231 lines
28 KiB
XML
231 lines
28 KiB
XML
<document id="D8BC47C884BAD186069BFCB5B7DBB41C" ID-DOI="10.4202/app.01054.2023" ID-ISSN="1732-2421" ID-Zenodo-Dep="10981125" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1713285152458" checkinUser="felipe" docAuthor="Zamora, Samuel & Gutiérrez-Marco, Juan Carlos" docDate="2023" docId="038887FFFFD5FF96FFD8CFACFC439D7F" docLanguage="en" docName="ActaPalaeontolPol.68.2.185-192.pdf" docOrigin="Acta Palaeontologica Polonica 68 (2)" docSource="http://dx.doi.org/10.4202/app.01054.2023" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.2:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="2" docTitle="Dehmicystis ariasi Zamora & Gutiérrez-Marco 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="190" masterDocId="FFB1FF87FFD6FF93FFBAC908FFE79B28" masterDocTitle="Filling the Silurian gap of solutan echinoderms with the description of new species of Dehmicystis from Spain" masterLastPageNumber="192" masterPageNumber="185" pageNumber="188" updateTime="1713359753001" updateUser="felipe" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5D5DD4211C9F3BC6B26A969511BD1C78">Filling the Silurian gap of solutan echinoderms with the description of new species of Dehmicystis from Spain</mods:title>
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<mods:namePart id="DD8991633B131A0185DDD8883A26F0C2">Zamora, Samuel</mods:namePart>
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<mods:namePart id="376154BC8F971AC12BA6A9C5EDDD8625">Gutiérrez-Marco, Juan Carlos</mods:namePart>
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<mods:typeOfResource id="581DA2C2830C2E425424AE16FA63AD35">text</mods:typeOfResource>
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<mods:title id="58ED30FF35C524545688452D6F1022DA">Acta Palaeontologica Polonica</mods:title>
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</mods:titleInfo>
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<mods:part id="15FE93F76FEC3012C4FC01284ECB5929">
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<mods:date id="D9571ED2DD308DF9F6A8F5579E978F84">2023</mods:date>
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<mods:number id="DE0AAB084F05B0849F1ECFD33ED08604">2023-05-09</mods:number>
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<mods:number id="C1868EA9045DE060DE4693B163DBE181">68</mods:number>
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<mods:url id="D20DF59DFF636A8DEFADF28D43E445F9">http://dx.doi.org/10.4202/app.01054.2023</mods:url>
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<mods:identifier id="1A01DBEA425839AB5929257DCE957BDE" type="DOI">10.4202/app.01054.2023</mods:identifier>
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<mods:identifier id="B4754BD24C16E8EFF752A5E61C51CC35" type="ISSN">1732-2421</mods:identifier>
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<treatment id="038887FFFFD5FF96FFD8CFACFC439D7F" LSID="urn:lsid:plazi:treatment:038887FFFFD5FF96FFD8CFACFC439D7F" httpUri="http://treatment.plazi.org/id/038887FFFFD5FF96FFD8CFACFC439D7F" lastPageId="5" lastPageNumber="190" pageId="3" pageNumber="188">
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<subSubSection id="C33B6562FFD5FF90FFD8CFACFE5C9DE8" box="[98,443,1700,1729]" pageId="3" pageNumber="188" type="nomenclature">
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<paragraph id="8B9E36E9FFD5FF90FFD8CFACFE5C9DE8" blockId="3.[98,443,1700,1729]" box="[98,443,1700,1729]" pageId="3" pageNumber="188">
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<heading id="D0D68185FFD5FF90FFD8CFACFE5C9DE8" box="[98,443,1700,1729]" fontSize="12" level="2" pageId="3" pageNumber="188" reason="2">
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<taxonomicName id="4C214D6AFFD5FF90FFD8CFACFEB69DE9" authority="Zamora & Gutiérrez-Marco, 2023" authorityName="Zamora & Gutiérrez-Marco" authorityYear="2023" box="[98,337,1700,1729]" family="Dendrocystitidae" genus="Dehmicystis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrocystitida" pageId="3" pageNumber="188" phylum="Echinodermata" rank="species" species="ariasi" status="sp. nov.">
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<emphasis id="B955EAFBFFD5FF90FFD8CFACFEB69DE9" box="[98,337,1700,1729]" italics="true" pageId="3" pageNumber="188">Dehmicystis ariasi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2665780FFD5FF90FEE3CFACFE5C9DE8" box="[345,443,1700,1728]" pageId="3" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6562FFD5FF96FFD8CFD8FC439D7F" lastPageId="5" lastPageNumber="190" pageId="3" pageNumber="188" type="description">
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<paragraph id="8B9E36E9FFD5FF90FFD8CFD8FF279DCE" blockId="3.[98,192,1744,1766]" box="[98,192,1744,1766]" pageId="3" pageNumber="188">
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<figureCitation id="131A2A6CFFD5FF90FFD8CFD8FF449DCE" box="[98,163,1744,1766]" captionStart="Fig" captionStartId="4.[151,184,1796,1818]" captionTargetBox="[153,1532,160,1771]" captionTargetId="figure-10@4.[153,1532,160,1771]" captionTargetPageId="4" captionText="Fig. 3. Solutan echinoderm Dehmicystis ariasi sp. nov. from the Llagarinos Formation, Ludlow, Silurian, northern Central Iberian Zone, Spain. A. MGM- 208S.A1, complete specimen in dorsal aspect preserving proxistele and feeding appendage above the body. A2, counterpart of same specimen.A3, detail of feeding appendage consisting on biserial floor plates and cover plates. Gonopore on single plate, and a possible hydropore close to the feeding appendage attachment. A4, detail of the periproct on right position. Note that the periproct is visible from the inside and was located on dorsal right part of the theca.A5, detail of the appendage preserving the complete proxistele and proximal dististele. Photographs are from latex casts whitened with ammonium chloride sublimated." figureDoi="http://doi.org/10.5281/zenodo.10981131" httpUri="https://zenodo.org/record/10981131/files/figure.png" pageId="3" pageNumber="188">Figs. 3</figureCitation>
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,
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<figureCitation id="131A2A6CFFD5FF90FF14CFD9FF5B9DCE" box="[174,188,1745,1766]" captionStart="Fig" captionStartId="5.[98,131,1052,1074]" captionTargetBox="[105,1478,161,1027]" captionTargetId="figure-285@5.[101,1478,161,1028]" captionTargetPageId="5" captionText="Fig. 4. Camera lucida drawings of solutan echinoderm Dehmicystis ariasi sp. nov. from the Llagarinos Formation, Ludlow, Silurian, northern Central Iberian Zone, Spain; with interpretation of main anatomical parts." figureDoi="http://doi.org/10.5281/zenodo.10981133" httpUri="https://zenodo.org/record/10981133/files/figure.png" pageId="3" pageNumber="188">4</figureCitation>
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.
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</paragraph>
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<paragraph id="8B9E36E9FFD5FF90FFD8CFF0FF1E9C01" blockId="3.[98,770,1783,1833]" pageId="3" pageNumber="188">
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<emphasis id="B955EAFBFFD5FF90FFD8CFF0FF179C25" box="[98,240,1784,1805]" italics="true" pageId="3" pageNumber="188">
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Zoobank
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<collectionCode id="ED30AE2CFFD5FF90FF00CFF0FF179C25" box="[186,240,1784,1805]" pageId="3" pageNumber="188">LCID</collectionCode>
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</emphasis>
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:
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<uri id="FFB03AEBFFD5FF90FF47CFFFFF1E9C01" pageId="3" pageNumber="188">
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urn:lsid:zoobank.org:act:
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<uuid id="FF870C3CFFD5FF90FE59CFFFFF1E9C01" pageId="3" pageNumber="188">88604815-85BD-4E73-8FA4- 2381B337484C</uuid>
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</uri>
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</paragraph>
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<paragraph id="8B9E36E9FFD5FF90FFD8CE3EFF7C9CAD" blockId="3.[98,771,1846,1925]" pageId="3" pageNumber="188">
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<emphasis id="B955EAFBFFD5FF90FFD8CE3EFF2E9C63" box="[98,201,1846,1867]" italics="true" pageId="3" pageNumber="188">Etymology</emphasis>
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: Dedicated to Francisco Arias Ferrero (Ponferrada,
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<collectingCountry id="F3367679FFD5FF90FD7ECE3EFD1B9C64" box="[708,764,1846,1868]" name="Spain" pageId="3" pageNumber="188">Spain</collectingCountry>
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) for collecting the only specimen known and making it available for study.
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</paragraph>
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<paragraph id="8B9E36E9FFD5FF90FFD8CE99FA56985A" blockId="3.[98,716,1937,1959]" lastBlockId="3.[808,1481,804,883]" pageId="3" pageNumber="188">
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<materialsCitation id="3B493CB4FFD5FF90FFD8CE99FA4A985A" collectionCode="MGM" country="Spain" elevation="946" latitude="42.45625" location="Poleares high" longLatPrecision="1" longitude="-6.865861" municipality="Rubia municipality" pageId="3" pageNumber="188" specimenCode="MGM-208S-a, MGM-208S-b" specimenCount="1" stateProvince="Ourense Province" typeStatus="holotype">
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<emphasis id="B955EAFBFFD5FF90FFD8CE99FF5D9C8E" box="[98,186,1937,1958]" italics="true" pageId="3" pageNumber="188">
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||
<typeStatus id="549A884BFFD5FF90FFD8CE99FF5D9C8E" box="[98,186,1937,1958]" pageId="3" pageNumber="188" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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:
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<specimenCode id="DB879E92FFD5FF90FF7CCE99FEA89C8E" box="[198,335,1937,1958]" pageId="3" pageNumber="188">MGM-208S-a</specimenCode>
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(part) and
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<specimenCode id="DB879E92FFD5FF90FE02CE99FDA59C8F" box="[440,578,1937,1959]" pageId="3" pageNumber="188">MGM-208S-b</specimenCode>
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(counterpart).
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<emphasis id="B955EAFBFFD5FF90FC92CA2CFC459811" box="[808,930,804,825]" italics="true" pageId="3" pageNumber="188">Type locality</emphasis>
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||
:
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<locationDeviation id="9A2C2FCBFFD5FF90FC17CA2CFBFD9811" box="[941,1050,804,825]" pageId="3" pageNumber="188">
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<quantity id="4CD99B0CFFD5FF90FC17CA2CFC0E9811" box="[941,1001,804,825]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="188" unit="m" value="340.0">340 m</quantity>
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west
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</locationDeviation>
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of
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<location id="8EFE6032FFD5FF90FB8DCA2CFB5E9812" LSID="urn:lsid:plazi:treatment:038887FFFFD5FF96FFD8CFACFC439D7F:8EFE6032FFD5FF90FB8DCA2CFB5E9812" box="[1079,1209,804,826]" country="Spain" latitude="42.45625" longLatPrecision="1" longitude="-6.865861" municipality="Rubia municipality" name="Poleares high" pageId="3" pageNumber="188" stateProvince="Ourense Province">Poleares high</location>
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(
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<quantity id="4CD99B0CFFD5FF90FB7ECA2CFAE49811" box="[1220,1283,804,826]" metricMagnitude="2" metricUnit="m" metricValue="9.459999999999999" pageId="3" pageNumber="188" unit="m" value="946.0">
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||
<elevation id="000CD1DAFFD5FF90FB7ECA2CFAE49811" box="[1220,1283,804,826]" metricMagnitude="2" metricUnit="m" metricValue="9.459999999999999" pageId="3" pageNumber="188" unit="m" value="946.0">946 m</elevation>
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</quantity>
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), about
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<quantity id="4CD99B0CFFD5FF90FAF4CA2CFA6D9811" box="[1358,1418,804,826]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="3" pageNumber="188" unit="m" value="960.0">960 m</quantity>
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northeast of O Castelo village in the
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<collectingMunicipality id="6BFAAC93FFD5FF90FBEFCA48FAE9987E" box="[1109,1294,832,854]" pageId="3" pageNumber="188">Rubiá municipality</collectingMunicipality>
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||
,
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<collectingRegion id="49E5F80BFFD5FF90FAA2CA48FA21987E" box="[1304,1478,832,854]" pageId="3" pageNumber="188">Ourense Province</collectingRegion>
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||
,
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<collectingRegion id="49E5F80BFFD5FF90FC92CA55FC97985B" box="[808,880,861,883]" country="Spain" name="Galicia" pageId="3" pageNumber="188">Galicia</collectingRegion>
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, NW
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<collectingCountry id="F3367679FFD5FF90FC1DCA55FC06985B" box="[935,993,861,883]" name="Spain" pageId="3" pageNumber="188">Spain</collectingCountry>
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. GPS coordinates:
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<geoCoordinate id="EE15502EFFD5FF90FB2DCA54FAC6985A" box="[1175,1313,860,883]" degrees="42" direction="north" minutes="27" orientation="latitude" pageId="3" pageNumber="188" precision="1" seconds="22.5" value="42.45625">42º27′22.5″ N</geoCoordinate>
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,
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<geoCoordinate id="EE15502EFFD5FF90FA91CA54FA4A985A" box="[1323,1453,860,883]" degrees="6" direction="west" minutes="51" orientation="longitude" pageId="3" pageNumber="188" precision="1" seconds="57.1" value="-6.865861">6º51′57.1″ W</geoCoordinate>
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</materialsCitation>
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||
.
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</paragraph>
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<paragraph id="8B9E36E9FFD5FF90FC92CA77FAAB98E6" blockId="3.[808,1487,895,974]" pageId="3" pageNumber="188">
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<emphasis id="B955EAFBFFD5FF90FC92CA77FC4198BC" box="[808,934,895,916]" italics="true" pageId="3" pageNumber="188">Type horizon</emphasis>
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: Shale bed with a slight chloritoid content in the upper part of the Llagarinos Formation. Bed of a probably Gorstian (early Ludlow) age by lateral correlation with a nearby locality.
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</paragraph>
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<paragraph id="8B9E36E9FFD5FF90FC92CAD7FBB298D1" blockId="3.[808,1109,991,1017]" box="[808,1109,991,1017]" pageId="3" pageNumber="188">
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<emphasis id="B955EAFBFFD5FF90FC92CAD7FC6D98D1" box="[808,906,991,1017]" italics="true" pageId="3" pageNumber="188">Material</emphasis>
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.—
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<typeStatus id="549A884BFFD5FF90FC11CAD7FBF198D1" box="[939,1046,991,1017]" pageId="3" pageNumber="188" type="holotype">Holotype</typeStatus>
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only.
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</paragraph>
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<paragraph id="8B9E36E9FFD5FF90FC92CD03FC929F4D" blockId="3.[808,1481,1034,1125]" pageId="3" pageNumber="188">
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<emphasis id="B955EAFBFFD5FF90FC92CD03FC459F0C" box="[808,930,1035,1060]" italics="true" pageId="3" pageNumber="188">Diagnosis.</emphasis>
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—A species of
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<taxonomicName id="4C214D6AFFD5FF90FBF3CD02FB339F0C" authorityName="Caster" authorityYear="1968" box="[1097,1236,1034,1060]" family="Dendrocystitidae" genus="Dehmicystis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="188" phylum="Echinodermata" rank="genus">
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<emphasis id="B955EAFBFFD5FF90FBF3CD02FB339F0C" box="[1097,1236,1034,1060]" italics="true" pageId="3" pageNumber="188">Dehmicystis</emphasis>
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</taxonomicName>
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with a globular theca, proxistele equal in length to the theca. Undifferentiated mesistele.
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</paragraph>
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<paragraph id="8B9E36E9FFD5FF90FC92CD7FFC789C74" blockId="3.[808,1481,1142,1884]" pageId="3" pageNumber="188">
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<emphasis id="B955EAFBFFD5FF90FC92CD7FFC559FB8" box="[808,946,1143,1168]" italics="true" pageId="3" pageNumber="188">Description</emphasis>
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.—Thecal shape rounded, large (
|
||
<quantity id="4CD99B0CFFD5FF90FAF0CD7EFA449FB9" box="[1354,1443,1142,1169]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="188" unit="mm" value="45.0">45 mm</quantity>
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||
in length,
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||
<quantity id="4CD99B0CFFD5FF90FCC5CD9FFC289F9A" box="[895,975,1175,1202]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="188" unit="mm" value="30.0">30 mm</quantity>
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||
in width), nearly bilaterally symmetric with the left side more developed than the right. Theca widening posteriorly with an invagination for the insertion of the homoiostele. Narrow peripheral ring as a result of compression rather than originally biological. Plates very thin, superimposed integuments (taphonomic), polygonal and relatively large. Boundaries between plates are poorly preserved. Feeding appendage off the apex, on an integument with attachment to the theca on left side; relatively large with wider rectangular basal plates (flooring plates) and thinner, cover plates. Feeding appendage thinning distally, with food groove oriented to the substrate (see interpretation below). Small gonopore on a circular tiny plate, associated with some rugose plates interpreted as hydropore. Both separated one another. Periproct large, on the opposite side of the theca, close to the appendage insertion but on the right position. Proxistele very large, consisting of eleven tetramerous rings, two on each side. Posterior side of each ring with a depressed side for the articulation of adjacent rows distally. Distal stele only preserved proximally, composed of two dimeres. Theca slightly longer than the proxistele.
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</paragraph>
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<paragraph id="8B9E36E9FFD5FF96FC92CE65FC529D23" blockId="3.[808,1481,1901,1960]" lastBlockId="5.[808,1481,1133,1547]" lastPageId="5" lastPageNumber="190" pageId="3" pageNumber="188">
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||
<emphasis id="B955EAFBFFD5FF90FC92CE65FC6D9CAF" box="[808,906,1901,1927]" italics="true" pageId="3" pageNumber="188">Remarks</emphasis>
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||
.—Comparison with the type species
|
||
<taxonomicName id="4C214D6AFFD5FF90FA84CE65FBCE9C80" authority="(Dehm, 1934)" baseAuthorityName="Dehm" baseAuthorityYear="1934" family="Dendrocystitidae" genus="Dehmicystis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="188" phylum="Echinodermata" rank="species" species="globulus">
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||
<emphasis id="B955EAFBFFD5FF90FA84CE65FC6E9C8F" italics="true" pageId="3" pageNumber="188">Dehmicystis globulus</emphasis>
|
||
(
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||
<bibRefCitation id="EFB04B18FFD5FF90FC2CCE85FBC69C80" author="Dehm, R." box="[918,1057,1933,1960]" pageId="3" pageNumber="188" pagination="19 - 43" refId="ref5088" refString="Dehm, R. 1934. Untersuchungen an Cystoideen des rheinischen Unterdevons. Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Abteilung der Bayerischen Akademie der Wissenschaften zu Munchen fur 1934: 19 - 43." type="journal article" year="1934">Dehm, 1934</bibRefCitation>
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||
)
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</taxonomicName>
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from
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<collectingCountry id="F3367679FFD5FF90FBD0CE86FB339C80" box="[1130,1236,1934,1960]" name="Germany" pageId="3" pageNumber="188">Germany</collectingCountry>
|
||
(
|
||
<figureCitation id="131A2A6CFFD5FF90FB58CE85FAC49C80" box="[1250,1315,1933,1960]" captionStart="Fig" captionStartId="3.[96,129,695,717]" captionTargetBox="[99,1479,161,669]" captionTargetId="figure-611@3.[99,1479,161,669]" captionTargetPageId="3" captionText="Fig. 2. Solutan echinoderm Dehmicystis globulus (Dehm, 1934) from the the Lower Devonian Hunsrück Slate, Germany. A. Holotype (SMF-HS 432), almost complete specimen showing the theca and posterior appendage. Note anus in on right side. B. SMF-HS 969, slab containing three individuals. Note that individual 1 has the anus on “left” position.Repository number assigned to a replica of the newly prepared slab.Photographs courtesy of Ulrich Jansen." figureDoi="http://doi.org/10.5281/zenodo.10981129" httpUri="https://zenodo.org/record/10981129/files/figure.png" pageId="3" pageNumber="188">Fig. 2</figureCitation>
|
||
), poses several problems; specially because German specimens are flattened and tectonically distorted, and also because none of the known specimens is totally complete (
|
||
<figureCitation id="131A2A6CFFD3FF96FDFBCDA7FD629FE2" box="[577,645,1199,1226]" captionStart="Fig" captionStartId="3.[96,129,695,717]" captionTargetBox="[99,1479,161,669]" captionTargetId="figure-611@3.[99,1479,161,669]" captionTargetPageId="3" captionText="Fig. 2. Solutan echinoderm Dehmicystis globulus (Dehm, 1934) from the the Lower Devonian Hunsrück Slate, Germany. A. Holotype (SMF-HS 432), almost complete specimen showing the theca and posterior appendage. Note anus in on right side. B. SMF-HS 969, slab containing three individuals. Note that individual 1 has the anus on “left” position.Repository number assigned to a replica of the newly prepared slab.Photographs courtesy of Ulrich Jansen." figureDoi="http://doi.org/10.5281/zenodo.10981129" httpUri="https://zenodo.org/record/10981129/files/figure.png" pageId="5" pageNumber="190">Fig. 2</figureCitation>
|
||
).
|
||
<typeStatus id="549A884BFFD3FF96FD23CDA7FCE59FE1" box="[665,770,1199,1225]" pageId="5" pageNumber="190" type="holotype">Holotype</typeStatus>
|
||
and individual 1 from Rahman and Linz (2012) have the anterior part of the theca incomplete, individual 2 preserves only the homoiostele and individual 3 preserves only the homoiostele and posterior part of the theca. Rahman and Linz (2012) indicated that the dimensions of the
|
||
<typeStatus id="549A884BFFD3FF96FD25CC5DFCE59E47" box="[671,770,1365,1391]" pageId="5" pageNumber="190" type="holotype">holotype</typeStatus>
|
||
are 70 ×
|
||
<quantity id="4CD99B0CFFD3FF96FF7DCC7EFEFE9EB8" box="[199,281,1398,1424]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" pageId="5" pageNumber="190" unit="mm" value="67.0">67 mm</quantity>
|
||
, but based on new observations, this is unlikely (see
|
||
<tableCitation id="C6A30352FFD3FF96FF67CC9FFEC99E99" box="[221,302,1431,1457]" captionStart="Table 1" captionStartId="5.[98,151,1656,1678]" captionTargetId="graphics-410@5.[278,1300,1724,1948]" captionTargetPageId="5" captionText="Table 1. Comparative measures (in mm) from Dehmicystis ariasi sp. nov. and Dehmicystis globulus (Dehm, 1934). L, length, W, width; (i), incomplete measure." pageId="5" pageNumber="190">Table 1</tableCitation>
|
||
). The length of the
|
||
<typeStatus id="549A884BFFD3FF96FDA8CC9FFD929E99" box="[530,629,1431,1457]" pageId="5" pageNumber="190" type="holotype">holotype</typeStatus>
|
||
is unknown because incomplete preservation of the anterior part and its width is about
|
||
<quantity id="4CD99B0CFFD3FF96FEAACCD1FE819EDC" box="[272,358,1497,1524]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="190" unit="mm" value="40.0">40 mm</quantity>
|
||
. So dimensions of the
|
||
<typeStatus id="549A884BFFD3FF96FDC9CCD1FD319EDB" box="[627,726,1497,1523]" pageId="5" pageNumber="190" type="holotype">holotype</typeStatus>
|
||
are comparable to
|
||
<taxonomicName id="4C214D6AFFD3FF96FEB5CCF3FE919D3C" authorityName="Zamora & Gutiérrez-Marco" authorityYear="2023" box="[271,374,1531,1556]" family="Dendrocystitidae" genus="Dehmicystis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="190" phylum="Echinodermata" rank="species" species="ariasi">
|
||
<emphasis id="B955EAFBFFD3FF96FEB5CCF3FE919D3C" box="[271,374,1531,1556]" italics="true" pageId="5" pageNumber="190">D. ariasi</emphasis>
|
||
</taxonomicName>
|
||
. The other individuals seem relatively larger. Shape of the theca between both species seems different, being more elongate in the German taxon, but this difference can be also due to the strong tectonic distortion affecting the Hunsrück material. Another important difference is the relation between thecal and proxistele lengths. Spanish material seems to have a longer proxistele which is almost equal than the length of the theca. For contrast
|
||
<taxonomicName id="4C214D6AFFD3FF96FC92CC18FC4A9E01" baseAuthorityName="Dehm" baseAuthorityYear="1934" box="[808,941,1295,1321]" family="Dendrocystitidae" genus="Dehmicystis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="190" phylum="Echinodermata" rank="species" species="globulus">
|
||
<emphasis id="B955EAFBFFD3FF96FC92CC18FC4A9E01" box="[808,941,1295,1321]" italics="true" pageId="5" pageNumber="190">D. globulus</emphasis>
|
||
</taxonomicName>
|
||
has a shorter proxistele. Mesistele is well preserved in the individual 2 of Rahman and Linz (2012), but it is absent in the Spanish material, where transition from the proxistele to the dististele is more abrupt. Further comparison with the type species is not possible due to incomplete preservation of material from both
|
||
<collectingCountry id="F3367679FFD3FF96FB09CCB9FB149EE3" box="[1203,1267,1457,1483]" name="Spain" pageId="5" pageNumber="190">Spain</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3367679FFD3FF96FA90CCB9FA729EE3" box="[1322,1429,1457,1483]" name="Germany" pageId="5" pageNumber="190">Germany</collectingCountry>
|
||
, and different preservation styles being German specimens more compressed.
|
||
</paragraph>
|
||
<caption id="DF5E6661FFD2FF97FF2DCE0CFEB59C8E" ID-DOI="http://doi.org/10.5281/zenodo.10981131" ID-Zenodo-Dep="10981131" httpUri="https://zenodo.org/record/10981131/files/figure.png" pageId="4" pageNumber="189" startId="4.[151,184,1796,1818]" targetBox="[153,1532,160,1771]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8B9E36E9FFD2FF97FF2DCE0CFEB59C8E" blockId="4.[151,1535,1796,1958]" pageId="4" pageNumber="189">
|
||
Fig. 3. Solutan echinoderm
|
||
<taxonomicName id="4C214D6AFFD2FF97FE2BCE0CFDDD9C31" authority="Zamora & Gutiérrez-Marco, 2023" authorityName="Zamora & Gutiérrez-Marco" authorityYear="2023" box="[401,570,1796,1817]" family="Dendrocystitidae" genus="Dehmicystis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="189" phylum="Echinodermata" rank="species" species="ariasi" status="sp. nov.">
|
||
<emphasis id="B955EAFBFFD2FF97FE2BCE0CFDDD9C31" box="[401,570,1796,1817]" italics="true" pageId="4" pageNumber="189">Dehmicystis ariasi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2665780FFD2FF97FD85CE0CFD629C31" box="[575,645,1796,1817]" pageId="4" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from the Llagarinos Formation, Ludlow, Silurian, northern Central Iberian Zone, Spain.
|
||
<emphasis id="B955EAFBFFD2FF97FA19CE0CFA549C31" bold="true" box="[1443,1459,1796,1817]" pageId="4" pageNumber="189">A</emphasis>
|
||
. MGM-208S. A
|
||
<subScript id="17A534ACFFD2FF97FF5ACE22FF0F9C10" attach="left" box="[224,232,1834,1848]" fontSize="6" pageId="4" pageNumber="189">1</subScript>
|
||
, complete specimen in dorsal aspect preserving proxistele and feeding appendage above the body. A
|
||
<subScript id="17A534ACFFD2FF97FBC1CE22FB649C10" attach="left" box="[1147,1155,1834,1848]" fontSize="6" pageId="4" pageNumber="189">2</subScript>
|
||
, counterpart of same specimen. A
|
||
<subScript id="17A534ACFFD2FF97FA03CE22FA269C10" attach="right" box="[1465,1473,1834,1848]" fontSize="6" pageId="4" pageNumber="189">3</subScript>
|
||
, detail of feeding appendage consisting on biserial floor plates and cover plates. Gonopore on single plate, and a possible hydropore close to the feeding appendage attachment. A
|
||
<subScript id="17A534ACFFD2FF97FE81CE6AFEA49C58" attach="left" box="[315,323,1890,1904]" fontSize="6" pageId="4" pageNumber="189">4</subScript>
|
||
, detail of the periproct on right position. Note that the periproct is visible from the inside and was located on dorsal right part of the theca. A
|
||
<subScript id="17A534ACFFD2FF97FF5BCE76FF0E9CA4" attach="left" box="[225,233,1918,1932]" fontSize="6" pageId="4" pageNumber="189">5</subScript>
|
||
, detail of the appendage preserving the complete proxistele and proximal dististele. Photographs are from latex casts whitened with ammonium chloride sublimated.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5E6661FFD3FF96FFD8CD14FD519F65" ID-DOI="http://doi.org/10.5281/zenodo.10981133" ID-Zenodo-Dep="10981133" httpUri="https://zenodo.org/record/10981133/files/figure.png" pageId="5" pageNumber="190" startId="5.[98,131,1052,1074]" targetBox="[105,1478,161,1027]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8B9E36E9FFD3FF96FFD8CD14FD519F65" blockId="5.[98,1481,1052,1102]" pageId="5" pageNumber="190">
|
||
Fig. 4. Camera lucida drawings of solutan echinoderm
|
||
<taxonomicName id="4C214D6AFFD3FF96FDDBCD14FCEB9F19" authority="Zamora & Gutiérrez-Marco, 2023" authorityName="Zamora & Gutiérrez-Marco" authorityYear="2023" box="[609,780,1052,1073]" family="Dendrocystitidae" genus="Dehmicystis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="190" phylum="Echinodermata" rank="species" species="ariasi" status="sp. nov.">
|
||
<emphasis id="B955EAFBFFD3FF96FDDBCD14FCEB9F19" box="[609,780,1052,1073]" italics="true" pageId="5" pageNumber="190">Dehmicystis ariasi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2665780FFD3FF96FCA9CD14FCBB9F19" box="[787,860,1052,1073]" pageId="5" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from the Llagarinos Formation, Ludlow, Silurian, northern Central Iberian Zone, Spain; with interpretation of main anatomical parts.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9E36E9FFD3FF96FC92CF15FC439D7F" blockId="5.[808,1481,1565,1623]" pageId="5" pageNumber="190">
|
||
<emphasis id="B955EAFBFFD3FF96FC92CF15FB5A9D1E" box="[808,1213,1565,1591]" italics="true" pageId="5" pageNumber="190">Stratigraphic and geographic range</emphasis>
|
||
.—
|
||
<typeStatus id="549A884BFFD3FF96FB67CF15FAF19D1F" box="[1245,1302,1565,1591]" pageId="5" pageNumber="190">Type</typeStatus>
|
||
horizon and locality only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |