339 lines
34 KiB
XML
339 lines
34 KiB
XML
<document id="341940F892C0203F6D88A8514CC026B4" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFFD7737732FAFC50E652FC32" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium endemicum Wesener 2009, new species" docType="treatment" docUuid="CB700299-60E2-468D-889E-0EF0791471CA" docUuidSource="ZooBank" docVersion="5" lastPageNumber="40" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="37" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="B0A57CD97E7AE5A895AF4CE7B32E0BD5">
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<mods:title id="EE17DD57DDC99E1516E367A33220029A">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
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<mods:name id="A157D60FAB160712B84F4DF7CF8F9447" type="personal">
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<mods:namePart id="6482082802F7011A2EC62A3096BB0E2D">Wesener, Thomas</mods:namePart>
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<mods:typeOfResource id="832D5A40636816806C131B08C76EFC30">text</mods:typeOfResource>
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<mods:title id="7D94320A28E7133DA1343A30BF038C1B">Zootaxa</mods:title>
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<mods:part id="7565A82F07F9FCC5A2D8841ED18DC7AF">
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<mods:date id="E3F7D7910C760050A969F36013DE5A0E">2009</mods:date>
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<mods:detail id="8938659691B83E7604EDC3783D87A9C7" type="pubDate">
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<mods:number id="530CF7F024D6ADC2BFD812B38EFFBE50">2009-05-11</mods:number>
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</mods:detail>
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<mods:detail id="301276549A88915AD0F0B582762287FE" type="volume">
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<mods:number id="FB03D02A33D172F06493D2371F067AE5">2097</mods:number>
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</mods:detail>
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<mods:detail id="8AF0FDA8BFD8F84F843CB2D42B1AC816" type="issue">
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<mods:number id="73F2A2265A7CE51896688CC54DC6BCF1">1</mods:number>
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<mods:url id="DC0305E7D9480D4A6177DC99C1CD4EAF">https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1</mods:url>
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</mods:location>
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<mods:classification id="F4C550DDB7F738620043CAC14D4D5D46">journal article</mods:classification>
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<mods:identifier id="505AF574D7A246C99E92F684AB06C2FC" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
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<mods:identifier id="E46236793C4D61A721F0BD739B3FDFBB" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="C17348D068FB0DBC44A5973F89C727CF" type="Zenodo-Dep">5316536</mods:identifier>
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<mods:identifier id="45A9B57CD2EF25548BC8B94932540614" type="ZooBank">0E18167A-78EB-4079-8BF4-972A4AC88C69</mods:identifier>
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</mods:mods>
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<treatment id="03FFBE1DFFD7737732FAFC50E652FC32" ID-DOI="http://doi.org/10.5281/zenodo.5325455" ID-Zenodo-Dep="5325455" LSID="urn:lsid:zoobank.org:act:CB700299-60E2-468D-889E-0EF0791471CA" httpUri="http://treatment.plazi.org/id/03FFBE1DFFD7737732FAFC50E652FC32" lastPageId="39" lastPageNumber="40" pageId="36" pageNumber="37">
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<subSubSection id="C34C5C80FFD7737432FAFC50E17FFBDD" box="[151,632,1002,1028]" pageId="36" pageNumber="37" type="nomenclature">
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<paragraph id="8BE90F0BFFD7737432FAFC50E17FFBDD" blockId="36.[151,632,1002,1028]" box="[151,632,1002,1028]" pageId="36" pageNumber="37">
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<heading id="D0A1B867FFD7737432FAFC50E17FFBDD" bold="true" box="[151,632,1002,1028]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
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||
<emphasis id="B922D319FFD7737432FAFC50E17FFBDD" bold="true" box="[151,632,1002,1028]" pageId="36" pageNumber="37">
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<taxonomicName id="4C567488FFD7737432FAFC50E2DDFBDD" ID-CoL="5D7GY" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,474,1002,1028]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="endemicum" status="sp. nov.">
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<emphasis id="B922D319FFD7737432FAFC50E2DDFBDD" bold="true" box="[151,474,1002,1028]" italics="true" pageId="36" pageNumber="37">Zoosphaerium endemicum</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2116E62FFD773743384FC50E17FFBDD" box="[489,632,1002,1028]" pageId="36" pageNumber="37" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C34C5C80FFD7737432FAFB89E3D0FAD6" pageId="36" pageNumber="37" type="etymology">
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||
<paragraph id="8BE90F0BFFD7737432FAFB89E2CDFBAD" blockId="36.[151,1436,1075,2030]" pageId="36" pageNumber="37">
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<emphasis id="B922D319FFD7737432FAFB89E285FB94" bold="true" box="[151,386,1075,1101]" pageId="36" pageNumber="37">Derivatio nominis:</emphasis>
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<taxonomicName id="4C567488FFD7737433E1FB8EE116FB94" authorityName="Wesener" authorityYear="2009" box="[396,529,1076,1101]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="endemicum">
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<emphasis id="B922D319FFD7737433E1FB8EE116FB94" box="[396,529,1076,1101]" italics="true" pageId="36" pageNumber="37">endemicum</emphasis>
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</taxonomicName>
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, adjective, refers to the restricted distribution of the species to the Western rainforest of Analavelona.
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</paragraph>
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<paragraph id="8BE90F0BFFD7737432ABFB3AE3D0FAD6" blockId="36.[151,1436,1075,2030]" pageId="36" pageNumber="37">
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<emphasis id="B922D319FFD7737432ABFB3AE2A4FB43" bold="true" box="[198,419,1152,1178]" pageId="36" pageNumber="37">Studied material:</emphasis>
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<materialsCitation id="3B3E0556FFD7737433C0FB3AE3D0FAD6" collectingDate="1998-03-09" collectingDateMax="1998-03-15" collectingDateMin="1998-03-09" collectionCode="FMMC" collectorName="S. Goodman" country="Madagascar" county="Locus" elevation="1050" latitude="-22.678333" location="Foret" longLatPrecision="125" longitude="44.191666" municipality="Province" pageId="36" pageNumber="37" specimenCode="FMMC 5420" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
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||
<emphasis id="B922D319FFD7737433C0FB3AE118FB43" bold="true" box="[429,543,1152,1178]" pageId="36" pageNumber="37">
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||
<typeStatus id="54EDB1A9FFD7737433C0FB3AE118FB43" box="[429,543,1152,1178]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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: 1 M (
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||
<quantity id="4CAEA2EEFFD773743015FB3BE1D7FB42" box="[632,720,1153,1179]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="36" pageNumber="37" unit="mm" value="24.0">24 mm</quantity>
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long),
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<emphasis id="B922D319FFD773743147FB3AE0CAFB43" bold="true" box="[810,973,1152,1178]" pageId="36" pageNumber="37">
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||
<specimenCode id="DBF0A770FFD773743147FB3AE0CAFB43" box="[810,973,1152,1178]" collectionCode="FMMC" pageId="36" pageNumber="37">FMMC 5420</specimenCode>
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||
</emphasis>
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||
.
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<emphasis id="B922D319FFD7737431B2FB3BE784FB43" box="[991,1155,1153,1178]" italics="true" pageId="36" pageNumber="37">
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<collectingCounty id="62887787FFD7737431B2FB3BE722FB43" box="[991,1061,1153,1178]" pageId="36" pageNumber="37">Locus</collectingCounty>
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typicus
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</emphasis>
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:
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<collectingCountry id="F3414F9BFFD7737436F8FB3BE622FB42" box="[1173,1317,1153,1179]" name="Madagascar" pageId="36" pageNumber="37">Madagascar</collectingCountry>
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,
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<collectingMunicipality id="6B8D9571FFD773743759FB3BE69BFB42" box="[1332,1436,1153,1179]" pageId="36" pageNumber="37">Province</collectingMunicipality>
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<collectingRegion id="4992C1E9FFD7737432FAFB12E3EAFB1B" box="[151,237,1192,1218]" country="Madagascar" name="Toliara" pageId="36" pageNumber="37">Toliara</collectingRegion>
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,
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<location id="8E8959D0FFD773743294FB12E230FB1B" LSID="urn:lsid:plazi:treatment:03FFBE1DFFD7737732FAFC50E652FC32:8E8959D0FFD773743294FB12E230FB1B" box="[249,311,1192,1218]" country="Madagascar" county="Locus" latitude="-22.678333" longLatPrecision="125" longitude="44.191666" municipality="Province" name="Foret" pageId="36" pageNumber="37" stateProvince="Toliara">Forêt</location>
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<taxonomicName id="4C567488FFD77374332DFB12E171FB1B" authority=", Antanimena" authorityName="Antanimena" box="[320,630,1192,1218]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="analavelona">Analavelona, Antanimena</taxonomicName>
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,
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<locationDeviation id="9A5B1629FFD7737430EEFB12E0C0FB1B" box="[643,967,1192,1218]" location="Andranoheza" pageId="36" pageNumber="37">
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||
<quantity id="4CAEA2EEFFD7737430EEFB12E1E3FB1B" box="[643,740,1192,1218]" metricMagnitude="4" metricUnit="m" metricValue="1.25" pageId="36" pageNumber="37" unit="km" value="12.5">12.5 km</quantity>
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NW Andranoheza
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</locationDeviation>
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,
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||
<geoCoordinate id="EE6269CCFFD7737431B9FB12E74FFB1B" box="[980,1096,1192,1218]" degrees="22" direction="south" minutes="40.7" orientation="latitude" pageId="36" pageNumber="37" precision="92" value="-22.678333">22°40.7'S</geoCoordinate>
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,
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<geoCoordinate id="EE6269CCFFD77374363BFB12E7CCFB1B" box="[1110,1227,1192,1218]" degrees="44" direction="east" minutes="11.5" orientation="longitude" pageId="36" pageNumber="37" precision="92" value="44.191666">44°11.5'E</geoCoordinate>
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, transitional midaltitude forest with elements of western and eastern forest,
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<quantity id="4CAEA2EEFFD773743131FB74E0BEFB31" box="[860,953,1230,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="36" pageNumber="37" unit="m" value="1050.0">
|
||
<elevation id="007BE838FFD773743131FB74E0BEFB31" box="[860,953,1230,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="36" pageNumber="37" unit="m" value="1050.0">1050 m</elevation>
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</quantity>
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||
, coll.
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||
<collectorName id="26A36ADDFFD773743669FB74E79BFB31" box="[1028,1180,1230,1256]" pageId="36" pageNumber="37">S. Goodman</collectorName>
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||
,
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||
<date id="FFE829CBFFD7737436C4FB74E64FFB31" box="[1193,1352,1230,1256]" pageId="36" pageNumber="37" value="1998-03-09" valueMax="1998-03-15" valueMin="1998-03-09">
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||
<collectingDate id="EFACD023FFD7737436C4FB74E64FFB31" box="[1193,1352,1230,1256]" pageId="36" pageNumber="37" value="1998-03-09" valueMax="1998-03-15" valueMin="1998-03-09">9–15.iii.1998</collectingDate>
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||
</date>
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||
,
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<collectingMethod id="5217771CFFD77374373BFB74E3D3FAD6" pageId="36" pageNumber="37">pitfall traps</collectingMethod>
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.
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34C5C80FFD7737432ABFAA1E7E8FAEF" box="[198,1263,1307,1334]" pageId="36" pageNumber="37" type="distribution">
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||
<paragraph id="8BE90F0BFFD7737432ABFAA1E7E8FAEF" blockId="36.[151,1436,1075,2030]" box="[198,1263,1307,1334]" pageId="36" pageNumber="37">
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<emphasis id="B922D319FFD7737432ABFAA1E2F4FAEC" bold="true" box="[198,499,1307,1333]" pageId="36" pageNumber="37">Distribution & Ecology:</emphasis>
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Only known from the transitional forest of Analavelona (Fig. 5).
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</paragraph>
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</subSubSection>
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<subSubSection id="C34C5C80FFD7737432ABFAF8E1E4F8DF" pageId="36" pageNumber="37" type="diagnosis">
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<paragraph id="8BE90F0BFFD7737432ABFAF8E7D3F9B2" blockId="36.[151,1436,1075,2030]" pageId="36" pageNumber="37">
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<emphasis id="B922D319FFD7737432ABFAF8E24FFA85" bold="true" box="[198,328,1346,1372]" pageId="36" pageNumber="37">Diagnosis:</emphasis>
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Small,
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||
<quantity id="4CAEA2EEFFD7737433CCFAF9E2F4FA84" box="[417,499,1347,1373]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="36" pageNumber="37" unit="mm" value="24.0">24 mm</quantity>
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long. Color green, head, legs and antennae also green. Tergites at margins thinly brown. Texture of tergites polished with glossy sheen. Disc of antenna with 11–13 apical cones (Fig. 25B). Movable finger of posterior telopods weakly curved, two large, non-sclerotized lobes and three sclerotized spines (Fig. 24D), posterior side with 24 sclerotized teeth (Fig. 24E). Fixed finger broad and stout. Third podomere of anterior telopods small, reaching only half of first podomeres height (Fig. 24C). Two stridulation ribs on male harp, lateral rib weakly developed (Fig. 24A). Anal shield with two locking carinae, first short, second twice as long as first. Locking carinae only weakly developed and difficult to detect. Anal shield slightly excavated on hindmost point, posteriorly covered with dense field of very short hair.
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</paragraph>
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<paragraph id="8BE90F0BFFD7737432ABF9CDE1E4F8DF" blockId="36.[151,1436,1075,2030]" pageId="36" pageNumber="37">
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<emphasis id="B922D319FFD7737432ABF9CDE28BF948" bold="true" box="[198,396,1655,1681]" pageId="36" pageNumber="37">Similar species:</emphasis>
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Although the unknown female makes it more difficult to separate
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<taxonomicName id="4C567488FFD7737436C1F9C2E655F948" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1196,1362,1656,1681]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="endemicum" status="sp. nov.">
|
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<emphasis id="B922D319FFD7737436C1F9C2E655F948" box="[1196,1362,1656,1681]" italics="true" pageId="36" pageNumber="37">Z. endemicum</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A2116E62FFD773743731F9C2E69BF94B" box="[1372,1436,1656,1682]" pageId="36" pageNumber="37" rank="species">n. sp.</taxonomicNameLabel>
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from other species, the unique combination of an excavation of the anal shield, the number of apical cones on the antenna, a green coloration, and the shape of anterior telopods and locking carinae separates this species unambiguously from other
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<taxonomicName id="4C567488FFD7737433BBF956E1E4F8DF" box="[470,739,1772,1798]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B922D319FFD7737433BBF956E178F8DC" box="[470,639,1772,1797]" italics="true" pageId="36" pageNumber="37">Zoosphaerium</emphasis>
|
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species.
|
||
</taxonomicName>
|
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</paragraph>
|
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</subSubSection>
|
||
<subSubSection id="C34C5C80FFD7737732ABF8A8E652FC32" lastPageId="39" lastPageNumber="40" pageId="36" pageNumber="37" type="description">
|
||
<paragraph id="8BE90F0BFFD7737432ABF8A8E10AF88A" blockId="36.[151,1436,1075,2030]" pageId="36" pageNumber="37">
|
||
<emphasis id="B922D319FFD7737432ABF8A8E267F8F5" bold="true" box="[198,352,1810,1836]" pageId="36" pageNumber="37">Description:</emphasis>
|
||
<emphasis id="B922D319FFD773743305F8A9E2DFF8F5" box="[360,472,1811,1836]" italics="true" pageId="36" pageNumber="37">Body size</emphasis>
|
||
,
|
||
<typeStatus id="54EDB1A9FFD77374338BF8A9E14BF8F4" box="[486,588,1811,1837]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
male: length up to
|
||
<quantity id="4CAEA2EEFFD77374315DF8A9E081F8F4" box="[816,902,1811,1837]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="36" pageNumber="37" unit="mm" value="24.0">24 mm</quantity>
|
||
, width of thoracic shield up to
|
||
<quantity id="4CAEA2EEFFD77374369CF8A9E641F8F4" box="[1265,1350,1811,1837]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="36" pageNumber="37" unit="mm" value="12.0">12 mm</quantity>
|
||
, height of thoracic shield up to
|
||
<quantity id="4CAEA2EEFFD7737433C1F883E10EF88A" box="[428,521,1849,1875]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="36" pageNumber="37" unit="mm" value="7.1">7.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD7737432AAF8DAE267F879" blockId="36.[151,1436,1075,2030]" pageId="36" pageNumber="37">
|
||
<emphasis id="B922D319FFD7737432AAF8DAE256F8A0" box="[199,337,1888,1913]" italics="true" pageId="36" pageNumber="37">Coloration:</emphasis>
|
||
Glossy. Shiny green, head collum, antenna and legs also green. Tergites with thin brown posterior margin.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD7737432AAF817E179F81E" blockId="36.[151,1436,1075,2030]" box="[199,638,1965,1991]" pageId="36" pageNumber="37">
|
||
<emphasis id="B922D319FFD7737432AAF817E217F81F" box="[199,272,1965,1990]" italics="true" pageId="36" pageNumber="37">Head:</emphasis>
|
||
Eyes with more than 75 ocelli.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD7737532AAF86EE03DFF01" blockId="36.[151,1436,1075,2030]" lastBlockId="37.[151,1436,152,216]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
|
||
<emphasis id="B922D319FFD7737432AAF86EE238F834" box="[199,319,2004,2029]" italics="true" pageId="36" pageNumber="37">Antennae:</emphasis>
|
||
Length of antennomeres: 1<2=3=4=5<6 (Fig. 25A), sixth antennomere bearing disc with 11–13 apical cones (Fig. 25B). First antennomere broadest, short and cylindrical, without sensilla or groove. Sclerotized teeth at base of antennomeres 1–3 (Fig. 25A).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD6737532FAF980E7F6F94F" blockId="37.[151,1436,1594,1687]" pageId="37" pageNumber="38">
|
||
<emphasis id="B922D319FFD6737532FAF980E23FF98B" bold="true" box="[151,312,1594,1618]" pageId="37" pageNumber="38">FIGURE 023.</emphasis>
|
||
Endoterga, underside of midbody tergites,
|
||
<typeStatus id="54EDB1A9FFD673753170F980E080F988" box="[797,903,1594,1617]" pageId="37" pageNumber="38" type="holotype">holotypes</typeStatus>
|
||
,
|
||
<emphasis id="B922D319FFD6737531FBF980E0AEF98B" bold="true" box="[918,937,1594,1618]" pageId="37" pageNumber="38">A</emphasis>
|
||
:
|
||
<taxonomicName id="4C567488FFD6737531D1F980E621F988" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[956,1318,1594,1617]" class="Diplopoda" family="Arthrosphaeridae" genus="Microsphaerotherium" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="anjozorobe" status="sp. nov.">
|
||
<emphasis id="B922D319FFD6737531D1F980E621F988" box="[956,1318,1594,1617]" italics="true" pageId="37" pageNumber="38">Microsphaerotherium anjozorobe</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B922D319FFD67375375DF980E677F98B" bold="true" box="[1328,1392,1594,1618]" pageId="37" pageNumber="38">
|
||
<taxonomicNameLabel id="A2116E62FFD67375375DF980E677F98B" box="[1328,1392,1594,1618]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<emphasis id="B922D319FFD6737537EEF980E692F98B" bold="true" box="[1411,1429,1594,1618]" pageId="37" pageNumber="38">B</emphasis>
|
||
:
|
||
<taxonomicName id="4C567488FFD6737532FAF9E7E290F9AD" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,407,1629,1652]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="haackeri" status="sp. nov.">
|
||
<emphasis id="B922D319FFD6737532FAF9E7E290F9AD" box="[151,407,1629,1652]" italics="true" pageId="37" pageNumber="38">Zoosphaerium haackeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B922D319FFD6737533CDF9E6E2D9F9AD" bold="true" box="[416,478,1628,1652]" pageId="37" pageNumber="38">
|
||
<taxonomicNameLabel id="A2116E62FFD6737533CDF9E6E2D9F9AD" box="[416,478,1628,1652]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<emphasis id="B922D319FFD673753382F9E6E105F9AD" bold="true" box="[495,514,1628,1652]" pageId="37" pageNumber="38">C</emphasis>
|
||
:
|
||
<taxonomicName id="4C567488FFD67375307FF9E7E173F9AD" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[530,628,1629,1652]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="album" status="sp. nov.">
|
||
<emphasis id="B922D319FFD67375307FF9E7E173F9AD" box="[530,628,1629,1652]" italics="true" pageId="37" pageNumber="38">Z. album</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B922D319FFD673753011F9E6E1BCF9AD" bold="true" box="[636,699,1628,1652]" pageId="37" pageNumber="38">
|
||
<taxonomicNameLabel id="A2116E62FFD673753011F9E6E1BCF9AD" box="[636,699,1628,1652]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<emphasis id="B922D319FFD6737530A6F9E6E1D9F9AD" bold="true" box="[715,734,1628,1652]" pageId="37" pageNumber="38">D</emphasis>
|
||
:
|
||
<taxonomicName id="4C567488FFD673753082F9E7E063F9AD" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[751,868,1629,1652]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="discolor" status="sp. nov.">
|
||
<emphasis id="B922D319FFD673753082F9E7E063F9AD" box="[751,868,1629,1652]" italics="true" pageId="37" pageNumber="38">Z. discolor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B922D319FFD673753103F9E6E0ABF9AD" bold="true" box="[878,940,1628,1652]" pageId="37" pageNumber="38">
|
||
<taxonomicNameLabel id="A2116E62FFD673753103F9E6E0ABF9AD" box="[878,940,1628,1652]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<emphasis id="B922D319FFD6737531D1F9E6E0C9F9AD" bold="true" box="[956,974,1628,1652]" pageId="37" pageNumber="38">E</emphasis>
|
||
:
|
||
<taxonomicName id="4C567488FFD6737531B2F9E7E740F9AD" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[991,1095,1629,1652]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="mitoho" status="sp. nov.">
|
||
<emphasis id="B922D319FFD6737531B2F9E7E740F9AD" box="[991,1095,1629,1652]" italics="true" pageId="37" pageNumber="38">Z. mitoho</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B922D319FFD67375363DF9E6E789F9AD" bold="true" box="[1104,1166,1628,1652]" pageId="37" pageNumber="38">
|
||
<taxonomicNameLabel id="A2116E62FFD67375363DF9E6E789F9AD" box="[1104,1166,1628,1652]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<emphasis id="B922D319FFD6737536F2F9E6E7A8F9AD" bold="true" box="[1183,1199,1628,1652]" pageId="37" pageNumber="38">F</emphasis>
|
||
:
|
||
<taxonomicName id="4C567488FFD6737536ADF9E7E653F9AD" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1216,1364,1629,1652]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pulchellum" status="sp. nov.">
|
||
<emphasis id="B922D319FFD6737536ADF9E7E653F9AD" box="[1216,1364,1629,1652]" italics="true" pageId="37" pageNumber="38">Z. pulchellum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B922D319FFD673753730F9E6E69BF9AD" bold="true" box="[1373,1436,1628,1652]" pageId="37" pageNumber="38">
|
||
<taxonomicNameLabel id="A2116E62FFD673753730F9E6E69BF9AD" box="[1373,1436,1628,1652]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Abbreviations:
|
||
<emphasis id="B922D319FFD673753351F9C5E248F94E" bold="true" box="[316,335,1663,1687]" pageId="37" pageNumber="38">ci</emphasis>
|
||
= cuticular impressions;
|
||
<emphasis id="B922D319FFD673753030F9C5E17DF94E" bold="true" box="[605,634,1663,1687]" pageId="37" pageNumber="38">IA</emphasis>
|
||
= inner area;
|
||
<emphasis id="B922D319FFD673753162F9C5E033F94E" bold="true" box="[783,820,1663,1687]" pageId="37" pageNumber="38">mb</emphasis>
|
||
= marginal bristles;
|
||
<emphasis id="B922D319FFD67375367DF9C5E735F94E" bold="true" box="[1040,1074,1663,1687]" pageId="37" pageNumber="38">mr</emphasis>
|
||
= marginal ridge.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD6737532AAF97FE2F0F906" blockId="37.[151,1436,1733,2029]" box="[199,503,1733,1759]" pageId="37" pageNumber="38">
|
||
<emphasis id="B922D319FFD6737532AAF97FE257F907" box="[199,336,1733,1758]" italics="true" pageId="37" pageNumber="38">Mouthparts</emphasis>
|
||
not dissected.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD6737532AAF956E1AEF8DC" blockId="37.[151,1436,1733,2029]" box="[199,681,1771,1797]" pageId="37" pageNumber="38">
|
||
<emphasis id="B922D319FFD6737532AAF956E220F8DC" box="[199,295,1772,1797]" italics="true" pageId="37" pageNumber="38">Collum:</emphasis>
|
||
Median part of collum glabrous.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD6737532AAF8A8E1C4F8F5" blockId="37.[151,1436,1733,2029]" box="[199,707,1810,1836]" pageId="37" pageNumber="38">
|
||
<emphasis id="B922D319FFD6737532AAF8A8E279F8F2" box="[199,382,1810,1835]" italics="true" pageId="37" pageNumber="38">Thoracic shield</emphasis>
|
||
: Surface similar to tergites.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD6737532AAF883E3D0F8A3" blockId="37.[151,1436,1733,2029]" pageId="37" pageNumber="38">
|
||
<emphasis id="B922D319FFD6737532AAF883E224F88B" box="[199,291,1849,1874]" italics="true" pageId="37" pageNumber="38">Tergites</emphasis>
|
||
: Paratergites not projecting posteriorly, straight. Tergites polished, covered with few isolated hairs.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD6737632AAF83DE143FF6B" blockId="37.[151,1436,1733,2029]" lastBlockId="38.[151,1436,152,444]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
|
||
<emphasis id="B922D319FFD6737532AAF83DE267F879" box="[199,352,1927,1952]" italics="true" pageId="37" pageNumber="38">Endotergum:</emphasis>
|
||
Inner section with short, elongated spines and few, isolated bristles. Between marginal ridge and inner area single row of small, rectangular-shaped cuticular impressions, standing closely to one another (Fig. 35A). Externally two rows of marginal bristles, located closely to one another. Bristles strongly extending beyond tergite (Fig. 35A).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD5737632AAFF05E2A6FEFF" blockId="38.[151,1436,152,444]" pageId="38" pageNumber="39">
|
||
<emphasis id="B922D319FFD5737632AAFF05E25BFF01" box="[199,348,191,216]" italics="true" pageId="38" pageNumber="39">Anal shield:</emphasis>
|
||
Excavated at hindmost point and covered posteriorly with dense field of minute hair. Underside carries two brownish, weakly developed locking carinae. Anterior one small, posterior carina 2.5 times longer than first.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD5737632FAFE88E3D0FE65" blockId="38.[151,1436,152,444]" pageId="38" pageNumber="39">
|
||
<emphasis id="B922D319FFD5737632FAFE88E3DFFE92" box="[151,216,306,331]" italics="true" pageId="38" pageNumber="39">Legs:</emphasis>
|
||
Tarsi of first pair with two, of second with two or three and of third pair of legs with seven or eight ventral spines. First two leg pairs with only weakly curved claws and without an apical spine. Tarsi of legs 4–21 with curved claws, 10 or 11 ventral spines and one apical spine. Femur 1.7 and tarsi 4 times longer than wide.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD5737632FAFA05E71FF9E7" blockId="38.[151,1436,1471,1599]" pageId="38" pageNumber="39">
|
||
<emphasis id="B922D319FFD5737632FAFA05E237FA0E" bold="true" box="[151,304,1471,1495]" pageId="38" pageNumber="39">FIGURE 024</emphasis>
|
||
.
|
||
<taxonomicName id="4C567488FFD573763352FA05E15CFA0F" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[319,603,1471,1494]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="endemicum" status="sp. nov.">
|
||
<emphasis id="B922D319FFD573763352FA05E15CFA0F" box="[319,603,1471,1494]" italics="true" pageId="38" pageNumber="39">Zoosphaerium endemicum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B922D319FFD573763009FA05E1A5FA0E" bold="true" box="[612,674,1471,1495]" pageId="38" pageNumber="39">
|
||
<taxonomicNameLabel id="A2116E62FFD573763009FA05E1A5FA0E" box="[612,674,1471,1495]" pageId="38" pageNumber="39" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male
|
||
<typeStatus id="54EDB1A9FFD573763082FA05E048FA0F" box="[751,847,1471,1494]" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
|
||
,
|
||
<emphasis id="B922D319FFD573763131FA05E068FA0E" bold="true" box="[860,879,1471,1495]" pageId="38" pageNumber="39">A</emphasis>
|
||
: right anterior telopod, anterior view;
|
||
<emphasis id="B922D319FFD57376377BFA05E62FFA0E" bold="true" box="[1302,1320,1471,1495]" pageId="38" pageNumber="39">B</emphasis>
|
||
: last three podomeres of anterior telopod, posterior view;
|
||
<emphasis id="B922D319FFD5737630E7FA5BE19AFA20" bold="true" box="[650,669,1505,1529]" pageId="38" pageNumber="39">C</emphasis>
|
||
: anterior telopod, podomeres II & III, lateral view;
|
||
<emphasis id="B922D319FFD5737636ADFA5BE7D4FA20" bold="true" box="[1216,1235,1505,1529]" pageId="38" pageNumber="39">D</emphasis>
|
||
: posterior telopod, anterior view;
|
||
<emphasis id="B922D319FFD573763356F9BEE24AF9C5" bold="true" box="[315,333,1540,1564]" pageId="38" pageNumber="39">E</emphasis>
|
||
: posterior telopod, posterior view. Abbreviations:
|
||
<emphasis id="B922D319FFD5737631EFF9BEE0ADF9C5" bold="true" box="[898,938,1540,1564]" pageId="38" pageNumber="39">cr-t</emphasis>
|
||
= crenulated teeth;
|
||
<emphasis id="B922D319FFD5737636FDF9BEE7A8F9C5" bold="true" box="[1168,1199,1540,1564]" pageId="38" pageNumber="39">IH</emphasis>
|
||
= inner horns;
|
||
<emphasis id="B922D319FFD573763709F9BEE686F9C5" bold="true" box="[1380,1409,1540,1564]" pageId="38" pageNumber="39">ml</emphasis>
|
||
= membranous lobe;
|
||
<emphasis id="B922D319FFD57376330EF99DE282F9E6" bold="true" box="[355,389,1575,1599]" pageId="38" pageNumber="39">s-p</emphasis>
|
||
= sclerotized spot;
|
||
<emphasis id="B922D319FFD57376303BF99DE17FF9E6" bold="true" box="[598,632,1575,1599]" pageId="38" pageNumber="39">SR</emphasis>
|
||
= stridulation ribs. Scale bars =
|
||
<quantity id="4CAEA2EEFFD5737631BEF99DE713F9E7" box="[979,1044,1575,1598]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="39" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD5737632AAF9CBE743F952" blockId="38.[151,1436,1649,2022]" box="[199,1092,1649,1675]" pageId="38" pageNumber="39">
|
||
<emphasis id="B922D319FFD5737632AAF9CBE28BF953" box="[199,396,1649,1674]" italics="true" pageId="38" pageNumber="39">Stigmatic plates:</emphasis>
|
||
First lobe curved, tip stout, protruding slightly above coxa.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD5737632AAF92DE29EF968" blockId="38.[151,1436,1649,2022]" box="[199,409,1687,1713]" pageId="38" pageNumber="39">Female unknown.</paragraph>
|
||
<paragraph id="8BE90F0BFFD5737632AAF904E13EF840" blockId="38.[151,1436,1649,2022]" pageId="38" pageNumber="39">
|
||
<emphasis id="B922D319FFD5737632AAF904E2D9F90E" box="[199,478,1726,1751]" italics="true" pageId="38" pageNumber="39">Male sexual characters</emphasis>
|
||
:
|
||
<emphasis id="B922D319FFD573763397F904E1CDF90E" box="[506,714,1726,1751]" italics="true" pageId="38" pageNumber="39">Anterior telopods</emphasis>
|
||
(Figs 24A–C): Podomeres on both sides with isolated, long hair. First podomere rectangular, width larger than height. Harp with two stridulation ribs, ribs straight and short, mesal rib longer and broader than lateral rib. Second podomere process lobe-like and little curved with rounded edges, protruding up to third podomere height. Third podomere cavity mesally with numerous sclerotized spots and
|
||
<emphasis id="B922D319FFD5737633F8F8E3E2D7F8AB" box="[405,464,1881,1906]" italics="true" pageId="38" pageNumber="39">circa</emphasis>
|
||
four sclerotized spines, laterally with
|
||
<emphasis id="B922D319FFD5737631F5F8E3E0D4F8AB" box="[920,979,1881,1906]" italics="true" pageId="38" pageNumber="39">circa</emphasis>
|
||
one crenulated, sclerotized tooth. One sclerotized spot located close to tip.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD5737732AAF81CE25EFF26" blockId="38.[151,1436,1649,2022]" lastBlockId="39.[151,1436,152,255]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
|
||
<emphasis id="B922D319FFD5737632AAF81CE2A1F866" box="[199,422,1958,1983]" italics="true" pageId="38" pageNumber="39">Posterior telopods</emphasis>
|
||
(Figs 24D, E): Movable finger of chela very weakly curved; hollowed-out inner margin with two non-sclerotized lobes and three smaller, sclerotized spines, posterior aspect with
|
||
<emphasis id="B922D319FFD573763754F877E673F83F" box="[1337,1396,1997,2022]" italics="true" pageId="38" pageNumber="39">circa</emphasis>
|
||
24 crenulated teeth. Fixed finger as broad as movable finger, tip weakly curved. Finger basally with single short spine, on inner margin broadly covered with numerous sclerotized spots. All three podomeres on both sides almost glabrous.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFD4737732FAFC6EE652FC32" blockId="39.[151,1365,980,1004]" box="[151,1365,980,1004]" pageId="39" pageNumber="40">
|
||
<emphasis id="B922D319FFD4737732FAFC6EE22AFC35" bold="true" box="[151,301,980,1004]" pageId="39" pageNumber="40">FIGURE 025</emphasis>
|
||
.
|
||
<taxonomicName id="4C567488FFD473773357FC6FE153FC35" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[314,596,981,1004]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="endemicum" status="sp. nov.">
|
||
<emphasis id="B922D319FFD473773357FC6FE153FC35" box="[314,596,981,1004]" italics="true" pageId="39" pageNumber="40">Zoosphaerium endemicum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B922D319FFD473773036FC6EE190FC35" bold="true" box="[603,663,980,1004]" pageId="39" pageNumber="40">
|
||
<taxonomicNameLabel id="A2116E62FFD473773036FC6EE190FC35" box="[603,663,980,1004]" pageId="39" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, SEM, antenna,
|
||
<typeStatus id="54EDB1A9FFD473773124FC6EE0AEFC32" box="[841,937,980,1003]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
|
||
,
|
||
<emphasis id="B922D319FFD4737731DEFC6EE0D7FC35" bold="true" box="[947,976,980,1004]" pageId="39" pageNumber="40">A:</emphasis>
|
||
lateral view;
|
||
<emphasis id="B922D319FFD47377360CFC6EE774FC35" bold="true" box="[1121,1139,980,1004]" pageId="39" pageNumber="40">B</emphasis>
|
||
: apical view on disc.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |