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<document id="DC964D0FC00748AC8B629683ECBA415D" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFFBC730232FAFAFCE622F88F" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium smaragdinum Wesener 2009, new species" docType="treatment" docUuid="0CA26D54-0E02-4A2F-B8E6-C3F166758A04" docUuidSource="ZooBank" docVersion="5" lastPageNumber="83" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="80" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="335B294CD29EE2D99A20DCA080871B6F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A193521EC27D0708E6A9F9CC28457856">
<mods:title id="135FAE24F5ABBA812BA63A3F90DE26A4">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
</mods:titleInfo>
<mods:name id="2900BEB23AA5B649B5EA778689F8242C" type="personal">
<mods:role id="27ACECC6D9FDC65706CBC9D1FB87B4DB">
<mods:roleTerm id="92640C638394272DED95ACFCCBD91B7B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BDCF6D7E8A933FF251C7E2B400F3E18D">Wesener, Thomas</mods:namePart>
</mods:name>
<mods:typeOfResource id="496BB81D6C7C074EB2F4655045204FBF">text</mods:typeOfResource>
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<mods:titleInfo id="C5646D1088F607DA36F3C5A0C2665B30">
<mods:title id="71B6675755F5AC570A49DCDF4930A0C9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5BEB14C869A1680F23670C94F75A6A91">
<mods:date id="5F9A35DB2189FC8F819468EC26405B6F">2009</mods:date>
<mods:detail id="5BCE5D6A7A145C841FC6864517148C13" type="pubDate">
<mods:number id="029331F69610322A76EFF3EB767134C0">2009-05-11</mods:number>
</mods:detail>
<mods:detail id="E523871681250C29325195925FF7AC2F" type="volume">
<mods:number id="80428CE7C4C5194E47F4F390192F5B45">2097</mods:number>
</mods:detail>
<mods:detail id="3391896700841C1702B5F3D4A82DC98C" type="issue">
<mods:number id="DDE24E0E37D8C7040E91D01BBFAEB299">1</mods:number>
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<mods:start id="E540A87D8F68BA0DBB3A7F7E6E9E26B2">1</mods:start>
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<mods:location id="786B2B3D8E5310F35505817218F8A4AE">
<mods:url id="F95C6C6F6D12BDCEFCAB47B568CDDE7B">https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1</mods:url>
</mods:location>
<mods:classification id="690EBD2F33D2AE63C4C23D64C0D72222">journal article</mods:classification>
<mods:identifier id="5FB1D023682BA85EFEA450627FEF63F6" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
<mods:identifier id="288A958FBEF364CAD0060351CB6E5BF6" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="4B55BBC5FFF3C5373192AAFBEC8201D9" type="Zenodo-Dep">5316536</mods:identifier>
<mods:identifier id="E5BDA222B8DC82342924307F750E35B9" type="ZooBank">0E18167A-78EB-4079-8BF4-972A4AC88C69</mods:identifier>
</mods:mods>
<treatment id="03FFBE1DFFBC730232FAFAFCE622F88F" ID-DOI="http://doi.org/10.5281/zenodo.5325499" ID-Zenodo-Dep="5325499" LSID="urn:lsid:zoobank.org:act:0CA26D54-0E02-4A2F-B8E6-C3F166758A04" httpUri="http://treatment.plazi.org/id/03FFBE1DFFBC730232FAFAFCE622F88F" lastPageId="82" lastPageNumber="83" pageId="79" pageNumber="80">
<subSubSection id="C34C5C80FFBC731F32FAFAFCE193FAB9" box="[151,660,1350,1376]" pageId="79" pageNumber="80" type="nomenclature">
<paragraph id="8BE90F0BFFBC731F32FAFAFCE193FAB9" blockId="79.[151,660,1350,1376]" box="[151,660,1350,1376]" pageId="79" pageNumber="80">
<heading id="D0A1B867FFBC731F32FAFAFCE193FAB9" bold="true" box="[151,660,1350,1376]" fontSize="11" level="1" pageId="79" pageNumber="80" reason="1">
<emphasis id="B922D319FFBC731F32FAFAFCE193FAB9" bold="true" box="[151,660,1350,1376]" pageId="79" pageNumber="80">
<emphasis id="B922D319FFBC731F32FAFAFCE2FBFAB9" bold="true" box="[151,508,1350,1376]" italics="true" pageId="79" pageNumber="80">
<taxonomicName id="4C567488FFBC731F32FAFAFCE2F1FAB9" ID-CoL="5D7J2" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,502,1350,1376]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaragdinum" status="sp. nov.">Zoosphaerium smaragdinum</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A2116E62FFBC731F3069FAFCE193FAB9" box="[516,660,1350,1376]" pageId="79" pageNumber="80" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFBC731F32FAFA35E10BFA09" pageId="79" pageNumber="80" type="etymology">
<paragraph id="8BE90F0BFFBC731F32FAFA35E7ADFA73" blockId="79.[151,1436,1423,2030]" box="[151,1194,1423,1450]" pageId="79" pageNumber="80">
<emphasis id="B922D319FFBC731F32FAFA35E287FA70" bold="true" box="[151,384,1423,1449]" pageId="79" pageNumber="80">Derivatio nominis:</emphasis>
<taxonomicName id="4C567488FFBC731F33E5FA2AE12EFA70" authorityName="Wesener" authorityYear="2009" box="[392,553,1424,1449]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaragdinum">
<emphasis id="B922D319FFBC731F33E5FA2AE12EFA70" box="[392,553,1424,1449]" italics="true" pageId="79" pageNumber="80">smaragdinum</emphasis>
</taxonomicName>
, adjective, refers to the shiny green coloration pattern.
</paragraph>
<paragraph id="8BE90F0BFFBC731F32ABFA0CE10BFA09" blockId="79.[151,1436,1423,2030]" box="[198,524,1462,1488]" pageId="79" pageNumber="80">
<emphasis id="B922D319FFBC731F32ABFA0CE2A7FA09" bold="true" box="[198,416,1462,1488]" pageId="79" pageNumber="80">Studied material:</emphasis>
1 M, 1 F
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFBC731F32AAFA67E055F9B2" pageId="79" pageNumber="80" type="materials_examined">
<paragraph id="8BE90F0BFFBC731F32AAFA67E1DCF99C" blockId="79.[151,1436,1423,2030]" pageId="79" pageNumber="80">
<materialsCitation id="3B3E0556FFBC731F32AAFA67E1DCF99C" collectingDate="2003-11-18" collectionCode="CAS" collectorName="B. L. Fisher" county="Province" elevation="1325" latitude="-14.443334" location="Antranohofa" longLatPrecision="21" longitude="49.743332" municipality="Parc National de Marojejy" pageId="79" pageNumber="80" specimenCode="BLF 9079" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
<emphasis id="B922D319FFBC731F32AAFA67E245FA2E" bold="true" box="[199,322,1501,1527]" pageId="79" pageNumber="80">
<typeStatus id="54EDB1A9FFBC731F32AAFA67E23AFA2E" box="[199,317,1501,1527]" pageId="79" pageNumber="80" type="holotype">Holotype</typeStatus>
:
</emphasis>
1 M (
<quantity id="4CAEA2EEFFBC731F33F0FA67E2F0FA2E" box="[413,503,1501,1527]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="79" pageNumber="80" unit="mm" value="30.0">30 mm</quantity>
long),
<emphasis id="B922D319FFBC731F303AFA67E027FA2E" bold="true" box="[599,800,1501,1527]" pageId="79" pageNumber="80">
<collectionCode id="ED4797CEFFBC731F303AFA67E196FA2E" box="[599,657,1501,1527]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="79" pageNumber="80" type="Museum">CAS</collectionCode>
<specimenCode id="DBF0A770FFBC731F30F2FA67E027FA2E" box="[671,800,1501,1527]" collectionCode="BLF" pageId="79" pageNumber="80">BLF 9079</specimenCode>
</emphasis>
,
<collectingCounty id="62887787FFBC731F3158FA67E09AFA2E" box="[821,925,1501,1527]" pageId="79" pageNumber="80">Province</collectingCounty>
<collectingRegion id="4992C1E9FFBC731F31C6FA67E73BFA2E" box="[939,1084,1501,1527]" country="Madagascar" name="Antsiranana" pageId="79" pageNumber="80">Antsiranana</collectingRegion>
,
<collectingMunicipality id="6B8D9571FFBC731F3620FA67E69FFA2E" box="[1101,1432,1501,1527]" pageId="79" pageNumber="80">Parc National de Marojejy</collectingMunicipality>
,
<location id="8E8959D0FFBC731F32FAF9BEE229F9C7" LSID="urn:lsid:plazi:treatment:03FFBE1DFFBC730232FAFAFCE622F88F:8E8959D0FFBC731F32FAF9BEE229F9C7" box="[151,302,1540,1566]" county="Province" latitude="-14.443334" longLatPrecision="21" longitude="49.743332" municipality="Parc National de Marojejy" name="Antranohofa" pageId="79" pageNumber="80" stateProvince="Antsiranana">Antranohofa</location>
,
<locationDeviation id="9A5B1629FFBC731F3357F9BEE175F9C7" box="[314,626,1540,1566]" location="Andapa" pageId="79" pageNumber="80">
<quantity id="4CAEA2EEFFBC731F3357F9BEE29EF9C7" box="[314,409,1540,1566]" metricMagnitude="4" metricUnit="m" metricValue="2.66" pageId="79" pageNumber="80" unit="km" value="26.6">26.6 km</quantity>
31° NNE Andapa
</locationDeviation>
,
<locationDeviation id="9A5B1629FFBC731F3013F9BEE0FEF9C7" box="[638,1017,1540,1566]" location="Manantenina" pageId="79" pageNumber="80">
<quantity id="4CAEA2EEFFBC731F3013F9BEE1D9F9C7" box="[638,734,1540,1566]" metricMagnitude="4" metricUnit="m" metricValue="1.0699999999999998" pageId="79" pageNumber="80" unit="km" value="10.7">10.7 km</quantity>
318° NW Manantenina
</locationDeviation>
,
<geoCoordinate id="EE6269CCFFBC731F3669F9BEE78CF9C7" box="[1028,1163,1540,1566]" degrees="14" direction="south" minutes="26" orientation="latitude" pageId="79" pageNumber="80" precision="15" seconds="36" value="-14.443334">14°26'36&quot;S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFFBC731F36F5F9BEE628F9C7" box="[1176,1327,1540,1566]" degrees="049" direction="east" minutes="44" orientation="longitude" pageId="79" pageNumber="80" precision="15" seconds="36" value="49.743332">049°44'36&quot;E</geoCoordinate>
,
<quantity id="4CAEA2EEFFBC731F3756F9BEE691F9C7" box="[1339,1430,1540,1566]" metricMagnitude="3" metricUnit="m" metricValue="1.325" pageId="79" pageNumber="80" unit="m" value="1325.0">
<elevation id="007BE838FFBC731F3756F9BEE691F9C7" box="[1339,1430,1540,1566]" metricMagnitude="3" metricUnit="m" metricValue="1.325" pageId="79" pageNumber="80" unit="m" value="1325.0">1325 m</elevation>
</quantity>
, montane rainforest, coll.
<collectorName id="26A36ADDFFBC731F33D1F991E14DF99C" box="[444,586,1579,1605]" pageId="79" pageNumber="80">B. L. Fisher</collectorName>
,
<date id="FFE829CBFFBC731F303BF991E1D1F99C" box="[598,726,1579,1605]" pageId="79" pageNumber="80" value="2003-11-18">
<collectingDate id="EFACD023FFBC731F303BF991E1D1F99C" box="[598,726,1579,1605]" pageId="79" pageNumber="80" value="2003-11-18">18.xi.2003</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE90F0BFFBC731F32AAF9EBE055F9B2" blockId="79.[151,1436,1423,2030]" box="[199,850,1617,1643]" pageId="79" pageNumber="80">
<materialsCitation id="3B3E0556FFBC731F32AAF9EBE04AF9B2" box="[199,845,1617,1643]" collectingDate="2003-11-18" collectionCode="CAS" collectorName="B. L. Fisher" county="Province" elevation="1325" latitude="-14.443334" location="Antranohofa" longLatPrecision="21" longitude="49.743332" municipality="Parc National de Marojejy" pageId="79" pageNumber="80" specimenCode="BLF 9079" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<emphasis id="B922D319FFBC731F32AAF9EBE245F9B2" bold="true" box="[199,322,1617,1643]" pageId="79" pageNumber="80">
<typeStatus id="54EDB1A9FFBC731F32AAF9EBE23AF9B2" box="[199,317,1617,1643]" pageId="79" pageNumber="80" type="paratype">Paratype</typeStatus>
:
</emphasis>
1 F,
<emphasis id="B922D319FFBC731F3311F9EBE13FF9B2" bold="true" box="[380,568,1617,1643]" pageId="79" pageNumber="80">
<collectionCode id="ED4797CEFFBC731F3311F9EBE2B1F9B2" box="[380,438,1617,1643]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="79" pageNumber="80" type="Museum">CAS</collectionCode>
<specimenCode id="DBF0A770FFBC731F33D3F9EBE13FF9B2" box="[446,568,1617,1643]" collectionCode="BLF" pageId="79" pageNumber="80">BLF 9079</specimenCode>
</emphasis>
, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFBC731F32ABF9CDE3C2F961" pageId="79" pageNumber="80" type="distribution">
<paragraph id="8BE90F0BFFBC731F32ABF9CDE3C2F961" blockId="79.[151,1436,1423,2030]" pageId="79" pageNumber="80">
<emphasis id="B922D319FFBC731F32ABF9CDE2F4F948" bold="true" box="[198,499,1655,1681]" pageId="79" pageNumber="80">Distribution &amp; Ecology:</emphasis>
Recorded from the montane rainforest of Marojejy from
<quantity id="4CAEA2EEFFBC731F36FFF9C2E7EDF94B" box="[1170,1258,1656,1682]" metricMagnitude="3" metricUnit="m" metricValue="1.325" pageId="79" pageNumber="80" unit="m" value="1325.0">1325 m</quantity>
elevation (Fig. 50).
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFBC730132ABF97FE2D5F8AE" lastPageId="81" lastPageNumber="82" pageId="79" pageNumber="80" type="diagnosis">
<paragraph id="8BE90F0BFFBC730032ABF97FE0A5FEFF" blockId="79.[151,1436,1423,2030]" lastBlockId="80.[151,1436,152,410]" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
<emphasis id="B922D319FFBC731F32ABF97FE24FF906" bold="true" box="[198,328,1733,1759]" pageId="79" pageNumber="80">Diagnosis:</emphasis>
Up to
<quantity id="4CAEA2EEFFBC731F33F6F97FE2EAF906" box="[411,493,1733,1759]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="79" pageNumber="80" unit="mm" value="30.0">30 mm</quantity>
long. Color faded in alcohol, best conserved specimens green, with light brown thoracic and anal shield. Head, legs and antennae also green. Texture of tergites orange-peel like, anterior part with some short hairs. Anal shield well-rounded, in males covered densely with numerous hairs. Disc of antenna with 10–15 apical cones (Fig. 55B). Tarsus of leg pairs 1–3 without apical spine, tarsi of legs 4–21 with only up to eight ventral spines. Movable finger of posterior telopods slender, with well-rounded tip, one triangular lobe and three sclerotized spines (Fig. 54D), posterior side with 10 sclerotized teeth (Fig. 54E). Fixed finger long and slender, weakly curved, basally with one small spine, distally at inner margin with numerous sclerotized spots (Fig. 54D). Fixed finger almost as long as movable finger. Two strongly developed stridulation ribs on each male harp (Fig 54B). Anal shield with two locking carinae, anterior small, posterior&gt;3 times longer than first. Female with two stridulation ribs on each side of subanal plate. Operculum of vulva protruding into two well-rounded, same sized tips, strongly extending beyond apical coxal margin (Fig. 54A). Lateral tip slightly sharper than inner tip.
</paragraph>
<paragraph id="8BE90F0BFFA3730032ABFE88E02BFE43" blockId="80.[151,1436,152,410]" pageId="80" pageNumber="81">
<emphasis id="B922D319FFA3730032ABFE88E28CFE95" bold="true" box="[198,395,306,332]" pageId="80" pageNumber="81">Similar species:</emphasis>
In no other species of the group is the posterior telopod movable finger as slender as in
<emphasis id="B922D319FFA3730032FAFEE3E25AFEAB" box="[151,349,345,370]" italics="true" pageId="80" pageNumber="81">
<taxonomicName id="4C567488FFA3730032FAFEE3E25EFEAB" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,345,345,370]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="smaragdinum" status="sp. nov.">Z. smaragdinum</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FFA373003309FEE2E2A1FEAB" bold="true" box="[356,422,344,370]" pageId="80" pageNumber="81">
<taxonomicNameLabel id="A2116E62FFA373003309FEE2E2A1FEAB" box="[356,422,344,370]" pageId="80" pageNumber="81" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
. The telopods of
<emphasis id="B922D319FFA37300301FFEE3E03FFEAB" box="[626,824,345,370]" italics="true" pageId="80" pageNumber="81">
<taxonomicName id="4C567488FFA37300301FFEE3E033FEAB" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[626,820,345,370]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="smaragdinum" status="sp. nov.">Z. smaragdinum</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FFA373003152FEE2E086FEAB" bold="true" box="[831,897,344,370]" pageId="80" pageNumber="81">
<taxonomicNameLabel id="A2116E62FFA373003152FEE2E086FEAB" box="[831,897,344,370]" pageId="80" pageNumber="81" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
feature in contrast to the coloration pattern no similarity to the sympatric living
<emphasis id="B922D319FFA373003073FE3AE1DCFE40" box="[542,731,384,409]" italics="true" pageId="80" pageNumber="81">
<taxonomicName id="4C567488FFA373003073FE3AE1D0FE40" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[542,727,384,409]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="viridissimum" status="sp. nov.">Z. viridissimum</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FFA37300308EFEC5E022FE40" bold="true" box="[739,805,383,409]" pageId="80" pageNumber="81">
<taxonomicNameLabel id="A2116E62FFA37300308EFEC5E022FE40" box="[739,805,383,409]" pageId="80" pageNumber="81" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
<paragraph id="8BE90F0BFFA3730032FAF9C9E3F6F8E1" blockId="80.[151,1436,1651,1848]" pageId="80" pageNumber="81">
<emphasis id="B922D319FFA3730032FAF9C9E229F952" bold="true" box="[151,302,1651,1675]" pageId="80" pageNumber="81">FIGURE 054</emphasis>
.
<taxonomicName id="4C567488FFA373003356F9CEE177F952" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[315,624,1652,1675]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="smaragdinum" status="sp. nov.">
<emphasis id="B922D319FFA373003356F9CEE177F952" box="[315,624,1652,1675]" italics="true" pageId="80" pageNumber="81">Zoosphaerium smaragdinum</emphasis>
</taxonomicName>
<emphasis id="B922D319FFA37300301BF9C9E1B5F952" bold="true" box="[630,690,1651,1675]" pageId="80" pageNumber="81">
<taxonomicNameLabel id="A2116E62FFA37300301BF9C9E1B5F952" box="[630,690,1651,1675]" pageId="80" pageNumber="81" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, A female
<typeStatus id="54EDB1A9FFA373003146F9CEE08EF952" box="[811,905,1652,1675]" pageId="80" pageNumber="81" type="paratype">paratype</typeStatus>
, B–E
<typeStatus id="54EDB1A9FFA3730031A6F9CEE72CF952" box="[971,1067,1652,1675]" pageId="80" pageNumber="81" type="holotype">holotype</typeStatus>
,
<emphasis id="B922D319FFA373003658F9C9E74FF952" bold="true" box="[1077,1096,1651,1675]" pageId="80" pageNumber="81">A</emphasis>
: second left coxa and prefemur with vulva, posterior view;
<emphasis id="B922D319FFA3730033AFF92CE2D3F977" bold="true" box="[450,468,1686,1710]" pageId="80" pageNumber="81">B</emphasis>
: right half of anterior telopods, anterior view;
<emphasis id="B922D319FFA3730031A2F92CE0E5F977" bold="true" box="[975,994,1686,1710]" pageId="80" pageNumber="81">C</emphasis>
: last two podomeres of anterior telopod, lateral view;
<emphasis id="B922D319FFA373003344F903E23BF908" bold="true" box="[297,316,1721,1745]" pageId="80" pageNumber="81">D</emphasis>
: posterior telopod, anterior view;
<emphasis id="B922D319FFA3730030D4F903E1CCF908" bold="true" box="[697,715,1721,1745]" pageId="80" pageNumber="81">E</emphasis>
: chela of posterior telopod, posterior view. Abbreviations:
<emphasis id="B922D319FFA373003731F903E684F908" bold="true" box="[1372,1411,1721,1745]" pageId="80" pageNumber="81">cav</emphasis>
= cavity;
<emphasis id="B922D319FFA37300328AF961E217F92A" bold="true" box="[231,272,1755,1779]" pageId="80" pageNumber="81">cr-t</emphasis>
= crenulated teeth;
<emphasis id="B922D319FFA373003387F961E10CF92A" bold="true" box="[490,523,1755,1779]" pageId="80" pageNumber="81">Cx</emphasis>
= coxa;
<emphasis id="B922D319FFA373003001F961E189F92A" bold="true" box="[620,654,1755,1779]" pageId="80" pageNumber="81">EP</emphasis>
= external plate of vulva;
<emphasis id="B922D319FFA3730031DDF961E0CDF92A" bold="true" box="[944,970,1755,1779]" pageId="80" pageNumber="81">IP</emphasis>
= mesal plate of vulva;
<emphasis id="B922D319FFA3730036B8F961E7F4F92A" bold="true" box="[1237,1267,1755,1779]" pageId="80" pageNumber="81">ml</emphasis>
= membranous lobe;
<emphasis id="B922D319FFA3730032BEF944E3EFF8CF" bold="true" box="[211,232,1790,1814]" pageId="80" pageNumber="81">O</emphasis>
= operculum;
<emphasis id="B922D319FFA3730033EBF944E2AAF8CF" bold="true" box="[390,429,1790,1814]" pageId="80" pageNumber="81">Pre</emphasis>
= prefemur;
<emphasis id="B922D319FFA373003056F944E15AF8CF" bold="true" box="[571,605,1790,1814]" pageId="80" pageNumber="81">s-p</emphasis>
= sclerotized spot;
<emphasis id="B922D319FFA373003142F944E058F8CF" bold="true" box="[815,863,1790,1814]" pageId="80" pageNumber="81">scl-t</emphasis>
= sclerotized teeth;
<emphasis id="B922D319FFA373003654F944E75CF8CF" bold="true" box="[1081,1115,1790,1814]" pageId="80" pageNumber="81">SR</emphasis>
= stridulation ribs. Scale bars =
<quantity id="4CAEA2EEFFA3730032C0F89BE3EAF8E1" box="[173,237,1825,1848]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="80" pageNumber="81" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFFA2730132FAF8A1E2D5F8AE" blockId="81.[151,1435,1819,1912]" pageId="81" pageNumber="82">
<emphasis id="B922D319FFA2730132FAF8A1E229F8EA" bold="true" box="[151,302,1819,1843]" pageId="81" pageNumber="82">FIGURE 055</emphasis>
.
<taxonomicName id="4C567488FFA273013351F8A1E177F8EB" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[316,624,1819,1842]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="smaragdinum" status="sp. nov.">
<emphasis id="B922D319FFA273013351F8A1E177F8EB" box="[316,624,1819,1842]" italics="true" pageId="81" pageNumber="82">Zoosphaerium smaragdinum</emphasis>
</taxonomicName>
<emphasis id="B922D319FFA27301301BF8A1E1B4F8EA" bold="true" box="[630,691,1819,1843]" pageId="81" pageNumber="82">
<taxonomicNameLabel id="A2116E62FFA27301301BF8A1E1B4F8EA" box="[630,691,1819,1843]" pageId="81" pageNumber="82" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FFA2730130ACF8A1E027F8EB" box="[705,800,1819,1842]" pageId="81" pageNumber="82" type="paratype">paratype</typeStatus>
, SEM,
<emphasis id="B922D319FFA273013102F8A1E085F8EA" bold="true" box="[879,898,1819,1843]" pageId="81" pageNumber="82">A</emphasis>
: antenna, lateral view;
<emphasis id="B922D319FFA273013613F8A1E797F8EA" bold="true" box="[1150,1168,1819,1843]" pageId="81" pageNumber="82">B</emphasis>
: antenna, apical view on disc;
<emphasis id="B922D319FFA2730132BDF887E3E4F88C" bold="true" box="[208,227,1853,1877]" pageId="81" pageNumber="82">C</emphasis>
: right mandible, mesal view. Abbreviations:
<emphasis id="B922D319FFA2730130AEF887E1EEF88C" bold="true" box="[707,745,1853,1877]" pageId="81" pageNumber="82">3iT</emphasis>
= 3 inner teeth;
<emphasis id="B922D319FFA2730131F5F887E0BFF88C" bold="true" box="[920,952,1853,1877]" pageId="81" pageNumber="82">Co</emphasis>
= condylus;
<emphasis id="B922D319FFA27301362FF887E767F88C" bold="true" box="[1090,1120,1853,1877]" pageId="81" pageNumber="82">eT</emphasis>
= external tooth;
<emphasis id="B922D319FFA273013770F887E645F88C" bold="true" box="[1309,1346,1853,1877]" pageId="81" pageNumber="82">mp</emphasis>
= molar plate;
<emphasis id="B922D319FFA2730132B4F8DAE3FEF8A1" bold="true" box="[217,249,1888,1912]" pageId="81" pageNumber="82">pL</emphasis>
= pectinate lamella.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFA2730232ABF81FE622F88F" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82" type="description">
<paragraph id="8BE90F0BFFA2730132ABF81FE159F83F" blockId="81.[151,1435,1957,2022]" pageId="81" pageNumber="82">
<emphasis id="B922D319FFA2730132ABF81FE267F866" bold="true" box="[198,352,1957,1983]" pageId="81" pageNumber="82">Description:</emphasis>
<emphasis id="B922D319FFA273013304F81CE2F1F866" box="[361,502,1958,1983]" italics="true" pageId="81" pageNumber="82">Body length</emphasis>
(
<typeStatus id="54EDB1A9FFA27301306AF81CE168F819" box="[519,623,1958,1984]" pageId="81" pageNumber="82" type="holotype">holotype</typeStatus>
only): Length up to
<quantity id="4CAEA2EEFFA273013108F81CE0BCF819" box="[869,955,1958,1984]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="81" pageNumber="82" unit="mm" value="30.0">30 mm</quantity>
, width of thoracic shield up to
<quantity id="4CAEA2EEFFA273013746F81CE69FF819" box="[1323,1432,1958,1984]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="81" pageNumber="82" unit="mm" value="14.5">14.5 mm</quantity>
, height of thoracic shield up to
<quantity id="4CAEA2EEFFA273013391F876E15EF83F" box="[508,601,1996,2022]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="81" pageNumber="82" unit="mm" value="7.1">7.1 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAFF22E2DBFEFF" blockId="82.[151,1436,152,1878]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAFF22E256FF68" box="[199,337,152,177]" italics="true" pageId="82" pageNumber="83">Coloration:</emphasis>
Possibly faded in alcohol. Quite different in male specimen in comparison to much smaller female. Both specimens with jade-like greenish colored legs, head, collum, antenna and posterior margin of tergites. Medial part of the thoracic shield and anal shield light brown. In larger male
<typeStatus id="54EDB1A9FFA173023614FF5FE7E5FF26" box="[1145,1250,229,255]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
, anterior part of tergites also light brownish.
</paragraph>
<paragraph id="8BE90F0BFFA1730232FAFE88E2D8FE95" blockId="82.[151,1436,152,1878]" box="[151,479,306,332]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232FAFE88E3D1FE92" box="[151,214,306,331]" italics="true" pageId="82" pageNumber="83">Head</emphasis>
: Eyes with&gt;60 ocelli.
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAFEE2E15BFE66" blockId="82.[151,1436,152,1878]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAFEE2E231FEA8" box="[199,310,344,369]" italics="true" pageId="82" pageNumber="83">Antennae</emphasis>
: Length of antennomeres: 1&gt;2=3&gt;4=5&lt;6 (Fig. 55A), sixth antennomere bearing disc with 11–14 sensory cones (Fig. 55B). First antennomere remarkably broader than others, with groove. Sclerotized teeth on antennomeres 1–4 (Fig. 55A).
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAFE76E21FFDEA" blockId="82.[151,1436,152,1878]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAFE76E230FE3C" box="[199,311,460,485]" italics="true" pageId="82" pageNumber="83">Mandible</emphasis>
: General appearance unusually slender (Fig. 55C). 5–6 rows of pectinate lamellae, number of teeth declining apically to proximally (Fig. 55C). Condylus with strongly developed step near apical margin (Fig. 55C).
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAFDFAE2E2FD59" blockId="82.[151,1436,152,1878]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAFDFAE296FD80" box="[199,401,576,601]" italics="true" pageId="82" pageNumber="83">Gnathochilarium</emphasis>
: Laterally of palpi four sensory cones, located together. Depression on posterior side of palpi without sensory cones.
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAFD37E0A6FD7E" blockId="82.[151,1436,152,1878]" box="[199,929,653,679]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAFD37E21AFD7F" box="[199,285,653,678]" italics="true" pageId="82" pageNumber="83">Collum</emphasis>
: Median part of collum with some long, isolated hairs.
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAFD0EE062FD17" blockId="82.[151,1436,152,1878]" box="[199,869,692,718]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAFD0EE279FD14" box="[199,382,692,717]" italics="true" pageId="82" pageNumber="83">Thoracic shield</emphasis>
shaped like in other species of the genus.
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAFD60E1C0FD2D" blockId="82.[151,1436,152,1878]" box="[199,711,730,756]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAFD60E224FD2A" box="[199,291,730,755]" italics="true" pageId="82" pageNumber="83">Tergites</emphasis>
: Paratergites projecting posteriorly.
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAFCBBE2C3FCB1" blockId="82.[151,1436,152,1878]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAFCBBE267FCC3" box="[199,352,769,794]" italics="true" pageId="82" pageNumber="83">Endotergum:</emphasis>
Inner area with thin conical spines and very long hair. Single row of large, circular cuticular impression, regularly distributed (Fig. 63C). One row of marginal bristles, reaching up to 1/2 of space towards tergite margin (Fig. 63C).
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAFCCFE61BFC05" blockId="82.[151,1436,152,1878]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAFCCFE254FC57" box="[199,339,885,910]" italics="true" pageId="82" pageNumber="83">Anal shield</emphasis>
: Well-rounded, neither bell-shaped nor tapered. In female without any hair. Underside carrying two well-developed black locking carinae, anterior one small, but well developed, projecting posteriorly strongly towards anal shield margin. Posterior carina&gt;3 times longer than anterior one.
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAFC53E69BFB89" blockId="82.[151,1436,152,1878]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAFC53E3F9FBDB" box="[199,254,1001,1026]" italics="true" pageId="82" pageNumber="83">Legs</emphasis>
: Tarsi of first pair of legs with four, second with four or five, third pair with six or seven long ventral spines. First three leg pairs with only weakly curved claws and without apical spine. Tarsi of legs 4–21 with curved claws, 8 ventral spines and an apical spine. In 9th leg femur 1.9 times, tarsi 4.9 times longer than wide.
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAFBE7E013FBAE" blockId="82.[151,1436,152,1878]" box="[199,788,1117,1143]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAFBE7E271FBAF" box="[199,374,1117,1142]" italics="true" pageId="82" pageNumber="83">Stigmatic plate</emphasis>
: First lobe short and well-rounded.
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAFB3EE783FA86" blockId="82.[151,1436,152,1878]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAFB3EE2FFFB44" box="[199,504,1156,1181]" italics="true" pageId="82" pageNumber="83">Female sexual characters</emphasis>
: Subanal plate with washboard, consisting of well-developed stridulation ribs, two ribs on each side. Stridulation ribs symmetrical, of medium length. Vulva large, covering slightly more than 2/3 of coxa. Operculum large, surmounting coxa up to basal 1/3 of prefemur (Fig. 54A). Apical margin of operculum medially notched, with two rounded lateral tips. Both tips of same height, space between them deeply constricted, dividing operculum in two almost same-sized parts. Lateral tip slightly sharper than inner tip (Fig. 54A). Mesal plate long and broad, protruding up to half of operculum height.
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAFAD6E712FA5F" blockId="82.[151,1436,152,1878]" box="[199,1045,1388,1414]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAFAD6E2E3FA5C" box="[199,484,1388,1413]" italics="true" pageId="82" pageNumber="83">Male sexual characters:</emphasis>
Anal shield covered with numerous short hairs.
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAFA29E0A2F9B7" blockId="82.[151,1436,152,1878]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAFA29E299FA75" box="[199,414,1427,1452]" italics="true" pageId="82" pageNumber="83">Anterior telopods</emphasis>
(Figs 54B, C): Syncoxite and first podomere almost glabrous. Harp with two stridulation ribs, straight, broad and well-developed. Second podomere process lobe-like and little curved with rounded edges, reaching 1/3 of third podomeres height, apically with sclerotized spots. Third podomere cavity mesally with several sclerotized spots and four sclerotized spines laterally basally ascending, with four sclerotized, crenulated teeth. Apical margin of third podomere well-rounded, one sclerotized spot located closely to tip. Second and third podomere on both sides with hairs.
</paragraph>
<paragraph id="8BE90F0BFFA1730232AAF9C1E622F88F" blockId="82.[151,1436,152,1878]" pageId="82" pageNumber="83">
<emphasis id="B922D319FFA1730232AAF9C1E2A7F94D" box="[199,416,1659,1684]" italics="true" pageId="82" pageNumber="83">Posterior telopods</emphasis>
(Figs 54D, E): Movable finger of chela relatively slender, 2.5 times longer than wide, weakly curved, slightly tapering apically. Hollowed-out inner margin with single triangular membranous lobe and three sclerotized spines; posterior aspect with
<emphasis id="B922D319FFA17302309FF972E02AF938" box="[754,813,1736,1761]" italics="true" pageId="82" pageNumber="83">circa</emphasis>
10 or 11 small, sclerotized teeth. Fixed finger more slender and slightly shorter than movable one, tip of former only weakly curved, anterior side basally with one short sclerotized spine, on tip with numerous sclerotized spots. Second and third podomere covered on both sides with numerous long hairs. Inner horns apically curved. Tips pointed, covered with short hairs.
</paragraph>
</subSubSection>
</treatment>
</document>
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