479 lines
57 KiB
XML
479 lines
57 KiB
XML
<document id="95F6AE0D228D3A9866F98C4D122C3B1E" ID-CLB-Dataset="244288" ID-DOI="10.11646/zootaxa.5263.3.8" ID-GBIF-Dataset="285bc709-1958-488b-964e-64b720b23149" ID-ISSN="1175-5326" ID-Zenodo-Dep="7804410" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1680761118113" checkinUser="plazi" docAuthor="Kantor, Yuri, Zvonareva, Sofia & Krylova, Elena" docDate="2023" docId="039ED47BB6481740FF09AD834059F89B" docLanguage="en" docName="zootaxa.5263.3.8.pdf" docOrigin="Zootaxa 5263 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Astyris axicostata Kantor & Zvonareva & Krylova 2023, n. sp." docType="treatment" docUuid="8199E15C-A73A-4828-AEED-9339C2B42E00" docUuidSource="ZooBank" docVersion="9" lastPageNumber="446" masterDocId="FFA7AC03B64A1745FF9EAA0F4114FFF4" masterDocTitle="New species of the genus Astyris (Gastropoda: Neogastropoda: Buccinoidea Columbellidae) from the vicinity of hydrothermal vents of the Piip Volcano southwestern Bering Sea" masterLastPageNumber="450" masterPageNumber="441" pageNumber="443" updateTime="1699515179009" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E9C8CC54C3667929B28B9AC0BB97780B">New species of the genus Astyris (Gastropoda: Neogastropoda: Buccinoidea Columbellidae) from the vicinity of hydrothermal vents of the Piip Volcano southwestern Bering Sea</mods:title>
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<mods:namePart id="CC0C0BBC8A1AEA6BD8E9CB68C287679D">Kantor, Yuri</mods:namePart>
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<mods:affiliation id="C5BF701192D168819C33BEE6681EF637">A. N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninski prospect 33, Moscow 119071, Russian Federation. & kantor. yuri 1956 @ gmail. com; https: // orcid. org / 0000 - 0002 - 3209 - 4940</mods:affiliation>
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<mods:nameIdentifier id="495477252230080DCF0292359F03EC6F" type="email">kantor.yuri1956@gmail.com</mods:nameIdentifier>
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<mods:namePart id="50DB40C608ADAED8E44882929DE0C2FA">Zvonareva, Sofia</mods:namePart>
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<mods:affiliation id="039B3E9AF9DAA6C76CF6FF89B972295C">A. N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninski prospect 33, Moscow 119071, Russian Federation. & sonyazv @ mail. ru; https: // orcid. org / 0000 - 0002 - 0372 - 9919</mods:affiliation>
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<mods:nameIdentifier id="4C83DDE7F6A1FD0944D71B5DF963AAAC" type="email">sonyazv@mail.ru</mods:nameIdentifier>
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<mods:namePart id="8A7B2E13F5BFB161349BC521F89B7546">Krylova, Elena</mods:namePart>
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<mods:affiliation id="2744D4730385D1248D54274AE642FEC7">Shirshov Institute of Oceanology, Russian Academy of Sciences, 36, Nahimovskiy prospekt, Moscow 117997, Russian Federation. elenamkr @ mail. ru; https: // orcid. org / 0000 - 0003 - 1341 - 1709</mods:affiliation>
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<mods:nameIdentifier id="8DD659473403936ECC0568C557317E31" type="email">elenamkr@mail.ru</mods:nameIdentifier>
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<mods:title id="0416AB2D38378C919B47E14C08FEC76D">Zootaxa</mods:title>
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<mods:date id="1C5890475832E406C69AF64364A69473">2023</mods:date>
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<treatment id="039ED47BB6481740FF09AD834059F89B" ID-DOI="http://doi.org/10.5281/zenodo.7805414" ID-GBIF-Taxon="209495683" ID-Zenodo-Dep="7805414" LSID="urn:lsid:zoobank.org:act:8199E15C-A73A-4828-AEED-9339C2B42E00" httpUri="http://treatment.plazi.org/id/039ED47BB6481740FF09AD834059F89B" lastPageId="5" lastPageNumber="446" pageId="2" pageNumber="443">
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<paragraph id="8B88656DB6481747FF09AD8340A2F852" blockId="2.[151,1171,1752,2030]" box="[151,438,1932,1958]" pageId="2" pageNumber="443">
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<emphasis id="B943B97FB6481747FF09AD8340A2F852" bold="true" box="[151,438,1932,1958]" pageId="2" pageNumber="443">
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<taxonomicName id="4C371EEEB6481747FF09AD834078F852" ID-CoL="c45744ae-20a7-4980-9734-607d8bf88350" authority="Kantor & Zvonareva & Krylova, 2023" authorityName="Kantor & Zvonareva & Krylova" authorityYear="2023" box="[151,364,1932,1958]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="443" phylum="Mollusca" rank="species" species="axicostata" status="sp. nov.">
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<emphasis id="B943B97FB6481747FF09AD834078F852" bold="true" box="[151,364,1932,1958]" italics="true" pageId="2" pageNumber="443">Astyris axicostata</emphasis>
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<taxonomicNameLabel id="A2700404B6481747FEEDAD8340A2F852" box="[371,438,1932,1958]" pageId="2" pageNumber="443" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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Zoobank registration:
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<subSubSection id="C32D36E6B6481747FF09ADD943B3F81A" box="[151,679,2006,2030]" pageId="2" pageNumber="443" type="reference_group">
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<paragraph id="8B88656DB6481747FF09ADD943B3F81A" blockId="2.[151,1171,1752,2030]" box="[151,679,2006,2030]" pageId="2" pageNumber="443">
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<taxonomicName id="4C371EEEB6481747FF09ADD9402EF819" ID-CoL="7NPZ9" authority="Kantor & Zvonareva & Krylova, 2023" authorityName="Kantor & Zvonareva & Krylova" authorityYear="2023" box="[151,314,2006,2029]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="443" phylum="Mollusca" rank="species" species="undetermined" status="sp. nov.">
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<emphasis id="B943B97FB6481747FF09ADD941F5F819" box="[151,225,2006,2029]" italics="true" pageId="2" pageNumber="443">Astyris</emphasis>
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<taxonomicNameLabel id="A2700404B6481747FF76ADD9402EF819" box="[232,314,2006,2029]" pageId="2" pageNumber="443" rank="species">sp. nov</taxonomicNameLabel>
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</taxonomicName>
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.:
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<treatmentCitation id="0A96437CB6481747FED6ADD94356F81A" author="Nekhaev, I. O. & Chaban, E. M. & Kantor, Yu. I. & Kuchsh, D. A. & Matveeva, K. & Rybakova, E." box="[328,578,2006,2030]" page="9" pageId="2" pageNumber="443" year="2022">
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<bibRefCitation id="EFA6189CB6481747FED6ADD94356F81A" author="Nekhaev, I. O. & Chaban, E. M. & Kantor, Yu. I. & Kuchsh, D. A. & Matveeva, K. & Rybakova, E." box="[328,578,2006,2030]" pageId="2" pageNumber="443" pagination="105164" refId="ref5921" refString="Nekhaev, I. O., Chaban, E. M., Kantor, Yu. I., Kuchsh, D. A., Matveeva, K. & Rybakova, E. (2022) Shell-bearing Gastropoda from the methane seeps and hydrothermal vents of the Bering Sea: A preliminary description. Deep Sea Research Part II: Topical Studies in Oceanography, 204, 105164. https: // doi. org / 10.1016 / j. dsr 2.2022.105164" type="journal article" year="2022">
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Nekhaev
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<emphasis id="B943B97FB6481747FE33ADD840F7F81A" box="[429,483,2006,2030]" italics="true" pageId="2" pageNumber="443">et al.</emphasis>
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, 2022: 9
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</bibRefCitation>
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</treatmentCitation>
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, Fig. 4D.
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</paragraph>
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<caption id="DF4835E5B6491746FF09AD4C42ADF857" ID-DOI="http://doi.org/10.5281/zenodo.7804412" ID-Zenodo-Dep="7804412" httpUri="https://zenodo.org/record/7804412/files/figure.png" pageId="3" pageNumber="444" startId="3.[151,250,1859,1883]" targetBox="[167,1414,190,1826]" targetPageId="3">
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<paragraph id="8B88656DB6491746FF09AD4C42ADF857" blockId="3.[151,1437,1859,1955]" pageId="3" pageNumber="444">
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<emphasis id="B943B97FB6491746FF09AD4C4002F8AF" bold="true" box="[151,278,1859,1883]" pageId="3" pageNumber="444">FIGURE 1.</emphasis>
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Phylogenetic reconstruction using the COI gene (658 bp) showing
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<taxonomicName id="4C371EEEB6491746FC40AD4B4581F8AF" authority="Kantor & Zvonareva & Krylova, 2023" authorityName="Kantor & Zvonareva & Krylova" authorityYear="2023" box="[990,1173,1860,1883]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="3" pageNumber="444" phylum="Mollusca" rank="species" species="axicostata" status="sp. nov.">
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<emphasis id="B943B97FB6491746FC40AD4B4581F8AF" box="[990,1173,1860,1883]" italics="true" pageId="3" pageNumber="444">Astyris axicostata</emphasis>
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</taxonomicName>
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<emphasis id="B943B97FB6491746FB01AD4C45CFF8AF" bold="true" box="[1183,1243,1859,1883]" pageId="3" pageNumber="444">
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<taxonomicNameLabel id="A2700404B6491746FB01AD4C45CFF8AF" box="[1183,1243,1859,1883]" pageId="3" pageNumber="444" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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forming the clade with
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<taxonomicName id="4C371EEEB6491746FF54AD674087F88B" box="[202,403,1895,1919]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="3" pageNumber="444" phylum="Mollusca" rank="species" species="thermophila">
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<emphasis id="B943B97FB6491746FF54AD674087F88B" box="[202,403,1895,1919]" italics="true" pageId="3" pageNumber="444">Astyris thermophila</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C371EEEB6491746FE5AAD67433BF88B" baseAuthorityName="Gould" baseAuthorityYear="1840" box="[452,559,1896,1919]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="3" pageNumber="444" phylum="Mollusca" rank="species" species="rosacea">
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<emphasis id="B943B97FB6491746FE5AAD67433BF88B" box="[452,559,1896,1919]" italics="true" pageId="3" pageNumber="444">A. rosacea</emphasis>
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</taxonomicName>
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nested within
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<taxonomicName id="4C371EEEB6491746FD5DAD684248F88B" authorityName="Swainson" authorityYear="1840" box="[707,860,1895,1919]" class="Gastropoda" family="Columbellidae" kingdom="Animalia" order="Neogastropoda" pageId="3" pageNumber="444" phylum="Mollusca" rank="family">Columbellidae</taxonomicName>
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. Node values indicate Bayesian posterior probability and Maximal Likelihood bootstrap values, those lower than 0.8 and 80 are omitted.
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</paragraph>
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</caption>
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<subSubSection id="C32D36E6B64E1740FF09AA9742B7FECB" lastPageId="5" lastPageNumber="446" pageId="4" pageNumber="445" type="materials_examined">
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<paragraph id="8B88656DB64E1741FF09AA9742F4FF20" blockId="4.[151,1437,151,284]" pageId="4" pageNumber="445">
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<emphasis id="B943B97FB64E1741FF09AA974044FF5B" box="[151,336,151,176]" inLineHeading="true" pageId="4" pageNumber="445" reason="1">
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<emphasis id="B943B97FB64E1741FF09AA97405EFF5B" bold="true" box="[151,330,151,176]" pageId="4" pageNumber="445">Type specimens</emphasis>
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.
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</emphasis>
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<materialsCitation id="3B5F6F30B64E1741FEC9AA9842F4FF20" ID-GBIF-Occurrence="4075342302" collectionCode="IORAS" pageId="4" pageNumber="445" specimenCode="MIMB 45730, MIMB 45731, INV0000847" specimenCount="1" typeStatus="holotype">
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<typeStatus id="548CDBCFB64E1741FEC9AA9840AEFF5B" box="[343,442,151,175]" pageId="4" pageNumber="445" type="holotype">Holotype</typeStatus>
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(
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<figureCitation id="130C79E8B64E1741FE57AA984357FF44" box="[457,579,151,176]" captionStart="FIGURE 2" captionStartId="4.[151,250,1843,1867]" captionTargetBox="[151,1435,308,1820]" captionTargetId="figure-87@4.[151,1435,308,1820]" captionTargetPageId="4" captionText="FIGURE 2. Astyris axicostata n. sp. (A–I) A–D. Holotype, SL 7.3 mm (A–C—dried shell, D—wet shell with body seen through transparency). E. Paratype 1, SL 7.4 mm. F–G. Paratype 2, SL 7.0 mm. H–I. Paratype 3, juvenile with intact protoconch, SL 3.0 mm. J. Astyris cf. rosacea (Gould, 1840), SL 12.1 mm. A–G—at the same scale, H–I—enlarged, J—reduced." figureDoi="http://doi.org/10.5281/zenodo.7804414" httpUri="https://zenodo.org/record/7804414/files/figure.png" pageId="4" pageNumber="445">Fig. 2 A–D</figureCitation>
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) [SL
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<quantity id="4CCFC888B64E1741FDE1AA9743C2FF5B" box="[639,726,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="4" pageNumber="445" unit="mm" value="7.3">7.3 mm</quantity>
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],
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<specimenCode id="DB91CD16B64E1741FD76AA97426AFF5B" box="[744,894,151,175]" collectionCode="MIMB" country="Russia" httpUri="http://grbio.org/cool/063z-3mpp" name="Museum of the Institute of Marine Biology" pageId="4" pageNumber="445">MIMB 45730</specimenCode>
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. Three paratypes [SL 7.4, 7.0, and 3.0 mm] (
|
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<figureCitation id="130C79E8B64E1741FAC3AA9841D4FF20" captionStart="FIGURE 2" captionStartId="4.[151,250,1843,1867]" captionTargetBox="[151,1435,308,1820]" captionTargetId="figure-87@4.[151,1435,308,1820]" captionTargetPageId="4" captionText="FIGURE 2. Astyris axicostata n. sp. (A–I) A–D. Holotype, SL 7.3 mm (A–C—dried shell, D—wet shell with body seen through transparency). E. Paratype 1, SL 7.4 mm. F–G. Paratype 2, SL 7.0 mm. H–I. Paratype 3, juvenile with intact protoconch, SL 3.0 mm. J. Astyris cf. rosacea (Gould, 1840), SL 12.1 mm. A–G—at the same scale, H–I—enlarged, J—reduced." figureDoi="http://doi.org/10.5281/zenodo.7804414" httpUri="https://zenodo.org/record/7804414/files/figure.png" pageId="4" pageNumber="445">Fig. 2 E–I</figureCitation>
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)
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<specimenCode id="DB91CD16B64E1741FF53AAB34070FF27" box="[205,356,187,211]" collectionCode="MIMB" country="Russia" httpUri="http://grbio.org/cool/063z-3mpp" name="Museum of the Institute of Marine Biology" pageId="4" pageNumber="445">MIMB 45731</specimenCode>
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;
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<specimenCount id="9D31AEE4B64E1741FEF1AAB34317FF20" box="[367,515,188,212]" pageId="4" pageNumber="445" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
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[SL 7.3, and
|
||
<quantity id="4CCFC888B64E1741FD0AAAB443FCFF27" box="[660,744,187,211]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="4" pageNumber="445" unit="mm" value="5.3">5.3 mm</quantity>
|
||
]
|
||
<collectionCode id="ED26FDA8B64E1741FD6BAAB34251FF20" box="[757,837,188,212]" pageId="4" pageNumber="445">IORAS</collectionCode>
|
||
<specimenCode id="DB91CD16B64E1741FCD5AAB342C8FF20" box="[843,988,188,212]" collectionCode="IORAS" pageId="4" pageNumber="445">INV0000847</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B88656DB64E1741FF59AAEF440DFEEF" blockId="4.[151,1437,151,284]" pageId="4" pageNumber="445">
|
||
<materialsCitation id="3B5F6F30B64E1741FF59AAEF4401FEEF" ID-GBIF-Occurrence="4075342304" collectionCode="R, V" collectorName="Akademik M. A. Lavrentyev & Hexacorallia" elevation="373" latitude="55.416" longLatPrecision="9" longitude="167.2744" pageId="4" pageNumber="445" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B943B97FB64E1741FF59AAEF404AFF03" bold="true" box="[199,350,223,248]" inLineHeading="true" pageId="4" pageNumber="445" reason="1">
|
||
<typeStatus id="548CDBCFB64E1741FF59AAEF41EBFF0C" box="[199,255,224,248]" pageId="4" pageNumber="445">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Northern summit of submarine Piip Volcano,
|
||
<geoCoordinate id="EE0303AAB64E1741FCD2AAD142DDFF03" box="[844,969,222,247]" degrees="55" direction="north" minutes="24.96" orientation="latitude" pageId="4" pageNumber="445" precision="9" value="55.416">55°24.96′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0303AAB64E1741FC4AAAD1457DFF03" box="[980,1129,222,247]" degrees="167" direction="east" minutes="16.464" orientation="longitude" pageId="4" pageNumber="445" precision="1" value="167.2744">167°16.464′E</geoCoordinate>
|
||
,
|
||
<quantity id="4CCFC888B64E1741FBEAAAEF45AEFF03" box="[1140,1210,224,248]" metricMagnitude="2" metricUnit="m" metricValue="3.7299999999999995" pageId="4" pageNumber="445" unit="m" value="373.0">
|
||
<elevation id="001A825EB64E1741FBEAAAEF45AEFF03" box="[1140,1210,224,248]" metricMagnitude="2" metricUnit="m" metricValue="3.7299999999999995" pageId="4" pageNumber="445" unit="m" value="373.0">373 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectionCode id="ED26FDA8B64E1741FB5BAAEF45C3FF0C" box="[1221,1239,224,248]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="445">R</collectionCode>
|
||
/
|
||
<collectionCode id="ED26FDA8B64E1741FB41AAEF45E6FF0C" box="[1247,1266,224,248]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="445" type="Museum">V</collectionCode>
|
||
<emphasis id="B943B97FB64E1741FB66AAD04018FEEF" italics="true" pageId="4" pageNumber="445">
|
||
<collectorName id="26C200BBB64E1741FB66AAD04018FEEF" pageId="4" pageNumber="445">Akademik M.A. Lavrentyev</collectorName>
|
||
</emphasis>
|
||
, cruise 82, sta LV82-1/7, 2018, rock with zoantharians (
|
||
<taxonomicName id="4C371EEEB64E1741FCF8AB0C451BFEE8" box="[870,1039,259,284]" class="Anthozoa" family="Epizoanthidae" genus="Epizoanthus" kingdom="Animalia" order="Zoantharia" pageId="4" pageNumber="445" phylum="Cnidaria" rank="species" species="undetermined">
|
||
<emphasis id="B943B97FB64E1741FCF8AB0C42FEFEEF" box="[870,1002,259,283]" italics="true" pageId="4" pageNumber="445">Epizoanthus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
) and other
|
||
<collectorName id="26C200BBB64E1741FB17AB0C4401FEEF" box="[1161,1301,259,283]" pageId="4" pageNumber="445">Hexacorallia</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF4835E5B64E1741FF09AD3C45FAF867" ID-DOI="http://doi.org/10.5281/zenodo.7804414" ID-Zenodo-Dep="7804414" httpUri="https://zenodo.org/record/7804414/files/figure.png" pageId="4" pageNumber="445" startId="4.[151,250,1843,1867]" targetBox="[151,1435,308,1820]" targetPageId="4">
|
||
<paragraph id="8B88656DB64E1741FF09AD3C45FAF867" blockId="4.[151,1437,1843,1939]" pageId="4" pageNumber="445">
|
||
<emphasis id="B943B97FB64E1741FF09AD3C4018F8BF" bold="true" box="[151,268,1843,1867]" pageId="4" pageNumber="445">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="4C371EEEB64E1741FE89AD3B40DEF8BF" authority="Kantor & Zvonareva & Krylova, 2023" authorityName="Kantor & Zvonareva & Krylova" authorityYear="2023" box="[279,458,1844,1867]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="445" phylum="Mollusca" rank="species" species="axicostata" status="sp. nov.">
|
||
<emphasis id="B943B97FB64E1741FE89AD3B40DEF8BF" box="[279,458,1844,1867]" italics="true" pageId="4" pageNumber="445">Astyris axicostata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B97FB64E1741FE51AD3C4313F8BF" bold="true" box="[463,519,1843,1867]" pageId="4" pageNumber="445">
|
||
<taxonomicNameLabel id="A2700404B64E1741FE51AD3C4313F8BF" box="[463,519,1843,1867]" pageId="4" pageNumber="445" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A–I)
|
||
<emphasis id="B943B97FB64E1741FDD7AD3C4395F8BF" bold="true" box="[585,641,1843,1867]" pageId="4" pageNumber="445">A–D.</emphasis>
|
||
Holotype, SL 7.3 mm (A–C—dried shell, D—wet shell with body seen through transparency).
|
||
<emphasis id="B943B97FB64E1741FEB3AD584050F89B" bold="true" box="[301,324,1879,1903]" pageId="4" pageNumber="445">E.</emphasis>
|
||
Paratype 1, SL 7.4 mm.
|
||
<emphasis id="B943B97FB64E1741FDA5AD584366F89B" bold="true" box="[571,626,1879,1903]" pageId="4" pageNumber="445">F–G.</emphasis>
|
||
Paratype 2, SL 7.0 mm.
|
||
<emphasis id="B943B97FB64E1741FCF6AD58428DF89B" bold="true" box="[872,921,1879,1903]" pageId="4" pageNumber="445">H–I.</emphasis>
|
||
Paratype 3, juvenile with intact protoconch, SL 3.0 mm.
|
||
<emphasis id="B943B97FB64E1741FF55AD7441CAF867" bold="true" box="[203,222,1915,1939]" pageId="4" pageNumber="445">J.</emphasis>
|
||
<taxonomicName id="4C371EEEB64E1741FF7BAD73432DF867" authority="(Gould, 1840)" baseAuthorityName="Gould" baseAuthorityYear="1840" box="[229,569,1915,1939]" class="Gastropoda" family="Columbellidae" genus="Astyris" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="445" phylum="Mollusca" rank="species" species="rosacea">
|
||
<emphasis id="B943B97FB64E1741FF7BAD73403FF867" box="[229,299,1916,1939]" italics="true" pageId="4" pageNumber="445">Astyris</emphasis>
|
||
cf.
|
||
<emphasis id="B943B97FB64E1741FECCAD7340B5F867" box="[338,417,1916,1939]" italics="true" pageId="4" pageNumber="445">rosacea</emphasis>
|
||
(Gould, 1840)
|
||
</taxonomicName>
|
||
, SL 12.1 mm. A–G—at the same scale, H–I—enlarged, J—reduced.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B88656DB64F1740FF59AA9842B7FECB" blockId="5.[151,1437,151,1903]" pageId="5" pageNumber="446">
|
||
<materialsCitation id="3B5F6F30B64F1740FF59AA98428BFF03" ID-GBIF-Occurrence="4075342303" collectionCode="R, V" collectorName="Akademik M. A. Lavrentyev" elevation="402" latitude="55.4166" longLatPrecision="1" longitude="167.2748" pageId="5" pageNumber="446" specimenCount="28">
|
||
<emphasis id="B943B97FB64F1740FF59AA98433FFF5B" bold="true" box="[199,555,151,175]" inLineHeading="true" pageId="5" pageNumber="446" reason="1">Additional material examined.</emphasis>
|
||
<specimenCount id="9D31AEE4B64F1740FDA8AA9743DEFF5B" box="[566,714,151,176]" pageId="5" pageNumber="446" type="generic">28 specimens</specimenCount>
|
||
from the
|
||
<typeStatus id="548CDBCFB64F1740FCA1AA974279FF44" box="[831,877,152,176]" pageId="5" pageNumber="446">type</typeStatus>
|
||
locality, one sequenced and examined for radular morphology (Buc298, shell partially destroyed). Northern summit of submarine Piip Volcano,
|
||
<geoCoordinate id="EE0303AAB64F1740FBEEAAB545EFFF27" box="[1136,1275,186,211]" degrees="55" direction="north" minutes="24.996" orientation="latitude" pageId="5" pageNumber="446" precision="1" value="55.4166">55°24.996′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0303AAB64F1740FA9DAAB5448CFF27" box="[1283,1432,186,211]" degrees="167" direction="east" minutes="16.488" orientation="longitude" pageId="5" pageNumber="446" precision="1" value="167.2748">167°16.488′E</geoCoordinate>
|
||
,
|
||
<quantity id="4CCFC888B64F1740FF09AAEF41CAFF03" box="[151,222,224,248]" metricMagnitude="2" metricUnit="m" metricValue="4.02" pageId="5" pageNumber="446" unit="m" value="402.0">
|
||
<elevation id="001A825EB64F1740FF09AAEF41CAFF03" box="[151,222,224,248]" metricMagnitude="2" metricUnit="m" metricValue="4.02" pageId="5" pageNumber="446" unit="m" value="402.0">402 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectionCode id="ED26FDA8B64F1740FF74AAEF41E8FF0C" box="[234,252,224,248]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="446">R</collectionCode>
|
||
/
|
||
<collectionCode id="ED26FDA8B64F1740FE9AAAEF4003FF0C" box="[260,279,224,248]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="446" type="Museum">V</collectionCode>
|
||
<emphasis id="B943B97FB64F1740FE80AAD04354FF03" box="[286,576,223,247]" italics="true" pageId="5" pageNumber="446">
|
||
<collectorName id="26C200BBB64F1740FE80AAD04354FF03" box="[286,576,223,247]" pageId="5" pageNumber="446">Akademik M.A. Lavrentyev</collectorName>
|
||
</emphasis>
|
||
, cruise 75, sta LV75-10/1, 2016
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B5F6F30B64F1740FC35AAEF401AFECB" ID-GBIF-Occurrence="4075342301" collectionCode="R, V" collectorName="Akademik M. A. Lavrentyev" elevation="468" latitude="55.3823" longLatPrecision="1" longitude="167.2616" pageId="5" pageNumber="446" specimenCount="1">
|
||
<specimenCount id="9D31AEE4B64F1740FC35AAEF4533FF03" box="[939,1063,223,248]" pageId="5" pageNumber="446" type="generic">1 specimen</specimenCount>
|
||
, rocks on bacterial mats. Southern summit of submarine Piip Volcano
|
||
<geoCoordinate id="EE0303AAB64F1740FD87AB0D43B7FEEF" box="[537,675,258,283]" degrees="55" direction="north" minutes="22.938" orientation="latitude" pageId="5" pageNumber="446" precision="1" value="55.3823">55°22.938′N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0303AAB64F1740FD2EAB0D4250FEEF" box="[688,836,258,283]" degrees="167" direction="east" minutes="15.696" orientation="longitude" pageId="5" pageNumber="446" precision="1" value="167.2616">167°15.696′E</geoCoordinate>
|
||
,
|
||
<quantity id="4CCFC888B64F1740FCCFAB0C428CFEEF" box="[849,920,259,283]" metricMagnitude="2" metricUnit="m" metricValue="4.68" pageId="5" pageNumber="446" unit="m" value="468.0">
|
||
<elevation id="001A825EB64F1740FCCFAB0C428CFEEF" box="[849,920,259,283]" metricMagnitude="2" metricUnit="m" metricValue="4.68" pageId="5" pageNumber="446" unit="m" value="468.0">468 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectionCode id="ED26FDA8B64F1740FC3BAB0B42A3FEE8" box="[933,951,260,284]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="446">R</collectionCode>
|
||
/
|
||
<collectionCode id="ED26FDA8B64F1740FC21AB0B42C6FEE8" box="[959,978,260,284]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="446" type="Museum">V</collectionCode>
|
||
<emphasis id="B943B97FB64F1740FC44AB0C45E9FEEF" box="[986,1277,259,283]" italics="true" pageId="5" pageNumber="446">
|
||
<collectorName id="26C200BBB64F1740FC44AB0C45E9FEEF" box="[986,1277,259,283]" pageId="5" pageNumber="446">Akademik M.A. Lavrentyev</collectorName>
|
||
</emphasis>
|
||
, cruise 75, sta LV75-15/2
|
||
</materialsCitation>
|
||
,
|
||
<specimenCount id="9D31AEE4B64F1740FE87AB274080FECB" box="[281,404,295,320]" pageId="5" pageNumber="446" type="generic">1 specimen</specimenCount>
|
||
, in vicinity of
|
||
<taxonomicName id="4C371EEEB64F1740FDB1AB2843A3FECB" authorityName="Dall" authorityYear="1891" box="[559,695,295,319]" class="Bivalvia" family="Vesicomyidae" genus="Calyptogena" kingdom="Animalia" order="Venerida" pageId="5" pageNumber="446" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B943B97FB64F1740FDB1AB2843A3FECB" box="[559,695,295,319]" italics="true" pageId="5" pageNumber="446">Calyptogena</emphasis>
|
||
</taxonomicName>
|
||
bed on soft sediment.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32D36E6B64F1740FF59AB43452DFE97" box="[199,1081,331,356]" pageId="5" pageNumber="446" type="etymology">
|
||
<paragraph id="8B88656DB64F1740FF59AB43452DFE97" blockId="5.[151,1437,151,1903]" box="[199,1081,331,356]" pageId="5" pageNumber="446">
|
||
<emphasis id="B943B97FB64F1740FF59AB43405CFE90" bold="true" box="[199,328,332,356]" inLineHeading="true" pageId="5" pageNumber="446" reason="1">Etymology.</emphasis>
|
||
The species epithet refers to characteristic axial ribs (‘costa’, L. ‘rib’).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32D36E6B64F1740FF59AB6041F3FC8B" pageId="5" pageNumber="446" type="description">
|
||
<paragraph id="8B88656DB64F1740FF59AB604321FD53" blockId="5.[151,1437,151,1903]" pageId="5" pageNumber="446">
|
||
<emphasis id="B943B97FB64F1740FF59AB6040D8FE73" bold="true" box="[199,460,367,391]" inLineHeading="true" pageId="5" pageNumber="446" reason="1">
|
||
Description (
|
||
<typeStatus id="548CDBCFB64F1740FEC7AB6040D4FE73" box="[345,448,367,391]" pageId="5" pageNumber="446" type="holotype">holotype</typeStatus>
|
||
).
|
||
</emphasis>
|
||
Shell (
|
||
<figureCitation id="130C79E8B64F1740FD89AB60439FFE7C" box="[535,651,367,392]" captionStart="FIGURE 2" captionStartId="4.[151,250,1843,1867]" captionTargetBox="[151,1435,308,1820]" captionTargetId="figure-87@4.[151,1435,308,1820]" captionTargetPageId="4" captionText="FIGURE 2. Astyris axicostata n. sp. (A–I) A–D. Holotype, SL 7.3 mm (A–C—dried shell, D—wet shell with body seen through transparency). E. Paratype 1, SL 7.4 mm. F–G. Paratype 2, SL 7.0 mm. H–I. Paratype 3, juvenile with intact protoconch, SL 3.0 mm. J. Astyris cf. rosacea (Gould, 1840), SL 12.1 mm. A–G—at the same scale, H–I—enlarged, J—reduced." figureDoi="http://doi.org/10.5281/zenodo.7804414" httpUri="https://zenodo.org/record/7804414/files/figure.png" pageId="5" pageNumber="446">Fig. 2A–D</figureCitation>
|
||
) moderate-sized for genus (up to SL
|
||
<quantity id="4CCFC888B64F1740FB88AB7F457FFE7C" box="[1046,1131,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="5" pageNumber="446" unit="mm" value="7.4">7.4 mm</quantity>
|
||
), tall-spired. Apex decollate, leaving 4.5 whorls remaining. Teleoconch whorls evenly rounded, convex, very thin and fragile, semi-transparent, very light yellow. Periostracum very thin, smooth, transparent, and tightly adherent to shell. Spiral sculpture of rather indistinct spiral cordlets slightly varying in width, best pronounced on penultimate whorl (16 on penultimate whorl) and less distinct on last whorl, very obscure on canal. Axial sculpture of very thin but distinct weakly sinuous axial ribs present on all teleoconch whorls, more distinct on penultimate and last whorls, and extending to canal, around 30 on last whorl. Aperture lacking dentition, semi-circular in shape, siphonate. Outer lip simple, not thickened, completely smooth on inside. Columella weakly concave, simple, with slightly thickened narrow callus. Siphonal canal short with weakly constricted but distinct siphonal notch.
|
||
</paragraph>
|
||
<paragraph id="8B88656DB64F1740FF59A8BC45CFFD3F" blockId="5.[151,1437,151,1903]" box="[199,1243,691,715]" pageId="5" pageNumber="446">Operculum corneous, small, irregularly oval, length about half of aperture height. Nucleus lateral.</paragraph>
|
||
<paragraph id="8B88656DB64F1740FF59A8D84514FCE7" blockId="5.[151,1437,151,1903]" pageId="5" pageNumber="446">
|
||
Protoconch persists in
|
||
<specimenCount id="9D31AEE4B64F1740FE27A8D74328FD1B" box="[441,572,727,751]" pageId="5" pageNumber="446" type="juvenile">one juvenile</specimenCount>
|
||
<typeStatus id="548CDBCFB64F1740FDDDA8D7438BFD04" box="[579,671,728,752]" pageId="5" pageNumber="446" type="paratype">paratype</typeStatus>
|
||
with SL 3.0 mm (
|
||
<figureCitation id="130C79E8B64F1740FCFEA8D842C2FD04" box="[864,982,727,752]" captionStart="FIGURE 2" captionStartId="4.[151,250,1843,1867]" captionTargetBox="[151,1435,308,1820]" captionTargetId="figure-87@4.[151,1435,308,1820]" captionTargetPageId="4" captionText="FIGURE 2. Astyris axicostata n. sp. (A–I) A–D. Holotype, SL 7.3 mm (A–C—dried shell, D—wet shell with body seen through transparency). E. Paratype 1, SL 7.4 mm. F–G. Paratype 2, SL 7.0 mm. H–I. Paratype 3, juvenile with intact protoconch, SL 3.0 mm. J. Astyris cf. rosacea (Gould, 1840), SL 12.1 mm. A–G—at the same scale, H–I—enlarged, J—reduced." figureDoi="http://doi.org/10.5281/zenodo.7804414" httpUri="https://zenodo.org/record/7804414/files/figure.png" pageId="5" pageNumber="446">Fig. 2 E–F</figureCitation>
|
||
), paucispiral, of about 1.5 whorls, smooth, diameter
|
||
<quantity id="4CCFC888B64F1740FF65A8F44048FCE7" box="[251,348,763,787]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="446" unit="mm" value="0.75">0.75 mm</quantity>
|
||
, transition to teleoconch is poorly demarcated by a weak varix.
|
||
</paragraph>
|
||
<paragraph id="8B88656DB64F1740FF59A91041F3FC8B" blockId="5.[151,1437,151,1903]" pageId="5" pageNumber="446">
|
||
Radula (
|
||
<figureCitation id="130C79E8B64F1740FEBCA9104061FCCC" box="[290,373,799,824]" captionStart="FIGURE 3" captionStartId="6.[151,250,1343,1367]" captionTargetBox="[151,1435,181,1320]" captionTargetId="figure-22@6.[151,1435,181,1320]" captionTargetPageId="6" captionText="FIGURE 3. A–B. Radulae of studied Astyris spp. A. Astyris axicostata n. sp., specimen from the type locality. B. Astyris cf. rosacea(shell on Fig. 2 J). C–D. Astyris axicostata n. sp. in situ, specimens of Astyris axicostatus are marked with red arrows, insert shows enlarged fragment of the photos. C. Sta LV82-2/4, northern summit of Piip volcano, 55.24′57′′N, 167.16′35′′E, 384 м, 13.06.2018; rocks with bacterial mats. In addition to the Astyris, numerous isopods Eurycope andreyi (Munnopsidae) and Corallimorphus cf. pilatus (Corallimorpharia,Anthozoa) are seen. D. Type locality, rock with dense aggregations of Epizoanthus sp. (Zoantharia, Anthozoa)." figureDoi="http://doi.org/10.5281/zenodo.7804416" httpUri="https://zenodo.org/record/7804416/files/figure.png" pageId="5" pageNumber="446">Fig. 3A</figureCitation>
|
||
) rachiglossan, typical of columbellids with acuspate smooth plate-like central tooth and one lateral tooth on each side. Laterals with narrow bases and three cusps, basal one obtuse and two distal ones sharp and closely spaced.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32D36E6B64F1740FF59A98440BCFC33" pageId="5" pageNumber="446" type="distribution">
|
||
<paragraph id="8B88656DB64F1740FF59A98440BCFC33" blockId="5.[151,1437,151,1903]" pageId="5" pageNumber="446">
|
||
<emphasis id="B943B97FB64F1740FF59A984404EFC57" bold="true" box="[199,346,907,931]" inLineHeading="true" pageId="5" pageNumber="446" reason="1">Distribution.</emphasis>
|
||
Presently known only from both summits of the submarine Piip Volcano (southwestern Bering Sea) at the depths of
|
||
<quantity id="4CCFC888B64F1740FEB6A9A040B0FC33" box="[296,420,943,967]" metricMagnitude="2" metricUnit="m" metricValue="4.205" metricValueMax="4.68" metricValueMin="3.7299999999999995" pageId="5" pageNumber="446" unit="m" value="420.5" valueMax="468.0" valueMin="373.0">
|
||
373–
|
||
<specimenCount id="9D31AEE4B64F1740FEC3A9A040B0FC33" box="[349,420,943,967]" pageId="5" pageNumber="446" type="generic">468 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32D36E6B64F1740FF59A9DB434DFB30" pageId="5" pageNumber="446" type="biology_ecology">
|
||
<paragraph id="8B88656DB64F1740FF59A9DB450AFBAC" blockId="5.[151,1437,151,1903]" pageId="5" pageNumber="446">
|
||
<emphasis id="B943B97FB64F1740FF59A9DB4033FC18" bold="true" box="[199,295,980,1004]" inLineHeading="true" pageId="5" pageNumber="446" reason="1">Ecology.</emphasis>
|
||
Most specimens were collected from a single station on a rock among dense population of
|
||
<taxonomicName id="4C371EEEB64F1740FA86A9DC41A2FBFB" class="Anthozoa" family="Epizoanthidae" genus="Epizoanthus" kingdom="Animalia" order="Zoantharia" pageId="5" pageNumber="446" phylum="Cnidaria" rank="species" species="undetermined">
|
||
<emphasis id="B943B97FB64F1740FA86A9DC4488FC1F" box="[1304,1436,979,1003]" italics="true" pageId="5" pageNumber="446">Epizoanthus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
in an area without visible hydrothermal vent activity (
|
||
<figureCitation id="130C79E8B64F1740FC9AA9F84296FBE4" box="[772,898,1015,1040]" captionStart="FIGURE 3" captionStartId="6.[151,250,1343,1367]" captionTargetBox="[151,1435,181,1320]" captionTargetId="figure-22@6.[151,1435,181,1320]" captionTargetPageId="6" captionText="FIGURE 3. A–B. Radulae of studied Astyris spp. A. Astyris axicostata n. sp., specimen from the type locality. B. Astyris cf. rosacea(shell on Fig. 2 J). C–D. Astyris axicostata n. sp. in situ, specimens of Astyris axicostatus are marked with red arrows, insert shows enlarged fragment of the photos. C. Sta LV82-2/4, northern summit of Piip volcano, 55.24′57′′N, 167.16′35′′E, 384 м, 13.06.2018; rocks with bacterial mats. In addition to the Astyris, numerous isopods Eurycope andreyi (Munnopsidae) and Corallimorphus cf. pilatus (Corallimorpharia,Anthozoa) are seen. D. Type locality, rock with dense aggregations of Epizoanthus sp. (Zoantharia, Anthozoa)." figureDoi="http://doi.org/10.5281/zenodo.7804416" httpUri="https://zenodo.org/record/7804416/files/figure.png" pageId="5" pageNumber="446">Fig. 3 C–D</figureCitation>
|
||
), however
|
||
<specimenCount id="9D31AEE4B64F1740FC64A9F7458DFBFB" box="[1018,1177,1015,1039]" pageId="5" pageNumber="446" type="generic">two specimens</specimenCount>
|
||
were obtained from the hydrothermal vent habitats in close proximity to
|
||
<taxonomicName id="4C371EEEB64F1740FD04AE144236FBC7" authorityName="Dall" authorityYear="1891" box="[666,802,1051,1075]" class="Bivalvia" family="Vesicomyidae" genus="Calyptogena" kingdom="Animalia" order="Venerida" pageId="5" pageNumber="446" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B943B97FB64F1740FD04AE144236FBC7" box="[666,802,1051,1075]" italics="true" pageId="5" pageNumber="446">Calyptogena</emphasis>
|
||
</taxonomicName>
|
||
beds and bacterial mats. In addition, a study of photographs shows that some molluscs located close to bacterial mats may be
|
||
<taxonomicName id="4C371EEEB64F1740FCCEAE4F42C8FBA3" authority="Kantor & Zvonareva & Krylova, 2023" authorityName="Kantor & Zvonareva & Krylova" authorityYear="2023" box="[848,988,1088,1111]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="axicostata" status="sp. nov.">
|
||
<emphasis id="B943B97FB64F1740FCCEAE4F42C8FBA3" box="[848,988,1088,1111]" italics="true" pageId="5" pageNumber="446">A. axicostata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B97FB64F1740FC7CAE4F450AFBAC" bold="true" box="[994,1054,1088,1112]" pageId="5" pageNumber="446">
|
||
<taxonomicNameLabel id="A2700404B64F1740FC7CAE4F450AFBAC" box="[994,1054,1088,1112]" pageId="5" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B88656DB64F1740FF59AE6C434DFB30" blockId="5.[151,1437,151,1903]" pageId="5" pageNumber="446">
|
||
The value of δ13C of a specimen of
|
||
<taxonomicName id="4C371EEEB64F1740FDFBAE6B43E7FB8F" authority="Kantor & Zvonareva & Krylova, 2023" authorityName="Kantor & Zvonareva & Krylova" authorityYear="2023" box="[613,755,1124,1147]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="axicostata" status="sp. nov.">
|
||
<emphasis id="B943B97FB64F1740FDFBAE6B43E7FB8F" box="[613,755,1124,1147]" italics="true" pageId="5" pageNumber="446">A. axicostata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B97FB64F1740FD63AE6B4228FB88" bold="true" box="[765,828,1124,1148]" pageId="5" pageNumber="446">
|
||
<taxonomicNameLabel id="A2700404B64F1740FD63AE6B4228FB88" box="[765,828,1124,1148]" pageId="5" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from a site without bacterial mats indicated prevailing contribution of photosynthetically derived carbon from sinking POM (particulated organic matter) in their nutrition (Kiyashko, Mordukhovich, pers. comm.)..
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32D36E6B64F1740FF59AEDF4059F89B" pageId="5" pageNumber="446" type="discussion">
|
||
<paragraph id="8B88656DB64F1740FF59AEDF4590FADB" blockId="5.[151,1437,151,1903]" pageId="5" pageNumber="446">
|
||
<emphasis id="B943B97FB64F1740FF59AEDF4022FB1C" bold="true" box="[199,310,1232,1256]" inLineHeading="true" pageId="5" pageNumber="446" reason="1">Remarks.</emphasis>
|
||
The present new species demonstrates little degree of conchological variation. The spiral sculpture can be more distinct in some specimens than in
|
||
<typeStatus id="548CDBCFB64F1740FDE8AEFC43C7FAFF" box="[630,723,1267,1291]" pageId="5" pageNumber="446" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="130C79E8B64F1740FD7AAEFC4275FAF8" box="[740,865,1267,1292]" captionStart="FIGURE 2" captionStartId="4.[151,250,1843,1867]" captionTargetBox="[151,1435,308,1820]" captionTargetId="figure-87@4.[151,1435,308,1820]" captionTargetPageId="4" captionText="FIGURE 2. Astyris axicostata n. sp. (A–I) A–D. Holotype, SL 7.3 mm (A–C—dried shell, D—wet shell with body seen through transparency). E. Paratype 1, SL 7.4 mm. F–G. Paratype 2, SL 7.0 mm. H–I. Paratype 3, juvenile with intact protoconch, SL 3.0 mm. J. Astyris cf. rosacea (Gould, 1840), SL 12.1 mm. A–G—at the same scale, H–I—enlarged, J—reduced." figureDoi="http://doi.org/10.5281/zenodo.7804414" httpUri="https://zenodo.org/record/7804414/files/figure.png" pageId="5" pageNumber="446">Fig. 2 F–G</figureCitation>
|
||
). The new species is most similar to
|
||
<taxonomicName id="4C371EEEB64F1740FB6AAEFB4481FAFF" box="[1268,1429,1267,1291]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="thermophila">
|
||
<emphasis id="B943B97FB64F1740FB6AAEFB4481FAFF" box="[1268,1429,1267,1291]" italics="true" pageId="5" pageNumber="446">A. thermophila</emphasis>
|
||
</taxonomicName>
|
||
, differing in more rounded teleoconch whorls and in the presence of thin but distinct axial ribs.
|
||
</paragraph>
|
||
<paragraph id="8B88656DB64F1740FF59AF334059F89B" blockId="5.[151,1437,151,1903]" pageId="5" pageNumber="446">
|
||
<taxonomicName id="4C371EEEB64F1740FF59AF334092FAA7" authority="Kantor & Zvonareva & Krylova, 2023" authorityName="Kantor & Zvonareva & Krylova" authorityYear="2023" box="[199,390,1340,1363]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="axicostata" status="sp. nov.">
|
||
<emphasis id="B943B97FB64F1740FF59AF334092FAA7" box="[199,390,1340,1363]" italics="true" pageId="5" pageNumber="446">Astyris axicostata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B97FB64F1740FE13AF3340DEFAA0" bold="true" box="[397,458,1340,1364]" pageId="5" pageNumber="446">
|
||
<taxonomicNameLabel id="A2700404B64F1740FE13AF3340DEFAA0" box="[397,458,1340,1364]" pageId="5" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is attributed to
|
||
<taxonomicName id="4C371EEEB64F1740FDEAAF3343AAFAA7" authorityName="Kantor & Zvonareva & Krylova" authorityYear="2023" box="[628,702,1340,1363]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B943B97FB64F1740FDEAAF3343AAFAA7" box="[628,702,1340,1363]" italics="true" pageId="5" pageNumber="446">Astyris</emphasis>
|
||
</taxonomicName>
|
||
since it belongs to the same molecularly defined clade as
|
||
<taxonomicName id="4C371EEEB64F1740FAAEAF3341FEFA83" baseAuthorityName="Gould" baseAuthorityYear="1840" class="Gastropoda" family="Columbellidae" genus="Astyris" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="rosacea">
|
||
<emphasis id="B943B97FB64F1740FAAEAF33446EFAA7" box="[1328,1402,1340,1363]" italics="true" pageId="5" pageNumber="446">Astyris</emphasis>
|
||
cf.
|
||
<emphasis id="B943B97FB64F1740FF09AF6F41FEFA83" box="[151,234,1376,1399]" italics="true" pageId="5" pageNumber="446">rosacea</emphasis>
|
||
</taxonomicName>
|
||
, very similar to the
|
||
<typeStatus id="548CDBCFB64F1740FE56AF6F40E2FA8C" box="[456,502,1376,1400]" pageId="5" pageNumber="446">type</typeStatus>
|
||
species of the genus. Correspondingly the tentative position of
|
||
<taxonomicName id="4C371EEEB64F1740FB35AF6F4458FA83" box="[1195,1356,1375,1399]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="thermophila">
|
||
<emphasis id="B943B97FB64F1740FB35AF6F4458FA83" box="[1195,1356,1375,1399]" italics="true" pageId="5" pageNumber="446">A. thermophila</emphasis>
|
||
</taxonomicName>
|
||
is here confirmed. The radulae of all three studied
|
||
<taxonomicName id="4C371EEEB64F1740FDF8AF8B4210FA6F" box="[614,772,1411,1435]" pageId="5" pageNumber="446">
|
||
<emphasis id="B943B97FB64F1740FDF8AF8B43A4FA6F" box="[614,688,1412,1435]" italics="true" pageId="5" pageNumber="446">Astyris</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
are very similar (
|
||
<figureCitation id="130C79E8B64F1740FC5FAF8C4515FA68" box="[961,1025,1411,1436]" captionStart="FIGURE 3" captionStartId="6.[151,250,1343,1367]" captionTargetBox="[151,1435,181,1320]" captionTargetId="figure-22@6.[151,1435,181,1320]" captionTargetPageId="6" captionText="FIGURE 3. A–B. Radulae of studied Astyris spp. A. Astyris axicostata n. sp., specimen from the type locality. B. Astyris cf. rosacea(shell on Fig. 2 J). C–D. Astyris axicostata n. sp. in situ, specimens of Astyris axicostatus are marked with red arrows, insert shows enlarged fragment of the photos. C. Sta LV82-2/4, northern summit of Piip volcano, 55.24′57′′N, 167.16′35′′E, 384 м, 13.06.2018; rocks with bacterial mats. In addition to the Astyris, numerous isopods Eurycope andreyi (Munnopsidae) and Corallimorphus cf. pilatus (Corallimorpharia,Anthozoa) are seen. D. Type locality, rock with dense aggregations of Epizoanthus sp. (Zoantharia, Anthozoa)." figureDoi="http://doi.org/10.5281/zenodo.7804416" httpUri="https://zenodo.org/record/7804416/files/figure.png" pageId="5" pageNumber="446">Fig. 3</figureCitation>
|
||
), although in
|
||
<taxonomicName id="4C371EEEB64F1740FB09AF8B4423FA6F" box="[1175,1335,1411,1435]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="thermophila">
|
||
<emphasis id="B943B97FB64F1740FB09AF8B4423FA6F" box="[1175,1335,1411,1435]" italics="true" pageId="5" pageNumber="446">A. thermophila</emphasis>
|
||
</taxonomicName>
|
||
the basal cusp of the lateral teeth is more distinct and better defined in comparison with
|
||
<taxonomicName id="4C371EEEB64F1740FC4DAFA7454AFA4B" authority="Kantor & Zvonareva & Krylova, 2023" authorityName="Kantor & Zvonareva & Krylova" authorityYear="2023" box="[979,1118,1448,1471]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="axicostata" status="sp. nov.">
|
||
<emphasis id="B943B97FB64F1740FC4DAFA7454AFA4B" box="[979,1118,1448,1471]" italics="true" pageId="5" pageNumber="446">A. axicostata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B97FB64F1740FBFDAFA7458AFA34" bold="true" box="[1123,1182,1448,1472]" pageId="5" pageNumber="446">
|
||
<taxonomicNameLabel id="A2700404B64F1740FBFDAFA7458AFA34" box="[1123,1182,1448,1472]" pageId="5" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C371EEEB64F1740FB4FAFA7448DFA4B" baseAuthorityName="Gould" baseAuthorityYear="1840" box="[1233,1433,1447,1471]" class="Gastropoda" family="Columbellidae" genus="Astyris" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="rosacea">
|
||
<emphasis id="B943B97FB64F1740FB4FAFA7440FFA4B" box="[1233,1307,1448,1471]" italics="true" pageId="5" pageNumber="446">Astyris</emphasis>
|
||
cf.
|
||
<emphasis id="B943B97FB64F1740FADCAFA7448DFA4B" box="[1346,1433,1448,1471]" italics="true" pageId="5" pageNumber="446">rosacea</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B97FB64F1740FA07AFA74043FA17" italics="true" pageId="5" pageNumber="446">
|
||
.
|
||
<taxonomicName id="4C371EEEB64F1740FF09AFC34043FA17" authority="Kantor & Zvonareva & Krylova, 2023" authorityName="Kantor & Zvonareva & Krylova" authorityYear="2023" box="[151,343,1484,1507]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="axicostata" status="sp. nov.">Astyris axicostata</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="B943B97FB64F1740FEC0AFC3408EFA10" bold="true" box="[350,410,1484,1508]" pageId="5" pageNumber="446">
|
||
<taxonomicNameLabel id="A2700404B64F1740FEC0AFC3408EFA10" box="[350,410,1484,1508]" pageId="5" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the fourth species of the genus recorded from reducing environments and the second species found in hydrothermal vent areas (
|
||
<tableCitation id="C6B550D6B64F1740FD8AAFE04373F9FC" box="[532,615,1519,1544]" captionStart="TABLE 1" captionStartId="1.[152,245,477,501]" captionTargetPageId="1" captionText="TABLE 1. Species of the genus Astyris found in reducing habitats." httpUri="http://table.plazi.org/id/DF4835E5B64B1744FF06ABD24272FE01" pageId="5" pageNumber="446" tableUuid="DF4835E5B64B1744FF06ABD24272FE01">Table 1</tableCitation>
|
||
). As for the feeding
|
||
<typeStatus id="548CDBCFB64F1740FCCFAFFF426BF9FC" box="[849,895,1520,1544]" pageId="5" pageNumber="446">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C371EEEB64F1740FC39AFFF4522F9F3" authority="Kantor & Zvonareva & Krylova, 2023" authorityName="Kantor & Zvonareva & Krylova" authorityYear="2023" box="[935,1078,1520,1543]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="axicostata" status="sp. nov.">
|
||
<emphasis id="B943B97FB64F1740FC39AFFF4522F9F3" box="[935,1078,1520,1543]" italics="true" pageId="5" pageNumber="446">A. axicostata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B97FB64F1740FBA1AFFF4560F9FC" bold="true" box="[1087,1140,1520,1544]" pageId="5" pageNumber="446">
|
||
<taxonomicNameLabel id="A2700404B64F1740FBA1AFFF4560F9FC" box="[1087,1140,1520,1544]" pageId="5" pageNumber="446" rank="species">n.sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, preliminary data on δ13C indicated organic matter mainly of photosynthetic origin in their diet (Kiyashko, Mordukhovich, pers. comm.). Thus, although found in the area of hydrothermal activity, the species is probably not specific for hydrothermal vents but rather has high tolerance to environmental factors allowing inhabiting areas of hydrothermal activity.
|
||
<taxonomicName id="4C371EEEB64F1740FB3FAC534461F987" box="[1185,1397,1627,1651]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="thermophila">
|
||
<emphasis id="B943B97FB64F1740FB3FAC534461F987" box="[1185,1397,1627,1651]" italics="true" pageId="5" pageNumber="446">Astyris thermophila</emphasis>
|
||
</taxonomicName>
|
||
, so far the only columbellid species of hydrothermal vent community, was found in tubeworm bushes attached on the tubes and in washings of
|
||
<taxonomicName id="4C371EEEB64F1740FEF2ACAC4318F94F" authorityName="Kenk & B. R. Wilson" authorityYear="1985" box="[364,524,1699,1723]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytilida" pageId="5" pageNumber="446" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B943B97FB64F1740FEF2ACAC4318F94F" box="[364,524,1699,1723]" italics="true" pageId="5" pageNumber="446">Bathymodiolus</emphasis>
|
||
</taxonomicName>
|
||
mussels. It was suggested that
|
||
<taxonomicName id="4C371EEEB64F1740FCFFACAB4516F94F" box="[865,1026,1699,1723]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="thermophila">
|
||
<emphasis id="B943B97FB64F1740FCFFACAB4516F94F" box="[865,1026,1699,1723]" italics="true" pageId="5" pageNumber="446">A. thermophila</emphasis>
|
||
</taxonomicName>
|
||
is a predator or ovophagous (
|
||
<bibRefCitation id="EFA6189CB64F1740FAD9ACAC41E7F914" author="Chen, C. & Watanabe, H. K. & Araya, J. F." pageId="5" pageNumber="446" pagination="592 - 596" refId="ref4587" refString="Chen, C., Watanabe, H. K. & Araya, J. F. (2017) First columbellid species (Gastropoda: Buccinoidea) from deep-sea hydrothermal vents, discovered in Okinawa Trough, Japan. Zootaxa, 4363 (4), 592 - 596. https: // doi. org / 10.11646 / zootaxa. 4363.4.13" type="journal article" year="2017">
|
||
Chen
|
||
<emphasis id="B943B97FB64F1740FA17ACAB41A7F92B" italics="true" pageId="5" pageNumber="446">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
). It is not clear whether
|
||
<taxonomicName id="4C371EEEB64F1740FE6CACC74386F92B" box="[498,658,1735,1759]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="thermophila">
|
||
<emphasis id="B943B97FB64F1740FE6CACC74386F92B" box="[498,658,1735,1759]" italics="true" pageId="5" pageNumber="446">A. thermophila</emphasis>
|
||
</taxonomicName>
|
||
is a species obligate to the hydrothermal vent habitats, since very similar specimens were recently collected at the methane seeps of the Koryak Slope of
|
||
<collectingRegion id="49F3AB8FB64F1740FC77ACE44540F8F7" box="[1001,1108,1771,1795]" country="Russia" name="Chukotka" pageId="5" pageNumber="446">Chukotka</collectingRegion>
|
||
, western Bering Sea (
|
||
<bibRefCitation id="EFA6189CB64F1740FAA2ACE44024F8DC" author="Nekhaev, I. O. & Chaban, E. M. & Kantor, Yu. I. & Kuchsh, D. A. & Matveeva, K. & Rybakova, E." pageId="5" pageNumber="446" pagination="105164" refId="ref5921" refString="Nekhaev, I. O., Chaban, E. M., Kantor, Yu. I., Kuchsh, D. A., Matveeva, K. & Rybakova, E. (2022) Shell-bearing Gastropoda from the methane seeps and hydrothermal vents of the Bering Sea: A preliminary description. Deep Sea Research Part II: Topical Studies in Oceanography, 204, 105164. https: // doi. org / 10.1016 / j. dsr 2.2022.105164" type="journal article" year="2022">
|
||
Nekhaev
|
||
<emphasis id="B943B97FB64F1740FF09AD1F41C4F8D3" box="[151,208,1807,1831]" italics="true" pageId="5" pageNumber="446">et al.</emphasis>
|
||
2022: 9
|
||
</bibRefCitation>
|
||
). We failed in sequencing those specimens and therefore their conspecifity with
|
||
<taxonomicName id="4C371EEEB64F1740FB2FAD1F4440F8D3" box="[1201,1364,1807,1831]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="thermophila">
|
||
<emphasis id="B943B97FB64F1740FB2FAD1F4440F8D3" box="[1201,1364,1807,1831]" italics="true" pageId="5" pageNumber="446">A. thermophila</emphasis>
|
||
</taxonomicName>
|
||
is not presently confirmed. But it may appear that both
|
||
<taxonomicName id="4C371EEEB64F1740FD34AD3B425FF8BF" box="[682,843,1843,1867]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="thermophila">
|
||
<emphasis id="B943B97FB64F1740FD34AD3B425FF8BF" box="[682,843,1843,1867]" italics="true" pageId="5" pageNumber="446">A. thermophila</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C371EEEB64F1740FC1EAD3B4519F8BF" authority="Kantor & Zvonareva & Krylova, 2023" authorityName="Kantor & Zvonareva & Krylova" authorityYear="2023" box="[896,1037,1844,1867]" class="Gastropoda" family="Columbellidae" genus="Astyris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="446" phylum="Mollusca" rank="species" species="axicostata" status="sp. nov.">
|
||
<emphasis id="B943B97FB64F1740FC1EAD3B4519F8BF" box="[896,1037,1844,1867]" italics="true" pageId="5" pageNumber="446">A. axicostata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B97FB64F1740FB8AAD3B4544F8B8" bold="true" box="[1044,1104,1844,1868]" pageId="5" pageNumber="446">
|
||
<taxonomicNameLabel id="A2700404B64F1740FB8AAD3B4544F8B8" box="[1044,1104,1844,1868]" pageId="5" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can dwell in rather wide
|
||
<typeStatus id="548CDBCFB64F1740FAFAAD3B4488F8B8" box="[1380,1436,1844,1868]" pageId="5" pageNumber="446">types</typeStatus>
|
||
of environments.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |