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<document id="E13432AA51962C956FD0470B146D028B" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFFD3737432FAF995E76DFC49" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium pulchellum Wesener 2009, new species" docType="treatment" docVersion="4" lastPageNumber="37" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="33" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="664AD38D64C57B593942255546B5C64E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="43BBF79A53E1E12F1718C6B3B03C996E">
<mods:title id="DAFD082769DE9768DAB7105A0D72C7A9">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
</mods:titleInfo>
<mods:name id="7DF22F579E0D993FA538E5920069DC85" type="personal">
<mods:role id="2FA145E6AFB8E3B0F9001591FAC3F219">
<mods:roleTerm id="064CCBAE644520C5AC31AA5C365202C3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="08BC22061F9F6248286FFDCF57F90CB4">Wesener, Thomas</mods:namePart>
</mods:name>
<mods:typeOfResource id="0FC65D30700E77DEC798BC308D79028B">text</mods:typeOfResource>
<mods:relatedItem id="8C4EA8596FE31CDDA855C5E9942EF86D" type="host">
<mods:titleInfo id="115D65E91070B26F08C1B4571BF2B94F">
<mods:title id="2906EC327B527ADF107A9FC4669CF45C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4A32FCB1B7FD530DF6D4A6EFD9E27EB9">
<mods:date id="BC96DA6BAD06B7739D14A2D37687BA04">2009</mods:date>
<mods:detail id="D75D322A65AC7A3407DC6375CD190C53" type="pubDate">
<mods:number id="4E4DDD913D8AF0229E31CCF415BE7A9C">2009-05-11</mods:number>
</mods:detail>
<mods:detail id="307EE85B358482B5B7C92E3085B8E2B5" type="volume">
<mods:number id="D2DA25323E4D8551CF8A34ADDAB00665">2097</mods:number>
</mods:detail>
<mods:detail id="1FFE0DF4B61CE471A261F38D58F19535" type="issue">
<mods:number id="AE45A155B06FC57B9DDC8DD42713A6F4">1</mods:number>
</mods:detail>
<mods:extent id="8BFE6E33DFF1765A4531364B06D7A51E" unit="page">
<mods:start id="E922030F23E11C309EA3619AE81F7594">1</mods:start>
<mods:end id="A5078670CB9168AB4E62399FB847D56A">134</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="4DED53409BD73DD6BF599AF31D5D369B">
<mods:url id="4F5D1249839290739D92282F81DF1FFE">https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1</mods:url>
</mods:location>
<mods:classification id="E3A7F073B58D0933F11A486BF30F02CA">journal article</mods:classification>
<mods:identifier id="DC183F728F69DD5D5B9138F6D90BB56F" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
<mods:identifier id="7A9B04E34E6317B555C8B4597D2247B3" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="66E4BA31140B12B8CE23130BA82AB041" type="Zenodo-Dep">5316536</mods:identifier>
<mods:identifier id="405FC5A2264D357F339E347A5FC0E81F" type="ZooBank">0E18167A-78EB-4079-8BF4-972A4AC88C69</mods:identifier>
</mods:mods>
<treatment id="03FFBE1DFFD3737432FAF995E76DFC49" ID-DOI="http://doi.org/10.5281/zenodo.5325453" ID-Zenodo-Dep="5325453" LSID="urn:lsid:plazi:treatment:03FFBE1DFFD3737432FAF995E76DFC49" httpUri="http://treatment.plazi.org/id/03FFBE1DFFD3737432FAF995E76DFC49" lastPageId="36" lastPageNumber="37" pageId="32" pageNumber="33">
<subSubSection id="C34C5C80FFD3737032FAF995E172F990" box="[151,629,1583,1609]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8BE90F0BFFD3737032FAF995E172F990" blockId="32.[151,629,1583,1609]" box="[151,629,1583,1609]" pageId="32" pageNumber="33">
<heading id="D0A1B867FFD3737032FAF995E172F990" bold="true" box="[151,629,1583,1609]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="B922D319FFD3737032FAF995E172F990" bold="true" box="[151,629,1583,1609]" pageId="32" pageNumber="33">
<taxonomicName id="4C567488FFD3737032FAF995E2D0F990" ID-CoL="7GH2R" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,471,1583,1609]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pulchellum" status="sp. nov.">
<emphasis id="B922D319FFD3737032FAF995E2D0F990" bold="true" box="[151,471,1583,1609]" italics="true" pageId="32" pageNumber="33">Zoosphaerium pulchellum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2116E62FFD373703388F995E172F990" box="[485,629,1583,1609]" pageId="32" pageNumber="33" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFD3737032FAF9C2E25CF8A3" pageId="32" pageNumber="33" type="etymology">
<paragraph id="8BE90F0BFFD3737032FAF9C2E7C4F94B" blockId="32.[151,1436,1656,1992]" box="[151,1219,1656,1682]" pageId="32" pageNumber="33">
<emphasis id="B922D319FFD3737032FAF9C2E287F94B" bold="true" box="[151,384,1656,1682]" pageId="32" pageNumber="33">Derivatio nominis:</emphasis>
<taxonomicName id="4C567488FFD3737033E5F9C3E10DF94B" authorityName="Wesener" authorityYear="2009" box="[392,522,1657,1682]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pulchellum">
<emphasis id="B922D319FFD3737033E5F9C3E10DF94B" box="[392,522,1657,1682]" italics="true" pageId="32" pageNumber="33">pulchellum</emphasis>
</taxonomicName>
, adjective, refers to the shiny coloration pattern (Fig. 17B).
</paragraph>
<paragraph id="8BE90F0BFFD3737032ABF925E2B3F8DF" blockId="32.[151,1436,1656,1992]" pageId="32" pageNumber="33">
<emphasis id="B922D319FFD3737032ABF925E2A5F960" bold="true" box="[198,418,1695,1721]" pageId="32" pageNumber="33">Studied material:</emphasis>
<materialsCitation id="3B3E0556FFD3737033C0F925E2B3F8DF" collectingDate="1995-11-17" collectingDateMax="1995-11-27" collectingDateMin="1995-11-17" collectionCode="FMMC" collectorName="NW Eminiminy &amp; S. Goodman" country="Madagascar" county="Locus" elevation="1500" latitude="-24.57" location="d'Andohahela" longLatPrecision="125" longitude="46.731667" municipality="Province" pageId="32" pageNumber="33" specimenCode="FMMC 5388" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<emphasis id="B922D319FFD3737033C0F925E118F960" bold="true" box="[429,543,1695,1721]" pageId="32" pageNumber="33">
<typeStatus id="54EDB1A9FFD3737033C0F925E118F960" box="[429,543,1695,1721]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
</emphasis>
: 1 M (
<quantity id="4CAEA2EEFFD37370301AF925E1C9F960" box="[631,718,1695,1721]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="32" pageNumber="33" unit="mm" value="30.0">30 mm</quantity>
long),
<emphasis id="B922D319FFD373703145F925E0CDF960" bold="true" box="[808,970,1695,1721]" pageId="32" pageNumber="33">
<specimenCode id="DBF0A770FFD373703145F925E0CDF960" box="[808,970,1695,1721]" collectionCode="FMMC" pageId="32" pageNumber="33">FMMC 5388</specimenCode>
</emphasis>
.
<collectingCounty id="62887787FFD3737031B0F925E722F960" box="[989,1061,1695,1721]" pageId="32" pageNumber="33">Locus</collectingCounty>
typicus:
<collectingCountry id="F3414F9BFFD3737036FBF925E622F960" box="[1174,1317,1695,1721]" name="Madagascar" pageId="32" pageNumber="33">Madagascar</collectingCountry>
,
<collectingMunicipality id="6B8D9571FFD373703759F925E69BF960" box="[1332,1436,1695,1721]" pageId="32" pageNumber="33">Province</collectingMunicipality>
<collectingRegion id="4992C1E9FFD3737032FAF97CE3EAF939" box="[151,237,1734,1760]" country="Madagascar" name="Toliara" pageId="32" pageNumber="33">Toliara</collectingRegion>
, RNI
<location id="8E8959D0FFD37370335DF97CE2D2F939" LSID="urn:lsid:plazi:treatment:03FFBE1DFFD3737432FAF995E76DFC49:8E8959D0FFD37370335DF97CE2D2F939" box="[304,469,1734,1760]" country="Madagascar" county="Locus" latitude="-24.57" longLatPrecision="125" longitude="46.731667" municipality="Province" name="d'Andohahela" pageId="32" pageNumber="33" stateProvince="Toliara">d'Andohahela</location>
, parcel 1, 15 km
<collectorName id="26A36ADDFFD3737030F6F97CE050F939" box="[667,855,1734,1760]" pageId="32" pageNumber="33">NW Eminiminy</collectorName>
,
<geoCoordinate id="EE6269CCFFD37370310FF97CE0D1F939" box="[866,982,1734,1760]" degrees="24" direction="south" minutes="34.2" orientation="latitude" pageId="32" pageNumber="33" precision="92" value="-24.57">24°34.2'S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFFD37370318FF97CE75FF939" box="[994,1112,1734,1760]" degrees="46" direction="east" minutes="43.9" orientation="longitude" pageId="32" pageNumber="33" precision="92" value="46.731667">46°43.9'E</geoCoordinate>
,
<quantity id="4CAEA2EEFFD37370360EF97CE7BAF939" box="[1123,1213,1734,1760]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="33" unit="m" value="1500.0">
<elevation id="007BE838FFD37370360EF97CE7BAF939" box="[1123,1213,1734,1760]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="33" unit="m" value="1500.0">1500 m</elevation>
</quantity>
, coll.
<collectorName id="26A36ADDFFD37370376EF97CE69FF939" box="[1283,1432,1734,1760]" pageId="32" pageNumber="33">S. Goodman</collectorName>
, camp 4,
<date id="FFE829CBFFD373703296F956E2B7F8DF" box="[251,432,1772,1798]" pageId="32" pageNumber="33" value="1995-11-17" valueMax="1995-11-27" valueMin="1995-11-17">
<collectingDate id="EFACD023FFD373703296F956E2B7F8DF" box="[251,432,1772,1798]" pageId="32" pageNumber="33" value="1995-11-17" valueMax="1995-11-27" valueMin="1995-11-17">17–27.XI.1995</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE90F0BFFD3737032AAF8A8E70FF8F4" blockId="32.[151,1436,1656,1992]" box="[199,1032,1810,1837]" pageId="32" pageNumber="33">
<materialsCitation id="3B3E0556FFD3737032AAF8A8E70FF8F4" box="[199,1032,1810,1837]" collectingDate="1995-11-17" collectingDateMax="1995-11-27" collectingDateMin="1995-11-17" collectionCode="FMMC" collectorName="NW Eminiminy &amp; S. Goodman" country="Madagascar" county="Locus" elevation="1500" latitude="-24.57" location="d'Andohahela" longLatPrecision="125" longitude="46.731667" municipality="Province" pageId="32" pageNumber="33" specimenCode="FMMC 5388" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
<emphasis id="B922D319FFD3737032AAF8A8E243F8F5" bold="true" box="[199,324,1810,1836]" pageId="32" pageNumber="33">
<typeStatus id="54EDB1A9FFD3737032AAF8A8E243F8F5" box="[199,324,1810,1836]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
</emphasis>
: 1 M, 1 F (with eggs),
<emphasis id="B922D319FFD37370303EF8A8E1F4F8F5" bold="true" box="[595,755,1810,1836]" pageId="32" pageNumber="33">
<specimenCode id="DBF0A770FFD37370303EF8A8E1F4F8F5" box="[595,755,1810,1836]" collectionCode="FMMC" pageId="32" pageNumber="33">FMMC 5388</specimenCode>
</emphasis>
. Same data as holotype
</materialsCitation>
</paragraph>
<paragraph id="8BE90F0BFFD3737032AAF883E25CF8A3" blockId="32.[151,1436,1656,1992]" pageId="32" pageNumber="33">
<materialsCitation id="3B3E0556FFD3737032AAF883E168F88A" box="[199,623,1849,1876]" collectionCode="FMMC" pageId="32" pageNumber="33" specimenCode="FMMC 5399" specimenCount="1">
<emphasis id="B922D319FFD3737032AAF883E284F88A" bold="true" box="[199,387,1849,1875]" pageId="32" pageNumber="33">Other material</emphasis>
: 1 M,
<emphasis id="B922D319FFD3737033A2F883E168F88A" bold="true" box="[463,623,1849,1875]" pageId="32" pageNumber="33">
<specimenCode id="DBF0A770FFD3737033A2F883E168F88A" box="[463,623,1849,1875]" collectionCode="FMMC" pageId="32" pageNumber="33">FMMC 5399</specimenCode>
</emphasis>
</materialsCitation>
; 1 F,
<materialsCitation id="3B3E0556FFD3737030DDF883E057F88A" box="[688,848,1849,1875]" collectionCode="FMMC" pageId="32" pageNumber="33" specimenCode="FMMC 5399" specimenCount="1">
<emphasis id="B922D319FFD3737030DDF883E057F88A" bold="true" box="[688,848,1849,1875]" pageId="32" pageNumber="33">
<specimenCode id="DBF0A770FFD3737030DDF883E057F88A" box="[688,848,1849,1875]" collectionCode="FMMC" pageId="32" pageNumber="33">FMMC 5399</specimenCode>
</emphasis>
</materialsCitation>
; 1 F,
<materialsCitation id="3B3E0556FFD3737031FCF883E736F88A" box="[913,1073,1849,1875]" collectionCode="FMMC" pageId="32" pageNumber="33" specimenCode="FMMC 5416" specimenCount="1">
<emphasis id="B922D319FFD3737031FCF883E736F88A" bold="true" box="[913,1073,1849,1875]" pageId="32" pageNumber="33">
<specimenCode id="DBF0A770FFD3737031FCF883E736F88A" box="[913,1073,1849,1875]" collectionCode="FMMC" pageId="32" pageNumber="33">FMMC 5416</specimenCode>
</emphasis>
</materialsCitation>
,
<materialsCitation id="3B3E0556FFD373703652F880E3ADF8A3" collectingDate="1995-11-17" collectingDateMax="1995-11-27" collectingDateMin="1995-11-17" collectionCode="FMMC" collectorName="NW Eminiminy &amp; S. Goodman" country="Madagascar" county="Locus" elevation="1500" latitude="-24.57" location="d'Andohahela" longLatPrecision="125" longitude="46.731667" municipality="Province" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Toliara">same data as holotype; 1 M, 3 F</materialsCitation>
,
<materialsCitation id="3B3E0556FFD3737032DEF8DAE253F8A3" box="[179,340,1888,1914]" collectionCode="FMMC" country="Madagascar" pageId="32" pageNumber="33" specimenCode="FMMC 5456" specimenCount="1" stateProvince="Toliara">
<emphasis id="B922D319FFD3737032DEF8DAE253F8A3" bold="true" box="[179,340,1888,1914]" pageId="32" pageNumber="33">
<specimenCode id="DBF0A770FFD3737032DEF8DAE253F8A3" box="[179,340,1888,1914]" collectionCode="FMMC" pageId="32" pageNumber="33">FMMC 5456</specimenCode>
</emphasis>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFD3737132AAF83DE235F96C" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="distribution">
<paragraph id="8BE90F0BFFD3737032AAF83DE25CF811" blockId="32.[151,1436,1656,1992]" pageId="32" pageNumber="33">
<emphasis id="B922D319FFD3737032AAF83DE267F878" bold="true" box="[199,352,1927,1953]" pageId="32" pageNumber="33">Distribution</emphasis>
: This species is known only from the
<quantity id="4CAEA2EEFFD37370314DF83DE07DF878" box="[800,890,1927,1953]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="33" unit="m" value="1500.0">1500 m</quantity>
montane rainforest range of RNI Andohahela, parcel 1 (Fig. 5).
</paragraph>
<paragraph id="8BE90F0BFFD2737132FAF98FE235F96C" blockId="33.[151,1436,1589,1717]" pageId="33" pageNumber="34">
<emphasis id="B922D319FFD2737132FAF98FE236F994" bold="true" box="[151,305,1589,1613]" pageId="33" pageNumber="34">FIGURE 020</emphasis>
.
<taxonomicName id="4C567488FFD27371332CF98CE128F994" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[321,559,1590,1613]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="mitoho" status="sp. nov.">
<emphasis id="B922D319FFD27371332CF98CE128F994" box="[321,559,1590,1613]" italics="true" pageId="33" pageNumber="34">Zoosphaerium mitoho</emphasis>
</taxonomicName>
<emphasis id="B922D319FFD273713054F98FE17FF994" bold="true" box="[569,632,1589,1613]" pageId="33" pageNumber="34">
<taxonomicNameLabel id="A2116E62FFD273713054F98FE17FF994" box="[569,632,1589,1613]" pageId="33" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, SEM,
<typeStatus id="54EDB1A9FFD2737130BCF98CE028F994" box="[721,815,1590,1613]" pageId="33" pageNumber="34" type="paratype">paratype</typeStatus>
,
<emphasis id="B922D319FFD273713153F98FE056F994" bold="true" box="[830,849,1589,1613]" pageId="33" pageNumber="34">A</emphasis>
: gnathochilarium, left inner palpus with rudimentary lateral palpus;
<emphasis id="B922D319FFD273713359F9E2E241F9A9" bold="true" box="[308,326,1624,1648]" pageId="33" pageNumber="34">B</emphasis>
: right mandible, mesal view. Abbreviations:
<emphasis id="B922D319FFD27371314AF9E2E049F9A9" bold="true" box="[807,846,1624,1648]" pageId="33" pageNumber="34">3iT</emphasis>
= 3 inner teeth;
<emphasis id="B922D319FFD273713190F9E2E719F9A9" bold="true" box="[1021,1054,1624,1648]" pageId="33" pageNumber="34">Co</emphasis>
= condylus;
<emphasis id="B922D319FFD2737136C4F9E2E7C0F9A9" bold="true" box="[1193,1223,1624,1648]" pageId="33" pageNumber="34">eT</emphasis>
= external tooth;
<emphasis id="B922D319FFD2737137E9F9E2E69BF9A9" bold="true" box="[1412,1436,1624,1648]" pageId="33" pageNumber="34">iP</emphasis>
= inner palpus;
<emphasis id="B922D319FFD27371332DF9C1E262F94A" bold="true" box="[320,357,1659,1683]" pageId="33" pageNumber="34">mp</emphasis>
= molar plate;
<emphasis id="B922D319FFD273713066F9C1E12BF94A" bold="true" box="[523,556,1659,1683]" pageId="33" pageNumber="34">pL</emphasis>
= pectinate lamella;
<emphasis id="B922D319FFD273713163F9C1E037F94A" bold="true" box="[782,816,1659,1683]" pageId="33" pageNumber="34">rlp</emphasis>
= rudimentary lateral palpus;
<emphasis id="B922D319FFD27371361BF9C1E784F94A" bold="true" box="[1142,1155,1659,1683]" pageId="33" pageNumber="34">v</emphasis>
= depression on posterior side of palpus.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFD2737232AAF95DE761F93A" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BE90F0BFFD2737232AAF95DE250FEAA" blockId="33.[151,1436,1767,2025]" lastBlockId="34.[151,1436,152,371]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<emphasis id="B922D319FFD2737132AAF95DE24EF8D8" bold="true" box="[199,329,1767,1793]" pageId="33" pageNumber="34">Diagnosis:</emphasis>
Up to
<quantity id="4CAEA2EEFFD2737133F1F95DE113F8D8" box="[412,532,1767,1793]" metricMagnitude="-2" metricUnit="m" metricValue="3.075" pageId="33" pageNumber="34" unit="mm" value="30.75">30.75 mm</quantity>
long. Color remarkable (Fig. 17B): tergites colored with two different kinds of green, one jade-like green and at anterior parts of tergites with darker green. Texture of tergites polished with glossy sheen (Fig. 17B), nevertheless covered with some long, isolated hairs and few minute setae. Disc of antenna with four apical cones (Fig. 22C). Movable finger of posterior telopods weakly curved, with two large, non-sclerotized lobes and three sclerotized spines (Fig. 21E), posterior side with up to 25 sclerotized teeth (Fig. 21F). Tip of fixed finger curved towards movable finger (Fig. 21F). Both fingers on both sides covered completely with numerous small pits, one short hair standing in each (Figs 21E, F). Two stridulation ribs on each male harp, mesal one very weakly developed, difficult to observe (Fig. 21B). Anal shield with two locking carinae, first short, second twice as long as first. Anal shield sexually dimorphic, in males weakly rounded and of rectangular shape, covered with dense field of very short hair, in large females apically excavated and weakly bell-shaped, sensorial hairs absent. Operculum small, ending before coxal margin. Apical margin of operculum not or only weakly notched (Fig. 21A). Washboard on each side with 2 or 3 stridulation ribs.
</paragraph>
<paragraph id="8BE90F0BFFD1737232FAFA3EE691F9FF" blockId="34.[151,1436,1412,1575]" pageId="34" pageNumber="35">
<emphasis id="B922D319FFD1737232FAFA3EE22AFA45" bold="true" box="[151,301,1412,1436]" pageId="34" pageNumber="35">FIGURE 021</emphasis>
.
<taxonomicName id="4C567488FFD173723354FA3FE148FA45" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[313,591,1413,1436]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pulchellum" status="sp. nov.">
<emphasis id="B922D319FFD173723354FA3FE148FA45" box="[313,591,1413,1436]" italics="true" pageId="34" pageNumber="35">Zoosphaerium pulchellum</emphasis>
</taxonomicName>
<emphasis id="B922D319FFD17372303BFA3EE196FA45" bold="true" box="[598,657,1412,1436]" pageId="34" pageNumber="35">
<taxonomicNameLabel id="A2116E62FFD17372303BFA3EE196FA45" box="[598,657,1412,1436]" pageId="34" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, A, female
<typeStatus id="54EDB1A9FFD173723160FA3EE06CFA42" box="[781,875,1412,1435]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
, B–F, male
<typeStatus id="54EDB1A9FFD173723185FA3EE74FFA42" box="[1000,1096,1412,1435]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
,
<emphasis id="B922D319FFD17372363FFA3EE762FA45" bold="true" box="[1106,1125,1412,1436]" pageId="34" pageNumber="35">A</emphasis>
: second left coxa with vulva, posterior view;
<emphasis id="B922D319FFD17372332EFA1DE259FA66" bold="true" box="[323,350,1447,1471]" pageId="34" pageNumber="35">B:</emphasis>
right side of anterior telopod, anterior view;
<emphasis id="B922D319FFD17372313EFA1DE061FA66" bold="true" box="[851,870,1447,1471]" pageId="34" pageNumber="35">C</emphasis>
: last three podomeres of anterior telopod, posterior view;
<emphasis id="B922D319FFD1737232B6FA73E3E9FA38" bold="true" box="[219,238,1481,1505]" pageId="34" pageNumber="35">D</emphasis>
: anterior telopod, last two podomeres, lateral view;
<emphasis id="B922D319FFD17372315CFA73E044FA38" bold="true" box="[817,835,1481,1505]" pageId="34" pageNumber="35">E</emphasis>
: posterior telopod, anterior view;
<emphasis id="B922D319FFD1737236D1FA73E7CBFA38" bold="true" box="[1212,1228,1481,1505]" pageId="34" pageNumber="35">F</emphasis>
: posterior telopod, posterior view. Abbreviations:
<emphasis id="B922D319FFD173723384FA56E115F9DD" bold="true" box="[489,530,1516,1540]" pageId="34" pageNumber="35">cr-t</emphasis>
= crenulated teeth;
<emphasis id="B922D319FFD17372309CFA56E016F9DD" bold="true" box="[753,785,1516,1540]" pageId="34" pageNumber="35">Cx</emphasis>
= coxa;
<emphasis id="B922D319FFD17372311AFA56E09EF9DD" bold="true" box="[887,921,1516,1540]" pageId="34" pageNumber="35">EP</emphasis>
= external plate of vulva;
<emphasis id="B922D319FFD1737236A9FA56E7D9F9DD" bold="true" box="[1220,1246,1516,1540]" pageId="34" pageNumber="35">IP</emphasis>
= mesal plate of vulva;
<emphasis id="B922D319FFD17372328DF9B5E3FAF9FE" bold="true" box="[224,253,1551,1575]" pageId="34" pageNumber="35">ml</emphasis>
= membranous lobe;
<emphasis id="B922D319FFD173723389F9B5E2FEF9FE" bold="true" box="[484,505,1551,1575]" pageId="34" pageNumber="35">O</emphasis>
= operculum of vulva;
<emphasis id="B922D319FFD17372309FF9B5E013F9FE" bold="true" box="[754,788,1551,1575]" pageId="34" pageNumber="35">s-p</emphasis>
= sclerotized spot;
<emphasis id="B922D319FFD17372318EF9B5E702F9FE" bold="true" box="[995,1029,1551,1575]" pageId="34" pageNumber="35">SR</emphasis>
= stridulation ribs. Scale bar =
<quantity id="4CAEA2EEFFD17372373FF9B5E695F9FF" box="[1362,1426,1551,1574]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="35" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFFD1737232A8F9EEE761F93A" blockId="34.[151,1436,1620,2033]" pageId="34" pageNumber="35">
<emphasis id="B922D319FFD1737232A8F9EEE278F9B7" bold="true" box="[197,383,1620,1646]" pageId="34" pageNumber="35">Similar species</emphasis>
: This species is probably closely related to
<taxonomicName id="4C567488FFD173723113F9EFE690F9B7" authority="(de Saussure &amp; Zehntner, 1897)" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[894,1431,1620,1646]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="anomalum">
<emphasis id="B922D319FFD173723113F9EFE71FF9B7" box="[894,1048,1621,1646]" italics="true" pageId="34" pageNumber="35">Z. anomalum</emphasis>
(
<bibRefCitation id="EFC772FAFFD173723645F9EEE689F9B7" author="de Saussure, H. &amp; Zehntner, L." box="[1064,1422,1620,1646]" pageId="34" pageNumber="35" pagination="1 - 12" refId="ref60128" refString="de Saussure, H. &amp; Zehntner, L. (1897) Atlas de l'histoire naturelle des Myriapodes, In: Grandidier (Ed.) Histoire physique, naturelle et politique de Madagascar. 27 (53), Alfred Grandidier, Paris, France, plates 1 - 12." type="book chapter" year="1897">de Saussure &amp; Zehntner, 1897</bibRefCitation>
)
</taxonomicName>
, as it shares unique pits and small sensorial hairs on the posterior telopods. Nevertheless does
<taxonomicName id="4C567488FFD1737236B1F9C1E67AF94D" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1244,1405,1659,1684]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pulchellum" status="sp. nov.">
<emphasis id="B922D319FFD1737236B1F9C1E67AF94D" box="[1244,1405,1659,1684]" italics="true" pageId="34" pageNumber="35">Z. pulchellum</emphasis>
</taxonomicName>
<emphasis id="B922D319FFD1737237E9F9C1E3BDF962" bold="true" pageId="34" pageNumber="35">
<taxonomicNameLabel id="A2116E62FFD1737237E9F9C1E3BDF962" pageId="34" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differ from
<taxonomicName id="4C567488FFD173723323F918E2EDF962" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[334,490,1698,1723]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="anomalum">
<emphasis id="B922D319FFD173723323F918E2EDF962" box="[334,490,1698,1723]" italics="true" pageId="34" pageNumber="35">Z. anomalum</emphasis>
</taxonomicName>
in the presence of two stridulation ribs on each male harp (
<quantity id="4CAEA2EEFFD1737236D4F918E7E1F965" box="[1209,1254,1698,1724]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="34" pageNumber="35" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="4C567488FFD173723683F918E68CF962" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[1262,1419,1698,1723]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="anomalum">
<emphasis id="B922D319FFD173723683F918E68CF962" box="[1262,1419,1698,1723]" italics="true" pageId="34" pageNumber="35">Z. anomalum</emphasis>
</taxonomicName>
), completely differently shaped anterior telopods and a different endotergum pattern.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFD1737432ABF955E76DFC49" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35" type="description">
<paragraph id="8BE90F0BFFD1737232ABF955E1C0F88F" blockId="34.[151,1436,1620,2033]" pageId="34" pageNumber="35">
<emphasis id="B922D319FFD1737232ABF955E250F8D0" bold="true" box="[198,343,1775,1801]" pageId="34" pageNumber="35">Description</emphasis>
:
<emphasis id="B922D319FFD17372330BF94AE2D2F8D0" box="[358,469,1776,1801]" italics="true" pageId="34" pageNumber="35">Body size</emphasis>
, males (
<specimenCount id="9D50C482FFD17372305AF955E1CAF8D0" box="[567,717,1775,1801]" pageId="34" pageNumber="35" type="generic">4 specimens</specimenCount>
): length up to
<quantity id="4CAEA2EEFFD17372311CF955E0C1F8D0" box="[881,966,1775,1801]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="34" pageNumber="35" unit="mm" value="31.0">31 mm</quantity>
, width of thoracic shield up to
<quantity id="4CAEA2EEFFD173723740F955E69FF8D0" box="[1325,1432,1775,1801]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="34" pageNumber="35" unit="mm" value="14.7">14.7 mm</quantity>
, height of thoracic shield up to
<quantity id="4CAEA2EEFFD173723064F8ACE16FF8E9" box="[521,616,1814,1840]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="34" pageNumber="35" unit="mm" value="8.7">8.7 mm</quantity>
. Females (
<specimenCount id="9D50C482FFD173723087F8ACE084F8E9" box="[746,899,1814,1840]" pageId="34" pageNumber="35" type="generic">6 specimens</specimenCount>
): length up to
<quantity id="4CAEA2EEFFD17372365FF8ACE78EF8E9" box="[1074,1161,1814,1840]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="34" pageNumber="35" unit="mm" value="31.0">31 mm</quantity>
, width up to
<quantity id="4CAEA2EEFFD173723747F8ACE69FF8E9" box="[1322,1432,1814,1840]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="34" pageNumber="35" unit="mm" value="12.5">12.5 mm</quantity>
, height up to
<quantity id="4CAEA2EEFFD173723347F886E280F88F" box="[298,391,1852,1878]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="34" pageNumber="35" unit="mm" value="8.3">8.3 mm</quantity>
. Both sexes of similar size.
</paragraph>
<paragraph id="8BE90F0BFFD1737232AAF8D9E1D3F8A4" blockId="34.[151,1436,1620,2033]" box="[199,724,1891,1917]" pageId="34" pageNumber="35">
<emphasis id="B922D319FFD1737232AAF8D9E223F8A5" box="[199,292,1891,1916]" italics="true" pageId="34" pageNumber="35">Habitus</emphasis>
: Body short and massive (Fig. 17B).
</paragraph>
<paragraph id="8BE90F0BFFD1737232AAF830E1F2F828" blockId="34.[151,1436,1620,2033]" pageId="34" pageNumber="35">
<emphasis id="B922D319FFD1737232AAF830E24FF87A" box="[199,328,1930,1955]" italics="true" pageId="34" pageNumber="35">Coloration</emphasis>
: Remarkable. Thoracic shield, paratergites, posterior half of tergites and anal shield with lightgreenish, jade-like coloration (Fig. 17B). Head, antenna and anterior half of tergites dark green. Legs light green with even lighter greenish coloration on tarsi.
</paragraph>
<paragraph id="8BE90F0BFFD0737332FAFB33E79EFB1A" blockId="35.[151,1435,1161,1219]" pageId="35" pageNumber="36">
<emphasis id="B922D319FFD0737332FAFB33E237FB78" bold="true" box="[151,304,1161,1185]" pageId="35" pageNumber="36">FIGURE 022</emphasis>
.
<taxonomicName id="4C567488FFD07373332DFB33E15DFB79" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[320,602,1161,1184]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pulchellum" status="sp. nov.">
<emphasis id="B922D319FFD07373332DFB33E15DFB79" box="[320,602,1161,1184]" italics="true" pageId="35" pageNumber="36">Zoosphaerium pulchellum</emphasis>
</taxonomicName>
<emphasis id="B922D319FFD073733009FB33E1A4FB78" bold="true" box="[612,675,1161,1185]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="A2116E62FFD073733009FB33E1A4FB78" box="[612,675,1161,1185]" pageId="35" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, SEM, antenna,
<typeStatus id="54EDB1A9FFD073733132FB33E0BAFB79" box="[863,957,1161,1184]" pageId="35" pageNumber="36" type="paratype">paratype</typeStatus>
,
<emphasis id="B922D319FFD0737331A7FB33E0E0FB78" bold="true" box="[970,999,1161,1185]" pageId="35" pageNumber="36">A:</emphasis>
lateral view;
<emphasis id="B922D319FFD0737336EFFB33E793FB78" bold="true" box="[1154,1172,1161,1185]" pageId="35" pageNumber="36">B</emphasis>
: first antennomere with
<emphasis id="B922D319FFD0737332FAFB16E265FB1A" box="[151,354,1196,1219]" italics="true" pageId="35" pageNumber="36">sensilla basiconica</emphasis>
, lateral view;
<emphasis id="B922D319FFD073733396FB11E109FB1A" bold="true" box="[507,526,1195,1219]" pageId="35" pageNumber="36">C</emphasis>
: apical view on disc. Abbreviation:
<emphasis id="B922D319FFD0737331FCFB11E0ADFB1A" bold="true" box="[913,938,1195,1219]" pageId="35" pageNumber="36">sb</emphasis>
=
<emphasis id="B922D319FFD0737331AAFB16E795FB1A" box="[967,1170,1196,1219]" italics="true" pageId="35" pageNumber="36">sensilla basiconica</emphasis>
.
</paragraph>
<paragraph id="8BE90F0BFFD0737332AAFB4BE17BFAD2" blockId="35.[151,1437,1265,2026]" box="[199,636,1265,1291]" pageId="35" pageNumber="36">
<emphasis id="B922D319FFD0737332AAFB4BE201FAD3" box="[199,262,1265,1290]" italics="true" pageId="35" pageNumber="36">Head</emphasis>
: Eyes with more than 75 ocelli.
</paragraph>
<paragraph id="8BE90F0BFFD0737332AAFAA2E7A0FAA6" blockId="35.[151,1437,1265,2026]" pageId="35" pageNumber="36">
<emphasis id="B922D319FFD0737332AAFAA2E231FAE8" box="[199,310,1304,1329]" italics="true" pageId="35" pageNumber="36">Antennae</emphasis>
: Length of antennomeres: 1&lt;2&lt;3&gt;4=5&gt;6 (Fig. 22A), sixth antennomere bearing disc with four apical cones (Fig. 22C). First antennomere broadest, short and cylindrical, with row of
<emphasis id="B922D319FFD0737336DFFA85E693FA81" box="[1202,1428,1343,1368]" italics="true" pageId="35" pageNumber="36">sensilla basiconica</emphasis>
; without groove (Fig. 22B). Sclerotized teeth at base of antennomeres 1 and 2 (Fig. 22A).
</paragraph>
<paragraph id="8BE90F0BFFD0737332AAFA36E2F4FA15" blockId="35.[151,1437,1265,2026]" pageId="35" pageNumber="36">
<emphasis id="B922D319FFD0737332AAFA36E296FA7C" box="[199,401,1420,1445]" italics="true" pageId="35" pageNumber="36">Gnathochilarium</emphasis>
: Lateral of palpi four sensory cones, located together. Depression on posterior side of palpi with few sensory cones.
</paragraph>
<paragraph id="8BE90F0BFFD0737332AAFA63E2D9FA2A" blockId="35.[151,1437,1265,2026]" box="[199,478,1497,1523]" pageId="35" pageNumber="36">
<emphasis id="B922D319FFD0737332AAFA63E230FA2B" box="[199,311,1497,1522]" italics="true" pageId="35" pageNumber="36">Mandible</emphasis>
not dissected.
</paragraph>
<paragraph id="8BE90F0BFFD0737332AAF9BAE1A0F9C3" blockId="35.[151,1437,1265,2026]" box="[199,679,1536,1562]" pageId="35" pageNumber="36">
<emphasis id="B922D319FFD0737332AAF9BAE21AF9C0" box="[199,285,1536,1561]" italics="true" pageId="35" pageNumber="36">Collum</emphasis>
: Median part of collum glabrous.
</paragraph>
<paragraph id="8BE90F0BFFD0737332AAF99DE03BF999" blockId="35.[151,1437,1265,2026]" box="[199,828,1574,1600]" pageId="35" pageNumber="36">
<emphasis id="B922D319FFD0737332AAF99DE279F999" box="[199,382,1575,1600]" italics="true" pageId="35" pageNumber="36">Thoracic shield</emphasis>
: Surface similar to tergites (Fig. 17B)
</paragraph>
<paragraph id="8BE90F0BFFD0737332AAF9F7E185F957" blockId="35.[151,1437,1265,2026]" pageId="35" pageNumber="36">
<emphasis id="B922D319FFD0737332AAF9F7E224F9BF" box="[199,291,1613,1638]" italics="true" pageId="35" pageNumber="36">Tergites</emphasis>
: Paratergites only slightly projecting posteriorly (Fig. 17B). Tergites glossy, covered with few isolated longer hairs and very small setae.
</paragraph>
<paragraph id="8BE90F0BFFD0737332AAF921E3D1F8F1" blockId="35.[151,1437,1265,2026]" pageId="35" pageNumber="36">
<emphasis id="B922D319FFD0737332AAF921E250F96D" box="[199,343,1691,1716]" italics="true" pageId="35" pageNumber="36">Endotergum</emphasis>
: Inner section with short, rounded spines and few, isolated bristles. Between marginal ridge and inner area no cuticular impressions (Fig. 23F). Single row of marginal bristles, regularly distributed. Bristles short, reaching 1/3 up to 1/2 of space between innervating point of bristles and tergite margin (Fig. 23F).
</paragraph>
<paragraph id="8BE90F0BFFD0737332AAF88FE181F845" blockId="35.[151,1437,1265,2026]" pageId="35" pageNumber="36">
<emphasis id="B922D319FFD0737332AAF88FE24CF897" box="[199,331,1845,1870]" italics="true" pageId="35" pageNumber="36">Anal shield</emphasis>
: Sexually dimorphic. In females weakly bell-shaped, excavated at hindmost point and covered with some isolated, longer hairs. Underside carrying two black locking carinae, anterior one small, but well developed, posterior twice as long as first.
</paragraph>
<paragraph id="8BE90F0BFFD0737432AAF813E7FCFF6B" blockId="35.[151,1437,1265,2026]" lastBlockId="36.[151,1436,152,912]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<emphasis id="B922D319FFD0737332AAF813E3F9F81B" box="[199,254,1961,1986]" italics="true" pageId="35" pageNumber="36">Legs</emphasis>
: Tarsi of first and second pair of legs with four to six, third with six or seven long ventral spines. First two leg pairs with only weakly curved claws and without apical spine. Tarsi of legs 3–21 with curved claws, eight ventral spines and one apical spine. Femur 1.7 and tarsi 4.7 times longer than wide.
</paragraph>
<paragraph id="8BE90F0BFFD7737432AAFF05E736FF01" blockId="36.[151,1436,152,912]" box="[199,1073,190,216]" pageId="36" pageNumber="37">
<emphasis id="B922D319FFD7737432AAFF05E285FF01" box="[199,386,191,216]" italics="true" pageId="36" pageNumber="37">Stigmatic plates</emphasis>
: First lobe straight, with stout tip and only weakly curved.
</paragraph>
<paragraph id="8BE90F0BFFD7737432AAFF5FE2BEFEAA" blockId="36.[151,1436,152,912]" pageId="36" pageNumber="37">
<emphasis id="B922D319FFD7737432AAFF5FE2F2FF27" box="[199,501,229,254]" italics="true" pageId="36" pageNumber="37">Female sexual characters</emphasis>
: Washboard with 2 or 3 short, but broad and well-developed stridulation ribs on each half. Vulva large, covering more than 2/3 of coxa surface. Operculum small and low, ending far before apical coxal margin. Apical margin of operculum weakly notched (Fig. 21A). Mesal plate long, reaching up to 2/3 of operculum height.
</paragraph>
<paragraph id="8BE90F0BFFD7737432AAFE3AE3BAFE3E" blockId="36.[151,1436,152,912]" pageId="36" pageNumber="37">
<emphasis id="B922D319FFD7737432AAFE3AE2DEFE40" box="[199,473,384,409]" italics="true" pageId="36" pageNumber="37">Male sexual characters</emphasis>
: Anal shield without any invagination, almost rectangular with steep slope. Shield covered with dense field of small pits especially around hindmost point, single minute setae standing in each pit.
</paragraph>
<paragraph id="8BE90F0BFFD7737432AAFE4EE61BFD71" blockId="36.[151,1436,152,912]" pageId="36" pageNumber="37">
<emphasis id="B922D319FFD7737432AAFE4EE291FDD4" box="[199,406,500,525]" italics="true" pageId="36" pageNumber="37">Anterior telopods</emphasis>
(Figs 21B–D): First podomere broad, rectangular, width as large as height. Harp with two stridulation ribs. Mesal rib very weakly developed, hardly visible. Lateral rib short, but well-developed. Second podomere process lobe-like and little curved with rounded edges, reaching almost to half of third podomere height; apically with sclerotized spots. Third podomere cavity mesally with numerous sclerotized spots and three small spines, all of them located closely towards tip. Tip with one crenulated tooth.
</paragraph>
<paragraph id="8BE90F0BFFD7737432AAFD0FE76DFC49" blockId="36.[151,1436,152,912]" pageId="36" pageNumber="37">
<emphasis id="B922D319FFD7737432AAFD0FE2A4FD17" box="[199,419,693,718]" italics="true" pageId="36" pageNumber="37">Posterior telopods</emphasis>
(Figs 21E, F): Movable finger weakly curved; hollowed-out inner margin with two non-sclerotized lobes and
<emphasis id="B922D319FFD7737433B9FD66E108FD2C" box="[468,527,732,757]" italics="true" pageId="36" pageNumber="37">circa</emphasis>
three smaller, sclerotized spines, posterior aspect with
<emphasis id="B922D319FFD7737436D9FD66E7E8FD2C" box="[1204,1263,732,757]" italics="true" pageId="36" pageNumber="37">circa</emphasis>
25 crenulated teeth. Fixed finger thinner than movable finger, with slightly curved tip; basally with single membranous lobe, on inner margin covered with numerous sclerotized spots. Second and third podomere without any long hair, but on both sides almost completely covered densely with very small pits, one very small seta standing in each pit. First podomere without those pits but with some isolated, standard longer hairs.
</paragraph>
</subSubSection>
</treatment>
</document>
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