treatments-xml/data/FF/67/FB/FF67FB0AFF8AC51FFF1601979E7FBE89.xml

<document id="ECF811ECF7DC1968545A75138A5BCF01" ID-CLB-Dataset="3378" ID-DOI="10.11646/zootaxa.5051.1.11" ID-GBIF-Dataset="90f8cfa4-8adf-42e7-b8ca-4acf39b7fc81" ID-ISSN="1175-5326" ID-Zenodo-Dep="5563475" ID-ZooBank="82B4BCC0-BE67-4BC2-BE3E-68D39F72CEEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634021034485" checkinUser="plazi" docAuthor="Gómez, Samuel" docDate="2021" docId="FF67FB0AFF8AC51FFF1601979E7FBE89" docLanguage="en" docName="zootaxa.5051.1.11.pdf" docOrigin="Zootaxa 5051 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudostenhelia bathyalis Gómez 2021, sp. nov." docType="treatment" docUuid="D86B0811-F015-43F6-BF3A-9397B4280D20" docUuidSource="ZooBank" docVersion="9" lastPageNumber="160" masterDocId="035E8372FF88C516FF81033A9A6ABB6C" masterDocTitle="On some deep-sea Stenheliinae from the Gulf of California and the west coast of the Baja California Peninsula (Mexico): Pseudostenhelia bathyalis sp. nov., and Beatricella calidafornax sp. nov. (Copepoda: Harpacticoida: Miraciidae)" masterLastPageNumber="170" masterPageNumber="151" pageNumber="153" updateTime="1699459449544" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="50B629E58AA316F9DC64185C945C7EAA">On some deep-sea Stenheliinae from the Gulf of California and the west coast of the Baja California Peninsula (Mexico): Pseudostenhelia bathyalis sp. nov., and Beatricella calidafornax sp. nov. (Copepoda: Harpacticoida: Miraciidae)</mods:title>
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<mods:namePart id="9E18E2A08DC05A127935DA07F720C033">Gómez, Samuel</mods:namePart>
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<treatment id="FF67FB0AFF8AC51FFF1601979E7FBE89" ID-DOI="http://doi.org/10.5281/zenodo.5715369" ID-GBIF-Taxon="190317046" ID-Zenodo-Dep="5715369" LSID="urn:lsid:zoobank.org:act:D86B0811-F015-43F6-BF3A-9397B4280D20" httpUri="http://treatment.plazi.org/id/FF67FB0AFF8AC51FFF1601979E7FBE89" lastPageId="9" lastPageNumber="160" pageId="2" pageNumber="153">
<subSubSection id="3FD41997FF8AC514FF160197985CB9A4" box="[151,566,685,712]" pageId="2" pageNumber="153" type="nomenclature">
<paragraph id="77714A1CFF8AC514FF160197985CB9A4" blockId="2.[151,953,685,785]" box="[151,566,685,712]" pageId="2" pageNumber="153">
<heading id="2C39FD70FF8AC514FF160197985CB9A4" bold="true" box="[151,566,685,712]" fontSize="11" level="1" pageId="2" pageNumber="153" reason="1">
<emphasis id="45BA960EFF8AC514FF160197985CB9A4" bold="true" box="[151,566,685,712]" pageId="2" pageNumber="153">
<taxonomicName id="B0CE319FFF8AC514FF1601979BBBB9A4" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[151,465,685,712]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="bathyalis" status="sp. nov.">
<emphasis id="45BA960EFF8AC514FF1601979BBBB9A4" bold="true" box="[151,465,685,712]" italics="true" pageId="2" pageNumber="153">Pseudostenhelia bathyalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5E892B75FF8AC514FE590194985CB9A4" box="[472,566,686,712]" pageId="2" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="3FD41997FF8AC514FF1601E899D3B87D" pageId="2" pageNumber="153" type="description">
<paragraph id="77714A1CFF8AC514FF1601E89B78B981" blockId="2.[151,953,685,785]" box="[151,274,722,749]" pageId="2" pageNumber="153">
(
<figureCitation id="EFF55699FF8AC514FF1E01E89B60B981" box="[159,266,722,749]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="3.[151,250,1943,1967]" captionStartId-1="4.[151,250,1928,1952]" captionStartId-2="5.[151,250,1930,1954]" captionStartId-3="7.[151,250,1921,1945]" captionStartId-4="8.[151,250,1904,1928]" captionTargetBox-0="[267,1320,190,1911]" captionTargetBox-1="[159,1424,352,1893]" captionTargetBox-2="[160,1433,604,1899]" captionTargetBox-3="[195,1390,497,1889]" captionTargetBox-4="[169,1414,190,1872]" captionTargetId-0="figure-15@3.[259,1329,181,1919]" captionTargetId-1="figure-95@4.[151,1436,345,1904]" captionTargetId-2="figure-198@5.[151,1436,597,1907]" captionTargetId-3="figure-178@7.[187,1401,489,1898]" captionTargetId-4="figure-20@8.[163,1423,181,1880]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." captionText-1="FIGURE 3. Pseudostenhelia bathyalis sp. nov., female: A, rostrum and antennule (microphotograph available in http:// metadata.icmyl.unam.mx/handle/20.500.12201/10578); B, antenna (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10569)." captionText-2="FIGURE 4. Pseudostenhelia bathyalissp. nov., female:A, mandible; B, coxa, basis, endopod and exopodof maxillule; C, maxilla (microphotograph available ion http://metadata.icmyl.unam.mx/handle/20.500.12201/10571); D, maxilliped (microphotograph available in http://metadata.icmyl.unam.mx/handle/20.500.12201/10572)." captionText-3="FIGURE 5. Pseudostenhelia bathyalis sp. nov., female: A, P1, anterior (microphotograph available in http://metadata. icmyl.unam.mx/handle/20.500.12201/10573); B, P2, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10574)." captionText-4="FIGURE 6. Pseudostenhelia bathyalis sp. nov., female: A, P3, anterior (microphotograph available in http://metadata. icmyl.unam.mx/handle/20.500.12201/10575); B, P4, anterior (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10576); C, P5, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10577)." figureDoi-0="http://doi.org/10.5281/zenodo.5563481" figureDoi-1="http://doi.org/10.5281/zenodo.5563485" figureDoi-2="http://doi.org/10.5281/zenodo.5563487" figureDoi-3="http://doi.org/10.5281/zenodo.5563489" figureDoi-4="http://doi.org/10.5281/zenodo.5563491" httpUri-0="https://zenodo.org/record/5563481/files/figure.png" httpUri-1="https://zenodo.org/record/5563485/files/figure.png" httpUri-2="https://zenodo.org/record/5563487/files/figure.png" httpUri-3="https://zenodo.org/record/5563489/files/figure.png" httpUri-4="https://zenodo.org/record/5563491/files/figure.png" pageId="2" pageNumber="153">Figs. 2–6</figureCitation>
)
</paragraph>
<paragraph id="77714A1CFF8AC514FF1601CC99D3B87D" blockId="2.[151,953,685,785]" box="[151,953,758,785]" pageId="2" pageNumber="153">
<uri id="035F461EFF8AC514FF1601CC99D3B87D" box="[151,953,758,785]" pageId="2" pageNumber="153">
urn:lsid:zoobank.org:act: 
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</paragraph>
</subSubSection>
<subSubSection id="3FD41997FF8AC514FF1600049F63B884" pageId="2" pageNumber="153" type="materials_examined">
<paragraph id="77714A1CFF8AC514FF1600049967B8CC" blockId="2.[151,1437,829,2008]" pageId="2" pageNumber="153">
<materialsCitation id="C7A64041FF8AC514FF1600049898B8CC" ID-GBIF-Occurrence="3409042301" country="Mexico" county="Talud" elevation="1752" latitude="29.81333" location="Southeastern San Quintin Basin" longLatPrecision="1" longitude="-116.12166" municipality="Bahia del Rosario" pageId="2" pageNumber="153" specimenCount="1" stateProvince="Baja California" typeStatus="holotype">
<emphasis id="45BA960EFF8AC514FF1600049B5FB83B" bold="true" box="[151,309,829,856]" pageId="2" pageNumber="153">
<typeStatus id="A875F4BEFF8AC514FF1600049AB8B834" box="[151,210,830,856]" pageId="2" pageNumber="153">Type</typeStatus>
locality.
</emphasis>
<location id="72111CC7FF8AC514FEBA000498CBB834" LSID="urn:lsid:plazi:treatment:FF67FB0AFF8AC51FFF1601979E7FBE89:72111CC7FF8AC514FEBA000498CBB834" box="[315,673,830,856]" country="Mexico" county="Talud" latitude="29.81333" longLatPrecision="1" longitude="-116.12166" municipality="Bahia del Rosario" name="Southeastern San Quintin Basin" pageId="2" pageNumber="153" stateProvince="Baja California">Southeastern San Quintin Basin</location>
, off 
<collectingMunicipality id="9715D066FF8AC514FD53000499F6B834" box="[722,924,830,856]" pageId="2" pageNumber="153">Bahia del Rosario</collectingMunicipality>
, 
<collectingRegion id="B50A84FEFF8AC514FC2700049E3EB834" box="[934,1108,830,856]" country="Mexico" name="Baja California" pageId="2" pageNumber="153">Baja California</collectingRegion>
, 
<collectingCountry id="0FD90A8CFF8AC514FBDE00049ED2B834" box="[1119,1208,830,856]" name="Mexico" pageId="2" pageNumber="153">Mexico</collectingCountry>
; 
<collectingCounty id="9E103290FF8AC514FB4300049F6BB834" box="[1218,1281,830,856]" pageId="2" pageNumber="153">Talud</collectingCounty>
XVIB cruise, sampling station 16 (
<geoCoordinate id="12FA2CDBFF8AC514FE0400589878B810" box="[389,530,866,892]" degrees="29.81333" direction="north" orientation="latitude" pageId="2" pageNumber="153" precision="1" value="29.81333">29.81333°N</geoCoordinate>
, 
<geoCoordinate id="12FA2CDBFF8AC514FD9E005898D5B811" box="[543,703,866,893]" degrees="116.12166" direction="west" orientation="longitude" pageId="2" pageNumber="153" precision="1" value="-116.12166">116.12166°W</geoCoordinate>
); depth, 
<quantity id="B036E7F9FF8AC514FCA0005899EBB810" box="[801,897,866,893]" metricMagnitude="3" metricUnit="m" metricValue="1.752" pageId="2" pageNumber="153" unit="m" value="1752.0">
<elevation id="FCE3AD2FFF8AC514FCA0005899EBB810" box="[801,897,866,893]" metricMagnitude="3" metricUnit="m" metricValue="1.752" pageId="2" pageNumber="153" unit="m" value="1752.0">1,752 m</elevation>
</quantity>
; organic carbon content, 4.08%; organic matter content, 7.01%; sand 5.45%; clay, 11.94%; silt, 82.61
</materialsCitation>
%.
</paragraph>
<paragraph id="77714A1CFF8AC514FF4600939F63B884" blockId="2.[151,1437,829,2008]" pageId="2" pageNumber="153">
<emphasis id="45BA960EFF8AC514FF4600939BA1B8AF" bold="true" box="[199,459,937,963]" pageId="2" pageNumber="153">Specimens examined.</emphasis>
Female 
<typeStatus id="A875F4BEFF8AC514FDB4009098FDB8A8" box="[565,663,938,964]" pageId="2" pageNumber="153" type="holotype">holotype</typeStatus>
(EMUCOP-290514-01) dissected and mounted onto seven slides; Talud XVIB cruise, sampling station 16 (
<geoCoordinate id="12FA2CDBFF8AC514FDE500F4989BB884" box="[612,753,974,1000]" degrees="29.81333" direction="north" orientation="latitude" pageId="2" pageNumber="153" precision="1" value="29.81333">29.81333°N</geoCoordinate>
, 
<geoCoordinate id="12FA2CDBFF8AC514FD7D00F499F7B885" box="[764,925,974,1001]" degrees="116.12166" direction="west" orientation="longitude" pageId="2" pageNumber="153" precision="1" value="-116.12166">116.12166°W</geoCoordinate>
); 
<date id="03706CDCFF8AC514FC3100F49E3AB884" box="[944,1104,974,1000]" pageId="2" pageNumber="153" value="2014-05-29">May 29, 2014</date>
; coll. S. Gómez.
</paragraph>
</subSubSection>
<subSubSection id="3FD41997FF8AC514FF4600C89BFEBF5C" pageId="2" pageNumber="153" type="etymology">
<paragraph id="77714A1CFF8AC514FF4600C89BFEBF5C" blockId="2.[151,1437,829,2008]" pageId="2" pageNumber="153">
<emphasis id="45BA960EFF8AC514FF4600C89B24BF60" bold="true" box="[199,334,1010,1036]" pageId="2" pageNumber="153">Etymology.</emphasis>
The specific epithet 
<taxonomicName id="B0CE319FFF8AC514FDB800C898CABF60" authorityName="Gómez" authorityYear="2021" box="[569,672,1010,1036]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="bathyalis">
<emphasis id="45BA960EFF8AC514FDB800C898CABF60" box="[569,672,1010,1036]" italics="true" pageId="2" pageNumber="153">bathyalis</emphasis>
</taxonomicName>
from the Greek βαθύς (bathys), deep, refers the environment where the species was found.
</paragraph>
</subSubSection>
<subSubSection id="3FD41997FF8AC510FF4607039BE2BCAB" lastPageId="6" lastPageNumber="157" pageId="2" pageNumber="153" type="description">
<paragraph id="77714A1CFF8AC514FF4607039B25BFF0" blockId="2.[151,1437,829,2008]" pageId="2" pageNumber="153">
<emphasis id="45BA960EFF8AC514FF4607039BA7BF3F" bold="true" box="[199,461,1081,1107]" pageId="2" pageNumber="153">Description of female.</emphasis>
Total body length measured from tip of rostrum to posterior margin of caudal rami 620 µm. Habitus pyriform, widest at posterior end of cephalothorax, tapering posteriad (
<figureCitation id="EFF55699FF8AC514FBCC07649EC3BF14" box="[1101,1193,1118,1144]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2A</figureCitation>
); cephalothorax/body length ratio, 0.3.
</paragraph>
<paragraph id="77714A1CFF8AC514FF46079C9F42BE3C" blockId="2.[151,1437,829,2008]" pageId="2" pageNumber="153">
Prosome consisting of cephalothorax with fused first pedigerous somite, and second to fourth free pedigerous somites. Cephalothorax (
<figureCitation id="EFF55699FF8AC514FE3707F0987BBF88" box="[438,529,1226,1252]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2A</figureCitation>
) slightly wider than long, no surface sensilla detected, hyaline fringe smooth, surface covered by small depressions. Free pedigerous somites without expansions laterally nor dorsally, without spinular ornamentation, no surface sensilla detected, integument covered by small depressions, weakly sclerotized, hyaline fringe smooth, width of second to fourth pedigerous somites decreasing progressively (
<figureCitation id="EFF55699FF8AC514FB41060C9F71BE3C" box="[1216,1307,1334,1360]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="77714A1CFF8AC514FF4606609F62BED0" blockId="2.[151,1437,829,2008]" pageId="2" pageNumber="153">
Urosome consisting of fifth pedigerous somite (first urosomite), genital double-somite (genital—second urosomite—and third urosomites fused), two free urosomites, and anal somite; urosomites without expansions laterally nor dorsally, integument weakly sclerotized, surface covered by small depressions (
<figureCitation id="EFF55699FF8AC514FB2006989E91BED0" box="[1185,1275,1442,1468]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="77714A1CFF8AC514FF4606FC9A86BD68" blockId="2.[151,1437,829,2008]" pageId="2" pageNumber="153">
Fifth pedigerous somite without spinular ornamentation, seemingly without sensilla (
<figureCitation id="EFF55699FF8AC514FB0C06FC9E83BE8C" box="[1165,1257,1478,1504]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2A</figureCitation>
), hyaline fringe narrow.
</paragraph>
<paragraph id="77714A1CFF8AC514FF46053498B3BD1C" blockId="2.[151,1437,829,2008]" pageId="2" pageNumber="153">
Second and third urosomites completely fused forming genital double-somite, without any trace of division, slightly longer than wide, widest part measured in the middle, no surface sensilla nor spinular ornamentation detected, posterior hyaline fringe smooth (
<figureCitation id="EFF55699FF8AC514FDF3056C98A6BD1C" box="[626,716,1622,1648]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="77714A1CFF8AC514FF4605409989BDD4" blockId="2.[151,1437,829,2008]" pageId="2" pageNumber="153">
Fourth and fifth urosomites seemingly without surface sensilla dorsally (
<figureCitation id="EFF55699FF8AC514FB9A05409E12BDF8" box="[1051,1144,1658,1684]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2A</figureCitation>
) nor ventrally (
<figureCitation id="EFF55699FF8AC514FAB205409FE5BDF8" box="[1331,1423,1658,1684]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2C</figureCitation>
), dorsally without (
<figureCitation id="EFF55699FF8AC514FEDC05A49BDDBDD4" box="[349,439,1694,1720]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2A</figureCitation>
), ventrally with medial row of spinules (
<figureCitation id="EFF55699FF8AC514FCFF05A499BCBDD4" box="[894,982,1694,1720]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="77714A1CFF8AC514FF4605F89F3BBC6C" blockId="2.[151,1437,829,2008]" pageId="2" pageNumber="153">
Anal somite 1.7 times as wide as long (
<figureCitation id="EFF55699FF8AC514FD0D05F89963BDB0" box="[652,777,1730,1756]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2A, C</figureCitation>
), with lateroventral spinules close to joint of caudal rami, seemingly without sensilla, with two lateral and two ventral pores (
<figureCitation id="EFF55699FF8AC514FC0905DC998ABC6C" box="[904,992,1766,1792]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2C</figureCitation>
); anal operculum inconspicuous.
</paragraph>
<paragraph id="77714A1CFF8AC514FF4604309927BCB4" blockId="2.[151,1437,829,2008]" pageId="2" pageNumber="153">
Caudal rami elongate, about five times as long as wide (
<figureCitation id="EFF55699FF8AC514FCB4043099C7BC48" box="[821,941,1802,1828]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2A–C</figureCitation>
) and about two times as long as anal somite, slightly divergent, each ramus with one subdistal pore ventrally, with dorsal spinules close to seta II, and on inner margin subdistally, with seven elements (
<figureCitation id="EFF55699FF8AC514FDE5046898B4BC00" box="[612,734,1874,1900]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2B–C</figureCitation>
); seta I and II arising subdistally, the former ventral to and half the length of the latter; seta III subdistal, arising ventrally (
<figureCitation id="EFF55699FF8AC514FCAE044C99E2BCFC" box="[815,904,1910,1936]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2B</figureCitation>
; lost in 
<figureCitation id="EFF55699FF8AC514FC60044C9E50BCFC" box="[993,1082,1910,1936]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[267,1320,190,1911]" captionTargetId="figure-15@3.[259,1329,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Pseudostenhelia bathyalis sp. nov., female:A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral." figureDoi="http://doi.org/10.5281/zenodo.5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="2" pageNumber="153">Fig. 2C</figureCitation>
); seta IV and V arising distally, both with fracture plane and rat-like, the former shorter; seta VI issuing at inner distal corner; dorsal seta VII triarticulate at base, situated subdistally close to inner margin.
</paragraph>
<caption id="23B11A94FF8BC515FF1604AD9F0EBCB8" ID-DOI="http://doi.org/10.5281/zenodo.5563481" ID-Zenodo-Dep="5563481" httpUri="https://zenodo.org/record/5563481/files/figure.png" pageId="3" pageNumber="154" startId="3.[151,250,1943,1967]" targetBox="[267,1320,190,1911]" targetPageId="3">
<paragraph id="77714A1CFF8BC515FF1604AD9F0EBCB8" blockId="3.[151,1437,1943,2004]" pageId="3" pageNumber="154">
<emphasis id="45BA960EFF8BC515FF1604AD9B78BCC3" bold="true" box="[151,274,1943,1967]" pageId="3" pageNumber="154">FIGURE 2.</emphasis>
<taxonomicName id="B0CE319FFF8BC515FE9904A2984ABCDC" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[280,544,1944,1968]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="bathyalis" status="sp. nov.">
<emphasis id="45BA960EFF8BC515FE9904A2984ABCDC" box="[280,544,1944,1968]" italics="true" pageId="3" pageNumber="154">Pseudostenhelia bathyalis</emphasis>
</taxonomicName>
<emphasis id="45BA960EFF8BC515FDA404A2981FBCDC" bold="true" box="[549,629,1944,1968]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="5E892B75FF8BC515FDA404A2981FBCDC" box="[549,629,1944,1968]" pageId="3" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A, habitus, dorsal (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10570); B, caudal rami, dorsal; C, fourth, fifth, and anal somite, and caudal rami, ventral.
</paragraph>
</caption>
<paragraph id="77714A1CFF8CC512FF4603AD9FF0BBDD" blockId="4.[151,1436,151,321]" box="[199,1434,151,177]" pageId="4" pageNumber="155">
Rostrum (
<figureCitation id="EFF55699FF8CC512FEB403AD9BE0BBDD" box="[309,394,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,1928,1952]" captionTargetBox="[159,1424,352,1893]" captionTargetId="figure-95@4.[151,1436,345,1904]" captionTargetPageId="4" captionText="FIGURE 3. Pseudostenhelia bathyalis sp. nov., female: A, rostrum and antennule (microphotograph available in http:// metadata.icmyl.unam.mx/handle/20.500.12201/10578); B, antenna (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10569)." figureDoi="http://doi.org/10.5281/zenodo.5563485" httpUri="https://zenodo.org/record/5563485/files/figure.png" pageId="4" pageNumber="155">Fig. 3A</figureCitation>
) trapezoidal, not fused to cephalothorax, slightly bifid, subdistal sensilla lost during dissection.
</paragraph>
<paragraph id="77714A1CFF8CC512FF460381985ABA2D" blockId="4.[151,1436,151,321]" pageId="4" pageNumber="155">
Antennule (
<figureCitation id="EFF55699FF8CC512FECF03819BC2BBB9" box="[334,424,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1928,1952]" captionTargetBox="[159,1424,352,1893]" captionTargetId="figure-95@4.[151,1436,345,1904]" captionTargetPageId="4" captionText="FIGURE 3. Pseudostenhelia bathyalis sp. nov., female: A, rostrum and antennule (microphotograph available in http:// metadata.icmyl.unam.mx/handle/20.500.12201/10578); B, antenna (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10569)." figureDoi="http://doi.org/10.5281/zenodo.5563485" httpUri="https://zenodo.org/record/5563485/files/figure.png" pageId="4" pageNumber="155">Fig. 3A</figureCitation>
) six-segmented, all segments smooth except for spinular row on first segment; all setae smooth; second segment with two, third segment with one seta with fracture plane; sixth segment with two articulated setae. Armature formula: 1(1); 2(9); 3(7); 4(4 + (1 + ae)), 5(2); 6(11 + acro). Acrothek consisting of two setae and one slender aesthetasc fused basally.
</paragraph>
<caption id="23B11A94FF8CC512FF1604B29B81BC84" ID-DOI="http://doi.org/10.5281/zenodo.5563485" ID-Zenodo-Dep="5563485" httpUri="https://zenodo.org/record/5563485/files/figure.png" pageId="4" pageNumber="155" startId="4.[151,250,1928,1952]" targetBox="[159,1424,352,1893]" targetPageId="4">
<paragraph id="77714A1CFF8CC512FF1604B29B81BC84" blockId="4.[151,1437,1928,2024]" pageId="4" pageNumber="155">
<emphasis id="45BA960EFF8CC512FF1604B29B72BCCC" bold="true" box="[151,280,1928,1952]" pageId="4" pageNumber="155">FIGURE 3.</emphasis>
<taxonomicName id="B0CE319FFF8CC512FEA504B29858BCCC" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[292,562,1928,1952]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="bathyalis" status="sp. nov.">
<emphasis id="45BA960EFF8CC512FEA504B29858BCCC" box="[292,562,1928,1952]" italics="true" pageId="4" pageNumber="155">Pseudostenhelia bathyalis</emphasis>
</taxonomicName>
<emphasis id="45BA960EFF8CC512FDBC04B298F9BCCC" bold="true" box="[573,659,1928,1952]" pageId="4" pageNumber="155">
<taxonomicNameLabel id="5E892B75FF8CC512FDBC04B298F9BCCC" box="[573,659,1928,1952]" pageId="4" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A, rostrum and antennule (microphotograph available in http:// metadata.icmyl.unam.mx/handle/20.500.12201/10578); B, antenna (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10569).
</paragraph>
</caption>
<paragraph id="77714A1CFF8DC513FF4603AD9B54BAE5" blockId="5.[151,1437,151,574]" pageId="5" pageNumber="156">
Antenna (
<figureCitation id="EFF55699FF8DC513FEB403AD9BE7BBDD" box="[309,397,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,1928,1952]" captionTargetBox="[159,1424,352,1893]" captionTargetId="figure-95@4.[151,1436,345,1904]" captionTargetPageId="4" captionText="FIGURE 3. Pseudostenhelia bathyalis sp. nov., female: A, rostrum and antennule (microphotograph available in http:// metadata.icmyl.unam.mx/handle/20.500.12201/10578); B, antenna (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10569)." figureDoi="http://doi.org/10.5281/zenodo.5563485" httpUri="https://zenodo.org/record/5563485/files/figure.png" pageId="5" pageNumber="156">Fig. 3B</figureCitation>
). Coxa short, with some slender outer spinules. Allobasis as long as free endopodal segment, with inner spinules proximally, with one slender abexopodal seta arising midway inner margin. Exopod threesegmented; first and third segments longest, each about three times as long as wide, and with spinules as shown; first and second segment with one pinnate seta each, third segment with one pinnate proximal and three smooth apical setae. Endopod with strong inner spinules, laterally with two spines and one slender seta, distally with six elements (outermost pinnate seta fused basally to strongly spinulose adjacent element), outer margin with two subdistal frills.
</paragraph>
<paragraph id="77714A1CFF8DC513FF4602A99905B952" blockId="5.[151,1437,151,574]" pageId="5" pageNumber="156">
Mandible (
<figureCitation id="EFF55699FF8DC513FEC002A99BFDBAC1" box="[321,407,403,429]" captionStart="FIGURE 4" captionStartId="5.[151,250,1930,1954]" captionTargetBox="[160,1433,604,1899]" captionTargetId="figure-198@5.[151,1436,597,1907]" captionTargetPageId="5" captionText="FIGURE 4. Pseudostenhelia bathyalissp. nov., female:A, mandible; B, coxa, basis, endopod and exopodof maxillule; C, maxilla (microphotograph available ion http://metadata.icmyl.unam.mx/handle/20.500.12201/10571); D, maxilliped (microphotograph available in http://metadata.icmyl.unam.mx/handle/20.500.12201/10572)." figureDoi="http://doi.org/10.5281/zenodo.5563487" httpUri="https://zenodo.org/record/5563487/files/figure.png" pageId="5" pageNumber="156">Fig. 4A</figureCitation>
) with relatively short coxa. Gnathobase wide, ventral distal corner produced into small sharp semi-hyaline process, with several acute teeth distally, and two smooth setae ventrally. Basis elongate, with narrow base, widest medially, with spinules as shown, with three subdistal setae. Exopod elongate, about four times as long as wide, tapering distally, with three lateral and three distal setae. Endopod recurved, twisted over exopod, with two lateral and six distal elements (two pairs of which fused basally).
</paragraph>
<caption id="23B11A94FF8DC513FF1604B099EEBC86" ID-DOI="http://doi.org/10.5281/zenodo.5563487" ID-Zenodo-Dep="5563487" httpUri="https://zenodo.org/record/5563487/files/figure.png" pageId="5" pageNumber="156" startId="5.[151,250,1930,1954]" targetBox="[160,1433,604,1899]" targetPageId="5">
<paragraph id="77714A1CFF8DC513FF1604B099EEBC86" blockId="5.[151,1436,1930,2026]" pageId="5" pageNumber="156">
<emphasis id="45BA960EFF8DC513FF1604B09B7BBCCE" bold="true" box="[151,273,1930,1954]" pageId="5" pageNumber="156">FIGURE 4.</emphasis>
<taxonomicName id="B0CE319FFF8DC513FE9404B09876BCCE" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[277,540,1930,1954]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="bathyalis" status="sp. nov.">
<emphasis id="45BA960EFF8DC513FE9404B09876BCCE" box="[277,540,1930,1954]" italics="true" pageId="5" pageNumber="156">Pseudostenhelia bathyalis</emphasis>
</taxonomicName>
<emphasis id="45BA960EFF8DC513FD9E04B09804BCCE" bold="true" box="[543,622,1930,1954]" pageId="5" pageNumber="156">
<taxonomicNameLabel id="5E892B75FF8DC513FD9E04B09804BCCE" box="[543,622,1930,1954]" pageId="5" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female:A, mandible; B, coxa, basis, endopod and exopod of maxillule; C, maxilla (microphotograph available ion http://metadata.icmyl.unam.mx/handle/20.500.12201/10571); D, maxilliped (microphotograph available in http://metadata.icmyl.unam.mx/handle/20.500.12201/10572).
</paragraph>
</caption>
<paragraph id="77714A1CFF8EC510FF4603AD9B2ABA71" blockId="6.[151,1437,151,1617]" pageId="6" pageNumber="157">
Maxillule (
<figureCitation id="EFF55699FF8EC510FEC903AD9BCFBBDD" box="[328,421,151,177]" captionStart="FIGURE 4" captionStartId="5.[151,250,1930,1954]" captionTargetBox="[160,1433,604,1899]" captionTargetId="figure-198@5.[151,1436,597,1907]" captionTargetPageId="5" captionText="FIGURE 4. Pseudostenhelia bathyalissp. nov., female:A, mandible; B, coxa, basis, endopod and exopodof maxillule; C, maxilla (microphotograph available ion http://metadata.icmyl.unam.mx/handle/20.500.12201/10571); D, maxilliped (microphotograph available in http://metadata.icmyl.unam.mx/handle/20.500.12201/10572)." figureDoi="http://doi.org/10.5281/zenodo.5563487" httpUri="https://zenodo.org/record/5563487/files/figure.png" pageId="6" pageNumber="157">Fig. 4B</figureCitation>
). Praecoxa and arthrite lost during dissection. Coxa with one pinnate and one smooth seta. Basis with small inner spinules, and with seven elements of which two pinnate. Exopod and endopod onesegmented, confluent, not fused to supporting basis; endopod larger than exopod, the former with four, the latter with two setae.
</paragraph>
<paragraph id="77714A1CFF8EC510FF46021D9B0BBABE" blockId="6.[151,1437,151,1617]" pageId="6" pageNumber="157">
Maxilla (
<figureCitation id="EFF55699FF8EC510FEAD021D9BE9BA2D" box="[300,387,295,321]" captionStart="FIGURE 4" captionStartId="5.[151,250,1930,1954]" captionTargetBox="[160,1433,604,1899]" captionTargetId="figure-198@5.[151,1436,597,1907]" captionTargetPageId="5" captionText="FIGURE 4. Pseudostenhelia bathyalissp. nov., female:A, mandible; B, coxa, basis, endopod and exopodof maxillule; C, maxilla (microphotograph available ion http://metadata.icmyl.unam.mx/handle/20.500.12201/10571); D, maxilliped (microphotograph available in http://metadata.icmyl.unam.mx/handle/20.500.12201/10572)." figureDoi="http://doi.org/10.5281/zenodo.5563487" httpUri="https://zenodo.org/record/5563487/files/figure.png" pageId="6" pageNumber="157">Fig. 4C</figureCitation>
). Large syncoxa seemingly without spinular ornamentation, with three endites; proximal endite smallest, with one seta; middle endite with one smooth and one spinulose seta; distal endite with one smooth and two spinulose elements. Basis drawn out into strong spinulose claw, additionally with strong naked spine and two slender setae. Endopod about 1.5 times as long as wide, with five slender setae (one arising basally, two medially, and two apically).
</paragraph>
<paragraph id="77714A1CFF8EC510FF4602E19E24B9E9" blockId="6.[151,1437,151,1617]" pageId="6" pageNumber="157">
Maxilliped (
<figureCitation id="EFF55699FF8EC510FED202E19BC7BA99" box="[339,429,475,501]" captionStart="FIGURE 4" captionStartId="5.[151,250,1930,1954]" captionTargetBox="[160,1433,604,1899]" captionTargetId="figure-198@5.[151,1436,597,1907]" captionTargetPageId="5" captionText="FIGURE 4. Pseudostenhelia bathyalissp. nov., female:A, mandible; B, coxa, basis, endopod and exopodof maxillule; C, maxilla (microphotograph available ion http://metadata.icmyl.unam.mx/handle/20.500.12201/10571); D, maxilliped (microphotograph available in http://metadata.icmyl.unam.mx/handle/20.500.12201/10572)." figureDoi="http://doi.org/10.5281/zenodo.5563487" httpUri="https://zenodo.org/record/5563487/files/figure.png" pageId="6" pageNumber="157">Fig. 4D</figureCitation>
) subchelate. Syncoxa elongate, rectangular; about 2.5 times as long as wide, with inner spinules subdistally, with two pinnate setae issuing subdistally at the same level, and one distal pinnate element arising from pedestal. Basis slightly shorter than syncoxa, oval, with arched row of outer spinules, with one anterior and one posterior inner spinular row as depicted, with two slender subdistal setae, one of which visibly longer. Endopod one-segmented, with one claw-like spinulose element and one smooth seta.
</paragraph>
<paragraph id="77714A1CFF8EC510FF4601B59FEEB881" blockId="6.[151,1437,151,1617]" pageId="6" pageNumber="157">
P1 (
<figureCitation id="EFF55699FF8EC510FF7901B59B3CB9C5" box="[248,342,654,681]" captionStart="FIGURE 5" captionStartId="7.[151,250,1921,1945]" captionTargetBox="[195,1390,497,1889]" captionTargetId="figure-178@7.[187,1401,489,1898]" captionTargetPageId="7" captionText="FIGURE 5. Pseudostenhelia bathyalis sp. nov., female: A, P1, anterior (microphotograph available in http://metadata. icmyl.unam.mx/handle/20.500.12201/10573); B, P2, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10574)." figureDoi="http://doi.org/10.5281/zenodo.5563489" httpUri="https://zenodo.org/record/5563489/files/figure.png" pageId="6" pageNumber="157">Fig. 5A</figureCitation>
). Coxa massive, trapezoidal, with proximal and subdistal outer spinules. Basis with few inner slender spinules, and with stronger spinules medially, between rami, and at base of outer and inner spines, of which the former smooth, the latter pinnate. Exopod three-segmented, reaching middle of ENP2; first segment longest, second segment shortest, length ratio of EXP1–EXP3 1, 0.5, 0.8; all segments without outer nor inner acute distal processes; EXP1 with outer and distal spinules, with outer spine, without inner armature; EXP2 with distal spinules, with outer and inner rat-like setae; EXP3 with two outer, one apical and one inner rat-like element, with spinules at base of outer and distal element. Endopod two-segmented, segments without inner nor outer acute distal processes; ENP1 reaching middle of EXP2, 1.8 times as long as wide and slightly shorter than ENP2, with slender long inner spinules, with strong outer and with small distal spinules, with one long inner seta; ENP2 slenderer than ENP1, 4.5 times as long as wide, with strong outer spinules, with one inner and one distal rat-like seta, and one outer spine.
</paragraph>
<paragraph id="77714A1CFF8EC510FF4600CD9F0BBE5D" blockId="6.[151,1437,151,1617]" pageId="6" pageNumber="157">
P2–P3 (
<figureCitation id="EFF55699FF8EC510FE9E00CD9BEBBF7D" box="[287,385,1014,1041]" captionStart="FIGURE 5" captionStartId="7.[151,250,1921,1945]" captionTargetBox="[195,1390,497,1889]" captionTargetId="figure-178@7.[187,1401,489,1898]" captionTargetPageId="7" captionText="FIGURE 5. Pseudostenhelia bathyalis sp. nov., female: A, P1, anterior (microphotograph available in http://metadata. icmyl.unam.mx/handle/20.500.12201/10573); B, P2, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10574)." figureDoi="http://doi.org/10.5281/zenodo.5563489" httpUri="https://zenodo.org/record/5563489/files/figure.png" pageId="6" pageNumber="157">Figs. 5B</figureCitation>
, 
<figureCitation id="EFF55699FF8EC510FE0C00CD9BD9BF7E" box="[397,435,1015,1042]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1928]" captionTargetBox="[169,1414,190,1872]" captionTargetId="figure-20@8.[163,1423,181,1880]" captionTargetPageId="8" captionText="FIGURE 6. Pseudostenhelia bathyalis sp. nov., female: A, P3, anterior (microphotograph available in http://metadata. icmyl.unam.mx/handle/20.500.12201/10575); B, P4, anterior (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10576); C, P5, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10577)." figureDoi="http://doi.org/10.5281/zenodo.5563491" httpUri="https://zenodo.org/record/5563491/files/figure.png" pageId="6" pageNumber="157">6A</figureCitation>
). Coxa shorter but wider than in P1, with one proximal and one subdistal row of spinules. Basis with acute inner process, with set of long inner spinules proximally, and with small spinules at base of outer seta. Exopod three-segmented, shorter than ENP; of P2 reaching middle, of P3 reaching distal third of ENP2; EXP1 longest, length ratio of P2/P3 EXP1–EXP3 1, 0.6, 0.7/1, 0.6, 0.8; EXP1 and EXP2 with outer and subdistal outer spinules, with outer acute process, EXP1 with, EXP2 seemingly without inner hyaline frill, EXP3 without spinular ornamentation; EXP1 and EXP2 without inner armature; EXP3 with two inner and two apical setae, and two outer spines. ENP arising from pedestal, at a higher level than EXP, two-segmented; ENP1 0.7 times as long as ENP2; ENP1 and ENP2 with longitudinal row of strong outer spinules; P2 and P3 ENP1 with one inner element, of P2 visibly shorter; P2 ENP2 with five, P3 ENP2 with seven setae/spines. No pores detected on EXP nor on ENP.
</paragraph>
<paragraph id="77714A1CFF8EC510FF4606019E9ABD41" blockId="6.[151,1437,151,1617]" pageId="6" pageNumber="157">
P4 (
<figureCitation id="EFF55699FF8EC510FF7506019B27BE3A" box="[244,333,1339,1366]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1928]" captionTargetBox="[169,1414,190,1872]" captionTargetId="figure-20@8.[163,1423,181,1880]" captionTargetPageId="8" captionText="FIGURE 6. Pseudostenhelia bathyalis sp. nov., female: A, P3, anterior (microphotograph available in http://metadata. icmyl.unam.mx/handle/20.500.12201/10575); B, P4, anterior (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10576); C, P5, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10577)." figureDoi="http://doi.org/10.5281/zenodo.5563491" httpUri="https://zenodo.org/record/5563491/files/figure.png" pageId="6" pageNumber="157">Fig. 6B</figureCitation>
). Coxa largely as in P2 and P3. Basis largely as in P2 and P3, but without inner acute process, and with both rami arising at the same level. Exopod three-segmented, two times as long as ENP; EXP1 longest, length ratio of EXP1–EXP3 1, 0.9, 0.9; EXP1 and EXP2 with outer and subdistal outer spinules, without acute outer process; EXP1 with, EXP2 seemingly without inner hyaline frill; EXP3 without spinular ornamentation; EXP1 and EXP2 without inner armature; EXP3 with three inner and two apical setae, and two outer spines. ENP two-segmented; ENP1 slightly shorter than ENP2; ENP1 and ENP2 with longitudinal row of strong outer spinules; ENP1 with one strong and stiff inner element; ENP2 with five setae/spines. No pores detected on EXP nor on ENP.
</paragraph>
<paragraph id="77714A1CFF8EC510FF46050D98DABD3D" blockId="6.[151,1437,151,1617]" box="[199,688,1591,1617]" pageId="6" pageNumber="157">Setal formula of swimming legs as follows:</paragraph>
<paragraph id="77714A1CFF8EC510FD8A05BF9F52BD81" pageId="6" pageNumber="157">
<table id="05CEB8BCFF8E3AE9FF2705BF9F2BBD81" box="[166,1345,1669,1773]" gridcols="5" gridrows="3" inLine="true" pageId="6" pageNumber="157">
<tr id="C9FE485EFF8E3AE9FF2705BF9F2BBDF1" box="[166,1345,1669,1693]" gridrow="0" pageId="6" pageNumber="157" rowspan-0="1">
<th id="8A2F2122FF8E3AE9FE7205BF9854BDF1" box="[499,574,1669,1693]" gridcol="1" gridrow="0" pageId="6" pageNumber="157">P1</th>
<th id="8A2F2122FF8E3AE9FD7505BF9955BDF1" box="[756,831,1669,1693]" gridcol="2" gridrow="0" pageId="6" pageNumber="157">P2</th>
<th id="8A2F2122FF8E3AE9FC7405BF9E2ABDF1" box="[1013,1088,1669,1693]" gridcol="3" gridrow="0" pageId="6" pageNumber="157">P3</th>
<th id="8A2F2122FF8E3AE9FB7705BF9F2BBDF1" box="[1270,1345,1669,1693]" gridcol="4" gridrow="0" pageId="6" pageNumber="157">P4</th>
</tr>
<tr id="C9FE485EFF8E3AE9FF2705979F2BBDA9" box="[166,1345,1709,1733]" gridrow="1" pageId="6" pageNumber="157">
<th id="8A2F2122FF8E3AE9FF2705979ABCBDA9" box="[166,214,1709,1733]" gridcol="0" gridrow="1" pageId="6" pageNumber="157">EXP</th>
<td id="8A2F2122FF8E3AE9FE7205979854BDA9" box="[499,574,1709,1733]" gridcol="1" gridrow="1" pageId="6" pageNumber="157">0,1,112</td>
<td id="8A2F2122FF8E3AE9FD7505979955BDA9" box="[756,831,1709,1733]" gridcol="2" gridrow="1" pageId="6" pageNumber="157">0,0,222</td>
<td id="8A2F2122FF8E3AE9FC7405979E2ABDA9" box="[1013,1088,1709,1733]" gridcol="3" gridrow="1" pageId="6" pageNumber="157">0,0,222</td>
<td id="8A2F2122FF8E3AE9FB7705979F2BBDA9" box="[1270,1345,1709,1733]" gridcol="4" gridrow="1" pageId="6" pageNumber="157">0,0,322</td>
</tr>
<tr id="C9FE485EFF8E3AE9FF2705EF9F2BBD81" box="[166,1345,1749,1773]" gridrow="2" pageId="6" pageNumber="157">
<th id="8A2F2122FF8E3AE9FF2705EF9ABCBD81" box="[166,214,1749,1773]" gridcol="0" gridrow="2" pageId="6" pageNumber="157">ENP</th>
<td id="8A2F2122FF8E3AE9FE7205EF9854BD81" box="[499,574,1749,1773]" gridcol="1" gridrow="2" pageId="6" pageNumber="157">1,111</td>
<td id="8A2F2122FF8E3AE9FD7505EF9955BD81" box="[756,831,1749,1773]" gridcol="2" gridrow="2" pageId="6" pageNumber="157">1,221</td>
<td id="8A2F2122FF8E3AE9FC7405EF9E2ABD81" box="[1013,1088,1749,1773]" gridcol="3" gridrow="2" pageId="6" pageNumber="157">1,421</td>
<td id="8A2F2122FF8E3AE9FB7705EF9F2BBD81" box="[1270,1345,1749,1773]" gridcol="4" gridrow="2" pageId="6" pageNumber="157">1,221</td>
</tr>
</table>
</paragraph>
<paragraph id="77714A1CFF8EC510FF4604269E4EBC13" blockId="6.[151,1437,1820,2027]" pageId="6" pageNumber="157">
P5 (
<figureCitation id="EFF55699FF8EC510FF7404279B24BC54" box="[245,334,1821,1848]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1928]" captionTargetBox="[169,1414,190,1872]" captionTargetId="figure-20@8.[163,1423,181,1880]" captionTargetPageId="8" captionText="FIGURE 6. Pseudostenhelia bathyalis sp. nov., female: A, P3, anterior (microphotograph available in http://metadata. icmyl.unam.mx/handle/20.500.12201/10575); B, P4, anterior (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10576); C, P5, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10577)." figureDoi="http://doi.org/10.5281/zenodo.5563491" httpUri="https://zenodo.org/record/5563491/files/figure.png" pageId="6" pageNumber="157">Fig. 6C</figureCitation>
). BENP pentagonal, poorly developed, without spinular ornamentation; endopodal lobe with four setae as shown; first and second outermost setae close to each other. Exopod oval, nearly as long as wide, without spinular ornamentation, with five setae. No pores detected on BENP nor on EXP.
</paragraph>
<paragraph id="77714A1CFF8EC510FF4604B39B89BCCF" blockId="6.[151,1437,1820,2027]" box="[199,483,1929,1955]" pageId="6" pageNumber="157">P6 lost during dissection.</paragraph>
<paragraph id="77714A1CFF8EC510FF4604979BE2BCAB" blockId="6.[151,1437,1820,2027]" box="[199,392,1965,1991]" pageId="6" pageNumber="157">
<emphasis id="45BA960EFF8EC510FF4604979B61BCAB" bold="true" box="[199,267,1965,1991]" pageId="6" pageNumber="157">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="3FD41997FF8EC51FFF4604EB9E7FBE89" lastPageId="9" lastPageNumber="160" pageId="6" pageNumber="157" type="discussion">
<paragraph id="77714A1CFF8EC511FF4604EB99A2BABD" blockId="6.[151,1437,1820,2027]" lastBlockId="7.[151,1436,151,465]" lastPageId="7" lastPageNumber="158" pageId="6" pageNumber="157">
<emphasis id="45BA960EFF8EC510FF4604EB9B9BBC86" bold="true" box="[199,497,2000,2027]" pageId="6" pageNumber="157">Remarks and discussion.</emphasis>
The genus 
<taxonomicName id="B0CE319FFF8EC510FDF504EB9946BC87" authorityName="Wells" authorityYear="1967" box="[628,812,2001,2027]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF8EC510FDF504EB9946BC87" box="[628,812,2001,2027]" italics="true" pageId="6" pageNumber="157">Pseudostenhelia</emphasis>
</taxonomicName>
accommodates three species, 
<taxonomicName id="B0CE319FFF8EC510FB0104EB9EB5BC86" authorityName="Wells" authorityYear="1967" box="[1152,1247,2001,2026]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="45BA960EFF8EC510FB0104EB9EB5BC86" box="[1152,1247,2001,2026]" italics="true" pageId="6" pageNumber="157">P. prima</emphasis>
</taxonomicName>
, 
<taxonomicName id="B0CE319FFF8EC510FB6C04EB9F0FBC87" authorityName="Wells" authorityYear="1971" box="[1261,1381,2001,2027]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="secunda">
<emphasis id="45BA960EFF8EC510FB6C04EB9F0FBC87" box="[1261,1381,2001,2027]" italics="true" pageId="6" pageNumber="157">P. secunda</emphasis>
</taxonomicName>
, and 
<taxonomicName id="B0CE319FFF8FC511FF1603AD9A99BBDD" authorityName="Coull &amp; Fleeger" authorityYear="1977" box="[151,243,151,177]" class="Hexanauplia" family="Rhizothrichidae" genus="Wellstenvalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="wellsi">
<emphasis id="45BA960EFF8FC511FF1603AD9A99BBDD" box="[151,243,151,177]" italics="true" pageId="7" pageNumber="158">P. wellsi</emphasis>
</taxonomicName>
. Members of the genus have a preference for shallow brackish conditions with detritus-rich fine sediments (mud and fine sand). The discovery of the female of 
<taxonomicName id="B0CE319FFF8FC511FD7F03819912BBB9" authorityName="Wells" authorityYear="1971" box="[766,888,187,213]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="secunda">
<emphasis id="45BA960EFF8FC511FD7F03819912BBB9" box="[766,888,187,213]" italics="true" pageId="7" pageNumber="158">P. secunda</emphasis>
</taxonomicName>
in plankton samples from Lake Kolleru (
<collectingCountry id="0FD90A8CFF8FC511FAD903819FFFBBBA" box="[1368,1429,187,214]" name="India" pageId="7" pageNumber="158">India</collectingCountry>
) (
<bibRefCitation id="135F37EDFF8FC511FF1E03E59B16BB95" author="Ranga Reddy" box="[159,380,223,249]" firstAuthor="Ranga Reddy" pageId="7" pageNumber="158" pagination="149 - 156" refId="ref9473" refString="Ranga Reddy, Y. (1984) The undescribed female of Pseudostenhelia secunda Wells, 1971 (Copepoda Harpacticoida) and keys to the genus. Hydrobiologia, 114, 149 - 156. https: // doi. org / 10.1007 / BF 00018111" type="journal article" year="1984">Ranga Reddy 1984</bibRefCitation>
) is, to the best of my knowledge, the only report of a species of this genus from freshwater habitats. 
<taxonomicName id="B0CE319FFF8FC511FE810239986CBA70" authorityName="Wells" authorityYear="1967" box="[256,518,259,285]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="prima">
<emphasis id="45BA960EFF8FC511FE810239986CBA70" box="[256,518,259,285]" italics="true" pageId="7" pageNumber="158">Pseudostenhelia prima</emphasis>
</taxonomicName>
, the first species assigned to this genus, was found in fine sediments (mud or muddy sand) from Saco da Inhaca, a shallow inlet south of Inhaca Island (
<collectingCountry id="0FD90A8CFF8FC511FC53021D9E03BA2E" box="[978,1129,295,322]" name="Mozambique" pageId="7" pageNumber="158">Mozambique</collectingCountry>
) (
<bibRefCitation id="135F37EDFF8FC511FBFE021D9F69BA2E" author="Wells" box="[1151,1283,295,322]" firstAuthor="Wells" pageId="7" pageNumber="158" pagination="189 - 358" refId="ref9752" refString="Wells, J. B. J. (1967) The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh, 67, 189 - 358. https: // doi. org / 10.1017 / S 0080456800024017" type="journal article" year="1967">Wells 1967</bibRefCitation>
). The male of 
<taxonomicName id="B0CE319FFF8FC511FF1602719B64BA09" authorityName="Wells" authorityYear="1971" box="[151,270,331,357]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="secunda">
<emphasis id="45BA960EFF8FC511FF1602719B64BA09" box="[151,270,331,357]" italics="true" pageId="7" pageNumber="158">P. secunda</emphasis>
</taxonomicName>
, the second species included in this genus, was found in muddy sediments from the estuary of the Vellar river near Parangipettai (formerly Porto Novo) in the 
<collectingRegion id="B50A84FEFF8FC511FD79025599D4BAE5" box="[760,958,367,393]" country="India" name="Tamil Nadu" pageId="7" pageNumber="158">Tamil Nadu State</collectingRegion>
(
<collectingCountry id="0FD90A8CFF8FC511FC4F02559E61BAE6" box="[974,1035,367,394]" name="India" pageId="7" pageNumber="158">India</collectingCountry>
) (
<bibRefCitation id="135F37EDFF8FC511FBA302559EBABAE5" author="McIntyre" box="[1058,1232,367,393]" firstAuthor="McIntyre" pageId="7" pageNumber="158" pagination="377 - 392" refId="ref9294" refString="McIntyre, A. D. (1968) The meiofauna and macrofauna of some tropical beaches. Journal of Zoology, 156, 377 - 392. https: // doi. org / 10.1111 / j. 1469 - 7998.1968. tb 04360. x" type="journal article" year="1968">McIntyre 1968</bibRefCitation>
; 
<bibRefCitation id="135F37EDFF8FC511FB5D02559F34BAE5" author="Wells" box="[1244,1374,367,393]" firstAuthor="Wells" pageId="7" pageNumber="158" pagination="507 - 520" refId="ref9800" refString="Wells, J. B. J. (1971) The Harpacticoida (Crustacea: Copepoda) of two beaches in south-east India. Journal of natural History, 5, 507 - 520. https: // doi. org / 10.1080 / 00222937100770381" type="journal article" year="1971">Wells 1971</bibRefCitation>
), and the female was found in a freshwater lake (see above; 
<bibRefCitation id="135F37EDFF8FC511FC8002A999B7BAC1" author="Ranga Reddy" box="[769,989,403,429]" firstAuthor="Ranga Reddy" pageId="7" pageNumber="158" pagination="149 - 156" refId="ref9473" refString="Ranga Reddy, Y. (1984) The undescribed female of Pseudostenhelia secunda Wells, 1971 (Copepoda Harpacticoida) and keys to the genus. Hydrobiologia, 114, 149 - 156. https: // doi. org / 10.1007 / BF 00018111" type="journal article" year="1984">Ranga Reddy 1984</bibRefCitation>
). Finally, 
<taxonomicName id="B0CE319FFF8FC511FBCF02A99EC6BAC1" authorityName="Coull &amp; Fleeger" authorityYear="1977" box="[1102,1196,403,429]" class="Hexanauplia" family="Rhizothrichidae" genus="Wellstenvalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="wellsi">
<emphasis id="45BA960EFF8FC511FBCF02A99EC6BAC1" box="[1102,1196,403,429]" italics="true" pageId="7" pageNumber="158">P. wellsi</emphasis>
</taxonomicName>
was found in muddy intertidal sediments from 
<collectingRegion id="B50A84FEFF8FC511FE39028D980ABABD" box="[440,608,439,465]" country="United States of America" name="South Carolina" pageId="7" pageNumber="158">South Carolina</collectingRegion>
(
<collectingCountry id="0FD90A8CFF8FC511FDEE028D98C0BABD" box="[623,682,439,465]" name="United States of America" pageId="7" pageNumber="158">USA</collectingCountry>
) (
<bibRefCitation id="135F37EDFF8FC511FD40028D99D1BABD" author="Coull &amp; Fleeger" box="[705,955,439,465]" firstAuthor="Coull" pageId="7" pageNumber="158" pagination="332 - 340" refId="ref8727" refString="Coull, B. C. &amp; Fleeger, J. W. (1977) A new species of Pseudostenhelia, and morphological variations in Nannopus palustris (Copepoda, Harpacticoida). Transactions of the American Microscopical Society, 96, 332 - 340. https: // doi. org / 10.2307 / 3225863" type="journal article" year="1977">Coull &amp; Fleeger 1977</bibRefCitation>
).
</paragraph>
<caption id="23B11A94FF8FC511FF1604BB9BA9BC8D" ID-DOI="http://doi.org/10.5281/zenodo.5563489" ID-Zenodo-Dep="5563489" httpUri="https://zenodo.org/record/5563489/files/figure.png" pageId="7" pageNumber="158" startId="7.[151,250,1921,1945]" targetBox="[195,1390,497,1889]" targetPageId="7">
<paragraph id="77714A1CFF8FC511FF1604BB9BA9BC8D" blockId="7.[151,1437,1921,2017]" pageId="7" pageNumber="158">
<emphasis id="45BA960EFF8FC511FF1604BB9B73BCF5" bold="true" box="[151,281,1921,1945]" pageId="7" pageNumber="158">FIGURE 5.</emphasis>
<taxonomicName id="B0CE319FFF8FC511FEA704BB985CBCF5" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[294,566,1921,1945]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="bathyalis" status="sp. nov.">
<emphasis id="45BA960EFF8FC511FEA704BB985CBCF5" box="[294,566,1921,1945]" italics="true" pageId="7" pageNumber="158">Pseudostenhelia bathyalis</emphasis>
</taxonomicName>
<emphasis id="45BA960EFF8FC511FDC304BB98F3BCF5" bold="true" box="[578,665,1921,1945]" pageId="7" pageNumber="158">
<taxonomicNameLabel id="5E892B75FF8FC511FDC304BB98F3BCF5" box="[578,665,1921,1945]" pageId="7" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A, P1, anterior (microphotograph available in http://metadata. icmyl.unam.mx/handle/20.500.12201/10573); B, P2, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10574).
</paragraph>
</caption>
<caption id="23B11A94FF80C51EFF16044A9BA9BC98" ID-DOI="http://doi.org/10.5281/zenodo.5563491" ID-Zenodo-Dep="5563491" httpUri="https://zenodo.org/record/5563491/files/figure.png" pageId="8" pageNumber="159" startId="8.[151,250,1904,1928]" targetBox="[169,1414,190,1872]" targetPageId="8">
<paragraph id="77714A1CFF80C51EFF16044A9BA9BC98" blockId="8.[151,1437,1904,2036]" pageId="8" pageNumber="159">
<emphasis id="45BA960EFF80C51EFF16044A9B73BCE4" bold="true" box="[151,281,1904,1928]" pageId="8" pageNumber="159">FIGURE 6.</emphasis>
<taxonomicName id="B0CE319FFF80C51EFEA7044A985CBCE4" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[294,566,1904,1928]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="bathyalis" status="sp. nov.">
<emphasis id="45BA960EFF80C51EFEA7044A985CBCE4" box="[294,566,1904,1928]" italics="true" pageId="8" pageNumber="159">Pseudostenhelia bathyalis</emphasis>
</taxonomicName>
<emphasis id="45BA960EFF80C51EFDC3044B98F3BCE5" bold="true" box="[578,665,1905,1929]" pageId="8" pageNumber="159">
<taxonomicNameLabel id="5E892B75FF80C51EFDC3044B98F3BCE5" box="[578,665,1905,1929]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A, P3, anterior (microphotograph available in http://metadata. icmyl.unam.mx/handle/20.500.12201/10575); B, P4, anterior (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10576); C, P5, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10577).
</paragraph>
</caption>
<paragraph id="77714A1CFF81C51FFF4603AD9983BAE5" blockId="9.[151,1437,150,1509]" pageId="9" pageNumber="160">
The relationships between 
<taxonomicName id="B0CE319FFF81C51FFE6A03AD98C9BBDD" authorityName="Wells" authorityYear="1967" box="[491,675,151,177]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFE6A03AD98C9BBDD" box="[491,675,151,177]" italics="true" pageId="9" pageNumber="160">Pseudostenhelia</emphasis>
</taxonomicName>
and some other stenheliins are not clear. 
<bibRefCitation id="135F37EDFF81C51FFBDB03AD9F58BBDD" author="Dahms" box="[1114,1330,150,177]" etAl="et al." firstAuthor="Dahms" pageId="9" pageNumber="160" pagination="1 - 14" refId="ref8784" refString="Dahms, H. - U., Schizas, N. V. &amp; Shirley, T. C. (2005) Naupliar evolutionary novelties of Stenhelia peniculata (Copepoda, Harpacticoida) from Alaska affirming taxa belonging to different categorial rank. Invertebrate Zoology, 2, 1 - 14. https: // doi. org / 10.15298 / invertzool. 02.1.01" type="journal article" year="2005">
Dahms 
<emphasis id="45BA960EFF81C51FFB2F03AD9E89BBDD" box="[1198,1251,151,177]" italics="true" pageId="9" pageNumber="160">et al.</emphasis>
(2005)
</bibRefCitation>
proposed some apomorphies for 
<taxonomicName id="B0CE319FFF81C51FFE1A03819839BBB9" authorityName="Wells" authorityYear="1967" box="[411,595,187,213]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFE1A03819839BBB9" box="[411,595,187,213]" italics="true" pageId="9" pageNumber="160">Pseudostenhelia</emphasis>
</taxonomicName>
(
<bibRefCitation id="135F37EDFF81C51FFDE00381990ABBB9" author="Dahms" box="[609,864,186,213]" etAl="et al." firstAuthor="Dahms" page="12" pageId="9" pageNumber="160" pagination="1 - 14" refId="ref8784" refString="Dahms, H. - U., Schizas, N. V. &amp; Shirley, T. C. (2005) Naupliar evolutionary novelties of Stenhelia peniculata (Copepoda, Harpacticoida) from Alaska affirming taxa belonging to different categorial rank. Invertebrate Zoology, 2, 1 - 14. https: // doi. org / 10.15298 / invertzool. 02.1.01" type="journal article" year="2005">
Dahms 
<emphasis id="45BA960EFF81C51FFD3B03819899BBB9" box="[698,755,187,213]" italics="true" pageId="9" pageNumber="160">et al.</emphasis>
2005: 12
</bibRefCitation>
, table 2) based on naupliar morphology. They also gave a list of characters shared by 
<taxonomicName id="B0CE319FFF81C51FFDB503E598F6BB95" authorityName="Boeck" authorityYear="1865" box="[564,668,223,249]" class="Hexanauplia" family="Miraciidae" genus="Stenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFDB503E598F6BB95" box="[564,668,223,249]" italics="true" pageId="9" pageNumber="160">Stenhelia</emphasis>
</taxonomicName>
, 
<taxonomicName id="B0CE319FFF81C51FFD2C03E59971BB95" authorityName="Brady" authorityYear="1869" box="[685,795,223,249]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFD2C03E59971BB95" box="[685,795,223,249]" italics="true" pageId="9" pageNumber="160">Delavalia</emphasis>
</taxonomicName>
, and 
<taxonomicName id="B0CE319FFF81C51FFCE003E59E73BB95" authorityName="Wells" authorityYear="1967" box="[865,1049,223,249]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFCE003E59E73BB95" box="[865,1049,223,249]" italics="true" pageId="9" pageNumber="160">Pseudostenhelia</emphasis>
</taxonomicName>
, 
<emphasis id="45BA960EFF81C51FFBAB03E59E3ABB94" box="[1066,1104,223,248]" italics="true" pageId="9" pageNumber="160">viz.</emphasis>
the extremely foreshortened cephalic shield, the presence of a spinous process on the distal third (middle?; 
<bibRefCitation id="135F37EDFF81C51FFBA502399E98BA71" author="Huys &amp; Mu" box="[1060,1266,259,285]" firstAuthor="Huys" pageId="9" pageNumber="160" pagination="51 - 68" refId="ref9099" refString="Huys, R. &amp; Mu, F. H. (2008) Description of a new species of Onychostenhelia Ito (Copepoda, Harpacticoida, Miraciidae) from the Bohai Sea, China. Zootaxa, 1706 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 1706.1.2" type="journal article" year="2008">Huys &amp; Mu 2008</bibRefCitation>
) of the second antennulary segment and a notched inner seta on the third antennulary segment, and a lobate outer protuberance of the first segment of the exopod of the antenna. 
<bibRefCitation id="135F37EDFF81C51FFD1F02719919BA09" author="Huys &amp; Mu" box="[670,883,331,357]" firstAuthor="Huys" pageId="9" pageNumber="160" pagination="51 - 68" refId="ref9099" refString="Huys, R. &amp; Mu, F. H. (2008) Description of a new species of Onychostenhelia Ito (Copepoda, Harpacticoida, Miraciidae) from the Bohai Sea, China. Zootaxa, 1706 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 1706.1.2" type="journal article" year="2008">Huys &amp; Mu (2008)</bibRefCitation>
questioned the phylogenetic significance of these characters, but admitted that they appear to be unique within 
<taxonomicName id="B0CE319FFF81C51FFCC50255998FBAE5" authorityName="Sars" authorityYear="1903" box="[836,997,367,393]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="order">Harpacticoida</taxonomicName>
.
</paragraph>
<paragraph id="77714A1CFF81C51FFF4602A99A8AB99D" blockId="9.[151,1437,150,1509]" pageId="9" pageNumber="160">
<bibRefCitation id="135F37EDFF81C51FFF4602A99BC4BAC1" author="Huys &amp; Mu" box="[199,430,403,429]" firstAuthor="Huys" pageId="9" pageNumber="160" pagination="51 - 68" refId="ref9099" refString="Huys, R. &amp; Mu, F. H. (2008) Description of a new species of Onychostenhelia Ito (Copepoda, Harpacticoida, Miraciidae) from the Bohai Sea, China. Zootaxa, 1706 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 1706.1.2" type="journal article" year="2008">Huys &amp; Mu (2008)</bibRefCitation>
drew attention to a core group of 
<taxonomicName id="B0CE319FFF81C51FFCD502A999B4BAC1" authorityName="Brady" authorityYear="1880" box="[852,990,403,429]" class="Hexanauplia" family="Miraciidae" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="153" phylum="Arthropoda" rank="subFamily" subFamily="Stenheliinae">Stenheliinae</taxonomicName>
(
<taxonomicName id="B0CE319FFF81C51FFC7502A99EDABAC1" authorityName="Ito" authorityYear="1979" box="[1012,1200,403,429]" class="Hexanauplia" family="Miraciidae" genus="Onychostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFC7502A99EDABAC1" box="[1012,1200,403,429]" italics="true" pageId="9" pageNumber="160">Onychostenhelia</emphasis>
</taxonomicName>
, 
<taxonomicName id="B0CE319FFF81C51FFB4202A99F0BBAC1" authorityName="Tai &amp; Song" authorityYear="1979" box="[1219,1377,403,429]" class="Hexanauplia" family="Diosaccidae" genus="Cladorostrata" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFB4202A99F0BBAC1" box="[1219,1377,403,429]" italics="true" pageId="9" pageNumber="160">Cladorostrata</emphasis>
</taxonomicName>
, and 
<taxonomicName id="B0CE319FFF81C51FFF16028D9B6FBABD" authorityName="Brady" authorityYear="1869" box="[151,261,439,465]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFF16028D9B6FBABD" box="[151,261,439,465]" italics="true" pageId="9" pageNumber="160">Delavalia</emphasis>
</taxonomicName>
) sharing a synapomorphic confluent maxillulary exopod and endopod not fused to the supporting basis. This condition is present also in 
<taxonomicName id="B0CE319FFF81C51FFDD002E1988DBA99" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[593,743,475,501]" class="Hexanauplia" family="Miraciidae" genus="Wellstenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFDD002E1988DBA99" box="[593,743,475,501]" italics="true" pageId="9" pageNumber="160">Wellstenhelia</emphasis>
</taxonomicName>
(
<bibRefCitation id="135F37EDFF81C51FFD7702E19E63BA99" author="Karanovic &amp; Kim" box="[758,1033,475,501]" firstAuthor="Karanovic" pageId="9" pageNumber="160" pagination="1 - 96" refId="ref9195" refString="Karanovic, T. &amp; Kim, K. (2014) New insights into polyphyly of the harpacticoid genus Delavalia (Crustacea, Copepoda) through morphological and molecular study of an unprecedented diversity of sympatric species in a small South Korean bay. Zootaxa, 3783 (1), 1 - 96. https: // doi. org / 10.11646 / zootaxa. 3783.1.1" type="journal article" year="2014">Karanovic &amp; Kim 2014</bibRefCitation>
; 
<bibRefCitation id="135F37EDFF81C51FFB9502E19F09BA99" author="Gomez &amp; Cruz-Barraza" box="[1044,1379,475,501]" firstAuthor="Gomez" pageId="9" pageNumber="160" pagination="117 - 150" refId="ref9015" refString="Gomez, S. &amp; Cruz-Barraza, J. A. (2021) On some deep-sea Stenheliinae from the Gulf of California and the west coast of the Baja California Peninsula (Mexico): Wellstenhelia euterpoides sp. nov., and Wellstenvalia wellsi gen. et sp. nov. (Copepoda: Harpacticoida: Miraciidae). Zootaxa, 5051 (1), 117 - 150. https: // doi. org / 10.11646 / zootaxa. 5051.1.10" type="journal article" year="2021">Gómez &amp; Cruz-Barraza 2021</bibRefCitation>
) and 
<taxonomicName id="B0CE319FFF81C51FFF1602C5985CB975" authority="(Gomez, 2020)" baseAuthorityName="Gomez" baseAuthorityYear="2020" box="[151,566,511,537]" class="Hexanauplia" family="Miraciidae" genus="Lonchoeidestenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFF1602C59B15B975" box="[151,383,511,537]" italics="true" pageId="9" pageNumber="160">Lonchoeidestenhelia</emphasis>
(
<bibRefCitation id="135F37EDFF81C51FFE1002C59844B975" author="Gomez" box="[401,558,511,537]" firstAuthor="Gomez" pageId="9" pageNumber="160" pagination="41 - 79" refId="ref8882" refString="Gomez, S. (2020) On some new species of Stenheliinae Brady, 1880 (Copepoda, Harpacticoida, Miraciidae) from north-western Mexico, with the proposal of Lonchoeidestenhelia gen. nov. Zookeys, 987, 41 - 79. https: // doi. org / 10.3897 / zookeys. 987.52906" type="journal article" year="2020">Gómez, 2020</bibRefCitation>
)
</taxonomicName>
and is confirmed here for 
<taxonomicName id="B0CE319FFF81C51FFCF102C59E42B975" authorityName="Wells" authorityYear="1967" box="[880,1064,511,537]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFCF102C59E42B975" box="[880,1064,511,537]" italics="true" pageId="9" pageNumber="160">Pseudostenhelia</emphasis>
</taxonomicName>
. Note that 
<bibRefCitation id="135F37EDFF81C51FFB2C02C59FF6B975" author="Ranga Reddy" box="[1197,1436,511,537]" firstAuthor="Ranga Reddy" pageId="9" pageNumber="160" pagination="149 - 156" refId="ref9473" refString="Ranga Reddy, Y. (1984) The undescribed female of Pseudostenhelia secunda Wells, 1971 (Copepoda Harpacticoida) and keys to the genus. Hydrobiologia, 114, 149 - 156. https: // doi. org / 10.1007 / BF 00018111" type="journal article" year="1984">Ranga Reddy (1984)</bibRefCitation>
described the maxillulary rami as discrete and separated from basis, but this is most probably erroneous. The significance of this condition in the maxillulary rami is not clear but 
<bibRefCitation id="135F37EDFF81C51FFC0D017D9E0AB90D" author="Huys &amp; Mu" box="[908,1120,583,609]" firstAuthor="Huys" pageId="9" pageNumber="160" pagination="51 - 68" refId="ref9099" refString="Huys, R. &amp; Mu, F. H. (2008) Description of a new species of Onychostenhelia Ito (Copepoda, Harpacticoida, Miraciidae) from the Bohai Sea, China. Zootaxa, 1706 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 1706.1.2" type="journal article" year="2008">Huys &amp; Mu (2008)</bibRefCitation>
suggested that the confluent maxillulary rami separated from the basis could be a strong synapomorphy for 
<taxonomicName id="B0CE319FFF81C51FFBB001519E87B9E9" authorityName="Ito" authorityYear="1979" box="[1073,1261,619,645]" class="Hexanauplia" family="Miraciidae" genus="Onychostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFBB001519E87B9E9" box="[1073,1261,619,645]" italics="true" pageId="9" pageNumber="160">Onychostenhelia</emphasis>
</taxonomicName>
, 
<taxonomicName id="B0CE319FFF81C51FFB7E01519FF7B9E9" authorityName="Tai &amp; Song" authorityYear="1979" box="[1279,1437,619,645]" class="Hexanauplia" family="Diosaccidae" genus="Cladorostrata" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFB7E01519FF7B9E9" box="[1279,1437,619,645]" italics="true" pageId="9" pageNumber="160">Cladorostrata</emphasis>
</taxonomicName>
and 
<taxonomicName id="B0CE319FFF81C51FFF4A01B59B53B9C5" authorityName="Brady" authorityYear="1869" box="[203,313,655,681]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFF4A01B59B53B9C5" box="[203,313,655,681]" italics="true" pageId="9" pageNumber="160">Delavalia</emphasis>
</taxonomicName>
, but also for 
<taxonomicName id="B0CE319FFF81C51FFE5C01B598AFB9C5" baseAuthorityName="Gomez" baseAuthorityYear="2020" box="[477,709,655,681]" class="Hexanauplia" family="Miraciidae" genus="Lonchoeidestenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFE5C01B598AFB9C5" box="[477,709,655,681]" italics="true" pageId="9" pageNumber="160">Lonchoeidestenhelia</emphasis>
</taxonomicName>
. However, its presence in some other stenheliin lineages (
<emphasis id="45BA960EFF81C51FFAF501B59B25B9A1" italics="true" pageId="9" pageNumber="160">
e.g. 
<taxonomicName id="B0CE319FFF81C51FFF1601899B25B9A1" authorityName="Wells" authorityYear="1967" box="[151,335,691,717]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">Pseudostenhelia</taxonomicName>
</emphasis>
, and 
<taxonomicName id="B0CE319FFF81C51FFE1701899846B9A1" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[406,556,691,717]" class="Hexanauplia" family="Miraciidae" genus="Wellstenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFE1701899846B9A1" box="[406,556,691,717]" italics="true" pageId="9" pageNumber="160">Wellstenhelia</emphasis>
</taxonomicName>
) seems to indicate convergent evolution (see also 
<bibRefCitation id="135F37EDFF81C51FFB0601899AB9B99D" author="Gomez &amp; Cruz-Barraza" firstAuthor="Gomez" pageId="9" pageNumber="160" pagination="117 - 150" refId="ref9015" refString="Gomez, S. &amp; Cruz-Barraza, J. A. (2021) On some deep-sea Stenheliinae from the Gulf of California and the west coast of the Baja California Peninsula (Mexico): Wellstenhelia euterpoides sp. nov., and Wellstenvalia wellsi gen. et sp. nov. (Copepoda: Harpacticoida: Miraciidae). Zootaxa, 5051 (1), 117 - 150. https: // doi. org / 10.11646 / zootaxa. 5051.1.10" type="journal article" year="2021">Gómez &amp; Cruz-Barraza 2021</bibRefCitation>
).
</paragraph>
<paragraph id="77714A1CFF81C51FFF4601C19853B8ED" blockId="9.[151,1437,150,1509]" pageId="9" pageNumber="160">
<taxonomicName id="B0CE319FFF81C51FFF4601C19B15B879" authorityName="Wells" authorityYear="1967" box="[199,383,763,789]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFF4601C19B15B879" box="[199,383,763,789]" italics="true" pageId="9" pageNumber="160">Pseudostenhelia</emphasis>
</taxonomicName>
shares a plesiomorphic anal operculum (semicircular, relatively small, weakly chitinized and flanked by a sensilla on each side) with 
<taxonomicName id="B0CE319FFF81C51FFDD8002598C5B855" authorityName="Por" authorityYear="1964" box="[601,687,799,825]" class="Hexanauplia" family="Miraciidae" genus="Melima" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFDD8002598C5B855" box="[601,687,799,825]" italics="true" pageId="9" pageNumber="160">Melima</emphasis>
</taxonomicName>
and 
<taxonomicName id="B0CE319FFF81C51FFD6400259990B855" authority="(Willen 2003)" baseAuthorityName="Willen" baseAuthorityYear="2003" box="[741,1018,799,825]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFD6400259939B855" box="[741,851,799,825]" italics="true" pageId="9" pageNumber="160">Delavalia</emphasis>
(
<bibRefCitation id="135F37EDFF81C51FFCE300259998B855" author="Willen" box="[866,1010,799,825]" firstAuthor="Willen" pageId="9" pageNumber="160" pagination="1691 - 1711" refId="ref9895" refString="Willen, E. (2003) A new species of Stenhelia (Copepoda, Harpacticoida) from a hydrothermal, active, submarine volcano in the New Ireland Fore-Arc system (Papua New Guinea) with notes on deep sea colonization within the Stenheliinae. Journal of natural History, 37, 1691 - 1711. https: // doi. org / 10.1080 / 00222930110114437" type="journal article" year="2003">Willen 2003</bibRefCitation>
)
</taxonomicName>
, but also with 
<taxonomicName id="B0CE319FFF81C51FFB1F00259F5EB855" authorityName="Karanovic &amp; Kim" authorityYear="2014" box="[1182,1332,799,825]" class="Hexanauplia" family="Miraciidae" genus="Wellstenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFB1F00259F5EB855" box="[1182,1332,799,825]" italics="true" pageId="9" pageNumber="160">Wellstenhelia</emphasis>
</taxonomicName>
(see also 
<bibRefCitation id="135F37EDFF81C51FFF1600799B8FB831" author="Gomez &amp; Cruz-Barraza" box="[151,485,835,861]" firstAuthor="Gomez" pageId="9" pageNumber="160" pagination="117 - 150" refId="ref9015" refString="Gomez, S. &amp; Cruz-Barraza, J. A. (2021) On some deep-sea Stenheliinae from the Gulf of California and the west coast of the Baja California Peninsula (Mexico): Wellstenhelia euterpoides sp. nov., and Wellstenvalia wellsi gen. et sp. nov. (Copepoda: Harpacticoida: Miraciidae). Zootaxa, 5051 (1), 117 - 150. https: // doi. org / 10.11646 / zootaxa. 5051.1.10" type="journal article" year="2021">Gómez &amp; Cruz-Barraza 2021</bibRefCitation>
), and it is completely absent in 
<taxonomicName id="B0CE319FFF81C51FFCC4007999C7B831" authorityName="Boeck" authorityYear="1865" box="[837,941,835,861]" class="Hexanauplia" family="Miraciidae" genus="Stenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFCC4007999C7B831" box="[837,941,835,861]" italics="true" pageId="9" pageNumber="160">Stenhelia</emphasis>
</taxonomicName>
and 
<taxonomicName id="B0CE319FFF81C51FFC6200799F03B831" authority="(Mu &amp; Huys 2002)" baseAuthorityName="Mu &amp; Huys" baseAuthorityYear="2002" box="[995,1385,835,861]" class="Hexanauplia" family="Miraciidae" genus="Anisostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFC6200799EE2B831" box="[995,1160,835,861]" italics="true" pageId="9" pageNumber="160">Anisostenhelia</emphasis>
(
<bibRefCitation id="135F37EDFF81C51FFB1600799F0BB831" author="Mu &amp; Huys" box="[1175,1377,835,861]" firstAuthor="Mu" pageId="9" pageNumber="160" pagination="179 - 206" refId="ref9341" refString="Mu, F. &amp; Huys, R. (2002) New species of Stenhelia (Copepoda, Harpacticoida, Diosaccidae) from the Bohai Sea (China) with notes on subgeneric division and phylogenetic relationships. Cahiers de Biologie marine, 43, 179 - 206. https: // doi. org / 10.21411 / CBM. A. C 482 EC 6 A" type="journal article" year="2002">Mu &amp; Huys 2002</bibRefCitation>
)
</taxonomicName>
, and 
<taxonomicName id="B0CE319FFF81C51FFF16005D985FB8ED" authority="(Huys &amp; Mu 2008)" baseAuthorityName="Huys &amp; Mu" baseAuthorityYear="2008" box="[151,565,871,897]" class="Hexanauplia" family="Miraciidae" genus="Onychostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFF16005D9B39B8ED" box="[151,339,871,897]" italics="true" pageId="9" pageNumber="160">Onychostenhelia</emphasis>
(
<bibRefCitation id="135F37EDFF81C51FFEE3005D9847B8ED" author="Huys &amp; Mu" box="[354,557,871,897]" firstAuthor="Huys" pageId="9" pageNumber="160" pagination="51 - 68" refId="ref9099" refString="Huys, R. &amp; Mu, F. H. (2008) Description of a new species of Onychostenhelia Ito (Copepoda, Harpacticoida, Miraciidae) from the Bohai Sea, China. Zootaxa, 1706 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 1706.1.2" type="journal article" year="2008">Huys &amp; Mu 2008</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="77714A1CFF81C51FFF4600B19933B881" blockId="9.[151,1437,150,1509]" pageId="9" pageNumber="160">
Although its significance is not clear, the caudal seta I is either a spine-like element or a reduced seta in most stenheliins, but it is a well-developed seta (most probably plesiomorphic) in 
<taxonomicName id="B0CE319FFF81C51FFC6D00959ECEB8A5" authorityName="Wells" authorityYear="1967" box="[1004,1188,943,969]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFC6D00959ECEB8A5" box="[1004,1188,943,969]" italics="true" pageId="9" pageNumber="160">Pseudostenhelia</emphasis>
</taxonomicName>
and 
<taxonomicName id="B0CE319FFF81C51FFB5800959FFFB8A5" baseAuthorityName="Huys &amp; Mu" baseAuthorityYear="2008" box="[1241,1429,943,969]" class="Hexanauplia" family="Miraciidae" genus="Onychostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFB5800959FFFB8A5" box="[1241,1429,943,969]" italics="true" pageId="9" pageNumber="160">Onychostenhelia</emphasis>
</taxonomicName>
, and some species of 
<taxonomicName id="B0CE319FFF81C51FFEFE00E9993FB881" authority="(Gomez &amp; Cruz-Barraza 2021)" baseAuthorityName="Gomez &amp; Cruz-Barraza" baseAuthorityYear="2021" box="[383,853,979,1005]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFEFE00E99B87B881" box="[383,493,979,1005]" italics="true" pageId="9" pageNumber="160">Delavalia</emphasis>
(
<bibRefCitation id="135F37EDFF81C51FFE7D00E99926B881" author="Gomez &amp; Cruz-Barraza" box="[508,844,979,1005]" firstAuthor="Gomez" pageId="9" pageNumber="160" pagination="117 - 150" refId="ref9015" refString="Gomez, S. &amp; Cruz-Barraza, J. A. (2021) On some deep-sea Stenheliinae from the Gulf of California and the west coast of the Baja California Peninsula (Mexico): Wellstenhelia euterpoides sp. nov., and Wellstenvalia wellsi gen. et sp. nov. (Copepoda: Harpacticoida: Miraciidae). Zootaxa, 5051 (1), 117 - 150. https: // doi. org / 10.11646 / zootaxa. 5051.1.10" type="journal article" year="2021">Gómez &amp; Cruz-Barraza 2021</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="77714A1CFF81C51FFF4600CD9F1FBE61" blockId="9.[151,1437,150,1509]" pageId="9" pageNumber="160">
In addition to the synapomorphic nature of the structure of the maxillulary rami in 
<taxonomicName id="B0CE319FFF81C51FFBB000CD9E87BF7D" baseAuthorityName="Huys &amp; Mu" baseAuthorityYear="2008" box="[1073,1261,1015,1041]" class="Hexanauplia" family="Miraciidae" genus="Onychostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFBB000CD9E87BF7D" box="[1073,1261,1015,1041]" italics="true" pageId="9" pageNumber="160">Onychostenhelia</emphasis>
</taxonomicName>
, 
<taxonomicName id="B0CE319FFF81C51FFB7900CD9FFCBF7D" authorityName="Tai &amp; Song" authorityYear="1979" box="[1272,1430,1015,1041]" class="Hexanauplia" family="Diosaccidae" genus="Cladorostrata" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFB7900CD9FFCBF7D" box="[1272,1430,1015,1041]" italics="true" pageId="9" pageNumber="160">Cladorostrata</emphasis>
</taxonomicName>
, and 
<taxonomicName id="B0CE319FFF81C51FFF4907219B5CBF59" baseAuthorityName="Gomez &amp; Cruz-Barraza" baseAuthorityYear="2021" box="[200,310,1051,1077]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFF4907219B5CBF59" box="[200,310,1051,1077]" italics="true" pageId="9" pageNumber="160">Delavalia</emphasis>
</taxonomicName>
(see above), 
<bibRefCitation id="135F37EDFF81C51FFE50072198C4BF59" author="Huys &amp; Mu" box="[465,686,1051,1077]" firstAuthor="Huys" pageId="9" pageNumber="160" pagination="51 - 68" refId="ref9099" refString="Huys, R. &amp; Mu, F. H. (2008) Description of a new species of Onychostenhelia Ito (Copepoda, Harpacticoida, Miraciidae) from the Bohai Sea, China. Zootaxa, 1706 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 1706.1.2" type="journal article" year="2008">Huys &amp; Mu (2008)</bibRefCitation>
suggested that the resemblance in leg segmentation and armature between 
<taxonomicName id="B0CE319FFF81C51FFF7807059BDFBF35" baseAuthorityName="Huys &amp; Mu" baseAuthorityYear="2008" box="[249,437,1087,1113]" class="Hexanauplia" family="Miraciidae" genus="Onychostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFF7807059BDFBF35" box="[249,437,1087,1113]" italics="true" pageId="9" pageNumber="160">Onychostenhelia</emphasis>
</taxonomicName>
and 
<taxonomicName id="B0CE319FFF81C51FFE66070598F5BF35" authorityName="Wells" authorityYear="1967" box="[487,671,1087,1113]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="45BA960EFF81C51FFE66070598F5BF35" box="[487,671,1087,1113]" italics="true" pageId="9" pageNumber="160">Pseudostenhelia</emphasis>
</taxonomicName>
observed by 
<bibRefCitation id="135F37EDFF81C51FFCAF070599F7BF35" author="Ito" box="[814,925,1087,1113]" firstAuthor="Ito" pageId="9" pageNumber="160" pagination="42 - 68" refId="ref9158" refString="Ito, T. (1979) Descriptions and records of marine harpacticoid copepods from Hokkaido, VII. Journal of the Faculty of Sciences, Hokkaido University, Zoology, 22, 42 - 68." type="journal article" year="1979">Itô (1979)</bibRefCitation>
had little phylogenetic significance. However, as noted above (see also Gómez &amp; Barraza 2020), the phylogenetic importance of the confluent maxillulary rami separated from the supporting basis, the shape of caudal seta I, and anal operculum is questionable. On the other hand, the shape of P1 EXP (EXP3 smaller than EXP2 and EXP3), giving the entire ramus a characteristic appearance (except for the somewhat longer P1 EXP 
<quantity id="B036E7F9FF81C51FFDDF07F598ECBF85" box="[606,646,1231,1257]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="160" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="B0CE319FFF81C51FFD0D07F59967BF85" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[652,781,1231,1257]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="bathyalis" status="sp. nov.">
<emphasis id="45BA960EFF81C51FFD0D07F59967BF85" box="[652,781,1231,1257]" italics="true" pageId="9" pageNumber="160">P. bathyalis</emphasis>
</taxonomicName>
<emphasis id="45BA960EFF81C51FFC9207F5991FBF84" bold="true" box="[787,885,1230,1257]" pageId="9" pageNumber="160">
<taxonomicNameLabel id="5E892B75FF81C51FFC9207F59907BF84" box="[787,877,1230,1257]" pageId="9" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
)
</emphasis>
, the exceedingly long outer elements of P1 EXP2 and EXP3, and the two-segmented endopod of P2–P4, seem to be strong synapomorphies for these two genera.
</paragraph>
<paragraph id="77714A1CFF81C51FFF46062D9E7FBE89" blockId="9.[151,1437,150,1509]" pageId="9" pageNumber="160">
Both sexes are known for all 
<taxonomicName id="B0CE319FFF81C51FFD99062D9943BE5D" box="[536,809,1303,1329]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="45BA960EFF81C51FFD99062D98BABE5D" box="[536,720,1303,1329]" italics="true" pageId="9" pageNumber="160">Pseudostenhelia</emphasis>
species
</taxonomicName>
except for 
<taxonomicName id="B0CE319FFF81C51FFC2D062D9E5ABE5D" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[940,1072,1303,1329]" class="Hexanauplia" family="Miraciidae" genus="Pseudostenhelia" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="bathyalis" status="sp. nov.">
<emphasis id="45BA960EFF81C51FFC2D062D9E5ABE5D" box="[940,1072,1303,1329]" italics="true" pageId="9" pageNumber="160">P. bathyalis</emphasis>
</taxonomicName>
<emphasis id="45BA960EFF81C51FFBB9062D9EF9BE5D" bold="true" box="[1080,1171,1303,1329]" pageId="9" pageNumber="160">
<taxonomicNameLabel id="5E892B75FF81C51FFBB9062D9EF9BE5D" box="[1080,1171,1303,1329]" pageId="9" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The female of the new deep-sea species can be readily separated from the females of its congeners by the segmentation of the antennule (with six segments in the new species, but with five segments in other species), and by its armature complement of P2 ENP2 and P3 ENP2, with five and seven setae instead of four and five setae respectively. The minute proximal endite of the maxillary syncoxa with one seta only, and the reduction of the armature complement on the P5 EXP from six to five setae seem to be the only potential apomorphies for the species.
</paragraph>
</subSubSection>
</treatment>
</document>