271 lines
34 KiB
XML
271 lines
34 KiB
XML
<document id="21B86D6B030AAAEA88050368D86A8BC3" ID-CLB-Dataset="298355" ID-DOI="10.5194/fr-24-19-2021" ID-GBIF-Dataset="cc5ca7c4-8577-4583-b28a-d9b10e9c5c36" ID-ISSN="2193-0074" ID-Zenodo-Dep="10965766" IM.bibliography_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1712928138748" checkinUser="felipe" docAuthor="Alekseev, Vitalii I., Mitchell, Jerit, McKellar, Ryan C., Barbi, Mauricio, Larsson, Hans C. E. & Bukejs, Andris" docDate="2021" docId="03901871FFD9FFCD65AEFF648EB2BC24" docLanguage="en" docName="FossRec.24.1.19-32.pdf" docOrigin="Fossil Record 24 (1)" docSource="http://dx.doi.org/10.5194/fr-24-19-2021" docStyle="DocumentStyle:1B3764FFAC190C6D25C29F616DE4358D.2:FossRec.2014-2021.journal_article" docStyleId="1B3764FFAC190C6D25C29F616DE4358D" docStyleName="FossRec.2014-2021.journal_article" docStyleVersion="2" docTitle="Chelonarium andabata Alekseev and Bukejs 2021, sp. nov." docType="treatment" docUuid="27596D45-C0C2-430B-A0AE-35DCF5FB116C" docUuidSource="ZooBank" docVersion="6" lastPageNumber="22" masterDocId="FFA96009FFDBFFC9651FFFD48A14BB23" masterDocTitle="The first described turtle beetles from Eocene Baltic amber, with notes on fossil Chelonariidae (Coleoptera: Byrrhoidea)" masterLastPageNumber="32" masterPageNumber="19" pageNumber="21" updateTime="1718077761551" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="18AF19064B38DB4F0F8DCFF2DB05C4AC">
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<mods:title id="2EDC895BBD08A500844981B6014B0081">The first described turtle beetles from Eocene Baltic amber, with notes on fossil Chelonariidae (Coleoptera: Byrrhoidea)</mods:title>
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<mods:name id="636FB6A37C32E74E17DC2910F7EEDB5F" type="personal">
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<mods:namePart id="3025A38B632F6B89C2B4266D4165B14D">Alekseev, Vitalii I.</mods:namePart>
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<mods:namePart id="CEA97170AA2CCB33D56140405A36157B">Mitchell, Jerit</mods:namePart>
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<mods:namePart id="3654D870D85132560513AA24ECFA4FE1">McKellar, Ryan C.</mods:namePart>
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<mods:roleTerm id="6DF564BF2FD87B9373497C6C53B9E3D0">Author</mods:roleTerm>
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<mods:namePart id="A44C99A2FFC7644223B338AEE5FD8CBB">Barbi, Mauricio</mods:namePart>
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<mods:namePart id="CE52628039C49986C7D6DC69F8ED9C9B">Larsson, Hans C. E.</mods:namePart>
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<mods:namePart id="A324C2A362B58D3D7138F931AE2C4C53">Bukejs, Andris</mods:namePart>
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<mods:title id="E3E08683D779F7561647728F8124F9D8">Fossil Record</mods:title>
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<mods:date id="F9583ACE73068D22FDF6FEE92D25EDCE">2021</mods:date>
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<mods:number id="1664F728B38CF3DD9A9CED118F94E633">2021-02-10</mods:number>
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<mods:classification id="5ABBCE899B60F4EDE8E4B53F6E091246">journal article</mods:classification>
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<treatment id="03901871FFD9FFCD65AEFF648EB2BC24" ID-DOI="http://doi.org/10.5281/zenodo.11558749" ID-GBIF-Taxon="232728041" ID-Zenodo-Dep="11558749" LSID="urn:lsid:zoobank.org:act:27596D45-C0C2-430B-A0AE-35DCF5FB116C" httpUri="http://treatment.plazi.org/id/03901871FFD9FFCD65AEFF648EB2BC24" lastPageId="4" lastPageNumber="22" pageId="2" pageNumber="21">
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<subSubSection id="C323FAECFFD9FFCB65AEFF648BA7BA31" pageId="2" pageNumber="21" type="nomenclature">
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<paragraph id="8B86A967FFD9FFCB65AEFF6488F3BBE4" blockId="2.[177,743,176,317]" box="[177,743,176,200]" pageId="2" pageNumber="21">
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<heading id="D0CE1E0BFFD9FFCB65AEFF6488F3BBE4" box="[177,743,176,200]" fontSize="10" level="2" pageId="2" pageNumber="21" reason="5">
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<taxonomicName id="4C39D2E4FFD9FFCB65AEFF64889BBBE4" authority="Alekseev and Bukejs" authorityName="Alekseev and Bukejs" authorityYear="2021" box="[177,655,176,200]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="andabata" status="sp. nov.">
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<emphasis id="B94D7575FFD9FFCB65AEFF648BBCBBEB" box="[177,424,176,200]" italics="true" pageId="2" pageNumber="21">Chelonarium andabata</emphasis>
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Alekseev and Bukejs
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</taxonomicName>
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<taxonomicNameLabel id="A27EC80EFFD9FFCB678AFF6488F3BBE4" box="[661,743,176,199]" pageId="2" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8B86A967FFD9FFCB65AEFF0F8BA7BA31" blockId="2.[177,743,176,317]" pageId="2" pageNumber="21">urn:lsid:zoobank.org:act:27596D45-C0C2-430BA0AE-35DCF5FB116C</paragraph>
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</subSubSection>
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<subSubSection id="C323FAECFFD9FFCB65AEFEF28B2FBA1E" box="[177,315,294,317]" pageId="2" pageNumber="21" type="description">
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<paragraph id="8B86A967FFD9FFCB65AEFEF28B2FBA1E" blockId="2.[177,743,176,317]" box="[177,315,294,317]" pageId="2" pageNumber="21">
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<figureCitation id="1302B5E2FFD9FFCB65AEFEF28B02BA1E" box="[177,278,294,317]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStartId-0="3.[124,191,1497,1519]" captionStartId-1="4.[124,191,1499,1521]" captionStartId-2="5.[124,191,1451,1473]" captionStartId-3="6.[124,191,787,809]" captionStartId-4="6.[124,191,1767,1789]" captionTargetBox-0="[226,1360,177,1466]" captionTargetBox-1="[283,1303,177,1468]" captionTargetBox-2="[283,1303,177,1420]" captionTargetBox-3="[339,1203,177,756]" captionTargetBox-4="[226,1360,885,1736]" captionTargetId-0="figure-15@3.[226,1360,177,1466]" captionTargetId-1="figure-15@4.[283,1303,177,1468]" captionTargetId-2="figure-15@5.[283,1303,177,1420]" captionTargetId-3="figure-15@6.[339,1246,177,756]" captionTargetId-4="figure-43@6.[226,1360,885,1736]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionText-0="Figure 1. Photomicrographs of Chelonarium andabata Alekseev and Bukejs sp. nov., holotype, RSKM_P3000.141 [RSKM]: (a) ventral habitus view; (b) right lateral habitus view; (c) detail of head with antenna (horizontal arrow) and protarsi (inclined arrows) in ventral view; and (d) detail of legs in ventral view with pro-, meso-, and metatarsus indicated by horizontal arrows (from top to bottom of view). Scale bars = 1.0 mm." captionText-1="Figure 2. X-ray micro-CT renderings of Chelonarium andabata Alekseev and Bukejs sp. nov., holotype, RSKM_P3000.141 [RSKM], habitus: (a) dorsal view, (b) ventral view and (c) left lateral view. Scale bar = 1.0 mm." captionText-2="Figure 3. X-ray micro-CT renderings of Chelonarium andabata Alekseev and Bukejs sp. nov., holotype, RSKM_P3000.141 [RSKM], habitus: (a) ventral view without legs, showing depressions for legs reception; (b) ventral view with antennae and legs in different colours; (c) frontal view; and (d) caudal view. Scale bar = 1.0 mm." captionText-3="Figure 4. Antennae of Chelonarium andabata Alekseev and Bukejs sp. nov., holotype, RSKM_P3000.141 [RSKM]: (a) X-ray micro-CT rendering and (b) reconstruction. Abbreviations: a1–a11 – antennomeres 1–11 respectively." captionText-4="Figure 5. X-ray micro-CT renderings of Chelonarium andabata Alekseev and Bukejs sp. nov., holotype, RSKM_P3000.141 [RSKM], aedeagus: (a) dorsal view, (b) lateral view and (c) ventral view. Scale bar = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10965768" figureDoi-1="http://doi.org/10.5281/zenodo.10965770" figureDoi-2="http://doi.org/10.5281/zenodo.10965772" figureDoi-3="http://doi.org/10.5281/zenodo.10965774" figureDoi-4="http://doi.org/10.5281/zenodo.10965776" httpUri-0="https://zenodo.org/record/10965768/files/figure.png" httpUri-1="https://zenodo.org/record/10965770/files/figure.png" httpUri-2="https://zenodo.org/record/10965772/files/figure.png" httpUri-3="https://zenodo.org/record/10965774/files/figure.png" httpUri-4="https://zenodo.org/record/10965776/files/figure.png" pageId="2" pageNumber="21">Figs. 1–5</figureCitation>
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,
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<figureCitation id="1302B5E2FFD9FFCB643EFEF28B2FBA1E" box="[289,315,294,317]" captionStart="Figure 8" captionStartId="9.[124,191,1243,1265]" captionTargetBox="[302,1285,177,1212]" captionTargetId="figure-15@9.[302,1285,177,1212]" captionTargetPageId="9" captionText="Figure 8. Elytral vestiture and setation shape in fossil Chelonarium: (a) Ch. dingansich Alekseev and Bukejs sp. nov., right elytron, and (b) Ch. andabata Alekseev and Bukejs sp. nov., left elytron. Scale bars = 0.5 mm. Abbreviation: pa – patches of paler and denser setae." figureDoi="http://doi.org/10.5281/zenodo.10965784" httpUri="https://zenodo.org/record/10965784/files/figure.png" pageId="2" pageNumber="21">8b</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C323FAECFFD9FFCB659CFEA88B6DB96D" pageId="2" pageNumber="21" type="etymology">
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<paragraph id="8B86A967FFD9FFCB659CFEA88B5CBAB7" blockId="2.[131,328,380,404]" box="[131,328,380,404]" pageId="2" pageNumber="21">
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<emphasis id="B94D7575FFD9FFCB659CFEA88B5CBAB7" bold="true" box="[131,328,380,404]" italics="true" pageId="2" pageNumber="21">Derivatio nominis</emphasis>
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</paragraph>
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<paragraph id="8B86A967FFD9FFCB6563FE6C8B6DB96D" blockId="2.[124,767,439,590]" pageId="2" pageNumber="21">
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The specific epithet
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<taxonomicName id="4C39D2E4FFD9FFCB6449FE6C8BD1BAEC" authorityName="Alekseev and Bukejs" authorityYear="2021" box="[342,453,439,463]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="andabata">
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“
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<emphasis id="B94D7575FFD9FFCB647EFE638BD1BAEC" box="[353,453,439,463]" italics="true" pageId="2" pageNumber="21">andabata</emphasis>
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</taxonomicName>
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” is a Latin word meaning “a
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<typeStatus id="548217C5FFD9FFCB6563FE038ABEBACD" box="[124,170,471,494]" pageId="2" pageNumber="21">type</typeStatus>
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of gladiator who fought blind because of a restrictive helmet” and refers to the strongly declined head of the beetle, which is concealed from above. The specific epithet is used as a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C323FAECFFD9FFCB6563FDA5884FBE0E" pageId="2" pageNumber="21" type="materials_examined">
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<paragraph id="8B86A967FFD9FFCB6563FDA58B0EB9AA" blockId="2.[124,282,625,649]" box="[124,282,625,649]" pageId="2" pageNumber="21">
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<emphasis id="B94D7575FFD9FFCB6563FDA58B0EB9AA" bold="true" box="[124,282,625,649]" pageId="2" pageNumber="21">Type material</emphasis>
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</paragraph>
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<paragraph id="8B86A967FFD9FFCB6563FD798B9EB8E0" blockId="2.[124,767,685,963]" pageId="2" pageNumber="21">
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<materialsCitation id="3B51A33AFFD9FFCB6563FD798B9EB8E0" collectionCode="RSKM" pageId="2" pageNumber="21" specimenCode="RSKM_P3000.141" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="548217C5FFD9FFCB6563FD798AF7B9E7" box="[124,227,685,708]" pageId="2" pageNumber="21" type="holotype">Holotype</typeStatus>
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: collection number “
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<specimenCode id="DB9F011CFFD9FFCB64D5FD798888B9E7" box="[458,668,685,708]" pageId="2" pageNumber="21">RSKM_P3000.141</specimenCode>
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”, “Holotype/
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<taxonomicName id="4C39D2E4FFD9FFCB65AEFD188BB9B9C7" authority="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs, 2021" authorityName="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs" authorityYear="2021" box="[177,429,716,740]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="andabata" status="sp. nov.">
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<emphasis id="B94D7575FFD9FFCB65AEFD188BB9B9C7" box="[177,429,716,740]" italics="true" pageId="2" pageNumber="21">Chelonarium andabata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27EC80EFFD9FFCB64A7FD188819B9C0" box="[440,525,716,739]" pageId="2" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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/Alekseev and Bukejs des. 2021” [red printed label] [RSKM]; adult,
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<specimenCount id="9D3F62EEFFD9FFCB678BFD3888DFB820" box="[660,715,748,771]" pageId="2" pageNumber="21" type="male">male</specimenCount>
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. Almost complete beetle (right antennomeres 10–11 lacking) included in a transparent, yellow amber piece with approximate dimensions of
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<quantity id="4CC10482FFD9FFCB6403FC988B43B840" box="[284,343,844,867]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="21" unit="mm" value="8.0">8 mm</quantity>
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×
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<quantity id="4CC10482FFD9FFCB646AFC988BA9B840" box="[373,445,844,867]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="21" unit="mm" value="16.0">16 mm</quantity>
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and a maximum thickness of
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<quantity id="4CC10482FFD9FFCB6563FCB88AAFB8A0" box="[124,187,876,899]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="21" unit="mm" value="8.0">8 mm</quantity>
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; preserved without supplementary fixation. The dorsal part of the specimen is partially obscured by milky amber. Syninclusions are absent.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B86A967FFD9FFCB6563FC318B00B8DE" blockId="2.[124,276,997,1021]" box="[124,276,997,1021]" pageId="2" pageNumber="21">
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<emphasis id="B94D7575FFD9FFCB6563FC318B00B8DE" bold="true" box="[124,276,997,1021]" pageId="2" pageNumber="21">Type stratum</emphasis>
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</paragraph>
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<paragraph id="8B86A967FFD9FFCB6563FBF58AFCBFBB" blockId="2.[124,766,1057,1176]" pageId="2" pageNumber="21">
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Baltic amber from Eocene amber-bearing Blue Earth layers; a predominantly Bartonian age has been interpreted for the extinct central European resin-producing forests (
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<bibRefCitation id="EFA8D496FFD9FFCB6788FBB58AC9BFBB" author="Bukejs, A. & Alekseev, V. I. & Pollock, D. A." pageId="2" pageNumber="21" pagination="261 - 273" refId="ref7344" refString="Bukejs, A., Alekseev, V. I., and Pollock, D. A.: Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location, Zootaxa, 4664, 261 - 273, https: // doi. org / 10.11646 / zootaxa. 4664.2.8, 2019." type="journal article" year="2019">Bukejs et al., 2019</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8B86A967FFD9FFCB6563FB6F8B19BFF0" blockId="2.[124,269,1211,1235]" box="[124,269,1211,1235]" pageId="2" pageNumber="21">
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<emphasis id="B94D7575FFD9FFCB6563FB6F8B19BFF0" bold="true" box="[124,269,1211,1235]" pageId="2" pageNumber="21">Type locality</emphasis>
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</paragraph>
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<paragraph id="8B86A967FFD9FFCB6563FB22884FBE0E" blockId="2.[124,767,1270,1325]" pageId="2" pageNumber="21">
|
||
Yantarny settlement (formerly Palmnicken), Sambian (Samland) Peninsula,
|
||
<collectingRegion id="49FD6785FFD9FFCB6431FAC28816BE0E" box="[302,514,1302,1325]" country="Russia" name="Kaliningrad" pageId="2" pageNumber="21">Kaliningrad Region</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32EE9F7FFD9FFCB6712FAC28843BE0E" box="[525,599,1302,1325]" name="Russia" pageId="2" pageNumber="21">Russia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C323FAECFFD9FFCA6563FA848B83BDDF" lastPageId="3" lastPageNumber="22" pageId="2" pageNumber="21" type="description">
|
||
<paragraph id="8B86A967FFD9FFCB6563FA848AEBBE4B" blockId="2.[124,255,1360,1384]" box="[124,255,1360,1384]" pageId="2" pageNumber="21">
|
||
<emphasis id="B94D7575FFD9FFCB6563FA848AEBBE4B" bold="true" box="[124,255,1360,1384]" pageId="2" pageNumber="21">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B86A967FFD9FFCB6563FA588B4DBD21" blockId="2.[124,767,1420,1921]" pageId="2" pageNumber="21">
|
||
Measurements: body length
|
||
<quantity id="4CC10482FFD9FFCB64AEFA588816BE80" box="[433,514,1420,1443]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="2" pageNumber="21" unit="mm" value="5.6">5.6 mm</quantity>
|
||
, maximum body width
|
||
<quantity id="4CC10482FFD9FFCB6563FA788ADBBEE0" box="[124,207,1452,1475]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="21" unit="mm" value="3.2">3.2 mm</quantity>
|
||
; pronotum length
|
||
<quantity id="4CC10482FFD9FFCB64ADFA788810BEE0" box="[434,516,1452,1475]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="21" unit="mm" value="1.2">1.2 mm</quantity>
|
||
, maximum pronotum width
|
||
<quantity id="4CC10482FFD9FFCB65DBFA1F8B03BEC1" box="[196,279,1483,1506]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="21" unit="mm" value="2.5">2.5 mm</quantity>
|
||
; elytra length
|
||
<quantity id="4CC10482FFD9FFCB64A2FA1F881ABEC1" box="[445,526,1483,1506]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="21" unit="mm" value="4.3">4.3 mm</quantity>
|
||
, maximum combined elytra width
|
||
<quantity id="4CC10482FFD9FFCB641DFA3F8B41BD21" box="[258,341,1515,1538]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="21" unit="mm" value="3.2">3.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B86A967FFD9FFCB6588F9DF8885BC22" blockId="2.[124,767,1420,1921]" pageId="2" pageNumber="21">
|
||
Body ovoid, widely oval, biconvex; total body length
|
||
<emphasis id="B94D7575FFD9FFCB65DCF9FF8ADBBD60" bold="true" box="[195,207,1579,1603]" italics="true" pageId="2" pageNumber="21">/</emphasis>
|
||
maximum body width equal to 1.75; integument unicolorous dark brown (as preserved). Pubescence unicolorous and dark: dorsum covered with short, semierect setae in moderate density; elytral vestiture uniform, simple, without any trichome-like structures, setae almost straight to slightly curved; setae at elytral lateral sides slightly stouter; venter and legs with dense, short recumbent setae.
|
||
</paragraph>
|
||
<paragraph id="8B86A967FFD9FFCB6588F8DE8F16BAE5" blockId="2.[124,767,1420,1921]" lastBlockId="2.[819,1463,176,1921]" pageId="2" pageNumber="21">Head declined, not visible from above, rounded pentagonal, slightly convex; densely punctate; punctures round and small, distance between punctures equal to 0.2–0.5× diameter of one puncture. Clypeus not distinct, frontoclypeal suture absent. Eyes small, entire, almost round, slightly convex, finely facetted, without interfacetal setation; interocular frontal distance equal to about 2× diameter of one eye. Antennal insertions hidden. Antennae rather long; scape hidden, rounded and short; pedicel and antennomeres 3–11 flattened; antennomere 3 longest, antennomere 4 shortest; antennomeres 2–3 and 11 elongate, antennomeres 5–10 triangular in outline. Relative length ratios of antennomeres 1–11 equal to 1.5: 3: 7: 0.7: 1.8: 2: 2: 2.2: 2.2: 2: 3.4.</paragraph>
|
||
<paragraph id="8B86A967FFD9FFCB6651FE1B8F78B846" blockId="2.[819,1463,176,1921]" pageId="2" pageNumber="21">Pronotum transverse, 2.1× as wide as long, widest at base, with fine and dense punctation. Posterior angles of pronotum acute. Anterior edge of pronotum almost semicircular in dorsal view; posterior edge bisinuate and crenulate. Pronotum margined anteriorly and laterally with sharp, raised carina dividing pronotum into upper (moderately convex dorsally) and lower (inclined posteriad) parts. Lower pronotal area (between prominent anterior carina and posterior edge of concealed head) wider than protibial width. Pro- and mesosternum with deep, elongate, intercoxal median excavation for reception of basal antennomeres. Hypomeron excavated to receive profemora; meso- and metaventrite with excavations for receiving meso- and metafemora as well as tibiae.</paragraph>
|
||
<paragraph id="8B86A967FFD9FFCB6651FCBA89C8BE00" blockId="2.[819,1463,176,1921]" pageId="2" pageNumber="21">Scutellum subpentagonal, almost as long as wide. Elytra moderately convex, widely oval, about 1.4× as long as combined width; distinctly wider than pronotal posterior margin, about 3.6× as long as pronotum. Elytral punctation irregular, dense and round along sides, sparser and finer on disc, distance between punctures equal to 0.3–3.0× diameter of one puncture, interspaces flat. Epipleura narrow, reaching abdominal ventrite 1. Metepisternum wide, about 2× as wide as maximum epipleural width, with dense, small punctation. Metaventrite slightly convex laterally, with almost flat disc; densely covered with small punctation; distance between punctures smaller than diameter of one puncture; interspaces slightly convex; discrimen distinct in anterior half of metaventrite.</paragraph>
|
||
<paragraph id="8B86A967FFD9FFCB6651FAF88E8CBC22" blockId="2.[819,1463,176,1921]" pageId="2" pageNumber="21">Legs rather short, flattened; densely covered with small punctures; distance between punctures smaller than diameter of one puncture. All coxae distinctly separated; proand mesocoxae widely oval, slightly transverse, about 1.3– 1.4× as wide as long; metacoxae narrow, strongly transverse, excavate, with short metacoxal plates and with triangular outer edge. Femora and tibiae subequal in length; proand mesofemur comparatively wider than metafemur. Tibiae narrower than femora, protibia about 0.5× as wide as profemur, mesotibia about 0.5× as wide mesofemur, metatibia about 0.6× as wide as metafemur; protibia spinose along inner margin. Tarsi 5 segmented, tarsomere 3 with long membranous lobe ventrally, tarsomere 4 minute. Relative length ratios of mesotarsomeres 1–5 equal to 15: 10: 7: 4: 25. Pretarsal claws with denticle basally.</paragraph>
|
||
<paragraph id="8B86A967FFD9FFCA6651F8DE88B0BD9F" blockId="2.[819,1463,176,1921]" lastBlockId="3.[124,766,1669,1788]" lastPageId="3" lastPageNumber="22" pageId="2" pageNumber="21">Abdomen with five visible ventrites; abdominal sutures entire, slightly concave; ventrite 5 simple, with widely rounded, semicircular apical margin; finely and densely punctate; distance between punctures equal to 0.5–1.0× diameter of one puncture. Relative length ratios of abdominal ventrites 1–5 equal to 7: 4.5: 4: 3.5: 6.5 (medially).</paragraph>
|
||
<caption id="DF46F9EFFFD8FFCA6563FA0D8B19BD65" ID-DOI="http://doi.org/10.5281/zenodo.10965768" ID-Zenodo-Dep="10965768" httpUri="https://zenodo.org/record/10965768/files/figure.png" pageId="3" pageNumber="22" startId="3.[124,191,1497,1519]" targetBox="[226,1360,177,1466]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8B86A967FFD8FFCA6563FA0D8B19BD65" blockId="3.[124,1462,1497,1606]" pageId="3" pageNumber="22">
|
||
<emphasis id="B94D7575FFD8FFCA6563FA0D8ACEBECC" bold="true" box="[124,218,1497,1519]" pageId="3" pageNumber="22">Figure 1.</emphasis>
|
||
Photomicrographs of
|
||
<taxonomicName id="4C39D2E4FFD8FFCA64AAFA0D897FBECC" authority="Alekseev and Bukejs" authorityName="Alekseev and Bukejs" authorityYear="2021" box="[437,875,1497,1519]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="andabata" status="sp. nov.">
|
||
<emphasis id="B94D7575FFD8FFCA64AAFA0D8881BECD" box="[437,661,1497,1518]" italics="true" pageId="3" pageNumber="22">Chelonarium andabata</emphasis>
|
||
Alekseev and Bukejs
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFD8FFCA666CFA0E89D6BECC" box="[883,962,1498,1519]" pageId="3" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, RSKM_P3000.141 [RSKM]:
|
||
<emphasis id="B94D7575FFD8FFCA604FFA0D8F7FBECC" bold="true" box="[1360,1387,1497,1519]" pageId="3" pageNumber="22">(a)</emphasis>
|
||
ventral habitus view;
|
||
<emphasis id="B94D7575FFD8FFCA641DFA228B0BBD2F" bold="true" box="[258,287,1526,1548]" pageId="3" pageNumber="22">(b)</emphasis>
|
||
right lateral habitus view;
|
||
<emphasis id="B94D7575FFD8FFCA673FFA22882EBD2F" bold="true" box="[544,570,1526,1548]" pageId="3" pageNumber="22">(c)</emphasis>
|
||
detail of head with antenna (horizontal arrow) and protarsi (inclined arrows) in ventral view; and
|
||
<emphasis id="B94D7575FFD8FFCA65B9F9C78AD7BD0A" bold="true" box="[166,195,1555,1577]" pageId="3" pageNumber="22">(d)</emphasis>
|
||
detail of legs in ventral view with pro-, meso-, and metatarsus indicated by horizontal arrows (from top to bottom of view). Scale bars = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B86A967FFD8FFCA6588F9118B83BDDF" blockId="3.[124,766,1669,1788]" pageId="3" pageNumber="22">
|
||
Aedeagus robust, with widely rounded base and gradually narrowed apically (
|
||
<figureCitation id="1302B5E2FFD8FFCA6456F9318B9EBDDF" box="[329,394,1765,1788]" captionStart="Figure 5" captionStartId="6.[124,191,1767,1789]" captionTargetBox="[226,1360,885,1736]" captionTargetId="figure-43@6.[226,1360,885,1736]" captionTargetPageId="6" captionText="Figure 5. X-ray micro-CT renderings of Chelonarium andabata Alekseev and Bukejs sp. nov., holotype, RSKM_P3000.141 [RSKM], aedeagus: (a) dorsal view, (b) lateral view and (c) ventral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10965776" httpUri="https://zenodo.org/record/10965776/files/figure.png" pageId="3" pageNumber="22">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C323FAECFFD8FFCD662CF9508EB2BC24" lastPageId="4" lastPageNumber="23" pageId="3" pageNumber="22" type="diagnosis">
|
||
<paragraph id="8B86A967FFD8FFCA662CF9508E32BDBF" blockId="3.[819,1062,1668,1692]" box="[819,1062,1668,1692]" pageId="3" pageNumber="22">
|
||
<emphasis id="B94D7575FFD8FFCA662CF9508E32BDBF" bold="true" box="[819,1062,1668,1692]" pageId="3" pageNumber="22">Differential diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B86A967FFD8FFCD662CF91E8EB2BC24" blockId="3.[819,1462,1738,1921]" lastBlockId="4.[819,1462,1616,1799]" lastPageId="4" lastPageNumber="23" pageId="3" pageNumber="22">
|
||
The genus
|
||
<taxonomicName id="4C39D2E4FFD8FFCA66B9F91E8E26BDC1" authorityName="Fabricius" authorityYear="1801" box="[934,1074,1738,1762]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D7575FFD8FFCA66B9F91E8E26BDC1" box="[934,1074,1738,1762]" italics="true" pageId="3" pageNumber="22">Chelonarium</emphasis>
|
||
</taxonomicName>
|
||
is very species-rich but poorly documented: it is in need of revision for many of its inadequately illustrated and briefly described species. The Eocene taxon
|
||
<taxonomicName id="4C39D2E4FFD8FFCA662CF8FE89D0BC61" authority="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs, 2021" authorityName="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs" authorityYear="2021" box="[819,964,1834,1858]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="andabata" status="sp. nov.">
|
||
<emphasis id="B94D7575FFD8FFCA662CF8FE89D0BC61" box="[819,964,1834,1858]" italics="true" pageId="3" pageNumber="22">Ch. andabata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFD8FFCA66D4F8FE8E09BC62" box="[971,1053,1834,1857]" pageId="3" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is similar to one Recent North American chelonariid species,
|
||
<taxonomicName id="4C39D2E4FFD8FFCA6129F89E8F37BC42" authority="Thomson" authorityName="Thomson" authorityYear="1867" box="[1078,1315,1866,1890]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="lecontei">
|
||
<emphasis id="B94D7575FFD8FFCA6129F89E8EA2BC41" box="[1078,1206,1866,1890]" italics="true" pageId="3" pageNumber="22">Ch. lecontei</emphasis>
|
||
Thomson
|
||
</taxonomicName>
|
||
, in its general antennal structure, but it differs in having smaller eyes (interocular distance in
|
||
<taxonomicName id="4C39D2E4FFDFFFCD6454F9848BD9BD4B" authorityName="Thomson" authorityYear="1867" box="[331,461,1616,1640]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="lecontei">
|
||
<emphasis id="B94D7575FFDFFFCD6454F9848BD9BD4B" box="[331,461,1616,1640]" italics="true" pageId="4" pageNumber="23">Ch. lecontei</emphasis>
|
||
</taxonomicName>
|
||
is slightly less than one eye diameter), denser head punctation and uniform elytral vestiture (without maculae). The newly described fossil species can be distinguished from the westernmost Palaearctic
|
||
<taxonomicName id="4C39D2E4FFDFFFCD67C6F97B8B27BDC4" authority="Mandl" authorityName="Mandl" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="vartianae">
|
||
<emphasis id="B94D7575FFDFFFCD67C6F97B8AF6BDC4" italics="true" pageId="4" pageNumber="23">Ch. vartianae</emphasis>
|
||
Mandl
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F32EE9F7FFDFFFCD6459F9048BDFBDC4" box="[326,459,1744,1767]" name="Afghanistan" pageId="4" pageNumber="23">Afghanistan</collectingCountry>
|
||
) based on having a pronotum without transverse impression.
|
||
<taxonomicName id="4C39D2E4FFDFFFCD671BF93B88EABC24" authority="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs, 2021" authorityName="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs" authorityYear="2021" box="[516,766,1775,1799]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="andabata" status="sp. nov.">
|
||
<emphasis id="B94D7575FFDFFFCD671BF93B88EABC24" box="[516,766,1775,1799]" italics="true" pageId="4" pageNumber="23">Chelonarium andabata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFDFFFCD6563F8C48AC5BC04" box="[124,209,1808,1831]" pageId="4" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
clearly differs from
|
||
<taxonomicName id="4C39D2E4FFDFFFCD64A6F8DB8896BC04" authority="Klug" authorityName="Klug" box="[441,642,1807,1831]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="ornatum">
|
||
<emphasis id="B94D7575FFDFFFCD64A6F8DB8857BC04" box="[441,579,1807,1831]" italics="true" pageId="4" pageNumber="23">Ch. ornatum</emphasis>
|
||
Klug
|
||
</taxonomicName>
|
||
in its ratio of four basal antennomere lengths; it differs from
|
||
<taxonomicName id="4C39D2E4FFDFFFCD6786F8FB8AE7BC45" authority="Sato" authorityName="Sato" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="kurosawai">
|
||
<emphasis id="B94D7575FFDFFFCD6786F8FB8AAEBC44" italics="true" pageId="4" pageNumber="23">Ch. kurosawai</emphasis>
|
||
Satô
|
||
</taxonomicName>
|
||
in possessing unicolorous elytra; and it differs from
|
||
<taxonomicName id="4C39D2E4FFDFFFCD6671F9848E72BD44" authority="Latreille" authorityName="Latreille" box="[878,1126,1616,1640]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="beauvoisi">
|
||
<emphasis id="B94D7575FFDFFFCD6671F9848E15BD4B" box="[878,1025,1616,1640]" italics="true" pageId="4" pageNumber="23">Ch. beauvoisi</emphasis>
|
||
Latreille
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C39D2E4FFDFFFCD616EF9848F13BD4B" box="[1137,1287,1616,1640]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="convexum">
|
||
<emphasis id="B94D7575FFDFFFCD616EF9848F13BD4B" box="[1137,1287,1616,1640]" italics="true" pageId="4" pageNumber="23">Ch. convexum</emphasis>
|
||
</taxonomicName>
|
||
MØquignon and
|
||
<taxonomicName id="4C39D2E4FFDFFFCD662CF9A489A9BDAB" box="[819,957,1648,1672]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="cupreum">
|
||
<emphasis id="B94D7575FFDFFFCD662CF9A489A9BDAB" box="[819,957,1648,1672]" italics="true" pageId="4" pageNumber="23">Ch. cupreum</emphasis>
|
||
</taxonomicName>
|
||
MØquignon in its smaller body size. The combination of characteristics mentioned in the species description, additional characteristics of body part ratios, and details of vestiture and punctation should distinguish the new fossil species from all Recent congeners.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |