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<document id="6CCFA28EE19B12829E05BC36886F5495" ID-CLB-Dataset="28430" ID-DOI="10.11646/zootaxa.4564.1.3" ID-GBIF-Dataset="4d361b19-8a22-4747-b7d7-57581cfdce26" ID-ISSN="1175-5326" ID-Zenodo-Dep="2588669" ID-ZooBank="B8991FC7-399B-4B00-B968-D3BC44F32020" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552167099920" checkinUser="plazi" docAuthor="Schubert, Joseph" docDate="2019" docId="03FE87F4970EFF98FF15F9CBFDB7BE25" docLanguage="en" docName="zootaxa.4564.1.3.pdf" docOrigin="Zootaxa 4564 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Maratus combustus Schubert 2019, sp. nov." docType="treatment" docUuid="15070D82-B56E-4880-A53B-75E0871ACA9D" docUuidSource="ZooBank" docVersion="8" lastPageNumber="94" masterDocId="FFC7FF8C9709FF95FF82FFBBFFC4BD30" masterDocTitle="Three new peacock spiders from Southwestern Australia (Araneae: Salticidae: Euophryini: Maratus Karsch, 1878)" masterLastPageNumber="100" masterPageNumber="81" pageNumber="88" updateTime="1698700571345" updateUser="ExternalLinkService">
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<mods:title id="92EA07B27A4B2CAB87C13908CF87E683">Three new peacock spiders from Southwestern Australia (Araneae: Salticidae: Euophryini: Maratus Karsch, 1878)</mods:title>
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<treatment id="03FE87F4970EFF98FF15F9CBFDB7BE25" ID-DOI="http://doi.org/10.5281/zenodo.5931341" ID-GBIF-Taxon="154366369" ID-Zenodo-Dep="5931341" LSID="urn:lsid:zoobank.org:act:15070D82-B56E-4880-A53B-75E0871ACA9D" httpUri="http://treatment.plazi.org/id/03FE87F4970EFF98FF15F9CBFDB7BE25" lastPageId="13" lastPageNumber="94" pageId="7" pageNumber="88">
<subSubSection id="C34D6569970EFF92FF15F9CBFE2EBBBB" box="[151,490,1648,1675]" pageId="7" pageNumber="88" type="nomenclature">
<paragraph id="8BE836E2970EFF92FF15F9CBFE2EBBBB" blockId="7.[151,981,1648,1708]" box="[151,490,1648,1675]" pageId="7" pageNumber="88">
<heading id="D0A0818E970EFF92FF15F9CBFE2EBBBB" bold="true" box="[151,490,1648,1675]" fontSize="11" level="1" pageId="7" pageNumber="88" reason="1">
<emphasis id="B923EAF0970EFF92FF15F9CBFE2EBBBB" bold="true" box="[151,490,1648,1675]" pageId="7" pageNumber="88">
<taxonomicName id="4C574D61970EFF92FF15F9CBFE42BBBA" ID-CoL="6R68P" authority="Schubert, 2019" authorityName="Schubert" authorityYear="2019" box="[151,390,1648,1674]" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="88" phylum="Arthropoda" rank="species" species="combustus" status="sp. nov.">
<emphasis id="B923EAF0970EFF92FF15F9CBFE42BBBA" bold="true" box="[151,390,1648,1674]" italics="true" pageId="7" pageNumber="88">Maratus combustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A210578B970EFF92FE0EF9CAFE2EBBBB" box="[396,490,1649,1675]" pageId="7" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34D6569970EFF92FF15F92FFC11BB9C" box="[151,981,1683,1708]" pageId="7" pageNumber="88" type="description">
<paragraph id="8BE836E2970EFF92FF15F92FFC11BB9C" blockId="7.[151,981,1648,1708]" box="[151,981,1683,1708]" pageId="7" pageNumber="88">
(
<figureCitation id="136C2A67970EFF92FF1CF92FFEFBBB9C" box="[158,319,1684,1708]" captionStart="FIGURE 2" captionStartId="2.[151,250,645,667]" captionTargetBox="[151,1435,208,604]" captionTargetId="figure@2.[151,1435,208,604]" captionTargetPageId="2" captionText="FIGURE 2. Living male paratype Maratus aquilus sp. nov., M. combustus sp. nov., and M. felinus sp. nov. performing courtship display to nearby female: A, Courtship display of male Maratus aquilus sp. nov.; B, courtship display of Maratus combustus sp. nov.; C, courtship display of Maratus felinus sp. nov." httpUri="https://zenodo.org/record/2588673/files/figure.png" pageId="7" pageNumber="88">FIGURES 2B</figureCitation>
,
<figureCitation id="136C2A67970EFF92FEC9F92FFEA1BB9C" box="[331,357,1684,1708]" captionStart="FIGURE 10" captionStartId="8.[151,250,1968,1990]" captionTargetBox="[168,1429,1095,1944]" captionTargetId="figure@8.[151,1436,1078,1952]" captionTargetPageId="8" captionText="FIGURE 10. Habitus images of living holotype Maratus combustus sp. nov. (WAMT147351): A, anterior view; B, dorsal view; C, lateral view; D: anterolateral view; E, dorsolateral view; F, lateral view." httpUri="https://zenodo.org/record/2588689/files/figure.png" pageId="7" pageNumber="88">10</figureCitation>
AF, 11AD, 12AC, 13AF, 14AD, 15AC, 16AB)
</paragraph>
</subSubSection>
<subSubSection id="C34D6569970EFF92FF15F960FDCEBA0C" pageId="7" pageNumber="88" type="materials_examined">
<paragraph id="8BE836E2970EFF92FF15F960FDCEBA0C" blockId="7.[151,1436,1755,2032]" pageId="7" pageNumber="88">
<emphasis id="B923EAF0970EFF92FF15F960FE46BBC4" bold="true" box="[151,386,1755,1780]" pageId="7" pageNumber="88">Material examined.</emphasis>
<materialsCitation id="3B3F3CBF970EFF92FE08F967FC9EBA28" ID-GBIF-Occurrence="2012749437" collectingDate="2018-09-25" collectionCode="MALE" collectorName="M. Duncan" country="Australia" elevation="123" latitude="-34.853054" location="Mount Romance" longLatPrecision="1" longitude="117.18667" pageId="7" pageNumber="88" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<collectionCode id="ED46AE27970EFF92FE08F967FE1DBBC4" box="[394,473,1756,1780]" pageId="7" pageNumber="88">MALE</collectionCode>
<typeStatus id="54EC8840970EFF92FE60F967FDB2BBC4" box="[482,630,1756,1780]" pageId="7" pageNumber="88" type="holotype">HOLOTYPE</typeStatus>
(WAMT147351) from
<collectingCountry id="F3407672970EFF92FC0DF960FC3DBBC4" box="[911,1017,1755,1780]" name="Australia" pageId="7" pageNumber="88">Australia</collectingCountry>
,
<collectingRegion id="4993F800970EFF92FB87F967FB14BBC4" box="[1029,1232,1755,1780]" country="Australia" name="Western Australia" pageId="7" pageNumber="88">Western Australia</collectingRegion>
<location id="8E886039970EFF92FB5AF967FA5CBBC3" LSID="urn:lsid:plazi:treatment:03FE87F4970EFF98FF15F9CBFDB7BE25:8E886039970EFF92FB5AF967FA5CBBC3" box="[1240,1432,1756,1779]" country="Australia" latitude="-34.853054" longLatPrecision="1" longitude="117.18667" name="Mount Romance" pageId="7" pageNumber="88" stateProvince="Western Australia">Mount Romance</location>
,
<geoCoordinate id="EE635025970EFF92FF15F944FEE8BA28" box="[151,300,1791,1816]" direction="south" orientation="latitude" pageId="7" pageNumber="88" precision="1" value="-34.853054">34°51'11.0&quot;S</geoCoordinate>
,
<geoCoordinate id="EE635025970EFF92FEBAF8BBFE18BA28" box="[312,476,1792,1816]" direction="east" orientation="longitude" pageId="7" pageNumber="88" precision="1" value="117.18667">117°11'12.0&quot;E</geoCoordinate>
,
<quantity id="4CAF9B07970EFF92FE6AF8BBFDEEBA28" box="[488,554,1792,1816]" metricMagnitude="2" metricUnit="m" metricValue="1.23" pageId="7" pageNumber="88" unit="m" value="123.0">
<elevation id="007AD1D1970EFF92FE6AF8BBFDEEBA28" box="[488,554,1792,1816]" metricMagnitude="2" metricUnit="m" metricValue="1.23" pageId="7" pageNumber="88" unit="m" value="123.0">123m</elevation>
</quantity>
,
<collectorName id="26A25334970EFF92FDB4F8BBFD73BA27" box="[566,695,1792,1815]" pageId="7" pageNumber="88">M. Duncan</collectorName>
,
<date id="FFE91022970EFF92FD40F944FC9EBA28" box="[706,858,1791,1816]" pageId="7" pageNumber="88" value="2018-09-25">
<collectingDate id="EFADE9CA970EFF92FD40F944FC9EBA28" box="[706,858,1791,1816]" pageId="7" pageNumber="88" value="2018-09-25">25 Sep. 2018</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B3F3CBF970EFF92FCE6F8BBFDC3BA0C" ID-GBIF-Occurrence="2012749436" collectingDate="2018-09-25" collectionCode="MALE" collectorName="M. Duncan" country="Australia" elevation="123" latitude="-34.853054" location="Mount Romance" longLatPrecision="1" longitude="117.18667" pageId="7" pageNumber="88" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Western Australia" typeStatus="paratype">
<typeStatus id="54EC8840970EFF92FCE6F8BBFBC7BA28" box="[868,1027,1792,1816]" pageId="7" pageNumber="88" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D51FD6B970EFF92FB8FF8BBFB9FBA28" box="[1037,1115,1791,1816]" pageId="7" pageNumber="88" type="male">2 male</specimenCount>
,
<specimenCount id="9D51FD6B970EFF92FBE5F8BBFB0CBA28" box="[1127,1224,1791,1816]" pageId="7" pageNumber="88" type="female">2 female</specimenCount>
(WAM-T147352), same collection data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D6569970EFF92FF45F8FCFCF8BAB4" pageId="7" pageNumber="88" type="etymology">
<paragraph id="8BE836E2970EFF92FF45F8FCFCF8BAB4" blockId="7.[151,1436,1755,2032]" pageId="7" pageNumber="88">
<emphasis id="B923EAF0970EFF92FF45F8FCFE8ABA50" bold="true" box="[199,334,1863,1888]" pageId="7" pageNumber="88">Etymology.</emphasis>
The specific name (
<taxonomicName id="4C574D61970EFF92FDB3F8FCFD37BA50" authority=", Latin" authorityName="Latin" box="[561,755,1863,1888]" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="88" phylum="Arthropoda" rank="species" species="combustus">
<emphasis id="B923EAF0970EFF92FDB3F8FCFD63BA6F" box="[561,679,1863,1887]" italics="true" pageId="7" pageNumber="88">combustus</emphasis>
, Latin
</taxonomicName>
, m., participle, English translation: burnt) refers to the burntorange colour of the scales on the dorsal opisthosomal plate.
</paragraph>
</subSubSection>
<subSubSection id="C34D6569970EFF9DFF45F834FE7FBC70" lastPageId="8" lastPageNumber="89" pageId="7" pageNumber="88" type="diagnosis">
<paragraph id="8BE836E2970EFF9DFF45F834FE7FBC70" blockId="7.[151,1436,1755,2032]" lastBlockId="8.[151,1437,151,1040]" lastPageId="8" lastPageNumber="89" pageId="7" pageNumber="88">
<emphasis id="B923EAF0970EFF92FF45F834FE85BA98" bold="true" box="[199,321,1935,1960]" pageId="7" pageNumber="88">Diagnosis.</emphasis>
This species belongs to the
<taxonomicName id="4C574D61970EFF92FDF8F82BFCDBBA97" authorityName="Otto &amp; Hill" authorityYear="2016" box="[634,799,1936,1959]" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="88" phylum="Arthropoda" rank="species" species="vespa">
<emphasis id="B923EAF0970EFF92FDF8F82BFCDBBA97" box="[634,799,1936,1959]" italics="true" pageId="7" pageNumber="88">Maratus vespa</emphasis>
</taxonomicName>
group (
<bibRefCitation id="EFC64B13970EFF92FCF7F82BFB90BA98" author="Otto, J. C. &amp; Hill, D. E." box="[885,1108,1935,1960]" pageId="7" pageNumber="88" pagination="1 - 23" refId="ref6047" refString="Otto, J. C. &amp; Hill, D. E. (2017 B) Catalogue of the Australian peacock spiders (Araneae: Salticidae: Euophryini: Maratus, Saratus), version 2. Peckhamia 148 (2), 1 - 23." type="journal article" year="2017">Otto &amp; Hill, 2017B</bibRefCitation>
), with respect to the presence of many long bristle-like setae on legs III of the male, and the courtship display of the male which rapidly turns from side to side with a raised opisthosoma and extended large, lobate lateral opisthosomal flaps, displaying to females at close proximity (within millimetres).
<taxonomicName id="4C574D619701FF9DFD3EFF22FC5DBD80" box="[700,921,152,176]" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="combustus">
<emphasis id="B923EAF09701FF9DFD3EFF22FC5DBD80" box="[700,921,152,176]" italics="true" pageId="8" pageNumber="89">Maratus combustus</emphasis>
</taxonomicName>
can be distinguished from other males of the group by its unique dorsal opisthosomal pattern. Females of
<taxonomicName id="4C574D619701FF9DFCCEFF06FC2DBDE4" box="[844,1001,188,212]" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="combustus">
<emphasis id="B923EAF09701FF9DFCCEFF06FC2DBDE4" box="[844,1001,188,212]" italics="true" pageId="8" pageNumber="89">M. combustus</emphasis>
</taxonomicName>
are similar to other females in the
<taxonomicName id="4C574D619701FF9DFAFFFF06FF13BDC8" authorityName="Otto &amp; Hill" authorityYear="2016" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="vespa">
<emphasis id="B923EAF09701FF9DFAFFFF06FF13BDC8" italics="true" pageId="8" pageNumber="89">M. vespa</emphasis>
</taxonomicName>
clade and identification is not possible without association with a male. The structures of the external male and female genitalia of
<taxonomicName id="4C574D619701FF9DFE2DFEBEFD8ABC2C" box="[431,590,260,284]" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="combustus">
<emphasis id="B923EAF09701FF9DFE2DFEBEFD8ABC2C" box="[431,590,260,284]" italics="true" pageId="8" pageNumber="89">M. combustus</emphasis>
</taxonomicName>
are of no use in distinguishing them from other Southwestern
<collectingCountry id="F34076729701FF9DFAB7FEBFFA58BC2D" box="[1333,1436,260,285]" name="Australia" pageId="8" pageNumber="89">Australia</collectingCountry>
endemic
<taxonomicName id="4C574D619701FF9DFF7EFE92FE9FBC70" authorityName="Karsch" authorityYear="1878" box="[252,347,297,320]" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B923EAF09701FF9DFF7EFE92FE9FBC70" box="[252,347,297,320]" italics="true" pageId="8" pageNumber="89">Maratus</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="C34D65699701FF98FF45FEF7FF17BF28" lastPageId="13" lastPageNumber="94" pageId="8" pageNumber="89" type="description">
<paragraph id="8BE836E29701FF9DFF45FEF7FCEFBF0C" blockId="8.[151,1437,151,1040]" pageId="8" pageNumber="89">
<emphasis id="B923EAF09701FF9DFF45FEF7FE5ABC55" bold="true" box="[199,414,332,357]" pageId="8" pageNumber="89">Description. Male</emphasis>
(WAMT147351). Carapace mostly black and glabrous; median thoracic tract comprised of white scales extends from rear slope of carapace to between the PLE (may be worn in some specimens, see
<figureCitation id="136C2A679701FF9DFF15FE2FFED8BC9D" box="[151,284,404,429]" captionStart="FIGURE 15" captionStartId="12.[151,250,986,1008]" captionTargetBox="[157,1418,388,950]" captionTargetId="figure@12.[151,1436,373,965]" captionTargetPageId="12" captionText="FIGURE 15. Variation in preserved holotype and paratype male Maratus combustus sp. nov. specimens: A, holotype male; B, paratype male #1; C, paratype male #2." httpUri="https://zenodo.org/record/2588699/files/figure.png" pageId="8" pageNumber="89">Figure 15C</figureCitation>
); carapace rimmed with well-defined marginal band comprised of white scales (
<figureCitation id="136C2A679701FF9DFB3EFE2FFA4BBC9C" box="[1212,1423,404,429]" captionStart="FIGURE 10" captionStartId="8.[151,250,1968,1990]" captionTargetBox="[168,1429,1095,1944]" captionTargetId="figure@8.[151,1436,1078,1952]" captionTargetPageId="8" captionText="FIGURE 10. Habitus images of living holotype Maratus combustus sp. nov. (WAMT147351): A, anterior view; B, dorsal view; C, lateral view; D: anterolateral view; E, dorsolateral view; F, lateral view." httpUri="https://zenodo.org/record/2588689/files/figure.png" pageId="8" pageNumber="89">Figure 10C and F</figureCitation>
); ocular quadrangle region with dense cover of grey scales and interrupted by seven patches of red scales; two patches of red scales at rear margin of ocular quadrangle with one smaller patch between them; two smaller patches of red scales at midline of ocular quadrangle positioned parallel to each of the AME; larger patches of red scales extend from directly behind each of the PLE (
<figureCitation id="136C2A679701FF9DFD1AFD9FFCDABF0C" box="[664,798,548,573]" captionStart="FIGURE 10" captionStartId="8.[151,250,1968,1990]" captionTargetBox="[168,1429,1095,1944]" captionTargetId="figure@8.[151,1436,1078,1952]" captionTargetPageId="8" captionText="FIGURE 10. Habitus images of living holotype Maratus combustus sp. nov. (WAMT147351): A, anterior view; B, dorsal view; C, lateral view; D: anterolateral view; E, dorsolateral view; F, lateral view." httpUri="https://zenodo.org/record/2588689/files/figure.png" pageId="8" pageNumber="89">Figure 10A</figureCitation>
).
</paragraph>
<paragraph id="8BE836E29701FF9DFF45FDF3FC0BBFDF" blockId="8.[151,1437,151,1040]" pageId="8" pageNumber="89">
PME closer to PLE than to ALE; AME bordered with grey-white scales; ALE bordered with bright red scales dorsally and grey-white scales laterally and below; long white setae project downwards from below anterior median eyes forming triangular shape; clypeus covered with short, white setae (
<figureCitation id="136C2A679701FF9DFB9CFD34FB6FBF98" box="[1054,1195,655,680]" captionStart="FIGURE 10" captionStartId="8.[151,250,1968,1990]" captionTargetBox="[168,1429,1095,1944]" captionTargetId="figure@8.[151,1436,1078,1952]" captionTargetPageId="8" captionText="FIGURE 10. Habitus images of living holotype Maratus combustus sp. nov. (WAMT147351): A, anterior view; B, dorsal view; C, lateral view; D: anterolateral view; E, dorsolateral view; F, lateral view." httpUri="https://zenodo.org/record/2588689/files/figure.png" pageId="8" pageNumber="89">Figures 10A</figureCitation>
and
<figureCitation id="136C2A679701FF9DFB66FD2BFAD1BF98" box="[1252,1301,656,680]" captionStart="FIGURE 11" captionStartId="9.[151,250,1521,1543]" captionTargetBox="[214,1386,233,1440]" captionTargetId="figure@9.[151,1435,193,1500]" captionTargetPageId="9" captionText="FIGURE 11. Habitus images of preserved holotype Maratus combustus sp. nov. (WAMT147351): A, dorsal view; B, ventral view; C, anterior view; D, lateral view." httpUri="https://zenodo.org/record/2588691/files/figure.png" pageId="8" pageNumber="89">11C</figureCitation>
); chelicerae dark brown to black and glabrous; coxae and labium dark brown and glabrous; endites dark brown and glabrous; sternum dark brown (
<figureCitation id="136C2A679701FF9DFE05FD6CFDC3BFC0" box="[391,519,727,752]" captionStart="FIGURE 11" captionStartId="9.[151,250,1521,1543]" captionTargetBox="[214,1386,233,1440]" captionTargetId="figure@9.[151,1435,193,1500]" captionTargetPageId="9" captionText="FIGURE 11. Habitus images of preserved holotype Maratus combustus sp. nov. (WAMT147351): A, dorsal view; B, ventral view; C, anterior view; D, lateral view." httpUri="https://zenodo.org/record/2588691/files/figure.png" pageId="8" pageNumber="89">Figure 11B</figureCitation>
) with light covering of fine, white setae.
</paragraph>
<paragraph id="8BE836E29701FF9DFF45FD47FC1FB920" blockId="8.[151,1437,151,1040]" pageId="8" pageNumber="89">
Dorsal opisthosomal plate distinctively patterned with thick cover of scales; five tracts of scales, alternating in colour between red and blue and bordered by black extend from each anterolateral edge of dorsal opisthosoma towards median dorsal opisthosoma; light cover of mixed white and tan setae between each set of tracts (may be worn in some specimens, see
<figureCitation id="136C2A679701FF9DFE6DFCDCFD98BEB0" box="[495,604,871,896]" captionStart="FIGURE 15" captionStartId="12.[151,250,986,1008]" captionTargetBox="[157,1418,388,950]" captionTargetId="figure@12.[151,1436,373,965]" captionTargetPageId="12" captionText="FIGURE 15. Variation in preserved holotype and paratype male Maratus combustus sp. nov. specimens: A, holotype male; B, paratype male #1; C, paratype male #2." httpUri="https://zenodo.org/record/2588699/files/figure.png" pageId="8" pageNumber="89">Figure 15</figureCitation>
AC); area below with thick cover of burnt orange/tan scales; triangular shape of gold scales bordered by black scales situated near posterior margin of dorsal opisthosomal plate; posterior margin of opisthosoma with distinctive stripes of scales alternating in colour between tan and white and bordered by black; prominent tufts of long setae extend beyond rear margin of dorsal opisthosomal plate, the colour of each of them corresponding with above posterior marginal stripes (
<figureCitation id="136C2A679701FF9DFCC8FC4CFC0AB920" box="[842,974,1015,1040]" captionStart="FIGURE 10" captionStartId="8.[151,250,1968,1990]" captionTargetBox="[168,1429,1095,1944]" captionTargetId="figure@8.[151,1436,1078,1952]" captionTargetPageId="8" captionText="FIGURE 10. Habitus images of living holotype Maratus combustus sp. nov. (WAMT147351): A, anterior view; B, dorsal view; C, lateral view; D: anterolateral view; E, dorsolateral view; F, lateral view." httpUri="https://zenodo.org/record/2588689/files/figure.png" pageId="8" pageNumber="89">Figure 10B</figureCitation>
).
</paragraph>
<caption id="DF28666A9701FF9DFF15F80BFC03BAD5" httpUri="https://zenodo.org/record/2588689/files/figure.png" pageId="8" pageNumber="89" startId="8.[151,250,1968,1990]" targetBox="[168,1429,1095,1944]" targetPageId="8">
<paragraph id="8BE836E29701FF9DFF15F80BFC03BAD5" blockId="8.[151,1436,1968,2021]" pageId="8" pageNumber="89">
<emphasis id="B923EAF09701FF9DFF15F80BFEE5BAF5" bold="true" box="[151,289,1968,1990]" pageId="8" pageNumber="89">FIGURE 10.</emphasis>
Habitus images of living holotype
<taxonomicName id="4C574D619701FF9DFD10F80AFC9FBAF6" authority="Schubert, 2019" authorityName="Schubert" authorityYear="2019" box="[658,859,1969,1990]" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="combustus" status="sp. nov.">
<emphasis id="B923EAF09701FF9DFD10F80AFC9FBAF6" box="[658,859,1969,1990]" italics="true" pageId="8" pageNumber="89">Maratus combustus</emphasis>
</taxonomicName>
<emphasis id="B923EAF09701FF9DFCE1F80AFC72BAF6" bold="true" box="[867,950,1969,1990]" pageId="8" pageNumber="89">
<taxonomicNameLabel id="A210578B9701FF9DFCE1F80AFC72BAF6" box="[867,950,1969,1990]" pageId="8" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(WAMT147351): A, anterior view; B, dorsal view; C, lateral view; D: anterolateral view; E, dorsolateral view; F, lateral view.
</paragraph>
</caption>
<caption id="DF28666A9700FF9CFF15FA4AFDE6BB16" httpUri="https://zenodo.org/record/2588691/files/figure.png" pageId="9" pageNumber="90" startId="9.[151,250,1521,1543]" targetBox="[214,1386,233,1440]" targetPageId="9">
<paragraph id="8BE836E29700FF9CFF15FA4AFDE6BB16" blockId="9.[151,1435,1521,1574]" pageId="9" pageNumber="90">
<emphasis id="B923EAF09700FF9CFF15FA4AFEDABB36" bold="true" box="[151,286,1521,1543]" pageId="9" pageNumber="90">FIGURE 11.</emphasis>
Habitus images of preserved holotype
<taxonomicName id="4C574D619700FF9CFD29FA49FCB6BB37" authority="Schubert, 2019" authorityName="Schubert" authorityYear="2019" box="[683,882,1522,1543]" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="combustus" status="sp. nov.">
<emphasis id="B923EAF09700FF9CFD29FA49FCB6BB37" box="[683,882,1522,1543]" italics="true" pageId="9" pageNumber="90">Maratus combustus</emphasis>
</taxonomicName>
<emphasis id="B923EAF09700FF9CFCFBFA49FC0DBB37" bold="true" box="[889,969,1522,1543]" pageId="9" pageNumber="90">
<taxonomicNameLabel id="A210578B9700FF9CFCFBFA49FC0DBB37" box="[889,969,1522,1543]" pageId="9" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(WAMT147351): A, dorsal view; B, ventral view; C, anterior view; D, lateral view.
</paragraph>
</caption>
<paragraph id="8BE836E29700FF9CFF45F9E8FC73BBE7" blockId="9.[151,1436,1619,2003]" pageId="9" pageNumber="90">
Lateral opisthosomal flaps large, lobate and ornamented with bright orange scales at upper margin, otherwise covered with dull purple scales; prominent tuft of long white setae project posteriorly from lateral margin of each opisthosomal flap, visible during courtship (
<figureCitation id="136C2A679700FF9CFD2AF920FCE0BB83" box="[680,804,1691,1716]" captionStart="FIGURE 2" captionStartId="2.[151,250,645,667]" captionTargetBox="[151,1435,208,604]" captionTargetId="figure@2.[151,1435,208,604]" captionTargetPageId="2" captionText="FIGURE 2. Living male paratype Maratus aquilus sp. nov., M. combustus sp. nov., and M. felinus sp. nov. performing courtship display to nearby female: A, Courtship display of male Maratus aquilus sp. nov.; B, courtship display of Maratus combustus sp. nov.; C, courtship display of Maratus felinus sp. nov." httpUri="https://zenodo.org/record/2588673/files/figure.png" pageId="9" pageNumber="90">Figure 2B</figureCitation>
); tuft of white setae above black spinnerets; ventral opisthosoma dark grey and scattered with short, creamy coloured setae.
</paragraph>
<paragraph id="8BE836E29700FF9CFF45F95FFF7EBAE3" blockId="9.[151,1436,1619,2003]" pageId="9" pageNumber="90">
Legs I and II subequal in length; legs III and IV longer; legs III by far longest. Legs I, II, and IV ringed with alternating bands of dark brown and white setae; legs III with less distinct and less frequent rings; anterior surface of femora of legs III with a thick cover of short, white setae; femora, patellae, and tibiae of legs III ornamented with heavy fringing of long white setae; tarsi of legs III tipped with white setae. Pedipalp covered dorsally with long off-white setae; relatively large palpal bulb with retrolateral sperm duct loop, large retrobasal tegular lobe, finger-like retrolateral tibial apophysis; embolus with pointed inner apex beneath thicker outer apex (
<figureCitation id="136C2A679700FF9CFA89F82CFABCBA9F" box="[1291,1400,1943,1968]" captionStart="FIGURE 12" captionStartId="10.[151,250,1038,1060]" captionTargetBox="[150,1436,344,998]" captionTargetId="figure@10.[150,1436,344,998]" captionTargetPageId="10" captionText="FIGURE 12. Images of preserved holotype Maratus combustus sp. nov. pedipalp structure (WAMT147351): A, retrolateral view; B, ventral view showing embolic disc and tegular lobe. C, prolateral view showing finger-like retrolateral tibial apophysis and retrolateral sperm duct loop." httpUri="https://zenodo.org/record/2588693/files/figure.png" pageId="9" pageNumber="90">Figure 12</figureCitation>
A C).
</paragraph>
<paragraph id="8BE836E29703FF9FFF45FF2CFDE6BC2C" blockId="10.[151,1436,151,285]" pageId="10" pageNumber="91">
<emphasis id="B923EAF09703FF9FFF45FF2CFEDABD80" bold="true" box="[199,286,151,176]" pageId="10" pageNumber="91">Female</emphasis>
(WAM-T147352). Ocular quadrangle region with incomplete cover of red-brown and white setae; area posterior to ocular quadrangle black and mostly glabrous; incomplete bands of white setae extend from PME to rear margin of carapace; area below dark brown to black and mostly glabrous; rim of carapace glabrous and lacking marginal band (
<figureCitation id="136C2A679703FF9FFEC4FEBFFDD7BC2C" box="[326,531,260,285]" captionStart="FIGURE 13" captionStartId="10.[151,250,1964,1986]" captionTargetBox="[151,1436,1186,1927]" captionTargetId="figure@10.[151,1436,1186,1927]" captionTargetPageId="10" captionText="FIGURE 13. Habitus images of living paratype #2 female Maratus combustus sp. nov. (WAM-T147352): A, anterior view; B, dorsal view; C, lateral view; D: anterodorsal view; E, dorsolateral view; F, dorsolateral view." httpUri="https://zenodo.org/record/2588695/files/figure.png" pageId="10" pageNumber="91">Figure 13A and B</figureCitation>
).
</paragraph>
<caption id="DF28666A9703FF9FFF15FBB5FD89B951" httpUri="https://zenodo.org/record/2588693/files/figure.png" pageId="10" pageNumber="91" startId="10.[151,250,1038,1060]" targetBox="[150,1436,344,998]" targetPageId="10">
<paragraph id="8BE836E29703FF9FFF15FBB5FD89B951" blockId="10.[151,1436,1038,1121]" pageId="10" pageNumber="91">
<emphasis id="B923EAF09703FF9FFF15FBB5FEE4B913" bold="true" box="[151,288,1038,1060]" pageId="10" pageNumber="91">FIGURE 12.</emphasis>
Images of preserved holotype
<taxonomicName id="4C574D619703FF9FFDDDFBB4FCE3B913" authority="Schubert, 2019" authorityName="Schubert" authorityYear="2019" box="[607,807,1038,1060]" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="combustus" status="sp. nov.">
<emphasis id="B923EAF09703FF9FFDDDFBB4FCE3B913" box="[607,807,1038,1060]" italics="true" pageId="10" pageNumber="91">Maratus combustus</emphasis>
</taxonomicName>
<emphasis id="B923EAF09703FF9FFCACFBB5FC44B913" bold="true" box="[814,896,1038,1059]" pageId="10" pageNumber="91">
<taxonomicNameLabel id="A210578B9703FF9FFCACFBB5FC44B913" box="[814,896,1038,1059]" pageId="10" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
pedipalp structure (WAMT147351): A, retrolateral view; B, ventral view showing embolic disc and tegular lobe. C, prolateral view showing finger-like retrolateral tibial apophysis and retrolateral sperm duct loop.
</paragraph>
</caption>
<caption id="DF28666A9703FF9FFF15F817FB87BAD1" httpUri="https://zenodo.org/record/2588695/files/figure.png" pageId="10" pageNumber="91" startId="10.[151,250,1964,1986]" targetBox="[151,1436,1186,1927]" targetPageId="10">
<paragraph id="8BE836E29703FF9FFF15F817FB87BAD1" blockId="10.[151,1435,1964,2017]" pageId="10" pageNumber="91">
<emphasis id="B923EAF09703FF9FFF15F817FEDBBAF1" bold="true" box="[151,287,1964,1986]" pageId="10" pageNumber="91">FIGURE 13.</emphasis>
Habitus images of living paratype #2 female
<taxonomicName id="4C574D619703FF9FFD6FF816FC70BAF1" authority="Schubert, 2019" authorityName="Schubert" authorityYear="2019" box="[749,948,1964,1986]" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="91" phylum="Arthropoda" rank="species" species="combustus" status="sp. nov.">
<emphasis id="B923EAF09703FF9FFD6FF816FC70BAF1" box="[749,948,1964,1986]" italics="true" pageId="10" pageNumber="91">Maratus combustus</emphasis>
</taxonomicName>
<emphasis id="B923EAF09703FF9FFC38F817FBCEBAF1" bold="true" box="[954,1034,1964,1985]" pageId="10" pageNumber="91">
<taxonomicNameLabel id="A210578B9703FF9FFC38F817FBCEBAF1" box="[954,1034,1964,1985]" pageId="10" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(WAM-T147352): A, anterior view; B, dorsal view; C, lateral view; D: anterodorsal view; E, dorsolateral view; F, dorsolateral view.
</paragraph>
</caption>
<caption id="DF28666A9702FF9EFF15F9A6FCBEBB62" httpUri="https://zenodo.org/record/2588697/files/figure.png" pageId="11" pageNumber="92" startId="11.[151,250,1565,1587]" targetBox="[157,1435,216,1540]" targetPageId="11">
<paragraph id="8BE836E29702FF9EFF15F9A6FCBEBB62" blockId="11.[151,1436,1565,1618]" pageId="11" pageNumber="92">
<emphasis id="B923EAF09702FF9EFF15F9A6FEE4BB03" bold="true" box="[151,288,1565,1587]" pageId="11" pageNumber="92">FIGURE 14.</emphasis>
Habitus and epigyne images of preserved paratype #2 female
<taxonomicName id="4C574D619702FF9EFC19F9A5FBA7BB03" authority="Schubert, 2019" authorityName="Schubert" authorityYear="2019" box="[923,1123,1566,1587]" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="92" phylum="Arthropoda" rank="species" species="combustus" status="sp. nov.">
<emphasis id="B923EAF09702FF9EFC19F9A5FBA7BB03" box="[923,1123,1566,1587]" italics="true" pageId="11" pageNumber="92">Maratus combustus</emphasis>
</taxonomicName>
<emphasis id="B923EAF09702FF9EFBEBF9A5FB7EBB03" bold="true" box="[1129,1210,1566,1587]" pageId="11" pageNumber="92">
<taxonomicNameLabel id="A210578B9702FF9EFBEBF9A5FB7EBB03" box="[1129,1210,1566,1587]" pageId="11" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(WAM-T147352): A, ventral view; B, lateral view; C, dorsal view; D, ventral view of epigyne.
</paragraph>
</caption>
<paragraph id="8BE836E29702FF9EFF45F9C2FE18BA11" blockId="11.[151,1436,1657,2007]" pageId="11" pageNumber="92">
Anterior eyes bordered with red-brown and white scales dorsally, and white scales laterally and below; PME closer to PLE than to ALE; long white setae project downwards from below AME forming triangular shape; clypeus covered with short, white setae (
<figureCitation id="136C2A679702FF9EFDDFF97AFD20BBEA" box="[605,740,1729,1754]" captionStart="FIGURE 13" captionStartId="10.[151,250,1964,1986]" captionTargetBox="[151,1436,1186,1927]" captionTargetId="figure@10.[151,1436,1186,1927]" captionTargetPageId="10" captionText="FIGURE 13. Habitus images of living paratype #2 female Maratus combustus sp. nov. (WAM-T147352): A, anterior view; B, dorsal view; C, lateral view; D: anterodorsal view; E, dorsolateral view; F, dorsolateral view." httpUri="https://zenodo.org/record/2588695/files/figure.png" pageId="11" pageNumber="92">Figure 13A</figureCitation>
); chelicerae dark brown to black and glabrous; coxae, endites, and labium dark brown, translucent and mostly glabrous; sternum medium-dark brown (
<figureCitation id="136C2A679702FF9EFB16F95EFADFBBCE" box="[1172,1307,1765,1790]" captionStart="FIGURE 14" captionStartId="11.[151,250,1565,1587]" captionTargetBox="[157,1435,216,1540]" captionTargetId="figure@11.[151,1435,193,1544]" captionTargetPageId="11" captionText="FIGURE 14. Habitus and epigyne images of preserved paratype #2 female Maratus combustus sp. nov. (WAM-T147352): A, ventral view; B, lateral view; C, dorsal view; D, ventral view of epigyne." httpUri="https://zenodo.org/record/2588697/files/figure.png" pageId="11" pageNumber="92">Figure 14A</figureCitation>
) with light covering of fine, white setae.
</paragraph>
<paragraph id="8BE836E29702FF9EFF45F896FD42BA82" blockId="11.[151,1436,1657,2007]" pageId="11" pageNumber="92">
Dorsal opisthosoma light brown with incomplete cover of off-white and dark brown setae; central spot of offwhite setae on dorsal opisthosoma; lateral and posterior edges of dorsal opisthosoma bordered with narrow band of light brown and white setae; spinnerets dark brown (
<figureCitation id="136C2A679702FF9EFD65F8CEFCA9BABE" box="[743,877,1909,1934]" captionStart="FIGURE 14" captionStartId="11.[151,250,1565,1587]" captionTargetBox="[157,1435,216,1540]" captionTargetId="figure@11.[151,1435,193,1544]" captionTargetPageId="11" captionText="FIGURE 14. Habitus and epigyne images of preserved paratype #2 female Maratus combustus sp. nov. (WAM-T147352): A, ventral view; B, lateral view; C, dorsal view; D, ventral view of epigyne." httpUri="https://zenodo.org/record/2588697/files/figure.png" pageId="11" pageNumber="92">Figure 14A</figureCitation>
); lateral and ventral opisthosoma light brown and covered with small dark spots (
<figureCitation id="136C2A679702FF9EFE76F822FDBEBA82" box="[500,634,1945,1970]" captionStart="FIGURE 14" captionStartId="11.[151,250,1565,1587]" captionTargetBox="[157,1435,216,1540]" captionTargetId="figure@11.[151,1435,193,1544]" captionTargetPageId="11" captionText="FIGURE 14. Habitus and epigyne images of preserved paratype #2 female Maratus combustus sp. nov. (WAM-T147352): A, ventral view; B, lateral view; C, dorsal view; D, ventral view of epigyne." httpUri="https://zenodo.org/record/2588697/files/figure.png" pageId="11" pageNumber="92">Figure 14A</figureCitation>
).
</paragraph>
<paragraph id="8BE836E29702FF99FF45F804FDBDBC70" blockId="11.[151,1436,1657,2007]" lastBlockId="12.[151,1437,151,320]" lastPageId="12" lastPageNumber="93" pageId="11" pageNumber="92">
Legs I and II subequal in length; legs III and IV longer; legs III longest. Integument of each leg with alternating bands of light brown and dark brown; ventral femoral surface of each leg distinctively marked with dark bands (
<figureCitation id="136C2A679705FF99FF1CFF07FEEDBDE4" box="[158,297,188,213]" captionStart="FIGURE 14" captionStartId="11.[151,250,1565,1587]" captionTargetBox="[157,1435,216,1540]" captionTargetId="figure@11.[151,1435,193,1544]" captionTargetPageId="11" captionText="FIGURE 14. Habitus and epigyne images of preserved paratype #2 female Maratus combustus sp. nov. (WAM-T147352): A, ventral view; B, lateral view; C, dorsal view; D, ventral view of epigyne." httpUri="https://zenodo.org/record/2588697/files/figure.png" pageId="12" pageNumber="93">Figure 14A</figureCitation>
); each leg with light cover of short white setae; tarsi of each leg tipped with dark brown setae; pedipalps light brown and translucent with incomplete cover of off-white setae (
<figureCitation id="136C2A679705FF99FBA2FF64FB63BDC8" box="[1056,1191,223,248]" captionStart="FIGURE 13" captionStartId="10.[151,250,1964,1986]" captionTargetBox="[151,1436,1186,1927]" captionTargetId="figure@10.[151,1436,1186,1927]" captionTargetPageId="10" captionText="FIGURE 13. Habitus images of living paratype #2 female Maratus combustus sp. nov. (WAM-T147352): A, anterior view; B, dorsal view; C, lateral view; D: anterodorsal view; E, dorsolateral view; F, dorsolateral view." httpUri="https://zenodo.org/record/2588695/files/figure.png" pageId="12" pageNumber="93">Figure 13A</figureCitation>
). Epigynum with pair of large ovate fossae separated by septum; ovate posterior spermatheca behind each fossa; sclerotized ducts present anterior to each spermatheca (
<figureCitation id="136C2A679705FF99FE64FE9CFDA9BC70" box="[486,621,295,320]" captionStart="FIGURE 14" captionStartId="11.[151,250,1565,1587]" captionTargetBox="[157,1435,216,1540]" captionTargetId="figure@11.[151,1435,193,1544]" captionTargetPageId="11" captionText="FIGURE 14. Habitus and epigyne images of preserved paratype #2 female Maratus combustus sp. nov. (WAM-T147352): A, ventral view; B, lateral view; C, dorsal view; D, ventral view of epigyne." httpUri="https://zenodo.org/record/2588697/files/figure.png" pageId="12" pageNumber="93">Figure 14D</figureCitation>
).
</paragraph>
<caption id="DF28666A9705FF99FF15FC61FDE1B93F" httpUri="https://zenodo.org/record/2588699/files/figure.png" pageId="12" pageNumber="93" startId="12.[151,250,986,1008]" targetBox="[157,1418,388,950]" targetPageId="12">
<paragraph id="8BE836E29705FF99FF15FC61FDE1B93F" blockId="12.[151,1435,986,1039]" pageId="12" pageNumber="93">
<emphasis id="B923EAF09705FF99FF15FC61FEDBBEDF" bold="true" box="[151,287,986,1008]" pageId="12" pageNumber="93">FIGURE 15.</emphasis>
Variation in preserved holotype and paratype male
<taxonomicName id="4C574D619705FF99FCABFC60FC34BEC0" authority="Schubert, 2019" authorityName="Schubert" authorityYear="2019" box="[809,1008,987,1008]" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="combustus" status="sp. nov.">
<emphasis id="B923EAF09705FF99FCABFC60FC34BEC0" box="[809,1008,987,1008]" italics="true" pageId="12" pageNumber="93">Maratus combustus</emphasis>
</taxonomicName>
<emphasis id="B923EAF09705FF99FC77FC60FB07BEC0" bold="true" box="[1013,1219,986,1008]" pageId="12" pageNumber="93">
<taxonomicNameLabel id="A210578B9705FF99FC77FC60FB81BEC0" box="[1013,1093,987,1008]" pageId="12" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
specimens:
</emphasis>
A, holotype male; B, paratype male #1; C, paratype male #2.
</paragraph>
</caption>
<caption id="DF28666A9705FF99FF15F8F4FEA4BAB3" httpUri="https://zenodo.org/record/2588701/files/figure.png" pageId="12" pageNumber="93" startId="12.[151,250,1871,1893]" targetBox="[172,1377,1090,1822]" targetPageId="12">
<paragraph id="8BE836E29705FF99FF15F8F4FEA4BAB3" blockId="12.[151,1435,1871,1923]" pageId="12" pageNumber="93">
<emphasis id="B923EAF09705FF99FF15F8F4FEE0BA54" bold="true" box="[151,292,1871,1893]" pageId="12" pageNumber="93">FIGURE 16.</emphasis>
Variation in preserved paratype female
<taxonomicName id="4C574D619705FF99FD51F8EBFC5BBA54" authority="Schubert, 2019" authorityName="Schubert" authorityYear="2019" box="[723,927,1871,1893]" class="Arachnida" family="Salticidae" genus="Maratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="combustus" status="sp. nov.">
<emphasis id="B923EAF09705FF99FD51F8EBFC5BBA54" box="[723,927,1871,1893]" italics="true" pageId="12" pageNumber="93">Maratus combustus</emphasis>
</taxonomicName>
<emphasis id="B923EAF09705FF99FC2BF8F4FC3ABA54" bold="true" box="[937,1022,1871,1892]" pageId="12" pageNumber="93">
<taxonomicNameLabel id="A210578B9705FF99FC2BF8F4FC3ABA54" box="[937,1022,1871,1892]" pageId="12" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens: A, paratype female #1; B, paratype female #2.
</paragraph>
</caption>
<paragraph id="8BE836E29705FF98FF45F80BFDACBDE5" blockId="12.[151,1436,1968,2029]" lastBlockId="13.[151,1437,151,789]" lastPageId="13" lastPageNumber="94" pageId="12" pageNumber="93">
<emphasis id="B923EAF09705FF99FF45F80BFEFBBAF9" bold="true" box="[199,319,1968,1993]" pageId="12" pageNumber="93">Variation.</emphasis>
The size of the triangular shape of scales on the lower dorsal opisthosoma of males may vary. The angle at which the tracts of scales at the anterior margin of the dorsal opisthosoma converge towards the middle varies in the additional
<typeStatus id="54EC88409704FF98FE28FF23FDCFBD80" box="[426,523,152,176]" pageId="13" pageNumber="94" type="paratype">paratype</typeStatus>
male specimens. The scales on the surface of the dorsal opisthosoma between these lines may be worn (
<figureCitation id="136C2A679704FF98FE36FF07FDE4BDE5" box="[436,544,188,213]" captionStart="FIGURE 15" captionStartId="12.[151,250,986,1008]" captionTargetBox="[157,1418,388,950]" captionTargetId="figure@12.[151,1436,373,965]" captionTargetPageId="12" captionText="FIGURE 15. Variation in preserved holotype and paratype male Maratus combustus sp. nov. specimens: A, holotype male; B, paratype male #1; C, paratype male #2." httpUri="https://zenodo.org/record/2588699/files/figure.png" pageId="13" pageNumber="94">Figure 15</figureCitation>
AC).
</paragraph>
<paragraph id="8BE836E29704FF98FF45FF64FC04BC2C" blockId="13.[151,1437,151,789]" pageId="13" pageNumber="94">The width of the light-coloured band on the lateral opisthosoma of females may vary. The median thoracic tract may be more or less distinct due to scales rubbing off (16A and B).</paragraph>
<paragraph id="8BE836E29704FF98FF45FE9CFAB4BCB8" blockId="13.[151,1437,151,789]" pageId="13" pageNumber="94">
<emphasis id="B923EAF09704FF98FF45FE9CFE62BC70" bold="true" box="[199,422,295,320]" pageId="13" pageNumber="94">Dimensions. Male.</emphasis>
Total length: 3.673.78 (3.72±0.03, n=3). Carapace length 1.831.96 (1.91±0.04, n=3). Opisthosoma length 1.721.86 (1.80± 0.04, n=3). Leg I length: 2.842.86 (2.85±0.01, n=3). Leg II length: 2.84 2.87 (2.85±0.01, n=3). Leg III length: 4.1 94.31 (4.25±0.03, n=3). Leg IV length: 3.743.81 (3.76±0.02, n=3).
</paragraph>
<paragraph id="8BE836E29704FF98FF45FE2FFF17BF28" blockId="13.[151,1437,151,789]" pageId="13" pageNumber="94">
<emphasis id="B923EAF09704FF98FF45FE2FFE7ABC9D" bold="true" box="[199,446,404,429]" pageId="13" pageNumber="94">Dimensions. Female.</emphasis>
Total length: 4.954.99 (4.97±0.02, n=2). Carapace length 2.112.27 (2.19 ±0.08, n=2). Opisthosoma length 2.722.84 (2.78± 0.06, n=2). Leg I length: 2.682.72 (2.70±0.02, n=2). Leg II length: 2.652.67 (2.66±0.01, n=2). Leg III length: 4.434.48 (4.46±0.03, n=2). Leg IV length: 3.924.00 (3.96±0.04, n=2).
</paragraph>
</subSubSection>
<subSubSection id="C34D65699704FF98FF45FD9FFDB7BE25" pageId="13" pageNumber="94" type="distribution">
<paragraph id="8BE836E29704FF98FF45FD9FFBCCBF0D" blockId="13.[151,1437,151,789]" box="[199,1032,548,573]" pageId="13" pageNumber="94">
<emphasis id="B923EAF09704FF98FF45FD9FFEA4BF0D" bold="true" box="[199,352,548,573]" pageId="13" pageNumber="94">Distribution.</emphasis>
Known only from near Mount Romance,
<collectingRegion id="4993F8009704FF98FCB5FD9EFBC7BF0D" box="[823,1027,548,573]" country="Australia" name="Western Australia" pageId="13" pageNumber="94">Western Australia</collectingRegion>
.
</paragraph>
<paragraph id="8BE836E29704FF98FF45FDFCFDB7BE25" blockId="13.[151,1437,151,789]" pageId="13" pageNumber="94">
<emphasis id="B923EAF09704FF98FF45FDFCFE60BF50" bold="true" box="[199,420,583,608]" pageId="13" pageNumber="94">Courtship display.</emphasis>
The male elevates legs III, holding them straight upwards and holding them in place for the duration of the courtship display. The pedipalps are held apart, exposing the chelicerae. The opisthosoma is elevated and waved, and the lateral opisthosoma flaps are extended. As the female approaches, the lateral opisthosomal flaps are repeatedly retracted and extended whist the opisthosoma remains elevated and waving. (
<figureCitation id="136C2A679704FF98FF1CFD6CFEDCBFC0" box="[158,280,727,752]" captionStart="FIGURE 2" captionStartId="2.[151,250,645,667]" captionTargetBox="[151,1435,208,604]" captionTargetId="figure@2.[151,1435,208,604]" captionTargetPageId="2" captionText="FIGURE 2. Living male paratype Maratus aquilus sp. nov., M. combustus sp. nov., and M. felinus sp. nov. performing courtship display to nearby female: A, Courtship display of male Maratus aquilus sp. nov.; B, courtship display of Maratus combustus sp. nov.; C, courtship display of Maratus felinus sp. nov." httpUri="https://zenodo.org/record/2588673/files/figure.png" pageId="13" pageNumber="94">Figure 2B</figureCitation>
). (Note: only partial courtship display observed, the males may exhibit a more complete courtship display with multiple modes of courtship).
</paragraph>
</subSubSection>
</treatment>
</document>