southern and western portions of the west-central area of endemism (Mts. Torompupu and Balease,
|
||
<collectingRegion id="49E7CCEC0158E94E4E83FD32FF4FB6EE" country="Indonesia" name="Sulawesi Tengah" pageId="65" pageNumber="65">Central Sulawesi Province</collectingRegion>
|
||
; Salu Tiwo,
|
||
<collectingRegion id="49E7CCEC0158E94E4E56FD50FDEAB6EE" box="[357,631,722,746]" country="Indonesia" name="Sulawesi Barat" pageId="65" pageNumber="65">West Sulawesi Province</collectingRegion>
|
||
; and Mt. Latimojong,
|
||
<collectingRegion id="49E7CCEC0158E94E4E6DFD71FDE9B70F" box="[350,628,755,779]" country="Indonesia" name="Sulawesi Selatan" pageId="65" pageNumber="65">South Sulawesi Province</collectingRegion>
|
||
) and in the south-east area of endemism (Mt. Mekongga,
|
||
<collectingRegion id="49E7CCEC0158E94E4FC5FCB7FDD5B749" box="[246,584,821,845]" country="Indonesia" name="Sulawesi Tenggara" pageId="65" pageNumber="65">Southeast Sulawesi Province</collectingRegion>
|
||
;
|
||
<figureCitation id="13181E8B0158E94E4D6AFCB7FF17B76A" captionStart="FIG" captionStartId="53.[110,152,1054,1075]" captionTargetId="graphics-333@53.[275,1045,225,1023]" captionTargetPageId="53" captionText="FIG. 25. Map of Sulawesi showing localities sampled for shrews. Colored areas enclose localities with known records of members of the Small-Bodied Group of Sulawesi Crocidura." figureDoi="http://doi.org/10.5281/zenodo.5789154" httpUri="https://zenodo.org/record/5789154/files/figure.png" pageId="65" pageNumber="65">fig. 25</figureCitation>
|
||
). We found this species from approximately
|
||
<quantity id="4CDBAFEB0158E94E4F5DFCF5FE92B78A" box="[110,271,887,911]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.9" metricValueMin="0.2" pageId="65" pageNumber="65" unit="m" value="1050.0" valueMax="1900.0" valueMin="200.0">200 to 1900 m</quantity>
|
||
elevation (
|
||
<figureCitation id="13181E8B0158E94E4EB8FCF5FE49B78B" box="[395,468,887,911]" captionStart="FIG" captionStartId="31.[104,146,1395,1419]" captionTargetBox="[104,1603,469,1356]" captionTargetId="graphics-9@31.[188,1530,466,1251]" captionTargetPageId="31" captionText="FIG. 13. Elevational records of all species of Crocidura known from Sulawesi. Each point represents a specimen. For specimens associated with a minimum and maximum elevation, we used the center of the given elevational range. Sample sizes are given above the x-axis. Species are grouped according to the species groups used in the text (Thick = Thick-Tailed Group)." figureDoi="http://doi.org/10.5281/zenodo.5788859" httpUri="https://zenodo.org/record/5788859/files/figure.png" pageId="65" pageNumber="65">fig. 13</figureCitation>
|
||
). Although the species was present at Salu Tiwo (ca.
|
||
<quantity id="4CDBAFEB0158E94E4D36FC1AFDD1B7B4" box="[517,588,920,944]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="65" pageNumber="65" unit="m" value="200.0">200 m</quantity>
|
||
), we did not find it at higher elevations on Mt. Gandang Dewata (
|
||
<tableCitation id="C6A137B50158E94E4E3DFC58FEC1B7F6" box="[270,348,986,1010]" captionStart="TABLE 3" captionStartId="27.[611,691,226,250]" captionTargetBox="[117,1200,329,1675]" captionTargetPageId="27" captionText="TABLE 3 Elevational Ranges (m) and Species Richness of Crocidura on Mountains of Sulawesia" httpUri="http://table.plazi.org/id/DF5C52860102E9144D50FF60FBDEB51A" pageId="65" pageNumber="65" tableUuid="DF5C52860102E9144D50FF60FBDEB51A">table 3</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33951850158E94E4FBFFC79FB1FB59B" pageId="65" pageNumber="65" type="diagnosis">
|
||
<paragraph id="8B9C020E0158E94E4FBFFC79FB1FB59B" blockId="65.[108,638,226,1738]" lastBlockId="65.[684,1215,226,1738]" pageId="65" pageNumber="65">
|
||
DIAGNOSIS:
|
||
<taxonomicName id="4C23798D0158E94E4E2AFC79FE66B017" box="[281,507,1019,1043]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="mediocris">
|
||
<emphasis id="B957DE1C0158E94E4E2AFC79FE66B017" box="[281,507,1019,1043]" italics="true" pageId="65" pageNumber="65">Crocidura mediocris</emphasis>
|
||
</taxonomicName>
|
||
is another small shrew (
|
||
<tableCitation id="C6A137B50158E94E4FCBFB9EFED1B030" box="[248,332,1052,1076]" captionStart="TABLE 2" captionStartId="25.[610,690,226,250]" captionTargetBox="[119,1184,329,1682]" captionTargetPageId="25" captionText="TABLE 2 Descriptive Statisticsa for External Measurements (mm) and Mass (g) for Species of Sulawesi Crocidura" httpUri="http://table.plazi.org/id/DF5C52860100E9164D51FF60FBE5B523" pageId="65" pageNumber="65" tableUuid="DF5C52860100E9164D51FF60FBE5B523">tables 2</tableCitation>
|
||
,
|
||
<tableCitation id="C6A137B50158E94E4E66FB9EFEFEB030" box="[341,355,1052,1076]" captionStart="TABLE 8" captionStartId="56.[609,689,226,250]" captionTargetBox="[117,1198,358,1166]" captionTargetPageId="56" captionText="TABLE 8 Descriptive Statisticsa for Craniodental Measurements (mm) for Species of the Small-Bodied Group of Sulawesi Crocidura" httpUri="http://table.plazi.org/id/DF5C52860121E9374D52FF60FC1DB540" pageId="65" pageNumber="65" tableUuid="DF5C52860121E9374D52FF60FC1DB540">8</tableCitation>
|
||
), with a delicate build and dark gray to medium brown pelage that is only slightly paler on the venter. The tail is shorter than head-and-body length and moderately dorsoventrally bicolored. The feet are somewhat paler than the pelage, more so around the digits (
|
||
<figureCitation id="13181E8B0158E94E4F46FB61FF50B0FF" box="[117,205,1251,1275]" captionStart="FIG" captionStartId="59.[108,150,1254,1278]" captionTargetBox="[104,1610,589,1228]" captionTargetId="figure-10@59.[104,1610,589,1228]" captionTargetPageId="59" captionText="FIG. 29. Images showing the ventral surface of the left hind foot and dorsal surfaces of the tail base (approximately 1 cm from rump) and tail tip from the three members of the Small-Bodied Group that are not from the northern peninsula: A, Crocidura levicula, FMNH 213271; B, C. mediocris, FMNH 210566; and C, C. parva, MVZ 237583. Where two scale bars are present within a panel, the upper applies to the foot and the lower to the tail." figureDoi="http://doi.org/10.5281/zenodo.5788887" httpUri="https://zenodo.org/record/5788887/files/figure.png" pageId="65" pageNumber="65">fig. 29B</figureCitation>
|
||
). The pelage covers the body completely, but individual hairs are short (
|
||
<quantity id="4CDBAFEB0158E94E4EF1FA86FDBDB11F" box="[450,544,1284,1308]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="65" pageNumber="65" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
|
||
at middorsum). Mystacial vibrissae are short relative to body size, and unpigmented along most of their length. The tail is covered in moderately dense bristles along the proximal two-thirds of its length and a relatively high density of short, dark, applied hairs along its entire length (
|
||
<figureCitation id="13181E8B0158E94E4EDEFA48FDDFB1E6" box="[493,578,1482,1506]" captionStart="FIG" captionStartId="59.[108,150,1254,1278]" captionTargetBox="[104,1610,589,1228]" captionTargetId="figure-10@59.[104,1610,589,1228]" captionTargetPageId="59" captionText="FIG. 29. Images showing the ventral surface of the left hind foot and dorsal surfaces of the tail base (approximately 1 cm from rump) and tail tip from the three members of the Small-Bodied Group that are not from the northern peninsula: A, Crocidura levicula, FMNH 213271; B, C. mediocris, FMNH 210566; and C, C. parva, MVZ 237583. Where two scale bars are present within a panel, the upper applies to the foot and the lower to the tail." figureDoi="http://doi.org/10.5281/zenodo.5788887" httpUri="https://zenodo.org/record/5788887/files/figure.png" pageId="65" pageNumber="65">fig. 29B</figureCitation>
|
||
). The dorsal surfaces of the digits and, in some specimens, the feet are light brown. The plantar and palmar surfaces are generally dark, but the ventral sides of the digits are white (
|
||
<figureCitation id="13181E8B0158E94E4EEAF9CCFDAEB263" box="[473,563,1614,1639]" captionStart="FIG" captionStartId="59.[108,150,1254,1278]" captionTargetBox="[104,1610,589,1228]" captionTargetId="figure-10@59.[104,1610,589,1228]" captionTargetPageId="59" captionText="FIG. 29. Images showing the ventral surface of the left hind foot and dorsal surfaces of the tail base (approximately 1 cm from rump) and tail tip from the three members of the Small-Bodied Group that are not from the northern peninsula: A, Crocidura levicula, FMNH 213271; B, C. mediocris, FMNH 210566; and C, C. parva, MVZ 237583. Where two scale bars are present within a panel, the upper applies to the foot and the lower to the tail." figureDoi="http://doi.org/10.5281/zenodo.5788887" httpUri="https://zenodo.org/record/5788887/files/figure.png" pageId="65" pageNumber="65">fig. 29B</figureCitation>
|
||
). As is typical, more pigment is present on the posterior and lateral portions of the palmar and plantar surfaces. The skull is small and delicate, with a relatively robust dentition (
|
||
<figureCitation id="13181E8B0158E94E4CEAFF60FBAFB4FE" box="[985,1074,226,250]" captionStart="FIG" captionStartId="64.[104,146,1434,1458]" captionTargetBox="[105,1612,439,1407]" captionTargetId="figure-65@64.[104,1612,439,1407]" captionTargetPageId="64" captionText="FIG. 31. Images showing dorsal, ventral, and lateral views of the skull and lateral and occlusal views of the dentary of the three members of the Small- Bodied Group that are not from the northern peninsula: A, Crocidura levicula, FMNH 213362; B, C. mediocris, FMNH 210603; and C, C. parva, MVZ 237577." figureDoi="http://doi.org/10.5281/zenodo.5788891" httpUri="https://zenodo.org/record/5788891/files/figure.png" pageId="65" pageNumber="65">fig. 31B</figureCitation>
|
||
). The braincase is somewhat narrow and the interobital region is narrow relative to skull length (
|
||
<figureCitation id="13181E8B0158E94E4B5BFEA6FB2DB538" box="[1128,1200,292,316]" captionStart="FIG" captionStartId="26.[108,150,1414,1435]" captionTargetId="graphics-12@26.[213,1196,224,1265]" captionTargetPageId="26" captionText="FIG. 10. Box plots of relative skull measures showing braincase breadth (BB), interorbital width (IOW), and rostral length (RL) divided by condyloincisive length (CIL) and BB divided by IOW for all species of Sulawesi shrew. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are ordered according to the species groups used in the text (Thick = Thick-Tailed Group)." figureDoi="http://doi.org/10.5281/zenodo.5789149" httpUri="https://zenodo.org/record/5789149/files/figure.png" pageId="65" pageNumber="65">fig. 10</figureCitation>
|
||
). The maxillary process is modest and the maxillary bridge is narrow (
|
||
<figureCitation id="13181E8B0158E94E4C90FEE4FC61B57A" box="[931,1020,358,382]" captionStart="FIG" captionStartId="64.[104,146,1434,1458]" captionTargetBox="[105,1612,439,1407]" captionTargetId="figure-65@64.[104,1612,439,1407]" captionTargetPageId="64" captionText="FIG. 31. Images showing dorsal, ventral, and lateral views of the skull and lateral and occlusal views of the dentary of the three members of the Small- Bodied Group that are not from the northern peninsula: A, Crocidura levicula, FMNH 213362; B, C. mediocris, FMNH 210603; and C, C. parva, MVZ 237577." figureDoi="http://doi.org/10.5281/zenodo.5788891" httpUri="https://zenodo.org/record/5788891/files/figure.png" pageId="65" pageNumber="65">fig. 31B</figureCitation>
|
||
). The lambdoidal ridge is prominent for such a small species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33951850158E94D4DFFFE2AFE80B538" lastPageId="66" lastPageNumber="66" pageId="65" pageNumber="65" type="description">
|
||
<paragraph id="8B9C020E0158E94D4DFFFE2AFE80B538" blockId="65.[684,1215,226,1738]" lastBlockId="66.[108,637,226,977]" lastPageId="66" lastPageNumber="66" pageId="65" pageNumber="65">
|
||
COMPARISONS:
|
||
<taxonomicName id="4C23798D0158E94E4C44FE2AFBCCB5C4" box="[887,1105,424,448]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="mediocris">
|
||
<emphasis id="B957DE1C0158E94E4C44FE2AFBCCB5C4" box="[887,1105,424,448]" italics="true" pageId="65" pageNumber="65">Crocidura mediocris</emphasis>
|
||
</taxonomicName>
|
||
is one of the smallest species on
|
||
<collectingRegion id="49E7CCEC0158E94E4C8DFE4BFB80B5E5" box="[958,1053,457,481]" country="Indonesia" name="Sulawesi Utara" pageId="65" pageNumber="65">Sulawesi</collectingRegion>
|
||
, easily distinguished on body size alone from all species outside the Small-Bodied Group (
|
||
<figureCitation id="13181E8B0158E94E4CC4FD89FBA2B627" box="[1015,1087,523,547]" captionStart="FIG" captionStartId="26.[108,150,1414,1435]" captionTargetId="graphics-12@26.[213,1196,224,1265]" captionTargetPageId="26" captionText="FIG. 10. Box plots of relative skull measures showing braincase breadth (BB), interorbital width (IOW), and rostral length (RL) divided by condyloincisive length (CIL) and BB divided by IOW for all species of Sulawesi shrew. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are ordered according to the species groups used in the text (Thick = Thick-Tailed Group)." figureDoi="http://doi.org/10.5281/zenodo.5789149" httpUri="https://zenodo.org/record/5789149/files/figure.png" pageId="65" pageNumber="65">fig. 10</figureCitation>
|
||
).
|
||
<taxonomicName id="4C23798D0158E94E4B66FD89FC94B640" authorityName="Esselstyn & Achmadi & Handika & Swanson & Giarla & Rowe" authorityYear="2021" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="normalis">
|
||
<emphasis id="B957DE1C0158E94E4B66FD89FC94B640" italics="true" pageId="65" pageNumber="65">Crocidura normalis</emphasis>
|
||
</taxonomicName>
|
||
is the smallest member of the Ordinary Group and may be closely related to
|
||
<taxonomicName id="4C23798D0158E94E4B09FDCCFB23B661" box="[1082,1214,589,613]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="mediocris">
|
||
<emphasis id="B957DE1C0158E94E4B09FDCCFB23B661" box="[1082,1214,589,613]" italics="true" pageId="65" pageNumber="65">C. mediocris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C23798D0158E94E4DD3FDEDFCDCB682" box="[736,833,623,646]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="parva">
|
||
<emphasis id="B957DE1C0158E94E4DD3FDEDFCDCB682" box="[736,833,623,646]" italics="true" pageId="65" pageNumber="65">C. parva</emphasis>
|
||
</taxonomicName>
|
||
, also of the Small-Bodied Group (
|
||
<figureCitation id="13181E8B0158E94E4D80FD0DFD69B6A3" box="[691,756,655,679]" captionStart="FIG" captionStartId="18.[108,150,1596,1617]" captionTargetBox="[111,1203,227,1567]" captionTargetId="graphics-32@18.[111,872,237,1559]" captionTargetPageId="18" captionText="FIG. 5. Maximum likelihood estimate of the mitochondrial gene tree derived from an analysis of 14,007 characters (representing protein-coding and rRNA genes) from 83 samples. Bootstrap support values <95 are shown at nodes. Tips are labeled with the species, locality, and voucher number." figureDoi="http://doi.org/10.5281/zenodo.5788845" httpUri="https://zenodo.org/record/5788845/files/figure.png" pageId="65" pageNumber="65">figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13181E8B0158E94E4DCCFD12FC93B6AC" box="[767,782,656,680]" captionStart="FIG" captionStartId="21.[108,149,1653,1674]" captionTargetBox="[209,1136,226,1678]" captionTargetId="graphics-38@20.[180,704,233,1669]" captionTargetPageId="20" captionText="FIG. 7. Estimated species tree from analysis of 3940 ultraconserved element loci in ASTRAL. Samples from Sulawesi are labeled with the species name, locality, and catalog number. Asterisks indicate type specimens from Miller and Hollister (1921). Local posterior probabilities <0.95 are shown. Tip branch lengths are arbitrary." figureDoi="http://doi.org/10.5281/zenodo.5789145" httpUri="https://zenodo.org/record/5789145/files/figure.png" pageId="65" pageNumber="65">7</figureCitation>
|
||
). Compared to
|
||
<taxonomicName id="4C23798D0158E94E4C87FD12FBB3B6A3" authorityName="Esselstyn & Achmadi & Handika & Swanson & Giarla & Rowe" authorityYear="2021" box="[948,1070,655,679]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="normalis">
|
||
<emphasis id="B957DE1C0158E94E4C87FD12FBB3B6A3" box="[948,1070,655,679]" italics="true" pageId="65" pageNumber="65">C. normalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C23798D0158E94E4B09FD12FB20B6A3" box="[1082,1213,655,679]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="mediocris">
|
||
<emphasis id="B957DE1C0158E94E4B09FD12FB20B6A3" box="[1082,1213,655,679]" italics="true" pageId="65" pageNumber="65">C. mediocris</emphasis>
|
||
</taxonomicName>
|
||
is smaller, paler colored, and has a shorter tail (
|
||
<figureCitation id="13181E8B0158E94E4BAAFD32FD5AB6EE" captionStart="FIG" captionStartId="26.[108,150,1414,1435]" captionTargetId="graphics-12@26.[213,1196,224,1265]" captionTargetPageId="26" captionText="FIG. 10. Box plots of relative skull measures showing braincase breadth (BB), interorbital width (IOW), and rostral length (RL) divided by condyloincisive length (CIL) and BB divided by IOW for all species of Sulawesi shrew. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are ordered according to the species groups used in the text (Thick = Thick-Tailed Group)." figureDoi="http://doi.org/10.5281/zenodo.5789149" httpUri="https://zenodo.org/record/5789149/files/figure.png" pageId="65" pageNumber="65">fig. 10</figureCitation>
|
||
;
|
||
<tableCitation id="C6A137B50158E94E4DE0FD50FC80B6EE" box="[723,797,722,746]" captionStart="TABLE 2" captionStartId="25.[610,690,226,250]" captionTargetBox="[119,1184,329,1682]" captionTargetPageId="25" captionText="TABLE 2 Descriptive Statisticsa for External Measurements (mm) and Mass (g) for Species of Sulawesi Crocidura" httpUri="http://table.plazi.org/id/DF5C52860100E9164D51FF60FBE5B523" pageId="65" pageNumber="65" tableUuid="DF5C52860100E9164D51FF60FBE5B523">table 2</tableCitation>
|
||
). Among the Small-Bodied species,
|
||
<taxonomicName id="4C23798D0158E94E4B95FD50FC8CB70F" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="mediocris">
|
||
<emphasis id="B957DE1C0158E94E4B95FD50FC8CB70F" italics="true" pageId="65" pageNumber="65">C. mediocris</emphasis>
|
||
</taxonomicName>
|
||
has the shortest average head-and-body length. Its tail is also short, and relative tail length is comparable to
|
||
<taxonomicName id="4C23798D0158E94E4C47FCB7FC40B749" box="[884,989,821,845]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="baletei">
|
||
<emphasis id="B957DE1C0158E94E4C47FCB7FC40B749" box="[884,989,821,845]" italics="true" pageId="65" pageNumber="65">C. baletei</emphasis>
|
||
</taxonomicName>
|
||
, greater than in
|
||
<taxonomicName id="4C23798D0158E94E4B97FCB7FD62B76A" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="levicula">
|
||
<emphasis id="B957DE1C0158E94E4B97FCB7FD62B76A" italics="true" pageId="65" pageNumber="65">C. levicula</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C23798D0158E94E4C22FCD4FC06B76A" authorityName="Esselstyn & Achmadi & Handika & Swanson & Giarla & Rowe" authorityYear="2021" box="[785,923,854,878]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="tenebrosa">
|
||
<emphasis id="B957DE1C0158E94E4C22FCD4FC06B76A" box="[785,923,854,878]" italics="true" pageId="65" pageNumber="65">C. tenebrosa</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C23798D0158E94E4CD1FCD4FBD9B76A" box="[994,1092,854,878]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="parva">
|
||
<emphasis id="B957DE1C0158E94E4CD1FCD4FBD9B76A" box="[994,1092,854,878]" italics="true" pageId="65" pageNumber="65">C. parva</emphasis>
|
||
</taxonomicName>
|
||
, and only slightly less than in
|
||
<taxonomicName id="4C23798D0158E94E4C4FFCF5FC25B78B" box="[892,952,887,911]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="lea">
|
||
<emphasis id="B957DE1C0158E94E4C4FFCF5FC25B78B" box="[892,952,887,911]" italics="true" pageId="65" pageNumber="65">C. lea</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13181E8B0158E94E4CF6FCF5FC60B78B" box="[965,1021,887,911]" captionStart="FIG" captionStartId="24.[108,150,1587,1608]" captionTargetBox="[137,1192,224,1561]" captionTargetId="graphics-9@24.[203,1191,224,1433]" captionTargetPageId="24" captionText="FIG. 9. Box plots showing variation in external measurements from all species of Sulawesi shrew. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are grouped according to the species groups used in the text (Thick = Thick- Tailed Group). All measurements in mm. HBL = head-and-body length." figureDoi="http://doi.org/10.5281/zenodo.5788853" httpUri="https://zenodo.org/record/5788853/files/figure.png" pageId="65" pageNumber="65">fig. 9</figureCitation>
|
||
). In color, the feet of
|
||
<taxonomicName id="4C23798D0158E94E4DF8FC1AFCCEB7B4" box="[715,851,920,944]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="mediocris">
|
||
<emphasis id="B957DE1C0158E94E4DF8FC1AFCCEB7B4" box="[715,851,920,944]" italics="true" pageId="65" pageNumber="65">C. mediocris</emphasis>
|
||
</taxonomicName>
|
||
are darker than those of
|
||
<taxonomicName id="4C23798D0158E94E4B45FC1AFB28B7B4" box="[1142,1205,920,944]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="lea">
|
||
<emphasis id="B957DE1C0158E94E4B45FC1AFB28B7B4" box="[1142,1205,920,944]" italics="true" pageId="65" pageNumber="65">C. lea</emphasis>
|
||
</taxonomicName>
|
||
. Aside from the relative tail-length differences,
|
||
<taxonomicName id="4C23798D0158E94E4B94FC3BFC8CB7F6" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="mediocris">
|
||
<emphasis id="B957DE1C0158E94E4B94FC3BFC8CB7F6" italics="true" pageId="65" pageNumber="65">C. mediocris</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C23798D0158E94E4C96FC59FB88B7F6" box="[933,1045,986,1010]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="levicula">
|
||
<emphasis id="B957DE1C0158E94E4C96FC59FB88B7F6" box="[933,1045,986,1010]" italics="true" pageId="65" pageNumber="65">C. levicula</emphasis>
|
||
</taxonomicName>
|
||
in overall proportions but differs in having a longer and slightly narrower skull (
|
||
<tableCitation id="C6A137B50158E94E4C53FB9EFC32B030" box="[864,943,1052,1076]" captionStart="TABLE 8" captionStartId="56.[609,689,226,250]" captionTargetBox="[117,1198,358,1166]" captionTargetPageId="56" captionText="TABLE 8 Descriptive Statisticsa for Craniodental Measurements (mm) for Species of the Small-Bodied Group of Sulawesi Crocidura" httpUri="http://table.plazi.org/id/DF5C52860121E9374D52FF60FC1DB540" pageId="65" pageNumber="65" tableUuid="DF5C52860121E9374D52FF60FC1DB540">table 8</tableCitation>
|
||
), paler pelage and feet, more rounded hypothenar on the hind foot (
|
||
<figureCitation id="13181E8B0158E94E4BAAFBBFFD54B073" captionStart="FIG" captionStartId="59.[108,150,1254,1278]" captionTargetBox="[104,1610,589,1228]" captionTargetId="figure-10@59.[104,1610,589,1228]" captionTargetPageId="59" captionText="FIG. 29. Images showing the ventral surface of the left hind foot and dorsal surfaces of the tail base (approximately 1 cm from rump) and tail tip from the three members of the Small-Bodied Group that are not from the northern peninsula: A, Crocidura levicula, FMNH 213271; B, C. mediocris, FMNH 210566; and C, C. parva, MVZ 237583. Where two scale bars are present within a panel, the upper applies to the foot and the lower to the tail." figureDoi="http://doi.org/10.5281/zenodo.5788887" httpUri="https://zenodo.org/record/5788887/files/figure.png" pageId="65" pageNumber="65">fig. 29</figureCitation>
|
||
), and on average, fewer tail bristles.
|
||
<taxonomicName id="4C23798D0158E94E4B66FBDCFC8CB093" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="mediocris">
|
||
<emphasis id="B957DE1C0158E94E4B66FBDCFC8CB093" italics="true" pageId="65" pageNumber="65">Crocidura mediocris</emphasis>
|
||
</taxonomicName>
|
||
is also paler than both
|
||
<taxonomicName id="4C23798D0158E94E4B38FB02FB11B093" authorityName="Esselstyn & Achmadi & Handika & Swanson & Giarla & Rowe" authorityYear="2021" box="[1035,1164,1151,1175]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="tenebrosa">
|
||
<emphasis id="B957DE1C0158E94E4B38FB02FB11B093" box="[1035,1164,1151,1175]" italics="true" pageId="65" pageNumber="65">C. tenebrosa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13181E8B0158E94E4BAAFBFDFD47B0BD" captionStart="FIG" captionStartId="55.[104,146,1261,1285]" captionTargetBox="[104,1615,582,1234]" captionTargetId="figure-102@55.[104,1616,582,1234]" captionTargetPageId="55" captionText="FIG. 27. Images showing the ventral surface of the hind foot and dorsal surfaces of the tail base (approximately 1 cm from rump) and tail tip from the three members of the Small-Bodied Group that are from the northern peninsula: A, Crocidura baletei, LSUMZ 36959 (right hind foot); B, C. lea, LSUMZ 38254 (left hind foot); and C, C. tenebrosa, LSUMZ 39268 (right hind foot). Scale bars represent 5 mm. Within each panel, the upper bar applies to the foot and the lower bar corresponds to both tail images." figureDoi="http://doi.org/10.5281/zenodo.5788883" httpUri="https://zenodo.org/record/5788883/files/figure.png" pageId="65" pageNumber="65">fig. 27C</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C23798D0158E94E4C28FB23FCE5B0BC" box="[795,888,1185,1208]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="parva">
|
||
<emphasis id="B957DE1C0158E94E4C28FB23FCE5B0BC" box="[795,888,1185,1208]" italics="true" pageId="65" pageNumber="65">C. parva</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13181E8B0158E94E4CBBFB22FC7EB0BD" box="[904,995,1184,1209]" captionStart="FIG" captionStartId="59.[108,150,1254,1278]" captionTargetBox="[104,1610,589,1228]" captionTargetId="figure-10@59.[104,1610,589,1228]" captionTargetPageId="59" captionText="FIG. 29. Images showing the ventral surface of the left hind foot and dorsal surfaces of the tail base (approximately 1 cm from rump) and tail tip from the three members of the Small-Bodied Group that are not from the northern peninsula: A, Crocidura levicula, FMNH 213271; B, C. mediocris, FMNH 210566; and C, C. parva, MVZ 237583. Where two scale bars are present within a panel, the upper applies to the foot and the lower to the tail." figureDoi="http://doi.org/10.5281/zenodo.5788887" httpUri="https://zenodo.org/record/5788887/files/figure.png" pageId="65" pageNumber="65">fig. 29C</figureCitation>
|
||
). In cranial proportions,
|
||
<taxonomicName id="4C23798D0158E94E4DC2FB40FCE5B0DE" box="[753,888,1218,1242]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="mediocris">
|
||
<emphasis id="B957DE1C0158E94E4DC2FB40FCE5B0DE" box="[753,888,1218,1242]" italics="true" pageId="65" pageNumber="65">C. mediocris</emphasis>
|
||
</taxonomicName>
|
||
has a greater average condyloincisive length than
|
||
<taxonomicName id="4C23798D0158E94E4CA0FB61FB9DB0FF" box="[915,1024,1251,1275]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="levicula">
|
||
<emphasis id="B957DE1C0158E94E4CA0FB61FB9DB0FF" box="[915,1024,1251,1275]" italics="true" pageId="65" pageNumber="65">C. levicula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C23798D0158E94E4B00FB61FB10B0FE" box="[1075,1165,1251,1274]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="parva">
|
||
<emphasis id="B957DE1C0158E94E4B00FB61FB10B0FE" box="[1075,1165,1251,1274]" italics="true" pageId="65" pageNumber="65">C. parva</emphasis>
|
||
</taxonomicName>
|
||
, but is comparable in this regard to
|
||
<taxonomicName id="4C23798D0158E94E4B36FA86FBDEB118" box="[1029,1091,1284,1308]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="lea">
|
||
<emphasis id="B957DE1C0158E94E4B36FA86FBDEB118" box="[1029,1091,1284,1308]" italics="true" pageId="65" pageNumber="65">C. lea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C23798D0158E94E4B61FA86FB2BB118" box="[1106,1206,1284,1308]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="baletei">
|
||
<emphasis id="B957DE1C0158E94E4B61FA86FB2BB118" box="[1106,1206,1284,1308]" italics="true" pageId="65" pageNumber="65">C. baletei</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C23798D0158E94E4DEFFAA7FCC2B139" authorityName="Esselstyn & Achmadi & Handika & Swanson & Giarla & Rowe" authorityYear="2021" box="[732,863,1317,1341]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="tenebrosa">
|
||
<emphasis id="B957DE1C0158E94E4DEFFAA7FCC2B139" box="[732,863,1317,1341]" italics="true" pageId="65" pageNumber="65">C. tenebrosa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C6A137B50158E94E4C43FAA7FC26B139" box="[880,955,1317,1341]" captionStart="TABLE 8" captionStartId="56.[609,689,226,250]" captionTargetBox="[117,1198,358,1166]" captionTargetPageId="56" captionText="TABLE 8 Descriptive Statisticsa for Craniodental Measurements (mm) for Species of the Small-Bodied Group of Sulawesi Crocidura" httpUri="http://table.plazi.org/id/DF5C52860121E9374D52FF60FC1DB540" pageId="65" pageNumber="65" tableUuid="DF5C52860121E9374D52FF60FC1DB540">table 8</tableCitation>
|
||
). The length of the rostrum, relative to skull length (RL/CIL), is greater on average than in
|
||
<taxonomicName id="4C23798D0158E94E4C47FAE5FC2DB17B" box="[884,944,1383,1407]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="lea">
|
||
<emphasis id="B957DE1C0158E94E4C47FAE5FC2DB17B" box="[884,944,1383,1407]" italics="true" pageId="65" pageNumber="65">C. lea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C23798D0158E94E4C88FAE5FBB4B17B" box="[955,1065,1383,1407]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="levicula">
|
||
<emphasis id="B957DE1C0158E94E4C88FAE5FBB4B17B" box="[955,1065,1383,1407]" italics="true" pageId="65" pageNumber="65">C. levicula</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C23798D0158E94E4B51FAE5FB21B17B" box="[1122,1212,1383,1407]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="parva">
|
||
<emphasis id="B957DE1C0158E94E4B51FAE5FB21B17B" box="[1122,1212,1383,1407]" italics="true" pageId="65" pageNumber="65">C. parva</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13181E8B0158E94E4D80FA0AFD6AB1A4" box="[691,759,1416,1440]" captionStart="FIG" captionStartId="26.[108,150,1414,1435]" captionTargetId="graphics-12@26.[213,1196,224,1265]" captionTargetPageId="26" captionText="FIG. 10. Box plots of relative skull measures showing braincase breadth (BB), interorbital width (IOW), and rostral length (RL) divided by condyloincisive length (CIL) and BB divided by IOW for all species of Sulawesi shrew. Plots show the median, 1st and 3rd quartiles, the maximum value within 1.5 × interquartile range (distance between 1st and 3rd quartiles; IQR), the minimum value within 1.5 × IQR, and outliers (black circles). Sample sizes are shown along the x-axis. Species are ordered according to the species groups used in the text (Thick = Thick-Tailed Group)." figureDoi="http://doi.org/10.5281/zenodo.5789149" httpUri="https://zenodo.org/record/5789149/files/figure.png" pageId="65" pageNumber="65">fig. 10</figureCitation>
|
||
). The relative breadth of the braincase (BB/ CIL) is greater than in
|
||
<taxonomicName id="4C23798D0158E94E4C82FA2BFC6DB1C5" box="[945,1008,1449,1473]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="lea">
|
||
<emphasis id="B957DE1C0158E94E4C82FA2BFC6DB1C5" box="[945,1008,1449,1473]" italics="true" pageId="65" pageNumber="65">C. lea</emphasis>
|
||
</taxonomicName>
|
||
and slightly more than in
|
||
<taxonomicName id="4C23798D0158E94E4C33FA49FCC7B1E6" box="[768,858,1483,1506]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="parva">
|
||
<emphasis id="B957DE1C0158E94E4C33FA49FCC7B1E6" box="[768,858,1483,1506]" italics="true" pageId="65" pageNumber="65">C. parva</emphasis>
|
||
</taxonomicName>
|
||
, but less than in the other Small- Bodied species. In morphometric space, a bivariate plot of condyloincisive length versus braincase breadth shows that
|
||
<taxonomicName id="4C23798D0158E94E4C49F9ACFC66B241" box="[890,1019,1581,1605]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="mediocris">
|
||
<emphasis id="B957DE1C0158E94E4C49F9ACFC66B241" box="[890,1019,1581,1605]" italics="true" pageId="65" pageNumber="65">C. mediocris</emphasis>
|
||
</taxonomicName>
|
||
occupies a widthto-length ratio between the narrow
|
||
<taxonomicName id="4C23798D0158E94E4B02F9CDFBF3B262" box="[1073,1134,1614,1638]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="lea">
|
||
<emphasis id="B957DE1C0158E94E4B02F9CDFBF3B262" box="[1073,1134,1614,1638]" italics="true" pageId="65" pageNumber="65">C. lea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C23798D0158E94E4B96F9CDFD74B283" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="parva">
|
||
<emphasis id="B957DE1C0158E94E4B96F9CDFD74B283" italics="true" pageId="65" pageNumber="65">C. parva</emphasis>
|
||
</taxonomicName>
|
||
and the wide
|
||
<taxonomicName id="4C23798D0158E94E4CBBF9F2FC68B283" box="[904,1013,1647,1671]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="levicula">
|
||
<emphasis id="B957DE1C0158E94E4CBBF9F2FC68B283" box="[904,1013,1647,1671]" italics="true" pageId="65" pageNumber="65">C. levicula</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C23798D0158E94E4B30F9F2FBFBB283" box="[1027,1126,1647,1671]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="baletei">
|
||
<emphasis id="B957DE1C0158E94E4B30F9F2FBFBB283" box="[1027,1126,1647,1671]" italics="true" pageId="65" pageNumber="65">C. baletei</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C23798D0158E94E4B94F9F2FC92B2AC" authorityName="Esselstyn & Achmadi & Handika & Swanson & Giarla & Rowe" authorityYear="2021" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="tenebrosa">
|
||
<emphasis id="B957DE1C0158E94E4B94F9F2FC92B2AC" italics="true" pageId="65" pageNumber="65">C. tenebrosa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13181E8B0158E94E4C2FF912FCE8B2AC" box="[796,885,1680,1704]" captionStart="FIG" captionStartId="54.[108,150,816,837]" captionTargetBox="[109,1204,234,787]" captionTargetId="graphics-423@54.[179,547,225,728]" captionTargetPageId="54" captionText="FIG. 26. Bivariate plots showing A, variation in the condyloincisive length and braincase breadth and B, the first two axes from a principal components analysis of 12 cranial measurements among all members of the Small-Bodied Group. Loadings and variance explained are given in table 9." figureDoi="http://doi.org/10.5281/zenodo.5788881" httpUri="https://zenodo.org/record/5788881/files/figure.png" pageId="65" pageNumber="65">fig. 26A</figureCitation>
|
||
). A PCA of cranial dimensions shows that specimens of
|
||
<taxonomicName id="4C23798D0158E94E4CE6F930FBFCB2CE" box="[981,1121,1714,1738]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="65" pageNumber="65" phylum="Chordata" rank="species" species="mediocris">
|
||
<emphasis id="B957DE1C0158E94E4CE6F930FBFCB2CE" box="[981,1121,1714,1738]" italics="true" pageId="65" pageNumber="65">C. mediocris</emphasis>
|
||
</taxonomicName>
|
||
occupy moderately distinct morphometric space, overlapping primarily with individuals of
|
||
<taxonomicName id="4C23798D015BE94D4ED8FE81FDB7B51F" box="[491,554,259,283]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="66" pageNumber="66" phylum="Chordata" rank="species" species="lea">
|
||
<emphasis id="B957DE1C015BE94D4ED8FE81FDB7B51F" box="[491,554,259,283]" italics="true" pageId="66" pageNumber="66">C. lea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C23798D015BE94D4D57FE81FF34B53F" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="66" pageNumber="66" phylum="Chordata" rank="species" species="parva">
|
||
<emphasis id="B957DE1C015BE94D4D57FE81FF34B53F" italics="true" pageId="66" pageNumber="66">C. parva</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13181E8B015BE94D4F84FEA6FE8FB538" box="[183,274,292,316]" captionStart="FIG" captionStartId="54.[108,150,816,837]" captionTargetBox="[109,1204,234,787]" captionTargetId="graphics-423@54.[179,547,225,728]" captionTargetPageId="54" captionText="FIG. 26. Bivariate plots showing A, variation in the condyloincisive length and braincase breadth and B, the first two axes from a principal components analysis of 12 cranial measurements among all members of the Small-Bodied Group. Loadings and variance explained are given in table 9." figureDoi="http://doi.org/10.5281/zenodo.5788881" httpUri="https://zenodo.org/record/5788881/files/figure.png" pageId="66" pageNumber="66">fig. 26A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3395185015BE94D4FBFFEC7FEC3B627" pageId="66" pageNumber="66" type="discussion">
|
||
<paragraph id="8B9C020E015BE94D4FBFFEC7FEC3B627" blockId="66.[108,637,226,977]" pageId="66" pageNumber="66">
|
||
COMMENTS:
|
||
<taxonomicName id="4C23798D015BE94D4E12FEC7FD98B559" box="[289,517,325,349]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="66" pageNumber="66" phylum="Chordata" rank="species" species="mediocris">
|
||
<emphasis id="B957DE1C015BE94D4E12FEC7FD98B559" box="[289,517,325,349]" italics="true" pageId="66" pageNumber="66">Crocidura mediocris</emphasis>
|
||
</taxonomicName>
|
||
is closely related to
|
||
<taxonomicName id="4C23798D015BE94D4FEFFEE4FEA5B57A" box="[220,312,358,382]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="66" pageNumber="66" phylum="Chordata" rank="species" species="parva">
|
||
<emphasis id="B957DE1C015BE94D4FEFFEE4FEA5B57A" box="[220,312,358,382]" italics="true" pageId="66" pageNumber="66">C. parva</emphasis>
|
||
</taxonomicName>
|
||
. See the
|
||
<taxonomicName id="4C23798D015BE94D4EAFFEE4FE64B57A" box="[412,505,358,382]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="66" pageNumber="66" phylum="Chordata" rank="species" species="parva">
|
||
<emphasis id="B957DE1C015BE94D4EAFFEE4FE64B57A" box="[412,505,358,382]" italics="true" pageId="66" pageNumber="66">C. parva</emphasis>
|
||
</taxonomicName>
|
||
account for a discussion of phylogenetic relationships between these two phenotypically similar species. For coalescent species delimitation results, see the
|
||
<taxonomicName id="4C23798D015BE94D4FFBFE69FED6B606" authorityName="Esselstyn & Achmadi & Handika & Swanson & Giarla & Rowe" authorityYear="2021" box="[200,331,490,514]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="66" pageNumber="66" phylum="Chordata" rank="species" species="normalis">
|
||
<emphasis id="B957DE1C015BE94D4FFBFE69FED6B606" box="[200,331,490,514]" italics="true" pageId="66" pageNumber="66">C. normalis</emphasis>
|
||
</taxonomicName>
|
||
(Ordinary Group) and
|
||
<taxonomicName id="4C23798D015BE94D4D57FE69FF37B627" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="66" pageNumber="66" phylum="Chordata" rank="species" species="parva">
|
||
<emphasis id="B957DE1C015BE94D4D57FE69FF37B627" italics="true" pageId="66" pageNumber="66">C. parva</emphasis>
|
||
</taxonomicName>
|
||
accounts below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3395185015BE94D4FBFFDAEFE0DB7D5" pageId="66" pageNumber="66" type="materials_examined">
|
||
<paragraph id="8B9C020E015BE94D4FBFFDAEFE0DB7D5" blockId="66.[108,637,226,977]" pageId="66" pageNumber="66">
|
||
SPECIMENS EXAMINED:
|
||
<materialsCitation id="3B4B0853015BE94D4EA5FDAFFDA7B661" ID-GBIF-Occurrence="3421079382" collectionCode="FMNH" location="Mt. Balease" pageId="66" pageNumber="66" specimenCode="FMNH 210563-210575, 210602, 210604-210606" specimenCount="1">
|
||
<location id="8EFC54D5015BE94D4EA5FDAFFD85B640" LSID="urn:lsid:plazi:treatment:038AB3180126E94D4DC9FA07FE0DB7D5:8EFC54D5015BE94D4EA5FDAFFD85B640" box="[406,536,556,580]" name="Mt. Balease" pageId="66" pageNumber="66">Mt. Balease</location>
|
||
(
|
||
<specimenCode id="DB85AA75015BE94D4D19FDAEFE86B661" pageId="66" pageNumber="66">FMNH 210563–210575</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4E1BFDCFFEE4B661" box="[296,377,589,613]" pageId="66" pageNumber="66">210602</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4EB4FDCFFDABB661" box="[391,566,589,613]" pageId="66" pageNumber="66">210604–210606</specimenCode>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4B0853015BE94D4D70FDCCFF23B683" ID-GBIF-Occurrence="3421079367" collectionCode="MZB" pageId="66" pageNumber="66" specimenCode="MZB 43007" specimenCount="1">
|
||
<specimenCode id="DB85AA75015BE94D4D70FDCCFF2FB683" pageId="66" pageNumber="66">MZB 43007</specimenCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4B0853015BE94D4FFBFDEDFE1CB6CC" ID-GBIF-Occurrence="3421079335" collectionCode="FMNH" location="Mt. Gandang Dewata" pageId="66" pageNumber="66" specimenCode="FMNH 218617, 218619, 218620, 218622-218638, 218640- 218642, 218644-218649" specimenCount="1">
|
||
<location id="8EFC54D5015BE94D4FFBFDEDFE24B682" LSID="urn:lsid:plazi:treatment:038AB3180126E94D4DC9FA07FE0DB7D5:8EFC54D5015BE94D4FFBFDEDFE24B682" box="[200,441,622,646]" name="Mt. Gandang Dewata" pageId="66" pageNumber="66">Mt. Gandang Dewata</location>
|
||
(
|
||
<specimenCode id="DB85AA75015BE94D4EF9FDEDFDEAB682" box="[458,631,622,647]" pageId="66" pageNumber="66">FMNH 218617</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4F5FFD0DFF5CB6A3" box="[108,193,655,679]" pageId="66" pageNumber="66">218619</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4FE8FD0DFEADB6A3" box="[219,304,655,679]" pageId="66" pageNumber="66">218620</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4E79FD0DFD9CB6A3" box="[330,513,655,679]" pageId="66" pageNumber="66">218622–218638</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4D28FD0DFF20B6CC" pageId="66" pageNumber="66">218640– 218642</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4FF5FD32FEEBB6CC" box="[198,374,688,712]" pageId="66" pageNumber="66">218644–218649</specimenCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4B0853015BE94D4EB4FD33FF55B6ED" ID-GBIF-Occurrence="3421079432" collectionCode="MVZ" location="Mt. Latimojong" pageId="66" pageNumber="66" specimenCode="MVZ 237574" specimenCount="1">
|
||
<location id="8EFC54D5015BE94D4EB4FD33FDACB6CC" LSID="urn:lsid:plazi:treatment:038AB3180126E94D4DC9FA07FE0DB7D5:8EFC54D5015BE94D4EB4FD33FDACB6CC" box="[391,561,688,712]" name="Mt. Latimojong" pageId="66" pageNumber="66">Mt. Latimojong</location>
|
||
(
|
||
<specimenCode id="DB85AA75015BE94D4D0DFD33FF5CB6ED" pageId="66" pageNumber="66">MVZ 237574</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B4B0853015BE94D4FEAFD50FDE1B728" ID-GBIF-Occurrence="3421079326" collectionCode="MWFB" location="Mt. Mekongga" pageId="66" pageNumber="66" specimenCode="MWFB 8092, 8104, 8121, 8131-8136, 8140-8142, 8144, 8145, 8148, 8149, 8152, 8153, 13508, 13510, 13513, 13514" specimenCount="1">
|
||
<location id="8EFC54D5015BE94D4FEAFD50FE1CB6EE" LSID="urn:lsid:plazi:treatment:038AB3180126E94D4DC9FA07FE0DB7D5:8EFC54D5015BE94D4FEAFD50FE1CB6EE" box="[217,385,722,746]" name="Mt. Mekongga" pageId="66" pageNumber="66">Mt. Mekongga</location>
|
||
(
|
||
<specimenCode id="DB85AA75015BE94D4EA5FD50FDB2B6EE" box="[406,559,722,746]" pageId="66" pageNumber="66">MWFB 8092</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4D73FD50FDEAB6EE" box="[576,631,722,746]" pageId="66" pageNumber="66">8104</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4F5FFD71FF3FB70F" box="[108,162,755,779]" pageId="66" pageNumber="66">8121</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4F9CFD71FEB5B70F" box="[175,296,755,779]" pageId="66" pageNumber="66">8131–8136</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4E06FD71FE33B70F" box="[309,430,755,779]" pageId="66" pageNumber="66">8140–8142</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4E88FD71FE6CB70F" box="[443,497,755,779]" pageId="66" pageNumber="66">8144</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4ECDFD71FDA9B70F" box="[510,564,755,779]" pageId="66" pageNumber="66">8145</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4D71FD71FDE5B70F" box="[578,632,755,779]" pageId="66" pageNumber="66">8148</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4F5FFC96FF3EB728" box="[108,163,788,812]" pageId="66" pageNumber="66">8149</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4F83FC96FF7AB728" box="[176,231,788,812]" pageId="66" pageNumber="66">8152</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4FC7FC96FEB6B728" box="[244,299,788,812]" pageId="66" pageNumber="66">8153</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4E0BFC96FEE1B728" box="[312,380,788,812]" pageId="66" pageNumber="66">13508</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4EBAFC96FE50B728" box="[393,461,788,812]" pageId="66" pageNumber="66">13510</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4EE9FC96FD83B728" box="[474,542,788,812]" pageId="66" pageNumber="66">13513</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4D18FC96FDEDB728" box="[555,624,788,812]" pageId="66" pageNumber="66">13514</specimenCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4B0853015BE94D4F5FFCB7FDD6B76A" ID-GBIF-Occurrence="3421079352" collectionCode="FMNH" location="Salu Tiwo" pageId="66" pageNumber="66" specimenCode="FMNH 218614-218616, 218618, 218621, 218639, 218643, 218650, 218651" specimenCount="1">
|
||
<location id="8EFC54D5015BE94D4F5FFCB7FF79B749" LSID="urn:lsid:plazi:treatment:038AB3180126E94D4DC9FA07FE0DB7D5:8EFC54D5015BE94D4F5FFCB7FF79B749" box="[108,228,821,845]" name="Salu Tiwo" pageId="66" pageNumber="66">Salu Tiwo</location>
|
||
(
|
||
<specimenCode id="DB85AA75015BE94D4FC8FCB7FD8DB749" box="[251,528,821,845]" pageId="66" pageNumber="66">FMNH 218614–218616</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4D17FCB7FDEAB749" box="[548,631,821,845]" pageId="66" pageNumber="66">218618</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4F5FFCD4FF23B76A" box="[108,190,854,878]" pageId="66" pageNumber="66">218621</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4FFFFCD4FE83B76A" box="[204,286,854,878]" pageId="66" pageNumber="66">218639</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4E1FFCD4FEE3B76A" box="[300,382,854,878]" pageId="66" pageNumber="66">218643</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4EBFFCD4FE43B76A" box="[396,478,854,878]" pageId="66" pageNumber="66">218650</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4EDFFCD4FDDDB76A" box="[492,576,854,878]" pageId="66" pageNumber="66">218651</specimenCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4B0853015BE94D4D65FCD4FE8AB7B4" ID-GBIF-Occurrence="3421079311" collectionCode="LSUMZ, MVZ" location="Mt. Torompupu" pageId="66" pageNumber="66" specimenCode="LSUMZ 39452-39457, 39555, MVZ 238109-238114" specimenCount="1">
|
||
<location id="8EFC54D5015BE94D4D65FCD4FF71B78A" LSID="urn:lsid:plazi:treatment:038AB3180126E94D4DC9FA07FE0DB7D5:8EFC54D5015BE94D4D65FCD4FF71B78A" name="Mt. Torompupu" pageId="66" pageNumber="66">Mt. Torompupu</location>
|
||
(
|
||
<specimenCode id="DB85AA75015BE94D4FCAFCF5FE77B78B" box="[249,490,887,911]" pageId="66" pageNumber="66">LSUMZ 39452–39457</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4EC7FCF5FDAAB78B" box="[500,567,887,911]" pageId="66" pageNumber="66">39555</specimenCode>
|
||
;
|
||
<specimenCode id="DB85AA75015BE94D4D72FCF5FE8AB7B4" pageId="66" pageNumber="66">MVZ 238109–238114</specimenCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B4B0853015BE94D4E12FC1BFE0DB7D5" ID-GBIF-Occurrence="3421079391" collectionCode="NMV, C, Z" pageId="66" pageNumber="66" specimenCode="NMV C40246, C40251, C40253, C40274, C40278, Z62199" specimenCount="1">
|
||
<specimenCode id="DB85AA75015BE94D4E12FC1BFE26B7B4" box="[289,443,920,944]" pageId="66" pageNumber="66">NMV C40246</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4EF6FC1AFD87B7B4" box="[453,538,920,944]" pageId="66" pageNumber="66">C40251</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4D10FC1AFDE5B7B4" box="[547,632,920,944]" pageId="66" pageNumber="66">C40253</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4F5FFC3BFF5FB7D5" box="[108,194,953,977]" pageId="66" pageNumber="66">C40274</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4FFEFC3BFEBEB7D5" box="[205,291,953,977]" pageId="66" pageNumber="66">C40278</specimenCode>
|
||
,
|
||
<specimenCode id="DB85AA75015BE94D4E1DFC3BFE18B7D5" box="[302,389,953,977]" pageId="66" pageNumber="66">Z62199</specimenCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |