397 lines
41 KiB
XML
397 lines
41 KiB
XML
<document id="2A007D091ABDB08B6893D3B404D1951D" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFFB4731832FAFAF4E69CF805" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium coquerelianum" docType="key" docVersion="4" lastPageNumber="73" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="72" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="81C63780A863B6E5040AB85C90EF0082">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
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<mods:namePart id="0154F16CB04BC94ADE8DB6D17698EC48">Wesener, Thomas</mods:namePart>
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<mods:title id="F5F5B211AC4E62F0B1D3FD97B2D2D14E">Zootaxa</mods:title>
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<mods:part id="F6E3BF17CE7A7056D5BD123B082D40E6">
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<mods:date id="E0FBE5172142130C951BDFC39A0C5F69">2009</mods:date>
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<mods:number id="95988B3B64703D74AAEFE442C233F7F7">2009-05-11</mods:number>
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<mods:number id="CF1B670E124DEECEC08D34401949009A">2097</mods:number>
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<mods:classification id="1D4528306E7DED9C3B3D458CCB6B2E74">journal article</mods:classification>
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<mods:identifier id="C210A0F68FAE4857FF8E8D803284D898" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
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<treatment id="03FFBE1DFFB4731832FAFAF4E69CF805" ID-DOI="http://doi.org/10.5281/zenodo.5325485" ID-Zenodo-Dep="5325485" LSID="urn:lsid:plazi:treatment:03FFBE1DFFB4731832FAFAF4E69CF805" httpUri="http://treatment.plazi.org/id/03FFBE1DFFB4731832FAFAF4E69CF805" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
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<subSubSection id="C34C5C80FFB4731732FAFAF4E023FAB1" box="[151,804,1358,1384]" pageId="71" pageNumber="72" type="nomenclature">
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<paragraph id="8BE90F0BFFB4731732FAFAF4E023FAB1" blockId="71.[151,804,1358,1384]" box="[151,804,1358,1384]" pageId="71" pageNumber="72">
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<heading id="D0A1B867FFB4731732FAFAF4E023FAB1" bold="true" box="[151,804,1358,1384]" fontSize="11" level="1" pageId="71" pageNumber="72" reason="1">
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<emphasis id="B922D319FFB4731732FAFAF4E023FAB1" bold="true" box="[151,804,1358,1384]" pageId="71" pageNumber="72">
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Key to species of the
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<taxonomicName id="4C567488FFB4731733F1FAF4E174FAB1" ID-CoL="5D7GQ" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[412,627,1358,1384]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="coquerelianum">
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<emphasis id="B922D319FFB4731733F1FAF4E174FAB1" bold="true" box="[412,627,1358,1384]" italics="true" pageId="71" pageNumber="72">Z. coquerelianum</emphasis>
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</taxonomicName>
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species-group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34C5C80FFB4731832FAFA34E69CF805" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72" type="key">
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<keyStep id="30A217AEFFB4731732FAFA34E69CFA10" pageId="71" pageNumber="72">
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<paragraph id="8BE90F0BFFB4731732FAFA34E69CFA7F" blockId="71.[151,1437,1422,2035]" box="[151,1435,1422,1446]" pageId="71" pageNumber="72">
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<keyLead id="30A7AC3EFFB4731732FAFA34E69CFA7F" box="[151,1435,1422,1446]" pageId="71" pageNumber="72">
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<emphasis id="B922D319FFB4731732FAFA34E3A3FA7F" bold="true" box="[151,164,1422,1446]" pageId="71" pageNumber="72">1</emphasis>
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. Male harp with two or three stridulation ribs...............................................................................................................
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<emphasis id="B922D319FFB4731737E3FA34E69CFA7F" bold="true" box="[1422,1435,1422,1446]" pageId="71" pageNumber="72">2</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="8BE90F0BFFB4731732FAFA0BE69CFA10" blockId="71.[151,1437,1422,2035]" box="[151,1435,1457,1481]" pageId="71" pageNumber="72">
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<keyLead id="30A7AC3EFFB4731732FAFA0BE69CFA10" box="[151,1435,1457,1481]" pageId="71" pageNumber="72">
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- Male harp with single stridulation rib. .........................................................................................................................
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<emphasis id="B922D319FFB4731737E3FA0BE69CFA10" bold="true" box="[1422,1435,1457,1481]" pageId="71" pageNumber="72">7</emphasis>
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</keyLead>
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</paragraph>
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||
</keyStep>
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<keyStep id="30A217AEFFB4731732FAFA69E69CF9AF" pageId="71" pageNumber="72">
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<paragraph id="8BE90F0BFFB4731732FAFA69E69CF9E8" blockId="71.[151,1437,1422,2035]" pageId="71" pageNumber="72">
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<keyLead id="30A7AC3EFFB4731732FAFA69E69CF9E8" pageId="71" pageNumber="72">
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<emphasis id="B922D319FFB4731732FAFA69E3A3FA32" bold="true" box="[151,164,1491,1515]" pageId="71" pageNumber="72">2</emphasis>
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. Tergites glossy, anteriorly shiny light-green, posteriorly dark green. Specimen larger than
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<quantity id="4CAEA2EEFFB4731736F4FA6EE7EEFA32" box="[1177,1257,1492,1515]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="71" pageNumber="72" unit="mm" value="50.0">50 mm</quantity>
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. Anal shield not well-rounded, with steep slope. Movable finger of posterior telopods apically with groove (Fig. 49E), finger 1.6 times longer than wide (Fig. 49D).
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<emphasis id="B922D319FFB47317302EF9A3E65EF9E9" box="[579,1369,1561,1584]" italics="true" pageId="71" pageNumber="72">
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....................................................................................................
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<taxonomicName id="4C567488FFB47317368DF9A3E653F9E9" ID-CoL="5D7GB" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1248,1364,1561,1584]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="amabile" status="sp. nov.">Z. amabile</taxonomicName>
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,
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</emphasis>
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<emphasis id="B922D319FFB47317370DF9A3E69CF9E8" bold="true" box="[1376,1435,1561,1585]" pageId="71" pageNumber="72">
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<taxonomicNameLabel id="A2116E62FFB47317370DF9A3E69CF9E8" box="[1376,1435,1561,1585]" pageId="71" pageNumber="72" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="8BE90F0BFFB4731732FAF986E69CF9AF" blockId="71.[151,1437,1422,2035]" pageId="71" pageNumber="72">
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<keyLead id="30A7AC3EFFB4731732FAF986E69CF9AF" pageId="71" pageNumber="72">
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- Coloration pattern different, or if similar then specimens smaller than
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<quantity id="4CAEA2EEFFB4731731A3F986E727F98A" box="[974,1056,1596,1619]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="71" pageNumber="72" unit="mm" value="35.0">35 mm</quantity>
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. Anal shield well-rounded or bellshaped. Movable finger without apical groove, podomere 2.0–2.5 times longer than wide. .....................................
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<emphasis id="B922D319FFB4731737E3F9E4E69CF9AF" bold="true" box="[1422,1435,1630,1654]" pageId="71" pageNumber="72">3</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30A217AEFFB4731732FAF93BE69CF962" pageId="71" pageNumber="72">
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<paragraph id="8BE90F0BFFB4731732FAF93BE69CF940" blockId="71.[151,1437,1422,2035]" box="[151,1435,1665,1689]" pageId="71" pageNumber="72">
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<keyLead id="30A7AC3EFFB4731732FAF93BE69CF940" box="[151,1435,1665,1689]" pageId="71" pageNumber="72">
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<emphasis id="B922D319FFB4731732FAF93BE3A3F940" bold="true" box="[151,164,1665,1689]" pageId="71" pageNumber="72">3</emphasis>
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. Antenna with four apical cones. ..................................................................................................................................
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<emphasis id="B922D319FFB4731737E3F93BE69CF940" bold="true" box="[1422,1435,1665,1689]" pageId="71" pageNumber="72">4</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="8BE90F0BFFB4731732FAF91EE69CF962" blockId="71.[151,1437,1422,2035]" box="[151,1435,1699,1723]" pageId="71" pageNumber="72">
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<keyLead id="30A7AC3EFFB4731732FAF91EE69CF962" box="[151,1435,1699,1723]" pageId="71" pageNumber="72">
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||
- Antenna with>10 apical cones (Fig. 52D). .................................................................................................................
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<emphasis id="B922D319FFB4731737E3F919E69CF962" bold="true" box="[1422,1435,1699,1723]" pageId="71" pageNumber="72">5</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30A217AEFFB4731732FAF97CE69BF89C" pageId="71" pageNumber="72">
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<paragraph id="8BE90F0BFFB4731732FAF97CE69BF8D9" blockId="71.[151,1437,1422,2035]" pageId="71" pageNumber="72">
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<keyLead id="30A7AC3EFFB4731732FAF97CE69BF8D9" pageId="71" pageNumber="72">
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<emphasis id="B922D319FFB4731732FAF97CE3A3F907" bold="true" box="[151,164,1734,1758]" pageId="71" pageNumber="72">4</emphasis>
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. Anal shield surface covered with short hairs. Posterior locking carina of anal shield 2.5–3 times longer than anterior carina
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<emphasis id="B922D319FFB47317337CF953E0C1F8D9" box="[273,966,1769,1792]" italics="true" pageId="71" pageNumber="72">........................................................................................................</emphasis>
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<taxonomicName id="4C567488FFB4731731A1F953E69BF8D9" ID-CoL="7GH2H" authority="(de Saussure & Zehntner, 1897)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[972,1436,1769,1792]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="alluaudi">
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<emphasis id="B922D319FFB4731731A1F953E746F8D9" box="[972,1089,1769,1792]" italics="true" pageId="71" pageNumber="72">Z. alluaudi</emphasis>
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(
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<bibRefCitation id="EFC772FAFFB473173622F953E693F8D9" author="de Saussure, H. & Zehntner, L." box="[1103,1428,1769,1792]" pageId="71" pageNumber="72" pagination="1 - 12" refId="ref60128" refString="de Saussure, H. & Zehntner, L. (1897) Atlas de l'histoire naturelle des Myriapodes, In: Grandidier (Ed.) Histoire physique, naturelle et politique de Madagascar. 27 (53), Alfred Grandidier, Paris, France, plates 1 - 12." type="book chapter" year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
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)
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BE90F0BFFB4731732FAF8B6E69BF89C" blockId="71.[151,1437,1422,2035]" pageId="71" pageNumber="72">
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<keyLead id="30A7AC3EFFB4731732FAF8B6E69BF89C" pageId="71" pageNumber="72">
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- Anal shield surface glabrous. Posterior locking carina of anal shield more than 3.5 times longer than anterior carina
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<emphasis id="B922D319FFB4731732AAF895E072F89F" box="[199,885,1839,1862]" italics="true" pageId="71" pageNumber="72">.......................................................................................................</emphasis>
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<taxonomicName id="4C567488FFB473173114F895E69BF89C" ID-CoL="5D7JF" authority="(de Saussure & Zehntner, 1897)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[889,1436,1838,1862]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="voeltzkowianum">
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<emphasis id="B922D319FFB473173114F895E746F89F" box="[889,1089,1839,1862]" italics="true" pageId="71" pageNumber="72">Z. voeltzkowianum</emphasis>
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(
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<bibRefCitation id="EFC772FAFFB473173622F894E693F89C" author="de Saussure, H. & Zehntner, L." box="[1103,1428,1838,1861]" pageId="71" pageNumber="72" pagination="1 - 12" refId="ref60128" refString="de Saussure, H. & Zehntner, L. (1897) Atlas de l'histoire naturelle des Myriapodes, In: Grandidier (Ed.) Histoire physique, naturelle et politique de Madagascar. 27 (53), Alfred Grandidier, Paris, France, plates 1 - 12." type="book chapter" year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
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)
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30A217AEFFB4731832FAF8EBE69CFF2D" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
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||
<paragraph id="8BE90F0BFFB4731732FAF8EBE69CF808" blockId="71.[151,1437,1422,2035]" pageId="71" pageNumber="72">
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<keyLead id="30A7AC3EFFB4731732FAF8EBE69CF808" pageId="71" pageNumber="72">
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<emphasis id="B922D319FFB4731732FAF8EBE3A3F8B0" bold="true" box="[151,164,1873,1897]" pageId="71" pageNumber="72">5</emphasis>
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. Male harp with three stridulation ribs (Fig. 51C). Movable finger of posterior telopods 2 times longer than wide (Fig. 51E). Endotergum with three rows of marginal bristles, bristles slightly extending beyond tergite (Fig. 63B). Posterior locking carina of anal shield four times longer than anterior carina. Male anal shield bell-shaped. Sclerotized teeth present on antennomeres 1–5 (Figs 52A, B).
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<emphasis id="B922D319FFB473173136F803E65EF809" box="[859,1369,1977,2000]" italics="true" pageId="71" pageNumber="72">
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.....................................................
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<taxonomicName id="4C567488FFB4731736D2F803E652F809" ID-CoL="5D7JB" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1215,1365,1977,2000]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="trichordum" status="sp. nov.">Z. trichordum</taxonomicName>
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,
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</emphasis>
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<emphasis id="B922D319FFB47317370DF803E69CF808" bold="true" box="[1376,1435,1977,2001]" pageId="71" pageNumber="72">
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||
<taxonomicNameLabel id="A2116E62FFB47317370DF803E69CF808" box="[1376,1435,1977,2001]" pageId="71" pageNumber="72" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="8BE90F0BFFB4731832FAF866E69CFF2D" blockId="71.[151,1437,1422,2035]" lastBlockId="72.[151,1437,152,2012]" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
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||
<keyLead id="30A7AC3EFFB4731832FAF866E69CFF2D" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
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||
- Male harp with two stridulation ribs. Movable finger 2.4–2.5 times longer than wide. Endotergum with one up to two rows of marginal bristles, bristles not reaching tergite margin. Posterior locking carina of anal shield three times longer than anterior carina. Male anal shield well-rounded. Sclerotized teeth only present up to antennomere 3 or 4. ......................................................................................................................................................................................
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||
<emphasis id="B922D319FFBB731837E3FF66E69CFF2D" bold="true" box="[1422,1435,220,244]" pageId="72" pageNumber="73">6</emphasis>
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</keyLead>
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||
</paragraph>
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||
</keyStep>
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||
<keyStep id="30A217AEFFBB731832FAFF45E69CFE3E" pageId="72" pageNumber="73">
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<paragraph id="8BE90F0BFFBB731832FAFF45E69BFE85" blockId="72.[151,1437,152,2012]" pageId="72" pageNumber="73">
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<keyLead id="30A7AC3EFFBB731832FAFF45E69BFE85" pageId="72" pageNumber="73">
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||
<emphasis id="B922D319FFBB731832FAFF45E3A3FECE" bold="true" box="[151,164,255,279]" pageId="72" pageNumber="73">6</emphasis>
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. Specimens longer than
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<quantity id="4CAEA2EEFFBB731833D2FEBAE10AFECE" box="[447,525,256,279]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="72" pageNumber="73" unit="mm" value="60.0">60 mm</quantity>
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. Tergites dark-green, covered with numerous large pits and short setae. Operculum of vulva not protruding above coxal margin. Sclerotized teeth present on antennomeres 1–3, disc with
|
||
<emphasis id="B922D319FFBB73183691FE98E635FEE0" box="[1276,1330,290,313]" italics="true" pageId="72" pageNumber="73">circa</emphasis>
|
||
45 apical cones. Endotergum with two rows of marginal bristles.
|
||
<emphasis id="B922D319FFBB7318317FFEFFE0E0FE85" box="[786,999,325,348]" italics="true" pageId="72" pageNumber="73">................................</emphasis>
|
||
<taxonomicName id="4C567488FFBB73183180FEFFE68FFE85" ID-CoL="5D7JD" authority="Wesener & Sierwald, 2005" authorityName="Wesener & Sierwald" authorityYear="2005" box="[1005,1416,325,348]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="villosum">
|
||
<emphasis id="B922D319FFBB73183180FEFFE763FE85" box="[1005,1124,325,348]" italics="true" pageId="72" pageNumber="73">Z. villosum</emphasis>
|
||
Wesener & Sierwald, 2005
|
||
</taxonomicName>
|
||
A
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFBB731832FAFED2E69CFE3E" blockId="72.[151,1437,152,2012]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAFED2E69CFE3E" pageId="72" pageNumber="73">
|
||
- Specimens only slightly longer than
|
||
<quantity id="4CAEA2EEFFBB7318300CFED2E1B4FEA6" box="[609,691,360,383]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="72" pageNumber="73" unit="mm" value="30.0">30 mm</quantity>
|
||
. Tergites jade-like green with thin brown posterior margin, texture glossy. Operculum of vulva deeply notched, strongly extending beyond coxa (Fig. 54A). Sclerotized teeth present on antennomeres 1–4 (Fig. 55A), disc with
|
||
<emphasis id="B922D319FFBB73183011FE17E1B5FE1D" box="[636,690,429,452]" italics="true" pageId="72" pageNumber="73">circa</emphasis>
|
||
14 apical cones (Fig. 55B). Endotergum with single row of marginal bristles (Fig. 63C).
|
||
<emphasis id="B922D319FFBB731833FBFE6AE65EFE3E" box="[406,1369,464,487]" italics="true" pageId="72" pageNumber="73">
|
||
......................................................................................................................
|
||
<taxonomicName id="4C567488FFBB731836C6FE6AE65EFE3E" ID-CoL="5D7J2" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1195,1369,464,487]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="smaragdinum" status="sp. nov.">Z. smaragdinum</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="B922D319FFBB7318370DFE75E69CFE3E" bold="true" box="[1376,1435,463,487]" pageId="72" pageNumber="73">
|
||
<taxonomicNameLabel id="A2116E62FFBB7318370DFE75E69CFE3E" box="[1376,1435,463,487]" pageId="72" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFBB731832FAFE48E69CFDF5" pageId="72" pageNumber="73">
|
||
<paragraph id="8BE90F0BFFBB731832FAFE48E69CFDD3" blockId="72.[151,1437,152,2012]" box="[151,1435,498,522]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAFE48E69CFDD3" box="[151,1435,498,522]" pageId="72" pageNumber="73">
|
||
<emphasis id="B922D319FFBB731832FAFE48E3A3FDD3" bold="true" box="[151,164,498,522]" pageId="72" pageNumber="73">7</emphasis>
|
||
. Movable finger 3 times longer than wide (Fig. 56C).
|
||
<emphasis id="B922D319FFBB7318309AFE48E65EFDD0" box="[759,1369,498,521]" italics="true" pageId="72" pageNumber="73">
|
||
......................................................................
|
||
<taxonomicName id="4C567488FFBB731836BDFE48E652FDD0" ID-CoL="5D7GC" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1232,1365,498,521]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="ambrense" status="sp. nov.">Z. ambrense</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFBB7318370DFE48E69CFDD3" bold="true" box="[1376,1435,498,522]" pageId="72" pageNumber="73">
|
||
<taxonomicNameLabel id="A2116E62FFBB7318370DFE48E69CFDD3" box="[1376,1435,498,522]" pageId="72" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFBB731832FAFDAFE69CFDF5" blockId="72.[151,1437,152,2012]" box="[151,1435,532,556]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAFDAFE69CFDF5" box="[151,1435,532,556]" pageId="72" pageNumber="73">
|
||
- Movable finger <2.5 times longer than wide. ..............................................................................................................
|
||
<emphasis id="B922D319FFBB731837E3FDAEE69CFDF5" bold="true" box="[1422,1435,532,556]" pageId="72" pageNumber="73">8</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFBB731832FAFD8DE69CFD6E" pageId="72" pageNumber="73">
|
||
<paragraph id="8BE90F0BFFBB731832FAFD8DE69CFDAB" blockId="72.[151,1437,152,2012]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAFD8DE69CFDAB" pageId="72" pageNumber="73">
|
||
<emphasis id="B922D319FFBB731832FAFD8DE3A3FD96" bold="true" box="[151,164,567,591]" pageId="72" pageNumber="73">8</emphasis>
|
||
. Antenna with four apical cones (Fig. 62C). Movable finger of posterior telopods at inner margin with two membranous lobes (Fig. 58C). ....................................................................................................................................
|
||
<emphasis id="B922D319FFBB731837E3FDE0E69CFDAB" bold="true" box="[1422,1435,602,626]" pageId="72" pageNumber="73">9</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFBB731832FAFDC7E69CFD6E" blockId="72.[151,1437,152,2012]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAFDC7E69CFD6E" pageId="72" pageNumber="73">
|
||
- Antenna with>10 apical cones (Fig. 65C). Movable finger of posterior telopods at inner margin with single membranous lobe. .....................................................................................................................................................
|
||
<emphasis id="B922D319FFBB731837ECFD25E69CFD6E" bold="true" box="[1409,1435,671,695]" pageId="72" pageNumber="73">10</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFBB731832FAFD78E69CFC73" pageId="72" pageNumber="73">
|
||
<paragraph id="8BE90F0BFFBB731832FAFD78E69CFCC6" blockId="72.[151,1437,152,2012]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAFD78E69CFCC6" pageId="72" pageNumber="73">
|
||
<emphasis id="B922D319FFBB731832FAFD78E3A3FD03" bold="true" box="[151,164,706,730]" pageId="72" pageNumber="73">9</emphasis>
|
||
. Collum glabrous. Anterior telopod second podomere process visible in anterior view (Fig. 58A). Fifth antennomere without
|
||
<emphasis id="B922D319FFBB73183372FD5FE2EDFD25" box="[287,490,741,764]" italics="true" pageId="72" pageNumber="73">sensilla basiconica</emphasis>
|
||
(Fig. 59B). Endotergum with single row of large cuticular impressions, marginal bristles slightly protruding above tergite (Fig. 63E). ..........................................................................................
|
||
<emphasis id="B922D319FFBB7318376EFCB2E65EFCC6" box="[1283,1369,776,799]" italics="true" pageId="72" pageNumber="73">
|
||
<taxonomicName id="4C567488FFBB7318376EFCB2E652FCC6" ID-CoL="5D7HC" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1283,1365,776,799]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="isalo" status="sp. nov.">Z. isalo</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFBB7318370DFCBDE69CFCC6" bold="true" box="[1376,1435,775,799]" pageId="72" pageNumber="73">
|
||
<taxonomicNameLabel id="A2116E62FFBB7318370DFCBDE69CFCC6" box="[1376,1435,775,799]" pageId="72" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFBB731832FAFC90E69CFC73" blockId="72.[151,1437,152,2012]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAFC90E69CFC73" pageId="72" pageNumber="73">
|
||
- Collum with isolated, long hair. Anterior telopod second podomere process not visible in anterior view (Fig. 61B). Fifth antennomere with field of
|
||
<emphasis id="B922D319FFBB73183073FCF7E1EDFCBD" box="[542,746,845,868]" italics="true" pageId="72" pageNumber="73">sensilla basiconica</emphasis>
|
||
(Fig. 62B). Endotergum with row of large cuticular impressions and second row of much smaller impressions, bristles long, strongly protruding above tergite (Fig. 63F). .................
|
||
<emphasis id="B922D319FFBB731832AAFC28E65EFC70" box="[199,1369,914,937]" italics="true" pageId="72" pageNumber="73">
|
||
.............................................................................................................................................................
|
||
<taxonomicName id="4C567488FFBB7318368CFC28E652FC70" ID-CoL="5D7GL" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1249,1365,914,937]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="bilobum" status="sp. nov.">Z. bilobum</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFBB7318370DFC28E69CFC73" bold="true" box="[1376,1435,914,938]" pageId="72" pageNumber="73">
|
||
<taxonomicNameLabel id="A2116E62FFBB7318370DFC28E69CFC73" box="[1376,1435,914,938]" pageId="72" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFBB731832FAFC0EE69CFC36" pageId="72" pageNumber="73">
|
||
<paragraph id="8BE90F0BFFBB731832FAFC0EE69BFC15" blockId="72.[151,1437,152,2012]" box="[151,1436,948,972]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAFC0EE69BFC15" box="[151,1436,948,972]" pageId="72" pageNumber="73">
|
||
<emphasis id="B922D319FFBB731832FAFC0EE3B5FC15" bold="true" box="[151,178,948,972]" pageId="72" pageNumber="73">10</emphasis>
|
||
. First pair of stigmatic plates greatly enlarged (Fig. 64A). .........................................................................................
|
||
<emphasis id="B922D319FFBB731837EEFC0EE69BFC15" bold="true" box="[1411,1436,948,972]" pageId="72" pageNumber="73">11</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFBB731832FAFC62E69CFC36" blockId="72.[151,1437,152,2012]" box="[151,1435,983,1007]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAFC62E69CFC36" box="[151,1435,983,1007]" pageId="72" pageNumber="73">
|
||
- First pair of stigmatic plates triangular, tapering towards tip. ..................................................................................
|
||
<emphasis id="B922D319FFBB731837ECFC6DE69CFC36" bold="true" box="[1409,1435,983,1007]" pageId="72" pageNumber="73">12</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFBB731832FAFC40E69CFA93" pageId="72" pageNumber="73">
|
||
<paragraph id="8BE90F0BFFBB731832FAFC40E69CFB45" blockId="72.[151,1437,152,2012]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAFC40E69CFB45" pageId="72" pageNumber="73">
|
||
<emphasis id="B922D319FFBB731832FAFC40E3B7FBCB" bold="true" box="[151,176,1018,1042]" pageId="72" pageNumber="73">11</emphasis>
|
||
. Operculum of vulva ending clearly before coxal margin, deeply notched, divided into two large, well-rounded samesized lobes (Fig. 64C). Third podomere of anterior telopod laterally with
|
||
<emphasis id="B922D319FFBB7318319BFBA7E72BFBED" box="[1014,1068,1053,1076]" italics="true" pageId="72" pageNumber="73">circa</emphasis>
|
||
five large, sclerotized teeth (Fig. 64E).
|
||
<emphasis id="B922D319FFBB73183364FBFAE2DFFB8E" box="[265,472,1088,1111]" italics="true" pageId="72" pageNumber="73">Sensilla basiconica</emphasis>
|
||
on first antennomere absent (Fig. 65B). Posterior locking carina of anal shield two times as long as anterior carina. Endotergum marginal bristles protruding up to tergite margin (Fig. 74A). Movable finger of posterior telopods thick, 1.85 times longer than wide (Fig. 64G).
|
||
<emphasis id="B922D319FFBB731831F3FB3FE65EFB45" box="[926,1369,1157,1180]" italics="true" pageId="72" pageNumber="73">
|
||
................................................
|
||
<taxonomicName id="4C567488FFBB7318368CFB3FE652FB45" ID-CoL="5D7J8" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1249,1365,1157,1180]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="tampolo" status="sp. nov.">Z. tampolo</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFBB7318370DFB3EE69CFB45" bold="true" box="[1376,1435,1156,1180]" pageId="72" pageNumber="73">
|
||
<taxonomicNameLabel id="A2116E62FFBB7318370DFB3EE69CFB45" box="[1376,1435,1156,1180]" pageId="72" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFBB731832FAFB12E69CFA93" blockId="72.[151,1437,152,2012]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAFB12E69CFA93" pageId="72" pageNumber="73">
|
||
- Operculum of vulva ending before coxal margin, elongated into single median lobe, lobe only shallowly notched at tip (Fig. 66C). Third podomere of anterior telopod without any large, sclerotized teeth (Fig. 66E).
|
||
<emphasis id="B922D319FFBB7318372AFB71E23DFADD" italics="true" pageId="72" pageNumber="73">Sensilla basiconica</emphasis>
|
||
on first antennomere present (Fig. 67A). Locking carinae of anal shield of equal length. Endotergum marginal bristles very short, ending far before tergite margin (Fig. 74B). Movable finger of posterior telopods thick, 2.1 times longer than wide (Fig. 66G). .............................................................................................
|
||
<emphasis id="B922D319FFBB731836B2FA89E65EFA93" box="[1247,1369,1331,1354]" italics="true" pageId="72" pageNumber="73">
|
||
<taxonomicName id="4C567488FFBB731836B2FA89E652FA93" ID-CoL="5D7H9" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1247,1365,1331,1354]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="ignotum" status="sp. nov.">Z. ignotum</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFBB7318370DFA88E69CFA93" bold="true" box="[1376,1435,1330,1354]" pageId="72" pageNumber="73">
|
||
<taxonomicNameLabel id="A2116E62FFBB7318370DFA88E69CFA93" box="[1376,1435,1330,1354]" pageId="72" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFBB731832FAFAEEE69CFA2E" pageId="72" pageNumber="73">
|
||
<paragraph id="8BE90F0BFFBB731832FAFAEEE69CFA6B" blockId="72.[151,1437,152,2012]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAFAEEE69CFA6B" pageId="72" pageNumber="73">
|
||
<emphasis id="B922D319FFBB731832FAFAEEE3B5FAB5" bold="true" box="[151,178,1364,1388]" pageId="72" pageNumber="73">12</emphasis>
|
||
. Tergite texture orange-peel like. Collum covered with isolated, long hair. Body size of adult female>
|
||
<quantity id="4CAEA2EEFFBB73183728FAEFE69EFAB5" box="[1349,1433,1365,1388]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="72" pageNumber="73" unit="mm" value="50.0">50 mm</quantity>
|
||
. Endotergum with two rows of marginal bristles and single row of large circular cuticular impressions (Fig. 74C). ... ....................................................................................................................................................................................
|
||
<emphasis id="B922D319FFBB731837ECFA20E69CFA6B" bold="true" box="[1409,1435,1434,1458]" pageId="72" pageNumber="73">13</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFBB731832FAFA07E69CFA2E" blockId="72.[151,1437,152,2012]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAFA07E69CFA2E" pageId="72" pageNumber="73">
|
||
- Tergite texture glossy or smooth. Collum glabrous. Body size of adults <
|
||
<quantity id="4CAEA2EEFFBB73183184FA07E73EFA0D" box="[1001,1081,1469,1492]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="72" pageNumber="73" unit="mm" value="35.0">35 mm</quantity>
|
||
. Endotergum with single row of marginal bristles and either without or with two rows of cuticular impressions. .....................................................
|
||
<emphasis id="B922D319FFBB731837ECFA65E69CFA2E" bold="true" box="[1409,1435,1503,1527]" pageId="72" pageNumber="73">14</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFBB731832FAF9B8E69CF91E" pageId="72" pageNumber="73">
|
||
<paragraph id="8BE90F0BFFBB731832FAF9B8E69BF986" blockId="72.[151,1437,152,2012]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAF9B8E69BF986" pageId="72" pageNumber="73">
|
||
<emphasis id="B922D319FFBB731832FAF9B8E3BFF9C3" bold="true" box="[151,184,1538,1562]" pageId="72" pageNumber="73">13.</emphasis>
|
||
Posterior locking carina on anal shield 2.5 times longer than anterior carina. Third podomere of anterior telopods without large, sclerotized teeth. Anal shield glabrous. Endotergum marginal bristles short, not protruding up to tergite margin.
|
||
<emphasis id="B922D319FFBB73183303F9F2E084F986" box="[366,899,1608,1631]" italics="true" pageId="72" pageNumber="73">................................................................................</emphasis>
|
||
<taxonomicName id="4C567488FFBB731831EBF9F2E69BF986" ID-CoL="5D7GQ" authority="(de Saussure & Zehntner, 1897)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[902,1436,1608,1631]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="coquerelianum">
|
||
<emphasis id="B922D319FFBB731831EBF9F2E746F986" box="[902,1089,1608,1631]" italics="true" pageId="72" pageNumber="73">Z. coquerelianum</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC772FAFFBB73183622F9F2E693F986" author="de Saussure, H. & Zehntner, L." box="[1103,1428,1608,1631]" pageId="72" pageNumber="73" pagination="1 - 12" refId="ref60128" refString="de Saussure, H. & Zehntner, L. (1897) Atlas de l'histoire naturelle des Myriapodes, In: Grandidier (Ed.) Histoire physique, naturelle et politique de Madagascar. 27 (53), Alfred Grandidier, Paris, France, plates 1 - 12." type="book chapter" year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFBB731832FAF9D0E69CF91E" blockId="72.[151,1437,152,2012]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAF9D0E69CF91E" pageId="72" pageNumber="73">
|
||
- Posterior locking carina on anal shield 4.5 times longer than anterior carina. Third podomere of anterior telopods with
|
||
<emphasis id="B922D319FFBB7318336CF937E230F97D" box="[257,311,1677,1700]" italics="true" pageId="72" pageNumber="73">circa</emphasis>
|
||
seven large, sclerotized teeth (Fig. 68C). Anal shield covered with short hair. Endotergum marginal bristles protruding up to tergite margin (Fig. 74C)..
|
||
<emphasis id="B922D319FFBB7318309CF90AE65EF91E" box="[753,1369,1712,1735]" italics="true" pageId="72" pageNumber="73">
|
||
.....................................................................
|
||
<taxonomicName id="4C567488FFBB731836D2F90AE652F91E" ID-CoL="5D7J7" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1215,1365,1712,1735]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="tainkintana" status="sp. nov.">Z. tainkintana</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFBB7318370DF915E69CF91E" bold="true" box="[1376,1435,1711,1735]" pageId="72" pageNumber="73">
|
||
<taxonomicNameLabel id="A2116E62FFBB7318370DF915E69CF91E" box="[1376,1435,1711,1735]" pageId="72" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A217AEFFBB731832FAF968E69CF805" pageId="72" pageNumber="73">
|
||
<paragraph id="8BE90F0BFFBB731832FAF968E69CF88B" blockId="72.[151,1437,152,2012]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAF968E69CF88B" pageId="72" pageNumber="73">
|
||
<emphasis id="B922D319FFBB731832FAF968E3B5F933" bold="true" box="[151,178,1746,1770]" pageId="72" pageNumber="73">14</emphasis>
|
||
. Tergites glossy. Female anal shield bell-shaped, male with steep slope. Posterior telopods fixed finger wide, only slightly curved towards movable finger (Fig. 70C). Movable finger 1.8 times longer than wide, on posterior side with ten small sclerotized teeth (Fig. 70D). Sclerotized teeth on antenna only present on first antennomere (Fig. 71B). Endotergum without cuticular impressions (Fig. 74D).
|
||
<emphasis id="B922D319FFBB73183123F880E65EF888" box="[846,1369,1850,1873]" italics="true" pageId="72" pageNumber="73">
|
||
....................................................
|
||
<taxonomicName id="4C567488FFBB731836C1F880E653F888" ID-CoL="7GGQS" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1196,1364,1850,1873]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="viridissimum" status="sp. nov.">Z. viridissimum</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFBB7318370DF880E69CF88B" bold="true" box="[1376,1435,1850,1874]" pageId="72" pageNumber="73">
|
||
<taxonomicNameLabel id="A2116E62FFBB7318370DF880E69CF88B" box="[1376,1435,1850,1874]" pageId="72" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFBB731832FAF8E7E69CF805" blockId="72.[151,1437,152,2012]" pageId="72" pageNumber="73">
|
||
<keyLead id="30A7AC3EFFBB731832FAF8E7E69CF805" pageId="72" pageNumber="73">
|
||
- Tergites matte. Anal shield well-rounded. Fixed finger slender, strongly curved towards movable finger (Fig. 72D). Movable finger 2.4 times longer than wide, on posterior side with
|
||
<emphasis id="B922D319FFBB731831D7F83AE0F7F84E" box="[954,1008,1920,1943]" italics="true" pageId="72" pageNumber="73">circa</emphasis>
|
||
25 small sclerotized teeth (Fig. 72E). Sclerotized teeth only present on antennomeres 1–3 (Fig. 73C). Endotergum with single row of large and second row of smaller circular cuticular impressions (Fig. 74E).
|
||
<emphasis id="B922D319FFBB7318309CF87FE65EF805" box="[753,1369,1989,2012]" italics="true" pageId="72" pageNumber="73">
|
||
..................................................................
|
||
<taxonomicName id="4C567488FFBB731836C6F87FE652F805" ID-CoL="7GGPZ" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1195,1365,1989,2012]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">Z. denticulatum</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFBB7318370DF87EE69CF805" bold="true" box="[1376,1435,1988,2012]" pageId="72" pageNumber="73">
|
||
<taxonomicNameLabel id="A2116E62FFBB7318370DF87EE69CF805" box="[1376,1435,1988,2012]" pageId="72" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |