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<document id="209C8828D4F293B5693789D8A781310B" ID-DOI="10.11646/zootaxa.5448.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="11231707" ID-ZooBank="79A27DBA-56C0-4D86-BA36-35D4070CED9A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1716279273055" checkinUser="plazi" docAuthor="Joshi, Shantanu, Sarkar, Vivek, Sawant, Dattaprasad, Pawar, Ujwala &amp; Kunte, Krushnamegh" docDate="2024" docId="039E2C42BD3EFF9F63F0F94EFD8D4CE6" docLanguage="en" docName="zootaxa.5448.3.2.pdf" docOrigin="Zootaxa 5448 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5448.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protosticta fraseri Kennedy 1936" docType="treatment" docVersion="1" lastPageNumber="352" masterDocId="FFA7543ABD3CFF9B6367FFB5FFF24E17" masterDocTitle="Description of Protosticta khasia sp. nov. and Yunnanosticta siangi sp. nov., with new records of P. samtsensis Gurung &amp; Phan, 2023 from Northeast India (Odonata: Zygoptera: Platystictidae)" masterLastPageNumber="370" masterPageNumber="348" pageNumber="350" updateTime="1716379561080" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="825A28D69DE78B57891400C3C2DBA280">Description of Protosticta khasia sp. nov. and Yunnanosticta siangi sp. nov., with new records of P. samtsensis Gurung &amp; Phan, 2023 from Northeast India (Odonata: Zygoptera: Platystictidae)</mods:title>
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<mods:namePart id="F49948644F5FB467EA77C9AC1F88722A">Joshi, Shantanu</mods:namePart>
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<mods:affiliation id="58E25A32921FECA9784A2BAEBC139A64">Department of Biological Sciences, University of Arkansas, Fayetteville, 72701, Arkansas, USA &amp; National Centre for Biological Sciences, GKVK Campus, Bellary Rd, Bengaluru, 560065, Karnataka, India</mods:affiliation>
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<mods:namePart id="EEAED34B35C791CE5FB6BAAE00D92BD2">Sarkar, Vivek</mods:namePart>
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<mods:affiliation id="0D134A24C91EDF14EE25ADDA2E7B3511">Wildlife Institute of India, Chandrabani, Dehradun, 248001, Uttarakhand, India</mods:affiliation>
<mods:nameIdentifier id="2118BF4A055B583F7B871D2779671DFD" type="email">viveksarkar87@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D6342B3ED1ADCE1D83C3EA0691CBCD98">Sawant, Dattaprasad</mods:namePart>
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<mods:affiliation id="C7343A0D9E88A87F8126DE8388E2F1B1">Government Medical College and Hospital, Sindhudurg, 416534, Maharashtra, India</mods:affiliation>
<mods:nameIdentifier id="033015BF4F34402B7859F6E35D6405F8" type="email">dattaprasad.101@gmail.com</mods:nameIdentifier>
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<mods:namePart id="36F71853E9238C1A9F209AB23DD631BE">Pawar, Ujwala</mods:namePart>
<mods:nameIdentifier id="58BFF99ABD8C8C32B667A0E5C00A43F3" type="ORCID">0000-0003-1281-2228</mods:nameIdentifier>
<mods:affiliation id="2E24A1BD19274BCD759468AC93C5EC13">National Centre for Biological Sciences, GKVK Campus, Bellary Rd, Bengaluru, 560065, Karnataka, India</mods:affiliation>
<mods:nameIdentifier id="8781CCE42E290720759F0F65D0273365" type="email">ujwalamp@ncbs.res.in</mods:nameIdentifier>
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<mods:namePart id="48F9208A06E7991F308611D36829AF10">Kunte, Krushnamegh</mods:namePart>
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<mods:affiliation id="802A092E14F2FFE3F368EFD0F5F34604">National Centre for Biological Sciences, GKVK Campus, Bellary Rd, Bengaluru, 560065, Karnataka, India</mods:affiliation>
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<mods:date id="11F0854E0188C5690DC2BA0B63EEE274">2024</mods:date>
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<treatment id="039E2C42BD3EFF9F63F0F94EFD8D4CE6" ID-DOI="http://doi.org/10.5281/zenodo.11231800" ID-Zenodo-Dep="11231800" LSID="urn:lsid:plazi:treatment:039E2C42BD3EFF9F63F0F94EFD8D4CE6" httpUri="http://treatment.plazi.org/id/039E2C42BD3EFF9F63F0F94EFD8D4CE6" lastPageId="4" lastPageNumber="352" pageId="2" pageNumber="350">
<subSubSection id="C32DCEDFBD3EFF9963F0F94EFE1A4902" box="[151,488,1787,1813]" pageId="2" pageNumber="350" type="nomenclature">
<paragraph id="8B889D54BD3EFF9963F0F94EFE1A4902" blockId="2.[151,488,1787,1849]" box="[151,488,1787,1813]" pageId="2" pageNumber="350">
<heading id="D0C02A38BD3EFF9963F0F94EFE1A4902" bold="true" box="[151,488,1787,1813]" fontSize="11" level="1" pageId="2" pageNumber="350" reason="1">
<taxonomicName id="4C37E6D7BD3EFF9963F0F94EFE1A4902" ID-CoL="77YKS" authority="Kennedy" authorityName="Kennedy" authorityYear="1936" box="[151,488,1787,1813]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="350" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B9434146BD3EFF9963F0F94EFE1A4902" bold="true" box="[151,488,1787,1813]" pageId="2" pageNumber="350">
<emphasis id="B9434146BD3EFF9963F0F94EFE834902" bold="true" box="[151,369,1787,1813]" italics="true" pageId="2" pageNumber="350">Protosticta fraseri</emphasis>
Kennedy
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32DCEDFBD3EFF9963F0F8AAFF1E492E" box="[151,236,1823,1849]" pageId="2" pageNumber="350" type="description">
<paragraph id="8B889D54BD3EFF9963F0F8AAFF1E492E" blockId="2.[151,488,1787,1849]" box="[151,236,1823,1849]" pageId="2" pageNumber="350">
(
<figureCitation id="130C81D1BD3EFF9963F8F8AAFF11492E" box="[159,227,1823,1849]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-20@3.[215,1373,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Protosticta fraseri holotype male (UMMZI-199248): (a) lateral habitus; (bc) thorax: (b) dorsal view; (c) lateral view; (de) caudal appendages: (d) dorsal view; (e) lateral view. Photo Courtesy: K. Eldredge / Museum of Zoology, University of Michigan." figureDoi="http://doi.org/10.5281/zenodo.11231711" httpUri="https://zenodo.org/record/11231711/files/figure.png" pageId="2" pageNumber="350">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32DCEDFBD3EFF9F63F0F8D2FEE84F0A" lastPageId="4" lastPageNumber="352" pageId="2" pageNumber="350" type="discussion">
<paragraph id="8B889D54BD3EFF9F63F0F8D2FEE84F0A" blockId="2.[151,1437,1895,2030]" lastBlockId="4.[151,1437,151,285]" lastPageId="4" lastPageNumber="352" pageId="2" pageNumber="350">
<emphasis id="B9434146BD3EFF9963F0F8D2FEED4996" bold="true" box="[151,287,1895,1921]" pageId="2" pageNumber="350">Comments.</emphasis>
<bibRefCitation id="EFA6E0A5BD3EFF99624EF8D2FE104996" author="Kennedy, C. H." box="[297,482,1895,1921]" pageId="2" pageNumber="350" pagination="67 - 71" refId="ref9164" refString="Kennedy, C. H. (1936) Protosticta fraseri, a new species of dragonfly (Platystictinae: Odonata) from Assam, India. Proceedings of the Royal Entomological Society of London, Series B (Taxonomy), 5 (4), 67 - 71. https: // doi. org / 10.1111 / j. 1365 - 3113.1936. tb 01317. x" type="journal article" year="1936">Kennedy (1936)</bibRefCitation>
described this species based on
<specimenCount id="9D3156DDBD3EFF99603CF8D2FC2F4996" box="[859,989,1895,1921]" count="3" pageId="2" pageNumber="350" type="male">three males</specimenCount>
and
<specimenCount id="9D3156DDBD3EFF99677EF8DDFB644996" box="[1049,1174,1895,1922]" count="1" pageId="2" pageNumber="350" type="female">one female</specimenCount>
which were part of “a small shipment of purchased dragonflies collected by U. Nissor Singh in
<collectingRegion id="49F353B6BD3EFF9960A4F83EFBE049B2" box="[963,1042,1931,1957]" country="India" name="Assam" pageId="2" pageNumber="350">Assam</collectingRegion>
,
<collectingCountry id="F320DDC4BD3EFF99677AF83EFBA549B2" box="[1053,1111,1931,1957]" name="India" pageId="2" pageNumber="350">India</collectingCountry>
during February, and March, 1935”. No precise locality was mentioned in the original description (
<collectingRegion id="49F353B6BD3EFF9960D4F81AFBF349DE" box="[947,1025,1967,1993]" country="India" name="Assam" pageId="2" pageNumber="350">Assam</collectingRegion>
referred to a large area of NE
<collectingCountry id="F320DDC4BD3EFF996604F81AFA6F49DE" box="[1379,1437,1967,1993]" name="India" pageId="2" pageNumber="350">India</collectingCountry>
during that era) and since then no further records have been published of the species.
<bibRefCitation id="EFA6E0A5BD3EFF99672DF866FB0D49FA" author="Kennedy, C. H." box="[1098,1279,2003,2029]" pageId="2" pageNumber="350" pagination="67 - 71" refId="ref9164" refString="Kennedy, C. H. (1936) Protosticta fraseri, a new species of dragonfly (Platystictinae: Odonata) from Assam, India. Proceedings of the Royal Entomological Society of London, Series B (Taxonomy), 5 (4), 67 - 71. https: // doi. org / 10.1111 / j. 1365 - 3113.1936. tb 01317. x" type="journal article" year="1936">Kennedy (1936)</bibRefCitation>
mentions that the types were deposited in Ohio State University.
<bibRefCitation id="EFA6E0A5BD38FF9F61B3FF22FC3A4EA6" author="Garrison, R. &amp; von Ellenrieder, N. &amp; O'Brien, M." box="[724,968,151,177]" pageId="4" pageNumber="352" pagination="1 - 73" refId="ref8746" refString="Garrison, R., von Ellenrieder, N. &amp; O'Brien, M. (2003) An annotated list of the name-bearing types of species-group names in Odonata preserved in the University of Michigan Museum of Zoology. Occasional Papers of the Museum of Zoology, University of Michigan, 736, 1 - 73." type="journal article" year="2003">
Garrison
<emphasis id="B9434146BD38FF9F605AFF22FC844EA6" box="[829,886,151,177]" italics="true" pageId="4" pageNumber="352">et al.</emphasis>
(2003)
</bibRefCitation>
published a list of name-bearing types of
<taxonomicName id="4C37E6D7BD38FF9F63F0FF0EFF044EC2" authorityName="Fabricius" authorityYear="1793" box="[151,246,187,213]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Odonata" pageId="4" pageNumber="352" phylum="Tracheophyta" rank="order">Odonata</taxonomicName>
in the University of Michigan Museum in which they included the
<typeStatus id="548C23F6BD38FF9F6087FF0EFBB04EC2" box="[992,1090,187,213]" pageId="4" pageNumber="352" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C37E6D7BD38FF9F6701FF0EFB394EC2" authorityName="Kennedy" authorityYear="1936" box="[1126,1227,187,213]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="352" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B9434146BD38FF9F6701FF0EFB394EC2" box="[1126,1227,187,213]" italics="true" pageId="4" pageNumber="352">P. fraseri</emphasis>
</taxonomicName>
. We could retrieve photos of
<typeStatus id="548C23F6BD38FF9F626BFF6AFE9C4EEE" box="[268,366,223,249]" pageId="4" pageNumber="352" type="holotype">holotype</typeStatus>
male from the University of Michigan-Museum of Zoology, which are reproduced here with permission.
</paragraph>
</subSubSection>
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<paragraph id="8B889D54BD3FFF9863FFF822FEE949E0" blockId="3.[151,1436,1943,2039]" pageId="3" pageNumber="351">
<emphasis id="B9434146BD3FFF9863FFF822FEEB49B8" bold="true" box="[152,281,1943,1967]" pageId="3" pageNumber="351">FIGURE 2.</emphasis>
<taxonomicName id="4C37E6D7BD3FFF986247F82DFE2949B8" authorityName="Kennedy" authorityYear="1936" box="[288,475,1943,1967]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="351" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B9434146BD3FFF986247F82DFE2949B8" box="[288,475,1943,1967]" italics="true" pageId="3" pageNumber="351">Protosticta fraseri</emphasis>
</taxonomicName>
holotype male (UMMZI-199248): (a) lateral habitus; (bc) thorax: (b) dorsal view; (c) lateral view; (de) caudal appendages: (d) dorsal view; (e) lateral view. Photo Courtesy: K. Eldredge / Museum of Zoology, University of Michigan.
</paragraph>
</caption>
<subSubSection id="C32DCEDFBD38FF9F63F0FEDBFD8D4CE6" pageId="4" pageNumber="352" type="diagnosis">
<paragraph id="8B889D54BD38FF9F63F0FEDBFEE24F9F" blockId="4.[151,272,366,392]" box="[151,272,366,392]" pageId="4" pageNumber="352">
<heading id="D0C02A38BD38FF9F63F0FEDBFEE24F9F" bold="true" box="[151,272,366,392]" fontSize="11" level="1" pageId="4" pageNumber="352" reason="1">
<emphasis id="B9434146BD38FF9F63F0FEDBFEE24F9F" bold="true" box="[151,272,366,392]" pageId="4" pageNumber="352">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B889D54BD38FF9F63F0FE02FC744C76" blockId="4.[151,1436,439,753]" pageId="4" pageNumber="352">
From other
<taxonomicName id="4C37E6D7BD38FF9F627BFE02FE3C4FC6" box="[284,462,439,465]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="352" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9434146BD38FF9F627BFE02FE654FC7" box="[284,407,439,464]" italics="true" pageId="4" pageNumber="352">Protosticta</emphasis>
spp.
</taxonomicName>
in Eastern Himalaya and NE
<collectingCountry id="F320DDC4BD38FF9F6078FE02FCAB4FC6" box="[799,857,439,465]" name="India" pageId="4" pageNumber="352">India</collectingCountry>
<taxonomicName id="4C37E6D7BD38FF9F6007FE02FC354FC6" authorityName="Kennedy" authorityYear="1936" box="[864,967,439,465]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="352" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B9434146BD38FF9F6007FE02FC354FC6" box="[864,967,439,465]" italics="true" pageId="4" pageNumber="352">P. fraseri</emphasis>
</taxonomicName>
can be distinguished by the form of male caudal appendages, especially the shape of paraprocts which are thin and curved, ending in a sharply pointed apex (
<figureCitation id="130C81D1BD38FF9F63F8FE4AFEE34C0E" box="[159,273,511,537]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-20@3.[215,1373,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Protosticta fraseri holotype male (UMMZI-199248): (a) lateral habitus; (bc) thorax: (b) dorsal view; (c) lateral view; (de) caudal appendages: (d) dorsal view; (e) lateral view. Photo Courtesy: K. Eldredge / Museum of Zoology, University of Michigan." figureDoi="http://doi.org/10.5281/zenodo.11231711" httpUri="https://zenodo.org/record/11231711/files/figure.png" pageId="4" pageNumber="352">Fig. 2de</figureCitation>
). Males of all other species in NE
<collectingCountry id="F320DDC4BD38FF9F61C3FE4AFD2C4C0E" box="[676,734,511,537]" name="India" pageId="4" pageNumber="352">India</collectingCountry>
have complex, lobed paraproct. The female and male have a characteristic dark thorax with a vaguely defined pale stripe (unique among known congeners in NE
<collectingCountry id="F320DDC4BD38FF9F6646FD96FAAD4C2A" box="[1313,1375,547,573]" name="India" pageId="4" pageNumber="352">India</collectingCountry>
), and S37 with medio-lateral white stripe and basal white rings (
<figureCitation id="130C81D1BD38FF9F6053FDF2FC8B4C76" box="[820,889,583,609]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-20@3.[215,1373,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Protosticta fraseri holotype male (UMMZI-199248): (a) lateral habitus; (bc) thorax: (b) dorsal view; (c) lateral view; (de) caudal appendages: (d) dorsal view; (e) lateral view. Photo Courtesy: K. Eldredge / Museum of Zoology, University of Michigan." figureDoi="http://doi.org/10.5281/zenodo.11231711" httpUri="https://zenodo.org/record/11231711/files/figure.png" pageId="4" pageNumber="352">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8B889D54BD38FF9F63A0FDDEFD8D4CE6" blockId="4.[151,1436,439,753]" pageId="4" pageNumber="352">
<taxonomicName id="4C37E6D7BD38FF9F63A0FDDEFE654C92" authorityName="Kennedy" authorityYear="1936" box="[199,407,619,645]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="352" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B9434146BD38FF9F63A0FDDEFE654C92" box="[199,407,619,645]" italics="true" pageId="4" pageNumber="352">Protosticta fraseri</emphasis>
</taxonomicName>
is comparable to
<taxonomicName id="4C37E6D7BD38FF9F6102FDDEFCCF4C92" authority="Asahina" authorityName="Asahina" authorityYear="1985" box="[613,829,619,645]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="352" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B9434146BD38FF9F6102FDDEFD2A4C92" box="[613,728,619,645]" italics="true" pageId="4" pageNumber="352">P. grandis</emphasis>
Asahina
</taxonomicName>
in overall morphology, size as well as the shape and color of caudal appendages.
<taxonomicName id="4C37E6D7BD38FF9F62B2FD3AFD5F4CBE" authorityName="Asahina" authorityYear="1985" box="[469,685,655,681]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="352" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B9434146BD38FF9F62B2FD3AFD5F4CBE" box="[469,685,655,681]" italics="true" pageId="4" pageNumber="352">Protosticta grandis</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="4C37E6D7BD38FF9F60BCFD3AFBB34CBE" authorityName="Kennedy" authorityYear="1936" box="[987,1089,655,681]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="352" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B9434146BD38FF9F60BCFD3AFBB34CBE" box="[987,1089,655,681]" italics="true" pageId="4" pageNumber="352">P. fraseri</emphasis>
</taxonomicName>
by a prominent lateral thoracic stripe (no lateral thoracic stripe in
<taxonomicName id="4C37E6D7BD38FF9F6171FD06FD894CDA" authorityName="Kennedy" authorityYear="1936" box="[534,635,691,717]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="352" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B9434146BD38FF9F6171FD06FD894CDA" box="[534,635,691,717]" italics="true" pageId="4" pageNumber="352">P. fraseri</emphasis>
</taxonomicName>
) (
<figureCitation id="130C81D1BD38FF9F61F4FD06FD164CDA" box="[659,740,691,717]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-20@3.[215,1373,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Protosticta fraseri holotype male (UMMZI-199248): (a) lateral habitus; (bc) thorax: (b) dorsal view; (c) lateral view; (de) caudal appendages: (d) dorsal view; (e) lateral view. Photo Courtesy: K. Eldredge / Museum of Zoology, University of Michigan." figureDoi="http://doi.org/10.5281/zenodo.11231711" httpUri="https://zenodo.org/record/11231711/files/figure.png" pageId="4" pageNumber="352">Fig. 2c</figureCitation>
), inner border of cerci more straight (sinuous in
<taxonomicName id="4C37E6D7BD38FF9F6799FD06FA964CDA" authorityName="Kennedy" authorityYear="1936" box="[1278,1380,691,717]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="352" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B9434146BD38FF9F6799FD06FA964CDA" box="[1278,1380,691,717]" italics="true" pageId="4" pageNumber="352">P. fraseri</emphasis>
</taxonomicName>
) and bulkier cerci apex (
<bibRefCitation id="EFA6E0A5BD38FF9F6209FD62FD814CE6" author="Phan, Q. T. &amp; Kompier, T." box="[366,627,727,753]" pageId="4" pageNumber="352" pagination="529 - 44" refId="ref9456" refString="Phan, Q. T. &amp; Kompier, T. (2016) A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae). Zootaxa, 4098 (3), 529 - 44. https: // doi. org / 10.11646 / zootaxa. 4098.3.6" type="journal article" year="2016">Phan &amp; Kompier 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>