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<document ID-DOI="http://doi.org/10.5281/zenodo.5131755" ID-GBIF-Dataset="09f66541-18e3-4044-b553-7aab04763396" ID-GBIF-Taxon="183733962" ID-ISSN="1982-0224" ID-Zenodo-Dep="5131755" checkinTime="1616760176976" checkinUser="carolina" docAuthor="Menezes, Naércio A. &amp; de Lucena, Carlos Alberto S." docDate="2014" docId="03C4D93834683F51FEECFAB9FABF6466" docLanguage="en" docName="NeotropIchthyol.12.2.193-228.pdf" docOrigin="Neotropical Ichthyology 12 (2)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252014000200193&amp;lng=en&amp;tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Charax metae Eigenmann 1922" docType="treatment" docVersion="6" lastPageId="19" lastPageNumber="212" masterDocId="FFFDA140347A3F42FFFBFE69FFCB6623" masterDocTitle="A taxonomic review of the species of Charax Scopoli, 1777 (Teleostei: Characidae: Characinae) with description of a new species from the rio Negro bearing superficial neuromasts on body scales, Amazon basin, Brazil" masterLastPageNumber="228" masterPageNumber="193" pageId="18" pageNumber="211" updateTime="1634698513480" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A taxonomic review of the species of Charax Scopoli, 1777 (Teleostei: Characidae: Characinae) with description of a new species from the rio Negro bearing superficial neuromasts on body scales, Amazon basin, Brazil</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Menezes, Naércio A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>de Lucena, Carlos Alberto S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>12</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>193</mods:start>
<mods:end>228</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252014000200193&amp;lng=en&amp;tlng=en</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/1982-0224-20130175</mods:identifier>
<mods:identifier type="GBIF-Dataset">09f66541-18e3-4044-b553-7aab04763396</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">4638995</mods:identifier>
<mods:identifier type="ZooBank">2CFAB1CE-D283-483B-8C2B-87D95AEADE70</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5131755" ID-GBIF-Taxon="183733962" ID-Zenodo-Dep="5131755" LSID="urn:lsid:plazi:treatment:03C4D93834683F51FEECFAB9FABF6466" httpUri="http://treatment.plazi.org/id/03C4D93834683F51FEECFAB9FABF6466" lastPageId="19" lastPageNumber="212" pageId="18" pageNumber="211">
<subSubSection box="[279,636,1232,1256]" pageId="18" pageNumber="211" type="nomenclature">
<paragraph blockId="18.[279,636,1232,1289]" box="[279,636,1232,1256]" pageId="18" pageNumber="211">
<heading box="[279,636,1232,1256]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="211" reason="2">
<taxonomicName authority="Eigenmann, 1922" authorityName="Eigenmann" authorityYear="1922" box="[279,636,1232,1256]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="metae">
<emphasis bold="true" box="[279,636,1232,1256]" pageId="18" pageNumber="211">
<emphasis bold="true" box="[279,434,1232,1256]" italics="true" pageId="18" pageNumber="211">Charax metae</emphasis>
Eigenmann, 1922
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="211" type="description">
<paragraph blockId="18.[279,636,1232,1289]" box="[422,492,1264,1289]" pageId="18" pageNumber="211">
<figureCitation box="[422,492,1264,1289]" captionStart="Fig" captionStartId="18.[136,175,767,791]" captionTargetBox="[138,1461,176,744]" captionTargetId="figure-544@18.[138,1461,176,744]" captionTargetPageId="18" captionText="Fig. 16. Charax metae, CAS 69117, 87 mm SL, Colômbia, Meta, Puerto Barrigona (=Barrigón) at head of navigation east of Bogota." figureDoi="http://doi.org/10.5281/zenodo.4639030" httpUri="https://zenodo.org/record/4639030/files/figure.png" pageId="18" pageNumber="211">Fig. 16</figureCitation>
</paragraph>
<paragraph blockId="18.[136,779,1329,1545]" pageId="18" pageNumber="211">
<taxonomicName authority="Eigenmann, 1922: 238" authorityName="Eigenmann" authorityPageNumber="238" authorityYear="1922" box="[136,514,1329,1353]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="metae">
<emphasis box="[136,280,1329,1353]" italics="true" pageId="18" pageNumber="211">Charax metae</emphasis>
Eigenmann, 1922: 238
</taxonomicName>
(original description,
<typeStatus box="[734,779,1329,1353]" pageId="18" pageNumber="211">type</typeStatus>
locality:
<collectingCountry box="[265,372,1361,1385]" name="Colombia" pageId="18" pageNumber="211">Colombia</collectingCountry>
, Barrigón, Orinoco system, río
<collectingRegion box="[695,754,1361,1385]" country="Colombia" name="Meta" pageId="18" pageNumber="211">Meta</collectingRegion>
). -
<collectorName box="[174,236,1393,1417]" pageId="18" pageNumber="211">Ibarra</collectorName>
&amp;
<collectorName box="[265,347,1393,1417]" pageId="18" pageNumber="211">Stewart</collectorName>
, 1987: 22 (listed in
<typeStatus box="[544,589,1393,1417]" pageId="18" pageNumber="211">type</typeStatus>
catalog). -Lucena, 1987: 44 (diagnosis; description; distribution). -Lucena, 1989: 201 (in key to species). -
<collectorName box="[476,521,1457,1481]" pageId="18" pageNumber="211">Vari</collectorName>
&amp;
<collectorName box="[550,614,1457,1480]" pageId="18" pageNumber="211">Howe</collectorName>
, 1991: 13 (listed in
<typeStatus box="[198,243,1489,1513]" pageId="18" pageNumber="211">type</typeStatus>
catalog). -
<collectorName box="[347,425,1489,1512]" pageId="18" pageNumber="211">Lucena</collectorName>
&amp;
<collectorName box="[455,550,1489,1512]" pageId="18" pageNumber="211">Menezes</collectorName>
, 2003: 201 (maximum length; distribution).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="211" type="diagnosis">
<paragraph blockId="18.[136,779,1584,1961]" lastBlockId="18.[816,1459,912,1033]" pageId="18" pageNumber="211">
<emphasis bold="true" box="[136,254,1584,1608]" pageId="18" pageNumber="211">Diagnosis.</emphasis>
<taxonomicName authorityName="Eigenmann" authorityYear="1922" box="[259,410,1585,1609]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="metae">
<emphasis box="[259,410,1585,1609]" italics="true" pageId="18" pageNumber="211">Charax metae</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Menezes &amp; de Lucena" authorityYear="2014" box="[423,590,1585,1609]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="delimai">
<emphasis box="[423,590,1585,1609]" italics="true" pageId="18" pageNumber="211">Charax delimai</emphasis>
</taxonomicName>
,
<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1870" box="[602,710,1585,1609]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="tectifer">
<emphasis box="[602,710,1585,1609]" italics="true" pageId="18" pageNumber="211">C. tectifer</emphasis>
</taxonomicName>
, some specimens of
<taxonomicName authorityName=", Lucena" authorityYear="1987" box="[289,419,1617,1641]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="gibbosus">
<emphasis box="[289,419,1617,1641]" italics="true" pageId="18" pageNumber="211">C. gibbosus</emphasis>
</taxonomicName>
,
<taxonomicName box="[434,567,1617,1640]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="rupununi">
<emphasis box="[434,567,1617,1640]" italics="true" pageId="18" pageNumber="211">C. rupununi</emphasis>
</taxonomicName>
, and
<taxonomicName box="[631,737,1617,1641]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="condei">
<emphasis box="[631,649,1617,1640]" italics="true" pageId="18" pageNumber="211">C</emphasis>
.
<emphasis box="[665,737,1617,1641]" italics="true" pageId="18" pageNumber="211">condei</emphasis>
</taxonomicName>
are the only species of the genus bearing teeth on the ectopterygoid bone.
<taxonomicName authorityName="Eigenmann" authorityYear="1922" box="[355,505,1681,1705]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="metae">
<emphasis box="[355,505,1681,1705]" italics="true" pageId="18" pageNumber="211">Charax metae</emphasis>
</taxonomicName>
differs from
<taxonomicName box="[644,746,1681,1705]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="condei">
<emphasis box="[644,746,1681,1705]" italics="true" pageId="18" pageNumber="211">C. condei</emphasis>
</taxonomicName>
by having the lateral line complete (
<emphasis box="[483,552,1713,1736]" italics="true" pageId="18" pageNumber="211">versus</emphasis>
incomplete), from
<taxonomicName authorityName=", Lucena" authorityYear="1987" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="gibbosus">
<emphasis box="[754,772,1713,1736]" italics="true" pageId="18" pageNumber="211">C</emphasis>
.
<emphasis box="[136,232,1745,1769]" italics="true" pageId="18" pageNumber="211">gibbosus</emphasis>
</taxonomicName>
and
<taxonomicName box="[292,426,1745,1768]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="rupununi">
<emphasis box="[292,310,1745,1768]" italics="true" pageId="18" pageNumber="211">C</emphasis>
.
<emphasis box="[327,426,1745,1768]" italics="true" pageId="18" pageNumber="211">rupununi</emphasis>
</taxonomicName>
which have the anal-fin origin anterior to the vertical through the dorsal-fin origin in having the anal-fin origin through vertical or slightly posterior to, dorsal fin-origin.
<taxonomicName authorityName="Eigenmann" authorityYear="1922" box="[324,474,1841,1865]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="metae">
<emphasis box="[324,474,1841,1865]" italics="true" pageId="18" pageNumber="211">Charax metae</emphasis>
</taxonomicName>
can be readily distinguished from
<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1870" box="[196,305,1873,1897]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="tectifer">
<emphasis box="[196,214,1873,1896]" italics="true" pageId="18" pageNumber="211">C</emphasis>
.
<emphasis box="[228,305,1873,1897]" italics="true" pageId="18" pageNumber="211">tectifer</emphasis>
</taxonomicName>
by the presence (
<emphasis box="[500,529,1873,1896]" italics="true" pageId="18" pageNumber="211">vs.</emphasis>
absence) of the dorsoventrally arranged superficial neuromasts on the trunk scales (
<figureCitation box="[144,208,1937,1961]" captionStart="Fig" captionStartId="3.[129,168,1194,1218]" captionTargetBox="[129,788,734,1181]" captionTargetId="figure-60@3.[129,772,745,1174]" captionText="Fig. 2. Superficial neuromasts (black arrow) on ventral body scales of Charax metae, CAS 69117, 59 mm SL." figureDoi="http://doi.org/10.5281/zenodo.5131735" httpUri="https://zenodo.org/record/5131735/files/figure.png" pageId="18" pageNumber="211">Fig. 2</figureCitation>
), except the lateral line and from
<taxonomicName authorityName="Menezes &amp; de Lucena" authorityYear="2014" box="[559,668,1937,1961]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="211" phylum="Chordata" rank="species" species="delimai">
<emphasis box="[559,668,1937,1961]" italics="true" pageId="18" pageNumber="211">C. delimai</emphasis>
</taxonomicName>
by having more ectopterygoid teeth (20-50
<emphasis box="[1172,1202,913,936]" italics="true" pageId="18" pageNumber="211">vs.</emphasis>
3-15), fewer transverse scale rows between the humeral spot and the supracleithrum (7-9
<emphasis box="[866,895,977,1000]" italics="true" pageId="18" pageNumber="211">vs.</emphasis>
10-12) and the humeral spot distance 41.6-47% of SL (
<emphasis box="[825,853,1009,1032]" italics="true" pageId="18" pageNumber="211">vs.</emphasis>
48-51% of SL,
<figureCitation box="[1011,1070,1009,1033]" captionStart="Fig" captionStartId="3.[807,846,1194,1218]" captionTargetBox="[129,1443,738,1174]" captionTargetPageId="3" captionText="Fig. 3. Humeral spot distance as a function of standard length for species of Charax." figureDoi="http://doi.org/10.5281/zenodo.4639001" httpUri="https://zenodo.org/record/4639001/files/figure.png" pageId="18" pageNumber="211">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="19" lastPageNumber="212" pageId="18" pageNumber="211" type="description">
<paragraph blockId="18.[816,1460,1072,1961]" pageId="18" pageNumber="211">
<emphasis bold="true" box="[816,961,1072,1096]" pageId="18" pageNumber="211">Description</emphasis>
. Morphometrics of examined specimens presented in
<tableCitation box="[950,1030,1105,1129]" captionStart="Table 9" captionStartId="19.[129,191,177,201]" captionTargetPageId="19" captionText="Table 9. Morphometrics of Charax metae. Specimens are from CAS 69117." pageId="18" pageNumber="211">Table 9</tableCitation>
. Body elongate, moderately large (
<quantity metricMagnitude="-2" metricUnit="m" metricValue="6.8" metricValueMax="9.9" metricValueMin="3.7" pageId="18" pageNumber="211" unit="mm" value="68.0" valueMax="99.0" valueMin="37.0">37-99 mm</quantity>
SL), compressed and moderately deep. Greatest body depth slightly in advance of dorsal-fin origin. Dorsal profile of head and body convex from tip of snout to anterior portion of fontanel, concave from that point to base of supraoccipital spine, convex from that point to dorsal-fin origin, nearly straight along dorsal-fin base, straight to slightly convex from end of dorsal-fin base to caudal peduncle and slightly concave along caudal peduncle. Ventral profile of head and body convex from tip of lower jaw to anal-fin origin, nearly straight along anal-fin base and slightly concave from end of anal-fin base to beginning of procurrent rays. Snout pointed. Lower jaw included in upper jaw when mouth closed. Maxilla extending slightly posterior to vertical through posterior border of orbit.
</paragraph>
<paragraph blockId="18.[816,1460,1072,1961]" pageId="18" pageNumber="211">
Dorsal-fin rays ii,
<quantity box="[1053,1094,1585,1609]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="18" pageNumber="211" unit="in" value="9.0">9 in</quantity>
all specimens; posterior most ray unbranched. Adipose fin present. Unbranched anal-fin rays iv or v, usually iv, branched rays 39-43, 41.2. Pectoral-fin rays i, 14-16, 14.7. Tips of longest pectoral-fin rays reaching to about vertical through middle of pelvic fin length. Pelvic-fin rays i, 7. Tips of longest pelvic-fin rays reaching slightly beyond anal-fin origin. Principal caudal-fin ray count 10/
<quantity box="[1420,1459,1777,1801]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="18" pageNumber="211" unit="in" value="9.0">9 in</quantity>
all specimens.
</paragraph>
<paragraph blockId="18.[816,1460,1072,1961]" lastBlockId="19.[129,772,944,1513]" lastPageId="19" lastPageNumber="212" pageId="18" pageNumber="211">
Lateral line complete; perforated scales 51-55, 53.5. Horizontal scale rows from dorsal-fin origin to lateral line 12- 13, 12.4. Horizontal scale rows from pelvic-fin origin to lateral line 9-10, 9.9. Horizontal scale rows from anal-fin origin to lateral line 11-13, 12.3. Predorsal scales 30-36, 33.3. Scale rows around caudal peduncle 19-21, 20. Scale row extending for about two thirds of anal-fin base. Basal portions of trunk scales except for those on lateral line bearing dorso-ventrally arranged superficial neuromasts in specimen with unidentified sex (
<figureCitation box="[176,237,1105,1129]" captionStart="Fig" captionStartId="3.[129,168,1194,1218]" captionTargetBox="[129,788,734,1181]" captionTargetId="figure-60@3.[129,772,745,1174]" captionText="Fig. 2. Superficial neuromasts (black arrow) on ventral body scales of Charax metae, CAS 69117, 59 mm SL." figureDoi="http://doi.org/10.5281/zenodo.5131735" httpUri="https://zenodo.org/record/5131735/files/figure.png" pageId="19" pageNumber="212">Fig. 2</figureCitation>
).
</paragraph>
<caption pageId="19" pageNumber="212" startId="19.[129,191,177,201]" targetBox="[142,756,271,856]" targetIsTable="true" targetPageId="19">
<paragraph blockId="19.[129,771,176,233]" pageId="19" pageNumber="212">
<emphasis bold="true" box="[129,214,176,201]" pageId="19" pageNumber="212">Table 9.</emphasis>
Morphometrics of
<taxonomicName authorityName="Eigenmann" authorityYear="1922" box="[411,555,177,201]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="19" pageNumber="212" phylum="Chordata" rank="species" species="metae">
<emphasis box="[411,555,177,201]" italics="true" pageId="19" pageNumber="212">Charax metae</emphasis>
</taxonomicName>
. Specimens are from CAS 69117.
</paragraph>
</caption>
<paragraph pageId="19" pageNumber="212">
<table box="[142,756,271,856]" gridcols="5" gridrows="24" pageId="19" pageNumber="212" rowsContinueFrom="17.[143,756,1350,1935]" rowsContinueIn="21.[823,1448,1380,1965]">
<tr box="[142,756,271,290]" gridrow="0" pageId="19" pageNumber="212">
<th box="[142,443,271,290]" gridcol="0" gridrow="0" pageId="19" pageNumber="212">Characters</th>
<th box="[478,499,271,290]" gridcol="1" gridrow="0" pageId="19" pageNumber="212">n</th>
<th box="[529,622,271,290]" gridcol="2" gridrow="0" pageId="19" pageNumber="212">range</th>
<th box="[653,699,271,290]" gridcol="3" gridrow="0" pageId="19" pageNumber="212">mean</th>
<th box="[728,756,271,290]" gridcol="4" gridrow="0" pageId="19" pageNumber="212">SD</th>
</tr>
<tr box="[142,756,297,316]" gridrow="1" pageId="19" pageNumber="212" rowspan-4="1">
<th box="[142,443,297,316]" gridcol="0" gridrow="1" pageId="19" pageNumber="212">Standard length</th>
<td box="[478,499,297,316]" gridcol="1" gridrow="1" pageId="19" pageNumber="212">16</td>
<td box="[529,622,297,316]" gridcol="2" gridrow="1" pageId="19" pageNumber="212">37.0 - 99.0</td>
<td box="[653,699,297,316]" gridcol="3" gridrow="1" pageId="19" pageNumber="212">62.0</td>
</tr>
<tr box="[142,756,321,341]" gridrow="2" pageId="19" pageNumber="212" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th box="[142,443,321,341]" gridcol="0" gridrow="2" pageId="19" pageNumber="212">Percents of standard length</th>
</tr>
<tr box="[142,756,346,365]" gridrow="3" pageId="19" pageNumber="212">
<th box="[142,443,346,365]" gridcol="0" gridrow="3" pageId="19" pageNumber="212">Depth at dorsal-fin origin</th>
<td box="[478,499,346,365]" gridcol="1" gridrow="3" pageId="19" pageNumber="212">16</td>
<td box="[529,622,346,365]" gridcol="2" gridrow="3" pageId="19" pageNumber="212">35.3 - 38.8</td>
<td box="[653,699,346,365]" gridcol="3" gridrow="3" pageId="19" pageNumber="212">37.0</td>
<td box="[728,756,346,365]" gridcol="4" gridrow="3" pageId="19" pageNumber="212">1.0</td>
</tr>
<tr box="[142,756,370,390]" gridrow="4" pageId="19" pageNumber="212">
<th box="[142,443,370,390]" gridcol="0" gridrow="4" pageId="19" pageNumber="212">Snout to dorsal-fin origin</th>
<td box="[478,499,370,390]" gridcol="1" gridrow="4" pageId="19" pageNumber="212">16</td>
<td box="[529,622,370,390]" gridcol="2" gridrow="4" pageId="19" pageNumber="212">52.8 - 56.8</td>
<td box="[653,699,370,390]" gridcol="3" gridrow="4" pageId="19" pageNumber="212">54.6</td>
<td box="[728,756,370,390]" gridcol="4" gridrow="4" pageId="19" pageNumber="212">1.2</td>
</tr>
<tr box="[142,756,395,414]" gridrow="5" pageId="19" pageNumber="212">
<th box="[142,443,395,414]" gridcol="0" gridrow="5" pageId="19" pageNumber="212">Snout to pectoral-fin origin</th>
<td box="[478,499,395,414]" gridcol="1" gridrow="5" pageId="19" pageNumber="212">16</td>
<td box="[529,622,395,414]" gridcol="2" gridrow="5" pageId="19" pageNumber="212">27.7 - 30.6</td>
<td box="[653,699,395,414]" gridcol="3" gridrow="5" pageId="19" pageNumber="212">29.2</td>
<td box="[728,756,395,414]" gridcol="4" gridrow="5" pageId="19" pageNumber="212">0.9</td>
</tr>
<tr box="[142,756,419,439]" gridrow="6" pageId="19" pageNumber="212">
<th box="[142,443,419,439]" gridcol="0" gridrow="6" pageId="19" pageNumber="212">Snout to pelvic-fin origin</th>
<td box="[478,499,419,439]" gridcol="1" gridrow="6" pageId="19" pageNumber="212">16</td>
<td box="[529,622,419,439]" gridcol="2" gridrow="6" pageId="19" pageNumber="212">36.1 - 41.6</td>
<td box="[653,699,419,439]" gridcol="3" gridrow="6" pageId="19" pageNumber="212">39.1</td>
<td box="[728,756,419,439]" gridcol="4" gridrow="6" pageId="19" pageNumber="212">1.5</td>
</tr>
<tr box="[142,756,444,463]" gridrow="7" pageId="19" pageNumber="212">
<th box="[142,443,444,463]" gridcol="0" gridrow="7" pageId="19" pageNumber="212">Snout to anal-fin origin</th>
<td box="[478,499,444,463]" gridcol="1" gridrow="7" pageId="19" pageNumber="212">16</td>
<td box="[529,622,444,463]" gridcol="2" gridrow="7" pageId="19" pageNumber="212">51.0 - 54.5</td>
<td box="[653,699,444,463]" gridcol="3" gridrow="7" pageId="19" pageNumber="212">52.8</td>
<td box="[728,756,444,463]" gridcol="4" gridrow="7" pageId="19" pageNumber="212">1.2</td>
</tr>
<tr box="[142,756,468,488]" gridrow="8" pageId="19" pageNumber="212">
<th box="[142,443,468,488]" gridcol="0" gridrow="8" pageId="19" pageNumber="212">Caudal peduncle depth</th>
<td box="[478,499,468,488]" gridcol="1" gridrow="8" pageId="19" pageNumber="212">15</td>
<td box="[529,622,468,488]" gridcol="2" gridrow="8" pageId="19" pageNumber="212">8.7 - 10.2</td>
<td box="[653,699,468,488]" gridcol="3" gridrow="8" pageId="19" pageNumber="212">9.6</td>
<td box="[728,756,468,488]" gridcol="4" gridrow="8" pageId="19" pageNumber="212">0.4</td>
</tr>
<tr box="[142,756,493,512]" gridrow="9" pageId="19" pageNumber="212">
<th box="[142,443,493,512]" gridcol="0" gridrow="9" pageId="19" pageNumber="212">Caudal peduncle length</th>
<td box="[478,499,493,512]" gridcol="1" gridrow="9" pageId="19" pageNumber="212">16</td>
<td box="[529,622,493,512]" gridcol="2" gridrow="9" pageId="19" pageNumber="212">6.2 - 8.2</td>
<td box="[653,699,493,512]" gridcol="3" gridrow="9" pageId="19" pageNumber="212">7.5</td>
<td box="[728,756,493,512]" gridcol="4" gridrow="9" pageId="19" pageNumber="212">0.4</td>
</tr>
<tr box="[142,756,517,537]" gridrow="10" pageId="19" pageNumber="212">
<th box="[142,443,517,537]" gridcol="0" gridrow="10" pageId="19" pageNumber="212">Pectoral-fin length</th>
<td box="[478,499,517,537]" gridcol="1" gridrow="10" pageId="19" pageNumber="212">13</td>
<td box="[529,622,517,537]" gridcol="2" gridrow="10" pageId="19" pageNumber="212">18.7 - 21.2</td>
<td box="[653,699,517,537]" gridcol="3" gridrow="10" pageId="19" pageNumber="212">20.0</td>
<td box="[728,756,517,537]" gridcol="4" gridrow="10" pageId="19" pageNumber="212">0.7</td>
</tr>
<tr box="[142,756,542,562]" gridrow="11" pageId="19" pageNumber="212">
<th box="[142,443,542,562]" gridcol="0" gridrow="11" pageId="19" pageNumber="212">Pelvic-fin length</th>
<td box="[478,499,542,562]" gridcol="1" gridrow="11" pageId="19" pageNumber="212">16</td>
<td box="[529,622,542,562]" gridcol="2" gridrow="11" pageId="19" pageNumber="212">17.6 - 21.6</td>
<td box="[653,699,542,562]" gridcol="3" gridrow="11" pageId="19" pageNumber="212">20.1</td>
<td box="[728,756,542,562]" gridcol="4" gridrow="11" pageId="19" pageNumber="212">1.2</td>
</tr>
<tr box="[142,756,566,586]" gridrow="12" pageId="19" pageNumber="212">
<th box="[142,443,566,586]" gridcol="0" gridrow="12" pageId="19" pageNumber="212">Dorsal-fin base length</th>
<td box="[478,499,566,586]" gridcol="1" gridrow="12" pageId="19" pageNumber="212">16</td>
<td box="[529,622,566,586]" gridcol="2" gridrow="12" pageId="19" pageNumber="212">10.3 - 13.0</td>
<td box="[653,699,566,586]" gridcol="3" gridrow="12" pageId="19" pageNumber="212">11.2</td>
<td box="[728,756,566,586]" gridcol="4" gridrow="12" pageId="19" pageNumber="212">0.7</td>
</tr>
<tr box="[142,756,591,610]" gridrow="13" pageId="19" pageNumber="212">
<th box="[142,443,591,610]" gridcol="0" gridrow="13" pageId="19" pageNumber="212">Dorsal-fin height</th>
<td box="[478,499,591,610]" gridcol="1" gridrow="13" pageId="19" pageNumber="212">14</td>
<td box="[529,622,591,610]" gridcol="2" gridrow="13" pageId="19" pageNumber="212">24.6 - 30.2</td>
<td box="[653,699,591,610]" gridcol="3" gridrow="13" pageId="19" pageNumber="212">28.0</td>
<td box="[728,756,591,610]" gridcol="4" gridrow="13" pageId="19" pageNumber="212">1.6</td>
</tr>
<tr box="[142,756,616,635]" gridrow="14" pageId="19" pageNumber="212">
<th box="[142,443,616,635]" gridcol="0" gridrow="14" pageId="19" pageNumber="212">Anal-fin base length</th>
<td box="[478,499,616,635]" gridcol="1" gridrow="14" pageId="19" pageNumber="212">16</td>
<td box="[529,622,616,635]" gridcol="2" gridrow="14" pageId="19" pageNumber="212">41.0 - 46.6</td>
<td box="[653,699,616,635]" gridcol="3" gridrow="14" pageId="19" pageNumber="212">44.6</td>
<td box="[728,756,616,635]" gridcol="4" gridrow="14" pageId="19" pageNumber="212">1.3</td>
</tr>
<tr box="[142,756,640,659]" gridrow="15" pageId="19" pageNumber="212">
<th box="[142,443,640,659]" gridcol="0" gridrow="15" pageId="19" pageNumber="212">Eye to dorsal-fin origin</th>
<td box="[478,499,640,659]" gridcol="1" gridrow="15" pageId="19" pageNumber="212">16</td>
<td box="[529,622,640,659]" gridcol="2" gridrow="15" pageId="19" pageNumber="212">37.7 - 41.7</td>
<td box="[653,699,640,659]" gridcol="3" gridrow="15" pageId="19" pageNumber="212">39.6</td>
<td box="[728,756,640,659]" gridcol="4" gridrow="15" pageId="19" pageNumber="212">1.4</td>
</tr>
<tr box="[142,756,665,684]" gridrow="16" pageId="19" pageNumber="212">
<th box="[142,443,665,684]" gridcol="0" gridrow="16" pageId="19" pageNumber="212">Dorsal-fin origin to caudal-fin base</th>
<td box="[478,499,665,684]" gridcol="1" gridrow="16" pageId="19" pageNumber="212">16</td>
<td box="[529,622,665,684]" gridcol="2" gridrow="16" pageId="19" pageNumber="212">49.5 - 55.0</td>
<td box="[653,699,665,684]" gridcol="3" gridrow="16" pageId="19" pageNumber="212">52.4</td>
<td box="[728,756,665,684]" gridcol="4" gridrow="16" pageId="19" pageNumber="212">1.5</td>
</tr>
<tr box="[142,756,689,709]" gridrow="17" pageId="19" pageNumber="212">
<th box="[142,443,689,709]" gridcol="0" gridrow="17" pageId="19" pageNumber="212">Humeral spot distance</th>
<td box="[478,499,689,709]" gridcol="1" gridrow="17" pageId="19" pageNumber="212">16</td>
<td box="[529,622,689,709]" gridcol="2" gridrow="17" pageId="19" pageNumber="212">41.6 - 47.0</td>
<td box="[653,699,689,709]" gridcol="3" gridrow="17" pageId="19" pageNumber="212">44.0</td>
<td box="[728,756,689,709]" gridcol="4" gridrow="17" pageId="19" pageNumber="212">1.6</td>
</tr>
<tr box="[142,756,713,733]" gridrow="18" pageId="19" pageNumber="212">
<th box="[142,443,713,733]" gridcol="0" gridrow="18" pageId="19" pageNumber="212">Bony head length</th>
<td box="[478,499,713,733]" gridcol="1" gridrow="18" pageId="19" pageNumber="212">16</td>
<td box="[529,622,713,733]" gridcol="2" gridrow="18" pageId="19" pageNumber="212">29.5 - 32.0</td>
<td box="[653,699,713,733]" gridcol="3" gridrow="18" pageId="19" pageNumber="212">31.0</td>
<td box="[728,756,713,733]" gridcol="4" gridrow="18" pageId="19" pageNumber="212">0.6</td>
</tr>
<tr box="[142,756,738,758]" gridrow="19" pageId="19" pageNumber="212" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th box="[142,443,738,758]" gridcol="0" gridrow="19" pageId="19" pageNumber="212">Percents of head length</th>
</tr>
<tr box="[142,756,763,782]" gridrow="20" pageId="19" pageNumber="212">
<th box="[142,443,763,782]" gridcol="0" gridrow="20" pageId="19" pageNumber="212">Horizontal orbital diameter</th>
<td box="[478,499,763,782]" gridcol="1" gridrow="20" pageId="19" pageNumber="212">16</td>
<td box="[529,622,763,782]" gridcol="2" gridrow="20" pageId="19" pageNumber="212">29.6 - 33.3</td>
<td box="[653,699,763,782]" gridcol="3" gridrow="20" pageId="19" pageNumber="212">31.5</td>
<td box="[728,756,763,782]" gridcol="4" gridrow="20" pageId="19" pageNumber="212">1.4</td>
</tr>
<tr box="[142,756,787,807]" gridrow="21" pageId="19" pageNumber="212">
<th box="[142,443,787,807]" gridcol="0" gridrow="21" pageId="19" pageNumber="212">Snout length</th>
<td box="[478,499,787,807]" gridcol="1" gridrow="21" pageId="19" pageNumber="212">16</td>
<td box="[529,622,787,807]" gridcol="2" gridrow="21" pageId="19" pageNumber="212">23.6 - 27.8</td>
<td box="[653,699,787,807]" gridcol="3" gridrow="21" pageId="19" pageNumber="212">25.5</td>
<td box="[728,756,787,807]" gridcol="4" gridrow="21" pageId="19" pageNumber="212">1.1</td>
</tr>
<tr box="[142,756,812,831]" gridrow="22" pageId="19" pageNumber="212">
<th box="[142,443,812,831]" gridcol="0" gridrow="22" pageId="19" pageNumber="212">Least interorbital width</th>
<td box="[478,499,812,831]" gridcol="1" gridrow="22" pageId="19" pageNumber="212">16</td>
<td box="[529,622,812,831]" gridcol="2" gridrow="22" pageId="19" pageNumber="212">23.0 - 26.7</td>
<td box="[653,699,812,831]" gridcol="3" gridrow="22" pageId="19" pageNumber="212">24.2</td>
<td box="[728,756,812,831]" gridcol="4" gridrow="22" pageId="19" pageNumber="212">1.0</td>
</tr>
<tr box="[142,756,836,856]" gridrow="23" pageId="19" pageNumber="212">
<th box="[142,443,836,856]" gridcol="0" gridrow="23" pageId="19" pageNumber="212">Upper jaw length</th>
<td box="[478,499,836,856]" gridcol="1" gridrow="23" pageId="19" pageNumber="212">16</td>
<td box="[529,622,836,856]" gridcol="2" gridrow="23" pageId="19" pageNumber="212">54.4 - 69.3</td>
<td box="[653,699,836,856]" gridcol="3" gridrow="23" pageId="19" pageNumber="212">60.5</td>
<td box="[728,756,836,856]" gridcol="4" gridrow="23" pageId="19" pageNumber="212">4.6</td>
</tr>
</table>
</paragraph>
<paragraph blockId="19.[129,772,944,1513]" pageId="19" pageNumber="212">Premaxillary with one anterior canine-like tooth followed by set of smaller conical teeth and another canine-like tooth followed by one or two small conical teeth. Total number of premaxillary teeth 11-14, 12.5. Maxillary teeth conical, 44-56, 48.7; larger specimens generally with higher counts. Dentary with one canine-like tooth followed by 6-8, 6.5 conical teeth, another canine-like tooth and a posterior row of 17-25, 21.2 conical teeth. Left ectopterygoid bone with 10-27, and right with 10-23 conical teeth. Total number of ectopterygoid teeth 20-50, 29.7.Vertebrae 33 and 34 (2). Gill-rakers on lower limb of first gill-arch 7-9, 8.3. Branchiostegal rays 4; 3 rays originating from anterior ceratohyal and 1 from posterior ceratohyal.</paragraph>
<paragraph blockId="19.[129,772,1552,1961]" lastBlockId="19.[809,1452,181,365]" pageId="19" pageNumber="212">
<emphasis bold="true" box="[129,314,1552,1577]" pageId="19" pageNumber="212">Color in alcohol.</emphasis>
Old preserved specimens show overall pale to light brown body coloration with scattered dark chromatophores especially concentrated on basal portion of scales on anterior midbody region. Superficial neuromasts vertically arranged along skin of basal portion of each scale in most specimens whitish and contrasting with background dark coloration of the scale basal portion (
<figureCitation box="[573,638,1745,1769]" captionStart="Fig" captionStartId="3.[129,168,1194,1218]" captionTargetBox="[129,788,734,1181]" captionTargetId="figure-60@3.[129,772,745,1174]" captionText="Fig. 2. Superficial neuromasts (black arrow) on ventral body scales of Charax metae, CAS 69117, 59 mm SL." figureDoi="http://doi.org/10.5281/zenodo.5131735" httpUri="https://zenodo.org/record/5131735/files/figure.png" pageId="19" pageNumber="212">Fig. 2</figureCitation>
). Irregularly shaped humeral dark blotch region, much closer to dorsal-fin origin than to posterior border of opercle, encompassing about four scales vertically and horizontally. Dark blotch on caudal peduncle approximately triangular shaped, higher posteriorly and extending over basal portions of median caudal-fin rays. Scattered dark chromatophores on the tips of the snout and the lower jaw. Tip of dorsal fin dark. Anal fin with faint basal dark stripe, separated by light stripe from more conspicuous dark wider distal stripe extending to end of fin. Tip of pelvic fins dark. Pectorals light with few scattered dark chromatophores. Marginal portion of caudal fin dark with scattered dark chromatophores.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="212" type="distribution">
<paragraph blockId="19.[809,1451,405,461]" pageId="19" pageNumber="212">
<emphasis bold="true" box="[809,957,405,429]" pageId="19" pageNumber="212">Distribution.</emphasis>
<taxonomicName authorityName="Eigenmann" authorityYear="1922" box="[967,1122,405,429]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="19" pageNumber="212" phylum="Chordata" rank="species" species="metae">
<emphasis box="[967,1122,405,429]" italics="true" pageId="19" pageNumber="212">Charax metae</emphasis>
</taxonomicName>
is known from the río
<collectingRegion box="[1389,1447,406,429]" country="Colombia" name="Meta" pageId="19" pageNumber="212">Meta</collectingRegion>
,
<collectingCountry box="[809,911,437,461]" name="Colombia" pageId="19" pageNumber="212">Colombia</collectingCountry>
(
<figureCitation box="[923,984,437,461]" captionStart="Fig" captionStartId="5.[809,847,1648,1672]" captionTargetBox="[809,1452,819,1618]" captionTargetId="figure-305@5.[809,1452,815,1618]" captionTargetPageId="5" captionText="Fig.7. Map of SouthAmerica showing the collecting sites of the Charax species, and the type locality of Charax delimai (purple circle). Charax apurensis - yellow square; C. caudimaculatus - white circle; C. condei - white square; C. gibbosus - purple triangle; C. hemigrammus - red star; C. leticiae - black circle;C. macrolepis-black triangle;C.metae-yellow triangle;C.michaeli - yellow circle;C.niger -blue triangle; C.notulatus - blue circle; C. pauciradiatus - red triangle; C. rupununi - red circle; C. stenopterus - green triangle; C. tectifer - green circle. Some symbols may represent more than one locality." figureDoi="http://doi.org/10.5281/zenodo.4639009" httpUri="https://zenodo.org/record/4639009/files/figure.png" pageId="19" pageNumber="212">Fig. 7</figureCitation>
).
</paragraph>
<paragraph blockId="19.[809,1452,501,581]" pageId="19" pageNumber="212">
<emphasis bold="true" box="[809,1032,501,523]" pageId="19" pageNumber="212">Specimens examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3326898415" accessionNumber="CAS 69117" collectionCode="CAS" country="Colombia" latitude="10.566667" location="Puerto Barrigona" longLatPrecision="1297" longitude="-73.75" pageId="19" pageNumber="212" specimenCount="16" stateProvince="Meta" typeStatus="paratype">
<accessionNumber box="[1040,1157,501,523]" httpUri="http://www.ncbi.nlm.nih.gov/protein/CAS69117" pageId="19" pageNumber="212">
<collectionCode box="[1040,1087,501,522]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="19" pageNumber="212" type="Museum">CAS</collectionCode>
69117
</accessionNumber>
,
<specimenCount box="[1169,1296,501,523]" pageId="19" pageNumber="212" type="generic" typeStatus="paratypes">16 paratypes</specimenCount>
,
<quantity box="[1308,1410,501,523]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" metricValueMax="8.7" metricValueMin="3.7" pageId="19" pageNumber="212" unit="mm" value="62.0" valueMax="87.0" valueMin="37.0">37-87 mm</quantity>
SL,
<collectingCountry box="[809,905,530,552]" name="Colombia" pageId="19" pageNumber="212">Colômbia</collectingCountry>
,
<collectingRegion box="[913,964,531,552]" country="Colombia" name="Meta" pageId="19" pageNumber="212">Meta</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03C4D93834683F51FEECFAB9FABF6466:8EB23EF534693F51FC37FC7AFBA5640B" box="[972,1134,530,552]" country="Colombia" latitude="10.566667" longLatPrecision="1297" longitude="-73.75" name="Puerto Barrigona" pageId="19" pageNumber="212" stateProvince="Meta">Puerto Barrigona</location>
(=Barrigón) at head of navigation east of
<collectingRegion box="[881,954,559,580]" country="Colombia" name="Bogota" pageId="19" pageNumber="212">Bogota</collectingRegion>
, approximately 10°34’N 75°11’W, 73°45’W
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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