257 lines
33 KiB
XML
257 lines
33 KiB
XML
<document id="6CFFC434AE2CCB8ECFF2FA7D0812859F" ID-DOI="10.1111/j.1096-3642.2008.00439.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5445958" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630459907035" checkinUser="felipe" docAuthor="Kriwet, Jürgen, Nunn, Elizabeth V. & Klug, Stefanie" docDate="2009" docId="038887F10F7CFFA48B33AADAFEE4FCDB" docLanguage="en" docName="j.1096-3642.2008.00439.x.pdf" docOrigin="Zoological Journal of the Linnean Society 155 (2)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00439.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Rhinobatos LINK 1790" docType="treatment" docVersion="4" lastPageNumber="329" masterDocId="FFB1FF890F77FFA98A26A94BFFB8FFBB" masterDocTitle="Neoselachians (Chondrichthyes, Elasmobranchii) from the Lower and lower Upper Cretaceous of north-eastern Spain" masterLastPageNumber="347" masterPageNumber="316" pageNumber="327" updateTime="1699278836986" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
|
||
<mods:mods id="80C702D9C70A0AFA6269BB95AF3A8815" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="5E4B73D6CC08B8110897E41967F4331B">
|
||
<mods:title id="0C67FB0C5C2F7953D77722B63FE3B442">Neoselachians (Chondrichthyes, Elasmobranchii) from the Lower and lower Upper Cretaceous of north-eastern Spain</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="7F83475AE5CD6A91E2D1FAE6EDD9047A" type="personal">
|
||
<mods:role id="CBE033423B4AE9110624563FCDA21519">
|
||
<mods:roleTerm id="ADA944912A48CEDF8F8AEAFB5BB176E2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8B6F2F53B0850DF391964B5046317667">Kriwet, Jürgen</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="FAA05A65F6DC821799AFE78F2EF9C95E" type="personal">
|
||
<mods:role id="7BAC155732F503B7DADFD24E76100FA2">
|
||
<mods:roleTerm id="59E0C861507C210437300314E8B53A06">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5942E889CAE06615F3FF293CFD126020">Nunn, Elizabeth V.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="22DCE3657102C7F14FD8DE0C5721ADB8" type="personal">
|
||
<mods:role id="2E4C86FBE45399AAA2D42A3AAF62D78D">
|
||
<mods:roleTerm id="3D90A3A985406DF32C86E060D859F685">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5E51C72B26F46ADC34E643078C12BF92">Klug, Stefanie</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="CA31F91354D1E0A1AFD3CA59DF07CDD1">text</mods:typeOfResource>
|
||
<mods:relatedItem id="EDE3525C2C1B67AC251BA2A2B22E1BAF" type="host">
|
||
<mods:titleInfo id="550022DFCCBC65453915AA264218D725">
|
||
<mods:title id="36EB18D12C2D9982D44E49FDFD507446">Zoological Journal of the Linnean Society</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="E74187A2A5CC448092C882F8207540BA">
|
||
<mods:date id="7A8E7CD2FC4083EAD5CBECC032B9C1E1">2009</mods:date>
|
||
<mods:detail id="865ACA9B9FB4C5608D5A32B161BBD6B5" type="pubDate">
|
||
<mods:number id="01BF918AF4FC8D4490AECF25E7EFEDA9">2009-02-28</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="F2131B5A09F95CED275F632B1CFF9E18" type="volume">
|
||
<mods:number id="BD6D0AB79BC7366937210DE68BE29C0C">155</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="B2F6FCB9939817266424940C6E8E4C62" type="issue">
|
||
<mods:number id="975CBA32272FC647548DFD3D3B8A369E">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="C9DB00309A777E7631A68A3385B530EE" unit="page">
|
||
<mods:start id="E4A0B843379A4E6D739C8BFB4B186A2E">316</mods:start>
|
||
<mods:end id="919332707397869BFEF6B1B9F38F64F8">347</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="AAB793D15D3E2D8B9D2CD7B0ACA08E05">
|
||
<mods:url id="A3B73661650B4000E3F13F8383D25765">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00439.x</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="A84E2763445674E7B1215EEC9D4CA969">journal article</mods:classification>
|
||
<mods:identifier id="D19E4B694AB3588F7B6C1F6B28A30931" type="DOI">10.1111/j.1096-3642.2008.00439.x</mods:identifier>
|
||
<mods:identifier id="52A7EFF2A0A0D5CBDA1F7E91420F0C43" type="ISSN">0024-4082</mods:identifier>
|
||
<mods:identifier id="8273BC0D1EB9AEF2D0BAFBFDC2708F9A" type="Zenodo-Dep">5445958</mods:identifier>
|
||
<mods:identifier id="38A30295311EFBDB9D2027331CE2B00A" type="ZooBank">AC1AA5F5-C49D-4768-95F1-90574BCB9B36</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="038887F10F7CFFA48B33AADAFEE4FCDB" LSID="urn:lsid:plazi:treatment:038887F10F7CFFA48B33AADAFEE4FCDB" httpUri="http://treatment.plazi.org/id/038887F10F7CFFA48B33AADAFEE4FCDB" lastPageId="13" lastPageNumber="329" pageId="11" pageNumber="327">
|
||
<subSubSection id="C33B656C0F7CFFA28B33AADAFDCAFC1C" box="[277,626,913,935]" pageId="11" pageNumber="327" type="nomenclature">
|
||
<paragraph id="8B9E36E70F7CFFA28B33AADAFDCAFC1C" blockId="11.[277,626,913,935]" box="[277,626,913,935]" pageId="11" pageNumber="327">
|
||
<heading id="D0D6818B0F7CFFA28B33AADAFDCAFC1C" box="[277,626,913,935]" centered="true" fontSize="9" level="2" pageId="11" pageNumber="327" reason="2">
|
||
‘
|
||
<taxonomicName id="4C214D640F7CFFA28B3CAAD9FE14FC1E" ID-CoL="78N6" ID-ENA="7861" authority="SP." authorityName="LINK" authorityYear="1790" box="[282,428,914,935]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rhinopristiformes" pageId="11" pageNumber="327" phylum="Chordata" rank="genus">
|
||
<emphasis id="B955EAF50F7CFFA28B3CAAD9FE14FC1E" box="[282,428,914,935]" italics="true" pageId="11" pageNumber="327">RHINOBATOS</emphasis>
|
||
</taxonomicName>
|
||
’ SP. (
|
||
<figureCitation id="131A2A620F7CFFA28BCBAADAFDD2FC1C" box="[493,618,913,935]" captionStart="Figure 6" captionStartId="10.[164,243,1685,1704]" captionTargetBox="[204,1404,194,1655]" captionTargetId="figure-0@10.[204,1404,194,1655]" captionTargetPageId="10" captionText="Figure 6. A–D, Belemnobatis sp., anterior tooth. A, labial view. B, lateral view. C, lingual view. D, occlusal view. E–L, Spathobatis sp. E–H, antero–lateral tooth. E, labial view. F, lateral view. G, lingual view. H, occlusal view. I–L, lateral tooth. I, labial view. J, lateral view. K, lingual view. L, occlusal view. M–P, Rhinobatoidei indet. M, labial view. N, lateral view. O, lingual view. P, occlusal view. Q–T, Rhinobatos sp. Q, labial view. R, lateral view. S, lingual view. T, occlusal view. All scale bars equal 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5445970" httpUri="https://zenodo.org/record/5445970/files/figure.png" pageId="11" pageNumber="327">FIG. 6M–T</figureCitation>
|
||
)
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33B656C0F7CFFA48AB6AAF3FEE4FCDB" lastPageId="13" lastPageNumber="329" pageId="11" pageNumber="327" type="materials_examined">
|
||
<paragraph id="8B9E36E70F7CFFA28AB6AAF3FDB7FBB0" blockId="11.[144,758,951,1035]" pageId="11" pageNumber="327">
|
||
<emphasis id="B955EAF50F7CFFA28AB6AAF3FF41FC76" box="[144,249,952,973]" italics="true" pageId="11" pageNumber="327">Material:</emphasis>
|
||
A well-preserved tooth,
|
||
<collectionCode id="ED30AE220F7CFFA28868AAF3FD3DFC76" box="[590,645,952,973]" country="Spain" httpUri="http://grbio.org/cool/3qqn-wszm" name="Museo Paleontologico de la Universidad de Zaragoza" pageId="11" pageNumber="327">MPZ</collectionCode>
|
||
2005-15 (
|
||
<figureCitation id="131A2A620F7CFFA28ABEAA9DFEA9FC57" box="[152,273,982,1004]" captionStart="Figure 6" captionStartId="10.[164,243,1685,1704]" captionTargetBox="[204,1404,194,1655]" captionTargetId="figure-0@10.[204,1404,194,1655]" captionTargetPageId="10" captionText="Figure 6. A–D, Belemnobatis sp., anterior tooth. A, labial view. B, lateral view. C, lingual view. D, occlusal view. E–L, Spathobatis sp. E–H, antero–lateral tooth. E, labial view. F, lateral view. G, lingual view. H, occlusal view. I–L, lateral tooth. I, labial view. J, lateral view. K, lingual view. L, occlusal view. M–P, Rhinobatoidei indet. M, labial view. N, lateral view. O, lingual view. P, occlusal view. Q–T, Rhinobatos sp. Q, labial view. R, lateral view. S, lingual view. T, occlusal view. All scale bars equal 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5445970" httpUri="https://zenodo.org/record/5445970/files/figure.png" pageId="11" pageNumber="327">Fig. 6M–P</figureCitation>
|
||
); an isolated tooth crown with root remnants,
|
||
<collectionCode id="ED30AE220F7CFFA28AC7AABEFEA0FBB1" box="[225,280,1013,1034]" country="Spain" httpUri="http://grbio.org/cool/3qqn-wszm" name="Museo Paleontologico de la Universidad de Zaragoza" pageId="11" pageNumber="327">MPZ</collectionCode>
|
||
2005-16 (
|
||
<figureCitation id="131A2A620F7CFFA28BA8AABEFDBCFBB0" box="[398,516,1013,1035]" captionStart="Figure 6" captionStartId="10.[164,243,1685,1704]" captionTargetBox="[204,1404,194,1655]" captionTargetId="figure-0@10.[204,1404,194,1655]" captionTargetPageId="10" captionText="Figure 6. A–D, Belemnobatis sp., anterior tooth. A, labial view. B, lateral view. C, lingual view. D, occlusal view. E–L, Spathobatis sp. E–H, antero–lateral tooth. E, labial view. F, lateral view. G, lingual view. H, occlusal view. I–L, lateral tooth. I, labial view. J, lateral view. K, lingual view. L, occlusal view. M–P, Rhinobatoidei indet. M, labial view. N, lateral view. O, lingual view. P, occlusal view. Q–T, Rhinobatos sp. Q, labial view. R, lateral view. S, lingual view. T, occlusal view. All scale bars equal 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5445970" httpUri="https://zenodo.org/record/5445970/files/figure.png" pageId="11" pageNumber="327">Fig. 6Q–T</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9E36E70F7CFFA28AB6AD7DFEE8FB13" blockId="11.[144,759,1078,1192]" pageId="11" pageNumber="327">
|
||
<emphasis id="B955EAF50F7CFFA28AB6AD7DFEACFBF0" box="[144,276,1078,1099]" italics="true" pageId="11" pageNumber="327">Occurrence:</emphasis>
|
||
Samples 4 and 6 from floodplain environments with channel deposits (facies 2 and 3, respectively) of the Mosqueruela Formation (Cenomanian), Aliaga subbasin.
|
||
</paragraph>
|
||
<paragraph id="8B9E36E70F7CFFA28AB6AD98FDD1F908" blockId="11.[144,759,1234,1715]" pageId="11" pageNumber="327">
|
||
<emphasis id="B955EAF50F7CFFA28AB6AD98FEA0FB53" box="[144,280,1235,1256]" italics="true" pageId="11" pageNumber="327">Description:</emphasis>
|
||
The tooth crown is broader than it is long, and is not cuspidate. The straight transverse ridge is sharp and lingually placed. The labial edge is rounded in occlusal view, with a slight central protuberance. The lingual face is abrupt with a marked concavity at the upper part of the median uvula in profile view. The central uvula maintains contact with the lingual contour, and is short and broad, with a rounded extremity not roofing the root notch. There is one pair of well-developed lateral uvulae. The root is slightly displaced lingually and extends laterally beyond the crown. It is narrow, with a pronounced lingual notch and broad nutritive groove separating two root lobes, which are subtriangular in basal view. There are a pair of margino-lingual foramina. A single foramen opens into the nutritive groove.
|
||
</paragraph>
|
||
<paragraph id="8B9E36E70F7CFFA28AB6AF95FA89FD97" blockId="11.[144,759,1758,1903]" lastBlockId="11.[806,1422,198,1536]" pageId="11" pageNumber="327">
|
||
<emphasis id="B955EAF50F7CFFA28AB6AF95FF43F948" box="[144,251,1758,1779]" italics="true" pageId="11" pageNumber="327">Remarks:</emphasis>
|
||
This species is assigned to rhinobatoids primarily on the basis of its lingually displaced tooth root, the lobes of which become narrower lingually on both sides of the groove, and a straight and sharp transverse crest dividing the tooth crowns into a labial and lingual portion. In this, they are very similar to teeth generally assigned to
|
||
<taxonomicName id="4C214D640F7CFFA28EA5A9AFFABCFF42" authorityName="LINK" authorityYear="1790" box="[1155,1284,228,249]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rhinopristiformes" pageId="11" pageNumber="327" phylum="Chordata" rank="genus">
|
||
<emphasis id="B955EAF50F7CFFA28EA5A9AFFABCFF42" box="[1155,1284,228,249]" italics="true" pageId="11" pageNumber="327">Rhinobatos</emphasis>
|
||
</taxonomicName>
|
||
and many similar fossil teeth of rhinobatoid-like appearance from the Cretaceous. Only a few specimens, mainly from the Upper Cretaceous limestones of
|
||
<collectingCountry id="F33676770F7CFFA28EDFA80BFAE6FEED" box="[1273,1374,320,342]" name="Lebanon" pageId="11" pageNumber="327">Lebanon</collectingCountry>
|
||
, are known by articulated and more or less complete skeletons. The differences found in the dental morphology of the described species indicate that the fossil genus
|
||
<taxonomicName id="4C214D640F7CFFA28900A8F0FC1FFE6B" authorityName="LINK" authorityYear="1790" box="[806,935,443,464]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rhinopristiformes" pageId="11" pageNumber="327" phylum="Chordata" rank="genus">
|
||
<emphasis id="B955EAF50F7CFFA28900A8F0FC1FFE6B" box="[806,935,443,464]" italics="true" pageId="11" pageNumber="327">Rhinobatos</emphasis>
|
||
</taxonomicName>
|
||
comprises different rhinobatid genera (
|
||
<bibRefCitation id="EFB04B160F7CFFA28908A892FC6BFE54" author="Kriwet J" box="[814,979,473,495]" pageId="11" pageNumber="327" pagination="113 - 143" refId="ref19270" refString="Kriwet J. 1999 a. Neoselachier (Pisces, Elasmobranchii) aus der Unterkreide (unteres Barremium) von Galve und Alciane (Spainien, Provinz Teruel). Palaeo Icthyologica 9: 113 - 143." type="journal article" year="1999">Kriwet, 1999a</bibRefCitation>
|
||
). This assessment is also supported by a phylogenetic analysis by
|
||
<bibRefCitation id="EFB04B160F7CFFA28E50A8B3FA35FDB5" author="Brito PM & Dutheil DB" box="[1142,1421,504,526]" pageId="11" pageNumber="327" pagination="101 - 109" refId="ref17022" refString="Brito PM, Dutheil DB. 2005. A preliminary systematic analysis of Cretaceous guitarfishes from Lebanon. In: Arratia G, Tintori A, eds. Mesozoic fishes 3 - systematics, paleoenvironments and biodiversity. Munich: Verlag Dr. Friedrich Pfeil, 101 - 109." type="book chapter" year="2005">Brito & Dutheil (2005)</bibRefCitation>
|
||
considering several extant and fossil batoids.
|
||
</paragraph>
|
||
<paragraph id="8B9E36E70F7CFFA28919AB7EFAF5FA44" blockId="11.[806,1422,198,1536]" pageId="11" pageNumber="327">
|
||
Nominal fossil species attributed to
|
||
<taxonomicName id="4C214D640F7CFFA28EF9AB7EFAD8FDF1" authorityName="LINK" authorityYear="1790" box="[1247,1376,565,586]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rhinopristiformes" pageId="11" pageNumber="327" phylum="Chordata" rank="genus">
|
||
<emphasis id="B955EAF50F7CFFA28EF9AB7EFAD8FDF1" box="[1247,1376,565,586]" italics="true" pageId="11" pageNumber="327">Rhinobatos</emphasis>
|
||
</taxonomicName>
|
||
are listed in supplementary Appendix S1. The large number of teeth assigned to
|
||
<taxonomicName id="4C214D640F7CFFA28E52AB39FB4DFD3C" authorityName="LINK" authorityYear="1790" box="[1140,1269,626,647]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rhinopristiformes" pageId="11" pageNumber="327" phylum="Chordata" rank="genus">
|
||
<emphasis id="B955EAF50F7CFFA28E52AB39FB4DFD3C" box="[1140,1269,626,647]" italics="true" pageId="11" pageNumber="327">Rhinobatos</emphasis>
|
||
</taxonomicName>
|
||
without specific identification (16) exemplifies the problems in dealing with isolated rhinobatoid teeth. Unfortunately, rhinobatoids are rather conservative in dental morphology, rendering the identification of true
|
||
<taxonomicName id="4C214D640F7CFFA28F02ABA6FC25FC9A" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rhinopristiformes" pageId="11" pageNumber="327" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B955EAF50F7CFFA28F02ABA6FCFEFC9A" italics="true" pageId="11" pageNumber="327">Rhinobatos</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
extremely difficult and a careful revision of fossil
|
||
<taxonomicName id="4C214D640F7CFFA2894CAA61FBFBFCFB" box="[874,1091,810,832]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rhinopristiformes" pageId="11" pageNumber="327" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B955EAF50F7CFFA2894CAA61FC53FC84" box="[874,1003,810,831]" italics="true" pageId="11" pageNumber="327">Rhinobatos</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
is necessary but beyond the scope of the present paper. Teeth of
|
||
<taxonomicName id="4C214D640F7CFFA28EDEAA02FB93FCC7" authority="Cappetta & Case, 1999" authorityName="Cappetta & Case" authorityYear="1999" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rhinopristiformes" pageId="11" pageNumber="327" phylum="Chordata" rank="species" species="kiestensis">
|
||
<emphasis id="B955EAF50F7CFFA28EDEAA02FA35FCE6" box="[1272,1421,840,862]" italics="true" pageId="11" pageNumber="327">R. kiestensis</emphasis>
|
||
<bibRefCitation id="EFB04B160F7CFFA28900AA2CFB93FCC7" author="Cappetta H & Case GR" box="[806,1067,871,892]" pageId="11" pageNumber="327" pagination="1 - 112" refId="ref17599" refString="Cappetta H, Case GR. 1999. Additions aux faunes de selaciens du Cretace du Texas. Palaeo Ichthyologica 9: 1 - 112." type="journal article" year="1999">Cappetta & Case, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
, which are very similar to teeth of the extant batoid genus
|
||
<taxonomicName id="4C214D640F7CFFA28E42AACDFAB9FC20" authorityName="Norman" authorityYear="1926" box="[1124,1281,902,923]" class="Elasmobranchii" family="Rhinobatidae" genus="Aptychotrema" kingdom="Animalia" order="Rhinopristiformes" pageId="11" pageNumber="327" phylum="Chordata" rank="genus">
|
||
<emphasis id="B955EAF50F7CFFA28E42AACDFAB9FC20" box="[1124,1281,902,923]" italics="true" pageId="11" pageNumber="327">Aptychotrema</emphasis>
|
||
</taxonomicName>
|
||
, also exemplify this problem very well. A preliminary phylogenetic analysis of batoids from the Cretaceous of
|
||
<collectingCountry id="F33676770F7CFFA28900AAAAFC30FC4C" box="[806,904,993,1015]" name="Lebanon" pageId="11" pageNumber="327">Lebanon</collectingCountry>
|
||
also supports the polyphyly of fossil
|
||
<taxonomicName id="4C214D640F7CFFA28F66AAAAFC04FBAD" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rhinopristiformes" pageId="11" pageNumber="327" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B955EAF50F7CFFA28F66AAAAFCDAFBAE" italics="true" pageId="11" pageNumber="327">Rhinobatos</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFB04B160F7CFFA289E8AD4BFB6DFBAD" author="Brito PM & Dutheil DB" box="[974,1237,1024,1046]" pageId="11" pageNumber="327" pagination="101 - 109" refId="ref17022" refString="Brito PM, Dutheil DB. 2005. A preliminary systematic analysis of Cretaceous guitarfishes from Lebanon. In: Arratia G, Tintori A, eds. Mesozoic fishes 3 - systematics, paleoenvironments and biodiversity. Munich: Verlag Dr. Friedrich Pfeil, 101 - 109." type="book chapter" year="2005">Brito & Dutheil, 2005</bibRefCitation>
|
||
). The few teeth described herein share certain characters with several of the species described from the Upper Cretaceous of
|
||
<collectingCountry id="F33676770F7CFFA28900AD17FC30FBC9" box="[806,904,1116,1138]" name="Lebanon" pageId="11" pageNumber="327">Lebanon</collectingCountry>
|
||
by
|
||
<bibRefCitation id="EFB04B160F7CFFA28990AD17FB38FBC9" author="Cappetta H" box="[950,1152,1116,1138]" pageId="11" pageNumber="327" pagination="149 - 229" refId="ref17446" refString="Cappetta H. 1980 b. Les selaciens du Cretace superieur du Liban. II. Batoides. Palaeontographica, Abteilung A 168: 149 - 229." type="journal article" year="1980">Cappetta (1980b)</bibRefCitation>
|
||
, most notably with ‘
|
||
<emphasis id="B955EAF50F7CFFA28F49AD17FA30FBCA" box="[1391,1416,1116,1137]" italics="true" pageId="11" pageNumber="327">
|
||
<collectionCode id="ED30AE220F7CFFA28F49AD17FA3CFBCA" box="[1391,1412,1116,1137]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="327">R</collectionCode>
|
||
.
|
||
</emphasis>
|
||
’
|
||
<emphasis id="B955EAF50F7CFFA28900AD30FC05FB2B" box="[806,957,1147,1168]" italics="true" pageId="11" pageNumber="327">primarmatus</emphasis>
|
||
, which has a similar crown shape in oral view, and development of the central and lateral uvulae, and with ‘
|
||
<emphasis id="B955EAF50F7CFFA289D5ADF3FBB4FB76" box="[1011,1036,1208,1229]" italics="true" pageId="11" pageNumber="327">
|
||
<collectionCode id="ED30AE220F7CFFA289D5ADF3FBB0FB76" box="[1011,1032,1208,1229]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="327">R</collectionCode>
|
||
.
|
||
</emphasis>
|
||
’
|
||
<emphasis id="B955EAF50F7CFFA28E31ADF3FB27FB76" box="[1047,1183,1208,1229]" italics="true" pageId="11" pageNumber="327">intermedius</emphasis>
|
||
, which has a similar structure of the root, especially in terms of the very well-developed labial notch. ‘
|
||
<emphasis id="B955EAF50F7CFFA28E92ADBEFB75FAB1" box="[1204,1229,1269,1290]" italics="true" pageId="11" pageNumber="327">
|
||
<collectionCode id="ED30AE220F7CFFA28E92ADBEFB71FAB1" box="[1204,1225,1269,1290]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="327">R</collectionCode>
|
||
.
|
||
</emphasis>
|
||
’
|
||
<emphasis id="B955EAF50F7CFFA28EC9ADBEFA3EFAB1" box="[1263,1414,1269,1290]" italics="true" pageId="11" pageNumber="327">primarmatus</emphasis>
|
||
, however, has a lower and more cuspidate crown than that of this specimen, whilst ‘
|
||
<emphasis id="B955EAF50F7CFFA28EA9AC78FB10FAF3" box="[1167,1192,1331,1352]" italics="true" pageId="11" pageNumber="327">
|
||
<collectionCode id="ED30AE220F7CFFA28EA9AC78FB1CFAF3" box="[1167,1188,1331,1352]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="327">R</collectionCode>
|
||
.
|
||
</emphasis>
|
||
’
|
||
<emphasis id="B955EAF50F7CFFA28E91AC79FA87FAFC" box="[1207,1343,1330,1351]" italics="true" pageId="11" pageNumber="327">intermedius</emphasis>
|
||
has a significantly narrower central uvula, and no lateral uvulae. Although the Spanish species seems likely to be a new taxon, the scarcity of material and unresolved systematic position of most Cretaceous rhinobatoids precludes defining a new species. We therefore tentatively assign the teeth to the genus
|
||
<taxonomicName id="4C214D640F7CFFA28EE3ACA1FAFEFA44" authorityName="LINK" authorityYear="1790" box="[1221,1350,1514,1535]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rhinopristiformes" pageId="11" pageNumber="327" phylum="Chordata" rank="genus">
|
||
<emphasis id="B955EAF50F7CFFA28EE3ACA1FAFEFA44" box="[1221,1350,1514,1535]" italics="true" pageId="11" pageNumber="327">Rhinobatos</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9E36E70F7CFFA2890DAF60FA30F9FA" blockId="11.[811,1416,1579,1601]" box="[811,1416,1579,1601]" pageId="11" pageNumber="327">
|
||
SUBORDER
|
||
<taxonomicName id="4C214D640F7CFFA28988AF67FB4DF9FA" authority="FAM. INDET." authorityName="FAM. INDET." box="[942,1269,1580,1601]" class="Elasmobranchii" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="327" phylum="Chordata" rank="subOrder" subOrder="Rhinobatoidei">RHINOBATOIDEI FAM. INDET.</taxonomicName>
|
||
(
|
||
<figureCitation id="131A2A620F7CFFA28F21AF60FA39F9FA" box="[1287,1409,1579,1601]" captionStart="Figure 7" captionStartId="12.[164,243,1595,1614]" captionTargetBox="[291,1316,194,1565]" captionTargetId="figure-0@12.[291,1316,194,1565]" captionTargetPageId="12" captionText="Figure 7. A–D, Rhinobatoidei indet. A, labial view. B, lateral view. C, lingual view. D, occlusal. E–T, Platypterix venustulus gen. et. sp. nov. E–H, anterior male tooth. E, labial view. F, lateral view. G, lingual view. H, occlusal view. I–L, lateral male tooth. I, labial view. J, lateral view. K, lingual view. L, occlusal view. M–P, anterior female tooth. M, labial view. N, lateral view. O, lingual view. P, occlusal view. Q–T, lateral female tooth. Q, labial view. R, lateral view. S, lingual view. T, occlusal view. U–X, Celtipristis herreroi (Kriwet, 1999b). U, labial view. V, lateral view. W, lingual view. X, occlusal view. All scale bars equal 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5445972" httpUri="https://zenodo.org/record/5445972/files/figure.png" pageId="11" pageNumber="327">FIG. 7A–D</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9E36E70F7CFFA28900AF19FAFCF9D3" blockId="11.[806,1348,1618,1640]" box="[806,1348,1618,1640]" pageId="11" pageNumber="327">
|
||
<emphasis id="B955EAF50F7CFFA28900AF19FC37F9DC" box="[806,911,1618,1639]" italics="true" pageId="11" pageNumber="327">Material:</emphasis>
|
||
A single lateral tooth,
|
||
<collectionCode id="ED30AE220F7CFFA28E84AF19FB61F9DC" box="[1186,1241,1618,1639]" country="Spain" httpUri="http://grbio.org/cool/3qqn-wszm" name="Museo Paleontologico de la Universidad de Zaragoza" pageId="11" pageNumber="327">MPZ</collectionCode>
|
||
2005-17.
|
||
</paragraph>
|
||
<paragraph id="8B9E36E70F7CFFA28900AFC2FCC0F940" blockId="11.[806,1421,1673,1787]" pageId="11" pageNumber="327">
|
||
<emphasis id="B955EAF50F7CFFA28900AFC2FC13F925" box="[806,939,1673,1694]" italics="true" pageId="11" pageNumber="327">Occurrence:</emphasis>
|
||
Sample 4 from floodplain environments with channel deposits (facies 2) of the lower part of the Mosqueruela Formation (lower Cenomanian), Aliaga.
|
||
</paragraph>
|
||
<paragraph id="8B9E36E70F7CFFA48900AE57FEEDFE6A" blockId="11.[806,1421,1820,1903]" lastBlockId="13.[144,759,198,465]" lastPageId="13" lastPageNumber="329" pageId="11" pageNumber="327">
|
||
<emphasis id="B955EAF50F7CFFA28900AE57FC16F88A" box="[806,942,1820,1841]" italics="true" pageId="11" pageNumber="327">Description:</emphasis>
|
||
This is a relatively large tooth (
|
||
<emphasis id="B955EAF50F7CFFA28F19AE57FAF5F88A" box="[1343,1357,1820,1841]" italics="true" pageId="11" pageNumber="327">></emphasis>
|
||
<quantity id="4CD99B020F7CFFA28F6BAE57FA35F88A" box="[1357,1421,1820,1841]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="327" unit="mm" value="2.0">2 mm</quantity>
|
||
wide), with a massive, high and globular crown with a sub-hexagonal outline in occlusal view and a flat occlusal surface. The transverse crest is almost absent and there is no clear boundary between the oral and the lingual face. The labial margin is straight and the labial face is fairly abrupt, with a broad, rounded apron overhanging the root. The lingual edge has a massive and wide-based central uvula that also juts out over the narrow, vertical root. The root has a margino-lingual and margino-labial pair of foramina.
|
||
</paragraph>
|
||
<caption id="DF5E666F0F7BFFA58A82AF70FD35F95A" ID-DOI="http://doi.org/10.5281/zenodo.5445972" ID-Zenodo-Dep="5445972" httpUri="https://zenodo.org/record/5445972/files/figure.png" pageId="12" pageNumber="328" startId="12.[164,243,1595,1614]" targetBox="[291,1316,194,1565]" targetPageId="12">
|
||
<paragraph id="8B9E36E70F7BFFA58A82AF70FD35F95A" blockId="12.[164,1442,1595,1761]" pageId="12" pageNumber="328">
|
||
<emphasis id="B955EAF50F7BFFA58A82AF70FEB4F9F5" bold="true" box="[164,268,1595,1614]" pageId="12" pageNumber="328">Figure 7.</emphasis>
|
||
A–D,
|
||
<taxonomicName id="4C214D640F7BFFA58B7CAF70FE54F9F5" box="[346,492,1595,1614]" class="Elasmobranchii" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="327" phylum="Chordata" rank="subOrder" subOrder="Rhinobatoidei">Rhinobatoidei</taxonomicName>
|
||
indet. A, labial view. B, lateral view. C, lingual view. D, occlusal. E–T,
|
||
<emphasis id="B955EAF50F7BFFA58F38AF70FE67F9D7" bold="true" pageId="12" pageNumber="328">
|
||
<taxonomicName id="4C214D640F7BFFA58F38AF70FE9AF9D0" authorityName="Kriwet & Nunn & Klug" authorityYear="2009" class="Chondrichthyes" genus="Platypterix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="328" phylum="Chordata" rank="species" species="venustulus">
|
||
<emphasis id="B955EAF50F7BFFA58F38AF70FE9AF9D0" bold="true" italics="true" pageId="12" pageNumber="328">Platypterix venustulus</emphasis>
|
||
</taxonomicName>
|
||
gen. et. sp. nov.
|
||
</emphasis>
|
||
E–H, anterior male tooth. E, labial view. F, lateral view. G, lingual view. H, occlusal view. I–L, lateral male tooth. I, labial view. J, lateral view. K, lingual view. L, occlusal view. M–P, anterior female tooth. M, labial view. N, lateral view. O, lingual view. P, occlusal view. Q–T, lateral female tooth. Q, labial view. R, lateral view. S, lingual view. T, occlusal view. U–X,
|
||
<taxonomicName id="4C214D640F7BFFA58810AFFBFC09F978" authority="(Kriwet, 1999 b)" baseAuthorityName="Kriwet" baseAuthorityYear="1999" box="[566,945,1712,1731]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Celtipristis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="328" phylum="Chordata" rank="species" species="herreroi">
|
||
<emphasis id="B955EAF50F7BFFA58810AFFBFCBCF978" box="[566,772,1712,1731]" italics="true" pageId="12" pageNumber="328">Celtipristis herreroi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB04B160F7BFFA58934AFFBFC12F978" author="Kriwet J" box="[786,938,1712,1731]" pageId="12" pageNumber="328" pagination="337 - 346" refId="ref19308" refString="Kriwet J. 1999 b. Ptychotrygon geyeri n. sp. (Chondrichthyes, Rajiformes) from the Utrillas Formation (Upper Albian) of the central Iberian Ranges (East-Spain). Profil 16: 337 - 346." type="journal article" year="1999">Kriwet, 1999b</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. U, labial view. V, lateral view. W, lingual view. X, occlusal view. All scale bars equal 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9E36E70F7AFFA48AB6A8B1FEE4FCDB" blockId="13.[144,759,506,864]" pageId="13" pageNumber="329">
|
||
<emphasis id="B955EAF50F7AFFA48AB6A8B1FF43FDB4" box="[144,251,506,527]" italics="true" pageId="13" pageNumber="329">Remarks:</emphasis>
|
||
This tooth shares many characters with the genus
|
||
<taxonomicName id="4C214D640F7AFFA48AC7AB53FD4BFD95" authority="(Cappetta & Case, 1975)" baseAuthorityName="Cappetta & Case" baseAuthorityYear="1975" box="[225,755,536,558]" class="Elasmobranchii" family="Myliobatidae" genus="Pseudohypolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="329" phylum="Chordata" rank="genus">
|
||
<emphasis id="B955EAF50F7AFFA48AC7AB53FE0BFD96" box="[225,435,536,557]" italics="true" pageId="13" pageNumber="329">Pseudohypolophus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB04B160F7AFFA48BECAB53FD53FD95" author="Cappetta H & Case GR" box="[458,747,536,558]" pageId="13" pageNumber="329" pagination="303 - 307" refId="ref17577" refString="Cappetta H, Case GR. 1975. Selaciens nouveaux du Cretace du Texas. Geobios 8: 303 - 307." type="journal article" year="1975">Cappetta & Case, 1975</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. However,
|
||
<taxonomicName id="4C214D640F7AFFA48B21AB7CFE61FDF7" baseAuthorityName="Cappetta & Case" baseAuthorityYear="1975" box="[263,473,567,588]" class="Elasmobranchii" family="Myliobatidae" genus="Pseudohypolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="329" phylum="Chordata" rank="genus">
|
||
<emphasis id="B955EAF50F7AFFA48B21AB7CFE61FDF7" box="[263,473,567,588]" italics="true" pageId="13" pageNumber="329">Pseudohypolophus</emphasis>
|
||
</taxonomicName>
|
||
has a lower and more distinctly hexagonal crown, compared with this tooth, which is much more globular, and with a more rounded lingual margin. The crown shape is more similar to that of
|
||
<taxonomicName id="4C214D640F7AFFA48B7CABFAFD1EFD7C" authority="(Case, 1978)" baseAuthorityName="Case" baseAuthorityYear="1978" box="[346,678,689,711]" class="Chondrichthyes" family="Rhinobatidae" genus="Protoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="329" phylum="Chordata" rank="genus">
|
||
<emphasis id="B955EAF50F7AFFA48B7CABFAFDB7FD7D" box="[346,527,689,710]" italics="true" pageId="13" pageNumber="329">Protoplatyrhina</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB04B160F7AFFA48839ABFAFD26FD7C" author="Case GR" box="[543,670,689,711]" pageId="13" pageNumber="329" pagination="176 - 205" refId="ref17775" refString="Case GR. 1978. A new selachian fauna from the Judith River Formation (Campanian) of Montana. Palaeontographica, Abeilung A 160: 176 - 205." type="journal article" year="1978">Case, 1978</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, which also has a similar short extension of the central uvula. However,
|
||
<taxonomicName id="4C214D640F7AFFA48B73ABA5FDB2FCB8" baseAuthorityName="Case" baseAuthorityYear="1978" box="[341,522,750,771]" class="Chondrichthyes" family="Rhinobatidae" genus="Protoplatyrhina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="329" phylum="Chordata" rank="genus">
|
||
<emphasis id="B955EAF50F7AFFA48B73ABA5FDB2FCB8" box="[341,522,750,771]" italics="true" pageId="13" pageNumber="329">Protoplatyrhina</emphasis>
|
||
</taxonomicName>
|
||
is only known from the Maastrichtian. Unfortunately, the single, poorly preserved specimen prevents an accurate identification of this tooth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |