628 lines
74 KiB
XML
628 lines
74 KiB
XML
<document id="1A7D0B3423B49ECDF32B7ED2256871D7" ID-CLB-Dataset="30792" ID-DOI="10.11646/zootaxa.4378.2.7" ID-GBIF-Dataset="2525291e-a481-499a-ab8d-df5a615cefa3" ID-ISSN="1175-5326" ID-Zenodo-Dep="1169533" ID-ZooBank="1B7051F6-FBA0-4FAF-88F0-4DD3FE823D4C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1518134972672" checkinUser="plazi" docAuthor="Kohlmann, Bert, Arriaga-Jiménez, Alfonsina & Rös, Matthias" docDate="2018" docId="038A366F762FFFF9FF5CFA43FB1EBD69" docLanguage="en" docName="zootaxa.4378.2.7.pdf" docOrigin="Zootaxa 4378 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Canthidium quercetorum Kohlmann & Arriaga-Jiménez & Rös 2018, new species" docType="treatment" docVersion="7" lastPageNumber="277" masterDocId="FFB34E17762FFFFDFFCBFFC8FFF8BE76" masterDocTitle="An unusual new species of Canthidium (Coleoptera: Scarabaeidae: Scarabaeinae) from Oaxaca, Mexico" masterLastPageNumber="278" masterPageNumber="273" pageNumber="273" updateTime="1698520576407" updateUser="ExternalLinkService">
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<mods:mods id="2CD574AFF16AB7F208C9B32D9876C445" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2AE379BD619046482C24E91F939A7216">An unusual new species of Canthidium (Coleoptera: Scarabaeidae: Scarabaeinae) from Oaxaca, Mexico</mods:title>
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<mods:namePart id="184DBCBD2A0F05B54BF3DF6CBF0EC397">Kohlmann, Bert</mods:namePart>
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<mods:namePart id="35E00470A75D0754EE5FFCE8FA93B58F">Arriaga-Jiménez, Alfonsina</mods:namePart>
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<mods:namePart id="D430D86201DDF3D3E6E22D39EC153840">Rös, Matthias</mods:namePart>
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<mods:title id="53115063AA84CE71FF0B996E98DA19A9">Zootaxa</mods:title>
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<mods:date id="7C1EA0A6561D4E76E974A43D2AA8AF9E">2018</mods:date>
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<mods:number id="740467A74C5261F1C58BCB46B49E0F9B">2018-02-08</mods:number>
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<treatment id="038A366F762FFFF9FF5CFA43FB1EBD69" ID-DOI="http://doi.org/10.5281/zenodo.5945537" ID-GBIF-Taxon="140569273" ID-Zenodo-Dep="5945537" LSID="urn:lsid:plazi:treatment:038A366F762FFFF9FF5CFA43FB1EBD69" httpUri="http://treatment.plazi.org/id/038A366F762FFFF9FF5CFA43FB1EBD69" lastPageId="4" lastPageNumber="277" pageId="0" pageNumber="273">
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<subSubSection id="C339D4F2762FFFFDFF5CFA43FB47BBD3" box="[151,1215,1419,1445]" pageId="0" pageNumber="273" type="nomenclature">
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<paragraph id="8B9C8779762FFFFDFF5CFA43FB47BBD3" blockId="0.[151,1436,1419,2031]" box="[151,1215,1419,1445]" pageId="0" pageNumber="273">
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<heading id="D0D43015762FFFFDFF5CFA43FB47BBD3" bold="true" box="[151,1215,1419,1445]" fontSize="11" level="1" pageId="0" pageNumber="273" reason="1">
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<emphasis id="B9575B6B762FFFFDFF5CFA43FB47BBD3" bold="true" box="[151,1215,1419,1445]" pageId="0" pageNumber="273">
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<taxonomicName id="4C23FCFA762FFFFDFF5CFA43FC69BBD3" ID-CoL="699F9" authority="Kohlmann, Arriaga-Jimenez & Ros" authorityName="Kohlmann & Arriaga-Jiménez & Rös" authorityYear="2018" box="[151,913,1419,1445]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="273" phylum="Arthropoda" rank="species" species="quercetorum" status="sp. nov.">
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<emphasis id="B9575B6B762FFFFDFF5CFA43FE32BBD3" bold="true" box="[151,458,1419,1445]" italics="true" pageId="0" pageNumber="273">Canthidium quercetorum</emphasis>
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Kohlmann, Arriaga-Jiménez & Rös
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</taxonomicName>
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,
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<taxonomicNameLabel id="A264E610762FFFFDFC55FA43FBD6BBD3" box="[926,1070,1419,1445]" pageId="0" pageNumber="273" rank="species">new species</taxonomicNameLabel>
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(
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<figureCitation id="13189BFC762FFFFDFBF6FA43FB48BBD3" box="[1085,1200,1419,1445]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,244,1602,1624]" captionStartId-1="2.[151,244,1546,1568]" captionStartId-2="3.[151,244,1442,1464]" captionTargetBox-0="[242,1343,219,1537]" captionTargetBox-1="[190,1402,227,1489]" captionTargetBox-2="[433,1154,376,1413]" captionTargetId-0="figure@1.[229,1358,194,1578]" captionTargetId-1="figure@2.[152,1435,191,1525]" captionTargetId-2="figure@3.[419,1168,359,1431]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1.Illustration of the dorsal habitus of a female Canthidium quercetorum new species." captionText-1="FIGURE 2. Canthidium quercetorum new species. A, the dorsal habitus; B, ventral habitus; C, lateral habitus; D, male protibia; E, female protibia;F, pygidium;and G, aedeagus." captionText-2="FIGURE 3. Sampling site, A, (grey point) of Canthidium quercetorum new species in Oaxaca (grey area), Mexico (white area). Satelliteimages ofCanthidium quercetorumsampling sites inadeciduous oakforest B,at the dry season (left)andC,at the endof the rainy season (right) (March 2017, November 2014). Bright colours on the left image indicate a forest with trees that have lost their leaves;whereas green vegetationon the same image shows pine-oakforest at higher altitudes,where Canthidium quercetorumhas not been collected.On the right image,oak forest appears withnearly maximum leafdensity at the endof the rainy season.Distributionof D, dry oak forest (dark brown area), situated between approximately 1800 and 2400 m. Sampling site of Canthidium quercetorum is represented by the white rectangle.On the left lower sideof the imagethe urbanarea of the Oaxaca Metropolitan Region can be seen. The dark green area on the right upper side of the image represents pine-oak and pine forest (highest elevation is around 3100 m). Satellite image was provided by GoogleEarth, and oak forest distribution by the Instituto Nacional de Estadística, Geografía e Informática." httpUri-0="https://zenodo.org/record/1169535/files/figure.png" httpUri-1="https://zenodo.org/record/1169537/files/figure.png" httpUri-2="https://zenodo.org/record/1169539/files/figure.png" pageId="0" pageNumber="273">Figs. 1–3</figureCitation>
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.)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C339D4F2762FFFFDFF5CFA65FAE2B804" pageId="0" pageNumber="273" type="diagnosis">
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<paragraph id="8B9C8779762FFFFDFF5CFA65FAE2B804" blockId="0.[151,1436,1419,2031]" pageId="0" pageNumber="273">
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<emphasis id="B9575B6B762FFFFDFF5CFA65FEF4BBB2" bold="true" box="[151,268,1453,1476]" pageId="0" pageNumber="273">Diagnosis.</emphasis>
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This species is distinguished by the following combination of characters: body sub-globose and dorsally dark green in colour; head and pronotum uniformly and strongly punctate; frons swollen; frontoclypeal region with a small, smooth swelling medial to each eye; eye dorsally at posterior end of gena two facets wide, interocular distance separated by approximately 14–16 times maximal eye width; antennal lamellae black and sericeous; pronotal surface smooth with sparse micropunctures, basal pronotal border not margined; elytra with eight clearly punctate striae, surface shagreen; pygidium shagreen and lightly punctate; the inner apical margin of the female protibia is not projected at all.
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</paragraph>
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</subSubSection>
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<subSubSection id="C339D4F2762FFFFDFF0CF9B5FC49B8E2" box="[199,945,1661,1684]" pageId="0" pageNumber="273" type="materials_examined">
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<paragraph id="8B9C8779762FFFFDFF0CF9B5FC49B8E2" blockId="0.[151,1436,1419,2031]" box="[199,945,1661,1684]" pageId="0" pageNumber="273">
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<emphasis id="B9575B6B762FFFFDFF0CF9B5FE38B8E2" bold="true" box="[199,448,1661,1684]" pageId="0" pageNumber="273">
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<typeStatus id="549839DB762FFFFDFF0CF9B5FF07B8E2" box="[199,255,1661,1684]" pageId="0" pageNumber="273">Type</typeStatus>
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locality.
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<collectingRegion id="49E7499B762FFFFDFEAFF9B5FE43B8E2" box="[356,443,1661,1684]" country="Mexico" name="Mexico" pageId="0" pageNumber="273">
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<collectingCountry id="F334C7E9762FFFFDFEAFF9B5FE43B8E2" box="[356,443,1661,1684]" name="Mexico" pageId="0" pageNumber="273">Mexico</collectingCountry>
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</collectingRegion>
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:
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</emphasis>
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<collectingRegion id="49E7499B762FFFFDFE0DF9B5FDE2B8E2" box="[454,538,1661,1684]" country="Mexico" name="Oaxaca" pageId="0" pageNumber="273">Oaxaca</collectingRegion>
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: Reserva Comunitaria San Pablo Etla.
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</paragraph>
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</subSubSection>
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<subSubSection id="C339D4F2762FFFFFFF0CF968FAD4B97C" lastPageId="2" lastPageNumber="275" pageId="0" pageNumber="273" type="description">
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<paragraph id="8B9C8779762FFFFDFF0CF968FE36B88A" blockId="0.[151,1436,1419,2031]" pageId="0" pageNumber="273">
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<emphasis id="B9575B6B762FFFFDFF0CF968FE21B8C1" bold="true" box="[199,473,1696,1719]" pageId="0" pageNumber="273">
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Description of
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<typeStatus id="549839DB762FFFFDFEBBF968FE2CB8C1" box="[368,468,1696,1719]" pageId="0" pageNumber="273" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Length:
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<quantity id="4CDB2A9C762FFFFDFDF0F968FD77B8C1" box="[571,655,1696,1719]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="0" pageNumber="273" unit="mm" value="5.1">5.1 mm</quantity>
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, humeral width:
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<quantity id="4CDB2A9C762FFFFDFC8DF968FC64B8C1" box="[838,924,1696,1719]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="0" pageNumber="273" unit="mm" value="3.5">3.5 mm</quantity>
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. Body form sub-globose (
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<figureCitation id="13189BFC762FFFFDFB7FF968FB06B8C1" box="[1204,1278,1696,1719]" captionStart="FIGURE 1" captionStartId="1.[151,244,1602,1624]" captionTargetBox="[242,1343,219,1537]" captionTargetId="figure@1.[229,1358,194,1578]" captionTargetPageId="1" captionText="FIGURE 1.Illustration of the dorsal habitus of a female Canthidium quercetorum new species." httpUri="https://zenodo.org/record/1169535/files/figure.png" pageId="0" pageNumber="273">Figs. 1</figureCitation>
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,
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<figureCitation id="13189BFC762FFFFDFAC6F968FAA8B8C1" box="[1293,1360,1696,1719]" captionStart="FIGURE 2" captionStartId="2.[151,244,1546,1568]" captionTargetBox="[190,1402,227,1489]" captionTargetId="figure@2.[152,1435,191,1525]" captionTargetPageId="2" captionText="FIGURE 2. Canthidium quercetorum new species. A, the dorsal habitus; B, ventral habitus; C, lateral habitus; D, male protibia; E, female protibia;F, pygidium;and G, aedeagus." httpUri="https://zenodo.org/record/1169537/files/figure.png" pageId="0" pageNumber="273">2A–C</figureCitation>
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). Head and pronotum dark green and shiny, elytra brownish black with green reflections (
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<figureCitation id="13189BFC762FFFFDFBCAF90BFBAFB8AC" box="[1025,1111,1731,1754]" captionStart="FIGURE 2" captionStartId="2.[151,244,1546,1568]" captionTargetBox="[190,1402,227,1489]" captionTargetId="figure@2.[152,1435,191,1525]" captionTargetPageId="2" captionText="FIGURE 2. Canthidium quercetorum new species. A, the dorsal habitus; B, ventral habitus; C, lateral habitus; D, male protibia; E, female protibia;F, pygidium;and G, aedeagus." httpUri="https://zenodo.org/record/1169537/files/figure.png" pageId="0" pageNumber="273">Fig. 2A</figureCitation>
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). Head and pronotum strongly punctate and devoid of setae.
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</paragraph>
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<paragraph id="8B9C8779762FFFFDFF0CF8C0FC90B9DC" blockId="0.[151,1436,1419,2031]" pageId="0" pageNumber="273">
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Clypeus rugosely punctate, apex strongly bidentate, median emargination broadly V-shaped; eye dorsally narrow 8– 9 facets at widest portion, at posterior end of gena very narrow, two facets wide, eyes separated approximately 14–16 times maximum eye width (
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<figureCitation id="13189BFC762FFFFDFE0AF885FDE0B912" box="[449,536,1869,1892]" captionStart="FIGURE 2" captionStartId="2.[151,244,1546,1568]" captionTargetBox="[190,1402,227,1489]" captionTargetId="figure@2.[152,1435,191,1525]" captionTargetPageId="2" captionText="FIGURE 2. Canthidium quercetorum new species. A, the dorsal habitus; B, ventral habitus; C, lateral habitus; D, male protibia; E, female protibia;F, pygidium;and G, aedeagus." httpUri="https://zenodo.org/record/1169537/files/figure.png" pageId="0" pageNumber="273">Fig. 2A</figureCitation>
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). Head, frons, and vertex strongly punctate; frons swollen; two small, black, smooth swellings between the eyes (
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<figureCitation id="13189BFC762FFFFDFE01F8B8FDD9B9F1" box="[458,545,1904,1927]" captionStart="FIGURE 2" captionStartId="2.[151,244,1546,1568]" captionTargetBox="[190,1402,227,1489]" captionTargetId="figure@2.[152,1435,191,1525]" captionTargetPageId="2" captionText="FIGURE 2. Canthidium quercetorum new species. A, the dorsal habitus; B, ventral habitus; C, lateral habitus; D, male protibia; E, female protibia;F, pygidium;and G, aedeagus." httpUri="https://zenodo.org/record/1169537/files/figure.png" pageId="0" pageNumber="273">Fig. 2A</figureCitation>
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). On either side of the vertex midline a small impunctate, shagreen area is present. Head surface lightly shagreen at base. Antennae blackish sericeous.
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</paragraph>
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<paragraph id="8B9C8779762FFFFDFF0CF87DFBBCB999" blockId="0.[151,1436,1419,2031]" pageId="0" pageNumber="273">
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Pronotum with smooth surface, sparse micropunctures, shiny, and strongly and uniformly punctate (
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<figureCitation id="13189BFC762FFFFDFB3BF87DFABEB9BA" box="[1264,1350,1973,1996]" captionStart="FIGURE 2" captionStartId="2.[151,244,1546,1568]" captionTargetBox="[190,1402,227,1489]" captionTargetId="figure@2.[152,1435,191,1525]" captionTargetPageId="2" captionText="FIGURE 2. Canthidium quercetorum new species. A, the dorsal habitus; B, ventral habitus; C, lateral habitus; D, male protibia; E, female protibia;F, pygidium;and G, aedeagus." httpUri="https://zenodo.org/record/1169537/files/figure.png" pageId="0" pageNumber="273">Fig. 2A</figureCitation>
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); lateral fovea inconspicuous and simple; posterior margin lacking elongate punctures or groove.
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</paragraph>
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<caption id="DF5CD7F1762EFFFCFF5CF98AFBD7B821" box="[151,1071,1602,1624]" httpUri="https://zenodo.org/record/1169535/files/figure.png" pageId="1" pageNumber="274" startId="1.[151,244,1602,1624]" targetBox="[242,1343,219,1537]" targetPageId="1">
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<paragraph id="8B9C8779762EFFFCFF5CF98AFBD7B821" blockId="1.[151,1071,1602,1624]" box="[151,1071,1602,1624]" pageId="1" pageNumber="274">
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<emphasis id="B9575B6B762EFFFCFF5CF98AFEF5B82E" bold="true" box="[151,269,1602,1624]" pageId="1" pageNumber="274">FIGURE 1.</emphasis>
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Illustration of the dorsal habitus of a female
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<taxonomicName id="4C23FCFA762EFFFCFD77F98AFC55B821" authority="Kohlmann & Arriaga-Jiménez & Rös, 2018" authorityName="Kohlmann & Arriaga-Jiménez & Rös" authorityYear="2018" box="[700,941,1602,1623]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="274" phylum="Arthropoda" rank="species" species="quercetorum" status="sp. nov.">
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<emphasis id="B9575B6B762EFFFCFD77F98AFC55B821" box="[700,941,1602,1623]" italics="true" pageId="1" pageNumber="274">Canthidium quercetorum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A264E610762EFFFCFC78F98AFBD1B82E" box="[947,1065,1602,1624]" pageId="1" pageNumber="274" rank="species">
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<emphasis id="B9575B6B762EFFFCFC78F98AFBD1B82E" bold="true" box="[947,1065,1602,1624]" pageId="1" pageNumber="274">new species</emphasis>
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</taxonomicNameLabel>
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.
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</paragraph>
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</caption>
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<paragraph id="8B9C8779762EFFFCFF0CF9B5FAACB8C1" blockId="1.[151,1436,1661,2031]" pageId="1" pageNumber="274">
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Elytral striae consisting of clear shallow, thin lines, interrupted by fairly evenly spaced oval punctures (spacing approximately at 3–5 the maximum length of one puncture); intervals finely punctate; surface shagreen (
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<figureCitation id="13189BFC762EFFFCFB39F968FAB0B8C1" box="[1266,1352,1696,1719]" captionStart="FIGURE 2" captionStartId="2.[151,244,1546,1568]" captionTargetBox="[190,1402,227,1489]" captionTargetId="figure@2.[152,1435,191,1525]" captionTargetPageId="2" captionText="FIGURE 2. Canthidium quercetorum new species. A, the dorsal habitus; B, ventral habitus; C, lateral habitus; D, male protibia; E, female protibia;F, pygidium;and G, aedeagus." httpUri="https://zenodo.org/record/1169537/files/figure.png" pageId="1" pageNumber="274">Fig. 2A</figureCitation>
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).
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</paragraph>
|
||
<paragraph id="8B9C8779762EFFFCFF0CF90BFD0BB969" blockId="1.[151,1436,1661,2031]" pageId="1" pageNumber="274">
|
||
Proepisternum excavate anteriorly, surface shagreen and finely punctate. Sternellum shagreen and with a line of setiferous punctures along base. Mesosternum shagreen and finely punctate. Metasternum punctate, less so at the midline (
|
||
<figureCitation id="13189BFC762EFFFCFF54F8C0FF0AB969" box="[159,242,1800,1823]" captionStart="FIGURE 2" captionStartId="2.[151,244,1546,1568]" captionTargetBox="[190,1402,227,1489]" captionTargetId="figure@2.[152,1435,191,1525]" captionTargetPageId="2" captionText="FIGURE 2. Canthidium quercetorum new species. A, the dorsal habitus; B, ventral habitus; C, lateral habitus; D, male protibia; E, female protibia;F, pygidium;and G, aedeagus." httpUri="https://zenodo.org/record/1169537/files/figure.png" pageId="1" pageNumber="274">Fig. 2B</figureCitation>
|
||
), lateral-lobe punctures forming coarse rugulae.
|
||
</paragraph>
|
||
<paragraph id="8B9C8779762EFFFCFF0CF8E3FB30B934" blockId="1.[151,1436,1661,2031]" box="[199,1224,1835,1858]" pageId="1" pageNumber="274">
|
||
Sternites 3–6 shagreen, sternites 3–5 with a row of small punctures along front border (
|
||
<figureCitation id="13189BFC762EFFFCFBA3F8E3FB43B934" box="[1128,1211,1835,1858]" captionStart="FIGURE 2" captionStartId="2.[151,244,1546,1568]" captionTargetBox="[190,1402,227,1489]" captionTargetId="figure@2.[152,1435,191,1525]" captionTargetPageId="2" captionText="FIGURE 2. Canthidium quercetorum new species. A, the dorsal habitus; B, ventral habitus; C, lateral habitus; D, male protibia; E, female protibia;F, pygidium;and G, aedeagus." httpUri="https://zenodo.org/record/1169537/files/figure.png" pageId="1" pageNumber="274">Fig. 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9C8779762EFFFCFF0CF885FC8DB9DC" blockId="1.[151,1436,1661,2031]" pageId="1" pageNumber="274">
|
||
Protibia with three teeth on external border, inner apical margin of protibia produced into a triangular anterior projection and slightly bent downwards; apical spur simple and straight (
|
||
<figureCitation id="13189BFC762EFFFCFC28F8B8FBB8B9F1" box="[995,1088,1904,1927]" captionStart="FIGURE 2" captionStartId="2.[151,244,1546,1568]" captionTargetBox="[190,1402,227,1489]" captionTargetId="figure@2.[152,1435,191,1525]" captionTargetPageId="2" captionText="FIGURE 2. Canthidium quercetorum new species. A, the dorsal habitus; B, ventral habitus; C, lateral habitus; D, male protibia; E, female protibia;F, pygidium;and G, aedeagus." httpUri="https://zenodo.org/record/1169537/files/figure.png" pageId="1" pageNumber="274">Fig. 2D</figureCitation>
|
||
). Profemora, mesofemora, and metafemora with ventral surface finely punctate and finely shagreen.
|
||
</paragraph>
|
||
<paragraph id="8B9C8779762EFFFCFF0CF87DFF44B999" blockId="1.[151,1436,1661,2031]" pageId="1" pageNumber="274">
|
||
Pygidium slightly convex, shagreen, and shiny with very fine and shallow punctures (
|
||
<figureCitation id="13189BFC762EFFFCFBAAF87DFB4CB9BA" box="[1121,1204,1973,1996]" captionStart="FIGURE 2" captionStartId="2.[151,244,1546,1568]" captionTargetBox="[190,1402,227,1489]" captionTargetId="figure@2.[152,1435,191,1525]" captionTargetPageId="2" captionText="FIGURE 2. Canthidium quercetorum new species. A, the dorsal habitus; B, ventral habitus; C, lateral habitus; D, male protibia; E, female protibia;F, pygidium;and G, aedeagus." httpUri="https://zenodo.org/record/1169537/files/figure.png" pageId="1" pageNumber="274">Fig. 2F</figureCitation>
|
||
). Aedeagus as in
|
||
<figureCitation id="13189BFC762EFFFCFABAF87DFF40B999" captionStart="FIGURE 2" captionStartId="2.[151,244,1546,1568]" captionTargetBox="[190,1402,227,1489]" captionTargetId="figure@2.[152,1435,191,1525]" captionTargetPageId="2" captionText="FIGURE 2. Canthidium quercetorum new species. A, the dorsal habitus; B, ventral habitus; C, lateral habitus; D, male protibia; E, female protibia;F, pygidium;and G, aedeagus." httpUri="https://zenodo.org/record/1169537/files/figure.png" pageId="1" pageNumber="274">Fig. 2G</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF5CD7F1762DFFFFFF5CF9C2FDA1B84B" httpUri="https://zenodo.org/record/1169537/files/figure.png" pageId="2" pageNumber="275" startId="2.[151,244,1546,1568]" targetBox="[190,1402,227,1489]" targetPageId="2">
|
||
<paragraph id="8B9C8779762DFFFFFF5CF9C2FDA1B84B" blockId="2.[151,1436,1546,1597]" pageId="2" pageNumber="275">
|
||
<emphasis id="B9575B6B762DFFFFFF5CF9C2FEF6B856" bold="true" box="[151,270,1546,1568]" pageId="2" pageNumber="275">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C23FCFA762DFFFFFEDFF9C3FDFFB856" authority="Kohlmann & Arriaga-Jiménez & Rös, 2018" authorityName="Kohlmann & Arriaga-Jiménez & Rös" authorityYear="2018" box="[276,519,1547,1568]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="275" phylum="Arthropoda" rank="species" species="quercetorum" status="sp. nov.">
|
||
<emphasis id="B9575B6B762DFFFFFEDFF9C3FDFFB856" box="[276,519,1547,1568]" italics="true" pageId="2" pageNumber="275">Canthidium quercetorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A264E610762DFFFFFDC5F9C2FD7DB856" box="[526,645,1546,1568]" pageId="2" pageNumber="275" rank="species">
|
||
<emphasis id="B9575B6B762DFFFFFDC5F9C2FD7DB856" bold="true" box="[526,645,1546,1568]" pageId="2" pageNumber="275">new species</emphasis>
|
||
</taxonomicNameLabel>
|
||
.
|
||
<emphasis id="B9575B6B762DFFFFFD59F9C2FD5BB856" bold="true" box="[658,675,1546,1568]" pageId="2" pageNumber="275">A</emphasis>
|
||
, the dorsal habitus;
|
||
<emphasis id="B9575B6B762DFFFFFCA1F9C2FC82B856" bold="true" box="[874,890,1546,1568]" pageId="2" pageNumber="275">B</emphasis>
|
||
, ventral habitus;
|
||
<emphasis id="B9575B6B762DFFFFFBEFF9C2FBCDB856" bold="true" box="[1060,1077,1546,1568]" pageId="2" pageNumber="275">C</emphasis>
|
||
, lateral habitus;
|
||
<emphasis id="B9575B6B762DFFFFFB12F9C2FB12B856" bold="true" box="[1241,1258,1546,1568]" pageId="2" pageNumber="275">D</emphasis>
|
||
, male protibia;
|
||
<emphasis id="B9575B6B762DFFFFFA4DF9C2FA6EB856" bold="true" box="[1414,1430,1546,1568]" pageId="2" pageNumber="275">E</emphasis>
|
||
, female protibia;
|
||
<emphasis id="B9575B6B762DFFFFFEFEF9EFFEBAB84B" bold="true" box="[309,322,1575,1597]" pageId="2" pageNumber="275">F</emphasis>
|
||
, pygidium; and
|
||
<emphasis id="B9575B6B762DFFFFFE14F9EFFE17B84B" bold="true" box="[479,495,1575,1597]" pageId="2" pageNumber="275">G</emphasis>
|
||
, aedeagus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9C8779762DFFFFFF0CF9A0FD34B8B2" blockId="2.[151,1436,1640,2010]" pageId="2" pageNumber="275">
|
||
<emphasis id="B9575B6B762DFFFFFF0CF9A0FED6B809" bold="true" box="[199,302,1640,1663]" pageId="2" pageNumber="275">
|
||
<typeStatus id="549839DB762DFFFFFF0CF9A0FED1B809" box="[199,297,1640,1663]" pageId="2" pageNumber="275" type="allotype">Allotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Female. Length:
|
||
<quantity id="4CDB2A9C762DFFFFFE31F9A0FDABB809" box="[506,595,1640,1663]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="275" unit="mm" value="4.6">4.6 mm</quantity>
|
||
, humeral width:
|
||
<quantity id="4CDB2A9C762DFFFFFCDCF9A0FC89B809" box="[791,881,1640,1663]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="275" unit="mm" value="3.4">3.4 mm</quantity>
|
||
. Differing from the male in the following major characters: shorter pygidium, last abdominal segment broader, inner apex of protibia not forming a triangular projection, and tip of apical spur slightly bent inwards (
|
||
<figureCitation id="13189BFC762DFFFFFDA6F965FD38B8B2" box="[621,704,1709,1732]" captionStart="FIGURE 2" captionStartId="2.[151,244,1546,1568]" captionTargetBox="[190,1402,227,1489]" captionTargetId="figure@2.[152,1435,191,1525]" captionTargetPageId="2" captionText="FIGURE 2. Canthidium quercetorum new species. A, the dorsal habitus; B, ventral habitus; C, lateral habitus; D, male protibia; E, female protibia;F, pygidium;and G, aedeagus." httpUri="https://zenodo.org/record/1169537/files/figure.png" pageId="2" pageNumber="275">Fig. 2E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9C8779762DFFFFFF0CF918FAD4B97C" blockId="2.[151,1436,1640,2010]" pageId="2" pageNumber="275">
|
||
<emphasis id="B9575B6B762DFFFFFF0CF918FEC1B891" bold="true" box="[199,313,1744,1767]" pageId="2" pageNumber="275">Variation.</emphasis>
|
||
Length:
|
||
<quantity id="4CDB2A9C762DFFFFFE6EF918FDD6B891" box="[421,558,1744,1767]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="5.4" metricValueMin="3.9" pageId="2" pageNumber="275" unit="mm" value="4.65" valueMax="5.4" valueMin="3.9">3.9–5.4 mm</quantity>
|
||
, humeral width:
|
||
<quantity id="4CDB2A9C762DFFFFFD38F918FC85B891" box="[755,893,1744,1767]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.7" metricValueMin="2.4" pageId="2" pageNumber="275" unit="mm" value="3.05" valueMax="3.7" valueMin="2.4">2.4–3.7 mm</quantity>
|
||
. Dorsal surface reflexions varies from green to coppery. Antennal lamella varies from black to dark brown. Elytral intervals vary from flat to slightly convex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C339D4F2762DFFF9FF0CF8DDFC9EBF91" lastPageId="4" lastPageNumber="277" pageId="2" pageNumber="275" type="materials_examined">
|
||
<paragraph id="8B9C8779762DFFFEFF0CF8DDFCD5BF4F" blockId="2.[151,1436,1640,2010]" lastBlockId="3.[151,1437,152,313]" lastPageId="3" lastPageNumber="276" pageId="2" pageNumber="275">
|
||
<materialsCitation id="3B4B8D24762DFFFFFF0CF8DDFC85B95A" ID-GBIF-Occurrence="1806521911" box="[199,893,1813,1836]" collectorName="Material" country="Mexico" location="Mexico" pageId="2" pageNumber="275" specimenCount="110" stateProvince="Mexico" typeStatus="holotype">
|
||
<emphasis id="B9575B6B762DFFFFFF0CF8DDFE67B95A" bold="true" box="[199,415,1813,1836]" pageId="2" pageNumber="275">
|
||
<collectorName id="26D6E2AF762DFFFFFF0CF8DDFED2B95A" box="[199,298,1813,1836]" pageId="2" pageNumber="275">Material</collectorName>
|
||
examined
|
||
</emphasis>
|
||
(
|
||
<specimenCount id="9D254CF0762DFFFFFE66F8DDFDB7B95A" box="[429,591,1813,1836]" pageId="2" pageNumber="275" type="generic">110 specimens</specimenCount>
|
||
).
|
||
<emphasis id="B9575B6B762DFFFFFDAAF8DDFD28B95A" bold="true" box="[609,720,1813,1836]" pageId="2" pageNumber="275">
|
||
<typeStatus id="549839DB762DFFFFFDAAF8DDFD33B95A" box="[609,715,1813,1836]" pageId="2" pageNumber="275" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male:
|
||
<emphasis id="B9575B6B762DFFFFFCD0F8DDFCD0B95A" bold="true" box="[795,808,1813,1836]" pageId="2" pageNumber="275">“</emphasis>
|
||
<collectingRegion id="49E7499B762DFFFFFCE3F8DDFC85B95A" box="[808,893,1813,1836]" country="Mexico" name="Mexico" pageId="2" pageNumber="275">
|
||
<collectingCountry id="F334C7E9762DFFFFFCE3F8DDFC85B95A" box="[808,893,1813,1836]" name="Mexico" pageId="2" pageNumber="275">México</collectingCountry>
|
||
</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B4B8D24762DFFFFFC4CF8DDFEA3B904" ID-GBIF-Occurrence="1806521914" collectingDate="2016-09-20" collectorName="Reserva Comunitaria San Pablo Etla. & Coprotrampa. & N. Bosque de Encino & Female." country="Mexico" elevation="2155" location="Oaxaca" pageId="2" pageNumber="275" specimenCount="1" stateProvince="Oaxaca" typeStatus="allotype">
|
||
<collectingRegion id="49E7499B762DFFFFFC4CF8DDFC23B95A" box="[903,987,1813,1836]" country="Mexico" name="Oaxaca" pageId="2" pageNumber="275">Oaxaca</collectingRegion>
|
||
:
|
||
<collectorName id="26D6E2AF762DFFFFFC2EF8DDFA8AB95A" box="[997,1394,1813,1836]" pageId="2" pageNumber="275">Reserva Comunitaria San Pablo Etla.</collectorName>
|
||
<collectingDate id="EFD95851762DFFFFFAB3F8DDFF0CB939" pageId="2" pageNumber="275" value="2016-09-20">
|
||
<date id="FF9DA1B9762DFFFFFAB3F8DDFF0CB939" pageId="2" pageNumber="275" value="2016-09-20">20- IX-2016</date>
|
||
</collectingDate>
|
||
.
|
||
<collectorName id="26D6E2AF762DFFFFFECBF8F0FE6CB939" box="[256,404,1848,1871]" pageId="2" pageNumber="275">Coprotrampa.</collectorName>
|
||
x- 96.732348’ W, y- 17.170968’
|
||
<collectorName id="26D6E2AF762DFFFFFCC9F8F0FC14B939" box="[770,1004,1848,1871]" pageId="2" pageNumber="275">N. Bosque de Encino</collectorName>
|
||
,
|
||
<elevation id="000E604A762DFFFFFC33F8F0FBB6B939" box="[1016,1102,1848,1871]" metricMagnitude="3" metricUnit="m" metricValue="2.155" pageId="2" pageNumber="275" unit="m" value="2155.0">
|
||
<quantity id="4CDB2A9C762DFFFFFC33F8F0FBB6B939" box="[1016,1102,1848,1871]" metricMagnitude="3" metricUnit="m" metricValue="2.155" pageId="2" pageNumber="275" unit="m" value="2155.0">2155 m</quantity>
|
||
</elevation>
|
||
. Arriaga, J.A., Col.”
|
||
<emphasis id="B9575B6B762DFFFFFAFEF8F0FA64B939" bold="true" box="[1333,1436,1848,1871]" pageId="2" pageNumber="275">
|
||
<typeStatus id="549839DB762DFFFFFAFEF8F0FA6FB939" box="[1333,1431,1848,1871]" pageId="2" pageNumber="275" type="allotype">Allotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectorName id="26D6E2AF762DFFFFFF5CF893FF15B904" box="[151,237,1883,1906]" pageId="2" pageNumber="275">Female.</collectorName>
|
||
<emphasis id="B9575B6B762DFFFFFF32F893FEFEB904" bold="true" box="[249,262,1883,1906]" pageId="2" pageNumber="275">“</emphasis>
|
||
<collectingRegion id="49E7499B762DFFFFFECDF893FEA3B904" box="[262,347,1883,1906]" country="Mexico" name="Mexico" pageId="2" pageNumber="275">
|
||
<collectingCountry id="F334C7E9762DFFFFFECDF893FEA3B904" box="[262,347,1883,1906]" name="Mexico" pageId="2" pageNumber="275">México</collectingCountry>
|
||
</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B4B8D24762DFFFFFEA0F893FBD8B9E2" ID-GBIF-Occurrence="1806521910" collectingDate="2016-09-20" collectorName="Reserva Comunitaria San Pablo Etla. & Coprotrampa. & N. Bosque de Encino & Arriaga, J. A. & Col." country="Mexico" elevation="2155" location="Oaxaca" pageId="2" pageNumber="275" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
|
||
<collectingRegion id="49E7499B762DFFFFFEA0F893FE47B904" box="[363,447,1883,1906]" country="Mexico" name="Oaxaca" pageId="2" pageNumber="275">Oaxaca</collectingRegion>
|
||
:
|
||
<collectorName id="26D6E2AF762DFFFFFE04F893FC8CB904" box="[463,884,1883,1906]" pageId="2" pageNumber="275">Reserva Comunitaria San Pablo Etla.</collectorName>
|
||
<collectingDate id="EFD95851762DFFFFFC4BF893FBF9B904" box="[896,1025,1883,1906]" pageId="2" pageNumber="275" value="2016-09-20">
|
||
<date id="FF9DA1B9762DFFFFFC4BF893FBF9B904" box="[896,1025,1883,1906]" pageId="2" pageNumber="275" value="2016-09-20">20-IX-2016</date>
|
||
</collectingDate>
|
||
.
|
||
<collectorName id="26D6E2AF762DFFFFFBDAF893FB5DB904" box="[1041,1189,1883,1906]" pageId="2" pageNumber="275">Coprotrampa.</collectorName>
|
||
x- 96.732348’ W, y- 17.170968’
|
||
<collectorName id="26D6E2AF762DFFFFFED2F8B5FDF9B9E2" box="[281,513,1917,1940]" pageId="2" pageNumber="275">N. Bosque de Encino</collectorName>
|
||
,
|
||
<elevation id="000E604A762DFFFFFDC7F8B5FD99B9E2" box="[524,609,1917,1940]" metricMagnitude="3" metricUnit="m" metricValue="2.155" pageId="2" pageNumber="275" unit="m" value="2155.0">
|
||
<quantity id="4CDB2A9C762DFFFFFDC7F8B5FD99B9E2" box="[524,609,1917,1940]" metricMagnitude="3" metricUnit="m" metricValue="2.155" pageId="2" pageNumber="275" unit="m" value="2155.0">2155 m</quantity>
|
||
</elevation>
|
||
.
|
||
<collectorName id="26D6E2AF762DFFFFFDA6F8B5FD01B9E2" box="[621,761,1917,1940]" pageId="2" pageNumber="275">Arriaga, J.A.</collectorName>
|
||
,
|
||
<collectorName id="26D6E2AF762DFFFFFCCDF8B5FCCCB9E2" box="[774,820,1917,1940]" pageId="2" pageNumber="275">Col.</collectorName>
|
||
”
|
||
<typeStatus id="549839DB762DFFFFFC8DF8B5FC4AB9E2" box="[838,946,1917,1940]" pageId="2" pageNumber="275" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<emphasis id="B9575B6B762DFFFFFC76F8B5FC32B9E2" bold="true" box="[957,970,1917,1940]" pageId="2" pageNumber="275">“</emphasis>
|
||
<collectingRegion id="49E7499B762DFFFFFC00F8B5FBD8B9E2" box="[971,1056,1917,1940]" country="Mexico" name="Mexico" pageId="2" pageNumber="275">
|
||
<collectingCountry id="F334C7E9762DFFFFFC00F8B5FBD8B9E2" box="[971,1056,1917,1940]" name="Mexico" pageId="2" pageNumber="275">México</collectingCountry>
|
||
</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B4B8D24762DFFFFFBE1F8B5FE07B9AC" ID-GBIF-Occurrence="1806521912" collectingDate="2016-09-20" collectionCode="IEXA" collectorName="Reserva Comunitaria San Pablo Etla. & Coprotrampa. & N. Bosque de Encino & Arriaga, J. A. & Col." country="Mexico" elevation="2155" location="Oaxaca" pageId="2" pageNumber="275" specimenCount="102" stateProvince="Oaxaca" typeStatus="paratype">
|
||
<collectingRegion id="49E7499B762DFFFFFBE1F8B5FB86B9E2" box="[1066,1150,1917,1940]" country="Mexico" name="Oaxaca" pageId="2" pageNumber="275">Oaxaca</collectingRegion>
|
||
:
|
||
<collectorName id="26D6E2AF762DFFFFFB42F8B5FEF7B9C1" pageId="2" pageNumber="275">Reserva Comunitaria San Pablo Etla.</collectorName>
|
||
<collectingDate id="EFD95851762DFFFFFEDCF868FE60B9C1" box="[279,408,1952,1975]" pageId="2" pageNumber="275" value="2016-09-20">
|
||
<date id="FF9DA1B9762DFFFFFEDCF868FE60B9C1" box="[279,408,1952,1975]" pageId="2" pageNumber="275" value="2016-09-20">20-IX-2016</date>
|
||
</collectingDate>
|
||
.
|
||
<collectorName id="26D6E2AF762DFFFFFE6EF868FDC1B9C1" box="[421,569,1952,1975]" pageId="2" pageNumber="275">Coprotrampa.</collectorName>
|
||
x- 96.732348’ W, y- 17.170968’
|
||
<collectorName id="26D6E2AF762DFFFFFC67F868FB61B9C1" box="[940,1177,1952,1975]" pageId="2" pageNumber="275">N. Bosque de Encino</collectorName>
|
||
,
|
||
<elevation id="000E604A762DFFFFFB6DF868FB04B9C1" box="[1190,1276,1952,1975]" metricMagnitude="3" metricUnit="m" metricValue="2.155" pageId="2" pageNumber="275" unit="m" value="2155.0">
|
||
<quantity id="4CDB2A9C762DFFFFFB6DF868FB04B9C1" box="[1190,1276,1952,1975]" metricMagnitude="3" metricUnit="m" metricValue="2.155" pageId="2" pageNumber="275" unit="m" value="2155.0">2155 m</quantity>
|
||
</elevation>
|
||
.
|
||
<collectorName id="26D6E2AF762DFFFFFAC2F868FA6FB9C1" box="[1289,1431,1952,1975]" pageId="2" pageNumber="275">Arriaga, J.A.</collectorName>
|
||
,
|
||
<collectorName id="26D6E2AF762DFFFFFF5CF80BFF3EB9AC" box="[151,198,1987,2010]" pageId="2" pageNumber="275">Col.</collectorName>
|
||
” (
|
||
<specimenCount id="9D254CF0762DFFFFFF15F80BFE78B9AC" box="[222,384,1987,2010]" pageId="2" pageNumber="275" type="generic">102 specimens</specimenCount>
|
||
,
|
||
<collectionCode id="ED321FBC762DFFFFFE40F80BFE33B9AC" box="[395,459,1987,2010]" pageId="2" pageNumber="275">IEXA</collectionCode>
|
||
(50)
|
||
</materialsCitation>
|
||
, MNHN (12), CMNC (20),
|
||
<materialsCitation id="3B4B8D24762DFFFFFCE7F80BFBDBB9AC" ID-GBIF-Occurrence="1806521913" box="[812,1059,1987,2010]" collectionCode="CEMT" country="Mexico" location="Mexico" pageId="2" pageNumber="275" specimenCount="1" stateProvince="Mexico" typeStatus="paratype">
|
||
<collectionCode id="ED321FBC762DFFFFFCE7F80BFC8EB9AC" box="[812,886,1987,2010]" pageId="2" pageNumber="275">CEMT</collectionCode>
|
||
(20));
|
||
<emphasis id="B9575B6B762DFFFFFC0AF80BFC36B9AC" bold="true" box="[961,974,1987,2010]" pageId="2" pageNumber="275">“</emphasis>
|
||
<collectingRegion id="49E7499B762DFFFFFC05F80BFBDBB9AC" box="[974,1059,1987,2010]" country="Mexico" name="Mexico" pageId="2" pageNumber="275">
|
||
<collectingCountry id="F334C7E9762DFFFFFC05F80BFBDBB9AC" box="[974,1059,1987,2010]" name="Mexico" pageId="2" pageNumber="275">México</collectingCountry>
|
||
</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B4B8D24762DFFFEFBE6F80BFDC3BEA7" ID-GBIF-Occurrence="1806521907" collectingDate="2016-09-20" collectionCode="IEXA" collectorName="Reserva Comunitaria San Pablo Etla. & Coprotrampa. & N. Bosque de Encino & Arriaga, J. A. & Col." country="Mexico" elevation="2151" lastPageId="3" lastPageNumber="276" location="Oaxaca" pageId="2" pageNumber="275" specimenCount="3" stateProvince="Oaxaca" typeStatus="paratype">
|
||
<collectingRegion id="49E7499B762DFFFFFBE6F80BFB79B9AC" box="[1069,1153,1987,2010]" country="Mexico" name="Oaxaca" pageId="2" pageNumber="275">Oaxaca</collectingRegion>
|
||
:
|
||
<collectorName id="26D6E2AF762DFFFEFB40F80BFEF7BED9" lastPageId="3" lastPageNumber="276" pageId="2" pageNumber="275">Reserva Comunitaria San Pablo Etla.</collectorName>
|
||
<collectingDate id="EFD95851762CFFFEFEDCFF50FE60BED9" box="[279,408,152,175]" pageId="3" pageNumber="276" value="2016-09-20">
|
||
<date id="FF9DA1B9762CFFFEFEDCFF50FE60BED9" box="[279,408,152,175]" pageId="3" pageNumber="276" value="2016-09-20">20-IX-2016</date>
|
||
</collectingDate>
|
||
.
|
||
<collectorName id="26D6E2AF762CFFFEFE6EFF50FDC1BED9" box="[421,569,152,175]" pageId="3" pageNumber="276">Coprotrampa.</collectorName>
|
||
x- 96.732386’ W, y- 17.170454’
|
||
<collectorName id="26D6E2AF762CFFFEFC67FF50FB61BED9" box="[940,1177,152,175]" pageId="3" pageNumber="276">N. Bosque de Encino</collectorName>
|
||
,
|
||
<elevation id="000E604A762CFFFEFB6DFF50FB04BED9" box="[1190,1276,152,175]" metricMagnitude="3" metricUnit="m" metricValue="2.151" pageId="3" pageNumber="276" unit="m" value="2151.0">
|
||
<quantity id="4CDB2A9C762CFFFEFB6DFF50FB04BED9" box="[1190,1276,152,175]" metricMagnitude="3" metricUnit="m" metricValue="2.151" pageId="3" pageNumber="276" unit="m" value="2151.0">2151 m</quantity>
|
||
</elevation>
|
||
.
|
||
<collectorName id="26D6E2AF762CFFFEFAC2FF50FA6FBED9" box="[1289,1431,152,175]" pageId="3" pageNumber="276">Arriaga, J.A.</collectorName>
|
||
,
|
||
<collectorName id="26D6E2AF762CFFFEFF5CFF72FF3EBEA7" box="[151,198,186,209]" pageId="3" pageNumber="276">Col.</collectorName>
|
||
” (
|
||
<specimenCount id="9D254CF0762CFFFEFF2FFF72FE97BEA7" box="[228,367,186,209]" pageId="3" pageNumber="276" type="generic">3 specimens</specimenCount>
|
||
,
|
||
<collectionCode id="ED321FBC762CFFFEFEB5FF72FE3ABEA7" box="[382,450,186,209]" pageId="3" pageNumber="276">IEXA</collectionCode>
|
||
);
|
||
<emphasis id="B9575B6B762CFFFEFE12FF72FE1EBEA7" bold="true" box="[473,486,186,209]" pageId="3" pageNumber="276">“</emphasis>
|
||
<collectingRegion id="49E7499B762CFFFEFE2DFF72FDC3BEA7" box="[486,571,186,209]" country="Mexico" name="Mexico" pageId="3" pageNumber="276">
|
||
<collectingCountry id="F334C7E9762CFFFEFE2DFF72FDC3BEA7" box="[486,571,186,209]" name="Mexico" pageId="3" pageNumber="276">México</collectingCountry>
|
||
</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B4B8D24762CFFFEFD81FF72FAC3BE82" ID-GBIF-Occurrence="1806521906" collectingDate="2016-09-20" collectionCode="IEXA" collectorName="Reserva Comunitaria San Pablo Etla. & Coprotrampa. & N. Bosque de Encino & Arriaga, J. A. & Col." country="Mexico" elevation="2133" location="Oaxaca" pageId="3" pageNumber="276" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
|
||
<collectingRegion id="49E7499B762CFFFEFD81FF72FD66BEA7" box="[586,670,186,209]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="276">Oaxaca</collectingRegion>
|
||
:
|
||
<collectorName id="26D6E2AF762CFFFEFD66FF72FBB4BEA7" box="[685,1100,186,209]" pageId="3" pageNumber="276">Reserva Comunitaria San Pablo Etla.</collectorName>
|
||
<collectingDate id="EFD95851762CFFFEFB9CFF72FB20BEA7" box="[1111,1240,186,209]" pageId="3" pageNumber="276" value="2016-09-20">
|
||
<date id="FF9DA1B9762CFFFEFB9CFF72FB20BEA7" box="[1111,1240,186,209]" pageId="3" pageNumber="276" value="2016-09-20">20-IX-2016</date>
|
||
</collectingDate>
|
||
.
|
||
<collectorName id="26D6E2AF762CFFFEFB2CFF72FA83BEA7" box="[1255,1403,186,209]" pageId="3" pageNumber="276">Coprotrampa.</collectorName>
|
||
x- 96.733014’ W, y- 17.170797’
|
||
<collectorName id="26D6E2AF762CFFFEFE1CFF15FD42BE82" box="[471,698,221,244]" pageId="3" pageNumber="276">N. Bosque de Encino</collectorName>
|
||
,
|
||
<elevation id="000E604A762CFFFEFD0FFF15FCEFBE82" box="[708,791,221,244]" metricMagnitude="3" metricUnit="m" metricValue="2.133" pageId="3" pageNumber="276" unit="m" value="2133.0">
|
||
<quantity id="4CDB2A9C762CFFFEFD0FFF15FCEFBE82" box="[708,791,221,244]" metricMagnitude="3" metricUnit="m" metricValue="2.133" pageId="3" pageNumber="276" unit="m" value="2133.0">2133 m</quantity>
|
||
</elevation>
|
||
.
|
||
<collectorName id="26D6E2AF762CFFFEFCEAFF15FC54BE82" box="[801,940,221,244]" pageId="3" pageNumber="276">Arriaga, J.A.</collectorName>
|
||
,
|
||
<collectorName id="26D6E2AF762CFFFEFC7DFF15FC1DBE82" box="[950,997,221,244]" pageId="3" pageNumber="276">Col.</collectorName>
|
||
” (
|
||
<specimenCount id="9D254CF0762CFFFEFBCBFF15FB81BE82" box="[1024,1145,221,244]" pageId="3" pageNumber="276" type="generic">1 specimen</specimenCount>
|
||
,
|
||
<collectionCode id="ED321FBC762CFFFEFB48FF15FB3FBE82" box="[1155,1223,221,244]" pageId="3" pageNumber="276">IEXA</collectionCode>
|
||
);
|
||
<emphasis id="B9575B6B762CFFFEFB12FF15FB1EBE82" bold="true" box="[1241,1254,221,244]" pageId="3" pageNumber="276">“</emphasis>
|
||
<collectingRegion id="49E7499B762CFFFEFB2DFF15FAC3BE82" box="[1254,1339,221,244]" country="Mexico" name="Mexico" pageId="3" pageNumber="276">
|
||
<collectingCountry id="F334C7E9762CFFFEFB2DFF15FAC3BE82" box="[1254,1339,221,244]" name="Mexico" pageId="3" pageNumber="276">México</collectingCountry>
|
||
</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B4B8D24762CFFFEFA8FFF15FCD5BF4F" ID-GBIF-Occurrence="1806521905" collectingDate="2016-09-20" collectingDateMax="2016-09-23" collectingDateMin="2016-09-20" collectionCode="IEXA" collectorName="Reserva Comunitaria San Pablo Etla. & Coprotrampa. & N. Bosque de Encino & Arriaga, J. A. & Col." country="Mexico" elevation="2271" location="Oaxaca" pageId="3" pageNumber="276" specimenCount="2" stateProvince="Oaxaca" typeStatus="paratype">
|
||
<collectingRegion id="49E7499B762CFFFEFA8FFF15FA60BE82" box="[1348,1432,221,244]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="276">Oaxaca</collectingRegion>
|
||
:
|
||
<collectorName id="26D6E2AF762CFFFEFF5CFEC8FDDFBF61" box="[151,551,256,279]" pageId="3" pageNumber="276">Reserva Comunitaria San Pablo Etla.</collectorName>
|
||
<collectingDate id="EFD95851762CFFFEFDE5FEC8FD57BF61" box="[558,687,256,279]" pageId="3" pageNumber="276" value="2016-09-20">
|
||
<date id="FF9DA1B9762CFFFEFDE5FEC8FD57BF61" box="[558,687,256,279]" pageId="3" pageNumber="276" value="2016-09-20">20-IX-2016</date>
|
||
</collectingDate>
|
||
.
|
||
<collectorName id="26D6E2AF762CFFFEFD71FEC8FCB6BF61" box="[698,846,256,279]" pageId="3" pageNumber="276">Coprotrampa.</collectorName>
|
||
<collectingDate id="EFD95851762CFFFEFC9EFEC8FC42BF61" box="[853,954,256,279]" pageId="3" pageNumber="276" value="2016-09-23">
|
||
<date id="FF9DA1B9762CFFFEFC9EFEC8FC42BF61" box="[853,954,256,279]" pageId="3" pageNumber="276" value="2016-09-23">23-IX-16</date>
|
||
</collectingDate>
|
||
, x- 96.725987’ W, y- 17.174158’
|
||
<collectorName id="26D6E2AF762CFFFEFAE1FEC8FEFDBF4F" pageId="3" pageNumber="276">N. Bosque de Encino</collectorName>
|
||
,
|
||
<elevation id="000E604A762CFFFEFEC4FEEAFE9BBF4F" box="[271,355,290,313]" metricMagnitude="3" metricUnit="m" metricValue="2.271" pageId="3" pageNumber="276" unit="m" value="2271.0">
|
||
<quantity id="4CDB2A9C762CFFFEFEC4FEEAFE9BBF4F" box="[271,355,290,313]" metricMagnitude="3" metricUnit="m" metricValue="2.271" pageId="3" pageNumber="276" unit="m" value="2271.0">2271 m</quantity>
|
||
</elevation>
|
||
.
|
||
<collectorName id="26D6E2AF762CFFFEFEA6FEEAFE01BF4F" box="[365,505,290,313]" pageId="3" pageNumber="276">Arriaga, J.A.</collectorName>
|
||
,
|
||
<collectorName id="26D6E2AF762CFFFEFDCFFEEAFDCBBF4F" box="[516,563,290,313]" pageId="3" pageNumber="276">Col.</collectorName>
|
||
” (
|
||
<specimenCount id="9D254CF0762CFFFEFD87FEEAFD2BBF4F" box="[588,723,290,313]" pageId="3" pageNumber="276" type="generic">2 specimens</specimenCount>
|
||
,
|
||
<collectionCode id="ED321FBC762CFFFEFD16FEEAFCD9BF4F" box="[733,801,290,313]" pageId="3" pageNumber="276">IEXA</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="DF5CD7F1762CFFFEFF5CFA6AFEF6B8C9" httpUri="https://zenodo.org/record/1169539/files/figure.png" pageId="3" pageNumber="276" startId="3.[151,244,1442,1464]" targetBox="[433,1154,376,1413]" targetPageId="3">
|
||
<paragraph id="8B9C8779762CFFFEFF5CFA6AFEF6B8C9" blockId="3.[151,1437,1442,1727]" pageId="3" pageNumber="276">
|
||
<emphasis id="B9575B6B762CFFFEFF5CFA6AFEF7BBCE" bold="true" box="[151,271,1442,1464]" pageId="3" pageNumber="276">FIGURE 3.</emphasis>
|
||
Sampling site,
|
||
<emphasis id="B9575B6B762CFFFEFE66FA6AFE46BBCE" bold="true" box="[429,446,1442,1464]" pageId="3" pageNumber="276">A</emphasis>
|
||
, (grey point) of
|
||
<taxonomicName id="4C23FCFA762CFFFEFDA3FA6AFCA4BBC1" authority="Kohlmann & Arriaga-Jiménez & Rös, 2018" authorityName="Kohlmann & Arriaga-Jiménez & Rös" authorityYear="2018" box="[616,860,1442,1463]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="276" phylum="Arthropoda" rank="species" species="quercetorum" status="sp. nov.">
|
||
<emphasis id="B9575B6B762CFFFEFDA3FA6AFCA4BBC1" box="[616,860,1442,1463]" italics="true" pageId="3" pageNumber="276">Canthidium quercetorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A264E610762CFFFEFCAEFA6AFC26BBCE" box="[869,990,1442,1464]" pageId="3" pageNumber="276" rank="species">
|
||
<emphasis id="B9575B6B762CFFFEFCAEFA6AFC26BBCE" bold="true" box="[869,990,1442,1464]" pageId="3" pageNumber="276">new species</emphasis>
|
||
</taxonomicNameLabel>
|
||
in Oaxaca (grey area), Mexico (white area). Satellite images of
|
||
<taxonomicName id="4C23FCFA762CFFFEFE85FA77FDC7BBA2" authorityName="Kohlmann & Arriaga-Jiménez & Rös" authorityYear="2018" box="[334,575,1471,1492]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="276" phylum="Arthropoda" rank="species" species="quercetorum">
|
||
<emphasis id="B9575B6B762CFFFEFE85FA77FDC7BBA2" box="[334,575,1471,1492]" italics="true" pageId="3" pageNumber="276">Canthidium quercetorum</emphasis>
|
||
</taxonomicName>
|
||
sampling sites in a deciduous oak forest
|
||
<emphasis id="B9575B6B762CFFFEFC0CFA77FC2FBBA3" bold="true" box="[967,983,1471,1493]" pageId="3" pageNumber="276">B</emphasis>
|
||
, at the dry season (left) and
|
||
<emphasis id="B9575B6B762CFFFEFB2EFA77FB0EBBA3" bold="true" box="[1253,1270,1471,1493]" pageId="3" pageNumber="276">C</emphasis>
|
||
, at the end of the rainy season (right) (March 2017, November 2014). Bright colours on the left image indicate a forest with trees that have lost their leaves; whereas green vegetation on the same image shows pine-oak forest at higher altitudes, where
|
||
<taxonomicName id="4C23FCFA762CFFFEFBAAFA32FAAAB879" authorityName="Kohlmann & Arriaga-Jiménez & Rös" authorityYear="2018" box="[1121,1362,1530,1551]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="276" phylum="Arthropoda" rank="species" species="quercetorum">
|
||
<emphasis id="B9575B6B762CFFFEFBAAFA32FAAAB879" box="[1121,1362,1530,1551]" italics="true" pageId="3" pageNumber="276">Canthidium quercetorum</emphasis>
|
||
</taxonomicName>
|
||
has not been collected. On the right image, oak forest appears with nearly maximum leaf density at the end of the rainy season. Distribution of
|
||
<emphasis id="B9575B6B762CFFFEFF5CF9FCFF50B83C" bold="true" box="[151,168,1588,1610]" pageId="3" pageNumber="276">D</emphasis>
|
||
, dry oak forest (dark brown area), situated between approximately 1800 and 2400 m. Sampling site of
|
||
<taxonomicName id="4C23FCFA762CFFFEFB58F9FDFA7DB83C" authorityName="Kohlmann & Arriaga-Jiménez & Rös" authorityYear="2018" box="[1171,1413,1589,1610]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="276" phylum="Arthropoda" rank="species" species="quercetorum">
|
||
<emphasis id="B9575B6B762CFFFEFB58F9FDFA7DB83C" box="[1171,1413,1589,1610]" italics="true" pageId="3" pageNumber="276">Canthidium quercetorum</emphasis>
|
||
</taxonomicName>
|
||
is represented by the white rectangle. On the left lower side of the image the urban area of the Oaxaca Metropolitan Region can be seen. The dark green area on the right upper side of the image represents pine-oak and pine forest (highest elevation is around 3100 m). Satellite image was provided by GoogleEarth, and oak forest distribution by the Instituto Nacional de Estadística, Geografía e Informática.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9C8779762CFFF9FF0CF922FB56BE82" blockId="3.[151,1436,1770,2036]" lastBlockId="4.[151,1436,152,799]" lastPageId="4" lastPageNumber="277" pageId="3" pageNumber="276">
|
||
<emphasis id="B9575B6B762CFFFEFF0CF922FE0FB977" bold="true" box="[199,503,1770,1793]" pageId="3" pageNumber="276">Geographical distribution.</emphasis>
|
||
The species is so far only known from Etla in the Sierra Norte in
|
||
<collectingRegion id="49E7499B762CFFFEFB1CF922FADFB977" box="[1239,1319,1770,1793]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="276">Oaxaca</collectingRegion>
|
||
(
|
||
<figureCitation id="13189BFC762CFFFEFAFCF922FA76B977" box="[1335,1422,1770,1793]" captionStart="FIGURE 3" captionStartId="3.[151,244,1442,1464]" captionTargetBox="[433,1154,376,1413]" captionTargetId="figure@3.[419,1168,359,1431]" captionTargetPageId="3" captionText="FIGURE 3. Sampling site, A, (grey point) of Canthidium quercetorum new species in Oaxaca (grey area), Mexico (white area). Satelliteimages ofCanthidium quercetorumsampling sites inadeciduous oakforest B,at the dry season (left)andC,at the endof the rainy season (right) (March 2017, November 2014). Bright colours on the left image indicate a forest with trees that have lost their leaves;whereas green vegetationon the same image shows pine-oakforest at higher altitudes,where Canthidium quercetorumhas not been collected.On the right image,oak forest appears withnearly maximum leafdensity at the endof the rainy season.Distributionof D, dry oak forest (dark brown area), situated between approximately 1800 and 2400 m. Sampling site of Canthidium quercetorum is represented by the white rectangle.On the left lower sideof the imagethe urbanarea of the Oaxaca Metropolitan Region can be seen. The dark green area on the right upper side of the image represents pine-oak and pine forest (highest elevation is around 3100 m). Satellite image was provided by GoogleEarth, and oak forest distribution by the Instituto Nacional de Estadística, Geografía e Informática." httpUri="https://zenodo.org/record/1169539/files/figure.png" pageId="3" pageNumber="276">Fig. 3A</figureCitation>
|
||
), along the internal dry slope facing the
|
||
<collectingRegion id="49E7499B762CFFFEFDF8F8C5FD7BB952" box="[563,643,1805,1828]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="276">Oaxaca</collectingRegion>
|
||
Valley. The dry deciduous oak forest where
|
||
<taxonomicName id="4C23FCFA762CFFFEFBABF8C5FA93B952" authorityName="Kohlmann & Arriaga-Jiménez & Rös" authorityYear="2018" box="[1120,1387,1805,1828]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="276" phylum="Arthropoda" rank="species" species="quercetorum">
|
||
<emphasis id="B9575B6B762CFFFEFBABF8C5FA93B952" box="[1120,1387,1805,1828]" italics="true" pageId="3" pageNumber="276">Canthidium quercetorum</emphasis>
|
||
</taxonomicName>
|
||
was found is characterized by trees between
|
||
<quantity id="4CDB2A9C762CFFFEFD8AF8F8FD6AB931" box="[577,658,1840,1863]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="3" pageNumber="276" unit="m" value="7.5" valueMax="10.0" valueMin="5.0">5–10 m</quantity>
|
||
in height, and with around
|
||
<quantity id="4CDB2A9C762CFFFEFC7EF8F8FC0FB931" box="[949,1015,1840,1863]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="276" unit="cm" value="12.0">12 cm</quantity>
|
||
diameter at breast height (
|
||
<figureCitation id="13189BFC762CFFFEFAD9F8F8FA77B931" box="[1298,1423,1840,1863]" captionStart="FIGURE 3" captionStartId="3.[151,244,1442,1464]" captionTargetBox="[433,1154,376,1413]" captionTargetId="figure@3.[419,1168,359,1431]" captionTargetPageId="3" captionText="FIGURE 3. Sampling site, A, (grey point) of Canthidium quercetorum new species in Oaxaca (grey area), Mexico (white area). Satelliteimages ofCanthidium quercetorumsampling sites inadeciduous oakforest B,at the dry season (left)andC,at the endof the rainy season (right) (March 2017, November 2014). Bright colours on the left image indicate a forest with trees that have lost their leaves;whereas green vegetationon the same image shows pine-oakforest at higher altitudes,where Canthidium quercetorumhas not been collected.On the right image,oak forest appears withnearly maximum leafdensity at the endof the rainy season.Distributionof D, dry oak forest (dark brown area), situated between approximately 1800 and 2400 m. Sampling site of Canthidium quercetorum is represented by the white rectangle.On the left lower sideof the imagethe urbanarea of the Oaxaca Metropolitan Region can be seen. The dark green area on the right upper side of the image represents pine-oak and pine forest (highest elevation is around 3100 m). Satellite image was provided by GoogleEarth, and oak forest distribution by the Instituto Nacional de Estadística, Geografía e Informática." httpUri="https://zenodo.org/record/1169539/files/figure.png" pageId="3" pageNumber="276">Figs. 3B–C</figureCitation>
|
||
). Abundant oak species limited to this vegetation
|
||
<typeStatus id="549839DB762CFFFEFD0BF89AFD16B91F" box="[704,750,1874,1897]" pageId="3" pageNumber="276">type</typeStatus>
|
||
are
|
||
<taxonomicName id="4C23FCFA762CFFFEFCE3F89AFBE5B91F" authority="Liebm." authorityName="Liebm." box="[808,1053,1874,1897]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="laeta">
|
||
<emphasis id="B9575B6B762CFFFEFCE3F89AFC3BB91F" box="[808,963,1874,1897]" italics="true" pageId="3" pageNumber="276">Quercus laeta</emphasis>
|
||
Liebm.
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C23FCFA762CFFFEFB97F89AFEF6B9FA" authority="Humb. & Bonpl." authorityName="Humb. & Bonpl." class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="laurina">
|
||
<emphasis id="B9575B6B762CFFFEFB97F89AFB29B91F" box="[1116,1233,1874,1897]" italics="true" pageId="3" pageNumber="276">Q. laurina</emphasis>
|
||
Humb. & Bonpl. (Fagaceae)
|
||
</taxonomicName>
|
||
, which in this region is distributed principally between
|
||
<quantity id="4CDB2A9C762CFFFEFC94F8BDFC0DB9FA" box="[863,1013,1909,1932]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="3" pageNumber="276" unit="m" value="2100.0" valueMax="2400.0" valueMin="1800.0">1800–2400 m</quantity>
|
||
. Other species dominating this forest in the sampling site are
|
||
<taxonomicName id="4C23FCFA762CFFFEFEBCF850FD60B9D9" authority="Mart. & Gal." authorityName="Mart. & Gal." box="[375,664,1944,1967]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="glaucoides">
|
||
<emphasis id="B9575B6B762CFFFEFEBCF850FDF2B9D9" box="[375,522,1944,1967]" italics="true" pageId="3" pageNumber="276">Q. glaucoides</emphasis>
|
||
Mart. & Gal.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C23FCFA762CFFFEFD6EF850FC6EB9D9" authority="Oersted." authorityName="Oersted." box="[677,918,1944,1967]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="liebmannii">
|
||
<emphasis id="B9575B6B762CFFFEFD6EF850FCCFB9D9" box="[677,823,1944,1967]" italics="true" pageId="3" pageNumber="276">Q. liebmannii</emphasis>
|
||
Oersted.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C23FCFA762CFFFEFC6FF850FBB9B9D9" authority="Nee" authorityName="Nee" box="[932,1089,1944,1967]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="rugosa">
|
||
<emphasis id="B9575B6B762CFFFEFC6FF850FBF6B9D9" box="[932,1038,1944,1967]" italics="true" pageId="3" pageNumber="276">Q. rugosa</emphasis>
|
||
Née
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C23FCFA762CFFFEFBB3F850FAD2B9D9" authority="Nee" authorityName="Nee" box="[1144,1322,1944,1967]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="276" phylum="Tracheophyta" rank="species" species="castanea">
|
||
<emphasis id="B9575B6B762CFFFEFBB3F850FB0FB9D9" box="[1144,1271,1944,1967]" italics="true" pageId="3" pageNumber="276">Q. castanea</emphasis>
|
||
Née
|
||
</taxonomicName>
|
||
, which are also found at higher or lower altitudes (John Williams,
|
||
<collectingRegion id="49E7499B762CFFFEFCC8F872FCB4B9A7" box="[771,844,1978,2001]" country="France" name="Centre" pageId="3" pageNumber="276">Centro</collectingRegion>
|
||
Interdisciplinario de Investigación para el Desarrollo Integral Regional,
|
||
<collectingRegion id="49E7499B762CFFFEFEAFF815FE4EB982" box="[356,438,2013,2036]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="276">Oaxaca</collectingRegion>
|
||
, personal communication; Valencia-Ávalos & Nixon 2004). This dry deciduous oak forest shows a strong seasonality, where most trees lose their leaves for around four to five months between December and May (
|
||
<figureCitation id="13189BFC762BFFF9FF54FF72FEE6BEA7" box="[159,286,186,209]" captionStart="FIGURE 3" captionStartId="3.[151,244,1442,1464]" captionTargetBox="[433,1154,376,1413]" captionTargetId="figure@3.[419,1168,359,1431]" captionTargetPageId="3" captionText="FIGURE 3. Sampling site, A, (grey point) of Canthidium quercetorum new species in Oaxaca (grey area), Mexico (white area). Satelliteimages ofCanthidium quercetorumsampling sites inadeciduous oakforest B,at the dry season (left)andC,at the endof the rainy season (right) (March 2017, November 2014). Bright colours on the left image indicate a forest with trees that have lost their leaves;whereas green vegetationon the same image shows pine-oakforest at higher altitudes,where Canthidium quercetorumhas not been collected.On the right image,oak forest appears withnearly maximum leafdensity at the endof the rainy season.Distributionof D, dry oak forest (dark brown area), situated between approximately 1800 and 2400 m. Sampling site of Canthidium quercetorum is represented by the white rectangle.On the left lower sideof the imagethe urbanarea of the Oaxaca Metropolitan Region can be seen. The dark green area on the right upper side of the image represents pine-oak and pine forest (highest elevation is around 3100 m). Satellite image was provided by GoogleEarth, and oak forest distribution by the Instituto Nacional de Estadística, Geografía e Informática." httpUri="https://zenodo.org/record/1169539/files/figure.png" pageId="4" pageNumber="277">Figs. 3B–C</figureCitation>
|
||
). As can be seen (
|
||
<figureCitation id="13189BFC762BFFF9FE23FF72FDC7BEA7" box="[488,575,186,209]" captionStart="FIGURE 3" captionStartId="3.[151,244,1442,1464]" captionTargetBox="[433,1154,376,1413]" captionTargetId="figure@3.[419,1168,359,1431]" captionTargetPageId="3" captionText="FIGURE 3. Sampling site, A, (grey point) of Canthidium quercetorum new species in Oaxaca (grey area), Mexico (white area). Satelliteimages ofCanthidium quercetorumsampling sites inadeciduous oakforest B,at the dry season (left)andC,at the endof the rainy season (right) (March 2017, November 2014). Bright colours on the left image indicate a forest with trees that have lost their leaves;whereas green vegetationon the same image shows pine-oakforest at higher altitudes,where Canthidium quercetorumhas not been collected.On the right image,oak forest appears withnearly maximum leafdensity at the endof the rainy season.Distributionof D, dry oak forest (dark brown area), situated between approximately 1800 and 2400 m. Sampling site of Canthidium quercetorum is represented by the white rectangle.On the left lower sideof the imagethe urbanarea of the Oaxaca Metropolitan Region can be seen. The dark green area on the right upper side of the image represents pine-oak and pine forest (highest elevation is around 3100 m). Satellite image was provided by GoogleEarth, and oak forest distribution by the Instituto Nacional de Estadística, Geografía e Informática." httpUri="https://zenodo.org/record/1169539/files/figure.png" pageId="4" pageNumber="277">Fig. 3D</figureCitation>
|
||
), the inferred area of distribution of this new species is adjacent to the
|
||
<collectingRegion id="49E7499B762BFFF9FA80FF72FA63BEA7" box="[1355,1435,186,209]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="277">Oaxaca</collectingRegion>
|
||
Metropolitan Area. Thankfully, the area is under protection as the San Pablo Community Reserve.
|
||
</paragraph>
|
||
<paragraph id="8B9C8779762BFFF9FF0CFEC8FEABBF09" blockId="4.[151,1436,152,799]" pageId="4" pageNumber="277">
|
||
This species has been collected in the forest in association with
|
||
<taxonomicName id="4C23FCFA762BFFF9FC53FEC8FB51BF61" authority="(Say)" authorityName="Say" baseAuthorityName="Say" baseAuthorityYear="1831" box="[920,1193,256,279]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="humectus">
|
||
<emphasis id="B9575B6B762BFFF9FC53FEC8FB9BBF61" box="[920,1123,256,279]" italics="true" pageId="4" pageNumber="277">Canthon humectus</emphasis>
|
||
(Say)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C23FCFA762BFFF9FB72FEC8FA60BF61" authority="Harold" authorityName="Harold" authorityYear="1869" box="[1209,1432,256,279]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="klugi">
|
||
<emphasis id="B9575B6B762BFFF9FB72FEC8FAB9BF61" box="[1209,1345,256,279]" italics="true" pageId="4" pageNumber="277">Copris klugi</emphasis>
|
||
Harold
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C23FCFA762BFFF9FF5CFEEAFDDEBF4F" authority="Burmeister" authorityName="Burmeister" authorityYear="1848" box="[151,550,290,313]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="mexicanum">
|
||
<emphasis id="B9575B6B762BFFF9FF5CFEEAFE59BF4F" box="[151,417,290,313]" italics="true" pageId="4" pageNumber="277">Deltochilum mexicanum</emphasis>
|
||
Burmeister
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C23FCFA762BFFF9FDFCFEEAFC81BF4F" authority="(Say)" authorityName="Say" baseAuthorityName="Say" baseAuthorityYear="1835" box="[567,889,290,313]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="colonicus">
|
||
<emphasis id="B9575B6B762BFFF9FDFCFEEAFCCABF4F" box="[567,818,290,313]" italics="true" pageId="4" pageNumber="277">Dichotomius colonicus</emphasis>
|
||
(Say)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C23FCFA762BFFF9FC47FEEAFB07BF4F" authority="Harold" authorityName="Harold" authorityYear="1873" box="[908,1279,290,313]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="anthracinus">
|
||
<emphasis id="B9575B6B762BFFF9FC47FEEAFB5FBF4F" box="[908,1191,290,313]" italics="true" pageId="4" pageNumber="277">Onthophagus anthracinus</emphasis>
|
||
Harold
|
||
</taxonomicName>
|
||
(or near),
|
||
<taxonomicName id="4C23FCFA762BFFF9FA49FEEAFEA6BF2A" authority="Bates" authorityName="Bates" authorityYear="1887" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="aureofuscus">
|
||
<emphasis id="B9575B6B762BFFF9FA49FEEAFEEFBF2A" italics="true" pageId="4" pageNumber="277">O. aureofuscus</emphasis>
|
||
Bates
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C23FCFA762BFFF9FEA0FE8DFD5CBF2A" authority="Harold" authorityName="Harold" authorityYear="1869" box="[363,676,325,348]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="subSpecies" species="chevrolati" subSpecies="retusus">
|
||
<emphasis id="B9575B6B762BFFF9FEA0FE8DFDB6BF2A" box="[363,590,325,348]" italics="true" pageId="4" pageNumber="277">O. chevrolati retusus</emphasis>
|
||
Harold
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C23FCFA762BFFF9FD7BFE8DFC73BF2A" authority="Bates" authorityName="Bates" authorityYear="1887" box="[688,907,325,348]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="mexicanus">
|
||
<emphasis id="B9575B6B762BFFF9FD7BFE8DFCBCBF2A" box="[688,836,325,348]" italics="true" pageId="4" pageNumber="277">O. mexicanus</emphasis>
|
||
Bates
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C23FCFA762BFFF9FC53FE8DFB0FBF2A" authority="Zunino & Halffter" authorityName="Zunino & Halffter" authorityYear="1988" box="[920,1271,325,348]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="zapotecus">
|
||
<emphasis id="B9575B6B762BFFF9FC53FE8DFBDCBF2A" box="[920,1060,325,348]" italics="true" pageId="4" pageNumber="277">O. zapotecus</emphasis>
|
||
Zunino & Halffter
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C23FCFA762BFFF9FAF8FE8DFEABBF09" authority="Harold." authorityName="Harold." class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="damocles">
|
||
<emphasis id="B9575B6B762BFFF9FAF8FE8DFF04BF09" italics="true" pageId="4" pageNumber="277">Phanaeus damocles</emphasis>
|
||
Harold.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8B9C8779762BFFF9FF0CFE42FC9EBF91" blockId="4.[151,1436,152,799]" pageId="4" pageNumber="277">
|
||
<emphasis id="B9575B6B762BFFF9FF0CFE42FE31BFD7" bold="true" box="[199,457,394,417]" pageId="4" pageNumber="277">Chorological affinities.</emphasis>
|
||
The known collection locality of
|
||
<taxonomicName id="4C23FCFA762BFFF9FCFEFE42FC24BFD7" authorityName="Kohlmann & Arriaga-Jiménez & Rös" authorityYear="2018" box="[821,988,394,417]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="quercetorum">
|
||
<emphasis id="B9575B6B762BFFF9FCFEFE42FC24BFD7" box="[821,988,394,417]" italics="true" pageId="4" pageNumber="277">C. quercetorum</emphasis>
|
||
</taxonomicName>
|
||
is widely separated from that of a similar species,
|
||
<taxonomicName id="4C23FCFA762BFFF9FF3AFE65FDC6BFB2" authority="Kohlmann & Solis" authorityName="Kohlmann & Solis" box="[241,574,429,452]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="delgadoi">
|
||
<emphasis id="B9575B6B762BFFF9FF3AFE65FE95BFB2" box="[241,365,429,452]" italics="true" pageId="4" pageNumber="277">C. delgadoi</emphasis>
|
||
Kohlmann & Solís
|
||
</taxonomicName>
|
||
, which is distributed in cloud forests on the Pacific slope of the Sierra Madre del Sur from
|
||
<collectingRegion id="49E7499B762BFFF9FF36FE18FEACBF91" box="[253,340,464,487]" country="American Samoa" name="Western" pageId="4" pageNumber="277">Western</collectingRegion>
|
||
<collectingRegion id="49E7499B762BFFF9FE91FE18FE42BF91" box="[346,442,464,487]" country="Mexico" name="Guerrero" pageId="4" pageNumber="277">Guerrero</collectingRegion>
|
||
to
|
||
<collectingRegion id="49E7499B762BFFF9FE17FE18FDDFBF91" box="[476,551,464,487]" country="Mexico" name="Jalisco" pageId="4" pageNumber="277">Jalisco</collectingRegion>
|
||
,
|
||
<collectingRegion id="49E7499B762BFFF9FDF9FE18FD7EBF91" box="[562,646,464,487]" country="Mexico" name="Mexico" pageId="4" pageNumber="277">
|
||
<collectingCountry id="F334C7E9762BFFF9FDF9FE18FD7EBF91" box="[562,646,464,487]" name="Mexico" pageId="4" pageNumber="277">Mexico</collectingCountry>
|
||
</collectingRegion>
|
||
, from
|
||
<quantity id="4CDB2A9C762BFFF9FD00FE18FC9ABF91" box="[715,866,464,487]" metricMagnitude="3" metricUnit="m" metricValue="1.775" metricValueMax="2.2" metricValueMin="1.35" pageId="4" pageNumber="277" unit="m" value="1775.0" valueMax="2200.0" valueMin="1350.0">1350–2200 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C339D4F2762BFFF9FF0CFE3AFD77BC8A" pageId="4" pageNumber="277" type="description">
|
||
<paragraph id="8B9C8779762BFFF9FF0CFE3AFDBCBC39" blockId="4.[151,1436,152,799]" pageId="4" pageNumber="277">
|
||
<emphasis id="B9575B6B762BFFF9FF0CFE3AFE1DBC7F" bold="true" box="[199,485,498,521]" pageId="4" pageNumber="277">Taxonomic relationships.</emphasis>
|
||
<taxonomicName id="4C23FCFA762BFFF9FE26FE3AFD02BC7F" authorityName="Kohlmann & Arriaga-Jiménez & Rös" authorityYear="2018" box="[493,762,498,521]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="quercetorum">
|
||
<emphasis id="B9575B6B762BFFF9FE26FE3AFD02BC7F" box="[493,762,498,521]" italics="true" pageId="4" pageNumber="277">Canthidium quercetorum</emphasis>
|
||
</taxonomicName>
|
||
is postulated to be the sister species of
|
||
<taxonomicName id="4C23FCFA762BFFF9FB64FE3AFAD5BC7F" box="[1199,1325,498,521]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="delgadoi">
|
||
<emphasis id="B9575B6B762BFFF9FB64FE3AFAD5BC7F" box="[1199,1325,498,521]" italics="true" pageId="4" pageNumber="277">C. delgadoi</emphasis>
|
||
</taxonomicName>
|
||
. They are both sub-globose in body shape; they also have a swollen frons, narrow eyes, punctate pronotum, shagreen elytra, and shagreen and lightly punctate pygidium.
|
||
</paragraph>
|
||
<paragraph id="8B9C8779762BFFF9FF0CFD92FD77BC8A" blockId="4.[151,1436,152,799]" pageId="4" pageNumber="277">
|
||
The two species are very similiar, but
|
||
<taxonomicName id="4C23FCFA762BFFF9FD4EFD92FCC9BC07" authorityName="Kohlmann & Arriaga-Jiménez & Rös" authorityYear="2018" box="[645,817,602,625]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="quercetorum">
|
||
<emphasis id="B9575B6B762BFFF9FD4EFD92FCC9BC07" box="[645,817,602,625]" italics="true" pageId="4" pageNumber="277">C. quercetorum</emphasis>
|
||
</taxonomicName>
|
||
can be easily separated from
|
||
<taxonomicName id="4C23FCFA762BFFF9FB5BFD92FAEABC07" box="[1168,1298,602,625]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="delgadoi">
|
||
<emphasis id="B9575B6B762BFFF9FB5BFD92FAEABC07" box="[1168,1298,602,625]" italics="true" pageId="4" pageNumber="277">C. delgadoi</emphasis>
|
||
</taxonomicName>
|
||
using these characteristics: antennal lamellae black (versus pale yellow); pronotum is lightly shagreen; punctures of the elytral striae well defined; aedeagus with a fine, transparent keel running smoothly without forming a step along the dorsal, central midline of the parameres (keel absent in
|
||
<taxonomicName id="4C23FCFA762BFFF9FD93FD0AFD2EBCAF" box="[600,726,706,729]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="delgadoi">
|
||
<emphasis id="B9575B6B762BFFF9FD93FD0AFD2EBCAF" box="[600,726,706,729]" italics="true" pageId="4" pageNumber="277">C. delgadoi</emphasis>
|
||
</taxonomicName>
|
||
); larger than average size (
|
||
<quantity id="4CDB2A9C762BFFF9FBCAFD0AFB7DBCAF" box="[1025,1157,706,729]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="5.4" metricValueMin="3.9" pageId="4" pageNumber="277" unit="mm" value="4.65" valueMax="5.4" valueMin="3.9">3.9–5.4 mm</quantity>
|
||
versus
|
||
<quantity id="4CDB2A9C762BFFF9FB11FD0AFA99BCAF" box="[1242,1377,706,729]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.4" metricValueMin="3.5" pageId="4" pageNumber="277" unit="mm" value="3.95" valueMax="4.4" valueMin="3.5">3.5–4.4 mm</quantity>
|
||
); and inhabits a dry oak forest and not a cloud-forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C339D4F2762BFFF9FF0CFCC0FB1EBD69" box="[199,1254,776,799]" pageId="4" pageNumber="277" type="etymology">
|
||
<paragraph id="8B9C8779762BFFF9FF0CFCC0FB1EBD69" blockId="4.[151,1436,152,799]" box="[199,1254,776,799]" pageId="4" pageNumber="277">
|
||
<emphasis id="B9575B6B762BFFF9FF0CFCC0FEBFBD69" bold="true" box="[199,327,776,799]" pageId="4" pageNumber="277">Etymology.</emphasis>
|
||
The name
|
||
<taxonomicName id="4C23FCFA762BFFF9FE75FCC0FDBDBD69" authorityName="Kohlmann & Arriaga-Jiménez & Rös" authorityYear="2018" box="[446,581,776,799]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="quercetorum">
|
||
<emphasis id="B9575B6B762BFFF9FE75FCC0FDBDBD69" box="[446,581,776,799]" italics="true" pageId="4" pageNumber="277">quercetorum</emphasis>
|
||
</taxonomicName>
|
||
is a latin word in genitive plural, meaning “of the oak woods”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |