322 lines
36 KiB
XML
322 lines
36 KiB
XML
<document ID-DOI="10.37520/aemnp.2022.022" ID-ISSN="1804-6487" ID-Zenodo-Dep="7617226" ID-ZooBank="B50198A3-A761-4BCE-B7EA-A892DBED6FD3" approvalRequired="11" approvalRequired_for_textStreams="9" approvalRequired_for_treatments="2" checkinTime="1675792646716" checkinUser="felipe" docAuthor="Šumpich, Jan" docDate="2022" docId="038887B4FFD5151DFE9FFCC9FEB5F863" docLanguage="en" docName="ActaEntMusNatPra.62.2.401-409.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 62 (2)" docSource="http://dx.doi.org/10.37520/aemnp.2022.022" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.6:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="6" docTitle="Megacraspedus bidzilyai Šumpich 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="404" masterDocId="FFB1FFCCFFD4151EFF8DFFDDFFD8FFB6" masterDocTitle="Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group" masterLastPageNumber="409" masterPageNumber="401" pageNumber="402" updateTime="1675793500789" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Megacraspedus (Lepidoptera: Gelechiidae) of the Altai Mountains with description of a new species belonging to the M. majorella group</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Šumpich, Jan</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>62</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>401</mods:start>
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<mods:end>409</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.37520/aemnp.2022.022</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.37520/aemnp.2022.022</mods:identifier>
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<mods:identifier type="ISSN">1804-6487</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7617226</mods:identifier>
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<mods:identifier type="ZooBank">B50198A3-A761-4BCE-B7EA-A892DBED6FD3</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:038887B4FFD5151DFE9FFCC9FEB5F863" httpUri="http://treatment.plazi.org/id/038887B4FFD5151DFE9FFCC9FEB5F863" lastPageId="3" lastPageNumber="404" pageId="1" pageNumber="402">
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<subSubSection box="[274,636,788,812]" pageId="1" pageNumber="402" type="nomenclature">
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<paragraph blockId="1.[274,636,788,812]" box="[274,636,788,812]" pageId="1" pageNumber="402">
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<heading box="[274,636,788,812]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="402" reason="3">
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<emphasis bold="true" box="[274,636,788,812]" pageId="1" pageNumber="402">
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<taxonomicName authority="Šumpich, 2022" authorityName="Šumpich" authorityYear="2022" box="[274,544,788,812]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="bidzilyai" status="sp. nov.">
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<emphasis bold="true" box="[274,544,788,812]" italics="true" pageId="1" pageNumber="402">Megacraspedus bidzilyai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[551,636,789,812]" pageId="1" pageNumber="402" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection box="[395,515,820,839]" pageId="1" pageNumber="402" type="description">
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<paragraph blockId="1.[150,760,820,1110]" box="[395,515,820,839]" pageId="1" pageNumber="402">
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(
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<figureCitation box="[400,477,820,839]" captionStart-0="Figs 1–4" captionStart-1="Figs 5–7" captionStartId-0="1.[150,187,2031,2050]" captionStartId-1="2.[151,188,2031,2050]" captionTargetBox-0="[155,1436,1162,2009]" captionTargetBox-1="[152,1435,461,2014]" captionTargetId-0="figure-488@1.[151,1436,1162,2021]" captionTargetId-1="figure-64@2.[151,1436,446,2021]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figs 1–4. Adults of Megacraspedus bidzilyai sp. nov., Russia, Altai Mts., details in text. 1–3 – males: 1 – holotype (barcoded); 2 – paratype; 3 – head (enlarged); 4 – female, paratype." captionText-1="Figs 5–7. Genitalia of Megacraspedus bidzilyai sp. nov., Russia, Altai Mts., details in text. 5–6 – male genitalia, paratypes: a – phallus; b – detail of apical part of phallus. 7 – female genitalia: c – detail of ostium; d – signum; e – oposite view of signum." figureDoi-0="http://doi.org/10.5281/zenodo.7617228" figureDoi-1="http://doi.org/10.5281/zenodo.7617230" httpUri-0="https://zenodo.org/record/7617228/files/figure.png" httpUri-1="https://zenodo.org/record/7617230/files/figure.png" pageId="1" pageNumber="402">Figs 1–7</figureCitation>
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,
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<figureCitation box="[486,508,820,839]" captionStart="Figs 19–21" captionStartId="7.[150,187,2028,2047]" captionTargetBox="[150,1435,506,2013]" captionTargetId="figure-12@7.[150,1435,506,2013]" captionTargetPageId="7" captionText="Figs 19–21. Habitats of Megacraspedus species. 19–20 – M. bidzilyai sp. nov., Russia,Altai Mts. 21 – M. latiuncus Huemer & Karsholt, 2018, Kyrgyzstan, North Tian-Shan Mts. (19 – photographed by Z. Růžičková, 21 – photographed by I. Dvořák)." figureDoi="http://doi.org/10.5281/zenodo.7617238" httpUri="https://zenodo.org/record/7617238/files/figure.png" pageId="1" pageNumber="402">19</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="402" type="materials_examined">
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<paragraph blockId="1.[150,760,820,1110]" pageId="1" pageNumber="402">
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<emphasis bold="true" box="[150,286,854,873]" pageId="1" pageNumber="402">Type material.</emphasis>
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<materialsCitation collectingDate="2017-07-27" collectingDateMax="2017-07-28" collectingDateMin="2017-07-27" collectionCode="NMPC, NW" collectorName="J. Sumpich" country="Russia" county="Belyashi" elevation="1400" latitude="49.865555" location="Karagem" longLatPrecision="18" longitude="87.172775" municipality="Argut" pageId="1" pageNumber="402" specimenCount="1" stateProvince="Altay" typeStatus="holotype">
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<typeStatus box="[294,384,854,873]" pageId="1" pageNumber="402">HOLOTYPE</typeStatus>
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: J (
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<collectionCode box="[427,493,854,873]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="402">NMPC</collectionCode>
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):
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<emphasis bold="true" box="[511,759,854,873]" pageId="1" pageNumber="402">
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<collectingCountry box="[511,596,854,873]" name="Russia" pageId="1" pageNumber="402">RUSSIA</collectingCountry>
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:
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<collectingRegion box="[607,657,854,873]" country="Russia" name="Altay" pageId="1" pageNumber="402">ALTAI</collectingRegion>
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REඉUBLIC:
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</emphasis>
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<collectingCounty box="[150,223,881,900]" pageId="1" pageNumber="402">Belyashi</collectingCounty>
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(Dzhazator) env.(
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<quantity box="[374,427,881,900]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="402" unit="km" value="25.0">25 km</quantity>
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<collectionCode box="[431,469,881,900]" pageId="1" pageNumber="402">NW</collectionCode>
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), confluence of
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<collectingMunicipality box="[596,645,881,900]" pageId="1" pageNumber="402">Argut</collectingMunicipality>
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and
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<location LSID="urn:lsid:plazi:treatment:038887B4FFD5151DFE9FFCC9FEB5F863:8EFE6079FFD5151FFD27FCACFD2FFC32" box="[682,759,881,900]" country="Russia" county="Belyashi" latitude="49.865555" longLatPrecision="18" longitude="87.172775" municipality="Argut" name="Karagem" pageId="1" pageNumber="402" stateProvince="Altay">Karagem</location>
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rivers,
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<geoCoordinate box="[213,316,907,926]" degrees="49" direction="north" minutes="51" orientation="latitude" pageId="1" pageNumber="402" precision="15" seconds="56" value="49.865555">49°51′56″N</geoCoordinate>
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,
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<geoCoordinate box="[328,429,907,926]" degrees="87" direction="east" minutes="10" orientation="longitude" pageId="1" pageNumber="402" precision="15" seconds="22" value="87.172775">87°10′22″E</geoCoordinate>
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, rocky steppe,
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<elevation box="[566,683,907,926]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="402" unit="m" value="1400.0">
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<quantity box="[566,634,907,926]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="402" unit="m" value="1400.0">1400 m</quantity>
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a.s.l.
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</elevation>
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,
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<date pageId="1" pageNumber="402" value="2017-07-27" valueMax="2017-07-28" valueMin="2017-07-27">
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<collectingDate pageId="1" pageNumber="402" value="2017-07-27" valueMax="2017-07-28" valueMin="2017-07-27">27.–28. vii.2017</collectingDate>
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</date>
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(Barcode NMPC-LEP-0381),
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<collectorName box="[490,588,934,953]" pageId="1" pageNumber="402">J. Šumpich</collectorName>
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leg.
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</materialsCitation>
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<typeStatus box="[635,724,934,953]" pageId="1" pageNumber="402">PARATYPES</typeStatus>
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: 51 JJ
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<specimenCount box="[187,218,954,980]" pageId="1" pageNumber="402" type="female">1 ♀</specimenCount>
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(49 JJ
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<specimenCount box="[292,323,954,980]" pageId="1" pageNumber="402" type="female">1 ♀</specimenCount>
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<collectionCode box="[329,392,961,980]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="402">NMPC</collectionCode>
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, 2 JJ
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<collectionCode box="[451,518,961,980]" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" pageId="1" pageNumber="402">ZMKU</collectionCode>
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): the same collecting data as
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<typeStatus box="[150,227,987,1006]" pageId="1" pageNumber="402">holotype</typeStatus>
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, gen. prep. Šumpich 18239, 21353, 21357.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="404" pageId="1" pageNumber="402" type="description">
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<paragraph blockId="1.[150,760,820,1110]" lastBlockId="1.[828,1437,190,1109]" pageId="1" pageNumber="402">
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<emphasis bold="true" box="[150,358,1022,1046]" pageId="1" pageNumber="402">
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Description.
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<emphasis bold="true" box="[291,351,1022,1046]" italics="true" pageId="1" pageNumber="402">Adult</emphasis>
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.
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</emphasis>
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<emphasis box="[362,416,1023,1046]" italics="true" pageId="1" pageNumber="402">Male</emphasis>
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(
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<figureCitation box="[428,521,1023,1046]" captionStart="Figs 1–4" captionStartId="1.[150,187,2031,2050]" captionTargetBox="[155,1436,1162,2009]" captionTargetId="figure-488@1.[151,1436,1162,2021]" captionTargetPageId="1" captionText="Figs 1–4. Adults of Megacraspedus bidzilyai sp. nov., Russia, Altai Mts., details in text. 1–3 – males: 1 – holotype (barcoded); 2 – paratype; 3 – head (enlarged); 4 – female, paratype." figureDoi="http://doi.org/10.5281/zenodo.7617228" httpUri="https://zenodo.org/record/7617228/files/figure.png" pageId="1" pageNumber="402">Figs 1–3</figureCitation>
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). Wingspan
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<quantity box="[647,759,1023,1046]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="1" pageNumber="402" unit="mm" value="19.0" valueMax="21.0" valueMin="17.0">17–21 mm</quantity>
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(on average
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<quantity box="[280,358,1055,1078]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="402" unit="mm" value="20.0">20 mm</quantity>
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). Segment 2 of labial palpus comparatively long, at upper border of outer surface white dusted with brown scales, lower part densely dusted with brown scales, exceptionally almost brown (
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<figureCitation box="[1214,1277,222,245]" captionStart="Figs 1–4" captionStartId="1.[150,187,2031,2050]" captionTargetBox="[155,1436,1162,2009]" captionTargetId="figure-488@1.[151,1436,1162,2021]" captionTargetPageId="1" captionText="Figs 1–4. Adults of Megacraspedus bidzilyai sp. nov., Russia, Altai Mts., details in text. 1–3 – males: 1 – holotype (barcoded); 2 – paratype; 3 – head (enlarged); 4 – female, paratype." figureDoi="http://doi.org/10.5281/zenodo.7617228" httpUri="https://zenodo.org/record/7617228/files/figure.png" pageId="1" pageNumber="402">Fig. 3</figureCitation>
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), inner surface similarly coloured. Segment 3 white, mottled with brown scales, especially near top. Antennal scape with several long pecten scales. Flagellum brown, distinctly ringed with creamy ochre to whitish. Head, thorax and tegula creamy whitish, weakly mottled with brown scales. Forewing grey, slightly darker near costa, two small blackish dots at 1/3 and 2/3 of wing, fringes light grey. Hindwing dark grey with grey fringes.
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</paragraph>
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<paragraph blockId="1.[828,1437,190,1109]" pageId="1" pageNumber="402">
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<emphasis box="[858,938,510,533]" italics="true" pageId="1" pageNumber="402">Female</emphasis>
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(
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<figureCitation box="[954,1022,510,533]" captionStart="Figs 1–4" captionStartId="1.[150,187,2031,2050]" captionTargetBox="[155,1436,1162,2009]" captionTargetId="figure-488@1.[151,1436,1162,2021]" captionTargetPageId="1" captionText="Figs 1–4. Adults of Megacraspedus bidzilyai sp. nov., Russia, Altai Mts., details in text. 1–3 – males: 1 – holotype (barcoded); 2 – paratype; 3 – head (enlarged); 4 – female, paratype." figureDoi="http://doi.org/10.5281/zenodo.7617228" httpUri="https://zenodo.org/record/7617228/files/figure.png" pageId="1" pageNumber="402">Fig. 4</figureCitation>
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). Wingspan
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<quantity box="[1157,1257,510,533]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="1" pageNumber="402" unit="mm" value="19.5">19.5 mm</quantity>
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. Similar to male but generally lighter, hindwings whitish. Head of the only available female is lost.
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</paragraph>
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<paragraph blockId="1.[828,1437,190,1109]" pageId="1" pageNumber="402">
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<emphasis bold="true" box="[858,967,605,629]" pageId="1" pageNumber="402">
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<emphasis bold="true" box="[858,960,605,629]" italics="true" pageId="1" pageNumber="402">Variation</emphasis>
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.
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</emphasis>
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The degree of variability is very low. Fresh specimens appear darker and more contrasting.
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</paragraph>
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<paragraph blockId="1.[828,1437,190,1109]" lastBlockId="2.[151,761,190,405]" lastPageId="2" lastPageNumber="403" pageId="1" pageNumber="402">
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<emphasis bold="true" box="[858,1015,669,693]" italics="true" pageId="1" pageNumber="402">Male genitalia</emphasis>
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(
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<figureCitation box="[1027,1123,670,693]" captionStart="Figs 5–7" captionStartId="2.[151,188,2031,2050]" captionTargetBox="[152,1435,461,2014]" captionTargetId="figure-64@2.[151,1436,446,2021]" captionTargetPageId="2" captionText="Figs 5–7. Genitalia of Megacraspedus bidzilyai sp. nov., Russia, Altai Mts., details in text. 5–6 – male genitalia, paratypes: a – phallus; b – detail of apical part of phallus. 7 – female genitalia: c – detail of ostium; d – signum; e – oposite view of signum." figureDoi="http://doi.org/10.5281/zenodo.7617230" httpUri="https://zenodo.org/record/7617230/files/figure.png" pageId="1" pageNumber="402">Figs 5–6</figureCitation>
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). Uncus broad, almost square, slightly rounded in corners of apical part. Gnathos stout, broad, slightly longer than uncus, middle part is straight, at end curved and sharply pointed.Tegumen broad, in shape of isosceles triangle, pedunculi broadly oval. Valva broad, regularly equal, extending to near top of uncus, apex distinctly rounded laterally and inwardly ended sharply, covered with setae, more densely in apex area. Separated sacculus not developed. Posterior margin of vinculum with comparatively deeper medial emargination. Saccus sub-triangular with weakly pointed apex, ratio maximum width to length approximately 0.7, lateral sclerites approximately 0.7 times maximum width of saccus. Phallus straight, bulbous coecum, in middle of phallus small sclerotised microplate with minute spines, ductus ejaculatorius twisted several times, apically with short interior sclerotisation.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7617228" ID-Zenodo-Dep="7617228" httpUri="https://zenodo.org/record/7617228/files/figure.png" pageId="1" pageNumber="402" startId="1.[150,187,2031,2050]" targetBox="[155,1436,1162,2009]" targetPageId="1">
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||
<paragraph blockId="1.[150,1436,2031,2077]" pageId="1" pageNumber="402">
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Figs 1–4. Adults of
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<taxonomicName authority="Šumpich, 2022" authorityName="Šumpich" authorityYear="2022" box="[322,536,2031,2050]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="bidzilyai" status="sp. nov.">
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<emphasis box="[322,536,2031,2050]" italics="true" pageId="1" pageNumber="402">Megacraspedus bidzilyai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[542,611,2031,2050]" pageId="1" pageNumber="402" rank="species">sp. nov.</taxonomicNameLabel>
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, Russia, Altai Mts., details in text. 1–3 – males: 1 – holotype (barcoded); 2 – paratype; 3 – head (enlarged); 4 – female, paratype.
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||
</paragraph>
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||
</caption>
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||
<paragraph blockId="2.[151,761,190,405]" lastBlockId="3.[150,760,190,2005]" lastPageId="3" lastPageNumber="404" pageId="2" pageNumber="403">
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<emphasis bold="true" box="[181,367,253,277]" italics="true" pageId="2" pageNumber="403">Female genitalia</emphasis>
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(
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<figureCitation box="[381,446,254,277]" captionStart="Figs 5–7" captionStartId="2.[151,188,2031,2050]" captionTargetBox="[152,1435,461,2014]" captionTargetId="figure-64@2.[151,1436,446,2021]" captionTargetPageId="2" captionText="Figs 5–7. Genitalia of Megacraspedus bidzilyai sp. nov., Russia, Altai Mts., details in text. 5–6 – male genitalia, paratypes: a – phallus; b – detail of apical part of phallus. 7 – female genitalia: c – detail of ostium; d – signum; e – oposite view of signum." figureDoi="http://doi.org/10.5281/zenodo.7617230" httpUri="https://zenodo.org/record/7617230/files/figure.png" pageId="2" pageNumber="403">Fig. 7</figureCitation>
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). Papilla analis large, elongated, apically rounded. Anterior apophysis very short, posterior apophysis 3.5 times longer than anterior apophysis. Segment VIII distinct, slightly sclerotized distally. Subgenital plate with triangular subostial sclerotisation, distally pointed, anteriorly delimiting oblong ostium bursae, anterior margin smooth, slightly concave, connecting with anterior apophysis. Colliculum comparatively short, tube-shaped, sclerotised, slightly expanding anteriorly. Ductus bursae gradually widening to weakly delimited, comparatively broad corpus bursae, entire length of ductus and corpus bursae only slightly longer than entire length of papilla analis and posterior apophysis. Signum small, in shape of rounded spiny plate.
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7617230" ID-Zenodo-Dep="7617230" httpUri="https://zenodo.org/record/7617230/files/figure.png" pageId="2" pageNumber="403" startId="2.[151,188,2031,2050]" targetBox="[152,1435,461,2014]" targetPageId="2">
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<paragraph blockId="2.[151,1437,2031,2077]" pageId="2" pageNumber="403">
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Figs 5–7. Genitalia of
|
||
<taxonomicName authority="Šumpich, 2022" authorityName="Šumpich" authorityYear="2022" box="[347,561,2031,2050]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="bidzilyai" status="sp. nov.">
|
||
<emphasis box="[347,561,2031,2050]" italics="true" pageId="2" pageNumber="403">Megacraspedus bidzilyai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[568,638,2031,2050]" pageId="2" pageNumber="403" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, Russia, Altai Mts., details in text. 5–6 – male genitalia, paratypes: a – phallus; b – detail of apical part of phallus. 7 – female genitalia: c – detail of ostium; d – signum; e – oposite view of signum.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="3" pageNumber="404" type="diagnosis">
|
||
<paragraph blockId="3.[150,760,190,2005]" pageId="3" pageNumber="404">
|
||
<emphasis bold="true" box="[150,398,254,277]" pageId="3" pageNumber="404">Differential diagnosis.</emphasis>
|
||
<taxonomicName authority="Šumpich, 2022" authorityName="Šumpich" authorityYear="2022" box="[404,671,254,277]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="bidzilyai" status="sp. nov.">
|
||
<emphasis box="[404,671,254,277]" italics="true" pageId="3" pageNumber="404">Megacraspedus bidzilyai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[677,760,254,277]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is characterised by grey coloration and two small black dots on the forewings in combination with a fairly large wingspan. It is most similar to
|
||
<taxonomicName authority="Caradja, 1920" authorityName="Caradja" authorityYear="1920" box="[466,755,350,373]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="majorella">
|
||
<emphasis box="[466,602,350,373]" italics="true" pageId="3" pageNumber="404">M. majorella</emphasis>
|
||
<bibRefCitation author="CARADJA A." box="[606,755,350,373]" pageId="3" pageNumber="404" pagination="75 - 179" refId="ref4406" refString="CARADJA A. 1920: Beitrag zur Kenntnis der geographischen Verbreitung der Mikrolepidopteren des palaearktischen Faunengebietes nebst Beschreibung neuer Formen. III. Teil. Deutsche Entomologische Zeitschrift, Iris 34: 75 - 179." type="journal article" year="1920">Caradja, 1920</bibRefCitation>
|
||
</taxonomicName>
|
||
, but
|
||
<taxonomicName authority="Šumpich, 2022" authorityName="Šumpich" authorityYear="2022" box="[193,323,382,405]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="bidzilyai" status="sp. nov.">
|
||
<emphasis box="[193,323,382,405]" italics="true" pageId="3" pageNumber="404">M. bidzilyai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[331,416,382,405]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is overall darker, a particularly distinct difference is in the hindwing, which is very light in
|
||
<taxonomicName authorityName="Caradja" authorityYear="1920" box="[180,321,446,469]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="majorella">
|
||
<emphasis box="[180,321,446,469]" italics="true" pageId="3" pageNumber="404">M. majorella</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, in
|
||
<taxonomicName authority="Šumpich, 2022" authorityName="Šumpich" authorityYear="2022" box="[486,617,446,469]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="408" phylum="Arthropoda" rank="species" species="bidzilyai" status="sp. nov.">
|
||
M.
|
||
<emphasis box="[525,617,446,469]" italics="true" pageId="3" pageNumber="404">bidzilyai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[626,714,446,469]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, the forewing is densely dusted with darker scales and appears finely speckled, while in
|
||
<taxonomicName authorityName="Caradja" authorityYear="1920" box="[411,548,510,533]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="majorella">
|
||
<emphasis box="[411,548,510,533]" italics="true" pageId="3" pageNumber="404">M. majorella</emphasis>
|
||
</taxonomicName>
|
||
the overall coloring gives a solid plain impression. Male genitalia of
|
||
<taxonomicName authority="Šumpich, 2022" authorityName="Šumpich" authorityYear="2022" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="bidzilyai" status="sp. nov.">
|
||
<emphasis italics="true" pageId="3" pageNumber="404">M. bidzilyai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[215,298,574,597]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are unique within the genus due to broadly sub-rectangular uncus, stout gnathos and a small sclerotised spined patch on the phallus. Similarity is evident only with species from the
|
||
<taxonomicName authorityName="Caradja" authorityYear="1920" box="[342,484,670,693]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="majorella">
|
||
<emphasis box="[342,484,670,693]" italics="true" pageId="3" pageNumber="404">M. majorella</emphasis>
|
||
</taxonomicName>
|
||
group, but
|
||
<taxonomicName authorityName="Caradja" authorityYear="1920" box="[617,759,670,693]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="majorella">
|
||
<emphasis box="[617,759,670,693]" italics="true" pageId="3" pageNumber="404">M. majorella</emphasis>
|
||
</taxonomicName>
|
||
has distinctly thinner valvae and broader saccus while
|
||
<taxonomicName authority="Huemer & Karsholt, 2018" authorityName="Huemer & Karsholt" authorityYear="2018" box="[150,582,734,757]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="latiuncus">
|
||
<emphasis box="[150,287,734,757]" italics="true" pageId="3" pageNumber="404">M. latiuncus</emphasis>
|
||
<bibRefCitation author="HUEMER P. & KARSHOLT O." box="[296,582,734,757]" pageId="3" pageNumber="404" pagination="1 - 278" refId="ref4652" refString="HUEMER P. & KARSHOLT O. 2018: Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1 - 278." type="journal article" year="2018">Huemer & Karsholt, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
has a less stout gnathos, posteriorly tapering valves and differently shaped spined patch on the phallus.
|
||
</paragraph>
|
||
<paragraph blockId="3.[150,760,190,2005]" pageId="3" pageNumber="404">
|
||
<emphasis bold="true" box="[150,338,830,853]" pageId="3" pageNumber="404">Molecular data.</emphasis>
|
||
BIN: BOLD:ADR7289. The average intraspecific distance of the barcode region is unknown (n = 1). The minimum distance to the nearest neighbour,
|
||
<taxonomicName authorityName="Huemer & Karsholt" authorityYear="2018" box="[150,436,926,949]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="latiuncus">
|
||
<emphasis box="[150,436,926,949]" italics="true" pageId="3" pageNumber="404">Megacraspedus latiuncus</emphasis>
|
||
</taxonomicName>
|
||
(BIN: BOLD:AEK4287), is 5.93% (p-dist) (
|
||
<figureCitation box="[319,398,958,981]" captionStart="Fig" captionStartId="6.[151,182,2028,2047]" captionTargetBox="[166,1423,758,2008]" captionTargetId="figure-13@6.[151,1436,747,2016]" captionTargetPageId="6" captionText="Fig. 18. Neighbor-joining tree of Megacraspedus species recorded in Altai Mts., M. latiuncus Huemer & Karsholt, 2018, and Caryocolum tetrameris (Meyrick, 1926) as an outgroup. Source: Barcode of Life Database, cf. RATNASINGHAM & HEBERT (2007)." figureDoi="http://doi.org/10.5281/zenodo.7617236" httpUri="https://zenodo.org/record/7617236/files/figure.png" pageId="3" pageNumber="404">Fig. 18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="404" type="etymology">
|
||
<paragraph blockId="3.[150,760,190,2005]" pageId="3" pageNumber="404">
|
||
<emphasis bold="true" box="[150,278,990,1013]" pageId="3" pageNumber="404">Etymology.</emphasis>
|
||
The species name is dedicated to the excellent Ukrainian lepidopterologist Oleksiy Bidzilya (Institute for Evolutionary Ecology of the National Academy of Sciences of
|
||
<collectingCountry box="[177,264,1086,1109]" name="Ukraine" pageId="3" pageNumber="404">Ukraine</collectingCountry>
|
||
,
|
||
<collectingRegion box="[272,325,1086,1109]" country="Ukraine" name="Kyiv" pageId="3" pageNumber="404">Kyiv</collectingRegion>
|
||
,
|
||
<collectingCountry box="[334,422,1086,1109]" name="Ukraine" pageId="3" pageNumber="404">Ukraine</collectingCountry>
|
||
), who has made an exceptionally significant contribution to the knowledge of the gelechiids of Siberia including the Russian Altai Mountains.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="404" type="biology_ecology">
|
||
<paragraph blockId="3.[150,760,190,2005]" pageId="3" pageNumber="404">
|
||
<emphasis bold="true" box="[150,242,1182,1205]" pageId="3" pageNumber="404">Biology.</emphasis>
|
||
Host plant and early stages are unknown. The adults were collected at an altitude of
|
||
<quantity box="[546,626,1214,1237]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="404" unit="m" value="1400.0">1400 m</quantity>
|
||
at the end of July. The
|
||
<typeStatus box="[255,301,1246,1269]" pageId="3" pageNumber="404">type</typeStatus>
|
||
locality is extremely low-lying in the geomorphological conditions of the high parts of the Russian
|
||
<collectingRegion box="[150,206,1310,1333]" country="Russia" name="Altay" pageId="3" pageNumber="404">Altai</collectingRegion>
|
||
, surrounded by several high mountain ridges with glacier-covered peaks, difficult to reach and substantially isolated from places with comparable natural conditions (
|
||
<figureCitation box="[158,283,1406,1429]" captionStart="Figs 19–21" captionStartId="7.[150,187,2028,2047]" captionTargetBox="[150,1435,506,2013]" captionTargetId="figure-12@7.[150,1435,506,2013]" captionTargetPageId="7" captionText="Figs 19–21. Habitats of Megacraspedus species. 19–20 – M. bidzilyai sp. nov., Russia,Altai Mts. 21 – M. latiuncus Huemer & Karsholt, 2018, Kyrgyzstan, North Tian-Shan Mts. (19 – photographed by Z. Růžičková, 21 – photographed by I. Dvořák)." figureDoi="http://doi.org/10.5281/zenodo.7617238" httpUri="https://zenodo.org/record/7617238/files/figure.png" pageId="3" pageNumber="404">Figs 19–20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[150,551,1438,1461]" pageId="3" pageNumber="404" type="distribution">
|
||
<paragraph blockId="3.[150,760,190,2005]" box="[150,551,1438,1461]" pageId="3" pageNumber="404">
|
||
<emphasis bold="true" box="[150,297,1438,1461]" pageId="3" pageNumber="404">Distribution.</emphasis>
|
||
<collectingCountry box="[303,378,1438,1461]" name="Russia" pageId="3" pageNumber="404">Russia</collectingCountry>
|
||
:
|
||
<collectingRegion box="[387,547,1438,1461]" country="Russia" name="Altay" pageId="3" pageNumber="404">Altai Republic</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="404" type="discussion">
|
||
<paragraph blockId="3.[150,760,190,2005]" pageId="3" pageNumber="404">
|
||
<emphasis bold="true" box="[150,264,1470,1493]" pageId="3" pageNumber="404">Remarks.</emphasis>
|
||
Based on male genitalia characters,
|
||
<taxonomicName authority="Šumpich, 2022" authorityName="Šumpich" authorityYear="2022" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="bidzilyai" status="sp. nov.">
|
||
<emphasis italics="true" pageId="3" pageNumber="404">Megacraspedus bidzilyai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[366,451,1502,1525]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
|
||
belongs to the
|
||
<taxonomicName authorityName="Caradja" authorityYear="1920" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="majorella">
|
||
<emphasis italics="true" pageId="3" pageNumber="404">Megacraspedus majorella</emphasis>
|
||
</taxonomicName>
|
||
species group, which was established by
|
||
<bibRefCitation author="HUEMER P. & KARSHOLT O." box="[150,463,1566,1589]" pageId="3" pageNumber="404" pagination="1 - 278" refId="ref4652" refString="HUEMER P. & KARSHOLT O. 2018: Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1 - 278." type="journal article" year="2018">HUEMER & KARSHOLT (2018)</bibRefCitation>
|
||
. This group comprised only two species so far, and no female is known from either of them.
|
||
<bibRefCitation author="HUEMER P. & KARSHOLT O." box="[217,527,1630,1653]" pageId="3" pageNumber="404" pagination="1 - 278" refId="ref4652" refString="HUEMER P. & KARSHOLT O. 2018: Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1 - 278." type="journal article" year="2018">HUEMER & KARSHOLT (2018)</bibRefCitation>
|
||
point to the uncertain systematic position of this species group due to the absence of supportive molecular data. However, since molecular data have been obtained for two species of this group, an updated annotated checklist of the
|
||
<taxonomicName authorityName="Caradja" authorityYear="1920" box="[525,664,1758,1781]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="majorella">
|
||
<emphasis box="[525,664,1758,1781]" italics="true" pageId="3" pageNumber="404">M. majorella</emphasis>
|
||
</taxonomicName>
|
||
group is attached. Specific structures in the male genitalia of all three species and the close genetic relationship between
|
||
<taxonomicName authorityName="Huemer & Karsholt" authorityYear="2018" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="latiuncus">
|
||
<emphasis italics="true" pageId="3" pageNumber="404">M. latiuncus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Šumpich, 2022" authorityName="Šumpich" authorityYear="2022" box="[297,423,1854,1877]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="bidzilyai" status="sp. nov.">
|
||
<emphasis box="[297,423,1854,1877]" italics="true" pageId="3" pageNumber="404">M. bidzilyai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[428,509,1854,1877]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
|
||
confirm the correctness of establishing a separate species group. The description of the female of
|
||
<taxonomicName authority="Šumpich, 2022" authorityName="Šumpich" authorityYear="2022" box="[334,463,1918,1941]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="bidzilyai" status="sp. nov.">
|
||
<emphasis box="[334,463,1918,1941]" italics="true" pageId="3" pageNumber="404">M. bidzilyai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[472,556,1918,1941]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be considered as the initial basis for the characteristics of the females of
|
||
<taxonomicName authorityName="Caradja" authorityYear="1920" box="[150,289,1982,2005]" class="Insecta" family="Gelechiidae" genus="Megacraspedus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="majorella">
|
||
<emphasis box="[150,289,1982,2005]" italics="true" pageId="3" pageNumber="404">M. majorella</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |