503 lines
68 KiB
XML
503 lines
68 KiB
XML
<document id="209E02AB26492FFA354433601016CF8F" ID-CLB-Dataset="31502" ID-DOI="10.11646/zootaxa.4079.4.9" ID-GBIF-Dataset="3712472b-2fc6-4695-b882-ecaf18b27145" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050875" ID-ZooBank="18895002-F0ED-45EE-A01A-919A7529C6D8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510811612384" checkinUser="plazi" docAuthor="Hämäläinen, Matti" docDate="2016" docId="0386D408FFBBBA74FF18E6ACFC3D6CBA" docLanguage="en" docName="zootaxa.4079.4.9.pdf" docOrigin="Zootaxa 4079 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heliocypha vantoli Hämäläinen 2016, spec. nov." docType="treatment" docVersion="7" lastPageNumber="499" masterDocId="FFBFAC70FFBBBA71FF8FE06FFFC26D6A" masterDocTitle="Description of Heliocypha vantoli spec. nov. from Siberut in the Mentawai Islands (Odonata: Chlorocyphidae)" masterLastPageNumber="500" masterPageNumber="495" pageNumber="495" updateTime="1698494180271" updateUser="ExternalLinkService">
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<mods:titleInfo id="94DF0FE48E214ADB41597B737BEB24D2">
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<mods:title id="342EBD330FB382F7F52529445EA75F84">Description of Heliocypha vantoli spec. nov. from Siberut in the Mentawai Islands (Odonata: Chlorocyphidae)</mods:title>
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<mods:name id="B1CB7C37B478206285D6F78D299CF877" type="personal">
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<mods:roleTerm id="2E44613E20F0EFE5BA9D35FAB73D3376">Author</mods:roleTerm>
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<mods:namePart id="5BADABD8729F58311DFD45A42011B30E">Hämäläinen, Matti</mods:namePart>
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<mods:title id="241A07DA447B43A6D985489BC2547D03">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="9088EA866D87E7CD1C624BB6A82E85F6">
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<mods:date id="0ADB036F07016B3BDEAE76BB71D59A56">2016</mods:date>
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<mods:detail id="37604D626DB7046DE696B9543EF09EA9" type="pubDate">
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<mods:number id="8A3BBC94E305BCCC34115884A1618404">2016-02-15</mods:number>
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<mods:number id="D331A364B434852128D16FED3DC88852">4079</mods:number>
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<mods:number id="37BA3C9A38DF72245F472A6510E773BE">4</mods:number>
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<mods:identifier id="BC2D3B57A9BAF2A7F2E6E614EAD504DA" type="DOI">10.11646/zootaxa.4079.4.9</mods:identifier>
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<treatment id="0386D408FFBBBA74FF18E6ACFC3D6CBA" ID-DOI="http://doi.org/10.5281/zenodo.6063283" ID-GBIF-Taxon="137307408" ID-Zenodo-Dep="6063283" LSID="urn:lsid:plazi:treatment:0386D408FFBBBA74FF18E6ACFC3D6CBA" httpUri="http://treatment.plazi.org/id/0386D408FFBBBA74FF18E6ACFC3D6CBA" lastPageId="5" lastPageNumber="499" pageId="0" pageNumber="495">
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<subSubSection id="C3353695FFBBBA71FF18E6ACFD6C6BB4" box="[151,686,1731,1758]" pageId="0" pageNumber="495" type="nomenclature">
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<paragraph id="8B90651EFFBBBA71FF18E6ACFD6C6BB4" blockId="0.[151,686,1731,1758]" box="[151,686,1731,1758]" pageId="0" pageNumber="495">
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<heading id="D0D8D272FFBBBA71FF18E6ACFD6C6BB4" bold="true" box="[151,686,1731,1758]" fontSize="11" level="1" pageId="0" pageNumber="495" reason="1">
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<emphasis id="B95BB90CFFBBBA71FF18E6ACFD6C6BB4" bold="true" box="[151,686,1731,1758]" pageId="0" pageNumber="495">
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<taxonomicName id="4C2F1E9DFFBBBA71FF18E6ACFEB96BB7" ID-CoL="9X2VX" authority="Hämäläinen, 2016" authorityName="Hämäläinen" authorityYear="2016" box="[151,379,1731,1757]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="495" phylum="Arthropoda" rank="species" species="vantoli" status="sp. nov.">
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<emphasis id="B95BB90CFFBBBA71FF18E6ACFEB96BB7" bold="true" box="[151,379,1731,1757]" italics="true" pageId="0" pageNumber="495">Heliocypha vantoli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2680477FFBBBA71FE0CE6ABFE386BB4" box="[387,506,1732,1758]" pageId="0" pageNumber="495" rank="species">spec. nov.</taxonomicNameLabel>
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(
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<figureCitation id="1314799BFFBBBA71FD85E6ABFD5F6BB4" box="[522,669,1732,1758]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,769,791]" captionStartId-1="2.[151,250,1217,1239]" captionStartId-2="2.[151,250,1816,1838]" captionTargetBox-0="[340,1247,195,745]" captionTargetBox-1="[272,1304,354,1182]" captionTargetBox-2="[173,1436,1352,1792]" captionTargetId-0="figure@1.[340,1247,195,745]" captionTargetId-1="figure@2.[264,1323,337,1195]" captionTargetId-2="figure@2.[151,1436,1352,1792]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionText-0="FIGURE 1. A male individual of Heliocypha vantoli spec. nov. photographed in the type locality (see Fig. 5). Photo by Kirill A. Kolesnichenko." captionText-1="FIGURE 2. Heliocypha vantoli spec. nov. male; a) lateral view of thorax of a paratype male; b) dorsal view of thorax of a paratype male; c) wings of holotype male." captionText-2="FIGURE 3. Heliocypha vantoli spec. nov. paratype female; a) habitus; b) details of head (dorsal view) and thorax and base of abdomen (lateral view)." httpUri-0="https://zenodo.org/record/1051716/files/figure.png" httpUri-1="https://zenodo.org/record/1051718/files/figure.png" httpUri-2="https://zenodo.org/record/1051720/files/figure.png" pageId="0" pageNumber="495">Figures 1–3</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3353695FFBBBA71FF18E765FC446A06" pageId="0" pageNumber="495" type="materials_examined">
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<paragraph id="8B90651EFFBBBA71FF18E765FC446A06" blockId="0.[151,1436,1799,2038]" pageId="0" pageNumber="495">
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<materialsCitation id="3B476F43FFBBBA71FF18E765FB9C6A23" ID-GBIF-Occurrence="1700783440" collectingDate="2013-01-29" collectingDateMax="2013-01-31" collectingDateMin="2013-01-29" collectorName="K. A. Kolesnichenko" country="Indonesia" county="Material" elevation="60" latitude="-1.5833333" location="Siberut Island" longLatPrecision="1308" longitude="99.15" municipality="Mentawai Islands" pageId="0" pageNumber="495" specimenCount="1" specimenCount-male="1" stateProvince="Sumatra" typeStatus="holotype">
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<emphasis id="B95BB90CFFBBBA71FF18E765FEAA6A49" bold="true" box="[151,360,1802,1827]" pageId="0" pageNumber="495">
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<collectingCounty id="62F11D92FFBBBA71FF18E765FF3D6A49" box="[151,255,1802,1827]" pageId="0" pageNumber="495">Material</collectingCounty>
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studied:
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</emphasis>
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<typeStatus id="5494DBBCFFBBBA71FEFFE765FE1A6A49" box="[368,472,1802,1827]" pageId="0" pageNumber="495" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D29AE97FFBBBA71FE6FE764FE3A6A48" box="[480,504,1803,1826]" pageId="0" pageNumber="495" type="male">♂</specimenCount>
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:
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<collectingCountry id="F338258EFFBBBA71FD8AE765FDB76A49" box="[517,629,1802,1827]" name="Indonesia" pageId="0" pageNumber="495">Indonesia</collectingCountry>
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,
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<collectingMunicipality id="6BF4FF64FFBBBA71FD0DE765FC8B6A49" box="[642,841,1802,1827]" pageId="0" pageNumber="495">Mentawai Islands</collectingMunicipality>
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(off
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<collectingRegion id="49EBABFCFFBBBA71FC0CE764FC276A49" box="[899,997,1803,1827]" country="Indonesia" name="Sumatera Utara" pageId="0" pageNumber="495">Sumatra</collectingRegion>
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),
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<location id="8EF033C5FFBBBA71FC76E765FB5B6A49" LSID="urn:lsid:plazi:treatment:0386D408FFBBBA74FF18E6ACFC3D6CBA:8EF033C5FFBBBA71FC76E765FB5B6A49" box="[1017,1177,1802,1827]" country="Indonesia" county="Material" latitude="-1.5833333" longLatPrecision="1308" longitude="99.15" municipality="Mentawai Islands" name="Siberut Island" pageId="0" pageNumber="495" stateProvince="Sumatra">Siberut Island</location>
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,
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<location id="8EF033C5FFBBBA71FB29E765FAC76A49" LSID="urn:lsid:plazi:treatment:0386D408FFBBBA74FF18E6ACFC3D6CBA:8EF033C5FFBBBA71FB29E765FAC76A49" box="[1190,1285,1802,1827]" country="Indonesia" county="Material" latitude="-1.5833333" longLatPrecision="1308" longitude="99.15" municipality="Mentawai Islands" name="Madoba" pageId="0" pageNumber="495" stateProvince="Sumatra">Madoba</location>
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; ca.
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<geoCoordinate id="EE1B03D9FFBBBA71FAB5E764FA546A49" box="[1338,1430,1799,1827]" direction="south" orientation="latitude" pageId="0" pageNumber="495" precision="925" value="-1.5833333">1o 35’ S</geoCoordinate>
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,
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<geoCoordinate id="EE1B03D9FFBBBA71FF18E75FFEC86A22" box="[151,266,1837,1864]" direction="east" orientation="longitude" pageId="0" pageNumber="495" precision="925" value="99.15">99o 09’ E</geoCoordinate>
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; alt.
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<elevation id="0002822DFFBBBA71FEC7E75FFE716A23" box="[328,435,1840,1865]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="0" pageNumber="495" unit="m" value="60.0">
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<quantity id="4CD7C8FBFFBBBA71FEC7E75FFE476A22" box="[328,389,1840,1865]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="0" pageNumber="495" unit="m" value="60.0">60 m</quantity>
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asl
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</elevation>
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.,
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<collectingDate id="EFD5BA36FFBBBA71FE48E75FFD746A22" box="[455,694,1840,1865]" pageId="0" pageNumber="495" value="2013-01-29" valueMax="2013-01-31" valueMin="2013-01-29">
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<date id="FF9143DEFFBBBA71FE48E75FFD746A22" box="[455,694,1840,1865]" pageId="0" pageNumber="495" value="2013-01-29" valueMax="2013-01-31" valueMin="2013-01-29">29–31 January 2013</date>
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||
</collectingDate>
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||
,
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||
<collectorName id="26DA00C8FFBBBA71FD4AE75EFC6C6A23" box="[709,942,1840,1865]" pageId="0" pageNumber="495">K.A. Kolesnichenko</collectorName>
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||
leg.
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<typeStatus id="5494DBBCFFBBBA71FC62E75EFB9C6A23" box="[1005,1118,1841,1865]" pageId="0" pageNumber="495" type="paratype">Paratypes</typeStatus>
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</materialsCitation>
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||
:
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<materialsCitation id="3B476F43FFBBBA71FBE2E75FFC446A06" ID-GBIF-Occurrence="1700783491" collectingDate="2013-01-29" collectingDateMax="2013-01-31" collectingDateMin="2013-01-29" collectionCode="RMNH" collectorName="K. A. Kolesnichenko" country="Netherlands" county="Material" elevation="60" latitude="-1.5833333" location="Leiden" longLatPrecision="1308" longitude="99.15" municipality="Mentawai Islands" pageId="0" pageNumber="495" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Sumatra" typeStatus="holotype">
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||
<specimenCount id="9D29AE97FFBBBA71FBE2E75FFB596A22" box="[1133,1179,1840,1864]" pageId="0" pageNumber="495" type="male">3 ♂</specimenCount>
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<specimenCount id="9D29AE97FFBBBA71FB29E75FFB166A22" box="[1190,1236,1840,1864]" pageId="0" pageNumber="495" type="female">1 ♀</specimenCount>
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, same data as in
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<typeStatus id="5494DBBCFFBBBA71FF18E73CFF3F6A06" box="[151,253,1875,1900]" pageId="0" pageNumber="495" type="holotype">holotype</typeStatus>
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.
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<typeStatus id="5494DBBCFFBBBA71FE88E73CFEAD6A06" box="[263,367,1875,1900]" pageId="0" pageNumber="495" type="holotype">Holotype</typeStatus>
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deposited at
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<collectionCode id="ED3EFDDBFFBBBA71FD8BE73BFD9A6A01" box="[516,600,1876,1899]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="0" pageNumber="495">RMNH</collectionCode>
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||
(
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<location id="8EF033C5FFBBBA71FDE9E73CFD7A6A06" LSID="urn:lsid:plazi:treatment:0386D408FFBBBA74FF18E6ACFC3D6CBA:8EF033C5FFBBBA71FDE9E73CFD7A6A06" box="[614,696,1875,1900]" country="Netherlands" name="Leiden" pageId="0" pageNumber="495">Leiden</location>
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,
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<collectingCountry id="F338258EFFBBBA71FD4BE73CFCBD6A06" box="[708,895,1875,1900]" name="Netherlands" pageId="0" pageNumber="495">The Netherlands</collectingCountry>
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)
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3353695FFBBBA71FF48E71AFE2E6A9C" pageId="0" pageNumber="495" type="etymology">
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||
<paragraph id="8B90651EFFBBBA71FF48E71AFE2E6A9C" blockId="0.[151,1436,1799,2038]" pageId="0" pageNumber="495">
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||
<emphasis id="B95BB90CFFBBBA71FF48E71AFE8C6AE4" bold="true" box="[199,334,1909,1934]" pageId="0" pageNumber="495">Etymology.</emphasis>
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This species is named after Dr Jan van Tol to honour his significant contributions to our knowledge of the Southeast Asian odonate fauna, particularly that of
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<collectingRegion id="49EBABFCFFBBBA71FC4CE7F7FBEA6ADB" box="[963,1064,1944,1969]" country="Indonesia" name="Sulawesi Utara" pageId="0" pageNumber="495">Sulawesi</collectingRegion>
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and nearby islands, and also in appreciation of his valued leadership of the ‘DAWN (Damselfly Workers Naturalis)—group’ of the Naturalis Biodiversity Center in Leiden.
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</paragraph>
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||
</subSubSection>
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||
<caption id="DF503596FFBABA70FF18E36EFE976E5C" httpUri="https://zenodo.org/record/1051716/files/figure.png" pageId="1" pageNumber="496" targetBox="[340,1247,195,745]" targetPageId="1">
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<paragraph id="8B90651EFFBABA70FF18E36EFE976E5C" blockId="1.[151,1436,769,822]" pageId="1" pageNumber="496">
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||
<emphasis id="B95BB90CFFBABA70FF18E36EFED16E7C" bold="true" box="[151,275,769,791]" pageId="1" pageNumber="496">FIGURE 1.</emphasis>
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A male individual of
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||
<taxonomicName id="4C2F1E9DFFBABA70FE7BE36EFD746E7D" authority="Hämäläinen, 2016" authorityName="Hämäläinen" authorityYear="2016" box="[500,694,769,791]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="496" phylum="Arthropoda" rank="species" species="vantoli" status="sp. nov.">
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<emphasis id="B95BB90CFFBABA70FE7BE36EFD746E7D" box="[500,694,769,791]" italics="true" pageId="1" pageNumber="496">Heliocypha vantoli</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A2680477FFBABA70FD33E36DFCE16E7D" box="[700,803,770,791]" pageId="1" pageNumber="496" rank="species">
|
||
<emphasis id="B95BB90CFFBABA70FD33E36DFCE16E7D" bold="true" box="[700,803,770,791]" pageId="1" pageNumber="496">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
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photographed in the type locality (see Fig. 5). Photo by Kirill A. Kolesnichenko.
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||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3353695FFBABA70FF48E332FC346B79" pageId="1" pageNumber="496" type="materials_examined">
|
||
<paragraph id="8B90651EFFBABA70FF48E332FD916EF3" blockId="1.[151,1437,861,2024]" pageId="1" pageNumber="496">
|
||
<emphasis id="B95BB90CFFBABA70FF48E332FEB16E1C" bold="true" box="[199,371,861,886]" pageId="1" pageNumber="496">
|
||
<typeStatus id="5494DBBCFFBABA70FF48E332FEF66E1C" box="[199,308,861,886]" pageId="1" pageNumber="496" type="holotype">Holotype</typeStatus>
|
||
male
|
||
</emphasis>
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||
. Head. Black throughout with small, oblong, ochreous spot beside each lateral ocellus and with tiny round dot on either side of occiput.
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||
</paragraph>
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||
<paragraph id="8B90651EFFBABA70FF48E3CCFAD169BB" blockId="1.[151,1437,861,2024]" pageId="1" pageNumber="496">
|
||
Thorax (cf.
|
||
<figureCitation id="1314799BFFBABA70FEDDE3CCFE116ED6" box="[338,467,931,956]" captionStart="FIGURE 2" captionStartId="2.[151,250,1217,1239]" captionTargetBox="[272,1304,354,1182]" captionTargetId="figure@2.[264,1323,337,1195]" captionTargetPageId="2" captionText="FIGURE 2. Heliocypha vantoli spec. nov. male; a) lateral view of thorax of a paratype male; b) dorsal view of thorax of a paratype male; c) wings of holotype male." httpUri="https://zenodo.org/record/1051718/files/figure.png" pageId="1" pageNumber="496">Fig. 2a, 2b</figureCitation>
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). Prothorax black with distinct pinkish magenta anterodorsal marking on hind lobe. Synthorax black with pinkish magenta dorsal triangle and with broad, pinkish magenta, irregularly shaped antehumeral stripe on mesepisternum. This stripe broadest in anterior half and narrowing towards wing base, bending ventrad along antealar ridge where it is narrowly tinged with pale blue. Small yellowish marking on mesostigmal laminae, not connected with antehumeral stripe. Irregularly shaped, pale cerulean lateral stripe on metepisternum, connected to similarly coloured marking on anterior edge of mesepimeron; this lateral stripe slightly narrower than antehumeral stripe. Small pale cerulean marking on metepimeron near wing base. Venter of thorax black with posterior and lower half of lateral borders of poststernum and poststernal membranes pale. Legs black, inner surfaces of middle and hind femora creamy yellow; those of middle and hind tibiae white.
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||
</paragraph>
|
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<paragraph id="8B90651EFFBABA70FF48E4B4FAC568EE" blockId="1.[151,1437,861,2024]" pageId="1" pageNumber="496">
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Wings. Shaped as in
|
||
<figureCitation id="1314799BFFBABA70FE3DE4B4FDC06999" box="[434,514,1243,1268]" captionStart="FIGURE 2" captionStartId="2.[151,250,1217,1239]" captionTargetBox="[272,1304,354,1182]" captionTargetId="figure@2.[264,1323,337,1195]" captionTargetPageId="2" captionText="FIGURE 2. Heliocypha vantoli spec. nov. male; a) lateral view of thorax of a paratype male; b) dorsal view of thorax of a paratype male; c) wings of holotype male." httpUri="https://zenodo.org/record/1051718/files/figure.png" pageId="1" pageNumber="496">Fig. 2c</figureCitation>
|
||
, wing apices distinctly rounded, especially in hindwing. Opaque area extensive in both wings, starting anterior to nodus. Hind wing with seven vitreous fenestrae (“windows”) arranged as in
|
||
<figureCitation id="1314799BFFBABA70FAC7E490FA5A687D" box="[1352,1432,1279,1304]" captionStart="FIGURE 2" captionStartId="2.[151,250,1217,1239]" captionTargetBox="[272,1304,354,1182]" captionTargetId="figure@2.[264,1323,337,1195]" captionTargetPageId="2" captionText="FIGURE 2. Heliocypha vantoli spec. nov. male; a) lateral view of thorax of a paratype male; b) dorsal view of thorax of a paratype male; c) wings of holotype male." httpUri="https://zenodo.org/record/1051718/files/figure.png" pageId="1" pageNumber="496">Fig. 2c</figureCitation>
|
||
. Basal fenestra long, covering 18 cells and extending to hyaline area at wing base. Lowest fenestra in middle of hindwing very long, covering 20 cells. At certain angles, dark background on either side of both wings reflects green, magenta or violet iridescence, especially striking in hindwing. Pterostigma dark in both wings.
|
||
</paragraph>
|
||
<paragraph id="8B90651EFFBABA70FF48E5E0FE36689A" blockId="1.[151,1437,861,2024]" pageId="1" pageNumber="496">Abdomen. Black, except with a small, mid-lateral pale blue spots near apices of S1, S2, S3 and S4. Spots on S1 and S2 a little larger than the others. Tiny remnant of pale ventrolateral stripe on S1 and S2. Appendages black, of the typical shape for the genus.</paragraph>
|
||
<paragraph id="8B90651EFFBABA70FF48E593FC346B79" blockId="1.[151,1437,861,2024]" box="[199,1014,1531,1556]" pageId="1" pageNumber="496">Measurements (in mm). Hindwing 21.5; abdomen (incl. appendages) 19.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3353695FFBABA74FF48E670FC3D6CBA" lastPageId="5" lastPageNumber="500" pageId="1" pageNumber="496" type="description">
|
||
<paragraph id="8B90651EFFBABA70FF48E670FE926BC9" blockId="1.[151,1437,861,2024]" pageId="1" pageNumber="496">
|
||
<emphasis id="B95BB90CFFBABA70FF48E670FDDC6B52" bold="true" box="[199,542,1567,1592]" pageId="1" pageNumber="496">
|
||
Variation in
|
||
<typeStatus id="5494DBBCFFBABA70FED0E670FE096B52" box="[351,459,1567,1592]" pageId="1" pageNumber="496" type="paratype">paratype</typeStatus>
|
||
males.
|
||
</emphasis>
|
||
The wing colour pattern is quite similar in all specimens. There is some small variation in the pale markings of thorax and abdomen, obviously partly age dependant. In two males the pale marking on the metepimeron is a little larger and longer than in
|
||
<typeStatus id="5494DBBCFFBABA70FCFBE608FC146BEA" box="[884,982,1639,1664]" pageId="1" pageNumber="496" type="holotype">holotype</typeStatus>
|
||
and in two males abdominal S4 lacks a lateral pale spot.
|
||
</paragraph>
|
||
<paragraph id="8B90651EFFBABA70FF11E6DFFBF46BAD" blockId="1.[151,1437,861,2024]" box="[158,1078,1711,1736]" pageId="1" pageNumber="496">Measurements (in mm). Hindwing 21.5–22.5; abdomen (incl. appendages) 19–20.</paragraph>
|
||
<paragraph id="8B90651EFFBABA70FF48E6BCFB656A5E" blockId="1.[151,1437,861,2024]" pageId="1" pageNumber="496">
|
||
<emphasis id="B95BB90CFFBABA70FF48E6BCFEDC6B86" bold="true" box="[199,286,1747,1772]" pageId="1" pageNumber="496">Female</emphasis>
|
||
. Head with orange markings much more extensive than in male. Middle and lateral lobes of labium pale at basal one third. Labrum with a pair of orange dots. Borders of eye narrowly orange up to level of lateral ocelli. Yellow markings on dorsal side of head as in
|
||
<figureCitation id="1314799BFFBABA70FD6EE774FCF16A5E" box="[737,819,1819,1844]" captionStart="FIGURE 3" captionStartId="2.[151,250,1816,1838]" captionTargetBox="[173,1436,1352,1792]" captionTargetId="figure@2.[151,1436,1352,1792]" captionTargetPageId="2" captionText="FIGURE 3. Heliocypha vantoli spec. nov. paratype female; a) habitus; b) details of head (dorsal view) and thorax and base of abdomen (lateral view)." httpUri="https://zenodo.org/record/1051720/files/figure.png" pageId="1" pageNumber="496">Fig. 3b</figureCitation>
|
||
. Venter of head partly pruinosed.
|
||
</paragraph>
|
||
<paragraph id="8B90651EFFBABA73FF48E750FD596C77" blockId="1.[151,1437,861,2024]" lastBlockId="2.[151,1436,151,285]" lastPageId="2" lastPageNumber="497" pageId="1" pageNumber="496">
|
||
Thorax (
|
||
<figureCitation id="1314799BFFBABA70FEA4E750FE406A32" box="[299,386,1855,1880]" captionStart="FIGURE 3" captionStartId="2.[151,250,1816,1838]" captionTargetBox="[173,1436,1352,1792]" captionTargetId="figure@2.[151,1436,1352,1792]" captionTargetPageId="2" captionText="FIGURE 3. Heliocypha vantoli spec. nov. paratype female; a) habitus; b) details of head (dorsal view) and thorax and base of abdomen (lateral view)." httpUri="https://zenodo.org/record/1051720/files/figure.png" pageId="1" pageNumber="496">Fig. 3b</figureCitation>
|
||
). Prothorax black with small pale/ochreous spots on sides of fore, middle and hind lobes. Distinct triangular posterodorsal marking on hind lobe connected with narrow orange stripe, which reaches anterior edge of hindlobe. Synthorax black with narrow, reddish brown antehumeral stripe on mesepisternum; much narrower than in male. This stripe narrowing towards wing base and turning ventrad along antealar ridge. Small pale marking on mesostigmal laminae, not connected with antehumeral stripe. Irregularly shaped, orange ochre lateral stripe on metepisternum, connected to similarly coloured marking on anterior edge of mesepimeron; this stripe resembling that in male, but a little broader. Elongated, slightly angulate, pale patch in posterior third of metepimeron near wing base. Venter of thorax coloured as in male. Legs black, trochanters and inner surface of middle and hind femora partly greyish brown.
|
||
</paragraph>
|
||
<caption id="DF503596FFB9BA73FF18E4AEFD80699F" httpUri="https://zenodo.org/record/1051718/files/figure.png" pageId="2" pageNumber="497" targetBox="[272,1304,354,1182]" targetPageId="2">
|
||
<paragraph id="8B90651EFFB9BA73FF18E4AEFD80699F" blockId="2.[151,1435,1217,1269]" pageId="2" pageNumber="497">
|
||
<emphasis id="B95BB90CFFB9BA73FF18E4AEFED669BC" bold="true" box="[151,276,1217,1239]" pageId="2" pageNumber="497">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C2F1E9DFFB9BA73FE92E4AEFE2269BD" authority="Hämäläinen, 2016" authorityName="Hämäläinen" authorityYear="2016" box="[285,480,1217,1239]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="497" phylum="Arthropoda" rank="species" species="vantoli" status="sp. nov.">
|
||
<emphasis id="B95BB90CFFB9BA73FE92E4AEFE2269BD" box="[285,480,1217,1239]" italics="true" pageId="2" pageNumber="497">Heliocypha vantoli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2680477FFB9BA73FE66E4AEFD9369BC" box="[489,593,1217,1238]" pageId="2" pageNumber="497" rank="species">
|
||
<emphasis id="B95BB90CFFB9BA73FE66E4AEFD9369BC" bold="true" box="[489,593,1217,1238]" pageId="2" pageNumber="497">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
male; a) lateral view of thorax of a paratype male; b) dorsal view of thorax of a paratype male; c) wings of holotype male.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF503596FFB9BA73FF18E777FE4B6A27" httpUri="https://zenodo.org/record/1051720/files/figure.png" pageId="2" pageNumber="497" targetBox="[173,1436,1352,1792]" targetPageId="2">
|
||
<paragraph id="8B90651EFFB9BA73FF18E777FE4B6A27" blockId="2.[151,1435,1816,1869]" pageId="2" pageNumber="497">
|
||
<emphasis id="B95BB90CFFB9BA73FF18E777FED16A47" bold="true" box="[151,275,1816,1838]" pageId="2" pageNumber="497">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C2F1E9DFFB9BA73FE95E777FE196A44" authority="Hämäläinen, 2016" authorityName="Hämäläinen" authorityYear="2016" box="[282,475,1816,1838]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="497" phylum="Arthropoda" rank="species" species="vantoli" status="sp. nov.">
|
||
<emphasis id="B95BB90CFFB9BA73FE95E777FE196A44" box="[282,475,1816,1838]" italics="true" pageId="2" pageNumber="497">Heliocypha vantoli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2680477FFB9BA73FE6DE776FD8A6A44" box="[482,584,1817,1838]" pageId="2" pageNumber="497" rank="species">
|
||
<emphasis id="B95BB90CFFB9BA73FE6DE776FD8A6A44" bold="true" box="[482,584,1817,1838]" pageId="2" pageNumber="497">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
paratype female; a) habitus; b) details of head (dorsal view) and thorax and base of abdomen (lateral view).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B90651EFFB9BA73FF48E71BFDFD6ABF" blockId="2.[151,1436,1908,2005]" pageId="2" pageNumber="497">
|
||
Wings. Both wings deeply suffused with brown (
|
||
<figureCitation id="1314799BFFB9BA73FD67E71BFCFB6AE7" box="[744,825,1908,1933]" captionStart="FIGURE 3" captionStartId="2.[151,250,1816,1838]" captionTargetBox="[173,1436,1352,1792]" captionTargetId="figure@2.[151,1436,1352,1792]" captionTargetPageId="2" captionText="FIGURE 3. Heliocypha vantoli spec. nov. paratype female; a) habitus; b) details of head (dorsal view) and thorax and base of abdomen (lateral view)." httpUri="https://zenodo.org/record/1051720/files/figure.png" pageId="2" pageNumber="497">Fig. 3a</figureCitation>
|
||
), especially strongly so in apical third and along costal margins distad from level of nodus in forewing and slightly further basad in hindwing. Pterostigma greyish white with anterior corner obscurely brown.
|
||
</paragraph>
|
||
<paragraph id="8B90651EFFB8BA72FF48E0F8FC0B6C2A" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="498">
|
||
Abdomen. Largely black with pale markings as follows (
|
||
<figureCitation id="1314799BFFB8BA72FCEDE0F8FC1C6DDA" box="[866,990,151,176]" captionStart="FIGURE 3" captionStartId="2.[151,250,1816,1838]" captionTargetBox="[173,1436,1352,1792]" captionTargetId="figure@2.[151,1436,1352,1792]" captionTargetPageId="2" captionText="FIGURE 3. Heliocypha vantoli spec. nov. paratype female; a) habitus; b) details of head (dorsal view) and thorax and base of abdomen (lateral view)." httpUri="https://zenodo.org/record/1051720/files/figure.png" pageId="3" pageNumber="498">Fig 3a, 3b</figureCitation>
|
||
). S1 with triangular marking, directed ventrad. S2, S3 and S4 with short mid-lateral stripe in apical half of each segment, with small round mid-lateral spot near apex and obscure short ventrolateral stipe in middle of segment. Pale markings on S4 tiny, almost obsolete. Dorsal carina of S3–S7 narrowly yellow, yellow colour not reaching either base or apex of each segments. Intersegmental rings, except the three most apical ones, narrowly yellow.
|
||
</paragraph>
|
||
<paragraph id="8B90651EFFB8BA72FF48E122FC346C0E" blockId="3.[151,1437,151,789]" box="[199,1014,332,357]" pageId="3" pageNumber="498">Measurements (in mm). Hindwing 24.5; abdomen (incl. appendages) 18.</paragraph>
|
||
<paragraph id="8B90651EFFB8BA72FF48E100FD8B6FEF" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="498">
|
||
<emphasis id="B95BB90CFFB8BA72FF48E100FDC16CE2" bold="true" box="[199,515,367,392]" pageId="3" pageNumber="498">Distinguishing characters.</emphasis>
|
||
Apart from this new species, only other two Sumatran chlorocyphid taxa have vitreous fenestrae in the partly opaque male hindwings. These are
|
||
<emphasis id="B95BB90CFFB8BA72FC0AE1FBFB466CC6" box="[901,1156,404,428]" italics="true" pageId="3" pageNumber="498">
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FC0AE1FBFBC46CC6" authorityName="Hämäläinen" authorityYear="2016" box="[901,1030,404,428]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="genus">Heliocypha</taxonomicName>
|
||
a.
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FBA4E1FAFB466CC6" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1853" box="[1067,1156,405,428]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="angusta">angusta</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FB33E1FAFA566CC6" baseAuthorityName="Lieftinck" baseAuthorityYear="1947" box="[1212,1428,405,428]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="angusta" subSpecies="oceanis">
|
||
<emphasis id="B95BB90CFFB8BA72FB33E1FAFA566CC6" box="[1212,1428,405,428]" italics="true" pageId="3" pageNumber="498">H. angusta oceanis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FF18E1D7FEAF6CBA" authority="Hämäläinen, 2016" authorityName="Hämäläinen" authorityYear="2016" box="[151,365,440,464]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="vantoli" status="sp. nov.">
|
||
<emphasis id="B95BB90CFFB8BA72FF18E1D7FEAF6CBA" box="[151,365,440,464]" italics="true" pageId="3" pageNumber="498">Heliocypha vantoli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2680477FFB8BA72FEFBE1D7FE266CA5" box="[372,484,440,463]" pageId="3" pageNumber="498" rank="species">spec. nov.</taxonomicNameLabel>
|
||
is easy to separate from these two by the shape of the wings. In
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FB37E1D6FAEA6CBA" authorityName="Hämäläinen" authorityYear="2016" box="[1208,1320,440,464]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="vantoli">
|
||
<emphasis id="B95BB90CFFB8BA72FB37E1D6FAEA6CBA" box="[1208,1320,440,464]" italics="true" pageId="3" pageNumber="498">H. vantoli</emphasis>
|
||
</taxonomicName>
|
||
the wings (
|
||
<figureCitation id="1314799BFFB8BA72FF10E1B3FF326C9E" box="[159,240,476,501]" captionStart="FIGURE 2" captionStartId="2.[151,250,1217,1239]" captionTargetBox="[272,1304,354,1182]" captionTargetId="figure@2.[264,1323,337,1195]" captionTargetPageId="2" captionText="FIGURE 2. Heliocypha vantoli spec. nov. male; a) lateral view of thorax of a paratype male; b) dorsal view of thorax of a paratype male; c) wings of holotype male." httpUri="https://zenodo.org/record/1051718/files/figure.png" pageId="3" pageNumber="498">Fig. 2c</figureCitation>
|
||
) are proportionally broader and shorter than in
|
||
<emphasis id="B95BB90CFFB8BA72FC9EE1B2FC696C9E" box="[785,939,477,500]" italics="true" pageId="3" pageNumber="498">
|
||
H. a.
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FCDDE1B2FC696C9E" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1853" box="[850,939,477,500]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="angusta">angusta</taxonomicName>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1314799BFFB8BA72FC33E1B3FBCF6C9E" box="[956,1037,476,501]" captionStart="FIGURE 4" captionStartId="3.[151,250,1659,1681]" captionTargetBox="[166,1405,863,1630]" captionTargetId="figure@3.[151,1436,844,1636]" captionTargetPageId="3" captionText="FIGURE 4. Heliocypha a. angusta and H. angusta oceanis males; a) wings of a H. a. angusta specimen from West Sumatra province, Sg. Jaring, between Bukittingi and Maninjau, 25 - v- 1997, M. Hämäläinen leg.; b) wings of holotype of H. angusta oceanis from Enggano Island; c) thorax of a H. a. angusta specimen from South Sumatra; d) thorax of a paratype of H. angusta oceanis from Enggano Island. Figures c and d are scanned from Lieftinck (1947, Figs. 2 and 4 on p. 221)." httpUri="https://zenodo.org/record/1051722/files/figure.png" pageId="3" pageNumber="498">Fig. 4a</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FBC1E1B2FAEB6C9E" baseAuthorityName="Lieftinck" baseAuthorityYear="1947" box="[1102,1321,477,500]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="angusta" subSpecies="oceanis">
|
||
<emphasis id="B95BB90CFFB8BA72FBC1E1B2FAEB6C9E" box="[1102,1321,477,500]" italics="true" pageId="3" pageNumber="498">H. angusta oceanis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1314799BFFB8BA72FAB5E1B3FA4D6C9F" box="[1338,1423,476,501]" captionStart="FIGURE 4" captionStartId="3.[151,250,1659,1681]" captionTargetBox="[166,1405,863,1630]" captionTargetId="figure@3.[151,1436,844,1636]" captionTargetPageId="3" captionText="FIGURE 4. Heliocypha a. angusta and H. angusta oceanis males; a) wings of a H. a. angusta specimen from West Sumatra province, Sg. Jaring, between Bukittingi and Maninjau, 25 - v- 1997, M. Hämäläinen leg.; b) wings of holotype of H. angusta oceanis from Enggano Island; c) thorax of a H. a. angusta specimen from South Sumatra; d) thorax of a paratype of H. angusta oceanis from Enggano Island. Figures c and d are scanned from Lieftinck (1947, Figs. 2 and 4 on p. 221)." httpUri="https://zenodo.org/record/1051722/files/figure.png" pageId="3" pageNumber="498">Fig. 4b</figureCitation>
|
||
). Moreover, the wing tips are distinctly more rounded in
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FC8FE26EFCB26F72" authorityName="Hämäläinen" authorityYear="2016" box="[768,880,512,536]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="vantoli">
|
||
<emphasis id="B95BB90CFFB8BA72FC8FE26EFCB26F72" box="[768,880,512,536]" italics="true" pageId="3" pageNumber="498">H. vantoli</emphasis>
|
||
</taxonomicName>
|
||
. The overall arrangement of the fenestrae is quite similar in these three taxa, but
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FD8AE24AFDBB6F56" authorityName="Hämäläinen" authorityYear="2016" box="[517,633,548,572]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="vantoli">
|
||
<emphasis id="B95BB90CFFB8BA72FD8AE24AFDBB6F56" box="[517,633,548,572]" italics="true" pageId="3" pageNumber="498">H. vantoli</emphasis>
|
||
</taxonomicName>
|
||
differs by having a much longer row of iridescent cells in the basal fenestra in the central part of the hindwing. In
|
||
<emphasis id="B95BB90CFFB8BA72FD13E226FCD06F0A" box="[668,786,584,608]" italics="true" pageId="3" pageNumber="498">
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FD13E226FCCC6F0A" authorityName="Hämäläinen" authorityYear="2016" box="[668,782,584,608]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="vantoli">H. vantoli</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the extent of opaque area in wings is much greater than in
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FF18E202FED06FEE" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1853" box="[151,274,621,644]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="angusta">
|
||
<emphasis id="B95BB90CFFB8BA72FF18E202FED06FEE" box="[151,274,621,644]" italics="true" pageId="3" pageNumber="498">H. angusta</emphasis>
|
||
</taxonomicName>
|
||
, especially in the forewing.
|
||
</paragraph>
|
||
<paragraph id="8B90651EFFB8BA72FF48E2E0FADE6E7F" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="498">
|
||
The colour pattern of the male thorax also differs. In
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FCBDE2FEFC676FC2" authorityName="Hämäläinen" authorityYear="2016" box="[818,933,656,680]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="vantoli">
|
||
<emphasis id="B95BB90CFFB8BA72FCBDE2FEFC676FC2" box="[818,933,656,680]" italics="true" pageId="3" pageNumber="498">H. vantoli</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1314799BFFB8BA72FC39E2E0FBC46FC2" box="[950,1030,655,680]" captionStart="FIGURE 1" captionStartId="1.[151,250,769,791]" captionTargetBox="[340,1247,195,745]" captionTargetId="figure@1.[340,1247,195,745]" captionTargetPageId="1" captionText="FIGURE 1. A male individual of Heliocypha vantoli spec. nov. photographed in the type locality (see Fig. 5). Photo by Kirill A. Kolesnichenko." httpUri="https://zenodo.org/record/1051716/files/figure.png" pageId="3" pageNumber="498">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="1314799BFFB8BA72FB98E2FFFB9D6FC2" box="[1047,1119,655,680]" captionStart="FIGURE 2" captionStartId="2.[151,250,1217,1239]" captionTargetBox="[272,1304,354,1182]" captionTargetId="figure@2.[264,1323,337,1195]" captionTargetPageId="2" captionText="FIGURE 2. Heliocypha vantoli spec. nov. male; a) lateral view of thorax of a paratype male; b) dorsal view of thorax of a paratype male; c) wings of holotype male." httpUri="https://zenodo.org/record/1051718/files/figure.png" pageId="3" pageNumber="498">2a, 2b</figureCitation>
|
||
) the pale markings are less extensive than in
|
||
<emphasis id="B95BB90CFFB8BA72FEEAE2DAFDC06FA6" box="[357,514,693,716]" italics="true" pageId="3" pageNumber="498">
|
||
H. a.
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FE26E2DAFDC06FA6" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1853" box="[425,514,693,716]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="angusta">angusta</taxonomicName>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1314799BFFB8BA72FD9AE2DBFDA86FA6" box="[533,618,692,717]" captionStart="FIGURE 4" captionStartId="3.[151,250,1659,1681]" captionTargetBox="[166,1405,863,1630]" captionTargetId="figure@3.[151,1436,844,1636]" captionTargetPageId="3" captionText="FIGURE 4. Heliocypha a. angusta and H. angusta oceanis males; a) wings of a H. a. angusta specimen from West Sumatra province, Sg. Jaring, between Bukittingi and Maninjau, 25 - v- 1997, M. Hämäläinen leg.; b) wings of holotype of H. angusta oceanis from Enggano Island; c) thorax of a H. a. angusta specimen from South Sumatra; d) thorax of a paratype of H. angusta oceanis from Enggano Island. Figures c and d are scanned from Lieftinck (1947, Figs. 2 and 4 on p. 221)." httpUri="https://zenodo.org/record/1051722/files/figure.png" pageId="3" pageNumber="498">Fig. 4c</figureCitation>
|
||
).
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FD0EE2DAFC9C6FA6" baseAuthorityName="Lieftinck" baseAuthorityYear="1947" box="[641,862,693,716]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="angusta" subSpecies="oceanis">
|
||
<emphasis id="B95BB90CFFB8BA72FD0EE2DAFC9C6FA6" box="[641,862,693,716]" italics="true" pageId="3" pageNumber="498">H. angusta oceanis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1314799BFFB8BA72FCFEE2DBFC046FA7" box="[881,966,692,717]" captionStart="FIGURE 4" captionStartId="3.[151,250,1659,1681]" captionTargetBox="[166,1405,863,1630]" captionTargetId="figure@3.[151,1436,844,1636]" captionTargetPageId="3" captionText="FIGURE 4. Heliocypha a. angusta and H. angusta oceanis males; a) wings of a H. a. angusta specimen from West Sumatra province, Sg. Jaring, between Bukittingi and Maninjau, 25 - v- 1997, M. Hämäläinen leg.; b) wings of holotype of H. angusta oceanis from Enggano Island; c) thorax of a H. a. angusta specimen from South Sumatra; d) thorax of a paratype of H. angusta oceanis from Enggano Island. Figures c and d are scanned from Lieftinck (1947, Figs. 2 and 4 on p. 221)." httpUri="https://zenodo.org/record/1051722/files/figure.png" pageId="3" pageNumber="498">Fig. 4d</figureCitation>
|
||
) differs from both
|
||
<emphasis id="B95BB90CFFB8BA72FB28E2DAFA866FA6" box="[1191,1348,693,716]" italics="true" pageId="3" pageNumber="498">
|
||
H. a.
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FB64E2DAFA866FA6" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1853" box="[1259,1348,693,716]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="angusta">angusta</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B95BB90CFFB8BA72FA0EE2DAFEC86F9A" italics="true" pageId="3" pageNumber="498">
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FA0EE2DAFF276F9A" authorityName="Hämäläinen" authorityYear="2016" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="vantoli">H. vantoli</taxonomicName>
|
||
by
|
||
</emphasis>
|
||
having very restricted pale lateral markings. The female of
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FC46E2B6FBFE6F9A" authorityName="Hämäläinen" authorityYear="2016" box="[969,1084,728,752]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="vantoli">
|
||
<emphasis id="B95BB90CFFB8BA72FC46E2B6FBFE6F9A" box="[969,1084,728,752]" italics="true" pageId="3" pageNumber="498">H. vantoli</emphasis>
|
||
</taxonomicName>
|
||
is readily distinguished by its strongly brown tinted wings; in females of
|
||
<emphasis id="B95BB90CFFB8BA72FDF5E292FCD36E7E" box="[634,785,765,788]" italics="true" pageId="3" pageNumber="498">
|
||
H. a.
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FD37E292FCD36E7E" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1853" box="[696,785,765,788]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="angusta">angusta</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FCC8E292FBDC6E7E" baseAuthorityName="Lieftinck" baseAuthorityYear="1947" box="[839,1054,765,788]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="angusta" subSpecies="oceanis">
|
||
<emphasis id="B95BB90CFFB8BA72FCC8E292FBDC6E7E" box="[839,1054,765,788]" italics="true" pageId="3" pageNumber="498">H. angusta oceanis</emphasis>
|
||
</taxonomicName>
|
||
the wings are hyaline.
|
||
</paragraph>
|
||
<caption id="DF503596FFB8BA72FF18E614FB076B87" httpUri="https://zenodo.org/record/1051722/files/figure.png" pageId="3" pageNumber="498" targetBox="[166,1405,863,1630]" targetPageId="3">
|
||
<paragraph id="8B90651EFFB8BA72FF18E614FB076B87" blockId="3.[151,1436,1659,1774]" pageId="3" pageNumber="498">
|
||
<emphasis id="B95BB90CFFB8BA72FF18E614FED66BFA" bold="true" box="[151,276,1659,1681]" pageId="3" pageNumber="498">FIGURE 4.</emphasis>
|
||
<emphasis id="B95BB90CFFB8BA72FE94E614FDC06BFB" box="[283,514,1659,1681]" italics="true" pageId="3" pageNumber="498">
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FE94E614FE4D6BFB" authorityName="Hämäläinen" authorityYear="2016" box="[283,399,1659,1681]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="genus">Heliocypha</taxonomicName>
|
||
a.
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FE3DE613FDC06BFB" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1853" box="[434,514,1660,1681]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="angusta">angusta</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FDB9E613FD396BFB" baseAuthorityName="Lieftinck" baseAuthorityYear="1947" box="[566,763,1660,1681]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="angusta" subSpecies="oceanis">
|
||
<emphasis id="B95BB90CFFB8BA72FDB9E613FD396BFB" box="[566,763,1660,1681]" italics="true" pageId="3" pageNumber="498">H. angusta oceanis</emphasis>
|
||
</taxonomicName>
|
||
males; a) wings of a
|
||
<emphasis id="B95BB90CFFB8BA72FC53E613FBA46BFB" box="[988,1126,1660,1681]" italics="true" pageId="3" pageNumber="498">
|
||
H. a.
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FB99E613FBA46BFB" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1853" box="[1046,1126,1660,1681]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="angusta">angusta</taxonomicName>
|
||
</emphasis>
|
||
specimen from West Sumatra province, Sg. Jaring, between Bukittingi and Maninjau, 25-v-1997, M. Hämäläinen leg.; b) wings of holotype of
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FAA4E6F4FF266BA5" baseAuthorityName="Lieftinck" baseAuthorityYear="1947" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="angusta" subSpecies="oceanis">
|
||
<emphasis id="B95BB90CFFB8BA72FAA4E6F4FF266BA5" italics="true" pageId="3" pageNumber="498">H. angusta oceanis</emphasis>
|
||
</taxonomicName>
|
||
from Enggano Island; c) thorax of a
|
||
<emphasis id="B95BB90CFFB8BA72FDD6E6D5FD236BA5" box="[601,737,1722,1743]" italics="true" pageId="3" pageNumber="498">
|
||
H. a.
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FD1EE6D5FD236BA5" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1853" box="[657,737,1722,1743]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="angusta">angusta</taxonomicName>
|
||
</emphasis>
|
||
specimen from South Sumatra; d) thorax of a paratype of
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FAA2E6D5FF266B84" baseAuthorityName="Lieftinck" baseAuthorityYear="1947" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="angusta" subSpecies="oceanis">
|
||
<emphasis id="B95BB90CFFB8BA72FAA2E6D5FF266B84" italics="true" pageId="3" pageNumber="498">H. angusta oceanis</emphasis>
|
||
</taxonomicName>
|
||
from Enggano Island. Figures c and d are scanned from Lieftinck (1947, Figs. 2 and 4 on p. 221).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B90651EFFB8BA75FF48E779FC5A6C76" blockId="3.[151,1436,1812,2018]" lastBlockId="4.[151,1436,151,357]" lastPageId="4" lastPageNumber="499" pageId="3" pageNumber="498">
|
||
<emphasis id="B95BB90CFFB8BA72FF48E779FEA46A47" box="[199,358,1814,1837]" italics="true" pageId="3" pageNumber="498">
|
||
H. a.
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FE82E779FEA46A47" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1853" box="[269,358,1814,1837]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="angusta">angusta</taxonomicName>
|
||
</emphasis>
|
||
is common and widespread in
|
||
<collectingRegion id="49EBABFCFFB8BA72FD57E77AFCF76A47" box="[728,821,1813,1837]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="498">Sumatra</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFBE18EFFFB8BA72FCC8E77BFC356A47" author="Lieftinck" box="[839,1015,1812,1837]" pageId="3" pageNumber="498" refString="Lieftinck, M. A. (1935) A synopsis of the Odonata (dragonflies) of Sumatra. Miscellanea zoologica sumatrana, 92 - 93, 1 - 23." type="book chapter" year="1935">Lieftinck 1935</bibRefCitation>
|
||
). The
|
||
<typeStatus id="5494DBBCFFB8BA72FBCBE779FBB66A44" box="[1092,1140,1814,1838]" pageId="3" pageNumber="498">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FA8FE779FABD6A47" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1853" box="[1280,1407,1814,1837]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="angusta">
|
||
<emphasis id="B95BB90CFFB8BA72FA8FE779FABD6A47" box="[1280,1407,1814,1837]" italics="true" pageId="3" pageNumber="498">H. angusta</emphasis>
|
||
</taxonomicName>
|
||
is Bencoolen [
|
||
<collectingRegion id="49EBABFCFFB8BA72FEADE756FE566A38" box="[290,404,1849,1874]" country="Indonesia" name="Bengkulu" pageId="3" pageNumber="498">Bengkulu</collectingRegion>
|
||
] at the west coast of southern
|
||
<collectingRegion id="49EBABFCFFB8BA72FC9CE756FCB16A3B" box="[787,883,1849,1873]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="498">Sumatra</collectingRegion>
|
||
, where the Danish naval officer I.K. Daldorff collected the teneral male
|
||
<typeStatus id="5494DBBCFFB8BA72FE41E732FDF26A1C" box="[462,560,1885,1910]" pageId="3" pageNumber="498" type="holotype">holotype</typeStatus>
|
||
in 1800. The specimen is still in good condition and preserved in ZMUC (Copenhagen).
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FEC9E7EDFDE06AF3" baseAuthorityName="Lieftinck" baseAuthorityYear="1947" box="[326,546,1922,1945]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="angusta" subSpecies="oceanis">
|
||
<emphasis id="B95BB90CFFB8BA72FEC9E7EDFDE06AF3" box="[326,546,1922,1945]" italics="true" pageId="3" pageNumber="498">H. angusta oceanis</emphasis>
|
||
</taxonomicName>
|
||
is endemic to Enggano. The
|
||
<typeStatus id="5494DBBCFFB8BA72FCF7E7EEFC186AF0" box="[888,986,1921,1946]" pageId="3" pageNumber="498" type="holotype">holotype</typeStatus>
|
||
male and a long series of
|
||
<typeStatus id="5494DBBCFFB8BA72FA9FE7EDFAB96AF0" box="[1296,1403,1922,1946]" pageId="3" pageNumber="498" type="paratype">paratypes</typeStatus>
|
||
of both sexes, collected by J.K. de Jong in 1936, are at RMNH (Leiden).
|
||
<bibRefCitation id="EFBE18EFFFB8BA72FC38E7CAFBA86AD7" author="Lieftinck" box="[951,1130,1957,1982]" pageId="3" pageNumber="498" refString="Lieftinck, M. A. (1947) Two interesting new insular Rhinocypha from Malaysia (Odon.). Tijdschrift voor Entomologie, 88, 215 - 224. [1945]" type="journal article" year="1947">Lieftinck (1947</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFBE18EFFFB8BA72FBF9E7CAFB6D6AD7" author="Lieftinck" box="[1142,1199,1957,1981]" pageId="3" pageNumber="498" refString="Lieftinck, M. A. (1948) The Odonata of Enggano with a survey of the dragonfly-fauna of the West Sumatran chain of islands. Treubia, 19, 279 - 304." type="journal article" year="1948">1948</bibRefCitation>
|
||
) ranked
|
||
<taxonomicName id="4C2F1E9DFFB8BA72FA9CE7C9FAAA6AD7" baseAuthorityName="Lieftinck" baseAuthorityYear="1947" box="[1299,1384,1958,1981]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="angusta" subSpecies="oceanis">
|
||
<emphasis id="B95BB90CFFB8BA72FA9CE7C9FAAA6AD7" box="[1299,1384,1958,1981]" italics="true" pageId="3" pageNumber="498">oceanis</emphasis>
|
||
</taxonomicName>
|
||
as a subspecies and pointed out that melanism is a typical modification among several other dragonfly species in Enggano, but there are also marked exceptions in this rule; some of the Enggano
|
||
<taxonomicName id="4C2F1E9DFFBFBA75FBD8E0F8FB726DDA" authorityName="Fabricius" authorityYear="1793" box="[1111,1200,151,176]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="500" phylum="Arthropoda" rank="order">odonata</taxonomicName>
|
||
forms are lighter in colour than in mainland
|
||
<collectingRegion id="49EBABFCFFBFBA75FE24E0D3FDCE6DBE" box="[427,524,188,212]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="499">Sumatra</collectingRegion>
|
||
. It could be argued that
|
||
<taxonomicName id="4C2F1E9DFFBFBA75FC93E0D2FC366DBE" baseAuthorityName="Lieftinck" baseAuthorityYear="1947" box="[796,1012,189,212]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="499" phylum="Arthropoda" rank="subSpecies" species="angusta" subSpecies="oceanis">
|
||
<emphasis id="B95BB90CFFBFBA75FC93E0D2FC366DBE" box="[796,1012,189,212]" italics="true" pageId="4" pageNumber="499">H. angusta oceanis</emphasis>
|
||
</taxonomicName>
|
||
deserves specific status. However, as its wing shape and arrangement of vitrious fenestrae match well with those in
|
||
<emphasis id="B95BB90CFFBFBA75FB87E08EFB626D92" box="[1032,1184,225,248]" italics="true" pageId="4" pageNumber="499">
|
||
H. a.
|
||
<taxonomicName id="4C2F1E9DFFBFBA75FBC8E08EFB626D92" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1853" box="[1095,1184,225,248]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="499" phylum="Arthropoda" rank="species" species="angusta">angusta</taxonomicName>
|
||
</emphasis>
|
||
, it is better to wait for further evidence before considering any change in the taxon’s status.
|
||
</paragraph>
|
||
<paragraph id="8B90651EFFBFBA75FF48E147FEBF6C0E" blockId="4.[151,1436,151,357]" pageId="4" pageNumber="499">
|
||
Most likely
|
||
<taxonomicName id="4C2F1E9DFFBFBA75FEDCE146FE076C2A" authorityName="Hämäläinen" authorityYear="2016" box="[339,453,296,320]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="499" phylum="Arthropoda" rank="species" species="vantoli">
|
||
<emphasis id="B95BB90CFFBFBA75FEDCE146FE076C2A" box="[339,453,296,320]" italics="true" pageId="4" pageNumber="499">H. vantoli</emphasis>
|
||
</taxonomicName>
|
||
is endemic to Siberut Island. Its occurrence in other major islands of Mentawai is of course not ruled out.
|
||
</paragraph>
|
||
<caption id="DF503596FFBFBA75FF18E64DFAC26B52" box="[151,1280,1570,1592]" httpUri="https://zenodo.org/record/1051724/files/figure.png" pageId="4" pageNumber="499" targetBox="[415,1171,411,1545]" targetPageId="4">
|
||
<paragraph id="8B90651EFFBFBA75FF18E64DFAC26B52" blockId="4.[151,1280,1570,1592]" box="[151,1280,1570,1592]" pageId="4" pageNumber="499">
|
||
<emphasis id="B95BB90CFFBFBA75FF18E64DFED06B5D" bold="true" box="[151,274,1570,1592]" pageId="4" pageNumber="499">FIGURE 5.</emphasis>
|
||
The stream habitat in Siberut where the new species was found. Photo by Kirill A. Kolesnichenko.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B90651EFFBFBA74FF48E60BFC3D6CBA" blockId="4.[151,1437,1636,2021]" lastBlockId="5.[151,1437,151,464]" lastPageId="5" lastPageNumber="500" pageId="4" pageNumber="499">
|
||
<emphasis id="B95BB90CFFBFBA75FF48E60BFD3F6B17" bold="true" box="[199,765,1636,1661]" pageId="4" pageNumber="499">
|
||
Notes on the knowledge of Mentawai
|
||
<taxonomicName id="4C2F1E9DFFBFBA75FD02E60BFD356B17" authorityName="Fabricius" authorityYear="1793" box="[653,759,1636,1661]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="499" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The dragonfly fauna of Mentawai islands (Siberut, Sipura, North Pagai, South Pagai) is still inadequately known. In September-November 1924 the entomologists Cecil Boden Kloss, Norman Smedley and Heinrich Karny collected in Siberut and Sipura.
|
||
<bibRefCitation id="EFBE18EFFFBFBA75FBD8E6C3FAC76BAF" author="Laidlaw" box="[1111,1285,1708,1733]" pageId="4" pageNumber="499" refString="Laidlaw, F. F. (1926) Spolia Mentawiensia. Dragonflies (Odonata). Journal of the Malayan Branch of the Royal Asiatic Society, 4 (2), 214 - 233." type="journal article" year="1926">Laidlaw (1926)</bibRefCitation>
|
||
identified 34 species from their material, including 27 species from Siberut, 18 from Sipura and 8 from the Pagai islands. In the earlier literature only one dragonfly specimen had been recorded from Mentawai, i.e. the
|
||
<typeStatus id="5494DBBCFFBFBA75FB5DE69BFAF66A67" box="[1234,1332,1780,1805]" pageId="4" pageNumber="499" type="holotype">holotype</typeStatus>
|
||
male of
|
||
<taxonomicName id="4C2F1E9DFFBFBA75FF18E776FDDB6A5B" authority="Selys, 1898" authorityName="Selys" authorityYear="1898" box="[151,537,1816,1841]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="499" phylum="Arthropoda" rank="species" species="modiagliani">
|
||
<emphasis id="B95BB90CFFBFBA75FF18E776FE486A5B" box="[151,394,1817,1841]" italics="true" pageId="4" pageNumber="499">Euphaea modiagliani</emphasis>
|
||
<bibRefCitation id="EFBE18EFFFBFBA75FE1DE777FDDB6A5B" author="Selys" box="[402,537,1816,1841]" pageId="4" pageNumber="499" refString="Selys Longchamps, E. de (1898) Causeries odonatologiques 11. Annales de la Societe Entomologique de Belgique, 42, 332 - 338." type="journal article" year="1898">Selys, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
, collected by Elio Modigliani on ‘Ile Metawei’ (
|
||
<bibRefCitation id="EFBE18EFFFBFBA75FBCEE777FA9E6A5B" author="Selys" box="[1089,1372,1816,1841]" pageId="4" pageNumber="499" refString="Selys Longchamps, E. de (1898) Causeries odonatologiques 11. Annales de la Societe Entomologique de Belgique, 42, 332 - 338." type="journal article" year="1898">Selys Longchamps 1898</bibRefCitation>
|
||
). It is known that the only island in Mentawai visited by Modigliani was Sipura, where he collected in 1894. Lieftinck’s (1954) handlist did not include any additions to those species listed by
|
||
<bibRefCitation id="EFBE18EFFFBFBA75FC3EE70FFBA06A13" author="Laidlaw" box="[945,1122,1888,1913]" pageId="4" pageNumber="499" refString="Laidlaw, F. F. (1926) Spolia Mentawiensia. Dragonflies (Odonata). Journal of the Malayan Branch of the Royal Asiatic Society, 4 (2), 214 - 233." type="journal article" year="1926">Laidlaw (1926)</bibRefCitation>
|
||
. As far as I know, later only one addition to the fauna has been published;
|
||
<bibRefCitation id="EFBE18EFFFBFBA75FD1BE7EBFC446AF7" author="Karube" box="[660,902,1924,1949]" pageId="4" pageNumber="499" refString="Karube, H. & Yeh, W. - C. (2001) Sarasaeschna gen. nov., with descriptions of female S. minuta (Asahina) and male penile structures of Linaeschna (Anisoptera: Aeshnidae). Tombo - Acta Odonatologica Japonica, 43, 1 - 8." type="journal article" year="2001">Karube & Yeh (2001)</bibRefCitation>
|
||
listed
|
||
<taxonomicName id="4C2F1E9DFFBFBA75FC40E7EAFA5E6AF7" authority="Lieftinck, 1953" authorityName="Lieftinck" authorityYear="1953" box="[975,1436,1924,1949]" class="Insecta" family="Aeshnidae" genus="Oligoaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="499" phylum="Arthropoda" rank="species" species="sumatrana">
|
||
<emphasis id="B95BB90CFFBFBA75FC40E7EAFB2B6AF6" box="[975,1257,1925,1949]" italics="true" pageId="4" pageNumber="499">Oligoaeschna sumatrana</emphasis>
|
||
Lieftinck, 1953
|
||
</taxonomicName>
|
||
from ‘Sibelute I.’ (misspelling of Siberut Island). In addition
|
||
<bibRefCitation id="EFBE18EFFFBFBA75FCF3E7C7FBF86AAB" author="Lieftinck" box="[892,1082,1960,1985]" pageId="4" pageNumber="499" refString="Lieftinck, M. A. (1968) A review of the genus Oligoaeschna Selys in Southeast Asia. Tijdschrift voor Entomologie, 111 (5), 137 - 184, pls. 12 - 13 excl." type="journal article" year="1968">Lieftinck (1968)</bibRefCitation>
|
||
reidentified Laidlaw’s (1926) ‘
|
||
<emphasis id="B95BB90CFFBFBA75FF2FE7A2FEBB6A8F" box="[160,377,1997,2021]" italics="true" pageId="4" pageNumber="499">
|
||
Jagoria
|
||
<taxonomicName id="4C2F1E9DFFBFBA75FE8FE7A2FEBB6A8F" box="[256,377,1997,2021]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="499" phylum="Arthropoda" rank="species" species="modigliani">modigliani</taxonomicName>
|
||
</emphasis>
|
||
Selys’ from Siberut and Sipura as a new species
|
||
<taxonomicName id="4C2F1E9DFFBFBA75FC20E7A2FABA6A8F" authority="Lieftinck, 1968" authorityName="Lieftinck" authorityYear="1968" box="[943,1400,1996,2021]" class="Insecta" family="Aeshnidae" genus="Oligoaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="499" phylum="Arthropoda" rank="species" species="uropetala">
|
||
<emphasis id="B95BB90CFFBFBA75FC20E7A2FB7D6A8F" box="[943,1215,1997,2021]" italics="true" pageId="4" pageNumber="499">Oligoaeschna uropetala</emphasis>
|
||
<bibRefCitation id="EFBE18EFFFBFBA75FB48E7A3FABA6A8F" author="Lieftinck" box="[1223,1400,1996,2021]" pageId="4" pageNumber="499" refString="Lieftinck, M. A. (1968) A review of the genus Oligoaeschna Selys in Southeast Asia. Tijdschrift voor Entomologie, 111 (5), 137 - 184, pls. 12 - 13 excl." type="journal article" year="1968">Lieftinck, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
. In earlier literature only three calopterygoid species are listed from the Mentawai islands:
|
||
<taxonomicName id="4C2F1E9DFFBEBA74FB23E0F7FF376DBF" authority="(Sipura)" authorityName="Sipura" class="Insecta" family="Chlorocyphidae" genus="Libellago" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="sumatrana">
|
||
<emphasis id="B95BB90CFFBEBA74FB23E0F7FA5E6DDA" box="[1196,1436,152,176]" italics="true" pageId="5" pageNumber="500">Libellago sumatrana</emphasis>
|
||
(Sipura)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2F1E9DFFBEBA74FE8BE0D3FD936DBF" authority="(Sipura)" authorityName="Sipura" box="[260,593,188,213]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="modigliani">
|
||
<emphasis id="B95BB90CFFBEBA74FE8BE0D3FE286DBE" box="[260,490,188,212]" italics="true" pageId="5" pageNumber="500">Euphaea modigliani</emphasis>
|
||
(Sipura)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2F1E9DFFBEBA74FD02E0D3FB8B6DBE" authority="Albarda in Selys, 1879" authorityName="Albarda in Selys" authorityYear="1879" box="[653,1097,188,213]" class="Insecta" family="Calopterygidae" genus="Vestalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B95BB90CFFBEBA74FD02E0D3FCF76DBE" box="[653,821,188,212]" italics="true" pageId="5" pageNumber="500">Vestalis lugens</emphasis>
|
||
Albarda
|
||
<emphasis id="B95BB90CFFBEBA74FC2BE0D2FC786DBE" box="[932,954,189,212]" italics="true" pageId="5" pageNumber="500">in</emphasis>
|
||
Selys, 1879
|
||
</taxonomicName>
|
||
(Siberut, Sipura). The recent calopterygoid material from Siberut, which I received for study, also includes specimens of
|
||
<taxonomicName id="4C2F1E9DFFBEBA74FB5FE08FFABB6D92" authorityName="Albarda in Selys" authorityYear="1879" box="[1232,1401,224,248]" class="Insecta" family="Calopterygidae" genus="Vestalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B95BB90CFFBEBA74FB5FE08FFABB6D92" box="[1232,1401,224,248]" italics="true" pageId="5" pageNumber="500">Vestalis lugens</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<specimenCount id="9D29AE97FFBEBA74FA02E08FFF196C77" pageId="5" pageNumber="500" type="male">2 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D29AE97FFBEBA74FF68E16BFE896C77" box="[231,331,260,285]" pageId="5" pageNumber="500" type="female">1 female</specimenCount>
|
||
) and
|
||
<taxonomicName id="4C2F1E9DFFBEBA74FE03E16BFD1B6C77" authority="Selys, 1853" authorityName="Selys" authorityYear="1853" box="[396,729,260,285]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="aspasia">
|
||
<emphasis id="B95BB90CFFBEBA74FE03E16BFD8F6C76" box="[396,589,260,284]" italics="true" pageId="5" pageNumber="500">Euphaea aspasia</emphasis>
|
||
Selys, 1853
|
||
</taxonomicName>
|
||
(
|
||
<specimenCount id="9D29AE97FFBEBA74FD65E16BFC976C77" box="[746,853,260,285]" pageId="5" pageNumber="500" type="male">10 males</specimenCount>
|
||
), with the same collecting data as for
|
||
<taxonomicName id="4C2F1E9DFFBEBA74FA8BE16AFAB76C76" authorityName="Hämäläinen" authorityYear="2016" box="[1284,1397,260,284]" class="Insecta" family="Chlorocyphidae" genus="Heliocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="vantoli">
|
||
<emphasis id="B95BB90CFFBEBA74FA8BE16AFAB76C76" box="[1284,1397,260,284]" italics="true" pageId="5" pageNumber="500">H. vantoli</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C2F1E9DFFBEBA74FA0BE16AFF2F6C55" authorityName="Selys" authorityYear="1853" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="aspasia">
|
||
<emphasis id="B95BB90CFFBEBA74FA0BE16AFF2F6C55" italics="true" pageId="5" pageNumber="500">E. aspasia</emphasis>
|
||
</taxonomicName>
|
||
is a new record to the Mentawai islands. The
|
||
<taxonomicName id="4C2F1E9DFFBEBA74FD64E146FCA26C55" authorityName="Selys" authorityYear="1853" box="[747,864,296,320]" class="Insecta" family="Euphaeidae" genus="Euphaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="aspasia">
|
||
<emphasis id="B95BB90CFFBEBA74FD64E146FCA26C55" box="[747,864,296,320]" italics="true" pageId="5" pageNumber="500">E. aspasia</emphasis>
|
||
</taxonomicName>
|
||
males from Siberut lack pale markings on the head and prothorax and the pale markings on the synthorax are restricted to narrow stripes on the metepisternum and metepimeron. The wings lack the distinct hyaline lower margins present in the mainland Sumatran specimens. The Siberut specimens are also slightly smaller. The Siberut form could qualify as a distinct insular subspecies. The two additions bring the number of
|
||
<taxonomicName id="4C2F1E9DFFBEBA74FE64E1D8FD886CBA" authorityName="Fabricius" authorityYear="1793" box="[491,586,439,464]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="500" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
|
||
species listed from these islands to 38.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |