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<document id="C2FB95801CE7842CAB7A94D677155F5D" ID-DOI="10.1046/j.1096-3642.2002.00016.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="10114642" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630432778602" checkinUser="carolina" docAuthor="Corbera, Jordi" docDate="2002" docId="03D1B43EFFABFFD6FE9E5AF8FB45C397" docLanguage="en" docName="j.1096-3642.2002.00016.x.pdf" docOrigin="Zoological Journal of the Linnean Society 134 (4)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1046/j.1096-3642.2002.00016.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Speleocuma guanche Corbera 2002, sp. nov." docType="treatment" docVersion="4" lastPageNumber="457" masterDocId="FFE8CC46FFA9FFD2FF9A5803FFBFC02C" masterDocTitle="Amphi-Atlantic distribution of the Mancocumatinae (Cumacea: Bodotriidae), with description of a new genus dwelling in marine lava caves of Tenerife (Canary Islands)" masterLastPageNumber="461" masterPageNumber="453" pageNumber="455" updateTime="1700038151179" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="B200B453FE7F762E9B009F3D62B3806E">Amphi-Atlantic distribution of the Mancocumatinae (Cumacea: Bodotriidae), with description of a new genus dwelling in marine lava caves of Tenerife (Canary Islands)</mods:title>
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<mods:namePart id="91175AEDFCEBB62929BF97BAF7F42CC6">Corbera, Jordi</mods:namePart>
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<mods:typeOfResource id="C643E33773A3A4702F793EE39205E0F7">text</mods:typeOfResource>
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<mods:title id="9483C6EF0616E620DB883143C0000C30">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="E8BE449C4895D32B4D3DDD889A6587E5">2002</mods:date>
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<mods:number id="2DDEEE4021A8E588A072E64BD46047B7">2002-04-30</mods:number>
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<mods:number id="876DE6F7242E530B6D4CA18C4DCB22A9">134</mods:number>
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<mods:number id="1DE48E45A450C3B49CFB1B793A4C891E">4</mods:number>
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<mods:url id="11B822B154C952FE57FAAADB81E82DC0">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1046/j.1096-3642.2002.00016.x</mods:url>
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<mods:classification id="69F2F838FFE48FD0E67EDB8674CCC3F0">journal article</mods:classification>
<mods:identifier id="067EEF05695B037E584DA3786B45611D" type="DOI">10.1046/j.1096-3642.2002.00016.x</mods:identifier>
<mods:identifier id="3794FB84AE8B948E60A7F9B2F8BE4F6B" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03D1B43EFFABFFD6FE9E5AF8FB45C397" LSID="urn:lsid:plazi:treatment:03D1B43EFFABFFD6FE9E5AF8FB45C397" httpUri="http://treatment.plazi.org/id/03D1B43EFFABFFD6FE9E5AF8FB45C397" lastPageId="4" lastPageNumber="457" pageId="2" pageNumber="455">
<subSubSection id="C36256A3FFABFFD0FE9E5AF8FD39C33F" box="[260,646,763,787]" pageId="2" pageNumber="455" type="nomenclature">
<paragraph id="8BC70528FFABFFD0FE9E5AF8FD39C33F" blockId="2.[260,646,763,787]" box="[260,646,763,787]" pageId="2" pageNumber="455">
<heading id="D08FB244FFABFFD0FE9E5AF8FD39C33F" box="[260,646,763,787]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="455" reason="2">
<emphasis id="B90CD93AFFABFFD0FE9E5AF8FD39C33F" bold="true" box="[260,646,763,787]" pageId="2" pageNumber="455">
<taxonomicName id="4C787EABFFABFFD0FE9E5AF8FDA7C33F" ID-CoL="6YX3R" authority="Corbera, 2002" authorityName="Corbera" authorityYear="2002" box="[260,536,763,787]" class="Malacostraca" family="Bodotriidae" genus="Speleocuma" kingdom="Animalia" order="Cumacea" pageId="2" pageNumber="455" phylum="Arthropoda" rank="species" species="guanche" status="sp. nov.">
<emphasis id="B90CD93AFFABFFD0FE9E5AF8FDA7C33F" bold="true" box="[260,536,763,787]" italics="true" pageId="2" pageNumber="455">Speleocuma guanche</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F6441FFABFFD0FDBB5AF8FD39C33F" box="[545,646,763,787]" pageId="2" pageNumber="455" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36256A3FFABFFD6FF0B5B28FB45C397" lastPageId="4" lastPageNumber="457" pageId="2" pageNumber="455" type="materials_examined">
<paragraph id="8BC70528FFABFFD0FF0B5B28FD51C3D4" blockId="2.[145,761,810,1016]" pageId="2" pageNumber="455">
<emphasis id="B90CD93AFFABFFD0FF0B5B28FEFEC313" box="[145,321,810,832]" italics="true" pageId="2" pageNumber="455">Type material.</emphasis>
<materialsCitation id="3B100F75FFABFFD0FECE5B28FE74C3D4" collectionCode="ICM, T" county="Cueva Escalon" location="Tenerife" municipality="Agua Dulce cave" pageId="2" pageNumber="455" specimenCode="CUM-0027, CUM-0028" specimenCount="21" specimenCount-adult="12" specimenCount-immature="2" stateProvince="Canarias" typeStatus="holotype">
<collectingCounty id="62A67DA4FFABFFD0FECE5B28FDB3C36C" box="[340,524,810,832]" pageId="2" pageNumber="455">Cueva Escalón</collectingCounty>
,
<collectingMunicipality id="6BA39F52FFABFFD0FDB85B28FD46C36C" box="[546,761,810,832]" pageId="2" pageNumber="455">Agua Dulce cave</collectingMunicipality>
complex,
<location id="8EA753F3FFABFFD0FE9E5B4AFEDBC373" LSID="urn:lsid:plazi:treatment:03D1B43EFFABFFD6FE9E5AF8FB45C397:8EA753F3FFABFFD0FE9E5B4AFEDBC373" box="[260,356,841,863]" county="Cueva Escalon" municipality="Agua Dulce cave" name="Tenerife" pageId="2" pageNumber="455" stateProvince="Canarias">Tenerife</location>
,
<collectingRegion id="49BCCBCAFFABFFD0FEED5B4AFD8EC373" box="[375,561,841,863]" country="Spain" name="Canarias" pageId="2" pageNumber="455">Canary Islands</collectingRegion>
, sta. AD 1,
<specimenCount id="9D7ECEA1FFABFFD0FD505B49FD46C373" box="[714,761,842,863]" pageId="2" pageNumber="455" type="generic">7 m.</specimenCount>
<typeStatus id="54C3BB8AFFABFFD0FF0B5B6BFF44C352" box="[145,251,872,894]" pageId="2" pageNumber="455" type="holotype">Holotype</typeStatus>
: ovigerous female dissected on two slides [
<collectionCode id="ED699DEDFFABFFD0FF0D5B84FF70C3B0" box="[151,207,903,924]" country="Spain" httpUri="http://grbio.org/cool/x81t-7j82" name="Instituto de Ciencias del Mar" pageId="2" pageNumber="455">ICM</collectionCode>
,
<specimenCode id="DBDEAD53FFABFFD0FF445B85FEDFC3B0" box="[222,352,902,924]" collectionCode="CUM-" pageId="2" pageNumber="455">CUM-0027</specimenCode>
].
<typeStatus id="54C3BB8AFFABFFD0FEEE5B84FE54C3B0" box="[372,491,903,924]" pageId="2" pageNumber="455" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7ECEA1FFABFFD0FE605B84FDE4C3B0" box="[506,603,902,924]" pageId="2" pageNumber="455" type="adult">11 adult</specimenCount>
females, two preadult females,
<specimenCount id="9D7ECEA1FFABFFD0FEF95BA6FDA6C397" box="[355,537,933,955]" pageId="2" pageNumber="455" type="immature">three immature</specimenCount>
females,
<specimenCount id="9D7ECEA1FFABFFD0FD105BA5FD46C397" box="[650,761,933,955]" pageId="2" pageNumber="455" type="adult">two adult</specimenCount>
males,
<specimenCount id="9D7ECEA1FFABFFD0FF705BC7FE2CC3F6" box="[234,403,964,986]" pageId="2" pageNumber="455" type="immature">one immature</specimenCount>
male and one manca [
<collectionCode id="ED699DEDFFABFFD0FD275BC7FD4AC3F5" box="[701,757,964,985]" country="Spain" httpUri="http://grbio.org/cool/x81t-7j82" name="Instituto de Ciencias del Mar" pageId="2" pageNumber="455">ICM</collectionCode>
,
<specimenCode id="DBDEAD53FFABFFD0FF0B5BE1FEACC3D4" box="[145,275,994,1016]" collectionCode="CUM-" pageId="2" pageNumber="455">CUM-0028</specimenCode>
]. Collected by
<collectionCode id="ED699DEDFFABFFD0FE235BE0FE74C3D4" box="[441,459,995,1016]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="455">T</collectionCode>
</materialsCitation>
. Cruz on
<date id="FFC623E8FFABFFD0FDA25BE0FD56C3D4" box="[568,745,995,1016]" pageId="2" pageNumber="455" value="1997-08-01">August 1, 1997</date>
.
</paragraph>
<paragraph id="8BC70528FFABFFD0FF0B5C13FEC1C493" blockId="2.[145,761,1040,1216]" pageId="2" pageNumber="455">
<emphasis id="B90CD93AFFABFFD0FF0B5C13FEFFC409" box="[145,320,1040,1061]" italics="true" pageId="2" pageNumber="455">Other material.</emphasis>
Cueva Sin Fin, sta. AD 3, three ovigerous females, four preadult females,
<specimenCount id="9D7ECEA1FFABFFD0FDA55C33FD11C469" box="[575,686,1071,1093]" pageId="2" pageNumber="455" type="adult">one adult</specimenCount>
male, three preadult males,
<specimenCount id="9D7ECEA1FFABFFD0FE0A5C4DFDA2C448" box="[400,541,1102,1124]" pageId="2" pageNumber="455" type="juvenile">one juvenile</specimenCount>
. Cueva Escachada, sta. AD 5, three ovigerous females, three preadult females,
<specimenCount id="9D7ECEA1FFABFFD0FF645C8FFECEC48D" box="[254,369,1163,1185]" pageId="2" pageNumber="455" type="adult">one adult</specimenCount>
male,
<specimenCount id="9D7ECEA1FFABFFD0FE575C8FFDCAC48D" box="[461,629,1163,1185]" pageId="2" pageNumber="455" type="immature">one immature</specimenCount>
male,
<specimenCount id="9D7ECEA1FFABFFD0FD4B5C8FFF50C4EC" pageId="2" pageNumber="455" type="juvenile">one juvenile</specimenCount>
, one manca.
</paragraph>
<caption id="DF0755A0FFABFFD0FCB35CC1FC54C503" ID-DOI="http://doi.org/10.5281/zenodo.10131657" ID-Zenodo-Dep="10131657" httpUri="https://zenodo.org/record/10131657/files/figure.png" pageId="2" pageNumber="455" startId="2.[809,888,1218,1239]" targetBox="[814,1422,195,1186]" targetPageId="2">
<paragraph id="8BC70528FFABFFD0FCB35CC1FC54C503" blockId="2.[809,1425,1218,1327]" pageId="2" pageNumber="455">
<emphasis id="B90CD93AFFABFFD0FCB35CC1FB14C4FB" bold="true" box="[809,1195,1218,1239]" pageId="2" pageNumber="455">
Figure 2.
<taxonomicName id="4C787EABFFABFFD0FC335CC1FB14C4FB" authority="Corbera, 2002" authorityName="Corbera" authorityYear="2002" box="[937,1195,1218,1239]" class="Malacostraca" family="Bodotriidae" genus="Speleocuma" kingdom="Animalia" order="Cumacea" pageId="2" pageNumber="455" phylum="Arthropoda" rank="species" species="guanche" status="gen. et sp. nov.">
<emphasis id="B90CD93AFFABFFD0FC335CC1FB14C4FB" bold="true" box="[937,1195,1218,1239]" italics="true" pageId="2" pageNumber="455">Speleocuma guanche</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A23F6441FFABFFD0FB245CC1FA34C4FB" box="[1214,1419,1218,1239]" pageId="2" pageNumber="455" rank="species">
<emphasis id="B90CD93AFFABFFD0FB245CC1FB50C4FB" bold="true" box="[1214,1263,1218,1239]" pageId="2" pageNumber="455">gen.</emphasis>
et
<emphasis id="B90CD93AFFABFFD0FAB25CC1FA34C4FB" bold="true" box="[1320,1419,1218,1239]" pageId="2" pageNumber="455">sp. nov.</emphasis>
</taxonomicNameLabel>
, preadult female, SEM micrographs. A, Anterior half of the carapace. B, Microstructure of the carapace showing the denticulate scales.
</paragraph>
</caption>
<paragraph id="8BC70528FFABFFD0FF0B5CDBFD9CC520" blockId="2.[145,761,1240,1292]" pageId="2" pageNumber="455">
<emphasis id="B90CD93AFFABFFD0FF0B5CDBFEB7C4C1" box="[145,264,1240,1261]" italics="true" pageId="2" pageNumber="455">Etymology</emphasis>
.
<taxonomicName id="4C787EABFFABFFD0FE8F5CDBFEC6C4C1" authorityName="Corbera" authorityYear="2002" box="[277,377,1240,1261]" class="Malacostraca" family="Bodotriidae" genus="Speleocuma" kingdom="Animalia" order="Cumacea" pageId="2" pageNumber="455" phylum="Arthropoda" rank="species" species="guanche">
<emphasis id="B90CD93AFFABFFD0FE8F5CDBFEC6C4C1" box="[277,377,1240,1261]" italics="true" pageId="2" pageNumber="455">Guanche</emphasis>
</taxonomicName>
was the name of the early human settlers of the Canary Archipelago.
</paragraph>
<paragraph id="8BC70528FFABFFD0FF0B5D26FDA1C662" blockId="2.[145,761,1316,1614]" pageId="2" pageNumber="455">
<emphasis id="B90CD93AFFABFFD0FF0B5D26FEA5C516" box="[145,282,1317,1338]" italics="true" pageId="2" pageNumber="455">Description.</emphasis>
Adult female,
<quantity id="4C80A8CDFFABFFD0FE6C5D26FDF6C516" box="[502,585,1317,1338]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="455" unit="mm" value="2.9">2.9 mm</quantity>
total length (
<figureCitation id="134319ADFFABFFD0FF035D40FF4FC575" box="[153,240,1347,1369]" captionStart="Figure 1" captionStartId="1.[159,238,1855,1876]" captionTargetBox="[178,1439,919,1833]" captionTargetId="figure-316@1.[164,1439,913,1833]" captionTargetPageId="1" captionText="Figure 1. Speleocuma guanche gen. et sp. nov. A, Ovigerous female, whole animal in lateral view. B, Adult male, whole animal in lateral view." figureDoi="http://doi.org/10.5281/zenodo.10131652" httpUri="https://zenodo.org/record/10131652/files/figure.png" pageId="2" pageNumber="455">Fig. 1a</figureCitation>
). Integument covered by scales that have their margin finely denticulate (
<figureCitation id="134319ADFFABFFD0FD905D61FDC8C554" box="[522,631,1378,1400]" captionStart="Figure 2" captionStartId="2.[809,888,1218,1239]" captionTargetBox="[814,1422,195,1186]" captionTargetId="figure-552@2.[814,1422,195,1186]" captionTargetPageId="2" captionText="Figure 2. Speleocuma guanche gen. et sp. nov., preadult female, SEM micrographs. A, Anterior half of the carapace. B, Microstructure of the carapace showing the denticulate scales." figureDoi="http://doi.org/10.5281/zenodo.10131657" httpUri="https://zenodo.org/record/10131657/files/figure.png" pageId="2" pageNumber="455">Figs 2a.b</figureCitation>
). Carapace slightly longer than high, about one-fourth total body length; middorsal with a double row of small denticles on anterior third, behind eyelobe. Pseudorostral lobes short, extending beyond and meeting anterior to eyelobe. Antennal notch wide, with anterolateral angle and anterolateral margin denticulate. Exopods on pereopod 1–3 and on maxilliped 3.
</paragraph>
<paragraph id="8BC70528FFABFFD6FF0B5E65FC53C0D6" blockId="2.[145,761,1638,1906]" lastBlockId="4.[809,1426,197,618]" lastPageId="4" lastPageNumber="457" pageId="2" pageNumber="455">
Peduncle of first antenna 3-segmented (
<figureCitation id="134319ADFFABFFD0FDFD5E65FD05C651" box="[615,698,1638,1661]" captionStart="Figure 3" captionStartId="3.[159,238,1687,1708]" captionTargetBox="[176,1424,192,1654]" captionTargetId="figure-27@3.[154,1434,181,1670]" captionTargetPageId="3" captionText="Figure 3. Speleocuma guanche gen. et sp. nov., ovigerous female A, Antenna 1. B, Maxilliped 2. C, Maxilliped 3. D, Pereopod 2. E, Pereopod 3. F, Pereopod 4. G, Last abdominal somite and uropod." figureDoi="http://doi.org/10.5281/zenodo.10131659" httpUri="https://zenodo.org/record/10131659/files/figure.png" pageId="2" pageNumber="455">Fig. 3a</figureCitation>
), 3rd segment a little longer than 2nd, main fagellum 2- segmented with two aesthetascs and long seta distally. Mandibles pars incisiva with four teeth; 13 long setae between pars incisa and truncate pars molaris (
<figureCitation id="134319ADFFABFFD0FD5D5EE2FF7CC73A" captionStart="Figure 4" captionStartId="4.[145,224,1854,1875]" captionTargetBox="[176,1400,1035,1823]" captionTargetId="figure-433@4.[150,1427,1027,1845]" captionTargetPageId="4" captionText="Figure 4. Speleocuma guanche gen. et sp. nov., ovigerous female.A, Right mandible. B, Pars incisa of the left mandible showing the lacinia mobilis. C, Maxillule. D, Maxilla. E, Maxilliped 1." figureDoi="http://doi.org/10.5281/zenodo.10131661" httpUri="https://zenodo.org/record/10131661/files/figure.png" pageId="2" pageNumber="455">Figs 4a.b</figureCitation>
). Maxilla 1 (
<figureCitation id="134319ADFFABFFD0FED45F03FE25C73A" box="[334,410,1792,1814]" captionStart="Figure 4" captionStartId="4.[145,224,1854,1875]" captionTargetBox="[176,1400,1035,1823]" captionTargetId="figure-433@4.[150,1427,1027,1845]" captionTargetPageId="4" captionText="Figure 4. Speleocuma guanche gen. et sp. nov., ovigerous female.A, Right mandible. B, Pars incisa of the left mandible showing the lacinia mobilis. C, Maxillule. D, Maxilla. E, Maxilliped 1." figureDoi="http://doi.org/10.5281/zenodo.10131661" httpUri="https://zenodo.org/record/10131661/files/figure.png" pageId="2" pageNumber="455">Fig. 4c</figureCitation>
) with simple robust acuminate setae on protopod, palp with two unequal filaments. Maxilla 2 (
<figureCitation id="134319ADFFABFFD0FE865F3EFECCC77F" box="[284,371,1853,1875]" captionStart="Figure 4" captionStartId="4.[145,224,1854,1875]" captionTargetBox="[176,1400,1035,1823]" captionTargetId="figure-433@4.[150,1427,1027,1845]" captionTargetPageId="4" captionText="Figure 4. Speleocuma guanche gen. et sp. nov., ovigerous female.A, Right mandible. B, Pars incisa of the left mandible showing the lacinia mobilis. C, Maxillule. D, Maxilla. E, Maxilliped 1." figureDoi="http://doi.org/10.5281/zenodo.10131661" httpUri="https://zenodo.org/record/10131661/files/figure.png" pageId="2" pageNumber="455">Fig. 4d</figureCitation>
) with simple setae on the two endites and inner margin of protopod. Maxilliped 1 (
<figureCitation id="134319ADFFABFFD0FCAB5D72FC39C5AB" box="[817,902,1393,1415]" captionStart="Figure 4" captionStartId="4.[145,224,1854,1875]" captionTargetBox="[176,1400,1035,1823]" captionTargetId="figure-433@4.[150,1427,1027,1845]" captionTargetPageId="4" captionText="Figure 4. Speleocuma guanche gen. et sp. nov., ovigerous female.A, Right mandible. B, Pars incisa of the left mandible showing the lacinia mobilis. C, Maxillule. D, Maxilla. E, Maxilliped 1." figureDoi="http://doi.org/10.5281/zenodo.10131661" httpUri="https://zenodo.org/record/10131661/files/figure.png" pageId="2" pageNumber="455">Fig. 4e</figureCitation>
) with distal process of basis reaching the carpus, with acuminate setae and a strong serrate seta; flattened hand-like setae on inner margin of carpus; propodus twice length of dactylus. Basis of maxilliped 2 (
<figureCitation id="134319ADFFABFFD0FC555DEFFB9CC62E" box="[975,1059,1516,1538]" captionStart="Figure 3" captionStartId="3.[159,238,1687,1708]" captionTargetBox="[176,1424,192,1654]" captionTargetId="figure-27@3.[154,1434,181,1670]" captionTargetPageId="3" captionText="Figure 3. Speleocuma guanche gen. et sp. nov., ovigerous female A, Antenna 1. B, Maxilliped 2. C, Maxilliped 3. D, Pereopod 2. E, Pereopod 3. F, Pereopod 4. G, Last abdominal somite and uropod." figureDoi="http://doi.org/10.5281/zenodo.10131659" httpUri="https://zenodo.org/record/10131659/files/figure.png" pageId="2" pageNumber="455">Fig. 3b</figureCitation>
) longer than the rest of limb, outer distal margin with one stout plumose seta and a simple seta, inner distal margin with one stout plumose seta; ischium short; carpus and propodus of similar length, both with setae on inner margin; dactylus half length of propodus with two distal claws. Basis of maxilliped 3 (
<figureCitation id="134319ADFFABFFD0FC7B5EA7FB92C696" box="[993,1069,1700,1722]" captionStart="Figure 3" captionStartId="3.[159,238,1687,1708]" captionTargetBox="[176,1424,192,1654]" captionTargetId="figure-27@3.[154,1434,181,1670]" captionTargetPageId="3" captionText="Figure 3. Speleocuma guanche gen. et sp. nov., ovigerous female A, Antenna 1. B, Maxilliped 2. C, Maxilliped 3. D, Pereopod 2. E, Pereopod 3. F, Pereopod 4. G, Last abdominal somite and uropod." figureDoi="http://doi.org/10.5281/zenodo.10131659" httpUri="https://zenodo.org/record/10131659/files/figure.png" pageId="2" pageNumber="455">Fig. 3c</figureCitation>
) slightly longer than combined length of other articles; outer distal process not developed, with three long plumose setae; inner margin with seven plumose setae; ischium very short and unarmed; merus and carpus with one and three inner plumose setae, respectively; propodus as long as carpus and twice as long as dactylus. Pereopod 1 only the basis remaining. Basis of pereopod 2 (
<figureCitation id="134319ADFFADFFD6FD0158C6FD4FC0F7" box="[667,752,197,219]" captionStart="Figure 3" captionStartId="3.[159,238,1687,1708]" captionTargetBox="[176,1424,192,1654]" captionTargetId="figure-27@3.[154,1434,181,1670]" captionTargetPageId="3" captionText="Figure 3. Speleocuma guanche gen. et sp. nov., ovigerous female A, Antenna 1. B, Maxilliped 2. C, Maxilliped 3. D, Pereopod 2. E, Pereopod 3. F, Pereopod 4. G, Last abdominal somite and uropod." figureDoi="http://doi.org/10.5281/zenodo.10131659" httpUri="https://zenodo.org/record/10131659/files/figure.png" pageId="4" pageNumber="457">Fig. 3d</figureCitation>
) slightly shorter than the combined lengths of other articles, with two spine-like setae on inner distal margin; ischium very short; merus twice as long as ischium; carpus longer than merus with some setae on distal margin; propodus about half length of carpus; dactylus as long as combined length of carpus and propodus, with a distal spine-like seta longer than article, three small setae on distal margin and some other on the inner margin. Basis of pereopod 3 (
<figureCitation id="134319ADFFADFFD6FD5659DAFF14C222" captionStart="Figure 3" captionStartId="3.[159,238,1687,1708]" captionTargetBox="[176,1424,192,1654]" captionTargetId="figure-27@3.[154,1434,181,1670]" captionTargetPageId="3" captionText="Figure 3. Speleocuma guanche gen. et sp. nov., ovigerous female A, Antenna 1. B, Maxilliped 2. C, Maxilliped 3. D, Pereopod 2. E, Pereopod 3. F, Pereopod 4. G, Last abdominal somite and uropod." figureDoi="http://doi.org/10.5281/zenodo.10131659" httpUri="https://zenodo.org/record/10131659/files/figure.png" pageId="4" pageNumber="457">Fig. 3e</figureCitation>
) about 1.3 times longer than rest of limb; ischium short with two setae; merus slightly longer than ischium; carpus as long as ischium and merus combined, with two distal long setae; propodus half long as carpus with two distal setae; dactylus very short with simple seta distally. Pereopod 4 (
<figureCitation id="134319ADFFADFFD6FDD65A92FD28C28B" box="[588,663,657,679]" captionStart="Figure 3" captionStartId="3.[159,238,1687,1708]" captionTargetBox="[176,1424,192,1654]" captionTargetId="figure-27@3.[154,1434,181,1670]" captionTargetPageId="3" captionText="Figure 3. Speleocuma guanche gen. et sp. nov., ovigerous female A, Antenna 1. B, Maxilliped 2. C, Maxilliped 3. D, Pereopod 2. E, Pereopod 3. F, Pereopod 4. G, Last abdominal somite and uropod." figureDoi="http://doi.org/10.5281/zenodo.10131659" httpUri="https://zenodo.org/record/10131659/files/figure.png" pageId="4" pageNumber="457">Fig. 3f</figureCitation>
) similar to 3 except for some differences in setation and lack of exopod. Peduncle of uropod (
<figureCitation id="134319ADFFADFFD6FE445ACDFD91C2C9" box="[478,558,718,741]" captionStart="Figure 3" captionStartId="3.[159,238,1687,1708]" captionTargetBox="[176,1424,192,1654]" captionTargetId="figure-27@3.[154,1434,181,1670]" captionTargetPageId="3" captionText="Figure 3. Speleocuma guanche gen. et sp. nov., ovigerous female A, Antenna 1. B, Maxilliped 2. C, Maxilliped 3. D, Pereopod 2. E, Pereopod 3. F, Pereopod 4. G, Last abdominal somite and uropod." figureDoi="http://doi.org/10.5281/zenodo.10131659" httpUri="https://zenodo.org/record/10131659/files/figure.png" pageId="4" pageNumber="457">
Fig.
<quantity id="4C80A8CDFFADFFD6FD895ACCFD91C2C9" box="[531,558,719,741]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="4" pageNumber="457" unit="g" value="3.0">3g</quantity>
</figureCitation>
) longer than last abdominal somite, with six acuminate setae on inner margin. Endopod as long as peduncle, 2-segmented; proximal segment twice as long as distal, with five acuminate setae on inner margin; distal segment with three acuminate setae on inner margin and a long distal spine-like seta. Exopod shorter than endopod, 2- segmented, second segment with two long distal spine- like setae, three setae on outer margin and other one on inner margin.
</paragraph>
<caption id="DF0755A0FFAAFFD1FF055E94FC48C6E5" ID-DOI="http://doi.org/10.5281/zenodo.10131659" ID-Zenodo-Dep="10131659" httpUri="https://zenodo.org/record/10131659/files/figure.png" pageId="3" pageNumber="456" startId="3.[159,238,1687,1708]" targetBox="[176,1424,192,1654]" targetPageId="3">
<paragraph id="8BC70528FFAAFFD1FF055E94FC48C6E5" blockId="3.[159,1439,1687,1737]" pageId="3" pageNumber="456">
<emphasis id="B90CD93AFFAAFFD1FF055E94FDB4C680" bold="true" box="[159,523,1687,1708]" pageId="3" pageNumber="456">
Figure 3.
<taxonomicName id="4C787EABFFAAFFD1FE8E5E94FDB4C680" authority="Corbera, 2002" authorityName="Corbera" authorityYear="2002" box="[276,523,1687,1708]" class="Malacostraca" family="Bodotriidae" genus="Speleocuma" kingdom="Animalia" order="Cumacea" pageId="3" pageNumber="456" phylum="Arthropoda" rank="species" species="guanche" status="gen. et sp. nov.">
<emphasis id="B90CD93AFFAAFFD1FE8E5E94FDB4C680" bold="true" box="[276,523,1687,1708]" italics="true" pageId="3" pageNumber="456">Speleocuma guanche</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A23F6441FFAAFFD1FD8E5E94FD7EC680" box="[532,705,1687,1708]" pageId="3" pageNumber="456" rank="species">
<emphasis id="B90CD93AFFAAFFD1FD8E5E94FDFAC680" bold="true" box="[532,581,1687,1708]" pageId="3" pageNumber="456">gen.</emphasis>
et
<emphasis id="B90CD93AFFAAFFD1FDF35E94FD7EC680" bold="true" box="[617,705,1687,1708]" pageId="3" pageNumber="456">sp. nov.</emphasis>
</taxonomicNameLabel>
, ovigerous female A, Antenna 1. B, Maxilliped 2. C, Maxilliped 3. D, Pereopod 2. E, Pereopod 3. F, Pereopod 4. G, Last abdominal somite and uropod.
</paragraph>
</caption>
<paragraph id="8BC70528FFADFFD6FCDB5901FBCDC246" blockId="4.[809,1426,197,618]" pageId="4" pageNumber="457">
Adult male (
<figureCitation id="134319ADFFADFFD6FC405901FB8BC134" box="[986,1076,258,280]" captionStart="Figure 1" captionStartId="1.[159,238,1855,1876]" captionTargetBox="[178,1439,919,1833]" captionTargetId="figure-316@1.[164,1439,913,1833]" captionTargetPageId="1" captionText="Figure 1. Speleocuma guanche gen. et sp. nov. A, Ovigerous female, whole animal in lateral view. B, Adult male, whole animal in lateral view." figureDoi="http://doi.org/10.5281/zenodo.10131652" httpUri="https://zenodo.org/record/10131652/files/figure.png" pageId="4" pageNumber="457">Figs 1b</figureCitation>
and
<figureCitation id="134319ADFFADFFD6FBED5901FB39C134" box="[1143,1158,258,280]" captionStart="Figure 5" captionStartId="5.[159,238,1787,1808]" captionTargetBox="[172,1433,194,1755]" captionTargetId="figure-24@5.[156,1436,186,1773]" captionTargetPageId="5" captionText="Figure 5. Speleocuma guanche gen. et sp. nov., adult male. A, Antennae 1 and 2. B, Mandible. C, Maxilula. D, Maxila. E, Maxilliped 2. F, Maxilliped 3. G, Pereopod 2. H, Pereopod 3. I, Pereopod 4. J, Pleopod. K, Uropod." figureDoi="http://doi.org/10.5281/zenodo.10131663" httpUri="https://zenodo.org/record/10131663/files/figure.png" pageId="4" pageNumber="457">5</figureCitation>
),
<quantity id="4C80A8CDFFADFFD6FB3B5900FB4CC134" box="[1185,1267,259,280]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="457" unit="mm" value="2.4">2.4 mm</quantity>
total length, considerably smaller than female. Differing from the female as follows: integument with small scattered hairs. Middorsal line of carapace smooth. Pseudorostral lobes very short, not reaching beyond eyelobe. Peduncle of the second antenna (
<figureCitation id="134319ADFFADFFD6FB07599FFB54C19E" box="[1181,1259,412,434]" captionStart="Figure 5" captionStartId="5.[159,238,1787,1808]" captionTargetBox="[172,1433,194,1755]" captionTargetId="figure-24@5.[156,1436,186,1773]" captionTargetPageId="5" captionText="Figure 5. Speleocuma guanche gen. et sp. nov., adult male. A, Antennae 1 and 2. B, Mandible. C, Maxilula. D, Maxila. E, Maxilliped 2. F, Maxilliped 3. G, Pereopod 2. H, Pereopod 3. I, Pereopod 4. J, Pleopod. K, Uropod." figureDoi="http://doi.org/10.5281/zenodo.10131663" httpUri="https://zenodo.org/record/10131663/files/figure.png" pageId="4" pageNumber="457">Fig. 5a</figureCitation>
) 5-segmented, with sensorial setae on fourth and fifth segments; flagellum short, scarcely reaching fourth free thoracic somite. Abdomen with two pairs of pleopods. Peduncle of uropod (
<figureCitation id="134319ADFFADFFD6FC3D5A15FC46C200" box="[935,1017,534,556]" captionStart="Figure 5" captionStartId="5.[159,238,1787,1808]" captionTargetBox="[172,1433,194,1755]" captionTargetId="figure-24@5.[156,1436,186,1773]" captionTargetPageId="5" captionText="Figure 5. Speleocuma guanche gen. et sp. nov., adult male. A, Antennae 1 and 2. B, Mandible. C, Maxilula. D, Maxila. E, Maxilliped 2. F, Maxilliped 3. G, Pereopod 2. H, Pereopod 3. I, Pereopod 4. J, Pleopod. K, Uropod." figureDoi="http://doi.org/10.5281/zenodo.10131663" httpUri="https://zenodo.org/record/10131663/files/figure.png" pageId="4" pageNumber="457">Fig. 5k</figureCitation>
) with 11 acuminate setae on inner edge, endopod with nine acuminate setae on first segment and four on second.
</paragraph>
<paragraph id="8BC70528FFADFFD6FCB35A92FB45C397" blockId="4.[809,1425,657,955]" pageId="4" pageNumber="457">
<emphasis id="B90CD93AFFADFFD6FCB35A92FC31C28A" box="[809,910,657,678]" italics="true" pageId="4" pageNumber="457">Remarks</emphasis>
. A previous record of a cumacean species from the Jameos del Agua, Lanzarote, assigned to the family
<taxonomicName id="4C787EABFFADFFD6FC335ACDFB94C2C8" authorityName="Sars" authorityYear="1878" box="[937,1067,718,740]" class="Malacostraca" family="Leuconidae" kingdom="Animalia" order="Cumacea" pageId="4" pageNumber="457" phylum="Arthropoda" rank="family">Leuconidae</taxonomicName>
(see
<bibRefCitation id="EFE978D9FFADFFD6FBF05ACDFA3BC2C8" author="Garcia-Valdecasas A" box="[1130,1412,718,740]" pageId="4" pageNumber="457" pagination="3 - 56" refId="ref3566" refString="Garcia-Valdecasas A. 1985. Estudio faunistico de la cueva submarina ' Tunel de la Atlantida', Jameos del Agua, Lanzarote. Naturalia Hispanica 27: 3 - 56." type="journal article" year="1985">García-Valdecasas, 1985</bibRefCitation>
), is not supported here because of the presence of a developed eye. Apparently, it could belong to the subfamily
<taxonomicName id="4C787EABFFADFFD6FC3F5B29FBD7C36C" box="[933,1128,810,832]" class="Malacostraca" family="Leuconidae" kingdom="Animalia" order="Cumacea" pageId="4" pageNumber="457" phylum="Arthropoda" rank="subFamily" subFamily="Mancocumatidae">Mancocumatidae</taxonomicName>
and perhaps to the new genus described above, since it displays two pairs of pleopods (see
<bibRefCitation id="EFE978D9FFADFFD6FB9E5B6BFA97C352" author="Garcia-Valdecasas A" box="[1028,1320,872,894]" pageId="4" pageNumber="457" pagination="3 - 56" refId="ref3566" refString="Garcia-Valdecasas A. 1985. Estudio faunistico de la cueva submarina ' Tunel de la Atlantida', Jameos del Agua, Lanzarote. Naturalia Hispanica 27: 3 - 56." type="journal article" year="1985">García-Valdecasas, 1985</bibRefCitation>
: fig. 8). However, a more concise identification is not possible due to the lack of a detailed description.
</paragraph>
</subSubSection>
</treatment>
</document>
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