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<document id="C2FB372C4C1A9717034FECA7A3A4A424" ID-DOI="10.11646/zootaxa.5297.3.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="8005225" ID-ZooBank="EC0AD75A-AB80-45CD-906F-39DFD6BCE6CC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685960276401" checkinUser="plazi" docAuthor="Alekseev, Vitalii I. &amp; Nabozhenko, Maxim V." docDate="2023" docId="F21687DACB337712FF4E33AAFB33FF73" docLanguage="en" docName="zootaxa.5297.3.8.pdf" docOrigin="Zootaxa 5297 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5297.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Yantaroxenos Nabozhenko, Kirejtshuk et Merkl 2016" docType="treatment" docVersion="2" lastPageNumber="442" masterDocId="0E2FFFA2CB34771AFFD9372CFF92FF8A" masterDocTitle="News from Baltic amber: a new tribe of extinct Paleogene ironclad beetles (Coleoptera: Zopheridae: Zopherinae)" masterLastPageNumber="445" masterPageNumber="435" pageNumber="442" updateTime="1685960731749" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8D5932330F52AD6E3BA305523FA813A5">News from Baltic amber: a new tribe of extinct Paleogene ironclad beetles (Coleoptera: Zopheridae: Zopherinae)</mods:title>
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<mods:namePart id="280DD2A7A402BE7FB70F1F1D55E3DA8C">Alekseev, Vitalii I.</mods:namePart>
<mods:affiliation id="9D2275FA93D9FDED192B7A985E90ED54">Kaliningrad Regional Amber Museum, Marshal Vasilevskii square 1, Kaliningrad 236016, Russia. &amp; Immanuel Kant Baltic Federal University, A. Nevskii str. 14, Kaliningrad 236016, Russia</mods:affiliation>
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<mods:namePart id="B40DAEF07638F432D62C23235DD1CAB7">Nabozhenko, Maxim V.</mods:namePart>
<mods:affiliation id="A8A37C301BF93EEEB580C64E567F291E">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str. 45, Makhachkala, Republic of Dagestan, 367000, Russia. &amp; Dagestan State University, M. Gadzhiev str. 43 a, Makhachkala, Republic of Dagestan, 367000, Russia</mods:affiliation>
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<mods:date id="2D195742F9F51AF4379614055F510DF7">2023</mods:date>
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<treatment id="F21687DACB337712FF4E33AAFB33FF73" LSID="urn:lsid:plazi:treatment:F21687DACB337712FF4E33AAFB33FF73" httpUri="http://treatment.plazi.org/id/F21687DACB337712FF4E33AAFB33FF73" lastPageId="8" lastPageNumber="442" pageId="7" pageNumber="442">
<subSubSection id="32A56547CB33771DFF4E33AAFC1EFB2A" box="[151,908,1158,1185]" pageId="7" pageNumber="442" type="nomenclature">
<paragraph id="7A0036CCCB33771DFF4E33AAFC1EFB2A" blockId="7.[151,908,1158,1185]" box="[151,908,1158,1185]" pageId="7" pageNumber="442">
<heading id="214881A0CB33771DFF4E33AAFC1EFB2A" bold="true" box="[151,908,1158,1185]" fontSize="11" level="1" pageId="7" pageNumber="442" reason="1">
<emphasis id="48CBEADECB33771DFF4E33AAFC1EFB2A" bold="true" box="[151,908,1158,1185]" pageId="7" pageNumber="442">
Genus
<taxonomicName id="BDBF4D4FCB33771DFF3733ABFC1EFB2A" authority="Nabozhenko, Kirejtshuk et Merkl, 2016" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[238,908,1158,1185]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="48CBEADECB33771DFF3733ABFE01FB2B" bold="true" box="[238,403,1159,1185]" italics="true" pageId="7" pageNumber="442">Yantaroxenos</emphasis>
<bibRefCitation id="1E2E4B3DCB33771DFE4233ABFC1EFB2A" author="Nabozhenko, M. &amp; Kirejtshuk, A. &amp; Merkl, O." box="[411,908,1158,1185]" pageId="7" pageNumber="442" pagination="563 - 566" refId="ref6010" refString="Nabozhenko, M., Kirejtshuk, A. &amp; Merkl, O. (2016) Yantaroxenos colydioides gen. et sp. nov. (Tenebrionidae: Lagriinae) from Baltic amber. Annales Zoologici, Warszawa, 66 (4), 563 - 566. https: // doi. org / 10.3161 / 00034541 ANZ 2016.66.4.008" type="journal article" year="2016">Nabozhenko, Kirejtshuk et Merkl, 2016</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="32A56547CB33771DFF4E33E0FC51FB69" box="[151,963,1227,1252]" pageId="7" pageNumber="442" type="materials_examined">
<paragraph id="7A0036CCCB33771DFF4E33E0FC51FB69" blockId="7.[151,963,1227,1252]" box="[151,963,1227,1252]" pageId="7" pageNumber="442">
<typeStatus id="A504886ECB33771DFF4E33E0FF5BFB6E" box="[151,201,1228,1252]" pageId="7" pageNumber="442">Type</typeStatus>
species:
<taxonomicName id="BDBF4D4FCB33771DFEFF33E0FC51FB69" authority="Nabozhenko, Kirejtshuk et Merkl, 2016" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[294,963,1227,1251]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="species" species="colydioides">
<emphasis id="48CBEADECB33771DFEFF33E0FDBAFB69" box="[294,552,1227,1251]" italics="true" pageId="7" pageNumber="442">Yantaroxenos colydioides</emphasis>
<bibRefCitation id="1E2E4B3DCB33771DFDF733E7FC51FB69" author="Nabozhenko, M. &amp; Kirejtshuk, A. &amp; Merkl, O." box="[558,963,1227,1251]" pageId="7" pageNumber="442" pagination="563 - 566" refId="ref6010" refString="Nabozhenko, M., Kirejtshuk, A. &amp; Merkl, O. (2016) Yantaroxenos colydioides gen. et sp. nov. (Tenebrionidae: Lagriinae) from Baltic amber. Annales Zoologici, Warszawa, 66 (4), 563 - 566. https: // doi. org / 10.3161 / 00034541 ANZ 2016.66.4.008" type="journal article" year="2016">Nabozhenko, Kirejtshuk et Merkl, 2016</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="32A56547CB337712FF4E323DFB33FF73" lastPageId="8" lastPageNumber="443" pageId="7" pageNumber="442" type="discussion">
<paragraph id="7A0036CCCB33771DFF4E323DFC19FA55" blockId="7.[151,1437,1297,2008]" pageId="7" pageNumber="442">
<emphasis id="48CBEADECB33771DFF4E323DFE85FAA1" bold="true" box="[151,279,1297,1323]" pageId="7" pageNumber="442">Comment:</emphasis>
The extinct monotypic genus
<taxonomicName id="BDBF4D4FCB33771DFDA7323EFB7CFAA1" authority="Nabozhenko, Kirejtshuk et Merkl, 2016" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[638,1262,1297,1324]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="48CBEADECB33771DFDA7323EFC86FAA1" box="[638,788,1298,1323]" italics="true" pageId="7" pageNumber="442">Yantaroxenos</emphasis>
<bibRefCitation id="1E2E4B3DCB33771DFCC6323DFB7CFAA1" author="Nabozhenko, M. &amp; Kirejtshuk, A. &amp; Merkl, O." box="[799,1262,1297,1324]" pageId="7" pageNumber="442" pagination="563 - 566" refId="ref6010" refString="Nabozhenko, M., Kirejtshuk, A. &amp; Merkl, O. (2016) Yantaroxenos colydioides gen. et sp. nov. (Tenebrionidae: Lagriinae) from Baltic amber. Annales Zoologici, Warszawa, 66 (4), 563 - 566. https: // doi. org / 10.3161 / 00034541 ANZ 2016.66.4.008" type="journal article" year="2016">Nabozhenko, Kirejtshuk et Merkl, 2016</bibRefCitation>
</taxonomicName>
was originally described from Baltic amber as a presumed representative of the tribe
<taxonomicName id="BDBF4D4FCB33771DFC1E3219FBB5FAC5" box="[967,1063,1333,1359]" class="Insecta" family="Lagriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="tribe" tribe="Belopini">Belopini</taxonomicName>
(
<taxonomicName id="BDBF4D4FCB33771DFBE03219FB4DFAC5" box="[1081,1247,1333,1359]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
:
<taxonomicName id="BDBF4D4FCB33771DFB353219FACFFAC5" authorityName="Latreille" authorityYear="1825" box="[1260,1373,1333,1359]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="subFamily" subFamily="Lagriinae">Lagriinae</taxonomicName>
). The
<typeStatus id="A504886ECB33771DFF4E3275FF6BFAF9" box="[151,249,1369,1395]" pageId="7" pageNumber="442">holotype</typeStatus>
was studied by optical microscopy. Since some important characters were not observed, the original description states: “parts of the integument of the specimen with milky cover,” “mesoventrite, meso- and metepisterna and metepimera not visible,” and “
<taxonomicName id="BDBF4D4FCB33771DFDF0328EFD2DFA31" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[553,703,1442,1467]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="48CBEADECB33771DFDF0328EFD2DFA31" box="[553,703,1442,1467]" italics="true" pageId="7" pageNumber="442">Yantaroxenos</emphasis>
</taxonomicName>
has unclear combination of tribal characters because diagnostic structures of the head are poorly visible” (
<bibRefCitation id="1E2E4B3DCB33771DFDB432E9FC12FA55" author="Nabozhenko, M. &amp; Kirejtshuk, A. &amp; Merkl, O." box="[621,896,1477,1503]" pageId="7" pageNumber="442" pagination="563 - 566" refId="ref6010" refString="Nabozhenko, M., Kirejtshuk, A. &amp; Merkl, O. (2016) Yantaroxenos colydioides gen. et sp. nov. (Tenebrionidae: Lagriinae) from Baltic amber. Annales Zoologici, Warszawa, 66 (4), 563 - 566. https: // doi. org / 10.3161 / 00034541 ANZ 2016.66.4.008" type="journal article" year="2016">
Nabozhenko
<emphasis id="48CBEADECB33771DFCDD32EAFCAEFA55" box="[772,828,1477,1503]" italics="true" pageId="7" pageNumber="442">et al.</emphasis>
2016
</bibRefCitation>
).
</paragraph>
<paragraph id="7A0036CCCB33771DFF1E32C5FC72F8A9" blockId="7.[151,1437,1297,2008]" pageId="7" pageNumber="442">
The familial placement of
<taxonomicName id="BDBF4D4FCB33771DFDDA32C6FD0BF989" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[515,665,1514,1539]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="48CBEADECB33771DFDDA32C6FD0BF989" box="[515,665,1514,1539]" italics="true" pageId="7" pageNumber="442">Yantaroxenos</emphasis>
</taxonomicName>
(with two logical alternatives,
<taxonomicName id="BDBF4D4FCB33771DFBD732C5FB1CF989" authorityName="Solier" authorityYear="1834" box="[1038,1166,1513,1539]" class="Insecta" family="Zopheridae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="family">Zopheridae</taxonomicName>
or
<taxonomicName id="BDBF4D4FCB33771DFB6532C5FACCF989" box="[1212,1374,1513,1539]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
) was initially the cause of much concern during the study of
<bibRefCitation id="1E2E4B3DCB33771DFCCA3121FBA8F9AD" author="Nabozhenko, M. &amp; Kirejtshuk, A. &amp; Merkl, O." box="[787,1082,1549,1575]" pageId="7" pageNumber="442" pagination="563 - 566" refId="ref6010" refString="Nabozhenko, M., Kirejtshuk, A. &amp; Merkl, O. (2016) Yantaroxenos colydioides gen. et sp. nov. (Tenebrionidae: Lagriinae) from Baltic amber. Annales Zoologici, Warszawa, 66 (4), 563 - 566. https: // doi. org / 10.3161 / 00034541 ANZ 2016.66.4.008" type="journal article" year="2016">
Nabozhenko
<emphasis id="48CBEADECB33771DFC733122FC76F9AD" box="[938,996,1549,1575]" italics="true" pageId="7" pageNumber="442">et al.</emphasis>
(2016)
</bibRefCitation>
. Finally, placement in the tribe
<taxonomicName id="BDBF4D4FCB33771DFF4E311DFF65F9C1" box="[151,247,1585,1611]" class="Insecta" family="Lagriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="tribe" tribe="Belopini">Belopini</taxonomicName>
was considered the most appropriate for this extinct Baltic amber beetle. One important character of
<taxonomicName id="BDBF4D4FCB33771DFF4E317AFEBFF9E5" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[151,301,1622,1647]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="48CBEADECB33771DFF4E317AFEBFF9E5" box="[151,301,1622,1647]" italics="true" pageId="7" pageNumber="442">Yantaroxenos</emphasis>
</taxonomicName>
, however, was regrettably misinterpreted: In reality, all abdominal ventrites in
<taxonomicName id="BDBF4D4FCB33771DFB01317AFAFCF9E5" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[1240,1390,1622,1647]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="48CBEADECB33771DFB01317AFAFCF9E5" box="[1240,1390,1622,1647]" italics="true" pageId="7" pageNumber="442">Yantaroxenos</emphasis>
</taxonomicName>
are unfused and apparently connected by narrow membranes (
<figureCitation id="E2842A49CB33771DFCF83155FCF7F919" box="[801,869,1657,1683]" captionStart="FIGURES 5–7" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[158,1430,181,1919]" captionTargetId="figure-25@6.[158,1430,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–7. Representatives of the tribe Zopheromimini trib. nov. (microphotographs): 5—head of Zopheromimus auriborussiensis; 6—abdomen of Z. auriborussiensis; 7—abdomen of Yantaroxenos colydioides. Abbreviations: mem— membranes between non-connate ventrites; pag—preapical groove on abdominal ventrite 5." figureDoi="http://doi.org/10.5281/zenodo.8005231" httpUri="https://zenodo.org/record/8005231/files/figure.png" pageId="7" pageNumber="442">Fig. 7</figureCitation>
), as is the case for
<taxonomicName id="BDBF4D4FCB33771DFBEF3155FA0EF919" authority="Alekseev &amp; Nabozhenko, 2023" authorityName="Alekseev &amp; Nabozhenko" authorityYear="2023" box="[1078,1436,1657,1683]" class="Insecta" family="Zopheridae" genus="Zopheromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="species" species="auriborussiensis" status="sp. nov.">
<emphasis id="48CBEADECB33771DFBEF3155FA0EF919" box="[1078,1436,1657,1683]" italics="true" pageId="7" pageNumber="442">Zopheromimus auriborussiensis</emphasis>
</taxonomicName>
<emphasis id="48CBEADECB33771DFF4E31B1FF61F93D" bold="true" box="[151,243,1693,1719]" pageId="7" pageNumber="442">
<taxonomicNameLabel id="53F857A5CB33771DFF4E31B1FF61F93D" box="[151,243,1693,1719]" pageId="7" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="E2842A49CB33771DFEDD31B1FED9F932" box="[260,331,1693,1720]" captionStart="FIGURES 5–7" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[158,1430,181,1919]" captionTargetId="figure-25@6.[158,1430,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–7. Representatives of the tribe Zopheromimini trib. nov. (microphotographs): 5—head of Zopheromimus auriborussiensis; 6—abdomen of Z. auriborussiensis; 7—abdomen of Yantaroxenos colydioides. Abbreviations: mem— membranes between non-connate ventrites; pag—preapical groove on abdominal ventrite 5." figureDoi="http://doi.org/10.5281/zenodo.8005231" httpUri="https://zenodo.org/record/8005231/files/figure.png" pageId="7" pageNumber="442">Fig. 5</figureCitation>
). Darkling beetles have connate abdominal ventrites 1 and 2. Ventrites 3–5 can be hinged in the middle or laterally, and membranes between ventrites 3–5 are visible in almost all representatives of tenebrionoid branch having defensive glands (
<bibRefCitation id="1E2E4B3DCB33771DFDCE31C9FCF4F975" author="Matthews, E. G. &amp; Bouchard, P." box="[535,870,1765,1791]" pageId="7" pageNumber="442" refId="ref5755" refString="Matthews, E. G. &amp; Bouchard, P. (2008) Tenebrionid Beetles of Australia: Descriptions of Tribes, Keys to Genera, Catalogue of Species. Australian Biological Resources Study, Canberra, 410 pp." type="book" year="2008">Matthews &amp; Bouchard 2008</bibRefCitation>
,
<bibRefCitation id="1E2E4B3DCB33771DFCAC31C9FBE0F975" author="Matthews, E. G. &amp; Lawrence, J. F. &amp; Bouchard, P. &amp; Steiner, W. E. &amp; Slipinski, S. A." box="[885,1138,1765,1791]" pageId="7" pageNumber="442" pagination="1802" refId="ref5796" refString="Matthews, E. G., Lawrence, J. F., Bouchard, P., Steiner, W. E. &amp; Slipinski, S. A. (2010) 11.14. Tenebrionidae Latreille, 1802. In: Leschen, R. A. B, Beutel, R. G. &amp; Lawrence, J. F. (Eds.), Handbook of zoology. Arthropoda: Insecta. Part 38. Coleoptera, beetles. Vol. 2. Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). Walter de Gruyter, Berlin, pp. 574 - 659. https: // doi. org / 10.1515 / 9783110911213.574" type="book chapter" year="2010">
Matthews
<emphasis id="48CBEADECB33771DFC3631CAFBB9F975" box="[1007,1067,1765,1791]" italics="true" pageId="7" pageNumber="442">et al.</emphasis>
2010
</bibRefCitation>
). By lacking any connate ventrites,
<taxonomicName id="BDBF4D4FCB33771DFEDF3026FE0EF8A9" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[262,412,1802,1827]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="48CBEADECB33771DFEDF3026FE0EF8A9" box="[262,412,1802,1827]" italics="true" pageId="7" pageNumber="442">Yantaroxenos</emphasis>
</taxonomicName>
should be excluded from the family
<taxonomicName id="BDBF4D4FCB33771DFCE13025FC4EF8A9" box="[824,988,1801,1827]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
.
</paragraph>
<paragraph id="7A0036CCCB337712FF1E3001FB33FF73" blockId="7.[151,1437,1297,2008]" lastBlockId="8.[151,1437,151,249]" lastPageId="8" lastPageNumber="443" pageId="7" pageNumber="442">
The new species
<taxonomicName id="BDBF4D4FCB33771DFE5C3001FD7EF8CD" authority="Alekseev &amp; Nabozhenko, 2023" authorityName="Alekseev &amp; Nabozhenko" authorityYear="2023" box="[389,748,1837,1863]" class="Insecta" family="Zopheridae" genus="Zopheromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="species" species="auriborussiensis" status="sp. nov.">
<emphasis id="48CBEADECB33771DFE5C3001FD7EF8CD" box="[389,748,1837,1863]" italics="true" pageId="7" pageNumber="442">Zopheromimus auriborussiensis</emphasis>
</taxonomicName>
<emphasis id="48CBEADECB33771DFD2B3001FCD9F8CD" bold="true" box="[754,843,1837,1863]" pageId="7" pageNumber="442">
<taxonomicNameLabel id="53F857A5CB33771DFD2B3001FCD9F8CD" box="[754,843,1837,1863]" pageId="7" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
allowed examination of the ventral side of the thorax and head in this group of beetles. Our findings supported the assignment of
<taxonomicName id="BDBF4D4FCB33771DFC26307DFB34F8E1" authority="Alekseev &amp; Nabozhenko, 2023" authorityName="Alekseev &amp; Nabozhenko" authorityYear="2023" box="[1023,1190,1873,1899]" class="Insecta" family="Zopheridae" genus="Zopheromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="48CBEADECB33771DFC26307DFB34F8E1" box="[1023,1190,1873,1899]" italics="true" pageId="7" pageNumber="442">Zopheromimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53F857A5CB33771DFB76307DFA89F8E6" box="[1199,1307,1873,1900]" pageId="7" pageNumber="442" rank="genus">
<emphasis id="48CBEADECB33771DFB76307DFA87F8E1" bold="true" box="[1199,1301,1873,1899]" pageId="7" pageNumber="442">gen. nov</emphasis>
.
</taxonomicNameLabel>
to the new tribe within the subfamily
<taxonomicName id="BDBF4D4FCB33771DFE113059FDD9F805" authorityName="Solier" authorityYear="1834" box="[456,587,1909,1935]" class="Insecta" family="Zopheridae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="subFamily" subFamily="Zopherinae">Zopherinae</taxonomicName>
.
<taxonomicName id="BDBF4D4FCB33771DFD8E305AFCE5F805" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[599,887,1909,1935]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="species" species="colydioides">
<emphasis id="48CBEADECB33771DFD8E305AFCE5F805" box="[599,887,1909,1935]" italics="true" pageId="7" pageNumber="442">Yantaroxenos colydioides</emphasis>
</taxonomicName>
shares with
<taxonomicName id="BDBF4D4FCB33771DFBD03059FAE0F805" authority="Alekseev &amp; Nabozhenko, 2023" authorityName="Alekseev &amp; Nabozhenko" authorityYear="2023" box="[1033,1394,1909,1935]" class="Insecta" family="Zopheridae" genus="Zopheromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="442" phylum="Arthropoda" rank="species" species="auriborussiensis" status="sp. nov.">
<emphasis id="48CBEADECB33771DFBD03059FAE0F805" box="[1033,1394,1909,1935]" italics="true" pageId="7" pageNumber="442">Zopheromimus auriborussiensis</emphasis>
</taxonomicName>
<emphasis id="48CBEADECB33771DFAA23059FF5AF839" bold="true" pageId="7" pageNumber="442">
<taxonomicNameLabel id="53F857A5CB33771DFAA23059FF5AF839" pageId="7" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
all of the external characters that are interpreted here to have tribal-level significance (including the paired preapical grooves on abdominal ventrite 5, which was not mentioned in original description, but is present, see
<figureCitation id="E2842A49CB337712FAA93091FF35FF3B" captionStart="FIGURES 5–7" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[158,1430,181,1919]" captionTargetId="figure-25@6.[158,1430,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–7. Representatives of the tribe Zopheromimini trib. nov. (microphotographs): 5—head of Zopheromimus auriborussiensis; 6—abdomen of Z. auriborussiensis; 7—abdomen of Yantaroxenos colydioides. Abbreviations: mem— membranes between non-connate ventrites; pag—preapical groove on abdominal ventrite 5." figureDoi="http://doi.org/10.5281/zenodo.8005231" httpUri="https://zenodo.org/record/8005231/files/figure.png" lastPageId="8" lastPageNumber="443" pageId="7" pageNumber="442">Fig. 7</figureCitation>
). The main differences between these two genera are in the pronotal shape and number of antennomeres. The features described above led us to conclude that the extinct genus
<taxonomicName id="BDBF4D4FCB3C7712FCAC3797FB99FF5E" authorityName="Nabozhenko, Kirejtshuk et Merkl" authorityYear="2016" box="[885,1035,187,212]" class="Insecta" family="Lagriidae" genus="Yantaroxenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="genus">
<emphasis id="48CBEADECB3C7712FCAC3797FB99FF5E" box="[885,1035,187,212]" italics="true" pageId="8" pageNumber="443">Yantaroxenos</emphasis>
</taxonomicName>
should be transferred from the tribe
<taxonomicName id="BDBF4D4FCB3C7712FF4E37F3FF65FF73" box="[151,247,223,249]" class="Insecta" family="Lagriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="tribe" tribe="Belopini">Belopini</taxonomicName>
(
<taxonomicName id="BDBF4D4FCB3C7712FEDF37F3FE3EFF73" box="[262,428,223,249]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
:
<taxonomicName id="BDBF4D4FCB3C7712FE6F37F3FDB1FF73" authorityName="Latreille" authorityYear="1825" box="[438,547,223,249]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="subFamily" subFamily="Lagriinae">Lagriinae</taxonomicName>
) to
<taxonomicName id="BDBF4D4FCB3C7712FD9637F3FD6DFF73" authority="Alekseev &amp; Nabozhenko, 2023" authorityName="Alekseev &amp; Nabozhenko" authorityYear="2023" box="[591,767,223,249]" class="Insecta" family="Zopheridae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Zopheromimini">Zopheromimini</taxonomicName>
<emphasis id="48CBEADECB3C7712FCDF37F2FCE0FF73" bold="true" box="[774,882,222,249]" pageId="8" pageNumber="443">
<taxonomicNameLabel id="53F857A5CB3C7712FCDF37F2FCE0FF73" box="[774,882,222,249]" pageId="8" pageNumber="443" rank="tribe">trib. nov.</taxonomicNameLabel>
</emphasis>
(
<taxonomicName id="BDBF4D4FCB3C7712FC5937F3FB94FF73" authorityName="Solier" authorityYear="1834" box="[896,1030,223,249]" class="Insecta" family="Zopheridae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="family">Zopheridae</taxonomicName>
:
<taxonomicName id="BDBF4D4FCB3C7712FBC937F3FB07FF73" authorityName="Solier" authorityYear="1834" box="[1040,1173,223,249]" class="Insecta" family="Zopheridae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="subFamily" subFamily="Zopherinae">Zopherinae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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