188 lines
20 KiB
XML
188 lines
20 KiB
XML
<document ID-DOI="10.1111/j.1096-3642.2008.00439.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5445958" approvalRequired="26" approvalRequired_for_taxonomicNames="8" approvalRequired_for_textStreams="17" approvalRequired_for_treatments="1" checkinTime="1630459907035" checkinUser="felipe" docAuthor="Kriwet, Jürgen, Nunn, Elizabeth V. & Klug, Stefanie" docDate="2009" docId="038887F10F66FFBA8958A8BBFD88FCB9" docLanguage="en" docName="j.1096-3642.2008.00439.x.pdf" docOrigin="Zoological Journal of the Linnean Society 155 (2)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00439.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.1:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="1" docTitle="Ptychotrygon Jaekel 1894" docType="treatment" docVersion="3" lastPageNumber="335" masterDocId="FFB1FF890F77FFA98A26A94BFFB8FFBB" masterDocTitle="Neoselachians (Chondrichthyes, Elasmobranchii) from the Lower and lower Upper Cretaceous of north-eastern Spain" masterLastPageNumber="347" masterPageNumber="316" pageNumber="333" updateTime="1631078048640" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo>
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<mods:title>Neoselachians (Chondrichthyes, Elasmobranchii) from the Lower and lower Upper Cretaceous of north-eastern Spain</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kriwet, Jürgen</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nunn, Elizabeth V.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Klug, Stefanie</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-02-28</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>155</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>316</mods:start>
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<mods:end>347</mods:end>
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</mods:extent>
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<mods:location>
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<mods:url>https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00439.x</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1111/j.1096-3642.2008.00439.x</mods:identifier>
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<mods:identifier type="ISSN">0024-4082</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5445958</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:038887F10F66FFBA8958A8BBFD88FCB9" httpUri="http://treatment.plazi.org/id/038887F10F66FFBA8958A8BBFD88FCB9" lastPageId="19" lastPageNumber="335" pageId="17" pageNumber="333">
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<subSubSection box="[894,1333,496,518]" pageId="17" pageNumber="333" type="nomenclature">
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<paragraph blockId="17.[894,1333,496,518]" box="[894,1333,496,518]" pageId="17" pageNumber="333">
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<heading box="[894,1333,496,518]" centered="true" fontSize="9" level="2" pageId="17" pageNumber="333" reason="2">
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GENUS
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<taxonomicName authority="JAEKEL, 1894" authorityName="Jaekel" authorityYear="1894" box="[983,1333,496,518]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="17" pageNumber="333" phylum="Chordata" rank="genus">
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<emphasis box="[983,1163,497,518]" italics="true" pageId="17" pageNumber="333">PTYCHOTRYGON</emphasis>
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<bibRefCitation author="Jaekel O" box="[1174,1333,496,518]" pageId="17" pageNumber="333" refId="ref19188" refString="Jaekel O. 1894. Die eocanen Selachier vom Monte Bolca - Ein Beitrag zur Morphogenie der Wirbeltiere. Berlin: Springer." type="book" year="1894">JAEKEL, 1894</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="19" lastPageNumber="335" pageId="17" pageNumber="333" type="materials_examined">
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<paragraph blockId="17.[806,1421,535,587]" pageId="17" pageNumber="333">
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<materialsCitation collectionCode="Genus" country="Bohemia, Czechoslovakia" location="Bohemia, Czechoslovakia" pageId="17" pageNumber="333" specimenCount="1" typeStatus="holotype">
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<emphasis box="[806,955,535,556]" italics="true" pageId="17" pageNumber="333">
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<typeStatus box="[806,858,535,556]" pageId="17" pageNumber="333">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName authority="Reuss, 1845" authorityName="Reuss" authorityYear="1845" box="[968,1421,535,557]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="17" pageNumber="333" phylum="Chordata" rank="species">
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<emphasis box="[968,1266,535,556]" italics="true" pageId="17" pageNumber="333">Ptychotrygon triangularis</emphasis>
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<bibRefCitation author="Reuss AE" box="[1279,1421,535,557]" pageId="17" pageNumber="333" refId="ref20527" refString="Reuss AE. 1845. Die Versteinerungen der Bohmischen Kreideformation. Abteilung I. Stuttgart." type="book" year="1845">Reuss, 1845</bibRefCitation>
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</taxonomicName>
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from the Turonian of
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<collectingCountry box="[1061,1356,565,587]" isEnumeration="true" pageId="17" pageNumber="333">Bohemia, Czechoslovakia</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="17.[806,1421,636,780]" pageId="17" pageNumber="333">
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<emphasis box="[806,914,636,657]" italics="true" pageId="17" pageNumber="333">Remarks:</emphasis>
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Up to now, 20 species of
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<taxonomicName authorityName="Jaekel" authorityYear="1894" box="[1210,1358,636,657]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="17" pageNumber="333" phylum="Chordata" rank="genus">
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<emphasis box="[1210,1358,636,657]" italics="true" pageId="17" pageNumber="333">Ptychotrygon</emphasis>
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</taxonomicName>
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have been described ranging from the Albian to the Maastrichtian. All species reported from the Palaeogene are either reworked specimens or misidentifications (Kriwet, J, pers. observ.).
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</paragraph>
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<paragraph blockId="17.[884,1343,843,897]" box="[884,1343,843,864]" pageId="17" pageNumber="333">
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<taxonomicName baseAuthorityName="Kriwet" baseAuthorityYear="1999" box="[884,1156,843,864]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="17" pageNumber="342" phylum="Chordata" rank="species">
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<emphasis box="[884,1156,843,864]" italics="true" pageId="17" pageNumber="333">PTYCHOTRYGON GEYERI</emphasis>
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</taxonomicName>
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<collectionCode box="[1166,1255,843,864]" pageId="17" pageNumber="333">KRIWET</collectionCode>
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, 1999B
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</paragraph>
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<paragraph blockId="17.[884,1343,843,897]" box="[993,1234,875,897]" pageId="17" pageNumber="333">
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(
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<figureCitation box="[1001,1130,875,896]" captionStart="Figure 9" captionStartId="16.[164,243,1595,1614]" captionTargetBox="[291,1316,194,1565]" captionTargetId="figure-0@16.[291,1316,194,1565]" captionTargetPageId="16" captionText="Figure 9. A–P, Celtipristis herreroi (Kriwet, 1999b). A–D, anterior tooth. A, labial view. B, lateral view. C, lingual view. D, occlusal view. E–H, antero–lateral (?) tooth. E, labial view. F, lateral view. G, lingual view. H, occlusal view. I–L, lateral tooth. I, labial view. J, lateral view. K, lingual view. L, occlusal view. M–P, anterior tooth. M, labial view. N, lateral view. O, lingual view. P, occlusal view. Q–X, Ptychotrygon geyeri (Kriwet, 1999a). Q–T, lateral tooth. Q, labial view. R, lateral view. S, lingual view. T, occlusal view. U–X, latero-posterior tooth. U, labial view. V, lateral view. W, lingual view. X, occlusal view. All scale bars equal 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5445976" httpUri="https://zenodo.org/record/5445976/files/figure.png" pageId="17" pageNumber="333">FIGS 9Q–X</figureCitation>
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,
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<figureCitation box="[1144,1226,875,897]" captionStart="Figure 10" captionStartId="18.[164,243,1558,1577]" captionTargetBox="[204,1404,196,1528]" captionTargetId="figure-63@18.[204,1404,196,1528]" captionTargetPageId="18" captionText="Figure 10. A–H, Ptychotrygon geyeri (Kriwet, 1999a). A–D, anterior tooth. A, labial view. B, lateral view. C, lingual view. D, occlusal view. E–H, lateral (juvenile?) tooth. E, labial view. F, lateral view. G, lingual view. H, occlusal view. I–L, Ptychotrygon sp., lateral tooth. I, labial view. J, lateral view. K, lingual view. L, occlusal view. M–X, Ptychotrygon pustulata sp. nov. M–P, antero-lateral tooth. M, labial view. N, lateral view. O, lingual view. P, occlusal view. Q–T, anterior tooth. Q, labial view. R, lateral view. S, lingual view. T, occlusal view. U–X, lateral tooth. U, labial view. V, lateral view. W, lingual view. X, occlusal view. All scale bars equal 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5445978" httpUri="https://zenodo.org/record/5445978/files/figure.png" pageId="17" pageNumber="333">10A–H</figureCitation>
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)
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</paragraph>
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<paragraph blockId="17.[806,1421,913,996]" pageId="17" pageNumber="333">
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<emphasis box="[806,911,913,934]" italics="true" pageId="17" pageNumber="333">Material:</emphasis>
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Six specimens,
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<collectionCode box="[1114,1169,914,935]" country="Spain" httpUri="http://grbio.org/cool/3qqn-wszm" name="Museo Paleontologico de la Universidad de Zaragoza" pageId="17" pageNumber="333">MPZ</collectionCode>
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2005–26. The specimens tend not to be complete, and often have large sections of the root missing.
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</paragraph>
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<paragraph blockId="17.[806,1421,1045,1159]" pageId="17" pageNumber="333">
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<emphasis box="[806,939,1045,1066]" italics="true" pageId="17" pageNumber="333">Occurrence:</emphasis>
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Samples 2 and 4 from floodplain environments with channel deposits (facies 1 and 2) of the Utrillas to Mosqueruela Formations, (upper Albian and lower Cenomanian), Aliaga subbasin.
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</paragraph>
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<paragraph blockId="17.[806,1422,1208,1903]" pageId="17" pageNumber="333">
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<emphasis box="[806,942,1208,1229]" italics="true" pageId="17" pageNumber="333">Description:</emphasis>
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The tooth crown is cuspidate, and broader than it is long. In occlusal view the crown is sub-triangular, and often has a fairly abrupt lingual face, with the main cusp lingually placed. The crown has a transverse occlusal crest and short vertical folds on the labial crown base. The apron is short and broad, and is well detached from the labial contour. The lingual face has a short uvula, with a rounded extremity, above which there is a weak central depression for tooth interlocking. The upper margin of the central depression is bounded by a weak ridge. The root is low, and appears not to extend below the crown. There are two root lobes separated by a well-developed median groove, with a large central foramen.
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</paragraph>
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<paragraph blockId="17.[806,1422,1208,1903]" pageId="17" pageNumber="333">
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A single specimen (
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<figureCitation box="[1060,1193,1667,1689]" captionStart="Figure 10" captionStartId="18.[164,243,1558,1577]" captionTargetBox="[204,1404,196,1528]" captionTargetId="figure-63@18.[204,1404,196,1528]" captionTargetPageId="18" captionText="Figure 10. A–H, Ptychotrygon geyeri (Kriwet, 1999a). A–D, anterior tooth. A, labial view. B, lateral view. C, lingual view. D, occlusal view. E–H, lateral (juvenile?) tooth. E, labial view. F, lateral view. G, lingual view. H, occlusal view. I–L, Ptychotrygon sp., lateral tooth. I, labial view. J, lateral view. K, lingual view. L, occlusal view. M–X, Ptychotrygon pustulata sp. nov. M–P, antero-lateral tooth. M, labial view. N, lateral view. O, lingual view. P, occlusal view. Q–T, anterior tooth. Q, labial view. R, lateral view. S, lingual view. T, occlusal view. U–X, lateral tooth. U, labial view. V, lateral view. W, lingual view. X, occlusal view. All scale bars equal 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5445978" httpUri="https://zenodo.org/record/5445978/files/figure.png" pageId="17" pageNumber="333">Fig. 10Q–X</figureCitation>
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), which may represent a juvenile, displays a cuspidate crown, with a clear transverse crest and a longitudinal centrolabial ridge, which divides into several v-shaped ridges away from the apex. The apron is fairly broad with a rounded extremity. It is clearly detached from the labial contour and bent significantly basally.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.5445978" ID-Zenodo-Dep="5445978" httpUri="https://zenodo.org/record/5445978/files/figure.png" pageId="18" pageNumber="334" startId="18.[164,243,1558,1577]" targetBox="[204,1404,196,1528]" targetPageId="18">
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<paragraph blockId="18.[164,1442,1558,1724]" pageId="18" pageNumber="334">
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<emphasis bold="true" box="[164,281,1558,1577]" pageId="18" pageNumber="334">Figure 10.</emphasis>
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A–H,
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<taxonomicName authority="(Kriwet, 1999 a)" baseAuthorityName="Kriwet" baseAuthorityYear="1999" box="[355,732,1558,1577]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="18" pageNumber="334" phylum="Chordata" rank="species">
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<emphasis box="[355,559,1558,1577]" italics="true" pageId="18" pageNumber="334">Ptychotrygon geyeri</emphasis>
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(
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<bibRefCitation author="Kriwet J" box="[573,725,1558,1577]" pageId="18" pageNumber="334" pagination="113 - 143" refId="ref19270" refString="Kriwet J. 1999 a. Neoselachier (Pisces, Elasmobranchii) aus der Unterkreide (unteres Barremium) von Galve und Alciane (Spainien, Provinz Teruel). Palaeo Icthyologica 9: 113 - 143." type="journal article" year="1999">Kriwet, 1999a</bibRefCitation>
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)
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</taxonomicName>
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. A–D, anterior tooth. A, labial view. B, lateral view. C, lingual view. D, occlusal view. E–H, lateral (juvenile?) tooth. E, labial view. F, lateral view. G, lingual view. H, occlusal view. I–L,
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<taxonomicName box="[164,343,1616,1636]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" kingdom="Animalia" order="Rajiformes" pageId="18" pageNumber="334" phylum="Chordata" rank="species">
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<emphasis box="[164,300,1616,1635]" italics="true" pageId="18" pageNumber="334">Ptychotrygon</emphasis>
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sp.
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</taxonomicName>
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, lateral tooth. I, labial view. J, lateral view. K, lingual view. L, occlusal view. M–X,
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<emphasis bold="true" pageId="18" pageNumber="334">
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<taxonomicName authority="Kriwet & Nunn & Klug, 2009" authorityName="Kriwet & Nunn & Klug" authorityYear="2009" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="18" pageNumber="341" phylum="Chordata" rank="species" status="sp.">
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<emphasis bold="true" italics="true" pageId="18" pageNumber="334">Ptychotrygon pustulata</emphasis>
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<taxonomicNameLabel box="[288,321,1646,1665]" pageId="18" pageNumber="334" rank="species">sp.</taxonomicNameLabel>
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</taxonomicName>
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nov.
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</emphasis>
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M–P, antero-lateral tooth. M, labial view. N, lateral view. O, lingual view. P, occlusal view. Q–T, anterior tooth. Q, labial view. R, lateral view. S, lingual view. T, occlusal view. U–X, lateral tooth. U, labial view. V, lateral view. W, lingual view. X, occlusal view. All scale bars equal 0.5 mm.
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</paragraph>
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</caption>
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<paragraph blockId="18.[164,779,1789,1903]" lastBlockId="19.[144,759,198,771]" lastPageId="19" lastPageNumber="335" pageId="18" pageNumber="334">
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<emphasis box="[164,271,1789,1810]" italics="true" pageId="18" pageNumber="334">Remarks:</emphasis>
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This species was described by
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<bibRefCitation author="Kriwet J" pageId="18" pageNumber="334" pagination="337 - 346" refId="ref19308" refString="Kriwet J. 1999 b. Ptychotrygon geyeri n. sp. (Chondrichthyes, Rajiformes) from the Utrillas Formation (Upper Albian) of the central Iberian Ranges (East-Spain). Profil 16: 337 - 346." type="journal article" year="1999">Kriwet (1999b)</bibRefCitation>
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from the Albian Utrillas Formation of the Aliaga subbasin. The teeth from the Cenomanian Mosqueruela Formation of the Aliaga subbasin display the characteristics of the species, e.g. short transversal ridges on the labial face on the crown shoulders, a short vertical ridge on the labial protuberance and a smooth lingual crown face except for some very delicate folds related to the lingual interlocking depression. The labial short transverse ridges below the transversal cutting edge are very faint and subtle in most specimens recovered from the Cenomanian. The occurrence of
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<taxonomicName baseAuthorityName="Kriwet" baseAuthorityYear="1999" box="[443,672,320,342]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="19" pageNumber="335" phylum="Chordata" rank="species">
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<emphasis box="[443,672,320,342]" italics="true" pageId="19" pageNumber="335">Ptychotrygon geyeri</emphasis>
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</taxonomicName>
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in the lower Cenomanian extends its range from the latest Early Cretaceous into the earliest Late Cretaceous.
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</paragraph>
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<paragraph blockId="19.[144,759,198,771]" pageId="19" pageNumber="335">
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The small, supposedly juvenile tooth resembles teeth of
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<taxonomicName authorityName="Cappetta" authorityYear="1980" box="[240,392,443,464]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Libanopristis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="19" pageNumber="335" phylum="Chordata" rank="genus">
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<emphasis box="[240,392,443,464]" italics="true" pageId="19" pageNumber="335">Libanopristis</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Cappetta" authorityYear="1980" box="[452,590,443,464]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Micropristis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="19" pageNumber="335" phylum="Chordata" rank="genus">
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<emphasis box="[452,590,443,464]" italics="true" pageId="19" pageNumber="335">Micropristis</emphasis>
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</taxonomicName>
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from the Cenomanian of
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<collectingCountry box="[287,385,473,495]" name="Lebanon" pageId="19" pageNumber="335">Lebanon</collectingCountry>
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(
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<bibRefCitation author="Cappetta H" box="[405,600,473,495]" pageId="19" pageNumber="335" pagination="149 - 229" refId="ref17446" refString="Cappetta H. 1980 b. Les selaciens du Cretace superieur du Liban. II. Batoides. Palaeontographica, Abteilung A 168: 149 - 229." type="journal article" year="1980">Cappetta, 1980b</bibRefCitation>
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). However, it differs from both in several major characteristics such as a root not extending below the crown, a comparably higher root, absent lingual ridge above the interlocking depression as in
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<taxonomicName authorityName="Cappetta" authorityYear="1980" box="[451,603,596,617]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Libanopristis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="19" pageNumber="335" phylum="Chordata" rank="genus">
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<emphasis box="[451,603,596,617]" italics="true" pageId="19" pageNumber="335">Libanopristis</emphasis>
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</taxonomicName>
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and a more pronounced labial ornamentation than in
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<taxonomicName authorityName="Cappetta" authorityYear="1980" class="Elasmobranchii" family="Sclerorhynchidae" genus="Micropristis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="19" pageNumber="335" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="19" pageNumber="335">Micropristis</emphasis>
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</taxonomicName>
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. The absence of well-marked but faint or even absent labial and lingual transversal crests and the presence of short labial ridges below the lateral heels attribute this specimen to
|
||
<taxonomicName baseAuthorityName="Kriwet" baseAuthorityYear="1999" box="[456,553,749,770]" class="Elasmobranchii" family="Sclerorhynchidae" genus="Ptychotrygon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="19" pageNumber="335" phylum="Chordata" rank="species">
|
||
<emphasis box="[456,553,749,770]" italics="true" pageId="19" pageNumber="335">P. geyeri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |