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<document id="1CE75C42CEEDD086F684F0107A71C9F3" ID-DOI="10.11646/zootaxa.5529.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="14022196" ID-ZooBank="598225C3-9B8F-4C0D-8E75-098F9943091D" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1730452285851" checkinUser="plazi" docAuthor="Hillert, Oliver, Král, David, Bagaturov, Mikhail &amp; Chuluunbaatar, Gantigmaa" docDate="2024" docId="004A87E8FFC9FFBFFF2AF992FD0DFEAA" docLanguage="en" docName="zootaxa.5529.1.6.pdf" docOrigin="Zootaxa 5529 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5529.1.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lethrus (Heteroplistodus) kucerai Hillert, Kral &amp; Bagaturov 2024, new species" docType="treatment" docVersion="3" lastPageNumber="126" masterDocId="FC73FF90FFC8FFBBFFBDFF9DF933FFB7" masterDocTitle="Lethrus (Heteroplistodus) kucerai (Coleoptera: Geotrupidae: Lethrinae), a new species from China" masterLastPageNumber="143" masterPageNumber="123" pageNumber="124" updateTime="1730831960486" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="B573075AB3066E20F9FB1DE2B2D9A7AA">Lethrus (Heteroplistodus) kucerai (Coleoptera: Geotrupidae: Lethrinae), a new species from China</mods:title>
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<mods:namePart id="6E01CE960590C4DD01615E155C49BAD5">Hillert, Oliver</mods:namePart>
<mods:affiliation id="9457B885AB89D57F1FC06AB4DD8DAFAD">Kieferndamm 10, D- 15566 Schöneiche bei Berlin, Germany.</mods:affiliation>
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<mods:namePart id="979DE303C5CABAE9BA9C33F34D35D2C4">Král, David</mods:namePart>
<mods:affiliation id="3A1C7185E5BD7ABABBD870A14DF9F7B0">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00 Praha 2, Czechia.</mods:affiliation>
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<mods:namePart id="53E14AF8A28137DB6FFFD6E0A2D888F7">Bagaturov, Mikhail</mods:namePart>
<mods:affiliation id="D4020BE111CD9F0A0E3B4315158293A5">Zoological Institute of Russian Academy of Sciences, St. Petersburg, Universitetskaya naberezhnaja 1, Russia; Athens Institute for Education and Research, 7 Eptanisou Street, Vrilissia, Athens, Greece.</mods:affiliation>
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<mods:namePart id="17A1CB1F5C5C3F907321248E1AB884E9">Chuluunbaatar, Gantigmaa</mods:namePart>
<mods:affiliation id="9C52EF3F9FFEF550DEDBA1E9C9552AE9">Kieferndamm 10, D- 15566 Schöneiche bei Berlin, Germany. &amp; Entomological Laboratory Institute of Biology, Mongolian Academy of Science, Peace Avenue- 54 B, Bayanzurkh, Ulaanbaatar- 13330, Mongolia. &amp; Kieferndamm 10, D- 15566 Schöneiche bei Berlin, Germany.</mods:affiliation>
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<mods:date id="6C82A8C596DD32C641DF65AEC98CD286">2024</mods:date>
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<treatment id="004A87E8FFC9FFBFFF2AF992FD0DFEAA" ID-DOI="http://doi.org/10.5281/zenodo.14022137" ID-Zenodo-Dep="14022137" LSID="urn:lsid:plazi:treatment:004A87E8FFC9FFBFFF2AF992FD0DFEAA" httpUri="http://treatment.plazi.org/id/004A87E8FFC9FFBFFF2AF992FD0DFEAA" lastPageId="4" lastPageNumber="126" pageId="1" pageNumber="124">
<subSubSection id="C0F96575FFC9FFBAFF2AF992FD2EF99E" box="[151,1053,1550,1578]" pageId="1" pageNumber="124" type="nomenclature">
<paragraph id="885C36FEFFC9FFBAFF2AF992FD2EF99E" blockId="1.[151,1053,1550,1609]" box="[151,1053,1550,1578]" pageId="1" pageNumber="124">
<heading id="D3148192FFC9FFBAFF2AF992FD2EF99E" bold="true" box="[151,1053,1550,1578]" fontSize="11" level="1" pageId="1" pageNumber="124" reason="1">
<emphasis id="BA97EAECFFC9FFBAFF2AF992FD2EF99E" bold="true" box="[151,1053,1550,1578]" pageId="1" pageNumber="124">
<taxonomicName id="4FE34D7DFFC9FFBAFF2AF992FAB2F99E" authority="Hillert, Kral &amp; Bagaturov" authorityName=" Hillert, Kral &amp; Bagaturov" authorityYear="2024" box="[151,897,1550,1578]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="124" phylum="Arthropoda" rank="species" species="kucerai" status="sp. nov." subGenus="Heteroplistodus">
<emphasis id="BA97EAECFFC9FFBAFF2AF992F9C5F99D" bold="true" box="[151,246,1551,1578]" italics="true" pageId="1" pageNumber="124">Lethrus</emphasis>
(
<emphasis id="BA97EAECFFC9FFBAFEBAF992F8F5F99D" bold="true" box="[263,454,1551,1578]" italics="true" pageId="1" pageNumber="124">Heteroplistodus</emphasis>
)
<emphasis id="BA97EAECFFC9FFBAFE6AF992FB01F99D" bold="true" box="[471,562,1551,1578]" italics="true" pageId="1" pageNumber="124">kucerai</emphasis>
Hillert, Král &amp; Bagaturov
</taxonomicName>
,
<taxonomicNameLabel id="A1A45797FFC9FFBAFC30F992FD2EF99E" box="[909,1053,1551,1577]" pageId="1" pageNumber="124" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C0F96575FFC9FFBAFF2AF9ACFBF0F9FE" box="[151,707,1585,1609]" pageId="1" pageNumber="124" type="description">
<paragraph id="885C36FEFFC9FFBAFF2AF9ACFBF0F9FE" blockId="1.[151,1053,1550,1609]" box="[151,707,1585,1609]" pageId="1" pageNumber="124">
(
<figureCitation id="10D82A7BFFC9FFBAFF22F9ACF836F9FE" box="[159,261,1585,1609]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1939,1963]" captionTargetBox="[246,1347,190,1890]" captionTargetId="figure-26@2.[226,1362,176,1914]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus of Lethrus (Heteroplistodus) potanini species group. 1–2, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species (1 holotype, 2 allotype); 3–4, L. (H.) potanini Jakovlev, 1889 (3 male, Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011, 4 female, Inner Mongolia, Bayan Hot [Alxa], 2.vii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022198" httpUri="https://zenodo.org/record/14022198/files/figure.png" pageId="1" pageNumber="124">Figs. 1–2</figureCitation>
,
<figureCitation id="10D82A7BFFC9FFBAFEACF9ACF866F9FE" box="[273,341,1585,1609]" captionStart="FIGURES 5–10" captionStartId="6.[151,264,1861,1885]" captionTargetBox="[169,1418,352,1825]" captionTargetId="figure-104@6.[156,1431,341,1837]" captionTargetPageId="6" captionText="FIGURES 5–10. Head of Lethrus (Heteroplistodus) potanini species group. 5–8 in lateral view. 5–6, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype (5 left, 6 right); 7–8, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011) (7 left, 8 right). 9–10 in dorsal view. 9, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 10, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022200" httpUri="https://zenodo.org/record/14022200/files/figure.png" pageId="1" pageNumber="124">5–6, 9</figureCitation>
,
<figureCitation id="10D82A7BFFC9FFBAFEDCF9ACF84FF9FE" box="[353,380,1585,1609]" captionStart="FIGURES 11–12" captionStartId="7.[151,264,1633,1657]" captionTargetBox="[159,1429,203,1609]" captionTargetId="figure-22@7.[158,1429,174,1609]" captionTargetPageId="7" captionText="FIGURES 11–12. Head in frontal view of Lethrus (Heteroplistodus) potanini species group. 11, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 12, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022202" httpUri="https://zenodo.org/record/14022202/files/figure.png" pageId="1" pageNumber="124">11</figureCitation>
,
<figureCitation id="10D82A7BFFC9FFBAFE35F9ACF8FEF9FE" box="[392,461,1585,1609]" captionStart="FIGURES 13–16" captionStartId="8.[151,264,1904,1928]" captionTargetBox="[199,1366,180,1867]" captionTargetId="figure-26@8.[190,1397,177,1880]" captionTargetPageId="8" captionText="FIGURES 13–16. Lethrus (Heteroplistodus) potanini species group in dorsal view. 13–16 genae. 13, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 14, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011). 15–16 elytra. 15, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 16, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022204" httpUri="https://zenodo.org/record/14022204/files/figure.png" pageId="1" pageNumber="124">13, 15</figureCitation>
,
<figureCitation id="10D82A7BFFC9FFBAFE65F9ACFB2EF9FE" box="[472,541,1585,1609]" captionStart="FIGURES 17–20" captionStartId="9.[151,264,1908,1932]" captionTargetBox="[201,1388,193,1883]" captionTargetId="figure-22@9.[187,1401,180,1883]" captionTargetPageId="9" captionText="FIGURES 17–20. Lethrus (Heteroplistodus) potanini species group. 17–18 half of pronotum in dorsal view. 17, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 18, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3– 6.viii.2011). 19–20 aedeagus, apical part of parameres in dorsal view.19, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 20, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022206" httpUri="https://zenodo.org/record/14022206/files/figure.png" pageId="1" pageNumber="124">17, 19</figureCitation>
,
<figureCitation id="10D82A7BFFC9FFBAFD95F9ACFB5EF9FE" box="[552,621,1585,1609]" captionStart="FIGURES 21–24" captionStartId="10.[151,264,1907,1931]" captionTargetBox="[198,1389,182,1878]" captionTargetId="figure-26@10.[186,1402,174,1886]" captionTargetPageId="10" captionText="FIGURES 21–24. Lethrus (Heteroplistodus) potanini species group. 21–22 aedeagus, apical part of parameres in lateral view. 21, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 22, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011). 23–24 apical spur of protibia. 23, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 24, L. (H.) potanini (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022208" httpUri="https://zenodo.org/record/14022208/files/figure.png" pageId="1" pageNumber="124">21, 23</figureCitation>
,
<figureCitation id="10D82A7BFFC9FFBAFDC5F9ACFB8FF9FE" box="[632,700,1585,1609]" captionStart="FIGURES 25–28" captionStartId="11.[151,264,1839,1863]" captionTargetBox="[220,1369,178,1801]" captionTargetId="figure-22@11.[210,1378,174,1816]" captionTargetPageId="11" captionText="FIGURES 25–28. Lethrus (Heteroplistodus) potanini species group. 25–27 habitus in dorsal view. 25, L. (H.) potanini Jakovlev, 1889 lectotype; 26, L. conformis Frivaldszky, 1890 lectotype; 27, L. (H.) potanini paralectotype [= L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species]. 28 labels of types. A, L. (H.) potanini Jakovlev, 1889 paralectotype [L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species]; B, L. (H.) potanini Jakovlev, 1889 paralectotype; C, Lethrus conformis Frivaldszky, 1890 lectotype; D, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; E, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species allotype." figureDoi="http://doi.org/10.5281/zenodo.14022210" httpUri="https://zenodo.org/record/14022210/files/figure.png" pageId="1" pageNumber="124">27–28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C0F96575FFC9FFBAFF2AF9EDF876F911" pageId="1" pageNumber="124" type="reference_group">
<paragraph id="885C36FEFFC9FFBAFF2AF9EDF876F911" blockId="1.[151,1436,1647,1702]" pageId="1" pageNumber="124">
<treatmentCitationGroup id="A8F311D0FFC9FFBAFF2AF9EDF876F911" pageId="1" pageNumber="124">
<treatmentCitation id="094210EFFFC9FFBAFF2AF9EDFB2AF930" author="Jakovlev, B. E." box="[151,537,1647,1672]" page="261" pageId="1" pageNumber="124" year="1889">
<taxonomicName id="4FE34D7DFFC9FFBAFF2AF9EDFB2AF930" ID-CoL="6PSDN" authority="Jakovlev, 1889: 261" authorityName="Jakovlev" authorityPageNumber="261" authorityYear="1889" box="[151,537,1647,1672]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="124" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="BA97EAECFFC9FFBAFF2AF9EDF870F930" box="[151,323,1648,1672]" italics="true" pageId="1" pageNumber="124">Lethrus Potanini</emphasis>
<bibRefCitation id="EC724B0FFFC9FFBAFEF6F9EDFB2AF930" author="Jakovlev, B. E." box="[331,537,1647,1672]" pageId="1" pageNumber="124" pagination="254 - 262" refId="ref9003" refString="Jakovlev, B. E. (1889) Insecta, a Cl. A. N. Potanin in China et in Mongolia novissime lecta XI. Coleoptera (Cymindis, Pseudopelta, Lethrus). Horae Societatis Entomologicae Rossicae, 24, 254 - 262." type="journal article" year="1889">Jakovlev, 1889: 261</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[partim, populations from Ordos plateau] (original description,
<typeStatus id="5758885CFFC9FFBAFB10F9EDFDEAF93F" box="[1197,1241,1648,1672]" pageId="1" pageNumber="124">type</typeStatus>
locality: “
<collectingCountry id="F0F4766EFFC9FFBAFAFEF9EDFCB1F93F" box="[1347,1410,1648,1672]" name="China" pageId="1" pageNumber="124">China</collectingCountry>
et
<collectingCountry id="F0F4766EFFC9FFBAFF7AF913F803F911" box="[199,304,1678,1702]" name="Mongolia" pageId="1" pageNumber="124">Mongolia</collectingCountry>
”).
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</paragraph>
</subSubSection>
<subSubSection id="C0F96575FFC9FFB8FF2AF953FD98FEF6" lastPageId="3" lastPageNumber="126" pageId="1" pageNumber="124" type="materials_examined">
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<typeStatus id="5758885CFFC9FFBAFF2AF953F9FCF951" box="[151,207,1742,1766]" pageId="1" pageNumber="124">Type</typeStatus>
locality.
</emphasis>
“
<collectingCountry id="F0F4766EFFC9FFBAFE83F953F8B1F951" box="[318,386,1742,1766]" name="China" pageId="1" pageNumber="124">China</collectingCountry>
,
<collectingRegion id="4A27F81CFFC9FFBAFE30F953F8D5F951" box="[397,486,1742,1766]" country="China" name="Shaanxi" pageId="1" pageNumber="124">Shaanxi</collectingRegion>
,
<location id="8D3C6025FFC9FFBAFE4DF953FB1CF951" LSID="urn:lsid:plazi:treatment:004A87E8FFC9FFBFFF2AF992FD0DFEAA:8D3C6025FFC9FFBAFE4DF953FB1CF951" box="[496,559,1742,1766]" country="China" name="Yulin" pageId="1" pageNumber="124" stateProvince="Shaanxi">Yulin</location>
”.
</paragraph>
<paragraph id="885C36FEFFC9FFBAFF7AF96CFD99F854" blockId="1.[151,1437,1742,2019]" pageId="1" pageNumber="124">
<emphasis id="BA97EAECFFC9FFBAFF7AF96CF8D9F8BD" bold="true" box="[199,490,1776,1803]" pageId="1" pageNumber="124">
<collectingCounty id="613D4E72FFC9FFBAFF7AF96CF831F8BC" box="[199,258,1777,1803]" pageId="1" pageNumber="124">Type</collectingCounty>
material examined
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(
<specimenCount id="9EE5FD77FFC9FFBAFE46F96CFBA5F8BD" box="[507,662,1776,1803]" count="19" pageId="1" pageNumber="124" type="generic">19 specimens</specimenCount>
).
<materialsCitation id="388B3CA3FFC9FFBAFD10F96DFA53F898" collectingDate="2009-07-18" collectionCode="ZSMC" collectorName="E. Kucera" country="China" county="Rd" location="Shaanxi" pageId="1" pageNumber="124" specimenCount="1" specimenCount-male="1" stateProvince="Shaanxi" typeStatus="holotype">
<emphasis id="BA97EAECFFC9FFBAFD10F96DFAD7F8BC" bold="true" box="[685,996,1776,1803]" pageId="1" pageNumber="124">
<collectingCountry id="F0F4766EFFC9FFBAFD10F96DFBCBF8BD" box="[685,760,1776,1802]" name="China" pageId="1" pageNumber="124">China</collectingCountry>
,
<collectingRegion id="4A27F81CFFC9FFBAFCB8F96DFA59F8BD" box="[773,874,1776,1802]" country="China" name="Shaanxi" pageId="1" pageNumber="124">Shaanxi</collectingRegion>
:
<typeStatus id="5758885CFFC9FFBAFCCAF96CFAD7F8BC" box="[887,996,1777,1803]" pageId="1" pageNumber="124" type="holotype">Holotype</typeStatus>
</emphasis>
,
<specimenCount id="9EE5FD77FFC9FFBAFC4EF96CFD3BF8BC" box="[1011,1032,1777,1803]" count="1" pageId="1" pageNumber="124" type="male">♂</specimenCount>
(
<collectionCode id="EEF2AE3BFFC9FFBAFBA5F96CFD5BF8BC" box="[1048,1128,1777,1803]" collectionName="Germany, Muenchen [= Munich], Zoologische Staatssammlung" country="Germany" httpUri="http://grbio.org/cool/a6ms-py1z" name="Zoologische Staatssammlung" pageId="1" pageNumber="124">ZSMC</collectionCode>
) (
<figureCitation id="10D82A7BFFC9FFBAFB3DF96DFDF1F8BC" box="[1152,1218,1776,1803]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1939,1963]" captionTargetBox="[246,1347,190,1890]" captionTargetId="figure-26@2.[226,1362,176,1914]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus of Lethrus (Heteroplistodus) potanini species group. 1–2, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species (1 holotype, 2 allotype); 3–4, L. (H.) potanini Jakovlev, 1889 (3 male, Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011, 4 female, Inner Mongolia, Bayan Hot [Alxa], 2.vii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022198" httpUri="https://zenodo.org/record/14022198/files/figure.png" pageId="1" pageNumber="124">Fig. 1</figureCitation>
), “China-Shaanxi | Yulin | 14. -
<date id="FC5D103EFFC9FFBAFEA7F888F8BFF898" box="[282,396,1813,1839]" pageId="1" pageNumber="124" value="2009-07-18">
<collectingDate id="EC19E9D6FFC9FFBAFEA7F888F8BFF898" box="[282,396,1813,1839]" pageId="1" pageNumber="124" value="2009-07-18">18.7.2009</collectingDate>
</date>
| leg.
<collectorName id="25165328FFC9FFBAFE6CF888FB71F898" box="[465,578,1813,1839]" pageId="1" pageNumber="124">E. Kučera</collectorName>
|| ex. coll. O. Hillert [p]”
</materialsCitation>
.
<materialsCitation id="388B3CA3FFC9FFBAFCD1F888F801F8E4" collectionCode="ZSMC" country="China" county="Rd" pageId="1" pageNumber="124" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Shaanxi" typeStatus="paratype">
<emphasis id="BA97EAECFFC9FFBAFCD1F888FAD9F898" bold="true" box="[876,1002,1813,1839]" pageId="1" pageNumber="124">
<typeStatus id="5758885CFFC9FFBAFCD1F888FAD6F898" box="[876,997,1813,1839]" pageId="1" pageNumber="124" type="paratype">Paratypes</typeStatus>
,
</emphasis>
allotype,
<specimenCount id="9EE5FD77FFC9FFBAFBE6F888FD43F898" box="[1115,1136,1813,1839]" count="1" pageId="1" pageNumber="124" type="female">♀</specimenCount>
(
<collectionCode id="EEF2AE3BFFC9FFBAFBC2F888FDFCF898" box="[1151,1231,1813,1839]" collectionName="Germany, Muenchen [= Munich], Zoologische Staatssammlung" country="Germany" httpUri="http://grbio.org/cool/a6ms-py1z" name="Zoologische Staatssammlung" pageId="1" pageNumber="124">ZSMC</collectionCode>
) (
<figureCitation id="10D82A7BFFC9FFBAFB5BF889FC18F898" box="[1254,1323,1812,1839]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1939,1963]" captionTargetBox="[246,1347,190,1890]" captionTargetId="figure-26@2.[226,1362,176,1914]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus of Lethrus (Heteroplistodus) potanini species group. 1–2, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species (1 holotype, 2 allotype); 3–4, L. (H.) potanini Jakovlev, 1889 (3 male, Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011, 4 female, Inner Mongolia, Bayan Hot [Alxa], 2.vii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022198" httpUri="https://zenodo.org/record/14022198/files/figure.png" pageId="1" pageNumber="124">Fig. 2</figureCitation>
),
<specimenCount id="9EE5FD77FFC9FFBAFAFDF888FCB1F898" box="[1344,1410,1813,1839]" count="2" pageId="1" pageNumber="124" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9EE5FD77FFC9FFBAFA33F889F9F1F8E4" count="5" pageId="1" pageNumber="124" type="female">5 ♀♀</specimenCount>
, (
<collectionCode id="EEF2AE3BFFC9FFBAFF65F8A4F819F8E4" box="[216,298,1849,1875]" collectionName="Germany, Muenchen [= Munich], Zoologische Staatssammlung" country="Germany" httpUri="http://grbio.org/cool/a6ms-py1z" name="Zoologische Staatssammlung" pageId="1" pageNumber="124">ZSMC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="388B3CA3FFC9FFBAFE83F8A4FB0CF8E4" box="[318,575,1849,1875]" collectionCode="NMPC" country="China" county="Rd" pageId="1" pageNumber="124" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Shaanxi" typeStatus="paratype">
<specimenCount id="9EE5FD77FFC9FFBAFE83F8A4F84CF8E4" box="[318,383,1849,1875]" count="3" pageId="1" pageNumber="124" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9EE5FD77FFC9FFBAFE31F8A4F8FEF8E4" box="[396,461,1849,1875]" count="2" pageId="1" pageNumber="124" type="female">2 ♀♀</specimenCount>
, (
<collectionCode id="EEF2AE3BFFC9FFBAFE5FF8A4FB04F8E4" box="[482,567,1849,1875]" collectionName="Czech Republic, Prague, National Museum (Natural History)" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="124">NMPC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="388B3CA3FFC9FFBAFDF6F8A4FBE5F8E4" box="[587,726,1849,1875]" collectionCode="JSCS" country="China" county="Rd" pageId="1" pageNumber="124" specimenCount="1" specimenCount-female="1" stateProvince="Shaanxi" typeStatus="paratype">
<specimenCount id="9EE5FD77FFC9FFBAFDF6F8A4FB45F8E4" box="[587,630,1849,1875]" count="1" pageId="1" pageNumber="124" type="female">1 ♀</specimenCount>
, (
<collectionCode id="EEF2AE3BFFC9FFBAFD30F8A4FBFCF8E4" box="[653,719,1849,1875]" pageId="1" pageNumber="124">JSCS</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="388B3CA3FFC9FFBAFD5EF8A4FDA9F8E5" box="[739,1178,1848,1875]" collectingDate="2009-07-18" collectionCode="SJCP" collectorName="E. Kucera &amp; O. Hillert" country="China" county="Rd" pageId="1" pageNumber="124" specimenCount="1" specimenCount-female="1" stateProvince="Shaanxi" typeStatus="paratype">
<specimenCount id="9EE5FD77FFC9FFBAFD5EF8A4FA3EF8E4" box="[739,781,1849,1875]" count="1" pageId="1" pageNumber="124" type="female">1 ♀</specimenCount>
, (
<collectionCode id="EEF2AE3BFFC9FFBAFC99F8A4FA57F8E4" box="[804,868,1849,1875]" pageId="1" pageNumber="124">SJCP</collectionCode>
) “[same data as holotype]”
</materialsCitation>
;
<materialsCitation id="388B3CA3FFC9FFBAFB1AF8A4F9C4F82D" collectingDate="1992-07-20" collectionCode="HBUM" collectorName="Guodong Ren" country="China" county="Rd" location="Rd" pageId="1" pageNumber="124" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Shaanxi" typeStatus="paratype">
<specimenCount id="9EE5FD77FFC9FFBAFB1AF8A4FDDBF8E4" box="[1191,1256,1849,1875]" count="2" pageId="1" pageNumber="124" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9EE5FD77FFC9FFBAFB48F8A4FC13F8E4" box="[1269,1312,1849,1875]" count="1" pageId="1" pageNumber="124" type="female">1 ♀</specimenCount>
, (
<collectionCode id="EEF2AE3BFFC9FFBAFA8BF8A4FCA3F8E4" box="[1334,1424,1849,1875]" country="China" httpUri="http://grbio.org/cool/6ww3-041h" name="College of Life Sciences Hebei Univesity, Baoding" pageId="1" pageNumber="124">HBUM</collectionCode>
), “
<date id="FC5D103EFFC9FFBAFF1FF8C0F82BF8C0" box="[162,280,1885,1911]" pageId="1" pageNumber="124" value="1992-07-20">
<collectingDate id="EC19E9D6FFC9FFBAFF1FF8C0F82BF8C0" box="[162,280,1885,1911]" pageId="1" pageNumber="124" value="1992-07-20">1992-7-20</collectingDate>
</date>
|
<emphasis id="BA97EAECFFC9FFBAFE97F8C0F8E1F8C3" bold="true" box="[298,466,1885,1908]" pageId="1" pageNumber="124">Ŕđ省榆µ市</emphasis>
|
<emphasis id="BA97EAECFFC9FFBAFE59F8C0FB67F8C3" bold="true" box="[484,596,1885,1908]" pageId="1" pageNumber="124">Ẻ国栋梁</emphasis>
|| HBU(E) || Shaanxi,
<date id="FC5D103EFFC9FFBAFCF4F8C1FAFFF8C1" box="[841,972,1884,1910]" pageId="1" pageNumber="124" value="1992-07-20">
<collectingDate id="EC19E9D6FFC9FFBAFCF4F8C1FAFFF8C1" box="[841,972,1884,1910]" pageId="1" pageNumber="124" value="1992-07-20">20.vii.1992</collectingDate>
</date>
|YULIN |
<collectorName id="25165328FFC9FFBAFB81F8C1FDE4F8C0" box="[1084,1239,1884,1911]" pageId="1" pageNumber="124">Guodong Ren</collectorName>
lgt. [p]”.
<taxonomicName id="4FE34D7DFFC9FFBAFAFFF8C1F9C4F82D" authorityName="Jakovlev" authorityYear="1889" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="124" phylum="Arthropoda" rank="species" species="potanini">Lethrus potanini</taxonomicName>
</materialsCitation>
<emphasis id="BA97EAECFFC9FFBAFF45F81CF9CCF82C" bold="true" box="[248,255,1921,1947]" pageId="1" pageNumber="124">.</emphasis>
<materialsCitation id="388B3CA3FFC9FFBAFEBBF81CFD96F854" collectionCode="ZMAS" collectorName="G. Patanin." country="China" county="Rd" determinerName="B. Jakowlew" location="Ordos" municipality="Ordos" pageId="1" pageNumber="124" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Shaanxi" typeStatus="paralectotype">
<emphasis id="BA97EAECFFC9FFBAFEBBF81CF88FF82C" bold="true" box="[262,444,1921,1947]" pageId="1" pageNumber="124">
<typeStatus id="5758885CFFC9FFBAFEBBF81CF884F82C" box="[262,439,1921,1947]" pageId="1" pageNumber="124" type="paralectotype">Paralectotypes</typeStatus>
,
</emphasis>
<specimenCount id="9EE5FD77FFC9FFBAFE7EF81CFB37F82C" box="[451,516,1921,1947]" count="3" pageId="1" pageNumber="124" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9EE5FD77FFC9FFBAFDADF81CFB09F82C" box="[528,570,1921,1947]" count="1" pageId="1" pageNumber="124" type="female">1 ♀</specimenCount>
(
<collectionCode id="EEF2AE3BFFC9FFBAFDF4F81CFBAFF82C" box="[585,668,1921,1947]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="1" pageNumber="124">ZMAS</collectionCode>
), (
<figureCitation id="10D82A7BFFC9FFBAFD05F81DFA3FF82C" box="[696,780,1920,1947]" captionStart="FIGURES 25–28" captionStartId="11.[151,264,1839,1863]" captionTargetBox="[220,1369,178,1801]" captionTargetId="figure-22@11.[210,1378,174,1816]" captionTargetPageId="11" captionText="FIGURES 25–28. Lethrus (Heteroplistodus) potanini species group. 25–27 habitus in dorsal view. 25, L. (H.) potanini Jakovlev, 1889 lectotype; 26, L. conformis Frivaldszky, 1890 lectotype; 27, L. (H.) potanini paralectotype [= L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species]. 28 labels of types. A, L. (H.) potanini Jakovlev, 1889 paralectotype [L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species]; B, L. (H.) potanini Jakovlev, 1889 paralectotype; C, Lethrus conformis Frivaldszky, 1890 lectotype; D, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; E, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species allotype." figureDoi="http://doi.org/10.5281/zenodo.14022210" httpUri="https://zenodo.org/record/14022210/files/figure.png" pageId="1" pageNumber="124">Fig. 27</figureCitation>
): “[round label, blue-blackish] ||
<collectingMunicipality id="6838AC84FFC9FFBAFBC1F81DFDF3F82D" box="[1148,1216,1920,1946]" pageId="1" pageNumber="124">Ordos</collectingMunicipality>
| 1884 |
<collectorName id="25165328FFC9FFBAFAA2F81CFCAEF82D" box="[1311,1437,1920,1947]" pageId="1" pageNumber="124">G. Patanin.</collectorName>
[p] || Heteroplistodes | typ.
<taxonomicName id="4FE34D7DFFC9FFBAFE7EF839FB61F809" authority="Jak." authorityName="Jakovlev" authorityYear="1889" box="[451,594,1956,1982]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="124" phylum="Arthropoda" rank="species" species="potanini">potanini Jak.</taxonomicName>
| B. Jakowlew det. [hw, p] || к. В. Яковлева [p] || Cotypus [p, red label] [p]” [all these specimens are not paratypes of
<taxonomicName id="4FE34D7DFFC9FFBAFDD9F854FBE5F855" authorityName="Hillert &amp; Král &amp; Bagaturov &amp; Chuluunbaatar" authorityYear="2024" box="[612,726,1992,2018]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="124" phylum="Arthropoda" rank="species" species="kucerai" status="new species">
<emphasis id="BA97EAECFFC9FFBAFDD9F854FBE5F855" box="[612,726,1992,2018]" italics="true" pageId="1" pageNumber="124">L. kucerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1A45797FFC9FFBAFD60F854FA55F855" box="[733,870,1992,2019]" pageId="1" pageNumber="124" rank="species">new species</taxonomicNameLabel>
,
<emphasis id="BA97EAECFFC9FFBAFCCCF854FAAAF855" box="[881,921,1993,2018]" italics="true" pageId="1" pageNumber="124">e.g.</emphasis>
note under
<taxonomicName id="4FE34D7DFFC9FFBAFBA2F854FDAAF855" authorityName="Jakovlev" authorityYear="1889" box="[1055,1177,1993,2018]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="124" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="BA97EAECFFC9FFBAFBA2F854FDAAF855" box="[1055,1177,1993,2018]" italics="true" pageId="1" pageNumber="124">L. potanini</emphasis>
</taxonomicName>
]
</materialsCitation>
.
</paragraph>
<caption id="DC9C6676FFCAFFB9FF2AF80EFAF3F844" ID-DOI="http://doi.org/10.5281/zenodo.14022198" ID-Zenodo-Dep="14022198" httpUri="https://zenodo.org/record/14022198/files/figure.png" pageId="2" pageNumber="125" startId="2.[151,264,1939,1963]" targetBox="[246,1347,190,1890]" targetPageId="2" targetType="figure">
<paragraph id="885C36FEFFCAFFB9FF2AF80EFAF3F844" blockId="2.[151,1437,1939,2036]" pageId="2" pageNumber="125">
<emphasis id="BA97EAECFFCAFFB9FF2AF80EF80CF81C" bold="true" box="[151,319,1939,1963]" pageId="2" pageNumber="125">FIGURES 1–4.</emphasis>
Dorsal habitus of
<taxonomicName id="4FE34D7DFFCAFFB9FDB6F80EFA40F81C" authorityName="Jakovlev" authorityYear="1889" box="[523,883,1939,1963]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="125" phylum="Arthropoda" rank="species" species="potanini" subGenus="Heteroplistodus">
<emphasis id="BA97EAECFFCAFFB9FDB6F80EFB6BF81C" box="[523,600,1939,1963]" italics="true" pageId="2" pageNumber="125">Lethrus</emphasis>
(
<emphasis id="BA97EAECFFCAFFB9FDD6F80EFA3FF81C" box="[619,780,1939,1963]" italics="true" pageId="2" pageNumber="125">Heteroplistodus</emphasis>
)
<emphasis id="BA97EAECFFCAFFB9FCA2F809FA40F81C" box="[799,883,1940,1963]" italics="true" pageId="2" pageNumber="125">potanini</emphasis>
</taxonomicName>
species group. 1–2,
<taxonomicName id="4FE34D7DFFCAFFB9FBE9F809F831F867" authority="Hillert, Kral &amp; Bagaturov" authorityName=", Hillert, Kral &amp; Bagaturov" authorityYear="2024" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="125" phylum="Arthropoda" rank="species" species="kucerai" status="new species" subGenus="Heteroplistodus">
<emphasis id="BA97EAECFFCAFFB9FBE9F809FD51F81C" box="[1108,1122,1940,1963]" italics="true" pageId="2" pageNumber="125">L</emphasis>
. (
<emphasis id="BA97EAECFFCAFFB9FBC1F809FDA6F81C" box="[1148,1173,1940,1963]" italics="true" pageId="2" pageNumber="125">H.</emphasis>
)
<emphasis id="BA97EAECFFCAFFB9FB1AF80EFDC0F81C" box="[1191,1267,1939,1963]" italics="true" pageId="2" pageNumber="125">kucerai</emphasis>
Hillert, Král &amp; Bagaturov
</taxonomicName>
,
<emphasis id="BA97EAECFFCAFFB9FEB3F82AF8BEF878" bold="true" box="[270,397,1975,1999]" pageId="2" pageNumber="125">
<taxonomicNameLabel id="A1A45797FFCAFFB9FEB3F82AF8BEF878" box="[270,397,1975,1999]" pageId="2" pageNumber="125" rank="species">new species</taxonomicNameLabel>
</emphasis>
(1 holotype, 2 allotype); 3–4,
<taxonomicName id="4FE34D7DFFCAFFB9FD6DF825FD24F878" authority="Jakovlev, 1889" authorityName="Jakovlev" authorityYear="1889" box="[720,1047,1975,1999]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="125" phylum="Arthropoda" rank="species" species="potanini" subGenus="Heteroplistodus">
<emphasis id="BA97EAECFFCAFFB9FD6DF825FBEDF878" box="[720,734,1976,1999]" italics="true" pageId="2" pageNumber="125">L</emphasis>
. (
<emphasis id="BA97EAECFFCAFFB9FD48F825FA3CF878" box="[757,783,1976,1999]" italics="true" pageId="2" pageNumber="125">H.</emphasis>
)
<emphasis id="BA97EAECFFCAFFB9FCA2F825FA40F878" box="[799,883,1976,1999]" italics="true" pageId="2" pageNumber="125">potanini</emphasis>
<bibRefCitation id="EC724B0FFFCAFFB9FCC1F82AFD24F878" author="Jakovlev, B. E." box="[892,1047,1975,1999]" pageId="2" pageNumber="125" pagination="254 - 262" refId="ref9003" refString="Jakovlev, B. E. (1889) Insecta, a Cl. A. N. Potanin in China et in Mongolia novissime lecta XI. Coleoptera (Cymindis, Pseudopelta, Lethrus). Horae Societatis Entomologicae Rossicae, 24, 254 - 262." type="journal article" year="1889">Jakovlev, 1889</bibRefCitation>
</taxonomicName>
(3 male, Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011, 4 female, Inner Mongolia, Bayan Hot [Alxa], 2.vii.2011).
</paragraph>
</caption>
<paragraph id="885C36FEFFCBFFB8FF7AFF0BFD98FEF6" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="126">
<emphasis id="BA97EAECFFCBFFB8FF7AFF0BFB1EFF07" bold="true" box="[199,557,150,176]" pageId="3" pageNumber="126">Additional material examined</emphasis>
(
<specimenCount id="9EE5FD77FFCBFFB8FD81FF0AFBE4FF06" box="[572,727,151,177]" count="22" pageId="3" pageNumber="126" type="generic">22 specimens</specimenCount>
).
<materialsCitation id="388B3CA3FFCBFFB8FD50FF0BFB86FF62" collectingDate="2007-07-18" collectingDateMax="2007-07-22" collectingDateMin="2007-07-18" collectionCode="NMPC" collectorName="E. Kucera" country="China" location="Genghis" municipality="Dongsheng" pageId="3" pageNumber="126" specimenCount="1" specimenCount-female="1" specimenCount-male="7" stateProvince="Inner Mongolia">
<emphasis id="BA97EAECFFCBFFB8FD50FF0BFD3AFF07" bold="true" box="[749,1033,150,177]" pageId="3" pageNumber="126">
<collectingCountry id="F0F4766EFFCBFFB8FD50FF0BFA0BFF07" box="[749,824,150,176]" name="China" pageId="3" pageNumber="126">China</collectingCountry>
,
<collectingRegion id="4A27F81CFFCBFFB8FCF9FF0AFD37FF07" box="[836,1028,150,177]" country="China" name="Nei Mongol" pageId="3" pageNumber="126">Inner Mongolia</collectingRegion>
:
</emphasis>
<specimenCount id="9EE5FD77FFCBFFB8FBADFF0AFD09FF06" box="[1040,1082,151,177]" count="1" pageId="3" pageNumber="126" type="female">1 ♀</specimenCount>
(
<collectionCode id="EEF2AE3BFFCBFFB8FBF4FF0AFDADFF06" box="[1097,1182,151,177]" collectionName="Czech Republic, Prague, National Museum (Natural History)" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="126">NMPC</collectionCode>
),
<collectingMunicipality id="6838AC84FFCBFFB8FB0CFF0AFC00FF06" box="[1201,1331,151,177]" pageId="3" pageNumber="126">Dongsheng</collectingMunicipality>
,
<location id="8D3C6025FFCBFFB8FA82FF0AFCAFFF06" LSID="urn:lsid:plazi:treatment:004A87E8FFC9FFBFFF2AF992FD0DFEAA:8D3C6025FFCBFFB8FA82FF0AFCAFFF06" box="[1343,1436,151,177]" country="China" municipality="Dongsheng" name="Genghis" pageId="3" pageNumber="126" stateProvince="Inner Mongolia">Genghis</location>
khan-memorial,
<date id="FC5D103EFFCBFFB8FEECFF26FB37FF62" box="[337,516,187,213]" pageId="3" pageNumber="126" value="2007-07-18" valueMax="2007-07-22" valueMin="2007-07-18">
<collectingDate id="EC19E9D6FFCBFFB8FEECFF26FB37FF62" box="[337,516,187,213]" pageId="3" pageNumber="126" value="2007-07-18" valueMax="2007-07-22" valueMin="2007-07-18">18.–22.vii.2007</collectingDate>
</date>
,
<collectorName id="25165328FFCBFFB8FDACFF26FBB1FF62" box="[529,642,187,213]" pageId="3" pageNumber="126">E. Kučera</collectorName>
lgt.,
</materialsCitation>
<materialsCitation id="388B3CA3FFCBFFB8FD00FF26FD59FF62" box="[701,1130,187,213]" collectingDate="2007-07-18" collectingDateMax="2013-06-09" collectingDateMin="2007-07-18" collectionCode="OHCB" collectorName="Peter Schneider" country="China" location="Genghis" municipality="Dongsheng" pageId="3" pageNumber="126" specimenCount="1" specimenCount-female="12" specimenCount-male="1" stateProvince="Nei Mongol">
<specimenCount id="9EE5FD77FFCBFFB8FD00FF26FBDBFF62" box="[701,744,187,213]" count="1" pageId="3" pageNumber="126" type="male">1 ♂</specimenCount>
(
<collectionCode id="EEF2AE3BFFCBFFB8FD45FF26FA7AFF62" box="[760,841,187,213]" pageId="3" pageNumber="126">OHCB</collectionCode>
) but
<collectorName id="25165328FFCBFFB8FC39FF26FD05FF62" box="[900,1078,187,213]" pageId="3" pageNumber="126">Peter Schneider</collectorName>
lgt.;
</materialsCitation>
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<specimenCount id="9EE5FD77FFCBFFB8FBCFFF26FDACFF62" box="[1138,1183,187,213]" count="1" pageId="3" pageNumber="126" type="male">1 ♂</specimenCount>
,
<specimenCount id="9EE5FD77FFCBFFB8FB11FF26FDDFFF62" box="[1196,1260,187,213]" count="2" pageId="3" pageNumber="126" type="female">2 ♀♀</specimenCount>
(
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),
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env.,
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<collectingDate id="EC19E9D6FFCBFFB8FE83FF42F8EEFF4E" box="[318,477,223,249]" pageId="3" pageNumber="126" value="2011-05-17" valueMax="2011-05-19" valueMin="2011-05-17">17.–19.v.2011</collectingDate>
</date>
,
<collectorName id="25165328FFCBFFB8FE54FF42FB61FF4E" box="[489,594,223,249]" pageId="3" pageNumber="126">J. S. Gull</collectorName>
lgt.;
</materialsCitation>
<materialsCitation id="388B3CA3FFCBFFB8FD30FF42F835FEAA" collectingDate="2013-06-09" collectingDateMax="2013-06-09" collectingDateMin="2007-07-18" collectionCode="HBUM" collectorName="Sun &amp; Chen" country="China" location="Zhungeerqi" municipality="Dongsheng" pageId="3" pageNumber="126" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Nei Mongol">
<specimenCount id="9EE5FD77FFCBFFB8FD30FF42FBFDFF4E" box="[653,718,223,249]" count="4" pageId="3" pageNumber="126" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="9EE5FD77FFCBFFB8FD66FF42FA29FF4E" box="[731,794,223,249]" count="6" pageId="3" pageNumber="126" type="female">6 ♀♀</specimenCount>
(
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),
<location id="8D3C6025FFCBFFB8FC2BFF42FD29FF4E" LSID="urn:lsid:plazi:treatment:004A87E8FFC9FFBFFF2AF992FD0DFEAA:8D3C6025FFCBFFB8FC2BFF42FD29FF4E" box="[918,1050,223,249]" country="China" municipality="Dongsheng" name="Zhungeerqi" pageId="3" pageNumber="126" stateProvince="Nei Mongol">Zhungeerqi</location>
,
<location id="8D3C6025FFCBFFB8FB98FF42FDE3FF4E" LSID="urn:lsid:plazi:treatment:004A87E8FFC9FFBFFF2AF992FD0DFEAA:8D3C6025FFCBFFB8FB98FF42FDE3FF4E" box="[1061,1232,223,249]" country="China" municipality="Dongsheng" name="Shierliancheng" pageId="3" pageNumber="126" stateProvince="Nei Mongol">Shierliancheng</location>
,
<date id="FC5D103EFFCBFFB8FB61FF42FC7BFF4E" box="[1244,1352,223,249]" pageId="3" pageNumber="126" value="2013-06-09">
<collectingDate id="EC19E9D6FFCBFFB8FB61FF42FC7BFF4E" box="[1244,1352,223,249]" pageId="3" pageNumber="126" value="2013-06-09">9.vi.2013</collectingDate>
</date>
,
<collectorName id="25165328FFCBFFB8FAE9FF42FCB3FF4E" box="[1364,1408,223,249]" pageId="3" pageNumber="126">Sun</collectorName>
&amp;
<collectorName id="25165328FFCBFFB8FF2AFE9EF9E1FEAA" box="[151,210,259,285]" pageId="3" pageNumber="126">Chen</collectorName>
lgt.;
</materialsCitation>
<materialsCitation id="388B3CA3FFCBFFB8FEB0FE9EFD18FEAA" box="[269,1067,258,285]" collectingDate="2007-07-18" collectingDateMax="2013-06-09" collectingDateMin="2007-07-18" collectionCode="MBCP" collectorName="E. Kucera &amp; J. S. Gull &amp; Sun &amp; Chen &amp; Ordos &amp; Boro-Balgasun &amp; G. Patanin" country="China" location="Ordos plateau" municipality="Dongsheng" pageId="3" pageNumber="126" specimenCount="1" specimenCount-female="12" specimenCount-male="1" stateProvince="Nei Mongol">
<specimenCount id="9EE5FD77FFCBFFB8FEB0FE9EF80BFEAA" box="[269,312,259,285]" count="1" pageId="3" pageNumber="126" type="male">1 ♂</specimenCount>
(
<collectionCode id="EEF2AE3BFFCBFFB8FEFAFE9EF8A9FEAA" box="[327,410,259,285]" pageId="3" pageNumber="126">MBCP</collectionCode>
),
<location id="8D3C6025FFCBFFB8FE13FE9EFB7EFEAA" LSID="urn:lsid:plazi:treatment:004A87E8FFC9FFBFFF2AF992FD0DFEAA:8D3C6025FFCBFFB8FE13FE9EFB7EFEAA" box="[430,589,259,285]" country="China" municipality="Dongsheng" name="Ordos plateau" pageId="3" pageNumber="126" stateProvince="Nei Mongol">Ordos plateau</location>
,
<location id="8D3C6025FFCBFFB8FDE4FE9EFA30FEAA" LSID="urn:lsid:plazi:treatment:004A87E8FFC9FFBFFF2AF992FD0DFEAA:8D3C6025FFCBFFB8FDE4FE9EFA30FEAA" box="[601,771,259,285]" country="China" municipality="Dongsheng" name="Boro-Balgasun" pageId="3" pageNumber="126" stateProvince="Nei Mongol">Boro-Balgasun</location>
env.
<emphasis id="BA97EAECFFCBFFB8FC83FE9FFA74FEAB" bold="true" box="[830,839,258,284]" pageId="3" pageNumber="126">(</emphasis>
Chengchuan), 1932;
</materialsCitation>
<materialsCitation id="388B3CA3FFCBFFB8FB8FFE9EF9DCFEF6" collectingDate="2007-07-18" collectingDateMax="2013-06-09" collectingDateMin="2007-07-18" collectionCode="ZMAS" collectorName="G. Patanin" country="China" location="Ordos" municipality="Dongsheng" pageId="3" pageNumber="126" specimenCount="3" specimenCount-female="3" specimenCount-male="7" stateProvince="Nei Mongol">
<specimenCount id="9EE5FD77FFCBFFB8FB8FFE9EFD41FEAA" box="[1074,1138,259,285]" count="3" pageId="3" pageNumber="126" type="female">3 ♀♀</specimenCount>
(
<collectionCode id="EEF2AE3BFFCBFFB8FB3CFE9EFDE7FEAA" box="[1153,1236,259,285]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" pageId="3" pageNumber="126">ZMAS</collectionCode>
),
<location id="8D3C6025FFCBFFB8FB55FE9EFC1CFEAA" LSID="urn:lsid:plazi:treatment:004A87E8FFC9FFBFFF2AF992FD0DFEAA:8D3C6025FFCBFFB8FB55FE9EFC1CFEAA" box="[1256,1327,259,285]" country="China" municipality="Dongsheng" name="Ordos" pageId="3" pageNumber="126" stateProvince="Nei Mongol">Ordos</location>
, 1884,
<collectorName id="25165328FFCBFFB8FA3CFE9EF9DCFEF6" pageId="3" pageNumber="126">G. Patanin</collectorName>
</materialsCitation>
.
<materialsCitation id="388B3CA3FFCBFFB8FF44FEBBFD94FEF6" box="[249,1191,294,321]" collectingDate="2007-05-04" collectionCode="HBUM" collectorName="Guodong Ren &amp; Wenjun Hou" country="China" location="Fugu" pageId="3" pageNumber="126" specimenCount="3" specimenCount-male="3" stateProvince="Shaanxi">
<emphasis id="BA97EAECFFCBFFB8FF44FEBBF889FEF7" bold="true" box="[249,442,294,320]" pageId="3" pageNumber="126">
<collectingCountry id="F0F4766EFFCBFFB8FF44FEBBF877FEF7" box="[249,324,294,320]" name="China" pageId="3" pageNumber="126">China</collectingCountry>
,
<collectingRegion id="4A27F81CFFCBFFB8FEEDFEBBF886FEF7" box="[336,437,294,320]" country="China" name="Shaanxi" pageId="3" pageNumber="126">Shaanxi</collectingRegion>
:
</emphasis>
<specimenCount id="9EE5FD77FFCBFFB8FE7CFEBAFB33FEF6" box="[449,512,295,321]" count="3" pageId="3" pageNumber="126" type="male">3 ♂♂</specimenCount>
(
<collectionCode id="EEF2AE3BFFCBFFB8FDB2FEBAFB5AFEF6" box="[527,617,295,321]" country="China" httpUri="http://grbio.org/cool/6ww3-041h" name="College of Life Sciences Hebei Univesity, Baoding" pageId="3" pageNumber="126">HBUM</collectionCode>
),
<location id="8D3C6025FFCBFFB8FDC1FEBAFB8BFEF6" LSID="urn:lsid:plazi:treatment:004A87E8FFC9FFBFFF2AF992FD0DFEAA:8D3C6025FFCBFFB8FDC1FEBAFB8BFEF6" box="[636,696,295,321]" country="China" name="Fugu" pageId="3" pageNumber="126" stateProvince="Shaanxi">Fugu</location>
,
<date id="FC5D103EFFCBFFB8FD7EFEBAFA15FEF6" box="[707,806,295,321]" pageId="3" pageNumber="126" value="2007-05-04">
<collectingDate id="EC19E9D6FFCBFFB8FD7EFEBAFA15FEF6" box="[707,806,295,321]" pageId="3" pageNumber="126" value="2007-05-04">4.v.2007</collectingDate>
</date>
,
<collectorName id="25165328FFCBFFB8FC8FFEBAFAFDFEF6" box="[818,974,295,321]" pageId="3" pageNumber="126">Guodong Ren</collectorName>
&amp;
<collectorName id="25165328FFCBFFB8FC4CFEBAFD4CFEF6" box="[1009,1151,295,321]" pageId="3" pageNumber="126">Wenjun Hou</collectorName>
lgt
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C0F96575FFCBFFB8FF7AFED7FCAFF92E" pageId="3" pageNumber="126" type="description">
<paragraph id="885C36FEFFCBFFB8FF7AFED7FA49FE1A" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="126">
<emphasis id="BA97EAECFFCBFFB8FF7AFED7F8C6FED3" bold="true" box="[199,501,330,357]" pageId="3" pageNumber="126">
Description of
<typeStatus id="5758885CFFCBFFB8FEC1FED6F8D0FED2" box="[380,483,331,357]" pageId="3" pageNumber="126" type="holotype">holotype</typeStatus>
(
</emphasis>
<specimenCount id="9EE5FD77FFCBFFB8FE48FED6FB39FED2" box="[501,522,331,357]" count="1" pageId="3" pageNumber="126" type="male">♂</specimenCount>
<emphasis id="BA97EAECFFCBFFB8FDB7FED7FB20FED3" bold="true" box="[522,531,330,356]" pageId="3" pageNumber="126">)</emphasis>
(
<figureCitation id="10D82A7BFFCBFFB8FD98FED6FB5AFED2" box="[549,617,331,357]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1939,1963]" captionTargetBox="[246,1347,190,1890]" captionTargetId="figure-26@2.[226,1362,176,1914]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus of Lethrus (Heteroplistodus) potanini species group. 1–2, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species (1 holotype, 2 allotype); 3–4, L. (H.) potanini Jakovlev, 1889 (3 male, Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011, 4 female, Inner Mongolia, Bayan Hot [Alxa], 2.vii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022198" httpUri="https://zenodo.org/record/14022198/files/figure.png" pageId="3" pageNumber="126">Fig. 1</figureCitation>
). Maximally developed (hyperthelic) male with well developed ventral mandible processes (
<figureCitation id="10D82A7BFFCBFFB8FEC2FEF2F88CFE3E" box="[383,447,367,393]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1939,1963]" captionTargetBox="[246,1347,190,1890]" captionTargetId="figure-26@2.[226,1362,176,1914]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus of Lethrus (Heteroplistodus) potanini species group. 1–2, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species (1 holotype, 2 allotype); 3–4, L. (H.) potanini Jakovlev, 1889 (3 male, Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011, 4 female, Inner Mongolia, Bayan Hot [Alxa], 2.vii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022198" httpUri="https://zenodo.org/record/14022198/files/figure.png" pageId="3" pageNumber="126">Fig. 1</figureCitation>
). Body oblong, length
<quantity id="4F1B9B1BFFCBFFB8FD03FEF2FA3DFE3E" box="[702,782,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="126" unit="mm" value="18.0">18 mm</quantity>
, strongly convex; dorsal surface blackish with distinct blue tinge, claws blackish brown, setation brownish; moderately shiny.
</paragraph>
<paragraph id="885C36FEFFCBFFB8FF7AFE2AF867FCA2" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="126">
<emphasis id="BA97EAECFFCBFFB8FF7AFE2AF837FE66" box="[199,260,439,465]" italics="true" pageId="3" pageNumber="126">Head</emphasis>
(
<figureCitation id="10D82A7BFFCBFFB8FEA8FE2AF850FE66" box="[277,355,439,465]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1939,1963]" captionTargetBox="[246,1347,190,1890]" captionTargetId="figure-26@2.[226,1362,176,1914]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus of Lethrus (Heteroplistodus) potanini species group. 1–2, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species (1 holotype, 2 allotype); 3–4, L. (H.) potanini Jakovlev, 1889 (3 male, Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011, 4 female, Inner Mongolia, Bayan Hot [Alxa], 2.vii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022198" httpUri="https://zenodo.org/record/14022198/files/figure.png" pageId="3" pageNumber="126">Figs. 1</figureCitation>
,
<figureCitation id="10D82A7BFFCBFFB8FECEFE2AF8B0FE66" box="[371,387,439,465]" captionStart="FIGURES 5–10" captionStartId="6.[151,264,1861,1885]" captionTargetBox="[169,1418,352,1825]" captionTargetId="figure-104@6.[156,1431,341,1837]" captionTargetPageId="6" captionText="FIGURES 5–10. Head of Lethrus (Heteroplistodus) potanini species group. 5–8 in lateral view. 5–6, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype (5 left, 6 right); 7–8, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011) (7 left, 8 right). 9–10 in dorsal view. 9, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 10, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022200" httpUri="https://zenodo.org/record/14022200/files/figure.png" pageId="3" pageNumber="126">9</figureCitation>
). Labrum bilobed, asymmetrical, right lobe remarkably more developed; surface rugose and shiny. Clypeus transverse, trapezoidal with anterior angles rounded, anterior margin strongly bent up at the side of the middle. Frontal impressions vague horseshoe-shaped to v-shaped, frontal tubercles elongate and strongly rounded. Frontoclypeal suture present only laterally only; keels separating eye canthus from frons distinctly, slightly divergent posteriad. Eye canthus exceeding eyes, projecting anterolaterad, almost pointed, lateral margins irregularly divergent anteriad, anterolateral angle moderate pointed and bend upward, oblique keel above eyes absent. Pleurostomal process rounded, not exceeding ventrolateral mandible outline. Punctation of frons coarse, composed of rough irregular indentations; coarse punctures separated by approximately one of their diameters, punctation becoming weakly denser posteriad on occiput; clypeus and eye canthus comparably punctured, weakly less pronounced.
</paragraph>
<paragraph id="885C36FEFFCBFFB8FF7AFC82F879FBA6" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="126">
<emphasis id="BA97EAECFFCBFFB8FF7AFC82F80EFC8E" box="[199,317,799,825]" italics="true" pageId="3" pageNumber="126">Mandibles</emphasis>
approximately symmetrical, external outline more or less parallel, almost semicircular apically (
<figureCitation id="10D82A7BFFCBFFB8FAD9FC82F996FCEA" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1939,1963]" captionTargetBox="[246,1347,190,1890]" captionTargetId="figure-26@2.[226,1362,176,1914]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus of Lethrus (Heteroplistodus) potanini species group. 1–2, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species (1 holotype, 2 allotype); 3–4, L. (H.) potanini Jakovlev, 1889 (3 male, Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011, 4 female, Inner Mongolia, Bayan Hot [Alxa], 2.vii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022198" httpUri="https://zenodo.org/record/14022198/files/figure.png" pageId="3" pageNumber="126">Figs. 1</figureCitation>
,
<figureCitation id="10D82A7BFFCBFFB8FF0FFCDEF9F2FCEA" box="[178,193,835,861]" captionStart="FIGURES 5–10" captionStartId="6.[151,264,1861,1885]" captionTargetBox="[169,1418,352,1825]" captionTargetId="figure-104@6.[156,1431,341,1837]" captionTargetPageId="6" captionText="FIGURES 5–10. Head of Lethrus (Heteroplistodus) potanini species group. 5–8 in lateral view. 5–6, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype (5 left, 6 right); 7–8, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011) (7 left, 8 right). 9–10 in dorsal view. 9, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 10, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022200" httpUri="https://zenodo.org/record/14022200/files/figure.png" pageId="3" pageNumber="126">9</figureCitation>
) with maximum width approximately at middle of mandibles length. Dorsal surface of mandible completely flat, without any slanting, transverse carina or hornlike protrusion. Ventral mandible processes (
<figureCitation id="10D82A7BFFCBFFB8FB26FCFAFC3BFC36" box="[1179,1288,870,897]" captionStart="FIGURES 5–10" captionStartId="6.[151,264,1861,1885]" captionTargetBox="[169,1418,352,1825]" captionTargetId="figure-104@6.[156,1431,341,1837]" captionTargetPageId="6" captionText="FIGURES 5–10. Head of Lethrus (Heteroplistodus) potanini species group. 5–8 in lateral view. 5–6, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype (5 left, 6 right); 7–8, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011) (7 left, 8 right). 9–10 in dorsal view. 9, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 10, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022200" httpUri="https://zenodo.org/record/14022200/files/figure.png" pageId="3" pageNumber="126">Figs. 5–6</figureCitation>
,
<figureCitation id="10D82A7BFFCBFFB8FAA9FCFAFC1CFC36" box="[1300,1327,871,897]" captionStart="FIGURES 11–12" captionStartId="7.[151,264,1633,1657]" captionTargetBox="[159,1429,203,1609]" captionTargetId="figure-22@7.[158,1429,174,1609]" captionTargetPageId="7" captionText="FIGURES 11–12. Head in frontal view of Lethrus (Heteroplistodus) potanini species group. 11, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 12, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022202" httpUri="https://zenodo.org/record/14022202/files/figure.png" pageId="3" pageNumber="126">11</figureCitation>
) strongly asymmetrical. Left process well developed, long and tapering, obliquely pointed without appendages or keels; directed from base of left mandible lower edge towards apical part of right mandible, not extending mandibular outline in dorsal view, subparallel with mandibles in frontal view. Right process strongly reduced, pointed, as short as thick basally.
</paragraph>
<paragraph id="885C36FEFFCBFFB8FF7AFB86FBCDFB72" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="126">
<emphasis id="BA97EAECFFCBFFB8FF7AFB86F805FB83" box="[199,310,1051,1076]" italics="true" pageId="3" pageNumber="126">Pronotum</emphasis>
(
<figureCitation id="10D82A7BFFCBFFB8FEF8FB86F8A0FB82" box="[325,403,1051,1077]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1939,1963]" captionTargetBox="[246,1347,190,1890]" captionTargetId="figure-26@2.[226,1362,176,1914]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus of Lethrus (Heteroplistodus) potanini species group. 1–2, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species (1 holotype, 2 allotype); 3–4, L. (H.) potanini Jakovlev, 1889 (3 male, Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011, 4 female, Inner Mongolia, Bayan Hot [Alxa], 2.vii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022198" httpUri="https://zenodo.org/record/14022198/files/figure.png" pageId="3" pageNumber="126">Figs. 1</figureCitation>
,
<figureCitation id="10D82A7BFFCBFFB8FE1CFB86F88DFB82" box="[417,446,1051,1077]" captionStart="FIGURES 17–20" captionStartId="9.[151,264,1908,1932]" captionTargetBox="[201,1388,193,1883]" captionTargetId="figure-22@9.[187,1401,180,1883]" captionTargetPageId="9" captionText="FIGURES 17–20. Lethrus (Heteroplistodus) potanini species group. 17–18 half of pronotum in dorsal view. 17, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 18, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3– 6.viii.2011). 19–20 aedeagus, apical part of parameres in dorsal view.19, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 20, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022206" httpUri="https://zenodo.org/record/14022206/files/figure.png" pageId="3" pageNumber="126">17</figureCitation>
) transverse, distinctly broader than base of elytra, broadest just at middle; lateral margin noticeable widened and entire weakly bordered, distinctly crenulate over whole length. Anterior angles broadly rounded, basal margin straight and finely bordered, anterior margin emarginate and entirely punctate. Punctation of dorsal surface coarse, deep and irregularly impressed, punctures separated by approximately their diameters discally; surface near lateral margins roughly punctate.
</paragraph>
<paragraph id="885C36FEFFCBFFB8FF7AFB52FABFFB5E" blockId="3.[151,1437,150,2013]" box="[199,908,1231,1257]" pageId="3" pageNumber="126">
<taxonomicName id="4FE34D7DFFCBFFB8FF7AFB52F806FB5E" box="[199,309,1231,1257]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="3" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="BA97EAECFFCBFFB8FF7AFB52F806FB5E" box="[199,309,1231,1257]" italics="true" pageId="3" pageNumber="126">Scutellum</emphasis>
</taxonomicName>
considerably short, almost as long as wide, glabrous.
</paragraph>
<paragraph id="885C36FEFFCBFFB8FF7AFB6EF8C6FACE" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="126">
<emphasis id="BA97EAECFFCBFFB8FF7AFB6EF83EFABA" box="[199,269,1267,1293]" italics="true" pageId="3" pageNumber="126">Elytra</emphasis>
(
<figureCitation id="10D82A7BFFCBFFB8FEA2FB6EF85CFABA" box="[287,367,1267,1293]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1939,1963]" captionTargetBox="[246,1347,190,1890]" captionTargetId="figure-26@2.[226,1362,176,1914]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus of Lethrus (Heteroplistodus) potanini species group. 1–2, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species (1 holotype, 2 allotype); 3–4, L. (H.) potanini Jakovlev, 1889 (3 male, Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011, 4 female, Inner Mongolia, Bayan Hot [Alxa], 2.vii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022198" httpUri="https://zenodo.org/record/14022198/files/figure.png" pageId="3" pageNumber="126">Figs. 1</figureCitation>
,
<figureCitation id="10D82A7BFFCBFFB8FE3DFB6FF8AEFABA" box="[384,413,1266,1293]" captionStart="FIGURES 13–16" captionStartId="8.[151,264,1904,1928]" captionTargetBox="[199,1366,180,1867]" captionTargetId="figure-26@8.[190,1397,177,1880]" captionTargetPageId="8" captionText="FIGURES 13–16. Lethrus (Heteroplistodus) potanini species group in dorsal view. 13–16 genae. 13, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 14, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011). 15–16 elytra. 15, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 16, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022204" httpUri="https://zenodo.org/record/14022204/files/figure.png" pageId="3" pageNumber="126">15</figureCitation>
) almost semicircular, apices not prominent, each apex forming independent semicircular tip. Striae visible, weakly indistinct and discontinuous in irregular wrinkled structure of surface. Lateral margin distinctly and broadly elevated in anterior quarter, strongly narrowed apicad. Epipleura distinctly and broadly elevated, reaching elytral apex.
</paragraph>
<paragraph id="885C36FEFFCBFFB8FF7AFA1EFBB4FA52" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="126">
<emphasis id="BA97EAECFFCBFFB8FF7AFA1EF9CFFA2B" box="[199,252,1411,1436]" italics="true" pageId="3" pageNumber="126">Legs</emphasis>
(
<figureCitation id="10D82A7BFFCBFFB8FEB1FA1EF86AFA2A" box="[268,345,1411,1437]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1939,1963]" captionTargetBox="[246,1347,190,1890]" captionTargetId="figure-26@2.[226,1362,176,1914]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus of Lethrus (Heteroplistodus) potanini species group. 1–2, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species (1 holotype, 2 allotype); 3–4, L. (H.) potanini Jakovlev, 1889 (3 male, Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011, 4 female, Inner Mongolia, Bayan Hot [Alxa], 2.vii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022198" httpUri="https://zenodo.org/record/14022198/files/figure.png" pageId="3" pageNumber="126">Figs. 1</figureCitation>
). Profemora with indistinct transversal edge anteroventrally; protibia with row of nine gradually proximad diminishing exterior denticles and with row of pointed or broadly tuberculate on ventromedial edge. Tip of apical spur of protibia bifurcate (
<figureCitation id="10D82A7BFFCBFFB8FD9BFA56FB49FA52" box="[550,634,1483,1509]" captionStart="FIGURES 21–24" captionStartId="10.[151,264,1907,1931]" captionTargetBox="[198,1389,182,1878]" captionTargetId="figure-26@10.[186,1402,174,1886]" captionTargetPageId="10" captionText="FIGURES 21–24. Lethrus (Heteroplistodus) potanini species group. 21–22 aedeagus, apical part of parameres in lateral view. 21, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 22, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011). 23–24 apical spur of protibia. 23, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 24, L. (H.) potanini (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022208" httpUri="https://zenodo.org/record/14022208/files/figure.png" pageId="3" pageNumber="126">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="885C36FEFFCBFFB8FF7AFA72FA4AF9BE" blockId="3.[151,1437,150,2013]" box="[199,889,1519,1545]" pageId="3" pageNumber="126">
<emphasis id="BA97EAECFFCBFFB8FF7AFA72F807F9BE" box="[199,308,1519,1545]" italics="true" pageId="3" pageNumber="126">Aedeagus</emphasis>
(
<figureCitation id="10D82A7BFFCBFFB8FEFEFA72F892F9BE" box="[323,417,1519,1545]" captionStart="FIGURES 17–20" captionStartId="9.[151,264,1908,1932]" captionTargetBox="[201,1388,193,1883]" captionTargetId="figure-22@9.[187,1401,180,1883]" captionTargetPageId="9" captionText="FIGURES 17–20. Lethrus (Heteroplistodus) potanini species group. 17–18 half of pronotum in dorsal view. 17, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 18, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3– 6.viii.2011). 19–20 aedeagus, apical part of parameres in dorsal view.19, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 20, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022206" httpUri="https://zenodo.org/record/14022206/files/figure.png" pageId="3" pageNumber="126">Figs. 19</figureCitation>
,
<figureCitation id="10D82A7BFFCBFFB8FE10FA72F8FFF9BE" box="[429,460,1519,1545]" captionStart="FIGURES 21–24" captionStartId="10.[151,264,1907,1931]" captionTargetBox="[198,1389,182,1878]" captionTargetId="figure-26@10.[186,1402,174,1886]" captionTargetPageId="10" captionText="FIGURES 21–24. Lethrus (Heteroplistodus) potanini species group. 21–22 aedeagus, apical part of parameres in lateral view. 21, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 22, L. (H.) potanini Jakovlev, 1889 (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011). 23–24 apical spur of protibia. 23, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species holotype; 24, L. (H.) potanini (Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022208" httpUri="https://zenodo.org/record/14022208/files/figure.png" pageId="3" pageNumber="126">21</figureCitation>
). Parameres broadly rounded apically.
</paragraph>
<paragraph id="885C36FEFFCBFFB8FF7AF98EFBE3F99A" blockId="3.[151,1437,150,2013]" box="[199,720,1554,1581]" pageId="3" pageNumber="126">
<emphasis id="BA97EAECFFCBFFB8FF7AF98EF84DF99A" bold="true" box="[199,382,1555,1581]" pageId="3" pageNumber="126">Measurements.</emphasis>
Total body length
<quantity id="4F1B9B1BFFCBFFB8FDEDF98FFBFFF99A" box="[592,716,1554,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="3" pageNumber="126" unit="mm" value="16.5" valueMax="18.0" valueMin="15.0">15–18 mm</quantity>
.
</paragraph>
<paragraph id="885C36FEFFCBFFB8FF7AF9ABFB68F9C2" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="126">
<emphasis id="BA97EAECFFCBFFB8FF7AF9ABF888F9E6" bold="true" box="[199,443,1590,1617]" pageId="3" pageNumber="126">Variability in males.</emphasis>
Left mandibular process in medium developed and underdeveloped (hypothelic) males shorter, right process minute or missing.
</paragraph>
<paragraph id="885C36FEFFCBFFB8FF7AF9E2FCAFF92E" blockId="3.[151,1437,150,2013]" box="[199,1436,1663,1689]" pageId="3" pageNumber="126">
<emphasis id="BA97EAECFFCBFFB8FF7AF9E2F82DF92E" bold="true" box="[199,286,1663,1689]" pageId="3" pageNumber="126">Female</emphasis>
(
<figureCitation id="10D82A7BFFCBFFB8FE93F9E2F842F92E" box="[302,369,1663,1689]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1939,1963]" captionTargetBox="[246,1347,190,1890]" captionTargetId="figure-26@2.[226,1362,176,1914]" captionTargetPageId="2" captionText="FIGURES 1–4. Dorsal habitus of Lethrus (Heteroplistodus) potanini species group. 1–2, L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species (1 holotype, 2 allotype); 3–4, L. (H.) potanini Jakovlev, 1889 (3 male, Inner Mongolia, Bayan Hot [Alxa], 3–6.viii.2011, 4 female, Inner Mongolia, Bayan Hot [Alxa], 2.vii.2011)." figureDoi="http://doi.org/10.5281/zenodo.14022198" httpUri="https://zenodo.org/record/14022198/files/figure.png" pageId="3" pageNumber="126">Fig. 2</figureCitation>
) differs from male only in the missing processes of mandibles; body length of
<typeStatus id="5758885CFFCBFFB8FB59F9E2FC0DF92E" box="[1252,1342,1663,1689]" pageId="3" pageNumber="126" type="allotype">allotype</typeStatus>
<quantity id="4F1B9B1BFFCBFFB8FAFBF9E2FCABF92E" box="[1350,1432,1663,1689]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="126" unit="mm" value="16.0">16 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C0F96575FFCBFFB8FF7AF93EF87CF956" pageId="3" pageNumber="126" type="etymology">
<paragraph id="885C36FEFFCBFFB8FF7AF93EF87CF956" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="126">
<emphasis id="BA97EAECFFCBFFB8FF7AF93EF87DF90A" bold="true" box="[199,334,1699,1725]" pageId="3" pageNumber="126">Etymology.</emphasis>
Patronymic; named in honor of Emil Kučera (Soběslav,
<collectingCountry id="F0F4766EFFCBFFB8FC72F93EFDB5F90A" box="[975,1158,1699,1725]" name="Czech Republic" pageId="3" pageNumber="126">Czech Republic</collectingCountry>
), collector of material of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C0F96575FFCBFFB8FF7AF977FD78F8B2" box="[199,1099,1770,1797]" pageId="3" pageNumber="126" type="distribution">
<paragraph id="885C36FEFFCBFFB8FF7AF977FD78F8B2" blockId="3.[151,1437,150,2013]" box="[199,1099,1770,1797]" pageId="3" pageNumber="126">
<emphasis id="BA97EAECFFCBFFB8FF7AF977F852F8B3" bold="true" box="[199,353,1770,1796]" pageId="3" pageNumber="126">Distribution.</emphasis>
Known from the
<collectingRegion id="4A27F81CFFCBFFB8FD95F976FBE4F8B2" box="[552,727,1771,1797]" country="China" name="Nei Mongol" pageId="3" pageNumber="126">Inner Mongolia</collectingRegion>
and
<collectingRegion id="4A27F81CFFCBFFB8FCB0F976FA54F8B2" box="[781,871,1771,1797]" country="China" name="Shaanxi" pageId="3" pageNumber="126">Shaanxi</collectingRegion>
provinces of
<collectingCountry id="F0F4766EFFCBFFB8FBBDF976FD74F8B2" box="[1024,1095,1771,1797]" name="China" pageId="3" pageNumber="126">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C0F96575FFCBFFBFFF7AF893FD0DFEAA" lastPageId="4" lastPageNumber="127" pageId="3" pageNumber="126" type="diagnosis">
<paragraph id="885C36FEFFCBFFBFFF7AF893FD0DFEAA" blockId="3.[151,1437,150,2013]" lastBlockId="4.[151,1437,151,285]" lastPageId="4" lastPageNumber="127" pageId="3" pageNumber="126">
<emphasis id="BA97EAECFFCBFFB8FF7AF893F8FFF89F" bold="true" box="[199,460,1806,1832]" pageId="3" pageNumber="126">Differential diagnosis.</emphasis>
<taxonomicName id="4FE34D7DFFCBFFB8FE6CF892FD59F89E" authority="Hillert, Kral &amp; Bagaturov" authorityName=", Hillert, Kral &amp; Bagaturov" authorityYear="2024" box="[465,1130,1807,1833]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="kucerai" status="sp. nov." subGenus="Heteroplistodus">
<emphasis id="BA97EAECFFCBFFB8FE6CF892FB14F89E" box="[465,551,1807,1833]" italics="true" pageId="3" pageNumber="126">Lethrus</emphasis>
(
<emphasis id="BA97EAECFFCBFFB8FD88F892FBD4F89E" box="[565,743,1807,1833]" italics="true" pageId="3" pageNumber="126">Heteroplistodus</emphasis>
)
<emphasis id="BA97EAECFFCBFFB8FD48F892FA7AF89E" box="[757,841,1807,1833]" italics="true" pageId="3" pageNumber="126">kucerai</emphasis>
Hillert, Král &amp; Bagaturov
</taxonomicName>
,
<emphasis id="BA97EAECFFCBFFB8FBCEF892FDC8F89F" bold="true" box="[1139,1275,1806,1833]" pageId="3" pageNumber="126">
<taxonomicNameLabel id="A1A45797FFCBFFB8FBCEF892FDC8F89F" box="[1139,1275,1806,1833]" pageId="3" pageNumber="126" rank="species">new species</taxonomicNameLabel>
</emphasis>
is classified in the
<taxonomicName id="4FE34D7DFFCBFFB8FF02F8AEF85EF8FB" authorityName="Jakovlev" authorityYear="1889" box="[191,365,1843,1869]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="potanini" subGenus="Heteroplistodus">
<emphasis id="BA97EAECFFCBFFB8FF02F8AEF9E5F8FB" box="[191,214,1843,1868]" italics="true" pageId="3" pageNumber="126">L.</emphasis>
(
<emphasis id="BA97EAECFFCBFFB8FF58F8AEF832F8FB" box="[229,257,1843,1868]" italics="true" pageId="3" pageNumber="126">H.</emphasis>
)
<emphasis id="BA97EAECFFCBFFB8FEADF8AEF85EF8FB" box="[272,365,1843,1868]" italics="true" pageId="3" pageNumber="126">potanini</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="EC724B0FFFCBFFB8FDA5F8AEFA6CF8FA" author="Bagaturov, M. F. &amp; Nikolajev, G. V." box="[536,863,1842,1869]" pageId="3" pageNumber="126" pagination="303 - 314" refId="ref8675" refString="Bagaturov, M. F. &amp; Nikolajev, G. V. (2015) Obzor rasprostraneniya zhukov-kravchikov roda Lethrus, 1777 (Coleoptera: Geotrupidae). Overview of distribution of the genus Lethrus Scopoli, 1777 (Coleoptera: Geotrupidae). Kavkazskiy Entomologicheskiy Byulleten, 11, 303 - 314. [in Russian, English abstract] https: // doi. org / 10.23885 / 1814 - 3326 - 2015 - 11 - 2 - 303 - 314" type="journal article" year="2015">Bagaturov &amp; Nikolajev 2015</bibRefCitation>
) due to the unique shape of mandibular appendices, which is strongly asymmetrical, left long and thin, obliquely pointed and located transversally under the mandibles (cf.
<bibRefCitation id="EC724B0FFFCBFFB8FF7AF8E6F894F822" author="Nikolajev, G. V." box="[199,423,1914,1941]" pageId="3" pageNumber="126" refId="ref9820" refString="Nikolajev, G. V. (2003 b) Zhuki-kravchiki (Scarabaeidae, Geotrupinae, Lethrini): biologiya, sistematika, rasprostranenie, opredelitel'. Lethrini (Coleoptera: Scarabaeidae, Geotrupinae): biology, systematics, distribution, keys. Kazak universiteti, Almaty, 254 pp. [in Russian, English summary]" type="book" year="2003">Nikolajev 2003: 75</bibRefCitation>
, figs. 45: 2–4). The species group comprises two species only,
<taxonomicName id="4FE34D7DFFCBFFB8FB35F8E6FC0DF823" authorityName="Jakovlev" authorityYear="1889" box="[1160,1342,1915,1941]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="potanini" subGenus="Heteroplistodus">
<emphasis id="BA97EAECFFCBFFB8FB35F8E6FDACF823" box="[1160,1183,1915,1940]" italics="true" pageId="3" pageNumber="126">L.</emphasis>
(
<emphasis id="BA97EAECFFCBFFB8FB0FF8E6FDFDF823" box="[1202,1230,1915,1940]" italics="true" pageId="3" pageNumber="126">H.</emphasis>
)
<emphasis id="BA97EAECFFCBFFB8FB5CF8E6FC0DF823" box="[1249,1342,1915,1940]" italics="true" pageId="3" pageNumber="126">potanini</emphasis>
</taxonomicName>
and the newly described species
<taxonomicName id="4FE34D7DFFCBFFB8FE13F802FAB3F80E" authority="Hillert, Kral &amp; Bagaturov" authorityName=", Hillert, Kral &amp; Bagaturov" authorityYear="2024" box="[430,896,1951,1977]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="kucerai" status="new species" subGenus="Heteroplistodus">
<emphasis id="BA97EAECFFCBFFB8FE13F802F8F6F80F" box="[430,453,1951,1976]" italics="true" pageId="3" pageNumber="126">L.</emphasis>
(
<emphasis id="BA97EAECFFCBFFB8FE68F802F8C2F80F" box="[469,497,1951,1976]" italics="true" pageId="3" pageNumber="126">H.</emphasis>
)
<emphasis id="BA97EAECFFCBFFB8FDBCF802FB66F80E" box="[513,597,1951,1977]" italics="true" pageId="3" pageNumber="126">kucerai</emphasis>
Hillert, Král &amp; Bagaturov
</taxonomicName>
,
<emphasis id="BA97EAECFFCBFFB8FC36F802FD26F80F" bold="true" box="[907,1045,1950,1977]" pageId="3" pageNumber="126">
<taxonomicNameLabel id="A1A45797FFCBFFB8FC36F802FD26F80F" box="[907,1045,1950,1977]" pageId="3" pageNumber="126" rank="species">new species</taxonomicNameLabel>
</emphasis>
. Both species are morphologically similar, for the differential diagnosis see character matrix on
<tableCitation id="C5610345FFCBFFB8FCF7F85EFAADF86A" box="[842,926,1987,2013]" captionStart="TABLE 1" captionStartId="13.[152,244,260,284]" captionText="TABLE 1. Character matrix for the differential diagnosis of males of the Lethrus (Heteroplistodus) potanini species" pageId="3" pageNumber="126">Table 1</tableCitation>
. The distribution areas seem to be allopatric, close to each other (see map on
<figureCitation id="10D82A7BFFCCFFBFFDBCFF0AFB67FF06" box="[513,596,151,177]" captionStart="FIGURE 29" captionStartId="12.[151,250,1164,1188]" captionTargetBox="[152,1436,177,1140]" captionTargetId="figure-26@12.[152,1436,177,1143]" captionTargetPageId="12" captionText="FIGURE 29. Map of distribution area of Lethrus (Heteroplistodus) potanini species group. Black circles represent records of L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species, red circles represent records of L. (H.) potanini Jakovlev, 1889, empty circles are type localities, The type locality of Lethrus conformis Frivaldszky, 1890 [= Lethrus (Heteroplistodus) potanini Jakovlev, 1889] is possibly a road leading from Shuozhou to Zhangye and is marked with a line (see the text, for details)." figureDoi="http://doi.org/10.5281/zenodo.14022212" httpUri="https://zenodo.org/record/14022212/files/figure.png" pageId="4" pageNumber="127">Fig. 29</figureCitation>
). Both species are distributed in the northcentral regions of
<collectingCountry id="F0F4766EFFCCFFBFFB48FF0AFC0FFF06" box="[1269,1340,151,177]" name="China" pageId="4" pageNumber="127">China</collectingCountry>
.
<taxonomicName id="4FE34D7DFFCCFFBFFAFAFF0AFB61FF62" authority="Hillert, Kral &amp; Bagaturov" authorityName=", Hillert, Kral &amp; Bagaturov" authorityYear="2024" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="species" species="kucerai" status="sp. nov." subGenus="Heteroplistodus">
<emphasis id="BA97EAECFFCCFFBFFAFAFF0AFCAEFF06" box="[1351,1437,151,177]" italics="true" pageId="4" pageNumber="127">Lethrus</emphasis>
(
<emphasis id="BA97EAECFFCCFFBFFF1CFF26F98FFF63" box="[161,188,187,212]" italics="true" pageId="4" pageNumber="127">H.</emphasis>
)
<emphasis id="BA97EAECFFCCFFBFFF71FF26F813FF62" box="[204,288,187,213]" italics="true" pageId="4" pageNumber="127">kucerai</emphasis>
, Hillert, Král &amp; Bagaturov
</taxonomicName>
,
<emphasis id="BA97EAECFFCCFFBFFDE0FF26FBD5FF63" bold="true" box="[605,742,186,213]" pageId="4" pageNumber="127">
<taxonomicNameLabel id="A1A45797FFCCFFBFFDE0FF26FBD5FF63" box="[605,742,186,213]" pageId="4" pageNumber="127" rank="species">new species</taxonomicNameLabel>
</emphasis>
is known from several localities in northwestern
<collectingRegion id="4A27F81CFFCCFFBFFAAFFF26FC5FFF62" box="[1298,1388,187,213]" country="China" name="Shaanxi" pageId="4" pageNumber="127">Shaanxi</collectingRegion>
and south of
<collectingRegion id="4A27F81CFFCCFFBFFF47FF42F899FF4E" box="[250,426,223,249]" country="China" name="Nei Mongol" pageId="4" pageNumber="127">Inner Mongolia</collectingRegion>
near the border to
<collectingRegion id="4A27F81CFFCCFFBFFD3CFF42FBECFF4E" box="[641,735,223,249]" country="China" name="Shaanxi" pageId="4" pageNumber="127">Shaanxi</collectingRegion>
.
<taxonomicName id="4FE34D7DFFCCFFBFFD54FF42FA0CFF4E" authorityName="Scopoli" authorityYear="1777" box="[745,831,223,249]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="BA97EAECFFCCFFBFFD54FF42FA0CFF4E" box="[745,831,223,249]" italics="true" pageId="4" pageNumber="127">Lethrus</emphasis>
</taxonomicName>
<taxonomicName id="4FE34D7DFFCCFFBFFCFBFF42FAEAFF4F" authorityName="Jakovlev" authorityYear="1889" box="[838,985,223,249]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="127" phylum="Arthropoda" rank="species" species="potanini" subGenus="Heteroplistodus">
(
<emphasis id="BA97EAECFFCCFFBFFCEDFF42FA57FF4F" box="[848,868,223,248]" italics="true" pageId="4" pageNumber="127">H</emphasis>
.)
<emphasis id="BA97EAECFFCCFFBFFCC1FF42FAEAFF4F" box="[892,985,223,248]" italics="true" pageId="4" pageNumber="127">potanini</emphasis>
</taxonomicName>
is distributed in southern
<collectingCountry id="F0F4766EFFCCFFBFFB42FF42FC5FFF4E" box="[1279,1388,223,249]" name="Mongolia" pageId="4" pageNumber="127">Mongolia</collectingCountry>
and the Chinese provinces of Gansu, Inner Mongolia and Ningxia (see map on
<figureCitation id="10D82A7BFFCCFFBFFC63FE9EFD01FEAA" box="[990,1074,259,285]" captionStart="FIGURE 29" captionStartId="12.[151,250,1164,1188]" captionTargetBox="[152,1436,177,1140]" captionTargetId="figure-26@12.[152,1436,177,1143]" captionTargetPageId="12" captionText="FIGURE 29. Map of distribution area of Lethrus (Heteroplistodus) potanini species group. Black circles represent records of L. (H.) kucerai Hillert, Král &amp; Bagaturov, new species, red circles represent records of L. (H.) potanini Jakovlev, 1889, empty circles are type localities, The type locality of Lethrus conformis Frivaldszky, 1890 [= Lethrus (Heteroplistodus) potanini Jakovlev, 1889] is possibly a road leading from Shuozhou to Zhangye and is marked with a line (see the text, for details)." figureDoi="http://doi.org/10.5281/zenodo.14022212" httpUri="https://zenodo.org/record/14022212/files/figure.png" pageId="4" pageNumber="127">Fig. 29</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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